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Hirnantian glaciation - Wikipedia

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class="vector-toc-list"> <li id="toc-Pre-Hirnantian_glaciations" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Pre-Hirnantian_glaciations"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Pre-Hirnantian glaciations</span> </div> </a> <ul id="toc-Pre-Hirnantian_glaciations-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Hirnantian_glaciation" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Hirnantian_glaciation"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2</span> <span>Hirnantian glaciation</span> </div> </a> <ul id="toc-Hirnantian_glaciation-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Silurian_glaciations" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Silurian_glaciations"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.3</span> <span>Silurian glaciations</span> </div> </a> <ul id="toc-Silurian_glaciations-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Evidence" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Evidence"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Evidence</span> </div> </a> <button aria-controls="toc-Evidence-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Evidence subsection</span> </button> <ul id="toc-Evidence-sublist" class="vector-toc-list"> <li id="toc-Lithologic" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Lithologic"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Lithologic</span> </div> </a> <ul id="toc-Lithologic-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Possible_causes" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Possible_causes"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Possible causes</span> </div> </a> <button aria-controls="toc-Possible_causes-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Possible causes subsection</span> </button> <ul id="toc-Possible_causes-sublist" class="vector-toc-list"> <li id="toc-CO2_depletion" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#CO2_depletion"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1</span> <span>CO<sub>2</sub> depletion</span> </div> </a> <ul id="toc-CO2_depletion-sublist" class="vector-toc-list"> <li id="toc-Silicate_weathering" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Silicate_weathering"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1.1</span> <span>Silicate weathering</span> </div> </a> <ul id="toc-Silicate_weathering-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Organic_carbon_burial" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Organic_carbon_burial"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1.2</span> <span>Organic carbon burial</span> </div> </a> <ul id="toc-Organic_carbon_burial-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Gamma-ray_burst" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Gamma-ray_burst"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1.3</span> <span>Gamma-ray burst</span> </div> </a> <ul id="toc-Gamma-ray_burst-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Asteroid_impact" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Asteroid_impact"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.2</span> <span>Asteroid impact</span> </div> </a> <ul id="toc-Asteroid_impact-sublist" class="vector-toc-list"> <li id="toc-Ordovician_meteor_event" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Ordovician_meteor_event"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.2.1</span> <span>Ordovician meteor event</span> </div> </a> <ul id="toc-Ordovician_meteor_event-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Deniliquin_impact_structure" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Deniliquin_impact_structure"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.2.2</span> <span>Deniliquin impact structure</span> </div> </a> <ul id="toc-Deniliquin_impact_structure-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Debris_ring" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Debris_ring"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.3</span> <span>Debris ring</span> </div> </a> <ul id="toc-Debris_ring-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Volcanic_aerosols" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Volcanic_aerosols"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.4</span> <span>Volcanic aerosols</span> </div> </a> <ul id="toc-Volcanic_aerosols-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Sea_level_change" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Sea_level_change"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.5</span> <span>Sea level change</span> </div> </a> <ul id="toc-Sea_level_change-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Palaeogeography" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Palaeogeography"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.6</span> <span>Palaeogeography</span> </div> </a> <ul id="toc-Palaeogeography-sublist" class="vector-toc-list"> <li id="toc-Poleward_ocean_heat_transport" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Poleward_ocean_heat_transport"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.6.1</span> <span>Poleward ocean heat transport</span> </div> </a> <ul id="toc-Poleward_ocean_heat_transport-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Orbital_parameters" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Orbital_parameters"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.7</span> <span>Orbital parameters</span> </div> </a> <ul id="toc-Orbital_parameters-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-End_of_the_event" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#End_of_the_event"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>End of the event</span> </div> </a> <button aria-controls="toc-End_of_the_event-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle End of the event subsection</span> </button> <ul id="toc-End_of_the_event-sublist" class="vector-toc-list"> <li id="toc-Ice_collapse" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Ice_collapse"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.1</span> <span>Ice collapse</span> </div> </a> <ul id="toc-Ice_collapse-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-CO2" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#CO2"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.2</span> <span>CO<sub>2</sub></span> </div> </a> <ul id="toc-CO2-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Significance" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Significance"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Significance</span> </div> </a> <ul id="toc-Significance-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> 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class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/Rhewlifiant_Andea-Saharaidd" title="Rhewlifiant Andea-Saharaidd – Welsh" lang="cy" hreflang="cy" data-title="Rhewlifiant Andea-Saharaidd" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Glaciaci%C3%B3n_Andino-Sahariana" title="Glaciación Andino-Sahariana – Spanish" lang="es" hreflang="es" data-title="Glaciación Andino-Sahariana" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Glaciation_de_l%27And%C3%A9en-Saharien" title="Glaciation de l&#039;Andéen-Saharien – French" lang="fr" 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title="Dinosaur">Dinosaurs</a></div></div></div><div class="notheme" style="font-size:100%;background-color:#ffcc00;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:0.000em;height:2.203em;left:8.580em;width:4.420em"><div style="font-size:100%;position:relative;top:0em;left:0em"><div class="notheme" style="position:relative;text-align:center;vertical-align:middle;display:block;bottom:-1.101em;transform:translateY(-50%);z-index:5">&#160;</div></div></div><div class="notheme" style="font-size:100%;background-color:#ffcc00;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:0.606em;height:1.597em;left:8.450em;width:4.290em"><div style="font-size:100%;position:relative;top:0.15em;left:0em"><div class="notheme" style="position:relative;text-align:center;font-size:90%;vertical-align:middle;display:block;bottom:-0.798em;transform:translateY(-50%);z-index:5"><a href="/wiki/Mammal" title="Mammal">Mammals</a></div></div></div><div class="notheme" style="font-size:100%;background-color:#e6b880;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:0.000em;height:1.333em;left:5.980em;width:2.600em"><div style="font-size:100%;position:relative;top:-0.225em;left:0em"><div class="notheme" style="position:relative;text-align:center;font-size:90%;vertical-align:middle;display:block;bottom:-0.666em;transform:translateY(-50%);z-index:5"><a href="/wiki/Bird" title="Bird">Birds</a></div></div></div><div class="notheme" style="font-size:100%;background-color:#ffbbbb;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:0.000em;height:0.606em;left:8.710em;width:4.030em"><div style="font-size:100%;position:relative;top:0.1em;left:0em"><div class="notheme" style="position:relative;text-align:center;font-size:90%;vertical-align:middle;display:block;bottom:-0.303em;transform:translateY(-50%);z-index:5"><a href="/wiki/Primate" title="Primate">Primates</a></div></div></div><div class="notheme" style="font-size:100%;background-color:#993c00;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:44.053em;height:5.947em;left:0.000em;width:0.910em"><div style="font-size:100%;position:relative;top:0em;left:-0.05em"><div class="notheme" style="position:relative;text-align:center;font-size:100%;vertical-align:middle;display:block;bottom:-2.974em;transform:translateY(-50%);z-index:5"><span style="display:block; text-align:center; line-height: 0.80em;"><a href="/wiki/Hadean" title="Hadean"><span style="color:#ffffff;">H<br />a<br />d<br />e<br />a<br />n</span></a></span></div></div></div><div class="notheme" style="font-size:100%;background-color:#936a00;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:27.533em;height:16.520em;left:0.000em;width:0.910em"><div style="font-size:100%;position:relative;top:-1.2em;left:-0.05em"><div class="notheme" style="position:relative;text-align:center;font-size:100%;vertical-align:middle;display:block;bottom:-8.260em;transform:translateY(-50%);z-index:5"><span