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Stanley B. Prusiner - Wikipedia

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Prusiner</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Ga naar een artikel in een andere taal. Beschikbaar in 55 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-55" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">55 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B3%D8%AA%D8%A7%D9%86%D9%84%D9%8A_%D8%A8%D8%B1%D9%88%D8%B3%D9%8A%D9%86%D8%B1" title="ستانلي بروسينر – Arabisch" lang="ar" hreflang="ar" data-title="ستانلي بروسينر" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%B3%D8%AA%D8%A7%D9%86%D9%84%D9%89_%D8%A8%D8%B1%D9%88%D8%B3%D9%8A%D9%86%D8%B1" title="ستانلى بروسينر – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="ستانلى بروسينر" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Stenli_Pruziner" title="Stenli Pruziner – Azerbeidzjaans" lang="az" hreflang="az" data-title="Stenli Pruziner" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbeidzjaans" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%A7%D8%B3%D8%AA%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%DB%8C%D8%B3%DB%8C%D9%86%D8%B1" title="استنلی پریسینر – South Azerbaijani" lang="azb" hreflang="azb" data-title="استنلی پریسینر" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D1%82%D1%8D%D0%BD%D0%BB%D1%96_%D0%91%D0%B5%D0%BD_%D0%9F%D1%80%D1%83%D0%B7%D1%96%D0%BD%D0%B5%D1%80" title="Стэнлі Бен Прузінер – Belarussisch" lang="be" hreflang="be" data-title="Стэнлі Бен Прузінер" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-be-x-old mw-list-item"><a href="https://be-tarask.wikipedia.org/wiki/%D0%A1%D1%82%D1%8D%D0%BD%D0%BB%D1%96_%D0%9F%D1%80%D1%83%D0%B7%D1%96%D0%BD%D1%8D%D1%80" title="Стэнлі Прузінэр – Belarusian (Taraškievica orthography)" lang="be-tarask" hreflang="be-tarask" data-title="Стэнлі Прузінэр" data-language-autonym="Беларуская (тарашкевіца)" data-language-local-name="Belarusian (Taraškievica orthography)" class="interlanguage-link-target"><span>Беларуская (тарашкевіца)</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8_%D0%9F%D1%80%D1%83%D1%81%D0%B8%D0%BD%D1%8A%D1%80" title="Стенли Прусинър – Bulgaars" lang="bg" hreflang="bg" data-title="Стенли Прусинър" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B8%E0%A7%8D%E0%A6%9F%E0%A6%BE%E0%A6%A8%E0%A6%B2%E0%A6%BF_%E0%A6%AC%E0%A6%BF_%E0%A6%AA%E0%A7%8D%E0%A6%B0%E0%A7%81%E0%A6%B8%E0%A6%BF%E0%A6%A8%E0%A6%BE%E0%A6%B0" title="স্টানলি বি প্রুসিনার – Bengaals" lang="bn" hreflang="bn" data-title="স্টানলি বি প্রুসিনার" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Stanley_Ben_Prusiner" title="Stanley Ben Prusiner – Catalaans" lang="ca" hreflang="ca" data-title="Stanley Ben Prusiner" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Tsjechisch" lang="cs" hreflang="cs" data-title="Stanley B. Prusiner" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Duits" lang="de" hreflang="de" data-title="Stanley Prusiner" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A3%CF%84%CE%AC%CE%BD%CE%BB%CE%B5%CF%8B_%CE%A0%CF%81%CE%BF%CF%8D%CE%B6%CE%B9%CE%BD%CE%B5%CF%81" title="Στάνλεϋ Προύζινερ – Grieks" lang="el" hreflang="el" data-title="Στάνλεϋ Προύζινερ" data-language-autonym="Ελληνικά" data-language-local-name="Grieks" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Engels" lang="en" hreflang="en" data-title="Stanley B. Prusiner" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Spaans" lang="es" hreflang="es" data-title="Stanley B. Prusiner" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Estisch" lang="et" hreflang="et" data-title="Stanley Prusiner" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Baskisch" lang="eu" hreflang="eu" data-title="Stanley Prusiner" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A7%D8%B3%D8%AA%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%DB%8C%D8%B3%DB%8C%D9%86%D8%B1" title="استنلی پریسینر – Perzisch" lang="fa" hreflang="fa" data-title="استنلی پریسینر" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Fins" lang="fi" hreflang="fi" data-title="Stanley Prusiner" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Frans" lang="fr" hreflang="fr" data-title="Stanley Prusiner" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Iers" lang="ga" hreflang="ga" data-title="Stanley