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id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Inhoudsopgave omschakelen" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Inhoudsopgave omschakelen</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Hamilton Othanel Smith</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Ga naar een artikel in een andere taal. Beschikbaar in 47 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-47" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">47 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%87%D8%A7%D9%85%D9%84%D8%AA%D9%88%D9%86_%D8%B3%D9%85%D9%8A%D8%AB" title="هاملتون سميث – Arabisch" lang="ar" hreflang="ar" data-title="هاملتون سميث" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%87%D8%A7%D9%85%D9%84%D8%AA%D9%88%D9%86_%D8%B3%D9%85%D9%8A%D8%AB" title="هاملتون سميث – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="هاملتون سميث" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%87%D9%85%DB%8C%D9%84%D8%AA%D9%88%D9%86_%D8%A7%D9%88._%D8%A7%D8%B3%D9%85%DB%8C%D8%AA" title="همیلتون او. اسمیت – South Azerbaijani" lang="azb" hreflang="azb" data-title="همیلتون او. اسمیت" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%93%D0%B0%D0%BC%D1%96%D0%BB%D1%8C%D1%82%D0%B0%D0%BD_%D0%A1%D0%BC%D1%96%D1%82" title="Гамільтан Сміт – Belarussisch" lang="be" hreflang="be" data-title="Гамільтан Сміт" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A5%D0%B0%D0%BC%D0%B8%D0%BB%D1%82%D1%8A%D0%BD_%D0%A1%D0%BC%D0%B8%D1%82" title="Хамилтън Смит – Bulgaars" lang="bg" hreflang="bg" data-title="Хамилтън Смит" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B9%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%AE%E0%A6%BF%E0%A6%B2%E0%A7%8D%E0%A6%9F%E0%A6%A8_%E0%A6%93._%E0%A6%B8%E0%A7%8D%E0%A6%AE%E0%A6%BF%E0%A6%A5" title="হ্যামিল্টন ও. স্মিথ – Bengaals" lang="bn" hreflang="bn" data-title="হ্যামিল্টন ও. স্মিথ" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith – Catalaans" lang="ca" hreflang="ca" data-title="Hamilton Othanel Smith" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%DA%BE%D8%A7%D9%85%DB%8C%DA%B5%D8%AA%DB%86%D9%86_%D8%B3%D9%85%DB%8C%D8%AA" title="ھامیڵتۆن سمیت – Soranî" lang="ckb" hreflang="ckb" data-title="ھامیڵتۆن سمیت" data-language-autonym="کوردی" data-language-local-name="Soranî" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Tsjechisch" lang="cs" hreflang="cs" data-title="Hamilton O. Smith" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith – Duits" lang="de" hreflang="de" data-title="Hamilton Othanel Smith" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Engels" lang="en" hreflang="en" data-title="Hamilton O. Smith" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Esperanto" lang="eo" hreflang="eo" data-title="Hamilton O. Smith" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Spaans" lang="es" hreflang="es" data-title="Hamilton O. Smith" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Estisch" lang="et" hreflang="et" data-title="Hamilton Smith" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Baskisch" lang="eu" hreflang="eu" data-title="Hamilton Smith" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%87%D9%85%DB%8C%D9%84%D8%AA%D9%88%D9%86_%D8%A7%D9%88._%D8%A7%D8%B3%D9%85%DB%8C%D8%AA" title="همیلتون او. اسمیت – Perzisch" lang="fa" hreflang="fa" data-title="همیلتون او. اسمیت" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Fins" lang="fi" hreflang="fi" data-title="Hamilton Smith" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Frans" lang="fr" hreflang="fr" data-title="Hamilton Smith" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%94%D7%9E%D7%99%D7%9C%D7%98%D7%95%D7%9F_%D7%90%D7%95._%D7%A1%D7%9E%D7%99%D7%AA%27" title="המילטון או. סמית' – Hebreeuws" lang="he" hreflang="he" data-title="המילטון או. סמית'" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B9%E0%A5%88%E0%A4%AE%E0%A4%BF%E0%A4%B2%E0%A5%8D%E0%A4%9F%E0%A4%A8_%E0%A4%B8%E0%A5%8D%E0%A4%AE%E0%A4%BF%E0%A4%A5" title="हैमिल्टन स्मिथ – Hindi" lang="hi" hreflang="hi" data-title="हैमिल्टन स्मिथ" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Kroatisch" lang="hr" hreflang="hr" data-title="Hamilton O. Smith" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Hongaars" lang="hu" hreflang="hu" data-title="Hamilton O. Smith" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith – Indonesisch" lang="id" hreflang="id" data-title="Hamilton Othanel Smith" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Ido" lang="io" hreflang="io" data-title="Hamilton O. Smith" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Italiaans" lang="it" hreflang="it" data-title="Hamilton Smith" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%8F%E3%83%9F%E3%83%AB%E3%83%88%E3%83%B3%E3%83%BB%E3%82%B9%E3%83%9F%E3%82%B9" title="ハミルトン・スミス – Japans" lang="ja" hreflang="ja" data-title="ハミルトン・スミス" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A5%D0%B0%D0%BC%D0%B8%D0%BB%D1%82%D0%BE%D0%BD_%D0%A1%D0%BC%D0%B8%D1%82" title="Хамилтон Смит – Kazachs" lang="kk" hreflang="kk" data-title="Хамилтон Смит" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%95%B4%EB%B0%80%ED%84%B4_O._%EC%8A%A4%EB%AF%B8%EC%8A%A4" title="해밀턴 O. 