style="display:block; text-align:center; line-height: 0.85em;"><a href="/wiki/Archean" title="Archean"><span style="color:#ffffff;"><br /><br /><br />A<br />r<br />c<br />h<br />e<br />a<br />n</span></a></span></div></div></div><div class="notheme" style="font-size:100%;background-color:#f1c309;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:5.958em;height:21.575em;left:0.000em;width:0.910em"><div style="font-size:100%;position:relative;top:0em;left:0em"><div class="notheme" style="position:relative;text-align:center;font-size:100%;vertical-align:middle;display:block;bottom:-10.787em;transform:translateY(-50%);z-index:5"><span style="display:block; text-align:center; line-height: 0.85em;"><a href="/wiki/Proterozoic" title="Proterozoic"><br /><br /><br /><br /><br /><br /><br />P<br />r<br />o<br />t<br />e<br />r<br />o<br />z<br />o<br />i<br />c</a></span></div></div></div><div class="notheme" style="font-size:100%;background-color:#c1e0c1;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:0.000em;height:5.958em;left:0.000em;width:0.910em"><div style="font-size:100%;position:relative;top:2em;left:0em"><div class="notheme" style="position:relative;text-align:center;font-size:100%;vertical-align:middle;display:block;bottom:-2.979em;transform:translateY(-50%);z-index:5"><span style="display:block; text-align:center; line-height: 0.85em;"><a href="/wiki/Phanerozoic" title="Phanerozoic">P<br />h<br />a<br />n<br />e<br />r<br />o<br />z<br />o<br />i<br />c</a></span></div></div></div><div class="notheme" style="font-size:100%;background-color:#ccbbaa;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:0.000em;height:8.260em;left:12.714em;width:0.286em"><div style="font-size:100%;position:relative;top:0em;left:0em"><div class="notheme" style="position:relative;text-align:center;vertical-align:middle;display:block;bottom:-4.130em;transform:translateY(-50%);z-index:5">&#160;</div></div></div><div class="notheme" style="font-size:100%;background-color:#ccbbaa;border-style:none none;position:absolute;text-align:center;margin:0;padding:0;top:0.000em;height:2.203em;left:12.714em;width:0.286em"><div style="font-size:100%;position:relative;top:0em;left:0em"><div class="notheme" style="position:relative;text-align:center;vertical-align:middle;display:block;bottom:-1.101em;transform:translateY(-50%);z-index:5">&#160;</div></div></div></td><td id="Annotations" style="padding:0;margin:0.7em 0 0.7em 0;float:left;position:relative;width:11.05em;height:50em"><table role="presentation" style="position:absolute;z-index:15;top:50.000em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Age_of_Earth" title="Age of Earth">Earth formed</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:48.568em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0.2em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Origin_of_water_on_Earth" title="Origin of water on Earth">Earliest water</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:47.907em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Last_universal_common_ancestor" title="Last universal common ancestor">LUCA</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:44.053em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Earliest_known_life_forms" title="Earliest known life forms">Earliest fossils</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:42.952em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Late_Heavy_Bombardment" title="Late Heavy Bombardment">LHB meteorites</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:38.546em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Geological_history_of_oxygen" title="Geological history of oxygen">Earliest oxygen</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:31.278em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Pongola_glaciation" title="Pongola glaciation">Pongola glaciation</a>*</div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:25.330em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0.2em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Great_Oxidation_Event" title="Great Oxidation Event">Atmospheric oxygen</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:24.780em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:-0.2em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Huronian_glaciation" title="Huronian glaciation">Huronian glaciation</a>*</div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:22.026em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Evolution_of_sexual_reproduction#Origin_of_sexual_reproduction" title="Evolution of sexual reproduction">Sexual reproduction</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:17.621em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Francevillian_biota" title="Francevillian biota">Earliest multicellular life</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:16.520em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Evolution_of_fungi" title="Evolution of fungi">Earliest fungi</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:9.361em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0.25em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Evolutionary_history_of_plants" title="Evolutionary history of plants">Earliest plants</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:8.260em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0.4em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Caveasphaera" title="Caveasphaera">Earliest animals</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:7.489em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0.2em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Cryogenian" title="Cryogenian">Cryogenian ice age</a>*</div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:6.993em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:-0.2em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Ediacaran_biota" title="Ediacaran biota">Ediacaran biota</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:5.958em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:-0.15em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Cambrian_explosion" title="Cambrian explosion">Cambrian explosion</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:4.956em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:-0.1em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a class="mw-selflink selflink">Hirnantian glaciation</a>*</div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:4.075em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:-0.1em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Tetrapod" title="Tetrapod">Earliest tetrapods</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:3.304em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:-0.2em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Late_Paleozoic_icehouse" title="Late Paleozoic icehouse">Karoo ice age</a>*</div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:0.275em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:0.3em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Ape" title="Ape">Earliest apes</a> / <a href="/wiki/Human" title="Human">humans</a></div></div></td></tr></tbody></table><table role="presentation" style="position:absolute;z-index:15;top:0.018em;left:0em;transform:translateY(-50%);padding:0;margin:0;font-size:100%"><tbody><tr><td style="padding:0;text-align:left;vertical-align:middle"><span style="color:var( --color-base, #000)">&#8592;</span></td><td style="padding:0;text-align:left;vertical-align:middle"><div class="annot-nudge" style="font-size:100%;float:left;position:relative;top:-0.4em"><div style="z-index:10;font-size:90%;color:var( --color-base, #000);display:block;line-height:105%;bottom:0"><a href="/wiki/Quaternary_glaciation" title="Quaternary glaciation">Quaternary ice age</a>*</div></div></td></tr></tbody></table></td></tr><tr><td id="Caption" colspan="3" style="padding:0;margin:0 0.2em 0.7em 0.2em"><div id="Caption" class="toccolours" style="margin:0 0.5em;border-style:none;clear:both;text-align:center;width:26.8em"><div style="float:left;font-size:85%;">(<a href="/wiki/Myr" class="mw-redirect" title="Myr">million years ago</a>)</div><div style="float:right;font-size:95%;">*<i><a href="/wiki/Timeline_of_glaciation" title="Timeline of glaciation">Ice Ages</a></i></div></div></td></tr></tbody></table> <p>The <b>Hirnantian <a href="/wiki/Glacial_period" title="Glacial period">glaciation</a></b>, also known as the <b>Andean-Saharan glaciation</b>, <b>Early Paleozoic Ice Age</b> (<b>EPIA</b>),<sup id="cite_ref-EPIA_1-0" class="reference"><a href="#cite_note-EPIA-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> the <b>Early Paleozoic Icehouse</b>,<sup id="cite_ref-GlacioeustasyBlackShalesEarlyPalaeozoicIcehouse_2-0" class="reference"><a href="#cite_note-GlacioeustasyBlackShalesEarlyPalaeozoicIcehouse-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> the <b>Late Ordovician glaciation</b>, or the <b>end-Ordovician glaciation</b>, occurred during the <a href="/wiki/Paleozoic" title="Paleozoic">Paleozoic</a> from approximately 460 <a href="/wiki/Annum" class="mw-redirect" title="Annum">Ma</a> to around 420 Ma, during the <a href="/wiki/Late_Ordovician" title="Late Ordovician">Late Ordovician</a> and the <a href="/wiki/Silurian" title="Silurian">Silurian</a> period. The major glaciation during this period was formerly thought only to consist of the Hirnantian glaciation itself but has now been recognized as a longer, more gradual event,<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Munnecke2010_5-0" class="reference"><a href="#cite_note-Munnecke2010-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> which began as early as the <a href="/wiki/Darriwilian" title="Darriwilian">Darriwilian</a>,<sup id="cite_ref-EPIA_1-1" class="reference"><a href="#cite_note-EPIA-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> and possibly even the <a href="/wiki/Floian" title="Floian">Floian</a>.<sup id="cite_ref-MayaElrick_6-0" class="reference"><a href="#cite_note-MayaElrick-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> Evidence of this glaciation can be seen in places such as <a href="/wiki/Arabia" class="mw-redirect" title="Arabia">Arabia</a>, <a href="/wiki/North_Africa" title="North Africa">North Africa</a>, <a href="/wiki/South_Africa" title="South Africa">South Africa</a>, <a href="/wiki/Brazil" title="Brazil">Brazil</a>, <a href="/wiki/Peru" title="Peru">Peru</a>, <a href="/wiki/Bolivia" title="Bolivia">Bolivia</a>, <a href="/wiki/Chile" title="Chile">Chile</a>, <a href="/wiki/Argentina" title="Argentina">Argentina</a>, and <a href="/wiki/Wyoming" title="Wyoming">Wyoming</a>.