Prusiner" data-language-autonym="Gaeilge" data-language-local-name="Iers" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Galicisch" lang="gl" hreflang="gl" data-title="Stanley B. Prusiner" data-language-autonym="Galego" data-language-local-name="Galicisch" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A1%D7%98%D7%A0%D7%9C%D7%99_%D7%A4%D7%A8%D7%95%D7%96%D7%99%D7%A0%D7%A8" title="סטנלי פרוזינר – Hebreeuws" lang="he" hreflang="he" data-title="סטנלי פרוזינר" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Kroatisch" lang="hr" hreflang="hr" data-title="Stanley B. Prusiner" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Hongaars" lang="hu" hreflang="hu" data-title="Stanley B. Prusiner" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Indonesisch" lang="id" hreflang="id" data-title="Stanley B. Prusiner" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Stanley_Ben_Prusiner" title="Stanley Ben Prusiner – Ido" lang="io" hreflang="io" data-title="Stanley Ben Prusiner" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Italiaans" lang="it" hreflang="it" data-title="Stanley Prusiner" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B9%E3%82%BF%E3%83%B3%E3%83%AA%E3%83%BC%E3%83%BBB%E3%83%BB%E3%83%97%E3%83%AB%E3%82%B7%E3%83%8A%E3%83%BC" title="スタンリー・B・プルシナー – Japans" lang="ja" hreflang="ja" data-title="スタンリー・B・プルシナー" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8_%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80" title="Стенли Прузинер – Kazachs" lang="kk" hreflang="kk" data-title="Стенли Прузинер" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%8A%A4%ED%83%A0%EB%A6%AC_B._%ED%94%84%EB%A3%A8%EC%8B%9C%EB%84%88" title="스탠리 B. 프루시너 – Koreaans" lang="ko" hreflang="ko" data-title="스탠리 B. 프루시너" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Latijn" lang="la" hreflang="la" data-title="Stanley B. Prusiner" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%B8%E0%B5%8D%E0%B4%B1%E0%B5%8D%E0%B4%B1%E0%B4%BE%E0%B5%BB%E0%B4%B2%E0%B4%BF_%E0%B4%AC%E0%B5%86%E0%B4%9E%E0%B5%8D%E0%B4%9A%E0%B4%AE%E0%B4%BF%E0%B5%BB_%E0%B4%AA%E0%B5%8D%E0%B4%B0%E0%B5%81%E0%B4%B8%E0%B4%BF%E0%B4%A8%E0%B5%BC" title="സ്റ്റാൻലി ബെഞ്ചമിൻ പ്രുസിനർ – Malayalam" lang="ml" hreflang="ml" data-title="സ്റ്റാൻലി ബെഞ്ചമിൻ പ്രുസിനർ" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Maleis" lang="ms" hreflang="ms" data-title="Stanley B. Prusiner" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%A7%D8%B3%D8%AA%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%DB%8C%D8%B3%DB%8C%D9%86%D8%B1" title="استنلی پریسینر – Mazanderani" lang="mzn" hreflang="mzn" data-title="استنلی پریسینر" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Stanley_Benjamin_Prusiner" title="Stanley Benjamin Prusiner – Nedersaksisch" lang="nds" hreflang="nds" data-title="Stanley Benjamin Prusiner" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Noors - Bokmål" lang="nb" hreflang="nb" data-title="Stanley B. Prusiner" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Occitaans" lang="oc" hreflang="oc" data-title="Stanley B. Prusiner" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Pools" lang="pl" hreflang="pl" data-title="Stanley Prusiner" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%B3%D9%B9%DB%8C%D9%86%D9%84%DB%8C_%D9%BE%D8%B1%D9%88%D8%B3%D9%86%D8%B1" title="سٹینلی پروسنر – Western Punjabi" lang="pnb" hreflang="pnb" data-title="سٹینلی پروسنر" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Portugees" lang="pt" hreflang="pt" data-title="Stanley Prusiner" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Roemeens" lang="ro" hreflang="ro" data-title="Stanley B. Prusiner" data-language-autonym="Română" data-language-local-name="Roemeens" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80,_%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8" title="Прузинер, Стенли – Russisch" lang="ru" hreflang="ru" data-title="Прузинер, Стенли" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Simple English" lang="en-simple" hreflang="en-simple" data-title="Stanley B. Prusiner" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Slowaaks" lang="sk" hreflang="sk" data-title="Stanley B. Prusiner" data-language-autonym="Slovenčina" data-language-local-name="Slowaaks" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Sloveens" lang="sl" hreflang="sl" data-title="Stanley B. Prusiner" data-language-autonym="Slovenščina" data-language-local-name="Sloveens" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D0%B8_%D0%91._%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80" title="Стенли