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Taxonomy (biology) - Wikipedia
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class="vector-toc-list"> <li id="toc-Monograph_and_taxonomic_revision" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Monograph_and_taxonomic_revision"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Monograph and taxonomic revision</span> </div> </a> <ul id="toc-Monograph_and_taxonomic_revision-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Taxonomic_characters" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Taxonomic_characters"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2</span> <span>Taxonomic characters</span> </div> </a> <ul id="toc-Taxonomic_characters-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Alpha_and_beta_taxonomy" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Alpha_and_beta_taxonomy"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.3</span> <span>Alpha and beta taxonomy</span> </div> </a> <ul id="toc-Alpha_and_beta_taxonomy-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Microtaxonomy_and_macrotaxonomy" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Microtaxonomy_and_macrotaxonomy"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.4</span> <span>Microtaxonomy and macrotaxonomy</span> </div> </a> <ul id="toc-Microtaxonomy_and_macrotaxonomy-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-History" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#History"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>History</span> </div> </a> <button aria-controls="toc-History-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle History subsection</span> </button> <ul id="toc-History-sublist" class="vector-toc-list"> <li id="toc-Pre-Linnaean" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Pre-Linnaean"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Pre-Linnaean</span> </div> </a> <ul id="toc-Pre-Linnaean-sublist" class="vector-toc-list"> <li id="toc-Early_taxonomists" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Early_taxonomists"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1.1</span> <span>Early taxonomists</span> </div> </a> <ul id="toc-Early_taxonomists-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ancient_times" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Ancient_times"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1.2</span> <span>Ancient times</span> </div> </a> <ul id="toc-Ancient_times-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Medieval" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Medieval"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1.3</span> <span>Medieval</span> </div> </a> <ul id="toc-Medieval-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Renaissance_and_early_modern" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Renaissance_and_early_modern"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1.4</span> <span>Renaissance and early modern</span> </div> </a> <ul id="toc-Renaissance_and_early_modern-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Linnaean_era" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Linnaean_era"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.2</span> <span>Linnaean era</span> </div> </a> <ul id="toc-Linnaean_era-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-The_digital_era_of_taxonomy" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#The_digital_era_of_taxonomy"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3</span> <span>The digital era of taxonomy</span> </div> </a> <ul id="toc-The_digital_era_of_taxonomy-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Modern_system_of_classification" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Modern_system_of_classification"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Modern system of classification</span> </div> </a> <button aria-controls="toc-Modern_system_of_classification-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Modern system of classification subsection</span> </button> <ul id="toc-Modern_system_of_classification-sublist" class="vector-toc-list"> <li id="toc-Kingdoms_and_domains" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Kingdoms_and_domains"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1</span> <span>Kingdoms and domains</span> </div> </a> <ul id="toc-Kingdoms_and_domains-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Recent_comprehensive_classifications" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Recent_comprehensive_classifications"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.2</span> <span>Recent comprehensive classifications</span> </div> </a> <ul id="toc-Recent_comprehensive_classifications-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Application" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Application"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Application</span> </div> </a> <button aria-controls="toc-Application-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Application subsection</span> </button> <ul id="toc-Application-sublist" class="vector-toc-list"> <li id="toc-Classifying_organisms" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Classifying_organisms"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.1</span> <span>Classifying organisms</span> </div> </a> <ul id="toc-Classifying_organisms-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Taxonomic_descriptions" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Taxonomic_descriptions"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.2</span> <span>Taxonomic descriptions</span> </div> </a> <ul id="toc-Taxonomic_descriptions-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Author_citation" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Author_citation"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.3</span> <span>Author citation</span> </div> </a> <ul id="toc-Author_citation-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Phenetics" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Phenetics"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Phenetics</span> </div> </a> <ul id="toc-Phenetics-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Databases" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Databases"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Databases</span> </div> </a> <ul id="toc-Databases-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notes" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notes"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>Notes</span> </div> </a> <ul id="toc-Notes-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">9</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Bibliography" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Bibliography"> <div class="vector-toc-text"> <span class="vector-toc-numb">10</span> <span>Bibliography</span> </div> </a> <ul id="toc-Bibliography-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">11</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Taxonomy (biology)</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 93 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-93" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">93 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Wetenskaplike_klassifikasie" title="Wetenskaplike klassifikasie – Afrikaans" lang="af" hreflang="af" data-title="Wetenskaplike klassifikasie" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-als mw-list-item"><a href="https://als.wikipedia.org/wiki/Taxonomie_(Biologie)" title="Taxonomie (Biologie) – Alemannic" lang="gsw" hreflang="gsw" data-title="Taxonomie (Biologie)" data-language-autonym="Alemannisch" data-language-local-name="Alemannic" class="interlanguage-link-target"><span>Alemannisch</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B9%D9%84%D9%85_%D8%A7%D9%84%D8%AA%D8%B5%D9%86%D9%8A%D9%81_(%D8%A3%D8%AD%D9%8A%D8%A7%D8%A1)" title="علم التصنيف (أحياء) – Arabic" lang="ar" hreflang="ar" data-title="علم التصنيف (أحياء)" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-an mw-list-item"><a href="https://an.wikipedia.org/wiki/Taxonom%C3%ADa" title="Taxonomía – Aragonese" lang="an" hreflang="an" data-title="Taxonomía" data-language-autonym="Aragonés" data-language-local-name="Aragonese" class="interlanguage-link-target"><span>Aragonés</span></a></li><li class="interlanguage-link interwiki-as mw-list-item"><a href="https://as.wikipedia.org/wiki/%E0%A6%9C%E0%A7%80%E0%A7%B1%E0%A6%AC%E0%A7%88%E0%A6%9C%E0%A7%8D%E0%A6%9E%E0%A6%BE%E0%A6%A8%E0%A6%BF%E0%A6%95_%E0%A6%AC%E0%A7%B0%E0%A7%8D%E0%A6%97%E0%A7%80%E0%A6%95%E0%A7%B0%E0%A6%A3" title="জীৱবৈজ্ঞানিক বৰ্গীকৰণ – Assamese" lang="as" hreflang="as" data-title="জীৱবৈজ্ঞানিক বৰ্গীকৰণ" data-language-autonym="অসমীয়া" data-language-local-name="Assamese" class="interlanguage-link-target"><span>অসমীয়া</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Taxonom%C3%ADa" title="Taxonomía – Asturian" lang="ast" hreflang="ast" data-title="Taxonomía" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%AA%D8%A7%DA%A9%D8%B3%D9%88%D9%86%D9%88%D9%85%DB%8C_(%D8%A8%DB%8C%D9%88%D9%84%D9%88%DA%98%DB%8C)" title="تاکسونومی (بیولوژی) – South Azerbaijani" lang="azb" hreflang="azb" data-title="تاکسونومی (بیولوژی)" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-ban mw-list-item"><a href="https://ban.wikipedia.org/wiki/Taksonomi_(biologi)" title="Taksonomi (biologi) – Balinese" lang="ban" hreflang="ban" data-title="Taksonomi (biologi)" data-language-autonym="Basa Bali" data-language-local-name="Balinese" class="interlanguage-link-target"><span>Basa Bali</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B6%E0%A7%8D%E0%A6%B0%E0%A7%87%E0%A6%A3%E0%A6%BF%E0%A6%AC%E0%A6%BF%E0%A6%A8%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%B8%E0%A6%AC%E0%A6%BF%E0%A6%A6%E0%A7%8D%E0%A6%AF%E0%A6%BE_(%E0%A6%9C%E0%A7%80%E0%A6%AC%E0%A6%AC%E0%A6%BF%E0%A6%9C%E0%A7%8D%E0%A6%9E%E0%A6%BE%E0%A6%A8)" title="শ্রেণিবিন্যাসবিদ্যা (জীববিজ্ঞান) – Bangla" lang="bn" hreflang="bn" data-title="শ্রেণিবিন্যাসবিদ্যা (জীববিজ্ঞান)" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Hun-l%C5%ABi-ha%CC%8Dk_(seng-bu%CC%8Dt-ha%CC%8Dk)" title="Hun-lūi-ha̍k (seng-bu̍t-ha̍k) – Minnan" lang="nan" hreflang="nan" data-title="Hun-lūi-ha̍k (seng-bu̍t-ha̍k)" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-bh mw-list-item"><a href="https://bh.wikipedia.org/wiki/%E0%A4%9F%E0%A5%88%E0%A4%95%E0%A5%8D%E0%A4%B8%E0%A5%8B%E0%A4%A8%E0%A5%89%E0%A4%AE%E0%A5%80_(%E0%A4%9C%E0%A5%80%E0%A4%B5_%E0%A4%AC%E0%A4%BF%E0%A4%9C%E0%A5%8D%E0%A4%9E%E0%A4%BE%E0%A4%A8)" title="टैक्सोनॉमी (जीव बिज्ञान) – Bhojpuri" lang="bh" hreflang="bh" data-title="टैक्सोनॉमी (जीव बिज्ञान)" data-language-autonym="भोजपुरी" data-language-local-name="Bhojpuri" class="interlanguage-link-target"><span>भोजपुरी</span></a></li><li class="interlanguage-link interwiki-bcl mw-list-item"><a href="https://bcl.wikipedia.org/wiki/Taksonomiya" title="Taksonomiya – Central Bikol" lang="bcl" hreflang="bcl" data-title="Taksonomiya" data-language-autonym="Bikol Central" data-language-local-name="Central Bikol" class="interlanguage-link-target"><span>Bikol Central</span></a></li><li class="interlanguage-link interwiki-ca badge-Q17437796 badge-featuredarticle mw-list-item" title="featured article badge"><a href="https://ca.wikipedia.org/wiki/Taxonomia" title="Taxonomia – Catalan" lang="ca" hreflang="ca" data-title="Taxonomia" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Taxonomie_(biologie)" title="Taxonomie (biologie) – Czech" lang="cs" hreflang="cs" data-title="Taxonomie (biologie)" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-sn mw-list-item"><a href="https://sn.wikipedia.org/wiki/Upangamutowo" title="Upangamutowo – Shona" lang="sn" hreflang="sn" data-title="Upangamutowo" data-language-autonym="ChiShona" data-language-local-name="Shona" class="interlanguage-link-target"><span>ChiShona</span></a></li><li class="interlanguage-link interwiki-de badge-Q70894304 mw-list-item" title=""><a href="https://de.wikipedia.org/wiki/Taxonomie_(Biologie)" title="Taxonomie (Biologie) – German" lang="de" hreflang="de" data-title="Taxonomie (Biologie)" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Taxonom%C3%ADa" title="Taxonomía – Spanish" lang="es" hreflang="es" data-title="Taxonomía" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Taksonomio" title="Taksonomio – Esperanto" lang="eo" hreflang="eo" data-title="Taksonomio" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Taxonomia" title="Taxonomia – Basque" lang="eu" hreflang="eu" data-title="Taxonomia" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A2%D8%B1%D8%A7%DB%8C%D9%87%E2%80%8C%D8%B4%D9%86%D8%A7%D8%B3%DB%8C_(%D8%B2%DB%8C%D8%B3%D8%AA%E2%80%8C%D8%B4%D9%86%D8%A7%D8%B3%DB%8C)" title="آرایهشناسی (زیستشناسی) – Persian" lang="fa" hreflang="fa" data-title="آرایهشناسی (زیستشناسی)" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Taxonomie" title="Taxonomie – French" lang="fr" hreflang="fr" data-title="Taxonomie" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Tacsanoma%C3%ADocht_(bitheola%C3%ADocht)" title="Tacsanomaíocht (bitheolaíocht) – Irish" lang="ga" hreflang="ga" data-title="Tacsanomaíocht (bitheolaíocht)" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gv mw-list-item"><a href="https://gv.wikipedia.org/wiki/Rang-oardraghey_(bea-oaylleeaght)" title="Rang-oardraghey (bea-oaylleeaght) – Manx" lang="gv" hreflang="gv" data-title="Rang-oardraghey (bea-oaylleeaght)" data-language-autonym="Gaelg" data-language-local-name="Manx" class="interlanguage-link-target"><span>Gaelg</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Taxonom%C3%ADa_(biolox%C3%ADa)" title="Taxonomía (bioloxía) – Galician" lang="gl" hreflang="gl" data-title="Taxonomía (bioloxía)" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-gu mw-list-item"><a href="https://gu.wikipedia.org/wiki/%E0%AA%B5%E0%AA%B0%E0%AB%8D%E0%AA%97%E0%AB%80%E0%AA%95%E0%AB%83%E0%AA%A4%E0%AA%A8%E0%AA%BE%E0%AA%AE%E0%AA%95%E0%AA%B0%E0%AA%A3_(%E0%AA%9C%E0%AB%80%E0%AA%B5%E0%AA%B5%E0%AA%BF%E0%AA%9C%E0%AB%8D%E0%AA%9E%E0%AA%BE%E0%AA%A8)" title="વર્ગીકૃતનામકરણ (જીવવિજ્ઞાન) – Gujarati" lang="gu" hreflang="gu" data-title="વર્ગીકૃતનામકરણ (જીવવિજ્ઞાન)" data-language-autonym="ગુજરાતી" data-language-local-name="Gujarati" class="interlanguage-link-target"><span>ગુજરાતી</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%B6%84%EB%A5%98%ED%95%99" title="분류학 – Korean" lang="ko" hreflang="ko" data-title="분류학" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D4%BF%D5%A5%D5%B6%D5%BD%D5%A1%D5%A2%D5%A1%D5%B6%D5%A1%D5%AF%D5%A1%D5%B6_%D5%A4%D5%A1%D5%BD%D5%A1%D5%AF%D5%A1%D6%80%D5%A3%D5%B8%D6%82%D5%B4" title="Կենսաբանական