<sup id="cite_ref-Díaz-MartínezGrahn2007_7-0" class="reference"><a href="#cite_note-Díaz-MartínezGrahn2007-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Bighorn_Dolomite,_Wyoming_10-0" class="reference"><a href="#cite_note-Bighorn_Dolomite,_Wyoming-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> More evidence derived from isotopic data is that during the Late Ordovician, tropical ocean temperatures were about 5&#160;°C cooler than present day; this would have been a major factor that aided in the glaciation process.<sup id="cite_ref-Magnitude_and_Duration_11-0" class="reference"><a href="#cite_note-Magnitude_and_Duration-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> </p><p>The Late Ordovician glaciation is widely considered to be the leading cause of the <a href="/wiki/Late_Ordovician_mass_extinction" title="Late Ordovician mass extinction">Late Ordovician mass extinction</a>,<sup id="cite_ref-GlacioeustasyBlackShalesEarlyPalaeozoicIcehouse_2-1" class="reference"><a href="#cite_note-GlacioeustasyBlackShalesEarlyPalaeozoicIcehouse-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-end-Ordovician_glaciation_12-0" class="reference"><a href="#cite_note-end-Ordovician_glaciation-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> and it is the only glacial episode that appears to have coincided with a major mass extinction of nearly 61% of marine life.<sup id="cite_ref-Mass_Extinction_13-0" class="reference"><a href="#cite_note-Mass_Extinction-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> Estimates of peak <a href="/wiki/Ice_sheet" title="Ice sheet">ice sheet</a> <a href="/wiki/Volume" title="Volume">volume</a> range from 50 to 250 million cubic kilometres, and its duration from 35 million to less than 1 million years. At its height during the <a href="/wiki/Hirnantian" title="Hirnantian">Hirnantian</a>, the ice age is believed to have been significantly more extreme than the <a href="/wiki/Last_Glacial_Maximum" title="Last Glacial Maximum">Last Glacial Maximum</a> occurring during the terminal <a href="/wiki/Pleistocene" title="Pleistocene">Pleistocene</a>.<sup id="cite_ref-Magnitude_and_Duration_11-1" class="reference"><a href="#cite_note-Magnitude_and_Duration-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> Glaciation of the <a href="/wiki/Northern_Hemisphere" title="Northern Hemisphere">Northern Hemisphere</a> was minimal because a large amount of the land was in the <a href="/wiki/Southern_Hemisphere" title="Southern Hemisphere">Southern Hemisphere</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Timeline">Timeline</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=1" title="Edit section: Timeline"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Pre-Hirnantian_glaciations">Pre-Hirnantian glaciations</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=2" title="Edit section: Pre-Hirnantian glaciations"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The earliest evidence for possible glaciation comes from Floian conodont apatite oxygen isotope fluctuations, which display a periodicity characteristic of Milankovitch cycles and have been interpreted as reflecting cyclic waxing and waning of polar ice caps.<sup id="cite_ref-MayaElrick_6-1" class="reference"><a href="#cite_note-MayaElrick-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> A speculated glaciation in the middle Darriwilian corresponds to the MDICE positive carbon isotope excursion.<sup id="cite_ref-ClimateChangesPreHirnantian_14-0" class="reference"><a href="#cite_note-ClimateChangesPreHirnantian-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> Sea level changes likely reflective of glacioeustasy are known from this geologic stage, around 467 Ma.<sup id="cite_ref-EPIA_1-2" class="reference"><a href="#cite_note-EPIA-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> However, there are no known Middle Ordovician glacial deposits that would provide direct geological evidence of glaciation.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup> Isotopic evidence from the Sandbian reveals three possible glaciations: an early Sandbian glaciation, a middle Sandbian glaciation, a late Sandbian glaciation.<sup id="cite_ref-ClimateChangesPreHirnantian_14-1" class="reference"><a href="#cite_note-ClimateChangesPreHirnantian-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> Although biostratigraphy dating the glacial deposits in Gondwana has been problematic, there is evidence suggesting the presence of glaciation by the <a href="/wiki/Sandbian" title="Sandbian">Sandbian</a> stage (approximately 451–461 Ma).<sup id="cite_ref-Bighorn_Dolomite,_Wyoming_10-1" class="reference"><a href="#cite_note-Bighorn_Dolomite,_Wyoming-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> Graptolite distribution during the time interval delineated by the <i>Nemacanthus gracilis</i> graptolite biozone indicates a latitudinal extent of the subtropics and tropics similar to that of today, as evidenced by a steep faunal gradient that is uncharacteristic of greenhouse periods, suggesting that Earth was in a mild icehouse state by the start of the Sandbian, around 460 Ma.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">&#91;</span>17<span class="cite-bracket">&#93;</span></a></sup> Many possible short glaciation occurred during the Katian: three very short glaciations during the early Katian, the Rakvere glaciation during the late early Katian, a middle Katian glaciation, the Early Ashgill glaciation of the early late Katian, and a latest Katian glaciation that was followed by a rapid warming event in the <i>Paraorthograptus pacificus</i> graptolite biozone immediately before the Hirnantian glaciation itself.<sup id="cite_ref-ClimateChangesPreHirnantian_14-2" class="reference"><a href="#cite_note-ClimateChangesPreHirnantian-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> Evidence of major changes in bottom water formation, which usually indicates a sudden change in global climate, is known from the Katian.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup> Shifts in isotopic ratios of carbon and neodymium that correspond to graptolite biostratigraphy lend further evidence in favour of the existence of glacioeustatic cycles during the Katian,<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> as do conodont apatite δ<sup>18</sup>O fluctuations from <a href="/wiki/Kentucky" title="Kentucky">Kentucky</a> and <a href="/wiki/Quebec" title="Quebec">Quebec</a> that likely reflect glacioeustatic sea level changes.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">&#91;</span>20<span class="cite-bracket">&#93;</span></a></sup> However, the existence of glacials during the Katian remains controversial.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup> Katian brachiopod and seawater δ<sup>18</sup>O values from Cincinnati Arch indicate ocean temperatures characteristic of a global greenhouse state.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">&#91;</span>23<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Hirnantian_glaciation">Hirnantian glaciation</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=3" title="Edit section: Hirnantian glaciation"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Carbon_13_and_time_scale_during_the_Ordovician.png" class="mw-file-description"><img alt="Ordovician Carbon 13 time scale" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Carbon_13_and_time_scale_during_the_Ordovician.png/220px-Carbon_13_and_time_scale_during_the_Ordovician.png" decoding="async" width="220" height="261" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Carbon_13_and_time_scale_during_the_Ordovician.png/330px-Carbon_13_and_time_scale_during_the_Ordovician.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Carbon_13_and_time_scale_during_the_Ordovician.png/440px-Carbon_13_and_time_scale_during_the_Ordovician.png 2x" data-file-width="881" data-file-height="1045" /></a><figcaption>In this graph the time period that represents the Late Ordovician is at the very top. There is a sharp shift in carbon 13, as well as a sharp decline in sea surface temperatures.<sup id="cite_ref-CO2_exit_glaciation_24-0" class="reference"><a href="#cite_note-CO2_exit_glaciation-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup></figcaption></figure> <p>At the Katian-Hirnantian boundary, a sudden cooling event caused a rapid expansion of glaciers, resulting in one of the most severe glaciations of the Phanerozoic, an extreme cooling event generally believed to be coincident with the first pulse of the Late Ordovician mass extinction.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">&#91;</span>25<span class="cite-bracket">&#93;</span></a></sup> An δ<sup>18</sup>O shift occurs at the start of the Hirnantian; the magnitude of this shift (+2-4‰) was extraordinary.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">&#91;</span>26<span class="cite-bracket">&#93;</span></a></sup> Its direction implies glacial cooling and possibly increases in ice-volume. The observed shifts in the δ<sup>18</sup>O isotopic indicator would require a sea-level fall of 100 meters and a drop of 10&#160;°C in tropical ocean temperatures to have occurred during this glacial episode.<sup id="cite_ref-Bathymetric_and_Isotopic_evidence_27-0" class="reference"><a href="#cite_note-Bathymetric_and_Isotopic_evidence-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> Sedimentological data shows that Late Ordovician ice sheets glacierized the <a href="/wiki/Al_Kufra" class="mw-redirect" title="Al Kufra">Al Kufrah Basin</a>. Ice sheets also probably formed continuous ice cover over North Africa and the Arabian Peninsula. In all areas of North Africa where <a href="/wiki/Llandovery_epoch" class="mw-redirect" title="Llandovery epoch">Early Silurian</a> shale occurs, Late Ordovician glaciogenic deposits occur beneath, likely due to the <a href="/wiki/Anoxic_waters" title="Anoxic waters">anoxia</a> promoted in these basins.