스미스 – Koreaans" lang="ko" hreflang="ko" data-title="해밀턴 O. 스미스" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Latijn" lang="la" hreflang="la" data-title="Hamilton Smith" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Maleis" lang="ms" hreflang="ms" data-title="Hamilton O. Smith" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%87%D9%85%DB%8C%D9%84%D8%AA%D9%88%D9%86_%D8%A7%D9%88._%D8%A7%D8%B3%D9%85%DB%8C%D8%AA" title="همیلتون او. اسمیت – Mazanderani" lang="mzn" hreflang="mzn" data-title="همیلتون او. اسمیت" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith – Nedersaksisch" lang="nds" hreflang="nds" data-title="Hamilton Othanel Smith" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Noors - Bokmål" lang="nb" hreflang="nb" data-title="Hamilton O. Smith" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith – Pools" lang="pl" hreflang="pl" data-title="Hamilton Othanel Smith" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DB%81%D9%85%D9%84%D9%B9%D9%86_%D8%B3%D9%85%D8%AA%DA%BE" title="ہملٹن سمتھ – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ہملٹن سمتھ" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Portugees" lang="pt" hreflang="pt" data-title="Hamilton Smith" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A1%D0%BC%D0%B8%D1%82,_%D0%A5%D0%B0%D0%BC%D0%B8%D0%BB%D1%82%D0%BE%D0%BD" title="Смит, Хамилтон – Russisch" lang="ru" hreflang="ru" data-title="Смит, Хамилтон" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Simple English" lang="en-simple" hreflang="en-simple" data-title="Hamilton Smith" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Zweeds" lang="sv" hreflang="sv" data-title="Hamilton O. Smith" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Hamilton_Smith" title="Hamilton Smith – Swahili" lang="sw" hreflang="sw" data-title="Hamilton Smith" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Turks" lang="tr" hreflang="tr" data-title="Hamilton O. Smith" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%93%D0%B5%D0%BC%D1%96%D0%BB%D1%82%D0%BE%D0%BD_%D0%A1%D0%BC%D1%96%D1%82" title="Гемілтон Сміт – Oekraïens" lang="uk" hreflang="uk" data-title="Гемілтон Сміт" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%DB%81%D9%85%D9%84%D9%B9%D9%86_%D8%A7%D9%88_%D8%B3%D9%85%D8%AA%DA%BE" title="ہملٹن او سمتھ – Urdu" lang="ur" hreflang="ur" data-title="ہملٹن او سمتھ" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Oezbeeks" lang="uz" hreflang="uz" data-title="Hamilton O. Smith" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Oezbeeks" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Vietnamees" lang="vi" hreflang="vi" data-title="Hamilton O. Smith" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Hamilton_O._Smith" title="Hamilton O. Smith – Yoruba" lang="yo" hreflang="yo" data-title="Hamilton O. Smith" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E6%BC%A2%E5%BD%8C%E7%88%BE%E9%A0%93%C2%B7%E5%8F%B2%E5%AF%86%E6%96%AF" title="漢彌爾頓·史密斯 – Chinees" lang="zh" hreflang="zh" data-title="漢彌爾頓·史密斯" data-language-autonym="中文" data-language-local-name="Chinees" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q108896#sitelinks-wikipedia" title="Taalkoppelingen bewerken" class="wbc-editpage">Koppelingen bewerken</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Naamruimten"> <div id="p-associated-pages" class="vector-menu 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dir="ltr"><table class="infobox" cellpadding="1" cellspacing="1"> <tbody><tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center; font-size:120%;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Nobel_prize_medal.svg" class="mw-file-description" title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  Hamilton Othanel Smith </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/23_augustus" title="23 augustus">23 augustus</a> <a href="/wiki/1931" title="1931">1931</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Hamilton_Smith.jpg" class="mw-file-description"><img alt="Hamilton Othanel Smith" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Hamilton_Smith.jpg/266px-Hamilton_Smith.jpg" decoding="async" width="266" height="200" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Hamilton_Smith.jpg/399px-Hamilton_Smith.