դասակարգում – Armenian" lang="hy" hreflang="hy" data-title="Կենսաբանական դասակարգում" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-hi badge-Q70893996 mw-list-item" title=""><a href="https://hi.wikipedia.org/wiki/%E0%A4%B5%E0%A4%B0%E0%A5%8D%E0%A4%97%E0%A4%BF%E0%A4%95%E0%A5%80_(%E0%A4%9C%E0%A5%80%E0%A4%B5%E0%A4%B5%E0%A4%BF%E0%A4%9C%E0%A5%8D%E0%A4%9E%E0%A4%BE%E0%A4%A8)" title="वर्गिकी (जीवविज्ञान) – Hindi" lang="hi" hreflang="hi" data-title="वर्गिकी (जीवविज्ञान)" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Taksonomija_(biologija)" title="Taksonomija (biologija) – Croatian" lang="hr" hreflang="hr" data-title="Taksonomija (biologija)" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-ilo mw-list-item"><a href="https://ilo.wikipedia.org/wiki/Taksonomia_(biolohia)" title="Taksonomia (biolohia) – Iloko" lang="ilo" hreflang="ilo" data-title="Taksonomia (biolohia)" data-language-autonym="Ilokano" data-language-local-name="Iloko" class="interlanguage-link-target"><span>Ilokano</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Taksonomi_(biologi)" title="Taksonomi (biologi) – Indonesian" lang="id" hreflang="id" data-title="Taksonomi (biologi)" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-zu mw-list-item"><a href="https://zu.wikipedia.org/wiki/Ingqwangqwa_(kumchazampilo)" title="Ingqwangqwa (kumchazampilo) – Zulu" lang="zu" hreflang="zu" data-title="Ingqwangqwa (kumchazampilo)" data-language-autonym="IsiZulu" data-language-local-name="Zulu" class="interlanguage-link-target"><span>IsiZulu</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%98%D7%A7%D7%A1%D7%95%D7%A0%D7%95%D7%9E%D7%99%D7%94_(%D7%91%D7%99%D7%95%D7%9C%D7%95%D7%92%D7%99%D7%94)" title="טקסונומיה (ביולוגיה) – Hebrew" lang="he" hreflang="he" data-title="טקסונומיה (ביולוגיה)" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Taksonomi_(biologi)" title="Taksonomi (biologi) – Javanese" lang="jv" hreflang="jv" data-title="Taksonomi (biologi)" data-language-autonym="Jawa" data-language-local-name="Javanese" class="interlanguage-link-target"><span>Jawa</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%9F%E0%B3%8D%E0%B2%AF%E0%B2%BE%E0%B2%95%E0%B3%8D%E0%B2%B8%E0%B2%BE%E0%B2%A8%E0%B2%AE%E0%B2%BF" title="ಟ್ಯಾಕ್ಸಾನಮಿ – Kannada" lang="kn" hreflang="kn" data-title="ಟ್ಯಾಕ್ಸಾನಮಿ" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-ks mw-list-item"><a href="https://ks.wikipedia.org/wiki/%D8%B1%D9%95%DA%98%D9%88%D9%B2%D9%86%D8%A0_(%D8%B2%DB%8C%D9%9B%D9%88%D9%97%D8%AF%D9%8F%D8%AA)" title="رٕژوٲنؠ (زیٛوٗدُت) – Kashmiri" lang="ks" hreflang="ks" data-title="رٕژوٲنؠ (زیٛوٗدُت)" data-language-autonym="कॉशुर / کٲشُر" data-language-local-name="Kashmiri" class="interlanguage-link-target"><span>कॉशुर / کٲشُر</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A2%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%BE%D0%BC%D0%B8%D1%8F_(%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F)" title="Таксономия (биология) – Kazakh" lang="kk" hreflang="kk" data-title="Таксономия (биология)" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ku mw-list-item"><a href="https://ku.wikipedia.org/wiki/Taksonom%C3%AE_(biyoloj%C3%AE)" title="Taksonomî (biyolojî) – Kurdish" lang="ku" hreflang="ku" data-title="Taksonomî (biyolojî)" data-language-autonym="Kurdî" data-language-local-name="Kurdish" class="interlanguage-link-target"><span>Kurdî</span></a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%91%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F%D0%BB%D1%8B%D0%BA_%D0%BA%D0%BB%D0%B0%D1%81%D1%81%D0%B8%D1%84%D0%B8%D0%BA%D0%B0%D1%86%D0%B8%D1%8F" title="Биологиялык классификация – Kyrgyz" lang="ky" hreflang="ky" data-title="Биологиялык классификация" data-language-autonym="Кыргызча" data-language-local-name="Kyrgyz" class="interlanguage-link-target"><span>Кыргызча</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Taksonomija" title="Taksonomija – Latvian" lang="lv" hreflang="lv" data-title="Taksonomija" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Taksonomija" title="Taksonomija – Lithuanian" lang="lt" hreflang="lt" data-title="Taksonomija" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-nia mw-list-item"><a href="https://nia.wikipedia.org/wiki/Taksonomi_(biologi)" title="Taksonomi (biologi) – Nias" lang="nia" hreflang="nia" data-title="Taksonomi (biologi)" data-language-autonym="Li Niha" data-language-local-name="Nias" class="interlanguage-link-target"><span>Li Niha</span></a></li><li class="interlanguage-link interwiki-lij mw-list-item"><a href="https://lij.wikipedia.org/wiki/Taxonom%C3%AFa" title="Taxonomïa – Ligurian" lang="lij" hreflang="lij" data-title="Taxonomïa" data-language-autonym="Ligure" data-language-local-name="Ligurian" class="interlanguage-link-target"><span>Ligure</span></a></li><li class="interlanguage-link interwiki-li mw-list-item"><a href="https://li.wikipedia.org/wiki/Taxonomie" title="Taxonomie – Limburgish" lang="li" hreflang="li" data-title="Taxonomie" data-language-autonym="Limburgs" data-language-local-name="Limburgish" class="interlanguage-link-target"><span>Limburgs</span></a></li><li class="interlanguage-link interwiki-lfn mw-list-item"><a href="https://lfn.wikipedia.org/wiki/Tasonomia" title="Tasonomia – Lingua Franca Nova" lang="lfn" hreflang="lfn" data-title="Tasonomia" data-language-autonym="Lingua Franca Nova" data-language-local-name="Lingua Franca Nova" class="interlanguage-link-target"><span>Lingua Franca Nova</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Rendszertan_(biol%C3%B3gia)" title="Rendszertan (biológia) – Hungarian" lang="hu" hreflang="hu" data-title="Rendszertan (biológia)" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mai mw-list-item"><a href="https://mai.wikipedia.org/wiki/%E0%A4%9F%E0%A5%8D%E0%A4%AF%E0%A4%BE%E0%A4%95%E0%A5%8D%E0%A4%B8%E0%A5%8B%E0%A4%A8%E0%A5%8B%E0%A4%AE%E0%A5%80_(%E0%A4%9C%E0%A5%80%E0%A4%B5_%E0%A4%B5%E0%A4%BF%E0%A4%9C%E0%A5%8D%E0%A4%9E%E0%A4%BE%E0%A4%A8)" title="ट्याक्सोनोमी (जीव विज्ञान) – Maithili" lang="mai" hreflang="mai" data-title="ट्याक्सोनोमी (जीव विज्ञान)" data-language-autonym="मैथिली" data-language-local-name="Maithili" class="interlanguage-link-target"><span>मैथिली</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%A2%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%BE%D0%BC%D0%B8%D1%98%D0%B0" title="Таксономија – Macedonian" lang="mk" hreflang="mk" data-title="Таксономија" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%9C%E0%B5%88%E0%B4%B5%E0%B4%B5%E0%B5%BC%E0%B4%97%E0%B5%8D%E0%B4%97%E0%B5%80%E0%B4%95%E0%B4%B0%E0%B4%A3%E0%B4%B6%E0%B4%BE%E0%B4%B8%E0%B5%8D%E0%B4%A4%E0%B5%8D%E0%B4%B0%E0%B4%82" title="ജൈവവർഗ്ഗീകരണശാസ്ത്രം – Malayalam" lang="ml" hreflang="ml" data-title="ജൈവവർഗ്ഗീകരണശാസ്ത്രം" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Taksonomi_(biologi)" title="Taksonomi (biologi) – Malay" lang="ms" hreflang="ms" data-title="Taksonomi (biologi)" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-my mw-list-item"><a href="https://my.wikipedia.org/wiki/%E1%80%99%E1%80%BB%E1%80%AD%E1%80%AF%E1%80%B8%E1%80%81%E1%80%BD%E1%80%B2%E1%80%97%E1%80%B1%E1%80%92" title="မျိုးခွဲဗေဒ – Burmese" lang="my" hreflang="my" data-title="မျိုးခွဲဗေဒ" data-language-autonym="မြန်မာဘာသာ" data-language-local-name="Burmese" class="interlanguage-link-target"><span>မြန်မာဘာသာ</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Taxonomie_(biologie)" title="Taxonomie (biologie) – Dutch" lang="nl" hreflang="nl" data-title="Taxonomie (biologie)" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ne mw-list-item"><a href="https://ne.wikipedia.org/wiki/%E0%A4%AA%E0%A5%8D%E0%A4%B0%E0%A4%BE%E0%A4%A3%E0%A5%80_%E0%A4%B5%E0%A4%B0%E0%A5%8D%E0%A4%97%E0%A5%80%E0%A4%95%E0%A4%B0%E0%A4%A3_%E0%A4%B5%E0%A4%BF%E0%A4%9C%E0%A5%8D%E0%A4%9E%E0%A4%BE%E0%A4%A8" title="प्राणी वर्गीकरण विज्ञान – Nepali" lang="ne" hreflang="ne" data-title="प्राणी वर्गीकरण विज्ञान" data-language-autonym="नेपाली" data-language-local-name="Nepali" class="interlanguage-link-target"><span>नेपाली</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%88%86%E9%A1%9E%E5%AD%A6" title="分類学 – Japanese" lang="ja" hreflang="ja" data-title="分類学" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-frr mw-list-item"><a href="https://frr.wikipedia.org/wiki/Taksonomii_faan_diarten_an_plaanten" title="Taksonomii faan diarten an plaanten – Northern Frisian" lang="frr" hreflang="frr" data-title="Taksonomii faan diarten an plaanten" data-language-autonym="Nordfriisk" data-language-local-name="Northern Frisian" class="interlanguage-link-target"><span>Nordfriisk</span></a></li><li class="interlanguage-link interwiki-om mw-list-item"><a href="https://om.wikipedia.org/wiki/Ramaddamee(Taxonomy)" title="Ramaddamee(Taxonomy) – Oromo" lang="om" hreflang="om" data-title="Ramaddamee(Taxonomy)" data-language-autonym="Oromoo" data-language-local-name="Oromo" class="interlanguage-link-target"><span>Oromoo</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Taksonomiya" title="Taksonomiya – Uzbek" lang="uz" hreflang="uz" data-title="Taksonomiya" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pap mw-list-item"><a href="https://pap.wikipedia.org/wiki/Taksonomia" title="Taksonomia – Papiamento" lang="pap" hreflang="pap" data-title="Taksonomia" data-language-autonym="Papiamentu" data-language-local-name="Papiamento" class="interlanguage-link-target"><span>Papiamentu</span></a></li><li class="interlanguage-link interwiki-ps mw-list-item"><a href="https://ps.wikipedia.org/wiki/%DA%89%D9%84%D8%A8%D9%86%D8%AF%D9%8A)_%D9%BC%DA%A9%D8%B3%D9%88%D9%86%D9%88%D9%85%D9%8A(" title="ډلبندي) ټکسونومي( – Pashto" lang="ps" hreflang="ps" data-title="ډلبندي) ټکسونومي(" data-language-autonym="پښتو" data-language-local-name="Pashto" class="interlanguage-link-target"><span>پښتو</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Taksonomia" title="Taksonomia – Polish" lang="pl" hreflang="pl" data-title="Taksonomia" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Taxonomia" title="Taxonomia – Portuguese" lang="pt" hreflang="pt" data-title="Taxonomia" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Taxonomie_(biologie)" title="Taxonomie (biologie) – Romanian" lang="ro" hreflang="ro" data-title="Taxonomie (biologie)" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-rue mw-list-item"><a href="https://rue.wikipedia.org/wiki/%D0%A2%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%BE%D0%BC%D0%B8%D1%8F_(%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F)" title="Таксономия (биология) – Rusyn" lang="rue" hreflang="rue" data-title="Таксономия (биология)" data-language-autonym="Русиньскый" data-language-local-name="Rusyn" class="interlanguage-link-target"><span>Русиньскый</span></a></li><li class="interlanguage-link interwiki-ru badge-Q70894304 mw-list-item" title=""><a href="https://ru.wikipedia.org/wiki/%D0%91%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%87%D0%B5%D1%81%D0%BA%D0%B0%D1%8F_%D1%82%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%BE%D0%BC%D0%B8%D1%8F" title="Биологическая таксономия – Russian" lang="ru" hreflang="ru" data-title="Биологическая таксономия" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sah mw-list-item"><a href="https://sah.wikipedia.org/wiki/%D0%91%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BA%D0%BB%D0%B0%D1%81%D1%81%D0%B8%D1%84%D0%B8%D0%BA%D0%B0%D1%86%D0%B8%D1%8F%D1%82%D0%B0" title="Биология классификацията – Yakut" lang="sah" hreflang="sah" data-title="Биология классификацията" data-language-autonym="Саха тыла" data-language-local-name="Yakut" class="interlanguage-link-target"><span>Саха тыла</span></a></li><li class="interlanguage-link interwiki-nso mw-list-item"><a href="https://nso.wikipedia.org/wiki/Taxonomy_(biology)" title="Taxonomy (biology) – Northern Sotho" lang="nso" hreflang="nso" data-title="Taxonomy (biology)" data-language-autonym="Sesotho sa Leboa" data-language-local-name="Northern Sotho" class="interlanguage-link-target"><span>Sesotho sa Leboa</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Taksonomia_(biologji)" title="Taksonomia (biologji) – Albanian" lang="sq" hreflang="sq" data-title="Taksonomia (biologji)" data-language-autonym="Shqip" data-language-local-name="Albanian" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-scn mw-list-item"><a href="https://scn.wikipedia.org/wiki/Tassinum%C3%ACa" title="Tassinumìa – Sicilian" lang="scn" hreflang="scn" data-title="Tassinumìa" data-language-autonym="Sicilianu" data-language-local-name="Sicilian" class="interlanguage-link-target"><span>Sicilianu</span></a></li><li class="interlanguage-link interwiki-si mw-list-item"><a href="https://si.wikipedia.org/wiki/%E0%B7%80%E0%B6%BB%E0%B7%8A%E0%B6%9C%E0%B7%93%E0%B6%9A%E0%B6%BB%E0%B6%AB%E0%B6%BA_(%E0%B6%A2%E0%B7%93%E0%B7%80_%E0%B7%80%E0%B7%92%E0%B6%AF%E0%B7%8A%E2%80%8D%E0%B6%BA%E0%B7%8F%E0%B7%80)" title="වර්ගීකරණය (ජීව විද්යාව) – Sinhala" lang="si" hreflang="si" data-title="වර්ගීකරණය (ජීව විද්යාව)" data-language-autonym="සිංහල" data-language-local-name="Sinhala" class="interlanguage-link-target"><span>සිංහල</span></a></li><li class="interlanguage-link interwiki-sd mw-list-item"><a href="https://sd.wikipedia.org/wiki/%D9%BD%D9%8A%DA%AA%D8%B3%D8%A7%D9%86%D9%88%D9%85%D9%8A" title="ٽيڪسانومي – Sindhi" lang="sd" hreflang="sd" data-title="ٽيڪسانومي" data-language-autonym="سنڌي" data-language-local-name="Sindhi" class="interlanguage-link-target"><span>سنڌي</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Taxon%C3%B3mia_(biol%C3%B3gia)" title="Taxonómia (biológia) – Slovak" lang="sk" hreflang="sk" data-title="Taxonómia (biológia)" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%A2%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%BE%D0%BC%D0%B8%D1%98%D0%B0_(%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%98%D0%B0)" title="Таксономија (биологија) – Serbian" lang="sr" hreflang="sr" data-title="Таксономија (биологија)" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Taksonomija" title="Taksonomija – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Taksonomija" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-su mw-list-item"><a href="https://su.wikipedia.org/wiki/Klasifikasi_ilmiah" title="Klasifikasi ilmiah – Sundanese" lang="su" hreflang="su" data-title="Klasifikasi ilmiah" data-language-autonym="Sunda" data-language-local-name="Sundanese" class="interlanguage-link-target"><span>Sunda</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Taksonomia" title="Taksonomia – Finnish" lang="fi" hreflang="fi" data-title="Taksonomia" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Taxonomi" title="Taxonomi – Swedish" lang="sv" hreflang="sv" data-title="Taxonomi" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tl mw-list-item"><a href="https://tl.wikipedia.org/wiki/Taksonomiya" title="Taksonomiya – Tagalog" lang="tl" hreflang="tl" data-title="Taksonomiya" data-language-autonym="Tagalog" data-language-local-name="Tagalog" class="interlanguage-link-target"><span>Tagalog</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%89%E0%AE%AF%E0%AE%BF%E0%AE%B0%E0%AE%BF%E0%AE%AF%E0%AE%B2%E0%AF%8D_%E0%AE%B5%E0%AE%95%E0%AF%88%E0%AE%AA%E0%AF%8D%E0%AE%AA%E0%AE%BE%E0%AE%9F%E0%AF%81" title="உயிரியல் வகைப்பாடு – Tamil" lang="ta" hreflang="ta" data-title="உயிரியல் வகைப்பாடு" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-shi mw-list-item"><a href="https://shi.wikipedia.org/wiki/Tasnmalayt" title="Tasnmalayt – Tachelhit" lang="shi" hreflang="shi" data-title="Tasnmalayt" data-language-autonym="Taclḥit" data-language-local-name="Tachelhit" class="interlanguage-link-target"><span>Taclḥit</span></a></li><li class="interlanguage-link interwiki-kab mw-list-item"><a href="https://kab.wikipedia.org/wiki/Tasensartut_(tasnudert)" title="Tasensartut (tasnudert) – Kabyle" lang="kab" hreflang="kab" data-title="Tasensartut (tasnudert)" data-language-autonym="Taqbaylit" data-language-local-name="Kabyle" class="interlanguage-link-target"><span>Taqbaylit</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a href="https://te.wikipedia.org/wiki/%E0%B0%9F%E0%B0%BE%E0%B0%95%E0%B1%8D%E0%B0%B8%E0%B0%BE%E0%B0%A8%E0%B0%AE%E0%B1%80_(%E0%B0%AC%E0%B0%AF%E0%B0%BE%E0%B0%B2%E0%B0%9C%E0%B1%80)" title="టాక్సానమీ (బయాలజీ) – Telugu" lang="te" hreflang="te" data-title="టాక్సానమీ (బయాలజీ)" data-language-autonym="తెలుగు" data-language-local-name="Telugu" class="interlanguage-link-target"><span>తెలుగు</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B8%AD%E0%B8%99%E0%B8%B8%E0%B8%81%E0%B8%A3%E0%B8%A1%E0%B8%A7%E0%B8%B4%E0%B8%98%E0%B8%B2%E0%B8%99" title="อนุกรมวิธาน – Thai" lang="th" hreflang="th" data-title="อนุกรมวิธาน" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-chr mw-list-item"><a href="https://chr.wikipedia.org/wiki/%E1%8F%8D%E1%8E%AA%E1%8F%AF_%E1%8F%97%E1%8F%83%E1%8E%B7%E1%8F%A9%E1%8F%98%E1%8F%8D%E1%8E%A9" title="ᏍᎪᏯ ᏗᏃᎷᏩᏘᏍᎩ – Cherokee" lang="chr" hreflang="chr" data-title="ᏍᎪᏯ ᏗᏃᎷᏩᏘᏍᎩ" data-language-autonym="ᏣᎳᎩ" data-language-local-name="Cherokee" class="interlanguage-link-target"><span>ᏣᎳᎩ</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Taksonomi_(biyoloji)" title="Taksonomi (biyoloji) – Turkish" lang="tr" hreflang="tr" data-title="Taksonomi (biyoloji)" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A2%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%BE%D0%BC%D1%96%D1%8F_(%D0%B1%D1%96%D0%BE%D0%BB%D0%BE%D0%B3%D1%96%D1%8F)" title="Таксономія (біологія) – Ukrainian" lang="uk" hreflang="uk" data-title="Таксономія (біологія)" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur badge-Q70893996 mw-list-item" title=""><a href="https://ur.wikipedia.org/wiki/%D8%AA%D8%B5%D9%86%DB%8C%D9%81%DB%8C%D8%A7%D8%AA_(%D8%AD%DB%8C%D8%A7%D8%AA%DB%8C%D8%A7%D8%AA)" title="تصنیفیات (حیاتیات) – Urdu" lang="ur" hreflang="ur" data-title="تصنیفیات (حیاتیات)" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Ph%C3%A2n_lo%E1%BA%A1i_sinh_h%E1%BB%8Dc" title="Phân loại sinh học – Vietnamese" lang="vi" hreflang="vi" data-title="Phân loại sinh học" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-vls mw-list-item"><a href="https://vls.wikipedia.org/wiki/Taxonomie_(biologie)" title="Taxonomie (biologie) – West Flemish" lang="vls" hreflang="vls" data-title="Taxonomie (biologie)" data-language-autonym="West-Vlams" data-language-local-name="West Flemish" class="interlanguage-link-target"><span>West-Vlams</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Taxonomiya_(biyolohiya)" title="Taxonomiya (biyolohiya) – Waray" lang="war" hreflang="war" data-title="Taxonomiya (biyolohiya)" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E7%94%9F%E7%89%A9%E5%88%86%E7%B1%BB%E5%AD%A6" title="生物分类学 – Wu" lang="wuu" hreflang="wuu" data-title="生物分类学" data-language-autonym="吴语" data-language-local-name="Wu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E7%94%9F%E7%89%A9%E5%88%86%E9%A1%9E%E5%AD%B8" title="生物分類學 – Cantonese" lang="yue" hreflang="yue" data-title="生物分類學" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E7%94%9F%E7%89%A9%E5%88%86%E7%B1%BB%E5%AD%A6" title="生物分类学 – Chinese" lang="zh" hreflang="zh" data-title="生物分类学" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-btm mw-list-item"><a href="https://btm.wikipedia.org/wiki/Klasifikasi_Biologi" title="Klasifikasi Biologi – Batak Mandailing" lang="btm" hreflang="btm" data-title="Klasifikasi Biologi" data-language-autonym="Batak Mandailing" data-language-local-name="Batak Mandailing" class="interlanguage-link-target"><span>Batak Mandailing</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q8269924#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div 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style="background:#90ee90;padding:0.2em;font-size:175%;font-weight:bold"><a href="/wiki/Evolutionary_biology" title="Evolutionary biology">Evolutionary biology</a></th></tr><tr><td class="sidebar-image"><span class="notpageimage" typeof="mw:File"><a href="/wiki/File:Darwin%27s_finches_by_Gould.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Darwin%27s_finches_by_Gould.jpg/180px-Darwin%27s_finches_by_Gould.jpg" decoding="async" width="180" height="136" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Darwin%27s_finches_by_Gould.jpg/270px-Darwin%27s_finches_by_Gould.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Darwin%27s_finches_by_Gould.jpg/360px-Darwin%27s_finches_by_Gould.jpg 2x" data-file-width="1748" data-file-height="1319" /></a></span><div class="sidebar-caption"><a href="/wiki/Darwin%27s_finches" title="Darwin's finches">Darwin's finches</a> by <a href="/wiki/John_Gould" title="John Gould">John Gould</a></div></td></tr><tr><td class="sidebar-above" style="display:block; margin-bottom:0.55em; background-color: transparent;"> <div class="hlist"> <ul><li><a href="/wiki/Index_of_evolutionary_biology_articles" title="Index of evolutionary biology articles">Index</a></li> <li><a href="/wiki/Introduction_to_evolution" title="Introduction to evolution">Introduction</a></li> <li><a href="/wiki/Evolution" title="Evolution">Main</a></li> <li><a href="/wiki/Outline_of_evolution" title="Outline of evolution">Outline</a></li></ul> <div class="hlist"> </div> <ul><li><a href="/wiki/Glossary_of_evolutionary_biology" class="mw-redirect" title="Glossary of evolutionary biology">Glossary</a></li> <li><a href="/wiki/Evidence_of_common_descent" title="Evidence of common descent">Evidence</a></li> <li><a href="/wiki/History_of_life" title="History of life">History</a></li></ul> </div></td></tr><tr><td class="sidebar-content" style="background:transparent;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90ee90;font-size:100%;font-weight:bold;;color: var(--color-base)">Processes and outcomes</div><div class="sidebar-list-content mw-collapsible-content plainlist"><div class="hlist"> <ul><li><a href="/wiki/Population_genetics" title="Population genetics">Population genetics</a></li> <li><a href="/wiki/Genetic_variation" title="Genetic variation">Variation</a></li> <li><a href="/wiki/Genetic_diversity" title="Genetic diversity">Diversity</a></li> <li><a href="/wiki/Mutation" title="Mutation">Mutation</a></li> <li><a href="/wiki/Natural_selection" title="Natural selection">Natural selection</a></li> <li><a href="/wiki/Adaptation" title="Adaptation">Adaptation</a></li> <li><a href="/wiki/Polymorphism_(biology)" title="Polymorphism (biology)">Polymorphism</a></li> <li><a href="/wiki/Genetic_drift" title="Genetic drift">Genetic drift</a></li> <li><a href="/wiki/Gene_flow" title="Gene flow">Gene flow</a></li> <li><a href="/wiki/Speciation" title="Speciation">Speciation</a></li> <li><a href="/wiki/Adaptive_radiation" title="Adaptive radiation">Adaptive radiation</a></li> <li><a href="/wiki/Cooperation_(evolution)" title="Cooperation (evolution)">Co-operation</a></li> <li><a href="/wiki/Coevolution" title="Coevolution">Coevolution</a></li> <li><a href="/wiki/Coextinction" title="Coextinction">Coextinction</a></li> <li><a href="/wiki/Contingency_(evolutionary_biology)" title="Contingency (evolutionary biology)">Contingency</a></li> <li><a href="/wiki/Divergent_evolution" title="Divergent evolution">Divergence</a></li> <li><a href="/wiki/Convergent_evolution" title="Convergent evolution">Convergence</a></li> <li><a href="/wiki/Parallel_evolution" title="Parallel evolution">Parallel evolution</a></li> <li><a href="/wiki/Extinction" title="Extinction">Extinction</a></li></ul> </div></div></div></td> </tr><tr><td class="sidebar-content" style="background:transparent;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90ee90;font-size:100%;font-weight:bold;;color: var(--color-base)">Natural history</div><div class="sidebar-list-content mw-collapsible-content plainlist"><div class="hlist"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Origin of life</a></li> <li><a href="/wiki/Common_descent" title="Common descent">Common descent</a></li> <li><a href="/wiki/History_of_life" title="History of life">History of life</a></li> <li><a href="/wiki/Timeline_of_the_evolutionary_history_of_life" title="Timeline of the evolutionary history of life">Timeline of evolution</a></li> <li><a href="/wiki/Human_evolution" title="Human evolution">Human evolution</a> <ul><li><a href="/wiki/Recent_human_evolution" title="Recent human evolution">Recent human evolution</a></li></ul></li> <li><a href="/wiki/Phylogenetic_tree" title="Phylogenetic tree">Phylogeny</a></li> <li><a href="/wiki/Biodiversity" title="Biodiversity">Biodiversity</a></li> <li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a class="mw-selflink selflink">Classification</a></li> <li><a href="/wiki/Evolutionary_taxonomy" title="Evolutionary taxonomy">Evolutionary taxonomy</a></li> <li><a href="/wiki/Cladistics" title="Cladistics">Cladistics</a></li> <li><a href="/wiki/Transitional_fossil" title="Transitional fossil">Transitional fossil</a></li> <li><a href="/wiki/Extinction_event" title="Extinction event">Extinction event</a></li></ul> </div></div></div></td> </tr><tr><td class="sidebar-content" style="background:transparent;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90ee90;font-size:100%;font-weight:bold;;color: var(--color-base)">History of evolutionary theory</div><div class="sidebar-list-content mw-collapsible-content plainlist"><div class="hlist"> <ul><li><a href="/wiki/History_of_evolutionary_thought" title="History of evolutionary thought">Overview</a></li> <li><a href="/wiki/Evolutionary_ideas_of_the_Renaissance_and_Enlightenment" title="Evolutionary ideas of the Renaissance and Enlightenment">Renaissance</a></li> <li><a href="/wiki/Transmutation_of_species" title="Transmutation of species">Before Darwin</a></li> <li><a href="/wiki/Charles_Darwin" title="Charles Darwin">Darwin</a></li> <li><i><a href="/wiki/On_the_Origin_of_Species" title="On the Origin of Species">Origin of Species</a></i></li> <li><a href="/wiki/The_eclipse_of_Darwinism" title="The eclipse of Darwinism">Before synthesis</a></li> <li><a href="/wiki/Modern_synthesis_(20th_century)" title="Modern synthesis (20th century)">Modern synthesis</a></li> <li><a href="/wiki/History_of_molecular_evolution" title="History of molecular evolution">Molecular evolution</a></li> <li><a href="/wiki/Evolutionary_developmental_biology" title="Evolutionary developmental biology">Evo-devo</a></li> <li><a href="/wiki/Current_research_in_evolutionary_biology" class="mw-redirect" title="Current research in evolutionary biology">Current research</a></li> <li><a href="/wiki/History_of_speciation" title="History of speciation">History of speciation</a></li> <li><a href="/wiki/History_of_paleontology" title="History of paleontology">History of paleontology</a> (<a href="/wiki/Timeline_of_paleontology" title="Timeline of paleontology">timeline</a>)</li></ul> </div></div></div></td> </tr><tr><td class="sidebar-content" style="background:transparent;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90ee90;font-size:100%;font-weight:bold;;color: var(--color-base)">Fields and applications</div><div class="sidebar-list-content mw-collapsible-content plainlist"> <ul><li><a href="/wiki/Applications_of_evolution" title="Applications of evolution">Applications of evolution</a></li> <li><a href="/wiki/Biosocial_criminology" title="Biosocial criminology">Biosocial criminology</a></li> <li><a href="/wiki/Ecological_genetics" title="Ecological genetics">Ecological genetics</a></li> <li><a href="/wiki/Evolutionary_aesthetics" title="Evolutionary aesthetics">Evolutionary aesthetics</a></li> <li><a href="/wiki/Evolutionary_anthropology" title="Evolutionary anthropology">Evolutionary anthropology</a></li> <li><a href="/wiki/Evolutionary_computation" title="Evolutionary computation">Evolutionary computation</a></li> <li><a href="/wiki/Evolutionary_ecology" title="Evolutionary ecology">Evolutionary ecology</a></li> <li><a href="/wiki/Evolutionary_economics" title="Evolutionary economics">Evolutionary economics</a></li> <li><a href="/wiki/Evolutionary_epistemology" title="Evolutionary epistemology">Evolutionary epistemology</a></li> <li><a href="/wiki/Evolutionary_ethics" title="Evolutionary ethics">Evolutionary ethics</a></li> 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breeding</a></li> <li><a href="/wiki/Laboratory_experiments_of_speciation" title="Laboratory experiments of speciation">Speciation experiments</a></li> <li><a href="/wiki/Sociobiology" title="Sociobiology">Sociobiology</a></li> <li><a href="/wiki/Island_biogeography" class="mw-redirect" title="Island biogeography">Island biogeography</a></li> <li><a href="/wiki/Systematics" title="Systematics">Systematics</a></li> <li><a href="/wiki/Universal_Darwinism" title="Universal Darwinism">Universal Darwinism</a></li></ul></div></div></td> </tr><tr><td class="sidebar-content" style="background:transparent;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90ee90;font-size:100%;font-weight:bold;;color: var(--color-base)">Social implications</div><div class="sidebar-list-content mw-collapsible-content plainlist"> <ul><li><a href="/wiki/Eugenics" title="Eugenics">Eugenics</a></li> <li><a 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class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Tree_of_life.svg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/09/Tree_of_life.svg/16px-Tree_of_life.svg.png" decoding="async" width="16" height="13" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/09/Tree_of_life.svg/24px-Tree_of_life.