<sup id="cite_ref-Evidence_Al_Kufrah_28-0" class="reference"><a href="#cite_note-Evidence_Al_Kufrah-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup> </p><p>At the end of the Hirnantian, an abrupt retreat of glaciers concurrent with the second pulse of the Late Ordovician mass extinction occurred,<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">&#91;</span>29<span class="cite-bracket">&#93;</span></a></sup> after which Earth receded back into a much warmer climate during the Rhuddanian.<sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">&#91;</span>30<span class="cite-bracket">&#93;</span></a></sup> Late Hirnantian warming was marked by a similarly meteoric shift in δ<sup>18</sup>O towards more negative values.<sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">&#91;</span>31<span class="cite-bracket">&#93;</span></a></sup> δ<sup>13</sup>C values likewise fall sharply at the beginning of the <a href="/wiki/Silurian" title="Silurian">Silurian</a>.<sup id="cite_ref-Bathymetric_and_Isotopic_evidence_27-1" class="reference"><a href="#cite_note-Bathymetric_and_Isotopic_evidence-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Silurian_glaciations">Silurian glaciations</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=4" title="Edit section: Silurian glaciations"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Following the relatively warm Rhuddanian, glacial events occurred during the early and latest Aeronian.<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">&#91;</span>32<span class="cite-bracket">&#93;</span></a></sup> A further glaciation occurred from the late Telychian to middle Sheinwoodian.<sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">&#91;</span>33<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">&#91;</span>34<span class="cite-bracket">&#93;</span></a></sup> From the early to late Homerian, Earth was in yet another glacial phase.<sup id="cite_ref-TrotterEtAl2016PPP_35-0" class="reference"><a href="#cite_note-TrotterEtAl2016PPP-35"><span class="cite-bracket">&#91;</span>35<span class="cite-bracket">&#93;</span></a></sup> The last major glaciation of the EPIA occurred during the Ludfordian and was associated with the Lau event.<sup id="cite_ref-LudfordianGlaciation_36-0" class="reference"><a href="#cite_note-LudfordianGlaciation-36"><span class="cite-bracket">&#91;</span>36<span class="cite-bracket">&#93;</span></a></sup> </p><p>During this period, glaciation is known from Arabia, Sahara, West Africa, the south Amazon, and the Andes, and the centre of glaciation is known to have migrated from the Sahara in the Ordovician (450–440 Ma) to South America in the Silurian (440–420 Ma). According to Eyles and Young, "A major glacial episode at c. 440 <a href="/wiki/Myr" class="mw-redirect" title="Myr">Ma</a>, is recorded in Late Ordovician <a href="/wiki/Strata" class="mw-redirect" title="Strata">strata</a> (predominantly <a href="/wiki/Ashgillian" class="mw-redirect" title="Ashgillian">Ashgillian</a>) in West Africa (Tamadjert Formation of the Sahara), in Morocco (<a href="/wiki/Tindouf_Basin" title="Tindouf Basin">Tindouf Basin</a>) and in west-central Saudi Arabia, all areas at polar latitudes at the time. From the Late Ordovician to the <a href="/wiki/Llandovery_epoch" class="mw-redirect" title="Llandovery epoch">Early Silurian</a> the centre of glaciation moved from northern Africa to southwestern South America."<sup id="cite_ref-Eyles_37-0" class="reference"><a href="#cite_note-Eyles-37"><span class="cite-bracket">&#91;</span>37<span class="cite-bracket">&#93;</span></a></sup> Continental glaciers developed in Africa and eastern Brazil, while alpine glaciers formed in the Andes.<sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">&#91;</span>38<span class="cite-bracket">&#93;</span></a></sup> In western South America (Peru, Bolivia and northern Argentina) were found glacio-marine diamictites interbedded with turbidites, shales, mud flows and debris flows, dated as early Silurian (Llandonvery), with a southward extension into northern Argentina and western Paraguay, and with a probably northern extension into Peru, Ecuador and Colombia.<sup id="cite_ref-Díaz-MartínezGrahn2007_7-1" class="reference"><a href="#cite_note-Díaz-MartínezGrahn2007-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> </p><p>A major ice age, the Andean-Saharan was preceded by the <a href="/wiki/Cryogenian" title="Cryogenian">Cryogenian</a> ice ages (720–630 Ma, the <a href="/wiki/Sturtian_glaciation" title="Sturtian glaciation">Sturtian</a> and <a href="/wiki/Marinoan_glaciation" title="Marinoan glaciation">Marinoan glaciations</a>), often referred to as <a href="/wiki/Snowball_Earth" title="Snowball Earth">Snowball Earth</a>, and followed by the <a href="/wiki/Karoo_Ice_Age" class="mw-redirect" title="Karoo Ice Age">Karoo Ice Age</a> (350–260 Ma).<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">&#91;</span>39<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Evidence">Evidence</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=5" title="Edit section: Evidence"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Lithologic">Lithologic</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=6" title="Edit section: Lithologic"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>The sequence of the stratigraphic architecture of the <a href="/wiki/Bighorn_Dolomite" title="Bighorn Dolomite">Bighorn Dolomite</a> (which represents end of the Ordovician period), is consistent with the gradual buildup of glacial ice. The sequences of the Bighorn Dolomite display systematic changes in their component cycles, and the changes in these cycles are interpreted as being a change from a greenhouse climate to a transitional ice house climate.<sup id="cite_ref-Bighorn_Dolomite,_Wyoming_10-2" class="reference"><a href="#cite_note-Bighorn_Dolomite,_Wyoming-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Possible_causes">Possible causes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=7" title="Edit section: Possible causes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="CO2_depletion">CO<sub>2</sub> depletion</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=8" title="Edit section: CO2 depletion"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>One of the factors that hindered glaciation during the early Paleozoic was atmospheric CO<sub>2</sub> concentrations, which at the time were somewhere between 8 and 20 times pre-industrial levels.<sup id="cite_ref-impacts_40-0" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> However, <a href="/wiki/Solar_irradiance" title="Solar irradiance">solar irradiance</a> was significantly lower during the Late Ordovician; 450 million years ago, solar irradiance of Earth was about 1312.00 W&#8239;m<sup>−2</sup> compared to 1360.89 W&#8239;m<sup>−2</sup> in the present day.<sup id="cite_ref-41" class="reference"><a href="#cite_note-41"><span class="cite-bracket">&#91;</span>41<span class="cite-bracket">&#93;</span></a></sup> Furthermore, CO<sub>2</sub> concentrations are thought to have dropped significantly in the Hirnantian, which could have induced widespread glaciation during an overall cooling trend.<sup id="cite_ref-42" class="reference"><a href="#cite_note-42"><span class="cite-bracket">&#91;</span>42<span class="cite-bracket">&#93;</span></a></sup> Methods for the removal of CO<sub>2</sub> during this time were not well known,<sup id="cite_ref-Bathymetric_and_Isotopic_evidence_27-2" class="reference"><a href="#cite_note-Bathymetric_and_Isotopic_evidence-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> and are still hotly debated, with the radiation of terrestrial plants,<sup id="cite_ref-Lenton2012_43-0" class="reference"><a href="#cite_note-Lenton2012-43"><span class="cite-bracket">&#91;</span>43<span class="cite-bracket">&#93;</span></a></sup> enhanced oceanic organic carbon burial,<sup id="cite_ref-44" class="reference"><a href="#cite_note-44"><span class="cite-bracket">&#91;</span>44<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-45" class="reference"><a href="#cite_note-45"><span class="cite-bracket">&#91;</span>45<span class="cite-bracket">&#93;</span></a></sup> and a reduction in volcanic outgassing of carbon dioxide having been proposed.<sup id="cite_ref-46" class="reference"><a href="#cite_note-46"><span class="cite-bracket">&#91;</span>46<span class="cite-bracket">&#93;</span></a></sup> It could have been possible for glaciation to initiate with high levels of CO<sub>2</sub>, but it would have depended highly on continental configuration.<sup id="cite_ref-impacts_40-1" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Silicate_weathering">Silicate weathering</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=9" title="Edit section: Silicate weathering"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Long-term <a href="/wiki/Carbonate%E2%80%93silicate_cycle" title="Carbonate–silicate cycle">silicate weathering</a> is a major mechanism through which CO<sub>2</sub> is removed from the atmosphere, converting it into <a href="/wiki/Bicarbonate" title="Bicarbonate">bicarbonate</a> which is stored in marine sediments. This has often been linked to the <a href="/wiki/Taconic_orogeny" title="Taconic orogeny">Taconic Orogeny</a>, a mountain-building event on the east coast of Laurentia (present-day North America).<sup id="cite_ref-:4_47-0" class="reference"><a href="#cite_note-:4-47"><span class="cite-bracket">&#91;</span>47<span class="cite-bracket">&#93;</span></a></sup> Another hypothesis is that a hypothetical large igneous province in the Katian led to basaltic flooding caused by high continental volcanic activity during that period. In the short term, this would have released a large amount of CO<sub>2</sub> into the atmosphere, which may explain a warming pulse in the Katian. However, in the long term flood basalts would have left behind plains of basaltic rock, replacing exposures of granitic rock. Basaltic rocks weather substantially faster than granitic rocks, which would quickly remove CO<sub>2</sub> from the atmosphere at a much faster rate than before the volcanic activity.<sup id="cite_ref-Katian_Large_Igneous_48-0" class="reference"><a href="#cite_note-Katian_Large_Igneous-48"><span class="cite-bracket">&#91;</span>48<span class="cite-bracket">&#93;</span></a></sup> CO<sub>2</sub> levels could also have decreased due to accelerated silicate weathering caused by the expansion of terrestrial non-vascular plants. Vascular plants only appeared 15 Ma after the glaciation.