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Hamilton_Smith.jpg/532px-Hamilton_Smith.jpg 2x" data-file-width="2816" data-file-height="2112" /></a></span> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Verenigde_Staten_(hoofdbetekenis)" class="mw-redirect" title="Verenigde Staten (hoofdbetekenis)">Verenigde Staten</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/New_York_(stad)" title="New York (stad)">New York</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2">Fysiologie of Geneeskunde </td></tr> <tr> <th>Jaar </th> <td colspan="2">1978 </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor de ontdekking van <a href="/wiki/Restrictie-enzym" title="Restrictie-enzym">restrictie-enzymen</a>, welke een instrument zijn geworden in de <a href="/wiki/Moleculaire_biologie" title="Moleculaire biologie">moleculaire biologie</a> </td></tr> <tr> <th>Samen met </th> <td colspan="2"><a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a><br /><a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a><br /><a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a><br /><a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Sussman Yalow</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Allan_Cormack" class="mw-redirect" title="Allan Cormack">Allan Cormack</a><br /><a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>Hamilton Othanel Smith</b> (<a href="/wiki/New_York_(stad)" title="New York (stad)">New York</a>, <a href="/wiki/23_augustus" title="23 augustus">23 augustus</a> <a href="/wiki/1931" title="1931">1931</a>) is een Amerikaans <a href="/wiki/Microbiologie" title="Microbiologie">microbioloog</a> en <a href="/wiki/Nobelprijs" title="Nobelprijs">Nobelprijswinnaar</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hamilton_Othanel_Smith&veaction=edit&section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Hamilton_Othanel_Smith&action=edit&section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Smith werd geboren in New York als zoon van professor Bunnie Othanel Smith en de onderwijzers Tommie Naomi Hackey.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Hij begon zijn studie aan de <a href="/w/index.php?title=University_Laboratory_High_School_of_Urbana,_Illinois&action=edit&redlink=1" class="new" title="University Laboratory High School of Urbana, Illinois (de pagina bestaat niet)">University Laboratory High School of Urbana, Illinois</a>. Daarna ging hij naar de <a href="/wiki/Universiteit_van_Illinois_te_Urbana-Champaign" title="Universiteit van Illinois te Urbana-Champaign">Universiteit van Illinois te Urbana-Champaign</a>, maar in 1950 stapte hij over naar de <a href="/wiki/Universiteit_van_Californi%C3%AB_-_Berkeley" title="Universiteit van Californië - Berkeley">Universiteit van Californië, Berkeley</a>. Hier behaalde hij in 1952 zijn <a href="/wiki/Bachelor" title="Bachelor">Bachelor</a> in <a href="/wiki/Wiskunde" title="Wiskunde">wiskunde</a>.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> In 1956 kreeg hij zijn medisch diploma van de <a href="/wiki/Johns_Hopkins-universiteit" title="Johns Hopkins-universiteit">Johns Hopkins-universiteit</a>. Hierna was hij werkzaam aan de <a href="/wiki/Washington-universiteit_te_Saint_Louis" title="Washington-universiteit te Saint Louis">Washington-universiteit</a> in <a href="/wiki/Saint_Louis_(Missouri)" title="Saint Louis (Missouri)">Saint Louis</a> en de <a href="/wiki/Universiteit_van_Michigan" title="Universiteit van Michigan">Universiteit van Michigan</a> voordat hij in 1967 terugkeerde naar Johns Hopkins. In 1973 werd hij aldaar benoemd tot hoogleraar microbiologie. </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hamilton_Othanel_Smith&veaction=edit&section=2" title="Bewerk dit kopje: Werk" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Hamilton_Othanel_Smith&action=edit&section=2" title="De broncode bewerken van de sectie: Werk"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Na zijn studie ging Smith zich bezighouden met onderzoek naar <a href="/wiki/Restrictie-enzym" title="Restrictie-enzym">restrictie-enzymen</a>. Voor zijn ontdekking van type II restrictie-enzymen won hij in 1978 samen met <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> en <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> de <a href="/wiki/Nobelprijs_voor_de_Fysiologie_of_Geneeskunde" class="mw-redirect" title="Nobelprijs voor de Fysiologie of Geneeskunde">Nobelprijs voor de Fysiologie of Geneeskunde</a>. </p><p>Later werd hij een vooraanstaand figuur op het gebied van <a href="/wiki/Genomica" title="Genomica">genomica</a>. In 1995 maakten hij en zijn team een sequentie van het eerste bacteriële <a href="/wiki/Genoom" title="Genoom">genoom</a>, te weten die van de <i><a href="/wiki/Haemophilus_influenzae" title="Haemophilus influenzae">Haemophilus influenzae</a></i>. <i>H. influenzae</i> was hetzelfde type organisme waarin Smith in de jaren zestig een restrictie-enzym had ontdekt dat de symmetrische <a href="/wiki/Nucleotide" title="Nucleotide">nucleotidevolgorde</a> in