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/09/Tree_of_life.svg/32px-Tree_of_life.svg.png 2x" data-file-width="915" data-file-height="760" /></a></span> </span><a href="/wiki/Portal:Evolutionary_biology" title="Portal:Evolutionary biology">Evolutionary biology portal</a></span></li> <li><span class="nowrap"><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Evolutionary_biology" title="Category:Evolutionary biology">Category</a></span></li></ul></td></tr><tr><td class="sidebar-navbar"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Evolutionary_biology" title="Template:Evolutionary biology"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Evolutionary_biology" title="Template talk:Evolutionary biology"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Evolutionary_biology" title="Special:EditPage/Template:Evolutionary biology"><abbr title="Edit this template">e</abbr></a></li></ul></div></td></tr></tbody></table> <p>In <a href="/wiki/Biology" title="Biology">biology</a>, <b>taxonomy</b> (from <a href="/wiki/Ancient_Greek_language" class="mw-redirect" title="Ancient Greek language">Ancient Greek</a> <i> </i><a href="https://en.wiktionary.org/wiki/%CF%84%CE%AC%CE%BE%CE%B9%CF%82" class="extiw" title="wikt:τάξις">τάξις</a><i> (<a href="/wiki/Taxis" title="Taxis">taxis</a>)</i> 'arrangement' and <i> </i><a href="https://en.wiktionary.org/wiki/%CE%BD%CF%8C%CE%BC%CE%BF%CF%82" class="extiw" title="wikt:νόμος">-νομία</a><i> (<a href="https://en.wiktionary.org/wiki/-nomy" class="extiw" title="wikt:-nomy">-nomia</a>)</i> '<a href="/wiki/Scientific_method" title="Scientific method">method</a>') is the <a href="/wiki/Science" title="Science">scientific</a> study of naming, defining (<a href="/wiki/Circumscription_(taxonomy)" title="Circumscription (taxonomy)">circumscribing</a>) and classifying groups of biological <a href="/wiki/Organism" title="Organism">organisms</a> based on shared characteristics. Organisms are grouped into <a href="/wiki/Taxon" title="Taxon">taxa</a> (singular: taxon) and these groups are given a <a href="/wiki/Taxonomic_rank" title="Taxonomic rank">taxonomic rank</a>; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are <a href="/wiki/Domain_(biology)" title="Domain (biology)">domain</a>, <a href="/wiki/Kingdom_(biology)" title="Kingdom (biology)">kingdom</a>, <a href="/wiki/Phylum" title="Phylum">phylum</a> (<i>division</i> is sometimes used in botany in place of <i>phylum</i>), <a href="/wiki/Class_(biology)" title="Class (biology)">class</a>, <a href="/wiki/Order_(biology)" title="Order (biology)">order</a>, <a href="/wiki/Family_(biology)" title="Family (biology)">family</a>, <a href="/wiki/Genus" title="Genus">genus</a>, and <a href="/wiki/Species" title="Species">species</a>. The Swedish botanist <a href="/wiki/Carl_Linnaeus" title="Carl Linnaeus">Carl Linnaeus</a> is regarded as the founder of the current system of taxonomy, as he developed a ranked system known as <a href="/wiki/Linnaean_taxonomy" title="Linnaean taxonomy">Linnaean taxonomy</a> for categorizing organisms and <a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">binomial nomenclature</a> for naming organisms. </p><p>With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transformed into a system of modern biological classification intended to reflect the <a href="/wiki/Evolution" title="Evolution">evolutionary</a> relationships among organisms, both living and extinct. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Definition">Definition</h2></div> <p>The exact definition of taxonomy varies from source to source, but the core of the discipline remains: the conception, naming, and classification of groups of organisms.<sup id="cite_ref-Wilkins2011_1-0" class="reference"><a href="#cite_note-Wilkins2011-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> As points of reference, recent definitions of taxonomy are presented below: </p> <ol><li>Theory and practice of grouping individuals into species, arranging species into larger groups, and giving those groups names, thus producing a classification.<sup id="cite_ref-Judd_2-0" class="reference"><a href="#cite_note-Judd-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li>A field of science (and a major component of <a href="/wiki/Systematics" title="Systematics">systematics</a>) that encompasses description, identification, <a href="/wiki/Nomenclature" title="Nomenclature">nomenclature</a>, and classification<sup id="cite_ref-Simpson_3-0" class="reference"><a href="#cite_note-Simpson-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></li> <li>The science of classification, in biology the arrangement of organisms into a classification<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li>"The science of classification as applied to living organisms, including the study of means of formation of species, etc."<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li> <li>"The analysis of an organism's characteristics for the purpose of classification"<sup id="cite_ref-Henderson_6-0" class="reference"><a href="#cite_note-Henderson-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></li> <li>"Systematics studies <a href="/wiki/Phylogeny" class="mw-redirect" title="Phylogeny">phylogeny</a> to provide a pattern that can be translated into the classification and names of the more inclusive field of taxonomy" (listed as a desirable but unusual definition)<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup></li></ol> <p>The varied definitions either place taxonomy as a sub-area of systematics (definition 2), invert that relationship (definition 6), or appear to consider the two terms synonymous. There is some disagreement as to whether <a href="/wiki/Biological_nomenclature" class="mw-redirect" title="Biological nomenclature">biological nomenclature</a> is considered a part of taxonomy (definitions 1 and 2), or a part of systematics outside taxonomy.<sup id="cite_ref-Herbarium_8-0" class="reference"><a href="#cite_note-Herbarium-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Laurin_2023_Advent_9-0" class="reference"><a href="#cite_note-Laurin_2023_Advent-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> For example, definition 6 is paired with the following definition of systematics that places nomenclature outside taxonomy:<sup id="cite_ref-Henderson_6-1" class="reference"><a href="#cite_note-Henderson-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <ul><li><i>Systematics</i>: "The study of the identification, taxonomy, and nomenclature of organisms, including the classification of living things with regard to their natural relationships and the study of variation and the evolution of taxa".</li></ul> <p>In 1970, Michener <i>et al.</i> defined "systematic biology" and "taxonomy" (terms that are often confused and used interchangeably) in relation to one another as follows:<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <blockquote> <p>Systematic biology (hereafter called simply systematics) is the field that (a) provides scientific names for organisms, (b) describes them, (c) preserves collections of them, (d) provides classifications for the organisms, keys for their identification, and data on their distributions, (e) investigates their evolutionary histories, and (f) considers their environmental adaptations. This is a field with a long history that in recent years has experienced a notable renaissance, principally with respect to theoretical content. Part of the theoretical material has to do with evolutionary areas (topics e and f above), the rest relates especially to the problem of classification. Taxonomy is that part of Systematics concerned with topics (a) to (d) above. </p> </blockquote> <p>A whole set of terms including taxonomy, systematic biology, <a href="/wiki/Systematics" title="Systematics">systematics</a>, scientific classification, biological classification, and <a href="/wiki/Phylogenetics" title="Phylogenetics">phylogenetics</a> have at times had overlapping meanings – sometimes the same, sometimes slightly different, but always related and intersecting.<sup id="cite_ref-Wilkins2011_1-1" class="reference"><a href="#cite_note-Wilkins2011-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Small1989_11-0" class="reference"><a href="#cite_note-Small1989-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> The broadest meaning of "taxonomy" is used here. The term itself was introduced in 1813 by <a href="/wiki/Augustin_Pyramus_de_Candolle" title="Augustin Pyramus de Candolle">de Candolle</a>, in his <i><a href="/wiki/Th%C3%A9orie_%C3%A9l%C3%A9mentaire_de_la_botanique" class="mw-redirect" title="Théorie élémentaire de la botanique">Théorie élémentaire de la botanique</a></i>.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> <a href="/wiki/John_Lindley" title="John Lindley">John Lindley</a> provided an early definition of systematics in 1830, although he wrote of "systematic botany" rather than using the term "systematics".<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Europeans tend to use the terms "systematics" and "biosystematics" for the study of biodiversity as a whole, whereas North Americans tend to use "taxonomy" more frequently.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> However, taxonomy, and in particular <a href="/wiki/Alpha_taxonomy" class="mw-redirect" title="Alpha taxonomy">alpha taxonomy</a>, is more specifically the identification, description, and naming (i.e., nomenclature) of organisms,<sup id="cite_ref-Fortey_15-0" class="reference"><a href="#cite_note-Fortey-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> while "classification" focuses on placing organisms within hierarchical groups that show their relationships to other organisms. </p> <div class="mw-heading mw-heading3"><h3 id="Monograph_and_taxonomic_revision">Monograph and taxonomic revision</h3></div> <p>A <b>taxonomic revision</b> or <b>taxonomic review</b> is a novel analysis of the variation patterns in a particular <a href="/wiki/Taxon" title="Taxon">taxon</a>. This analysis may be executed on the basis of any combination of the various available kinds of characters, such as morphological, <a href="/wiki/Anatomical" class="mw-redirect" title="Anatomical">anatomical</a>, <a href="/wiki/Palynological" class="mw-redirect" title="Palynological">palynological</a>, <a href="/wiki/Biochemistry" title="Biochemistry">biochemical</a> and <a href="/wiki/Genetics" title="Genetics">genetic</a>. A <a href="/wiki/Monograph#In_biology" title="Monograph">monograph</a> or complete revision is a revision that is comprehensive for a taxon for the information given at a particular time, and for the entire world. Other (partial) revisions may be restricted in the sense that they may only use some of the available character sets or have a limited spatial scope. A revision results in a conformation of or new insights in the relationships between the subtaxa within the taxon under study, which may lead to a change in the classification of these subtaxa, the identification of new subtaxa, or the merger of previous subtaxa.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Taxonomic_characters">Taxonomic characters</h3></div> <p>Taxonomic characters are the taxonomic attributes that can be used to provide the evidence from which relationships (the <a href="/wiki/Phylogeny" class="mw-redirect" title="Phylogeny">phylogeny</a>) between taxa are inferred.<sup id="cite_ref-Hennig_1965_17-0" class="reference"><a href="#cite_note-Hennig_1965-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> Kinds of taxonomic characters include:<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> </p> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 35em;"> <ul><li><a href="/wiki/Morphology_(biology)" title="Morphology (biology)">Morphological</a> characters <ul><li>General external <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphology</a></li> <li>Special structures (e.g., <a href="/wiki/Sex_organ" title="Sex organ">genitalia</a>)</li> <li>Internal morphology (<a href="/wiki/Anatomy" title="Anatomy">anatomy</a>)</li> <li><a href="/wiki/Embryology" title="Embryology">Embryology</a></li> <li><a href="/wiki/Karyotype" title="Karyotype">Karyology</a> and other <a href="/wiki/Cell_biology" title="Cell biology">cytological</a> factors</li></ul></li> <li><a href="/wiki/Physiology" title="Physiology">Physiological</a> characters <ul><li><a href="/wiki/Metabolism" title="Metabolism">Metabolic factors</a></li> <li>Body secretions</li> <li>Genic sterility factors</li></ul></li> <li><a href="/wiki/Molecular_biology" title="Molecular biology">Molecular</a> characters <ul><li>Immunological distance</li> <li>Electrophoretic differences</li> <li>Amino acid sequences of proteins</li> <li>DNA hybridization</li> <li>DNA and RNA sequences</li> <li>Restriction endonuclease analyses</li> <li>Other molecular differences</li></ul></li> <li><a href="/wiki/Behavior" title="Behavior">Behavioral</a> characters <ul><li>Courtship and other ethological isolating mechanisms</li> <li>Other behavior patterns</li></ul></li> <li><a href="/wiki/Ecology" title="Ecology">Ecological</a> characters <ul><li>Habit and habitats</li> <li>Food</li> <li>Seasonal variations</li> <li>Parasites and hosts</li></ul></li> <li><a href="/wiki/Geography" title="Geography">Geographic</a> characters <ul><li>General <a href="/wiki/Species_distribution" title="Species distribution">biogeographic distribution</a> patterns</li> <li><a href="/wiki/Sympatry" title="Sympatry">Sympatric</a>-<a href="/wiki/Allopatric_speciation" title="Allopatric speciation">allopatric</a> relationship of populations</li></ul></li></ul> </div> <div class="mw-heading mw-heading3"><h3 id="Alpha_and_beta_taxonomy">Alpha and beta taxonomy</h3></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Not to be confused with <a href="/wiki/Alpha_diversity" title="Alpha diversity">Alpha diversity</a>.</div> <p>The term "<b>alpha taxonomy</b>" is primarily used to refer to the discipline of finding, describing, and naming <a href="/wiki/Taxon" title="Taxon">taxa</a>, particularly species.<sup id="cite_ref-BiologyDiscussion_20-0" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> In earlier literature, the term had a different meaning, referring to morphological taxonomy, and the products of research through the end of the 19th century.