<sup id="cite_ref-49" class="reference"><a href="#cite_note-49"><span class="cite-bracket">&#91;</span>49<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Lenton2012_43-1" class="reference"><a href="#cite_note-Lenton2012-43"><span class="cite-bracket">&#91;</span>43<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Organic_carbon_burial">Organic carbon burial</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=10" title="Edit section: Organic carbon burial"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Isotopic evidence points to a global <a href="/wiki/Hirnantian" title="Hirnantian">Hirnantian</a> positive shift in δ<sup>13</sup>C at nearly the same time as the positive shift in marine carbonate δ<sup>18</sup>O.<sup id="cite_ref-Wang1997_50-0" class="reference"><a href="#cite_note-Wang1997-50"><span class="cite-bracket">&#91;</span>50<span class="cite-bracket">&#93;</span></a></sup> This shift is known as the <a href="/wiki/Hirnantian_Isotopic_Carbon_Excursion" title="Hirnantian Isotopic Carbon Excursion">Hirnantian Isotopic Carbon Excursion</a> (HICE).<sup id="cite_ref-Men2022_51-0" class="reference"><a href="#cite_note-Men2022-51"><span class="cite-bracket">&#91;</span>51<span class="cite-bracket">&#93;</span></a></sup> The positive shift in δ<sup>13</sup>C implies a change in the <a href="/wiki/Carbon_cycle" title="Carbon cycle">carbon cycle</a> leading to more burial of organic carbon,<sup id="cite_ref-Men2022_51-1" class="reference"><a href="#cite_note-Men2022-51"><span class="cite-bracket">&#91;</span>51<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-52" class="reference"><a href="#cite_note-52"><span class="cite-bracket">&#91;</span>52<span class="cite-bracket">&#93;</span></a></sup> though some researchers hold a conflicting interpretation of this δ<sup>13</sup>C change as being caused by increased weathering of carbonate platforms exposed by sea level fall.<sup id="cite_ref-53" class="reference"><a href="#cite_note-53"><span class="cite-bracket">&#91;</span>53<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-54" class="reference"><a href="#cite_note-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup> This enhanced organic carbon burial resulted in a decrease in the atmospheric CO<sub>2</sub> levels and an inverse greenhouse effect, allowing glaciation to occur more readily.<sup id="cite_ref-Bathymetric_and_Isotopic_evidence_27-3" class="reference"><a href="#cite_note-Bathymetric_and_Isotopic_evidence-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Gamma-ray_burst">Gamma-ray burst</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=11" title="Edit section: Gamma-ray burst"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>A gamma-ray burst (GRB) has been suggested by some researchers as the cause of the abrupt glaciation at the beginning of the Hirnantian.<sup id="cite_ref-GammaRayBurstLOME_55-0" class="reference"><a href="#cite_note-GammaRayBurstLOME-55"><span class="cite-bracket">&#91;</span>55<span class="cite-bracket">&#93;</span></a></sup> The effects of a ten second GRB occurring within two <a href="/wiki/Parsec" title="Parsec">kiloparsecs</a> of Earth would have delivered it a fluence of 100 kilojoules per square metre. This would have resulted in large amounts of <a href="/wiki/Nitric_acid" title="Nitric acid">nitric acid</a> raining down on Earth's surface in the aftermath of the gamma-ray burst, causing blooms of nitrate-limited photosynthesisers that would have sequestered large amounts of carbon dioxide from the atmosphere. Additionally, the GRB would have initiated a major depletion of <a href="/wiki/Ozone" title="Ozone">ozone</a>, another potent greenhouse gas, through its reaction with <a href="/wiki/Nitric_oxide" title="Nitric oxide">nitric oxide</a> produced as a result of the GRB's dissociation of diatomic nitrogen and subsequent reaction of nitrogen atoms with oxygen.<sup id="cite_ref-TerrestrialOzoneDepletion_56-0" class="reference"><a href="#cite_note-TerrestrialOzoneDepletion-56"><span class="cite-bracket">&#91;</span>56<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Asteroid_impact">Asteroid impact</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=12" title="Edit section: Asteroid impact"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading4"><h4 id="Ordovician_meteor_event">Ordovician meteor event</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=13" title="Edit section: Ordovician meteor event"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The breakup of the L-chondrite parent body caused a rain of extraterrestrial material onto the Earth called the <a href="/wiki/Ordovician_meteor_event" title="Ordovician meteor event">Ordovician meteor event</a>. This event increased stratospheric dust by 3 or 4 orders of magnitude and may have triggered the ice age by reflecting sunlight back into space.<sup id="cite_ref-57" class="reference"><a href="#cite_note-57"><span class="cite-bracket">&#91;</span>57<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Deniliquin_impact_structure">Deniliquin impact structure</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=14" title="Edit section: Deniliquin impact structure"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>A 2023 paper has proposed that the Hirnantian glaciation could have come about due to an <a href="/wiki/Impact_winter" title="Impact winter">impact winter</a> generated by the impact that formed the <a href="/wiki/Deniliquin_multiple-ring_feature" title="Deniliquin multiple-ring feature">Deniliquin multiple-ring feature</a> in what is now southeastern Australia, although this hypothesis currently remains untested.<sup id="cite_ref-AsteroidImpactOrigin_58-0" class="reference"><a href="#cite_note-AsteroidImpactOrigin-58"><span class="cite-bracket">&#91;</span>58<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Debris_ring">Debris ring</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=15" title="Edit section: Debris ring"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Rings_of_Earth" title="Rings of Earth">Rings of Earth</a></div> <p>A 2024 study suggests that rather than a complete breakup or outright impact, the L-chondrite parent body may have had a near-miss encounter with Earth, causing a part of it to break off from Earth's gravitational pull. This debris may have formed a <a href="/wiki/Ring_system" title="Ring system">planetary ring</a>, and down-falling debris from the ring may have shaded Earth from the sun's rays and triggering significant cooling. Evidence for this comes from the fact that craters dating from the <a href="/wiki/Ordovician_meteor_event" title="Ordovician meteor event">Ordovician meteor event</a> appear to cluster in a distinctive band around the Earth instead of being randomly scattered, which may have come from debris falling to Earth from the ring. This ring may have lasted for nearly 40 million years.<sup id="cite_ref-59" class="reference"><a href="#cite_note-59"><span class="cite-bracket">&#91;</span>59<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Volcanic_aerosols">Volcanic aerosols</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=16" title="Edit section: Volcanic aerosols"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Although volcanic activity often leads to warming through the release of greenhouse gasses, it may also lead to cooling via the production of <a href="/wiki/Aerosol" title="Aerosol">aerosols</a>, light-blocking particles. There is good evidence for elevated volcanic activity through the Hirnantian, based on anomalously high concentrations of mercury (Hg) in many areas. <a href="/wiki/Sulphur_dioxide" class="mw-redirect" title="Sulphur dioxide">Sulphur dioxide</a> (SO<sub>2</sub>) and other sulphurous volcanic gasses are converted into <a href="/wiki/Sulfate_aerosol" class="mw-redirect" title="Sulfate aerosol">sulphate aerosols</a> in the <a href="/wiki/Stratosphere" title="Stratosphere">stratosphere</a>, and short, periodic large igneous province eruptions may be able to account for cooling in this way.<sup id="cite_ref-60" class="reference"><a href="#cite_note-60"><span class="cite-bracket">&#91;</span>60<span class="cite-bracket">&#93;</span></a></sup> Although there is no direct evidence for a large igneous province during the Hirnantian, volcanism could still be a major factor. Explosive volcanic eruptions, which regularly send debris and volatiles into the stratosphere, would be even more effective at producing sulfate aerosols. Ash beds are common in the Late Ordovician, and Hirnantian pyrite records sulphur isotope anomalies consistent with stratospheric eruptions.<sup id="cite_ref-61" class="reference"><a href="#cite_note-61"><span class="cite-bracket">&#91;</span>61<span class="cite-bracket">&#93;</span></a></sup> The enormous megaeruption that formed the <a href="/wiki/Deicke_and_Millbrig_bentonite_layers" title="Deicke and Millbrig bentonite layers">Deicke bentonite layer</a> in particular has been linked to global cooling due to it coinciding with a major positive oxygen isotope excursion and the high sulphur concentration observed in its bentonite layer.<sup id="cite_ref-62" class="reference"><a href="#cite_note-62"><span class="cite-bracket">&#91;</span>62<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Sea_level_change">Sea level change</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=17" title="Edit section: Sea level change"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>One of the possible causes for the temperature drop during this period is a drop in sea level. Sea level must drop prior to the initiation of extensive ice sheets in order for it to be a possible trigger. A drop in sea level allows more land to become available for ice sheet growth. There is wide debate on the timing of sea level change, but there is some evidence that a sea level drop started before the <a href="/wiki/Ashgillian" class="mw-redirect" title="Ashgillian">Ashgillian</a>, which would have made it a contributing factor to glaciation.