Б. Прузинер – Servisch" lang="sr" hreflang="sr" data-title="Стенли Б. Прузинер" data-language-autonym="Српски / srpski" data-language-local-name="Servisch" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Zweeds" lang="sv" hreflang="sv" data-title="Stanley B. Prusiner" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Stanley_Prusiner" title="Stanley Prusiner – Swahili" lang="sw" hreflang="sw" data-title="Stanley Prusiner" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Turks" lang="tr" hreflang="tr" data-title="Stanley B. Prusiner" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%A1%D1%82%D3%99%D0%BD%D0%BB%D0%B8_%D0%9F%D1%80%D1%83%D0%B7%D0%B8%D0%BD%D0%B5%D1%80" title="Стәнли Прузинер – Tataars" lang="tt" hreflang="tt" data-title="Стәнли Прузинер" data-language-autonym="Татарча / tatarça" data-language-local-name="Tataars" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A1%D1%82%D0%B5%D0%BD%D0%BB%D1%96_%D0%9F%D1%80%D1%83%D0%B7%D1%96%D0%BD%D0%B5%D1%80" title="Стенлі Прузінер – Oekraïens" lang="uk" hreflang="uk" data-title="Стенлі Прузінер" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%B3%D9%B9%DB%8C%D9%86%D9%84%DB%8C_%D8%A8%DB%8C_%D9%BE%D8%B1%D9%88%D8%B3%DB%8C%D9%86%D8%B1" title="سٹینلی بی پروسینر – Urdu" lang="ur" hreflang="ur" data-title="سٹینلی بی پروسینر" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Vietnamees" lang="vi" hreflang="vi" data-title="Stanley B. Prusiner" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner – Yoruba" lang="yo" hreflang="yo" data-title="Stanley B. Prusiner" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%8F%B2%E5%9D%A6%E5%88%A9%C2%B7%E5%B8%83%E9%AD%AF%E5%B8%8C%E7%B4%8D" title="史坦利·布魯希納 – Chinees" lang="zh" hreflang="zh" data-title="史坦利·布魯希納" data-language-autonym="中文" data-language-local-name="Chinees" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q155380#sitelinks-wikipedia" title="Taalkoppelingen bewerken" class="wbc-editpage">Koppelingen bewerken</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Naamruimten"> <div id="p-associated-pages" class="vector-menu 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font-size:120%;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Nobel_prize_medal.svg" class="mw-file-description" title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>&#160;&#160;Stanley B. Prusiner </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/28_mei" title="28 mei">28 mei</a> <a href="/wiki/1942" title="1942">1942</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Prusiner_1.JPG" class="mw-file-description"><img alt="Stanley Prusiner in 2007" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a2/Prusiner_1.JPG/266px-Prusiner_1.JPG" decoding="async" width="266" height="200" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a2/Prusiner_1.JPG/399px-Prusiner_1.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a2/Prusiner_1.JPG/532px-Prusiner_1.JPG 2x" data-file-width="1280" data-file-height="960" /></a></span> </td></tr> <tr> <td style="text-align:center;" colspan="3"><div style="font-size:90%">Stanley Prusiner in 2007</div> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Verenigde_Staten" title="Verenigde Staten">Verenigde Staten</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/Des_Moines_(Iowa)" title="Des Moines (Iowa)">Des Moines</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2">Fysiologie of Geneeskunde </td></tr> <tr> <th>Jaar </th> <td colspan="2">1997 </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor zijn ontdekking van <a href="/wiki/Prion" title="Prion">prionen</a> - een nieuw biologisch infectieprincipe </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Peter C. Doherty</a><br /><a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Robert_Furchgott" class="mw-redirect" title="Robert Furchgott">Robert Furchgott</a><br /><a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a><br /><a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>&#160; <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span> &#160; </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>Stanley Benjamin Prusiner</b> (<a href="/wiki/Des_Moines_(Iowa)" title="Des Moines (Iowa)">Des Moines</a>, <a href="/wiki/28_mei" title="28 mei">28 mei</a> <a href="/wiki/1942" title="1942">1942</a>) is een <a href="/wiki/Joden" title="Joden">Joods</a>-<a