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> </p> <p><a href="/wiki/William_Bertram_Turrill" title="William Bertram Turrill">William Bertram Turrill</a> introduced the term "alpha taxonomy" in a series of papers published in 1935 and 1937 in which he discussed the philosophy and possible future directions of the discipline of taxonomy.<sup id="cite_ref-FOOTNOTETurrill1938_22-0" class="reference"><a href="#cite_note-FOOTNOTETurrill1938-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup></p><blockquote><p> ... there is an increasing desire amongst taxonomists to consider their problems from wider viewpoints, to investigate the possibilities of closer co-operation with their cytological, ecological and genetics colleagues and to acknowledge that some revision or expansion, perhaps of a drastic nature, of their aims and methods, may be desirable ... Turrill (1935) has suggested that while accepting the older invaluable taxonomy, based on structure, and conveniently designated "alpha", it is possible to glimpse a far-distant taxonomy built upon as wide a basis of morphological and physiological facts as possible, and one in which "place is found for all observational and experimental data relating, even if indirectly, to the constitution, subdivision, origin, and behaviour of species and other taxonomic groups". Ideals can, it may be said, never be completely realized. They have, however, a great value of acting as permanent stimulants, and if we have some, even vague, ideal of an "omega" taxonomy we may progress a little way down the Greek alphabet. Some of us please ourselves by thinking we are now groping in a "beta" taxonomy.<sup id="cite_ref-FOOTNOTETurrill1938_22-1" class="reference"><a href="#cite_note-FOOTNOTETurrill1938-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup></p></blockquote> <p>Turrill thus explicitly excludes from alpha taxonomy various areas of study that he includes within taxonomy as a whole, such as ecology, physiology, genetics, and cytology. He further excludes phylogenetic reconstruction from alpha taxonomy.<sup id="cite_ref-FOOTNOTETurrill1938365–366_23-0" class="reference"><a href="#cite_note-FOOTNOTETurrill1938365–366-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p><p> Later authors have used the term in a different sense, to mean the delimitation of species (not subspecies or taxa of other ranks), using whatever investigative techniques are available, and including sophisticated computational or laboratory techniques.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-BiologyDiscussion_20-1" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> Thus, <a href="/wiki/Ernst_Mayr" title="Ernst Mayr">Ernst Mayr</a> in 1968 defined "<b>beta taxonomy</b>" as the classification of ranks higher than species.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup></p><blockquote><p>An understanding of the biological meaning of variation and of the evolutionary origin of groups of related species is even more important for the second stage of taxonomic activity, the sorting of species into groups of relatives ("taxa") and their arrangement in a hierarchy of higher categories. This activity is what the term classification denotes; it is also referred to as "beta taxonomy".</p></blockquote> <div class="mw-heading mw-heading3"><h3 id="Microtaxonomy_and_macrotaxonomy">Microtaxonomy and macrotaxonomy</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Species_problem" class="mw-redirect" title="Species problem">Species problem</a></div> <p>How species should be defined in a particular group of organisms gives rise to practical and theoretical problems that are referred to as the <a href="/wiki/Species_problem" class="mw-redirect" title="Species problem">species problem</a>. The scientific work of deciding how to define species has been called microtaxonomy.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-BiologyDiscussion_20-2" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> By extension, macrotaxonomy is the study of groups at the higher <a href="/wiki/Taxonomic_rank" title="Taxonomic rank">taxonomic ranks</a> subgenus and above,<sup id="cite_ref-BiologyDiscussion_20-3" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> or simply in clades that include more than one taxon considered a species, expressed in terms of <a href="/wiki/Phylogenetic_nomenclature" title="Phylogenetic nomenclature">phylogenetic nomenclature</a>.<sup id="cite_ref-Cantino_&_de_Queiroz_2020_28-0" class="reference"><a href="#cite_note-Cantino_&_de_Queiroz_2020-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="History">History</h2></div> <p>While some descriptions of taxonomic history attempt to date taxonomy to ancient civilizations, a truly scientific attempt to classify organisms did not occur until the 18th century, with the possible exception of Aristotle, whose works hint at a taxonomy.<sup id="cite_ref-Voultsiadou_&_Vafidis_2007_29-0" class="reference"><a href="#cite_note-Voultsiadou_&_Vafidis_2007-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Voultsiadou_et_al._2017_30-0" class="reference"><a href="#cite_note-Voultsiadou_et_al._2017-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> Earlier works were primarily descriptive and focused on plants that were useful in agriculture or medicine. </p><p>There are a number of stages in this scientific thinking. Early taxonomy was based on arbitrary criteria, the so-called "artificial systems", including <a href="/wiki/Carl_Linnaeus" title="Carl Linnaeus">Linnaeus</a>'s system of sexual classification for plants (Linnaeus's 1735 classification of animals was entitled "<a href="/wiki/Systema_Naturae" title="Systema Naturae">Systema Naturae</a>" ("the System of Nature"), implying that he, at least, believed that it was more than an "artificial system"). </p><p>Later came systems based on a more complete consideration of the characteristics of taxa, referred to as "natural systems", such as those of <a href="/wiki/Antoine_Laurent_de_Jussieu" title="Antoine Laurent de Jussieu">de Jussieu</a> (1789), de Candolle (1813) and <a href="/wiki/Bentham_and_Hooker" class="mw-redirect" title="Bentham and Hooker">Bentham and Hooker</a> (1862–1863). These classifications described empirical patterns and were pre-<a href="/wiki/Evolution" title="Evolution">evolutionary</a> in thinking. </p><p>The publication of <a href="/wiki/Charles_Darwin" title="Charles Darwin">Charles Darwin</a>'s <i><a href="/wiki/On_the_Origin_of_Species" title="On the Origin of Species">On the Origin of Species</a></i> (1859) led to a new explanation for classifications, based on evolutionary relationships. This was the concept of <a href="/wiki/Phyletic" class="mw-redirect" title="Phyletic">phyletic</a> systems, from 1883 onwards. This approach was typified by those of <a href="/wiki/August_Eichler" class="mw-redirect" title="August Eichler">Eichler</a> (1883) and <a href="/wiki/Adolf_Engler" title="Adolf Engler">Engler</a> (1886–1892). </p><p>The advent of <a href="/wiki/Cladistic" class="mw-redirect" title="Cladistic">cladistic</a> methodology in the 1970s led to classifications based on the sole criterion of <a href="/wiki/Monophyly" title="Monophyly">monophyly</a>, supported by the presence of <a href="/wiki/Synapomorphies" class="mw-redirect" title="Synapomorphies">synapomorphies</a>. Since then, the evidentiary basis has been expanded with data from <a href="/wiki/Molecular_genetics" title="Molecular genetics">molecular genetics</a> that for the most part complements traditional <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphology</a>.<sup id="cite_ref-FOOTNOTEDatta1988_31-0" class="reference"><a href="#cite_note-FOOTNOTEDatta1988-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup><sup class="noprint Inline-Template" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citing_sources" title="Wikipedia:Citing sources"><span title="This citation requires a reference to the specific page or range of pages in which the material appears. (July 2019)">page needed</span></a></i>]</sup><sup id="cite_ref-FOOTNOTEStace1989_32-0" class="reference"><a href="#cite_note-FOOTNOTEStace1989-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup><sup class="noprint Inline-Template" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citing_sources" title="Wikipedia:Citing sources"><span title="This citation requires a reference to the specific page or range of pages in which the material appears. (July 2019)">page needed</span></a></i>]</sup><sup id="cite_ref-FOOTNOTEStuessy2009_33-0" class="reference"><a href="#cite_note-FOOTNOTEStuessy2009-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup><sup class="noprint Inline-Template" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citing_sources" title="Wikipedia:Citing sources"><span title="This citation requires a reference to the specific page or range of pages in which the material appears. (July 2019)">page needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading3"><h3 id="Pre-Linnaean">Pre-Linnaean</h3></div> <div class="mw-heading mw-heading4"><h4 id="Early_taxonomists">Early taxonomists</h4></div> <p>Naming and classifying human surroundings likely began with the onset of language. Distinguishing poisonous plants from edible plants is integral to the survival of human communities. Medicinal plant illustrations show up in Egyptian wall paintings from <abbr title="circa">c.</abbr><span style="white-space:nowrap;"> 1500 BC</span>, indicating that the uses of different species were understood and that a basic taxonomy was in place.<sup id="cite_ref-Manktelow_34-0" class="reference"><a href="#cite_note-Manktelow-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Ancient_times">Ancient times</h4></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Aristotle%27s_biology#Classification" title="Aristotle's biology">Aristotle's biology § Classification</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Huang-Quan-Xie-sheng-zhen-qin-tu.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/b/bf/Huang-Quan-Xie-sheng-zhen-qin-tu.jpg/370px-Huang-Quan-Xie-sheng-zhen-qin-tu.jpg" decoding="async" width="370" height="217" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/bf/Huang-Quan-Xie-sheng-zhen-qin-tu.jpg/555px-Huang-Quan-Xie-sheng-zhen-qin-tu.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/bf/Huang-Quan-Xie-sheng-zhen-qin-tu.jpg/740px-Huang-Quan-Xie-sheng-zhen-qin-tu.jpg 2x" data-file-width="6826" data-file-height="4000" /></a><figcaption><i>Description of rare animals</i> (写生珍禽图), by <a href="/wiki/Song_dynasty" title="Song dynasty">Song dynasty</a> painter <a href="/wiki/Huang_Quan_(painter)" title="Huang Quan (painter)">Huang Quan</a> (903–965)</figcaption></figure> <p>Organisms were first classified by <a href="/wiki/Aristotle" title="Aristotle">Aristotle</a> (<a href="/wiki/Greece" title="Greece">Greece</a>, 384–322 BC) during his stay on the <a href="/wiki/Lesbos" title="Lesbos">Island of Lesbos</a>.<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Palaeos_36-0" class="reference"><a href="#cite_note-Palaeos-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Britannica_–_Taxonomy_37-0" class="reference"><a href="#cite_note-Britannica_–_Taxonomy-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> He classified beings by their parts, or in modern terms <i>attributes</i>, such as having live birth, having four legs, laying eggs, having blood, or being warm-bodied.<sup id="cite_ref-Roanoke_38-0" class="reference"><a href="#cite_note-Roanoke-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> He divided all living things into two groups: <a href="/wiki/Plant" title="Plant">plants</a> and <a href="/wiki/Animal" title="Animal">animals</a>.<sup id="cite_ref-Palaeos_36-1" class="reference"><a href="#cite_note-Palaeos-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p><p>Some of his groups of animals, such as <i>Anhaima</i> (animals without blood, translated as <a href="/wiki/Invertebrate" title="Invertebrate">invertebrates</a>) and <i>Enhaima</i> (animals with blood, roughly the <a href="/wiki/Vertebrate" title="Vertebrate">vertebrates</a>), as well as groups like the <a href="/wiki/Shark" title="Shark">sharks</a> and <a href="/wiki/Cetacean" class="mw-redirect" title="Cetacean">cetaceans</a>, are commonly used.<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">[</span>39<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-von_Lieven_&_Humar_2008_40-0" class="reference"><a href="#cite_note-von_Lieven_&_Humar_2008-40"><span class="cite-bracket">[</span>40<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Laurin_&_Humar_2022_41-0" class="reference"><a href="#cite_note-Laurin_&_Humar_2022-41"><span class="cite-bracket">[</span>41<span class="cite-bracket">]</span></a></sup> </p><p>His student <a href="/wiki/Theophrastus" title="Theophrastus">Theophrastus</a> (Greece, 370–285 BC) carried on this tradition, mentioning some 500 plants and their uses in his <i><a href="/wiki/Historia_Plantarum_(Theophrastus)" title="Historia Plantarum (Theophrastus)">Historia Plantarum</a></i>. Several plant <a href="/wiki/Genus" title="Genus">genera</a> can be traced back to Theophrastus, such as <i><a href="/wiki/Cornus_(genus)" class="mw-redirect" title="Cornus (genus)">Cornus</a></i>, <i><a href="/wiki/Crocus" title="Crocus">Crocus</a></i>, and <i><a href="/wiki/Narcissus_(plant)" title="Narcissus (plant)">Narcissus</a></i>.<sup id="cite_ref-Palaeos_36-2" class="reference"><a href="#cite_note-Palaeos-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Medieval">Medieval</h4></div> <p>Taxonomy in the <a href="/wiki/Middle_Ages" title="Middle Ages">Middle Ages</a> was largely based on the <a href="/wiki/Aristotelian_system" class="mw-redirect" title="Aristotelian system">Aristotelian system</a>,<sup id="cite_ref-Roanoke_38-1" class="reference"><a href="#cite_note-Roanoke-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> with additions concerning the philosophical and existential order of creatures. This included concepts such as the <a href="/wiki/Great_chain_of_being" title="Great chain of being">great chain of being</a> in the Western <a href="/wiki/Scholasticism" title="Scholasticism">scholastic</a> tradition,<sup id="cite_ref-Roanoke_38-2" class="reference"><a href="#cite_note-Roanoke-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> again deriving ultimately from Aristotle. </p><p>The Aristotelian system did not classify plants or <a href="/wiki/Fungi" class="mw-redirect" title="Fungi">fungi</a>, due to the lack of microscopes at the time,<sup id="cite_ref-Britannica_–_Taxonomy_37-1" class="reference"><a href="#cite_note-Britannica_–_Taxonomy-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> as his ideas were based on arranging the complete world in a single continuum, as per the <i>scala naturae</i> (the Natural Ladder).