<sup id="cite_ref-impacts_40-2" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Palaeogeography">Palaeogeography</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=18" title="Edit section: Palaeogeography"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The possible setup of the paleogeography during the period from 460 Ma to 440 Ma falls in a range between the Caradocian and the Ashgillian. The choice of setup is important, because the Caradocian setup is more likely to produce glacial ice at high CO<sub>2</sub> concentrations, and the Ashgillian is more likely to produce glacial ice at low CO<sub>2</sub> concentrations.<sup id="cite_ref-impacts_40-3" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> </p><p>The height of the land mass above sea level also plays an important role, especially after ice sheets have been established. A higher elevation allows ice sheets to remain with more stability, but a lower elevation allows ice sheets to develop more readily. The Caradocian is considered to have a lower surface elevation, and though it would be better for initiation during high CO<sub>2</sub>, it would have a harder time maintaining glacial coverage.<sup id="cite_ref-revised_world_maps_63-0" class="reference"><a href="#cite_note-revised_world_maps-63"><span class="cite-bracket">&#91;</span>63<span class="cite-bracket">&#93;</span></a></sup> </p><p>From what we know about <a href="/wiki/Tectonic" class="mw-redirect" title="Tectonic">tectonic</a> movement, the time span required to allow the southward movement of <a href="/wiki/Gondwana" title="Gondwana">Gondwana</a> toward the South Pole would have been too long to trigger this glaciation. Tectonic movement tends to take several million years, but the scale of the glaciation seems to have occurred in less than 1 million years, but the exact time frame of glaciation ranges from less than 1 million years to 35 million years, so it could still be possible for tectonic movement to have triggered this glacial period.<sup id="cite_ref-impacts_40-4" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> Alternatively, <a href="/wiki/True_polar_wander" title="True polar wander">true polar wander</a> (TPW) and not conventional plate motion may have been responsible for the initiation of the Hirnantian glaciation. Palaeomagnetic data from between 450 and 440 Ma indicates a TPW of around ~50˚ occurring at a maximum speed of ~55 cm per year, which better explains the rapid motion of the continents than conventional plate tectonics.<sup id="cite_ref-64" class="reference"><a href="#cite_note-64"><span class="cite-bracket">&#91;</span>64<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Poleward_ocean_heat_transport">Poleward ocean heat transport</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=19" title="Edit section: Poleward ocean heat transport"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Ocean heat transport is a major driver in the warming of the poles, taking warm water from the equator and distributing it to higher latitudes. A weakening of this heat transport may have allowed the poles to cool enough to form ice under high CO<sub>2</sub> conditions.<sup id="cite_ref-impacts_40-5" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> Due to the paleogeographic configuration of the continents, global ocean heat transport is thought to have been stronger in the Late Ordovician.<sup id="cite_ref-glaciation_under_high_CO2_65-0" class="reference"><a href="#cite_note-glaciation_under_high_CO2-65"><span class="cite-bracket">&#91;</span>65<span class="cite-bracket">&#93;</span></a></sup> However, research shows that in order for glaciation to occur, poleward heat transport had to be lower, which creates a discrepancy in what is known.<sup id="cite_ref-impacts_40-6" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Orbital_parameters">Orbital parameters</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=20" title="Edit section: Orbital parameters"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Orbital parameters may have acted in conjunction with some of the above parameters to help start glaciation. The variation of the earth's precession, and eccentricity, could have set the off the tipping point for initiation of glaciation.<sup id="cite_ref-impacts_40-7" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> The Orbit at this time is thought to have been in a cold summer orbit for the Southern Hemisphere.<sup id="cite_ref-impacts_40-8" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> This type of orbital configuration is a change in the <a href="/wiki/Apsidal_precession" title="Apsidal precession">orbital precession</a> such that during the summer when the hemisphere is tilted toward the sun (in this case the earth) the earth is furthest away from the sun, and <a href="/wiki/Orbital_eccentricity" title="Orbital eccentricity">orbital eccentricity</a> such that the orbit of the earth is more elongated which would enhance the effect of precession. </p><p>Coupled models have shown that in order to maintain ice at the pole in the Southern Hemisphere, the earth would have to be in a cold summer configuration.<sup id="cite_ref-glaciation_under_high_CO2_65-1" class="reference"><a href="#cite_note-glaciation_under_high_CO2-65"><span class="cite-bracket">&#91;</span>65<span class="cite-bracket">&#93;</span></a></sup> The glaciation was most likely to start during a cold summer period because this configuration enhances the chance of snow and ice surviving throughout the summer.<sup id="cite_ref-impacts_40-9" class="reference"><a href="#cite_note-impacts-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="End_of_the_event">End of the event</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=21" title="Edit section: End of the event"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The cause for the end of the Late Ordovician Glaciation is a matter of intense research, but evidence shows that the deglaciation in the terminal Hirnantian may have occurred abruptly, as Silurian strata marks a significant change from the glacial deposits left during the Late Ordovician.<sup id="cite_ref-deglaciation_sequence_66-0" class="reference"><a href="#cite_note-deglaciation_sequence-66"><span class="cite-bracket">&#91;</span>66<span class="cite-bracket">&#93;</span></a></sup> Though the Hirnantian glaciation ended rapidly, milder glaciations continued to occur throughout the subsequent Silurian period,<sup id="cite_ref-TrotterEtAl2016PPP_35-1" class="reference"><a href="#cite_note-TrotterEtAl2016PPP-35"><span class="cite-bracket">&#91;</span>35<span class="cite-bracket">&#93;</span></a></sup> with the last glacial phase occurring in the Late Silurian.<sup id="cite_ref-LudfordianGlaciation_36-1" class="reference"><a href="#cite_note-LudfordianGlaciation-36"><span class="cite-bracket">&#91;</span>36<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Ice_collapse">Ice collapse</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=22" title="Edit section: Ice collapse"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>One of the possible causes for the end of the Hirnantian glaciation is that during the glacial maximum, the ice reached out too far and began collapsing on itself. The ice sheet initially stabilized once it reached as far north as <a href="/wiki/Ghat,_Libya" title="Ghat, Libya">Ghat, Libya</a> and developed a large proglacial fan-delta system. A glaciotectonic fold and thrust belt began to form from repeated small-scale fluctuations in the ice. The glaciotectonic fold and thrust belt eventually led to ice sheet collapse and retreat of the ice to south of Ghat. Once stabilized south of Ghat, the ice sheet began advancing north again. This cycle slowly shrank more south each time which lead to further retreat and further collapse of glacial conditions. This recursion allowed the melting of the ice sheet, and rising sea level. This hypothesis is supported by glacial deposits and large land formations found in Ghat, Libya which is part of the <a href="/wiki/Murzuq_Basin" class="mw-redirect" title="Murzuq Basin">Murzuq Basin</a>.<sup id="cite_ref-deglaciation_sequence_66-1" class="reference"><a href="#cite_note-deglaciation_sequence-66"><span class="cite-bracket">&#91;</span>66<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="CO2">CO<sub>2</sub></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=23" title="Edit section: CO2"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>As the Ice sheets began to increase the weathering of <a href="/wiki/Silicate" title="Silicate">silicate</a> rocks and basaltic important to <a href="/wiki/Carbon_sequestration" title="Carbon sequestration">carbon sequestration</a> (the silicates through the <a href="/wiki/Carbonate%E2%80%93silicate_cycle" title="Carbonate–silicate cycle">Carbonate–silicate cycle</a>, the <a href="/wiki/Basalt" title="Basalt">basalt</a> through forming <a href="/wiki/Calcium_carbonate" title="Calcium carbonate">calcium carbonate</a>) decreased, which caused CO<sub style="font-size: 80%;vertical-align: -0.35em">2</sub> levels to rise again, this in turned helped push deglaciation. This deglaciation cause the transformation of silicates exposed to the air (thus given the opportunity to bind to its CO<sub style="font-size: 80%;vertical-align: -0.35em">2</sub>) and weathering of basaltic rock to start back up which caused glaciation to occur again.<sup id="cite_ref-CO2_exit_glaciation_24-1" class="reference"><a href="#cite_note-CO2_exit_glaciation-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Significance">Significance</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=24" title="Edit section: Significance"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Even before the <a href="/wiki/Mass_extinction" class="mw-redirect" title="Mass extinction">mass extinction</a> at the end of the Ordovician, which resulted in a significant drop in chitinozoan diversity and abundance,<sup id="cite_ref-67" class="reference"><a href="#cite_note-67"><span class="cite-bracket">&#91;</span>67<span class="cite-bracket">&#93;</span></a></sup> the biodiversity of <a href="/wiki/Chitinozoan" title="Chitinozoan">chitinozoans</a> was adversely impacted by the onset of the Andean-Saharan glaciation. Following a peak in diversity in the late Darriwilian, chitinozoans declined in diversity as the Late Ordovician progressed. An exception to this declining trend of chitinozoan diversity was exhibited in Laurentia due to its low latitude position and warmer climate.