href="/wiki/Verenigde_Staten" title="Verenigde Staten">Amerikaanse</a> <a href="/wiki/Neurologie" title="Neurologie">neuroloog</a> en <a href="/wiki/Biochemie" title="Biochemie">biochemicus</a> aan de <a href="/wiki/Universiteit_van_Californi%C3%AB" title="Universiteit van Californië">Universiteit van Californië</a> te <a href="/wiki/San_Francisco" title="San Francisco">San Francisco</a> (UCSF). Hij kreeg in <a href="/wiki/1997" title="1997">1997</a> de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a> voor zijn ontdekking van de <a href="/wiki/Prion" title="Prion">prionen</a>, een klasse van besmettelijke ziekteverwekkende <a href="/wiki/Prote%C3%AFne" class="mw-redirect" title="Proteïne">eiwitten</a>, die onder andere de <a href="/wiki/Boviene_spongiforme_encefalopathie" title="Boviene spongiforme encefalopathie">gekke-koeienziekte BSE</a>, <a href="/wiki/Scrapie" title="Scrapie">scrapie</a> en de <a href="/wiki/Ziekte_van_Creutzfeldt-Jakob" title="Ziekte van Creutzfeldt-Jakob">ziekte van Creutzfeldt-Jakob</a> kunnen veroorzaken. </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Stanley_B._Prusiner&amp;veaction=edit&amp;section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Stanley_B._Prusiner&amp;action=edit&amp;section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Prusiner werd geboren in Des Moines, Iowa, als zoon van Mariam (Spigel) en Lawrence Prusiner, een architect. Hij bracht zijn jeugd door in Des Moines en <a href="/wiki/Cincinnati_(Ohio)" class="mw-redirect" title="Cincinnati (Ohio)">Cincinnati</a>, waar hij naar de <a href="/w/index.php?title=Walnut_Hills_High_School&amp;action=edit&amp;redlink=1" class="new" title="Walnut Hills High School (de pagina bestaat niet)">Walnut Hills High School</a> ging. Hij studeerde <a href="/wiki/Scheikunde" title="Scheikunde">scheikunde</a> aan de <a href="/wiki/Universiteit_van_Pennsylvania" title="Universiteit van Pennsylvania">Universiteit van Pennsylvania</a>, behaalde zijn <a href="/wiki/Doctor_of_Medicine" title="Doctor of Medicine">M.D.</a> aan de <a href="/w/index.php?title=University_of_Pennsylvania_School_of_Medicine&amp;action=edit&amp;redlink=1" class="new" title="University of Pennsylvania School of Medicine (de pagina bestaat niet)">University of Pennsylvania School of Medicine</a> en promoveerde er in 1968. Hij voltooide daarna een <a href="/wiki/Stage" title="Stage">stage</a> medicijnen aan de <a href="/wiki/Universiteit_van_Californi%C3%AB" title="Universiteit van Californië">Universiteit van Californië</a>, San Francisco. </p><p>Na zijn studies ging Prusiner werken aan het <a href="/wiki/National_Institutes_of_Health" title="National Institutes of Health">National Institutes of Health</a>, waar hij zich bezighield met de studie van <i><a href="/wiki/Escherichia_coli" title="Escherichia coli">E. coli</a></i> in het laboratorium van <a href="/w/index.php?title=Earl_Stadtman&amp;action=edit&amp;redlink=1" class="new" title="Earl Stadtman (de pagina bestaat niet)">Earl Stadtman</a>. Na drie jaar bij het NIH, keerde hij terug naar de UCSF om zijn stage af te ronden. In 1974 werd hij op de universiteit lid van de faculteit <a href="/wiki/Neurologie" title="Neurologie">neurologie</a>. Sindsdien heeft hij verschillende posities gehad op de faculteit. In 1980 werd hij aldaar benoemd tot hoogleraar in de neurologie. </p><p>In 1992 werd Prusiner verkozen tot lid van de <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Stanley_B._Prusiner&amp;veaction=edit&amp;section=2" title="Bewerk dit kopje: Werk" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Stanley_B._Prusiner&amp;action=edit&amp;section=2" title="De broncode bewerken van de sectie: Werk"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Begin jaren 1970 begon Prusiner aan zijn onderzoek om de oorzaak te achterhalen van de ziekte van Creutzfeldt-Jakob, waarvan wetenschappers vermoedden dat dit werd veroorzaakt door een onbekend, nog niet ontdekt virus of bacterie. Dit werd nog eens benadrukt doordat de Nobelprijs in 1976 werd uitgereikt aan de Amerikaanse arts <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a> voor zijn theorie dat <a href="/wiki/Kuru_(ziekte)" title="Kuru (ziekte)">kuru</a>, een aan Creutzfelt-Jacob verwante hersenziekte, werd veroorzaakt door een virus. </p><p>De doorbraak kwam in 1982 toen Prusiner in het vakblad <i>Science</i> een