<sup id="cite_ref-Palaeos_36-3" class="reference"><a href="#cite_note-Palaeos-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> This, as well, was taken into consideration in the great chain of being.<sup id="cite_ref-Palaeos_36-4" class="reference"><a href="#cite_note-Palaeos-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p><p>Advances were made by scholars such as <a href="/wiki/Procopius" title="Procopius">Procopius</a>, <a href="/wiki/Timotheus_of_Gaza" title="Timotheus of Gaza">Timotheus of Gaza</a>, <a href="/wiki/Demetrios_Pepagomenos" title="Demetrios Pepagomenos">Demetrios Pepagomenos</a>, and <a href="/wiki/Thomas_Aquinas" title="Thomas Aquinas">Thomas Aquinas</a>. Medieval thinkers used abstract philosophical and logical categorizations more suited to abstract philosophy than to pragmatic taxonomy.<sup id="cite_ref-Palaeos_36-5" class="reference"><a href="#cite_note-Palaeos-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Renaissance_and_early_modern">Renaissance and early modern</h4></div> <p>During the <a href="/wiki/Renaissance" title="Renaissance">Renaissance</a> and the <a href="/wiki/Age_of_Enlightenment" title="Age of Enlightenment">Age of Enlightenment</a>, categorizing organisms became more prevalent,<sup id="cite_ref-Palaeos_36-6" class="reference"><a href="#cite_note-Palaeos-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> and taxonomic works became ambitious enough to replace the ancient texts. This is sometimes credited to the development of sophisticated optical lenses, which allowed the morphology of organisms to be studied in much greater detail. </p><p>One of the earliest authors to take advantage of this leap in technology was the Italian physician <a href="/wiki/Andrea_Cesalpino" title="Andrea Cesalpino">Andrea Cesalpino</a> (1519–1603), who has been called "the first taxonomist".<sup id="cite_ref-42" class="reference"><a href="#cite_note-42"><span class="cite-bracket">[</span>42<span class="cite-bracket">]</span></a></sup> His <a href="/wiki/Masterpiece" title="Masterpiece">magnum opus</a> <i>De Plantis</i> came out in 1583, and described more than 1500 plant species.<sup id="cite_ref-43" class="reference"><a href="#cite_note-43"><span class="cite-bracket">[</span>43<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-44" class="reference"><a href="#cite_note-44"><span class="cite-bracket">[</span>44<span class="cite-bracket">]</span></a></sup> Two large plant families that he first recognized are in use: the <a href="/wiki/Asteraceae" title="Asteraceae">Asteraceae</a> and <a href="/wiki/Brassicaceae" title="Brassicaceae">Brassicaceae</a>.<sup id="cite_ref-45" class="reference"><a href="#cite_note-45"><span class="cite-bracket">[</span>45<span class="cite-bracket">]</span></a></sup> </p><p>In the 17th century <a href="/wiki/John_Ray" title="John Ray">John Ray</a> (<a href="/wiki/England" title="England">England</a>, 1627–1705) wrote many important taxonomic works.<sup id="cite_ref-Britannica_–_Taxonomy_37-2" class="reference"><a href="#cite_note-Britannica_–_Taxonomy-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> Arguably his greatest accomplishment was <i>Methodus Plantarum Nova</i> (1682),<sup id="cite_ref-46" class="reference"><a href="#cite_note-46"><span class="cite-bracket">[</span>46<span class="cite-bracket">]</span></a></sup> in which he published details of over 18,000 plant species. At the time, his classifications were perhaps the most complex yet produced by any taxonomist, as he based his taxa on many combined characters. </p><p>The next major taxonomic works were produced by <a href="/wiki/Joseph_Pitton_de_Tournefort" title="Joseph Pitton de Tournefort">Joseph Pitton de Tournefort</a> (France, 1656–1708).<sup id="cite_ref-47" class="reference"><a href="#cite_note-47"><span class="cite-bracket">[</span>47<span class="cite-bracket">]</span></a></sup> His work from 1700, <i>Institutiones Rei Herbariae</i>, included more than 9000 species in 698 genera, which directly influenced Linnaeus, as it was the text he used as a young student.<sup id="cite_ref-Manktelow_34-1" class="reference"><a href="#cite_note-Manktelow-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Linnaean_era">Linnaean era</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Linnaean_taxonomy" title="Linnaean taxonomy">Linnaean taxonomy</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Linn%C3%A9-Systema_Naturae_1735.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Linn%C3%A9-Systema_Naturae_1735.jpg/170px-Linn%C3%A9-Systema_Naturae_1735.jpg" decoding="async" width="170" height="213" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Linn%C3%A9-Systema_Naturae_1735.jpg/255px-Linn%C3%A9-Systema_Naturae_1735.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/54/Linn%C3%A9-Systema_Naturae_1735.jpg/340px-Linn%C3%A9-Systema_Naturae_1735.jpg 2x" data-file-width="910" data-file-height="1141" /></a><figcaption>Title page of <i><a href="/wiki/Systema_Naturae" title="Systema Naturae">Systema Naturae</a></i>, Leiden, 1735</figcaption></figure> <p>The Swedish botanist <a href="/wiki/Carl_Linnaeus" title="Carl Linnaeus">Carl Linnaeus</a> (1707–1778)<sup id="cite_ref-Roanoke_38-3" class="reference"><a href="#cite_note-Roanoke-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> ushered in a new era of taxonomy. With his major works <i><a href="/wiki/Systema_Naturae" title="Systema Naturae">Systema Naturae</a></i> 1st Edition in 1735,<sup id="cite_ref-48" class="reference"><a href="#cite_note-48"><span class="cite-bracket">[</span>48<span class="cite-bracket">]</span></a></sup> <i><a href="/wiki/Species_Plantarum" title="Species Plantarum">Species Plantarum</a></i> in 1753,<sup id="cite_ref-49" class="reference"><a href="#cite_note-49"><span class="cite-bracket">[</span>49<span class="cite-bracket">]</span></a></sup> and <a href="/wiki/10th_edition_of_Systema_Naturae" title="10th edition of Systema Naturae"><i>Systema Naturae</i> 10th Edition</a>,<sup id="cite_ref-50" class="reference"><a href="#cite_note-50"><span class="cite-bracket">[</span>50<span class="cite-bracket">]</span></a></sup> he revolutionized modern taxonomy. His works implemented a standardized binomial naming system for animal and plant species,<sup id="cite_ref-Britannica_–_Linnaean_51-0" class="reference"><a href="#cite_note-Britannica_–_Linnaean-51"><span class="cite-bracket">[</span>51<span class="cite-bracket">]</span></a></sup> which proved to be an elegant solution to a chaotic and disorganized taxonomic literature. He not only introduced the standard of class, order, genus, and species, but also made it possible to identify plants and animals from his book, by using the smaller parts of the flower (known as the <a href="/wiki/Linnaean_system" class="mw-redirect" title="Linnaean system">Linnaean system</a>).<sup id="cite_ref-Britannica_–_Linnaean_51-1" class="reference"><a href="#cite_note-Britannica_–_Linnaean-51"><span class="cite-bracket">[</span>51<span class="cite-bracket">]</span></a></sup> </p><p>Plant and animal taxonomists regard Linnaeus' work as the "starting point" for valid names (at 1753 and 1758 respectively).<sup id="cite_ref-52" class="reference"><a href="#cite_note-52"><span class="cite-bracket">[</span>52<span class="cite-bracket">]</span></a></sup> Names published before these dates are referred to as "pre-Linnaean", and not considered valid (with the exception of spiders published in <i><a href="/wiki/Svenska_Spindlar" title="Svenska Spindlar">Svenska Spindlar</a></i><sup id="cite_ref-53" class="reference"><a href="#cite_note-53"><span class="cite-bracket">[</span>53<span class="cite-bracket">]</span></a></sup>). Even taxonomic names published by Linnaeus himself before these dates are considered pre-Linnaean.<sup id="cite_ref-Manktelow_34-2" class="reference"><a href="#cite_note-Manktelow-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="The_digital_era_of_taxonomy">The digital era of taxonomy</h3></div> <p>Modern taxonomy is heavily influenced by technology such as <a href="/wiki/DNA_sequencing" title="DNA sequencing">DNA sequencing</a>, <a href="/wiki/Bioinformatics" title="Bioinformatics">bioinformatics</a>, <a href="/wiki/Biological_database" title="Biological database">databases</a>, and <a href="/wiki/Imaging" title="Imaging">imaging</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Modern_system_of_classification">Modern system of classification</h2></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main articles: <a href="/wiki/Evolutionary_taxonomy" title="Evolutionary taxonomy">Evolutionary taxonomy</a> and <a href="/wiki/Phylogenetic_nomenclature" title="Phylogenetic nomenclature">Phylogenetic nomenclature</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Spindle_diagram.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/76/Spindle_diagram.jpg/280px-Spindle_diagram.jpg" decoding="async" width="280" height="194" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/76/Spindle_diagram.jpg/420px-Spindle_diagram.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/76/Spindle_diagram.jpg/560px-Spindle_diagram.jpg 2x" data-file-width="734" data-file-height="509" /></a><figcaption>Evolution of the <a href="/wiki/Vertebrates" class="mw-redirect" title="Vertebrates">vertebrates</a> at class level, width of spindles indicating number of families. Spindle diagrams are typical for <a href="/wiki/Evolutionary_taxonomy" title="Evolutionary taxonomy">evolutionary taxonomy</a>.</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Cladogram_vertebrata.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Cladogram_vertebrata.jpg/280px-Cladogram_vertebrata.jpg" decoding="async" width="280" height="129" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Cladogram_vertebrata.jpg/420px-Cladogram_vertebrata.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Cladogram_vertebrata.jpg/560px-Cladogram_vertebrata.jpg 2x" data-file-width="734" data-file-height="337" /></a><figcaption>The same relationship, expressed as a <a href="/wiki/Cladogram" title="Cladogram">cladogram</a> typical for <a href="/wiki/Cladistics" title="Cladistics">cladistics</a></figcaption></figure> <p>A pattern of groups nested within groups was specified by Linnaeus' classifications of plants and animals, and these patterns began to be represented as <a href="/wiki/Dendrograms" class="mw-redirect" title="Dendrograms">dendrograms</a> of the animal and plant <a href="/wiki/Kingdom_(biology)" title="Kingdom (biology)">kingdoms</a> toward the end of the 18th century, well before Charles Darwin's <i>On the Origin of Species</i> was published.<sup id="cite_ref-Britannica_–_Taxonomy_37-3" class="reference"><a href="#cite_note-Britannica_–_Taxonomy-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> The pattern of the "Natural System" did not entail a generating process, such as evolution, but may have implied it, inspiring early transmutationist thinkers. Among early works exploring the idea of a <a href="/wiki/Transmutation_of_species" title="Transmutation of species">transmutation of species</a> were <i><a href="/wiki/Zoonomia" title="Zoonomia">Zoonomia</a></i> in 1796 by <a href="/wiki/Erasmus_Darwin" title="Erasmus Darwin">Erasmus Darwin</a> (Charles Darwin's grandfather), and <a href="/wiki/Jean-Baptiste_Lamarck" title="Jean-Baptiste Lamarck">Jean-Baptiste Lamarck</a>'s <i><a href="/wiki/Philosophie_zoologique" title="Philosophie zoologique">Philosophie zoologique</a></i> of 1809.<sup id="cite_ref-BiologyDiscussion_20-4" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> The idea was popularized in the Anglophone world by the speculative but widely read <i><a href="/wiki/Vestiges_of_the_Natural_History_of_Creation" title="Vestiges of the Natural History of Creation">Vestiges of the Natural History of Creation</a></i>, published anonymously by <a href="/wiki/Robert_Chambers_(publisher,_born_1802)" title="Robert Chambers (publisher, born 1802)">Robert Chambers</a> in 1844.<sup id="cite_ref-54" class="reference"><a href="#cite_note-54"><span class="cite-bracket">[</span>54<span class="cite-bracket">]</span></a></sup> </p><p>With Darwin's theory, a general acceptance quickly appeared that a classification should reflect the Darwinian principle of <a href="/wiki/Common_descent" title="Common descent">common descent</a>.<sup id="cite_ref-Britannica_–_since_Linnaeus_55-0" class="reference"><a href="#cite_note-Britannica_–_since_Linnaeus-55"><span class="cite-bracket">[</span>55<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Tree_of_life_(science)" class="mw-redirect" title="Tree of life (science)">Tree of life</a> representations became popular in scientific works, with known fossil groups incorporated. One of the first modern groups tied to fossil ancestors was birds.<sup id="cite_ref-56" class="reference"><a href="#cite_note-56"><span class="cite-bracket">[</span>56<span class="cite-bracket">]</span></a></sup> Using the then newly discovered fossils of <i><a href="/wiki/Archaeopteryx" title="Archaeopteryx">Archaeopteryx</a></i> and <i><a href="/wiki/Hesperornis" title="Hesperornis">Hesperornis</a></i>, <a href="/wiki/Thomas_Henry_Huxley" title="Thomas Henry Huxley">Thomas Henry Huxley</a> pronounced that they had evolved from dinosaurs, a group formally named by <a href="/wiki/Richard_Owen" title="Richard Owen">Richard Owen</a> in 1842.<sup id="cite_ref-57" class="reference"><a href="#cite_note-57"><span class="cite-bracket">[</span>57<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-58" class="reference"><a href="#cite_note-58"><span class="cite-bracket">[</span>58<span class="cite-bracket">]</span></a></sup> The resulting description, that of dinosaurs "giving rise to" or being "the ancestors of" birds, is the essential hallmark of <a href="/wiki/Evolutionary_taxonomy" title="Evolutionary taxonomy">evolutionary taxonomic</a> thinking. As more and more fossil groups were found and recognized in the late 19th and early 20th centuries, <a href="/wiki/Paleontology" title="Paleontology">palaeontologists</a> worked to understand the history of animals through the ages by linking together known groups.<sup id="cite_ref-59" class="reference"><a href="#cite_note-59"><span class="cite-bracket">[</span>59<span class="cite-bracket">]</span></a></sup> With the <a href="/wiki/Modern_synthesis_(20th_century)" title="Modern synthesis (20th century)">modern evolutionary synthesis</a> of the early 1940s, an essentially modern understanding of the evolution of the major groups was in place. As evolutionary taxonomy is based on Linnaean taxonomic ranks, the two terms are largely interchangeable in modern use.