dubbelstrengs-DNA kon doorbreken. Deze breukplaatsen bleken op vaste plaatsten te liggen waar de nucleotidevolgorde rotatiesymmetrie vertoond. Deze ontdekking gaf aanleiding tot het vinden van restrictie-enzymen bij allerlei bacteria en bleek van grote betekenis te zijn in het onderzoek naar <a href="/wiki/Recombinant_DNA" title="Recombinant DNA">recombinant DNA</a>. Smith speelde ook een belangrijke rol bij het onderzoek naar het <a href="/wiki/Menselijk_genoom" class="mw-redirect" title="Menselijk genoom">menselijk genoom</a>. In 1998 ging hij werken bij <a href="/w/index.php?title=Celera_Genomics&action=edit&redlink=1" class="new" title="Celera Genomics (de pagina bestaat niet)">Celera Genomics</a>, waarvan hij mede-oprichter was. </p><p>Smith kreeg in 2002 de leiding over een team aan het <a href="/w/index.php?title=J._Craig_Venter_Institute&action=edit&redlink=1" class="new" title="J. Craig Venter Institute (de pagina bestaat niet)">J. Craig Venter Institute</a>, waarmee hij werkt aan het maken van een <a href="/wiki/Synthetische_biologie" title="Synthetische biologie">synthetische bacterie</a>, <i><a href="/w/index.php?title=Mycoplasma_laboratorium&action=edit&redlink=1" class="new" title="Mycoplasma laboratorium (de pagina bestaat niet)">Mycoplasma laboratorium</a></i>. </p> <div class="mw-heading mw-heading2"><h2 id="Externe_link">Externe link</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hamilton_Othanel_Smith&veaction=edit&section=3" title="Bewerk dit kopje: Externe link" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Hamilton_Othanel_Smith&action=edit&section=3" title="De broncode bewerken van de sectie: Externe link"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="http://nobelprize.org/medicine/laureates/1978/smith-autobio.html">Autobiografie op de Nobelprijssite</a></li></ul> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <div class="reflist" style="list-style-type: decimal;"><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="http://www.nndb.com/people/480/000131087/">Biografie Hamilton O. Smith</a> op Notable Names Database (NNDb).<small> Geraadpleegd op 25 juni 2015</small></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://cal.berkeley.edu">@cal, UC Berkeley’s alumni community</a></span> </li> </ol></div></div> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output .navigatie-afb-rechts{position:absolute}.mw-parser-output .navigatie-afb-rechts{right:2px}.mw-parser-output .navigatie-afb-groot{float:right;padding-left:0.5em}.mw-parser-output .navigatie-titel{background-color:#ddeef8;padding:2px 5.5em;font-weight:bold}.mw-parser-output .navigatie-inhoud{padding:0.5em}.mw-parser-output .navigatie-inhoud p:first-child{margin:0}.mw-parser-output .navigatie div[style*="background-color"],.mw-parser-output .navigatie div[style*="background"]{color:inherit}@media screen{html.skin-theme-clientpref-night .mw-parser-output .navigatie-titel{background-color:var(--background-color-interactive,#eaecf0)!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navigatie-titel{background-color:var(--background-color-interactive,#eaecf0)!important}}</style><div class="navigatie mw-collapsible mw-collapsed" role="navigation" aria-labelledby="Bibliografische informatie" data-collapsetext="Inklappen" data-expandtext="Uitklappen"><div id="Bibliografische_informatie" class="navigatie-titel"><a href="/wiki/Wikipedia:Bibliografische_informatie" title="Wikipedia:Bibliografische informatie">Bibliografische informatie</a></div><div class="navigatie-inhoud mw-collapsible-content plainlist ta-left"><ul style="column-width:36em"><li><b><a href="/wiki/Gemeinsame_Normdatei" title="Gemeinsame Normdatei">Gemeinsame Normdatei</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/1219832405">1219832405</a></span></li><li><b><a href="/wiki/International_Standard_Name_Identifier" title="International Standard Name Identifier">International Standard Name Identifier</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://isni.org/isni/0000000058899718">0000 0000 5889 9718</a></span></li><li><b><a href="/wiki/Library_of_Congress_Control_Number" title="Library of Congress Control Number">Library of Congress Control Number</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/names/n2016043671">n2016043671</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Tsjechi%C3%AB" title="Nationale Bibliotheek van Tsjechië">Nationale Bibliotheek van Tsjechië</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=xx0193295&CON_LNG=ENG">xx0193295</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Isra%C3%ABl" title="Nationale Bibliotheek van Israël">Nationale Bibliotheek van Israël</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&local_base=NLX10&find_code=UID&request=987007405897505171">987007405897505171</a></span></li><li><b><a