artikel publiceerde met de ontdekking dat niet een virus, schimmel of bacterie verantwoordelijk waren voor deze tot dan toe onbegrijpelijke ziektes, maar infectueuze eiwitten. Voor deze eiwitten introduceerde hij de term prion, een samentrekking voor <i>proteinaceous infectious particle</i>. Zelfs nadat het prion het jaar erop positief werd geïdentificeerd in Prusiners laboratorium zou het bijna tien jaar duren voordat deze nieuwe ziekteverwekker algemeen werd geaccepteerd. De reden was dat zijn idee dwars tegen het heersende dogma inging, namelijk dat alle besmettelijke ziekten ontstaan door dragers van erfelijk materiaal, zoals virussen en bacteriën. Aangezien eiwitten noch <a href="/wiki/Desoxyribonucle%C3%AFnezuur" class="mw-redirect" title="Desoxyribonucleïnezuur">DNA</a>, noch <a href="/wiki/Ribonucle%C3%AFnezuur" title="Ribonucleïnezuur">RNA</a> bevatten, werd het onmogelijk geacht dat zij een infectie konden veroorzaken. </p><p>Het onderzoek naar prionen kwam in een stroomversnelling terecht toen eind jaren 1990 een aantal jonge Britten waren overleden aan een nieuwe <a href="/wiki/VCJD" title="VCJD">variant van Creutzfeldt-Jacob</a> door, naar wat later bleek, de consumptie van vlees van runderen met <i><a href="/wiki/Boviene_spongiforme_encefalopathie" title="Boviene spongiforme encefalopathie">boviene spongiforme encefalopathie</a></i> (BSE), ook wel de gekkekoeienziekte genoemd. </p><p>Behalve de Nobelprijs ontving Prusiner voor zijn werk ook de <a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award</a> voor basaal medisch onderzoek in 1994, de <a href="/w/index.php?title=Potamkinprijs&amp;action=edit&amp;redlink=1" class="new" title="Potamkinprijs (de pagina bestaat niet)">Potamkinprijs</a>, de <a href="/wiki/Wolfprijs" title="Wolfprijs">Wolfprijs</a> voor medisch onderzoek en de Gairdner Foundation Award. </p> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="http://www.nobelprize.org/nobel_prizes/medicine/laureates/1997/prusiner-bio.html">Autobiografie Stanley B. Prusiner</a> op Nobelprize.org</li> <li><span class="citation" id="CITEREFS.B._Prusiner1982"><span style="font-variant:small-caps;">S.B. Prusiner</span>&#32;(1982).&#32;<cite><a rel="nofollow" class="external text" href="https://archive.org/details/sim_science_1982-04-09_216_4542/page/n38">Novel infectious particles cause scrapie</a></cite>. <i>Science</i>&#32;<b>216</b>&#32;(4542): 136-144. <a href="/wiki/PubMed" title="PubMed">PMID</a> <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/6801762">6801762</a>. <a href="/wiki/Digital_object_identifier" title="Digital object identifier">DOI</a>: <a rel="nofollow" class="external text" href="https://doi.org/10.1126/science.6801762">10.1126/science.6801762</a>. </span></li> <li><span class="citation" id="CITEREFS.B._Prusiner1991"><span style="font-variant:small-caps;">S.B. Prusiner</span>&#32;(1991).&#32;<cite><a rel="nofollow" class="external text" href="https://archive.org/details/sim_science_1991-06-14_252_5012/page/n58">Molecular biology of prion diseases</a></cite>. <i>Science</i>&#32;<b>252</b>&#32;(5012): 1515-1522. <a href="/wiki/PubMed" title="PubMed">PMID</a> <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/1675487">1675487</a>. <a href="/wiki/Digital_object_identifier" title="Digital object identifier">DOI</a>: <a rel="nofollow" class="external text" href="https://doi.org/10.1126/science.1675487">10.1126/science.1675487</a>. </span></li></ul> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output .navigatie-afb-rechts{position:absolute}.mw-parser-output .navigatie-afb-rechts{right:2px}.mw-parser-output .navigatie-afb-groot{float:right;padding-left:0.5em}.mw-parser-output .navigatie-titel{background-color:#ddeef8;padding:2px 5.5em;font-weight:bold}.mw-parser-output .navigatie-inhoud{padding:0.5em}.mw-parser-output .navigatie-inhoud p:first-child{margin:0}.mw-parser-output .navigatie div[style*="background-color"],.mw-parser-output .navigatie div[style*="background"]{color:inherit}@media screen{html.skin-theme-clientpref-night .mw-parser-output .navigatie-titel{background-color:var(--background-color-interactive,#eaecf0)!