<sup id="cite_ref-60" class="reference"><a href="#cite_note-60"><span class="cite-bracket">[</span>60<span class="cite-bracket">]</span></a></sup> </p><p>The <a href="/wiki/Cladistic" class="mw-redirect" title="Cladistic">cladistic</a> method has emerged since the 1960s.<sup id="cite_ref-Britannica_–_since_Linnaeus_55-1" class="reference"><a href="#cite_note-Britannica_–_since_Linnaeus-55"><span class="cite-bracket">[</span>55<span class="cite-bracket">]</span></a></sup> In 1958, <a href="/wiki/Julian_Huxley" title="Julian Huxley">Julian Huxley</a> used the term <i>clade</i>.<sup id="cite_ref-BiologyDiscussion_20-5" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> Later, in 1960, Cain and Harrison introduced the term <i>cladistic</i>.<sup id="cite_ref-BiologyDiscussion_20-6" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> The salient feature is arranging taxa in a hierarchical <a href="/wiki/Evolutionary_tree" class="mw-redirect" title="Evolutionary tree">evolutionary tree</a>, with the desideratum that all named taxa are monophyletic.<sup id="cite_ref-Britannica_–_since_Linnaeus_55-2" class="reference"><a href="#cite_note-Britannica_–_since_Linnaeus-55"><span class="cite-bracket">[</span>55<span class="cite-bracket">]</span></a></sup> A taxon is called monophyletic if it includes all the descendants of an ancestral form.<sup id="cite_ref-Mike_Taylor_61-0" class="reference"><a href="#cite_note-Mike_Taylor-61"><span class="cite-bracket">[</span>61<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-NCSE_62-0" class="reference"><a href="#cite_note-NCSE-62"><span class="cite-bracket">[</span>62<span class="cite-bracket">]</span></a></sup> Groups that have descendant groups removed from them are termed <a href="/wiki/Paraphyletic" class="mw-redirect" title="Paraphyletic">paraphyletic</a>,<sup id="cite_ref-Mike_Taylor_61-1" class="reference"><a href="#cite_note-Mike_Taylor-61"><span class="cite-bracket">[</span>61<span class="cite-bracket">]</span></a></sup> while groups representing more than one branch from the tree of life are called <a href="/wiki/Polyphyletic" class="mw-redirect" title="Polyphyletic">polyphyletic</a>.<sup id="cite_ref-Mike_Taylor_61-2" class="reference"><a href="#cite_note-Mike_Taylor-61"><span class="cite-bracket">[</span>61<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-NCSE_62-1" class="reference"><a href="#cite_note-NCSE-62"><span class="cite-bracket">[</span>62<span class="cite-bracket">]</span></a></sup> Monophyletic groups are recognized and diagnosed on the basis of <a href="/wiki/Synapomorphies" class="mw-redirect" title="Synapomorphies">synapomorphies</a>, shared derived character states.<sup id="cite_ref-63" class="reference"><a href="#cite_note-63"><span class="cite-bracket">[</span>63<span class="cite-bracket">]</span></a></sup> </p><p>Cladistic classifications are compatible with traditional Linnean taxonomy and the Codes of <a href="/wiki/Zoological_nomenclature" class="mw-redirect" title="Zoological nomenclature">Zoological</a> and <a href="/wiki/Botanical_nomenclature" title="Botanical nomenclature">Botanical nomenclature</a>, to a certain extent.<sup id="cite_ref-64" class="reference"><a href="#cite_note-64"><span class="cite-bracket">[</span>64<span class="cite-bracket">]</span></a></sup> An alternative system of nomenclature, the <i><a href="/wiki/PhyloCode" title="PhyloCode">International Code of Phylogenetic Nomenclature</a></i> or <i>PhyloCode</i> has been proposed, which regulates the formal naming of clades.<sup id="cite_ref-65" class="reference"><a href="#cite_note-65"><span class="cite-bracket">[</span>65<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Cantino_&_de_Queiroz_2020_28-1" class="reference"><a href="#cite_note-Cantino_&_de_Queiroz_2020-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Laurin_2023_Advent_9-1" class="reference"><a href="#cite_note-Laurin_2023_Advent-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Linnaean ranks are optional and have no formal standing under the <i>PhyloCode</i>, which is intended to coexist with the current, rank-based codes.<sup id="cite_ref-Cantino_&_de_Queiroz_2020_28-2" class="reference"><a href="#cite_note-Cantino_&_de_Queiroz_2020-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> While popularity of phylogenetic nomenclature has grown steadily in the last few decades,<sup id="cite_ref-Laurin_2023_Advent_9-2" class="reference"><a href="#cite_note-Laurin_2023_Advent-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> it remains to be seen whether a majority of systematists will eventually adopt the <i>PhyloCode</i> or continue using the current systems of nomenclature that have been employed (and modified, but arguably not as much as some systematists wish)<sup id="cite_ref-Dubois_2007_66-0" class="reference"><a href="#cite_note-Dubois_2007-66"><span class="cite-bracket">[</span>66<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Dubois_et_al._2019_67-0" class="reference"><a href="#cite_note-Dubois_et_al._2019-67"><span class="cite-bracket">[</span>67<span class="cite-bracket">]</span></a></sup> for over 250 years. </p> <div class="mw-heading mw-heading3"><h3 id="Kingdoms_and_domains">Kingdoms and domains</h3></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Biological_classification_L_Pengo_vflip.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Biological_classification_L_Pengo_vflip.svg/170px-Biological_classification_L_Pengo_vflip.svg.png" decoding="async" width="170" height="425" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Biological_classification_L_Pengo_vflip.svg/255px-Biological_classification_L_Pengo_vflip.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Biological_classification_L_Pengo_vflip.svg/340px-Biological_classification_L_Pengo_vflip.svg.png 2x" data-file-width="240" data-file-height="600" /></a><figcaption>The basic scheme of modern classification. Many other levels can be used; domain, the highest level within life, is both new and disputed.</figcaption></figure> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main articles: <a href="/wiki/Kingdom_(biology)" title="Kingdom (biology)">Kingdom (biology)</a> and <a href="/wiki/Domain_(biology)" title="Domain (biology)">Domain (biology)</a></div> <p>Well before Linnaeus, plants and animals were considered separate Kingdoms.<sup id="cite_ref-Kingdom_classification_68-0" class="reference"><a href="#cite_note-Kingdom_classification-68"><span class="cite-bracket">[</span>68<span class="cite-bracket">]</span></a></sup><sup class="noprint Inline-Template" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Reliable_sources" title="Wikipedia:Reliable sources"><span title="The material near this tag may rely on an unreliable source. (April 2017)">unreliable source?</span></a></i>]</sup> Linnaeus used this as the top rank, dividing the physical world into the vegetable, animal and mineral kingdoms. As advances in microscopy made the classification of microorganisms possible, the number of kingdoms increased, five- and six-kingdom systems being the most common. </p><p><a href="/wiki/Domain_(biology)" title="Domain (biology)">Domains</a> are a relatively new grouping. First proposed in 1977, <a href="/wiki/Carl_Woese" title="Carl Woese">Carl Woese</a>'s <a href="/wiki/Three-domain_system" title="Three-domain system">three-domain system</a> was not generally accepted until later.<sup id="cite_ref-69" class="reference"><a href="#cite_note-69"><span class="cite-bracket">[</span>69<span class="cite-bracket">]</span></a></sup> One main characteristic of the three-domain method is the separation of <a href="/wiki/Archaea" title="Archaea">Archaea</a> and <a href="/wiki/Bacteria" title="Bacteria">Bacteria</a>, previously grouped into the single kingdom Bacteria (a kingdom also sometimes called <a href="/wiki/Monera" title="Monera">Monera</a>),<sup id="cite_ref-Kingdom_classification_68-1" class="reference"><a href="#cite_note-Kingdom_classification-68"><span class="cite-bracket">[</span>68<span class="cite-bracket">]</span></a></sup> with the <a href="/wiki/Eukaryota" class="mw-redirect" title="Eukaryota">Eukaryota</a> for all organisms whose cells contain a <a href="/wiki/Nucleus_(biology)" class="mw-redirect" title="Nucleus (biology)">nucleus</a>.<sup id="cite_ref-70" class="reference"><a href="#cite_note-70"><span class="cite-bracket">[</span>70<span class="cite-bracket">]</span></a></sup> A small number of scientists include a sixth kingdom, Archaea, but do not accept the domain method.<sup id="cite_ref-Kingdom_classification_68-2" class="reference"><a href="#cite_note-Kingdom_classification-68"><span class="cite-bracket">[</span>68<span class="cite-bracket">]</span></a></sup> </p><p><a href="/wiki/Thomas_Cavalier-Smith" title="Thomas Cavalier-Smith">Thomas Cavalier-Smith</a>, who published extensively on the classification of <a href="/wiki/Protist" title="Protist">protists</a>, in 2002<sup id="cite_ref-pmid11931142_71-0" class="reference"><a href="#cite_note-pmid11931142-71"><span class="cite-bracket">[</span>71<span class="cite-bracket">]</span></a></sup> proposed that the <a href="/wiki/Neomura" title="Neomura">Neomura</a>, the clade that groups together the Archaea and <a href="/wiki/Eukaryote" title="Eukaryote">Eucarya</a>, would have evolved from Bacteria, more precisely from <a href="/wiki/Actinomycetota" title="Actinomycetota">Actinomycetota</a>. His 2004 classification treated the <a href="/wiki/Archaeobacteria" class="mw-redirect" title="Archaeobacteria">archaeobacteria</a> as part of a subkingdom of the kingdom Bacteria, i.e., he rejected the three-domain system entirely.<sup id="cite_ref-CavalierSmith2004_72-0" class="reference"><a href="#cite_note-CavalierSmith2004-72"><span class="cite-bracket">[</span>72<span class="cite-bracket">]</span></a></sup> Stefan Luketa in 2012 proposed a five "dominion" system, adding <a href="/wiki/Prion" title="Prion">Prionobiota</a> (<a href="/wiki/Non-cellular_life" title="Non-cellular life">acellular</a> and without <a href="/wiki/Nucleic_acid" title="Nucleic acid">nucleic acid</a>) and <a href="/wiki/Virus" title="Virus">Virusobiota</a> (acellular but <i>with</i> nucleic acid) to the traditional three domains.<sup id="cite_ref-Luketa2012_73-0" class="reference"><a href="#cite_note-Luketa2012-73"><span class="cite-bracket">[</span>73<span class="cite-bracket">]</span></a></sup> </p> <div class="center" style="width:auto; margin-left:auto; margin-right:auto;"> <table style="margin-left: auto; margin-right: auto; border-collapse:collapse;"> <tbody><tr> <th style="padding:4px; border-bottom:solid black 1px;"><a href="/wiki/Carl_Linnaeus" title="Carl Linnaeus">Linnaeus</a><br />1735<sup id="cite_ref-Linnaeus1735_74-0" class="reference"><a href="#cite_note-Linnaeus1735-74"><span class="cite-bracket">[</span>74<span class="cite-bracket">]</span></a></sup> </th> <th style="padding:4px; border-bottom:solid black 1px;"><a href="/wiki/Ernst_Haeckel" title="Ernst Haeckel">Haeckel</a><br />1866<sup id="cite_ref-Haeckel_75-0" class="reference"><a href="#cite_note-Haeckel-75"><span class="cite-bracket">[</span>75<span class="cite-bracket">]</span></a></sup> </th> <th style="padding:4px; border-bottom:solid black 1px;"><a href="/wiki/%C3%89douard_Chatton" title="Édouard Chatton">Chatton</a><br />1925<sup id="cite_ref-Chatton1925_76-0" class="reference"><a href="#cite_note-Chatton1925-76"><span class="cite-bracket">[</span>76<span class="cite-bracket">]</span></a></sup> </th> <th style="padding:4px; border-bottom:solid black 1px;"><a href="/wiki/Herbert_Copeland_(biologist)" title="Herbert Copeland (biologist)">Copeland</a><br />1938<sup id="cite_ref-Copeland1938_77-0" class="reference"><a href="#cite_note-Copeland1938-77"><span class="cite-bracket">[</span>77<span class="cite-bracket">]</span></a></sup> </th> <th style="padding:4px; border-bottom:solid black 1px;"><a href="/wiki/Robert_Whittaker_(ecologist)" title="Robert Whittaker (ecologist)">Whittaker</a><br />1969<sup id="cite_ref-Whittaker1969_78-0" class="reference"><a href="#cite_note-Whittaker1969-78"><span class="cite-bracket">[</span>78<span class="cite-bracket">]</span></a></sup> </th> <th style="padding:4px; border-bottom:solid black 1px;"><a href="/wiki/Carl_Woese" title="Carl Woese">Woese</a> et al.