href="/wiki/LIBRIS" title="LIBRIS">LIBRIS</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://libris.kb.se/auth/285086">285086</a></span></li><li><b><a href="/wiki/Syst%C3%A8me_universitaire_de_documentation" title="Système universitaire de documentation">Système universitaire de documentation</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/237641615">237641615</a></span></li><li><b><a href="/wiki/Virtual_International_Authority_File" title="Virtual International Authority File">Virtual International Authority File</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/305857637">305857637</a></span></li><li><b></b> <b><a href="/wiki/WorldCat" title="WorldCat">WorldCat</a></b> (via VIAF): <a rel="nofollow" class="external text" href="https://www.worldcat.org/identities/containsVIAFID/305857637">305857637</a></li></ul></div></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67837862"> <div class="navigatie mw-collapsible mw-collapsed" role="navigation" aria-labelledby="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" data-expandtext="Uitklappen" data-collapsetext="Inklappen"> <div class="navigatie-afb-links noviewer"><div class="navigatie-bewerken show-autoconfirmed nomobile"><span class="nowrap skin-invert-image"><span typeof="mw:File"><a href="/wiki/Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Sjabloon:Navigatie winnaars van de Nobelprijs voor Fysiologie of Geneeskunde"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/16px-OOjs_UI_icon_article-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/24px-OOjs_UI_icon_article-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/32px-OOjs_UI_icon_article-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> · <span class="ui-state-disabled" typeof="mw:File"><span title="Overleg sjabloon (de pagina bestaat niet)"><img alt="Overleg sjabloon (de pagina bestaat niet)" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/16px-OOjs_UI_icon_speechBubbles-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/24px-OOjs_UI_icon_speechBubbles-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/32px-OOjs_UI_icon_speechBubbles-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></span></span> · <span typeof="mw:File"><a href="//nl.wikipedia.org/w/index.php?title=Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde&action=edit" title="Sjabloon bewerken"><img alt="Sjabloon bewerken" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/16px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/24px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/32px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span></div></div> <div id="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" class="navigatie-titel">Winnaars van de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a></div> <div class="navigatie-inhoud mw-collapsible-content"> <p>1901: <a href="/wiki/Emil_Adolf_von_Behring" title="Emil Adolf von Behring">Behring</a> · 1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> · 1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> · 1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Pavlov</a> · 1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> · 1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a>, <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Ramón y Cajal</a> · 1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a class="mw-selflink selflink">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1&useformat=desktop" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Overgenomen van "<a dir="ltr" href="https://nl.wikipedia.org/w/index.php?title=Hamilton_Othanel_Smith&oldid=66182355">https://nl.wikipedia.org/w/index.php?title=Hamilton_Othanel_Smith&oldid=66182355</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Categorie:Alles" title="Categorie:Alles">Categorieën</a>: <ul><li><a href="/wiki/Categorie:Amerikaans_microbioloog" title="Categorie:Amerikaans microbioloog">Amerikaans microbioloog</a></li><li><a href="/wiki/Categorie:Winnaar_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Categorie:Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde">Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde</a></li><li><a href="/wiki/Categorie:20e-eeuws_medicus" title="Categorie:20e-eeuws medicus">20e-eeuws medicus</a></li><li><a href="/wiki/Categorie:20e-eeuws_bioloog" title="Categorie:20e-eeuws bioloog">20e-eeuws bioloog</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> Deze pagina is voor het laatst bewerkt op 22 okt 2023 om 13:43.</li> <li id="footer-info-copyright">De tekst is beschikbaar onder de licentie <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/deed.nl">Creative Commons Naamsvermelding/Gelijk delen</a>, er kunnen aanvullende voorwaarden van toepassing zijn. 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