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output 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mw-collapsible-content"> <p>1901:&#160;<a href="/wiki/Emil_Adolf_von_Behring" title="Emil Adolf von Behring">Behring</a> · 1902:&#160;<a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> · 1903:&#160;<a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> · 1904:&#160;<a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Pavlov</a> · 1905:&#160;<a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> · 1906:&#160;<a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a>, <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Ramón&#160;y&#160;Cajal</a> · 1907:&#160;<a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908:&#160;<a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909:&#160;<a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910:&#160;<a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911:&#160;<a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912:&#160;<a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913:&#160;<a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914:&#160;<a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919:&#160;<a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920:&#160;<a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922:&#160;<a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923:&#160;<a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924:&#160;<a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926:&#160;<a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927:&#160;<a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928:&#160;<a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929:&#160;<a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930:&#160;<a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931:&#160;<a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932:&#160;<a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933:&#160;<a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934:&#160;<a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935:&#160;<a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936:&#160;<a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937:&#160;<a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938:&#160;<a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939:&#160;<a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943:&#160;<a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944:&#160;<a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945:&#160;<a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946:&#160;<a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947:&#160;<span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948:&#160;<a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949:&#160;<a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950:&#160;<a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951:&#160;<a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952:&#160;<a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953:&#160;<a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954:&#160;<a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955:&#160;<a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956:&#160;<a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957:&#160;<a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958:&#160;<a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959:&#160;<a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960:&#160;<a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961:&#160;<a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962:&#160;<a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963:&#160;<a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964:&#160;<a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965:&#160;<a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966:&#160;<a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967:&#160;<a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968:&#160;<a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969:&#160;<a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970:&#160;<a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971:&#160;<a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972:&#160;<a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973:&#160;<a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974:&#160;<a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975:&#160;<a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976:&#160;<a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