<sup id="cite_ref-68" class="reference"><a href="#cite_note-68"><span class="cite-bracket">&#91;</span>68<span class="cite-bracket">&#93;</span></a></sup> </p><p>The Late Ordovician Glaciation coincided with the second largest of the five major <a href="/wiki/Extinction_events" class="mw-redirect" title="Extinction events">extinction events</a>, known as the <a href="/wiki/Late_Ordovician_mass_extinction" title="Late Ordovician mass extinction">Late Ordovician mass extinction</a>. This period is the only known glaciation to occur alongside of a mass extinction event. The extinction event consisted of two discrete pulses. The first pulse of extinctions is thought to have taken place because of the rapid cooling, and increased oxygenation of the water column. This first pulse was the larger of the two and caused the extinction of most of the marine animal species that existed in the shallow and deep oceans. The second phase of extinction was associated with strong <a href="/wiki/Sea_level_rise" title="Sea level rise">sea level rise</a>, and due to the atmospheric conditions, namely oxygen levels being at or below 50% of present-day levels, high levels of anoxic waters would have been common. This anoxia would have killed off many of the survivors of the first extinction pulse. In all the extinction event of the Late Ordovician saw a loss of 85% of marine animal species and 26% of animal families.<sup id="cite_ref-Sulfidic_driver_69-0" class="reference"><a href="#cite_note-Sulfidic_driver-69"><span class="cite-bracket">&#91;</span>69<span class="cite-bracket">&#93;</span></a></sup> </p><p>The deglaciation at the end of the Homerian glacial interval was coeval with the first major radiation of trilete spore-producing plants, harbingering the dawn of the <a href="/wiki/Silurian-Devonian_Terrestrial_Revolution" title="Silurian-Devonian Terrestrial Revolution">Silurian-Devonian Terrestrial Revolution</a>. The later middle Ludfordian glaciation caused a sea level drop that created vast areas of new terrestrial habitats that were promptly colonised by land plants, further facilitating their diversification.<sup id="cite_ref-SilurianPlantDynamics_70-0" class="reference"><a href="#cite_note-SilurianPlantDynamics-70"><span class="cite-bracket">&#91;</span>70<span class="cite-bracket">&#93;</span></a></sup> The warming during the <a href="/wiki/Pridoli_Epoch" title="Pridoli Epoch">Pridoli</a> that marked the end of the Andean-Saharan glaciation saw further floral expansion.<sup id="cite_ref-71" class="reference"><a href="#cite_note-71"><span class="cite-bracket">&#91;</span>71<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=25" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Timeline_of_glaciation" title="Timeline of glaciation">Timeline of glaciation</a></li> <li><a href="/wiki/Ordovician%E2%80%93Silurian_extinction_events" class="mw-redirect" title="Ordovician–Silurian extinction events">Ordovician–Silurian extinction events</a></li> <li><a href="/wiki/Late_Paleozoic_icehouse" title="Late Paleozoic icehouse">Late Paleozoic icehouse</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hirnantian_glaciation&amp;action=edit&amp;section=26" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-EPIA-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-EPIA_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-EPIA_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-EPIA_1-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFPohlDonnadieuLe_HirLadant2016" class="citation journal cs1">Pohl, Alexandre; Donnadieu, Yannick; Le Hir, Guillaume; Ladant, Jean-Baptiste; Dumas, Christophe; Alvarez-Solas, Jorge; Vandenbroucke, Thijs R. A. (28 May 2016). <a rel="nofollow" class="external text" href="https://doi.org/10.1002%2F2016PA002928">"Glacial onset predated Late Ordovician climate cooling"</a>. <i><a href="/wiki/Paleoceanography_and_Paleoclimatology" title="Paleoceanography and Paleoclimatology">Paleoceanography and Paleoclimatology</a></i>. <b>31</b> (6): 800–821. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2016PalOc..31..800P">2016PalOc..31..800P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1002%2F2016PA002928">10.1002/2016PA002928</a></span>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/1854%2FLU-8057556">1854/LU-8057556</a></span>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:133243759">133243759</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Paleoceanography+and+Paleoclimatology&amp;rft.atitle=Glacial+onset+predated+Late+Ordovician+climate+cooling&amp;rft.volume=31&amp;rft.issue=6&amp;rft.pages=800-821&amp;rft.date=2016-05-28&amp;rft_id=info%3Ahdl%2F1854%2FLU-8057556&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A133243759%23id-name%3DS2CID&amp;rft_id=info%3Adoi%2F10.1002%2F2016PA002928&amp;rft_id=info%3Abibcode%2F2016PalOc..31..800P&amp;rft.aulast=Pohl&amp;rft.aufirst=Alexandre&amp;rft.au=Donnadieu%2C+Yannick&amp;rft.au=Le+Hir%2C+Guillaume&amp;rft.au=Ladant%2C+Jean-Baptiste&amp;rft.au=Dumas%2C+Christophe&amp;rft.au=Alvarez-Solas%2C+Jorge&amp;rft.au=Vandenbroucke%2C+Thijs+R.+A.&amp;rft_id=https%3A%2F%2Fdoi.org%2F10.1002%252F2016PA002928&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHirnantian+glaciation" class="Z3988"></span></span> </li> <li id="cite_note-GlacioeustasyBlackShalesEarlyPalaeozoicIcehouse-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-GlacioeustasyBlackShalesEarlyPalaeozoicIcehouse_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-GlacioeustasyBlackShalesEarlyPalaeozoicIcehouse_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPage,_A.Zalasiewicz,_J.Williams,_M.Popov,_L.2007" class="citation book cs1">Page, A.; Zalasiewicz, J.; Williams, M.; Popov, L. (2007). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=j4eCj1J8k0IC&amp;pg=PA123">"Were transgressive black shales a negative feedback modulating glacioeustasy in the Early Palaeozoic Icehouse?"</a>. In Williams, Mark; Haywood, A. M.; Gregory, J.; et&#160;al. (eds.). <i>Deep-time perspectives on climate change: marrying the signal from computer models and biological proxies</i>. Special Publication of the Geological Society of London. 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A.; Armstrong, Howard A.; Williams, Mark; Paris, Florentin; Sabbe, Koen; Zalasiewicz, Jan A.; Nõlvak, Jaak; Verniers, Jacques (15 August 2010). <a rel="nofollow" class="external text" href="https://www.sciencedirect.com/science/article/abs/pii/S0031018209005215">"Epipelagic chitinozoan biotopes map a steep latitudinal temperature gradient for earliest Late Ordovician seas: Implications for a cooling Late Ordovician climate"</a>. <i><a href="/wiki/Palaeogeography,_Palaeoclimatology,_Palaeoecology" title="Palaeogeography, Palaeoclimatology, Palaeoecology">Palaeogeography, Palaeoclimatology, Palaeoecology</a></i>. <b>294</b> (3–4): 202–219. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2010PPP...294..202V">2010PPP...294..202V</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.palaeo.2009.11.026">10.1016/j.palaeo.2009.11.026</a><span class="reference-accessdate">. Retrieved <span class="nowrap">29 December</span> 2022</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Palaeogeography%2C+Palaeoclimatology%2C+Palaeoecology&amp;rft.atitle=Epipelagic+chitinozoan+biotopes+map+a+steep+latitudinal+temperature+gradient+for+earliest+Late+Ordovician+seas%3A+Implications+for+a+cooling+Late+Ordovician+climate&amp;rft.volume=294&amp;rft.issue=3%E2%80%934&amp;rft.pages=202-219&amp;rft.date=2010-08-15&amp;rft_id=info%3Adoi%2F10.1016%2Fj.palaeo.2009.11.026&amp;rft_id=info%3Abibcode%2F2010PPP...294..202V&amp;rft.aulast=Vandenbroucke&amp;rft.aufirst=Thijs+R.+A.&amp;rft.au=Armstrong%2C+Howard+A.&amp;rft.au=Williams%2C+Mark&amp;rft.au=Paris%2C+Florentin&amp;rft.au=Sabbe%2C+Koen&amp;rft.au=Zalasiewicz%2C+Jan+A.&amp;rft.au=N%C3%B5lvak%2C+Jaak&amp;rft.au=Verniers%2C+Jacques&amp;rft_id=https%3A%2F%2Fwww.sciencedirect.com%2Fscience%2Farticle%2Fabs%2Fpii%2FS0031018209005215&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHirnantian+glaciation" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRosenauHermannLeslie2012" class="citation journal cs1 cs1-prop-long-vol">Rosenau, Nicholas A.; Hermann, Achim D.; Leslie, Stephen A. 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Retrieved <span class="nowrap">29 December</span> 2022</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Palaeogeography%2C+Palaeoclimatology%2C+Palaeoecology&amp;rft.atitle=Conodont+apatite+%CE%B418O+values+from+a+platform+margin+setting%2C+Oklahoma%2C+USA%3A+Implications+for+initiation+of+Late+Ordovician+icehouse+conditions&amp;rft.volume=315-316&amp;rft.pages=172-180&amp;rft.date=2012-01-15&amp;rft_id=info%3Adoi%2F10.1016%2Fj.palaeo.2011.12.003&amp;rft_id=info%3Abibcode%2F2012PPP...315..172R&amp;rft.aulast=Rosenau&amp;rft.aufirst=Nicholas+A.&amp;rft.au=Hermann%2C+Achim+D.&amp;rft.au=Leslie%2C+Stephen+A.&amp;rft_id=https%3A%2F%2Fwww.sciencedirect.com%2Fscience%2Farticle%2Fabs%2Fpii%2FS0031018211005839&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHirnantian+glaciation" class="Z3988"></span></span> </li> <li id="cite_note-Munnecke2010-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-Munnecke2010_5-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMunneckeCalnerHarperServais2010" class="citation journal cs1">Munnecke, Axel; Calner, Mikael; <a href="/wiki/David_Harper_(palaeontologist)" title="David Harper (palaeontologist)">Harper, David A. 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Retrieved <span class="nowrap">10 January</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Review+of+Palaeobotany+and+Palynology&amp;rft.atitle=The+effects+of+the+final+stages+of+the+Late+Ordovician+glaciation+on+marine+palynomorphs+%28chitinozoans%2C+acritarchs%2C+leiospheres%29+in+well+Nl-2+%28NE+Algerian+Sahara%29&amp;rft.volume=113&amp;rft.issue=1%E2%80%933&amp;rft.pages=87-104&amp;rft.date=2000-12&amp;rft_id=info%3Apmid%2F11164214&amp;rft_id=info%3Adoi%2F10.1016%2FS0034-6667%2800%2900054-3&amp;rft_id=info%3Abibcode%2F2000RPaPa.113...87P&amp;rft.aulast=Paris&amp;rft.aufirst=F.&amp;rft.au=Bourahrouh%2C+A.&amp;rft.au=H%C3%A9riss%C3%A9%2C+A.+L.