977:&#160;<a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978:&#160;<a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979:&#160;<a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980:&#160;<a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981:&#160;<a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982:&#160;<a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983:&#160;<a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984:&#160;<a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985:&#160;<a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986:&#160;<a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987:&#160;<a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988:&#160;<a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989:&#160;<a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990:&#160;<a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991:&#160;<a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992:&#160;<a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993:&#160;<a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994:&#160;<a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995:&#160;<a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996:&#160;<a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997:&#160;<a class="mw-selflink selflink">Prusiner</a> · 1998:&#160;<a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999:&#160;<a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000:&#160;<a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001:&#160;<a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002:&#160;<a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003:&#160;<a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004:&#160;<a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005:&#160;<a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006:&#160;<a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007:&#160;<a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008:&#160;<a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009:&#160;<a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010:&#160;<a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011:&#160;<a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012:&#160;<a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013:&#160;<a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014:&#160;<a href="/wiki/John_O%27Keefe" title="John O&#39;Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015:&#160;<a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>,&#160;<a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016:&#160;<a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017:&#160;<a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018:&#160;<a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019:&#160;<a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020:&#160;<a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021:&#160;<a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022:&#160;<a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023:&#160;<a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024:&#160;<a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div> <!-- NewPP limit report Parsed by mw‐api‐ext.codfw.main‐9884d96b7‐gvpdz Cached time: 20241127045147 Cache expiry: 2592000 Reduced expiry: false Complications: [] CPU time usage: 0.100 seconds Real time usage: 0.237 seconds Preprocessor visited node count: 1541/1000000 Post‐expand include size: 32294/2097152 bytes Template argument size: 13285/2097152 bytes Highest expansion depth: 10/100 Expensive parser function count: 1/500 Unstrip recursion depth: 0/20 Unstrip post‐expand size: 1688/5000000 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 76.849 1 -total 57.10% 43.878 1 Sjabloon:Infobox_Nobelprijswinnaar 51.83% 39.831 1 Sjabloon:Infobox_generiek 29.38% 22.576 1 Sjabloon:Wikidata_afbeelding 24.35% 18.714 1 Sjabloon:Appendix 18.35% 14.099 1 Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde 15.34% 11.790 1 Sjabloon:Navigatie_uitklapbaar 12.40% 9.531 2 Sjabloon:Citeer_journal 9.40% 7.223 1 Sjabloon:En 6.52% 5.009 1 Sjabloon:Taalaanduiding --> <!-- Saved in parser cache with key nlwiki:pcache:idhash:710086-0!canonical and timestamp 20241127045146 and revision id 68438564. 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