&amp;rft_id=https%3A%2F%2Fwww.sciencedirect.com%2Fscience%2Farticle%2Fabs%2Fpii%2FS0034666700000543&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHirnantian+glaciation" class="Z3988"></span></span> </li> <li id="cite_note-68"><span class="mw-cite-backlink"><b><a href="#cite_ref-68">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAchabParis2007" class="citation journal cs1">Achab, Aïcha; Paris, Florentin (7 March 2007). <a rel="nofollow" class="external text" href="https://www.sciencedirect.com/science/article/abs/pii/S003101820600335X">"The Ordovician chitinozoan biodiversification and its leading factors"</a>. <i><a href="/wiki/Palaeogeography,_Palaeoclimatology,_Palaeoecology" title="Palaeogeography, Palaeoclimatology, Palaeoecology">Palaeogeography, Palaeoclimatology, Palaeoecology</a></i>. <b>245</b> (1–2): 5–19. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2007PPP...245....5A">2007PPP...245....5A</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.palaeo.2006.02.030">10.1016/j.palaeo.2006.02.030</a><span class="reference-accessdate">. Retrieved <span class="nowrap">16 October</span> 2022</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Palaeogeography%2C+Palaeoclimatology%2C+Palaeoecology&amp;rft.atitle=The+Ordovician+chitinozoan+biodiversification+and+its+leading+factors&amp;rft.volume=245&amp;rft.issue=1%E2%80%932&amp;rft.pages=5-19&amp;rft.date=2007-03-07&amp;rft_id=info%3Adoi%2F10.1016%2Fj.palaeo.2006.02.030&amp;rft_id=info%3Abibcode%2F2007PPP...245....5A&amp;rft.aulast=Achab&amp;rft.aufirst=A%C3%AFcha&amp;rft.au=Paris%2C+Florentin&amp;rft_id=https%3A%2F%2Fwww.sciencedirect.com%2Fscience%2Farticle%2Fabs%2Fpii%2FS003101820600335X&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHirnantian+glaciation" class="Z3988"></span></span> </li> <li id="cite_note-Sulfidic_driver-69"><span class="mw-cite-backlink"><b><a href="#cite_ref-Sulfidic_driver_69-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHammarlund2012" class="citation journal cs1 cs1-prop-long-vol">Hammarlund, E. 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Retrieved <span class="nowrap">14 August</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Bulletin+of+Geosciences&amp;rft.atitle=Early+Silurian+%28mid-Sheinwoodian%29+palynomorphs+from+the+Lod%C4%9Bnice-%C5%A0pi%C4%8Dat%C3%BD+vrch%2C+Prague+Basin%2C+Czech+Republic.&amp;rft.volume=97&amp;rft.issue=3&amp;rft.pages=385-396&amp;rft.date=2022&amp;rft_id=info%3Adoi%2F10.3140%2Fbull.geosci.1831&amp;rft.aulast=Bek&amp;rft.aufirst=Ji%C5%99%C3%AD&amp;rft.au=%C5%A0torch%2C+Petr&amp;rft.au=Tonarov%C3%A1%2C+Petra&amp;rft.au=Libert%C3%ADn%2C+Milan&amp;rft_id=https%3A%2F%2Fweb.p.ebscohost.com%2Fabstract%3Fdirect%3Dtrue%26profile%3Dehost%26scope%3Dsite%26authtype%3Dcrawler%26jrnl%3D12141119%26AN%3D163982020%26h%3DJ4hndCjVAEf8HgMrCxVarnB4jXNouJXI2MLEt1KUW9fo%252bXOJzJ9XohMEXUpR8bihoJuQJOAQHCSTgI%252bEs3PATQ%253d%253d%26crl%3Dc%26resultNs%3DAdminWebAuth%26resultLocal%3DErrCrlNotAuth%26crlhashurl%3Dlogin.aspx%253fdirect%253dtrue%2526profile%253dehost%2526scope%253dsite%2526authtype%253dcrawler%2526jrnl%253d12141119%2526AN%253d163982020&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHirnantian+glaciation" class="Z3988"></span></span> </li> </ol></div></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output 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href="/wiki/Template_talk:Ice_ages" title="Template talk:Ice ages"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Ice_ages" title="Special:EditPage/Template:Ice ages"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Ice_ages" style="font-size:114%;margin:0 4em"><a href="/wiki/Ice_age" title="Ice age">Ice ages</a></div></th></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Quaternary_glaciation" title="Quaternary glaciation">Quaternary</a> / <br /> <a href="/wiki/Late_Cenozoic_Ice_Age" title="Late Cenozoic Ice Age">Late Cenozoic</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Quaternary" title="Quaternary">Quaternary</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Antarctica" title="Antarctica">Antarctica</a></li> <li><a href="/wiki/Greenland_ice_sheet" title="Greenland ice sheet">Greenland</a></li> <li><a href="/wiki/Penultimate_Glacial_Period" title="Penultimate Glacial Period">Penultimate Glacial Period</a></li> <li><a href="/wiki/Last_Glacial_Period" title="Last Glacial Period">Last Glacial Period</a> (<a href="/wiki/Last_Glacial_Maximum" title="Last Glacial Maximum">Last Glacial Maximum</a>, <a href="/wiki/Older_Dryas" title="Older Dryas">Older</a> and <a href="/wiki/Younger_Dryas" title="Younger Dryas">Younger Dryas</a>)</li> <li>1st: <a href="/wiki/W%C3%BCrm_glaciation" title="Würm glaciation">Würm</a>, <a href="/wiki/Wisconsin_glaciation" title="Wisconsin glaciation">Wisconsin</a>, <a href="/wiki/Weichselian_glaciation" title="Weichselian glaciation">Weichselian</a>, <a href="/wiki/Devensian_glaciation" class="mw-redirect" title="Devensian glaciation">Devensian</a>/<a href="/w/index.php?title=Midlandian_glaciation&amp;action=edit&amp;redlink=1" class="new" title="Midlandian glaciation (page does not exist)">Midlandian</a>, <a href="/wiki/Pinedale_glaciation" class="mw-redirect" title="Pinedale glaciation">Pinedale</a>/<a href="/wiki/Fraser_glaciation" class="mw-redirect" title="Fraser glaciation">Fraser</a>, <a href="/wiki/Merida_glaciation" class="mw-redirect" title="Merida glaciation">Merida</a>, <a href="/wiki/Llanquihue_glaciation" title="Llanquihue glaciation">Llanquihue</a></li> <li>2nd: <a href="/wiki/Riss_glaciation" title="Riss glaciation">Riss</a>, <a href="/wiki/Illinoian_(stage)" title="Illinoian (stage)">Illinoian</a>, <a href="/wiki/Saale_glaciation" title="Saale glaciation">Saale</a>, <a href="/wiki/Wolstonian_Stage" title="Wolstonian Stage">Wolstonian</a>, <a href="/wiki/Santa_Mar%C3%ADa_glaciation" class="mw-redirect" title="Santa María glaciation">Santa María</a></li> <li>3rd–6th: <a href="/wiki/Mindel_glaciation" title="Mindel glaciation">Mindel</a>, <a href="/wiki/Pre-Illinoian" title="Pre-Illinoian">Pre-Illinoian</a>, <a href="/wiki/Elster_glaciation" title="Elster glaciation">Elster</a>, <a href="/wiki/Anglian_stage" title="Anglian stage">Anglian</a>, <a href="/w/index.php?title=Rio_Llico_glaciation&amp;action=edit&amp;redlink=1" class="new" title="Rio Llico glaciation (page does not exist)">Rio Llico</a></li> <li>7th–8th: <a href="/wiki/Gunz_glaciation" class="mw-redirect" title="Gunz glaciation">Günz</a>, <a href="/wiki/Pre-Illinoian" title="Pre-Illinoian">Pre-Illinoian</a>, <a href="/wiki/Elbe_glaciation" class="mw-redirect" title="Elbe glaciation">Elbe</a> or <a href="/wiki/Menapian_glaciation" class="mw-redirect" title="Menapian glaciation">Menapian</a>, <a href="/wiki/Beestonian_stage" title="Beestonian stage">Beestonian</a>, <a href="/w/index.php?title=Caracol_glaciation&amp;action=edit&amp;redlink=1" class="new" title="Caracol glaciation (page does not exist)">Caracol</a> (2.5 to 0 Mya)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Pliocene" title="Pliocene">Pliocene</a><br /><a href="/wiki/Miocene" title="Miocene">Miocene</a><br /><a href="/wiki/Oligocene" title="Oligocene">Oligocene</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Late_Cenozoic_Ice_Age" title="Late Cenozoic Ice Age">Antarctic</a> (34 to 2.5 Mya)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Paleozoic" title="Paleozoic">Paleozoic</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Hirnantian</a> (460 Mya to 430 Mya)</li> <li><a href="/wiki/Late_Paleozoic_icehouse" title="Late Paleozoic icehouse">Late Paleozoic icehouse</a> (360 Mya to 260 Mya)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Ediacaran" title="Ediacaran">Ediacaran</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Gaskiers_glaciation" title="Gaskiers glaciation">Gaskiers</a> (579.88 to 579.63 Mya)</li> <li><a href="/wiki/Baykonurian_glaciation" title="Baykonurian glaciation">Baykonurian</a> (547 to 541.5 Mya)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Cryogenian" title="Cryogenian">Cryogenian</a>-<a href="/wiki/Snowball_Earth" title="Snowball Earth">Snowball Earth</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Sturtian_glaciation" title="Sturtian glaciation">Sturtian</a> (717 to 660 Mya); <a href="/wiki/Marinoan_glaciation" title="Marinoan glaciation">Marinoan</a> (650 to 635 Mya)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Paleoproterozoic" title="Paleoproterozoic">Paleoproterozoic</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Huronian_glaciation" title="Huronian glaciation">Huronian</a> (2.4 to 2.1 Gya)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%"><a href="/wiki/Mesoarchean" title="Mesoarchean">Mesoarchean</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pongola_glaciation" title="Pongola glaciation">Pongola</a> (2.9 to 2.78 Gya)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background:#bbab85;;width:1%">Related topics</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glacial_period" title="Glacial period">Glacial period</a></li> <li><a href="/wiki/Greenhouse_and_icehouse_Earth" title="Greenhouse and icehouse Earth">Greenhouse and icehouse Earth</a></li> <li><a href="/wiki/Great_Oxidation_Event" title="Great Oxidation Event">Great Oxidation Event</a></li> <li><a href="/wiki/Little_Ice_Age" title="Little Ice Age">Little Ice Age</a></li> <li><a href="/wiki/Snowball_Earth" title="Snowball Earth">Snowball Earth</a></li> <li><a href="/wiki/Interglacial" title="Interglacial">Interglacial</a></li> <li><a href="/wiki/Milankovitch_cycles" title="Milankovitch cycles">Milankovitch cycles</a></li> <li><a href="/wiki/Stadial" class="mw-redirect" title="Stadial">Stadial</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background:#bbab85;"><div><b><a href="/wiki/Timeline_of_glaciation" title="Timeline of glaciation">Timeline of glaciation</a></b></div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐76dfdc59c8‐ts6xs Cached time: 20241126121333 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 1.347 seconds Real time usage: 1.460 seconds Preprocessor visited node count: 6196/1000000 Post‐expand include size: 339476/2097152 bytes Template argument size: 876/2097152 bytes Highest expansion depth: 8/100 Expensive parser function count: 2/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 322613/5000000 bytes Lua time usage: 0.919/10.000 seconds Lua memory usage: 5866097/52428800 bytes Number of Wikibase entities loaded: 0/400 --> <!-- Transclusion 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