<br />1990<sup id="cite_ref-Woese1990_79-0" class="reference"><a href="#cite_note-Woese1990-79"><span class="cite-bracket">[</span>79<span class="cite-bracket">]</span></a></sup> </th> <th style="padding:4px; border-bottom:solid black 1px;"><a href="/wiki/Thomas_Cavalier-Smith" title="Thomas Cavalier-Smith">Cavalier-Smith</a><br />1998,<sup id="cite_ref-CavalierSmith2004_72-1" class="reference"><a href="#cite_note-CavalierSmith2004-72"><span class="cite-bracket">[</span>72<span class="cite-bracket">]</span></a></sup> 2015<sup id="cite_ref-80" class="reference"><a href="#cite_note-80"><span class="cite-bracket">[</span>80<span class="cite-bracket">]</span></a></sup> </th></tr> <tr> <td style="text-align:center; border-bottom:double black 3px; padding-left:4px; padding-right:4px;">2 kingdoms </td> <td style="text-align:center; border-bottom:double black 3px; border-left:solid black 1px; padding-left:4px; padding-right:4px;">3 kingdoms </td> <td style="text-align:center; border-bottom:double black 3px; border-left:solid black 1px; padding-left:4px; padding-right:4px;"><a href="/wiki/Two-empire_system" title="Two-empire system">2 empires</a> </td> <td style="text-align:center; border-bottom:double black 3px; border-left:solid black 1px; padding-left:4px; padding-right:4px;"><a href="/wiki/Kingdom_(biology)#Four_kingdoms" title="Kingdom (biology)">4 kingdoms</a> </td> <td style="text-align:center; border-bottom:double black 3px; border-left:solid black 1px; padding-left:4px; padding-right:4px;">5 kingdoms </td> <td style="text-align:center; border-bottom:double black 3px; border-left:solid black 1px; padding-left:4px; padding-right:4px;"><a href="/wiki/Three-domain_system" title="Three-domain system">3 domains</a> </td> <td style="text-align:center; border-bottom:double black 3px; border-left:solid black 1px; padding-left:4px; padding-right:4px;">2 empires,<br />6/7 kingdoms </td></tr> <tr> <td rowspan="4" style="padding-left:4px;"><i>(not treated)</i> </td> <td rowspan="4" style="background-color:khaki; padding-left:4px; padding-right:4px; border-left:dotted black 1px;"><a href="/wiki/Protist" title="Protist">Protista</a> </td> <td rowspan="2" style="background-color:lightgrey; padding-left:4px; padding-right:4px; border-left:dotted black 1px;"><a href="/wiki/Prokaryota" class="mw-redirect" title="Prokaryota">Prokaryota</a> </td> <td rowspan="2" style="background-color:lightgrey; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Monera" title="Monera">Monera</a> </td> <td rowspan="2" style="background-color:lightgrey; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Monera" title="Monera">Monera</a> </td> <td style="background-color:lightgrey; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Bacteria" title="Bacteria">Bacteria</a> </td> <td style="background-color:lightgrey; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Bacteria" title="Bacteria">Bacteria</a> </td></tr> <tr> <td style="background-color:darkgray; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Archaea" title="Archaea">Archaea</a> </td> <td style="background-color:darkgray; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Archaea" title="Archaea">Archaea</a> (2015) </td></tr> <tr> <td rowspan="5" style="background-color:#e0d0b0; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Eukaryota" class="mw-redirect" title="Eukaryota">Eukaryota</a> </td> <td rowspan="2" style="background-color:khaki; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Protista" class="mw-redirect" title="Protista">Protoctista</a> </td> <td rowspan="2" style="background-color:khaki; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Protist" title="Protist">Protista</a> </td> <td rowspan="5" style="background-color:#e0d0b0; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Eukaryote" title="Eukaryote">Eucarya</a> </td> <td rowspan="1" style="background-color:khaki; padding-left:4px; border-left:dotted black 1px;">"<a href="/wiki/Protozoa" title="Protozoa">Protozoa</a>" </td></tr> <tr> <td rowspan="1" style="background-color:burlywood; padding-left:4px; border-left:dotted black 1px;">"<a href="/wiki/Chromista" title="Chromista">Chromista</a>" </td></tr> <tr> <td rowspan="2" style="background-color:lightgreen; padding-left:4px;"><a href="/wiki/Vegetabilia" class="mw-redirect" title="Vegetabilia">Vegetabilia</a> </td> <td rowspan="2" style="background-color:lightgreen; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Plant" title="Plant">Plantae</a> </td> <td rowspan="2" style="background-color:lightgreen; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Plant" title="Plant">Plantae</a> </td> <td style="background-color:lightgreen; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Plant" title="Plant">Plantae</a> </td> <td style="background-color:lightgreen; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Plant" title="Plant">Plantae</a> </td></tr> <tr> <td style="background-color:lightblue; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Fungus" title="Fungus">Fungi</a> </td> <td style="background-color:lightblue; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Fungus" title="Fungus">Fungi</a> </td></tr> <tr> <td style="background-color:pink; padding-left:4px;"><a href="/wiki/Animal" title="Animal">Animalia</a> </td> <td style="background-color:pink; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Animal" title="Animal">Animalia</a> </td> <td style="background-color:pink; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Animal" title="Animal">Animalia</a> </td> <td style="background-color:pink; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Animal" title="Animal">Animalia</a> </td> <td style="background-color:pink; padding-left:4px; border-left:dotted black 1px;"><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr></tbody></table></div> <div class="mw-heading mw-heading3"><h3 id="Recent_comprehensive_classifications">Recent comprehensive classifications</h3></div> <p>Partial classifications exist for many individual groups of organisms and are revised and replaced as new information becomes available; however, comprehensive, published treatments of most or all life are rarer; recent examples are that of Adl et al., 2012 and 2019,<sup id="cite_ref-Adl-et-al-2012_81-0" class="reference"><a href="#cite_note-Adl-et-al-2012-81"><span class="cite-bracket">[</span>81<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Adl-et-al-2019_82-0" class="reference"><a href="#cite_note-Adl-et-al-2019-82"><span class="cite-bracket">[</span>82<span class="cite-bracket">]</span></a></sup> which covers eukaryotes only with an emphasis on protists, and Ruggiero et al., 2015,<sup id="cite_ref-Ruggiero-et-al-2015_83-0" class="reference"><a href="#cite_note-Ruggiero-et-al-2015-83"><span class="cite-bracket">[</span>83<span class="cite-bracket">]</span></a></sup> covering both eukaryotes and <a href="/wiki/Prokaryote" title="Prokaryote">prokaryotes</a> to the rank of Order, although both exclude fossil representatives.<sup id="cite_ref-Ruggiero-et-al-2015_83-1" class="reference"><a href="#cite_note-Ruggiero-et-al-2015-83"><span class="cite-bracket">[</span>83<span class="cite-bracket">]</span></a></sup> A separate compilation (Ruggiero, 2014)<sup id="cite_ref-84" class="reference"><a href="#cite_note-84"><span class="cite-bracket">[</span>84<span class="cite-bracket">]</span></a></sup> covers extant taxa to the rank of Family. Other, database-driven treatments include the <a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">Encyclopedia of Life</a>, the <a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">Global Biodiversity Information Facility</a>, the <a href="/wiki/NCBI_taxonomy_database" class="mw-redirect" title="NCBI taxonomy database">NCBI taxonomy database</a>, the <a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">Interim Register of Marine and Nonmarine Genera</a>, the <a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>, and the <a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">Catalogue of Life</a>. The <a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a> is a resource for fossils. </p> <div class="mw-heading mw-heading2"><h2 id="Application">Application</h2></div> <p>Biological taxonomy is a sub-discipline of <a href="/wiki/Biology" title="Biology">biology</a>, and is generally practiced by biologists known as "taxonomists", though enthusiastic <a href="/wiki/Natural_history" title="Natural history">naturalists</a> are also frequently involved in the publication of new taxa.<sup id="cite_ref-85" class="reference"><a href="#cite_note-85"><span class="cite-bracket">[</span>85<span class="cite-bracket">]</span></a></sup> Because taxonomy aims to describe and organize <a href="/wiki/Life" title="Life">life</a>, the work conducted by taxonomists is essential for the study of <a href="/wiki/Biodiversity" title="Biodiversity">biodiversity</a> and the resulting field of <a href="/wiki/Conservation_biology" title="Conservation biology">conservation biology</a>.<sup id="cite_ref-86" class="reference"><a href="#cite_note-86"><span class="cite-bracket">[</span>86<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-87" class="reference"><a href="#cite_note-87"><span class="cite-bracket">[</span>87<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Classifying_organisms">Classifying organisms</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Taxonomic_rank" title="Taxonomic rank">Taxonomic rank</a></div> <p>Biological classification is a critical component of the taxonomic process. As a result, it informs the user as to what the relatives of the taxon are hypothesized to be. Biological classification uses taxonomic ranks, including among others (in order from most inclusive to least inclusive): <a href="/wiki/Domain_(biology)" title="Domain (biology)">Domain</a>, <a href="/wiki/Kingdom_(biology)" title="Kingdom (biology)">Kingdom</a>, <a href="/wiki/Phylum" title="Phylum">Phylum</a>, <a href="/wiki/Class_(biology)" title="Class (biology)">Class</a>, <a href="/wiki/Order_(biology)" title="Order (biology)">Order</a>, <a href="/wiki/Family_(biology)" title="Family (biology)">Family</a>, <a href="/wiki/Genus" title="Genus">Genus</a>, <a href="/wiki/Species" title="Species">Species</a>, and <a href="/wiki/Strain_(biology)" title="Strain (biology)">Strain</a>.<sup id="cite_ref-88" class="reference"><a href="#cite_note-88"><span class="cite-bracket">[</span>88<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-89" class="reference"><a href="#cite_note-89"><span class="cite-bracket">[</span>note 1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Taxonomic_descriptions">Taxonomic descriptions</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Species_description" title="Species description">Species description</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Nepenthes_smilesii_type_specimen.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Nepenthes_smilesii_type_specimen.jpg/170px-Nepenthes_smilesii_type_specimen.jpg" decoding="async" width="170" height="227" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Nepenthes_smilesii_type_specimen.jpg/255px-Nepenthes_smilesii_type_specimen.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Nepenthes_smilesii_type_specimen.jpg/340px-Nepenthes_smilesii_type_specimen.jpg 2x" data-file-width="1944" data-file-height="2592" /></a><figcaption><a href="/wiki/Type_(biology)" title="Type (biology)">Type</a> specimen for <i><a href="/wiki/Nepenthes_smilesii" title="Nepenthes smilesii">Nepenthes smilesii</a></i>, a tropical <a href="/wiki/Pitcher_plant" title="Pitcher plant">pitcher plant</a></figcaption></figure> <p>The "definition" of a taxon is encapsulated by its description or its diagnosis or by both combined. There are no set rules governing the definition of taxa, but the naming and publication of new taxa is governed by sets of rules.<sup id="cite_ref-Herbarium_8-1" class="reference"><a href="#cite_note-Herbarium-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> In <a href="/wiki/Zoology" title="Zoology">zoology</a>, the <a href="/wiki/Nomenclature" title="Nomenclature">nomenclature</a> for the more commonly used ranks (<a href="/wiki/Superfamily_(taxonomy)" class="mw-redirect" title="Superfamily (taxonomy)">superfamily</a> to <a href="/wiki/Subspecies" title="Subspecies">subspecies</a>), is regulated by the <i><a href="/wiki/International_Code_of_Zoological_Nomenclature" title="International Code of Zoological Nomenclature">International Code of Zoological Nomenclature</a></i> (<i>ICZN Code</i>).<sup id="cite_ref-90" class="reference"><a href="#cite_note-90"><span class="cite-bracket">[</span>89<span class="cite-bracket">]</span></a></sup> In the fields of <a href="/wiki/Phycology" title="Phycology">phycology</a>, <a href="/wiki/Mycology" title="Mycology">mycology</a>, and <a href="/wiki/Botany" title="Botany">botany</a>, the naming of taxa is governed by the <i><a href="/wiki/International_Code_of_Nomenclature_for_algae,_fungi,_and_plants" title="International Code of Nomenclature for algae, fungi, and plants">International Code of Nomenclature for algae, fungi, and plants</a></i> (<i>ICN</i>).<sup id="cite_ref-91" class="reference"><a href="#cite_note-91"><span class="cite-bracket">[</span>90<span class="cite-bracket">]</span></a></sup> </p><p>The initial description of a taxon involves five main requirements:<sup id="cite_ref-92" class="reference"><a href="#cite_note-92"><span class="cite-bracket">[</span>91<span class="cite-bracket">]</span></a></sup> </p> <ol><li>The taxon must be given a name based on the 26 letters of the Latin alphabet (a <a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">binomial</a> for new species, or uninomial for other ranks).</li> <li>The name must be unique (i.e. not a <a href="/wiki/Homonym_(biology)" title="Homonym (biology)">homonym</a>).</li> <li>The description must be based on at least one name-bearing <a href="/wiki/Type_species" title="Type species">type specimen</a>.</li> <li>It should include statements about appropriate attributes either to describe (define) the taxon or to differentiate it from other taxa (the diagnosis, <i>ICZN Code</i>, Article 13.1.1, <i>ICN</i>, Article 38, which may or may not be based on morphology<sup id="cite_ref-93" class="reference"><a href="#cite_note-93"><span class="cite-bracket">[</span>92<span class="cite-bracket">]</span></a></sup>). Both codes deliberately separate defining the content of a taxon (its <a href="/wiki/Circumscription_(taxonomy)" title="Circumscription (taxonomy)">circumscription</a>) from defining its name.</li> <li>These first four requirements must be published in a work that is obtainable in numerous identical copies, as a permanent scientific record.</li></ol> <p>However, often much more information is included, like the geographic range of the taxon, ecological notes, chemistry, behavior, etc. How researchers arrive at their taxa varies: depending on the available data, and resources, methods vary from simple <a href="/wiki/Quantitative_property" class="mw-redirect" title="Quantitative property">quantitative</a> or <a href="/wiki/Qualitative_property" title="Qualitative property">qualitative</a> comparisons of striking features, to elaborate computer analyses of large amounts of <a href="/wiki/DNA_sequence" class="mw-redirect" title="DNA sequence">DNA sequence</a> data.<sup id="cite_ref-94" class="reference"><a href="#cite_note-94"><span class="cite-bracket">[</span>93<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Author_citation">Author citation</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main articles: <a href="/wiki/Author_citation_(botany)" title="Author citation (botany)">Author citation (botany)</a> and <a href="/wiki/Author_citation_(zoology)" title="Author citation (zoology)">Author citation (zoology)</a></div> <p>An "authority" may be placed after a scientific name.<sup id="cite_ref-AJE_95-0" class="reference"><a href="#cite_note-AJE-95"><span class="cite-bracket">[</span>94<span class="cite-bracket">]</span></a></sup> The authority is the name of the scientist or scientists who first validly published the name.<sup id="cite_ref-AJE_95-1" class="reference"><a href="#cite_note-AJE-95"><span class="cite-bracket">[</span>94<span class="cite-bracket">]</span></a></sup> For example, in 1758 Linnaeus gave the <a href="/wiki/Asian_elephant" title="Asian elephant">Asian elephant</a> the scientific name <i>Elephas maximus</i>, so the name is sometimes written as "<i>Elephas maximus</i> Linnaeus, 1758".<sup id="cite_ref-96" class="reference"><a href="#cite_note-96"><span class="cite-bracket">[</span>95<span class="cite-bracket">]</span></a></sup> The names of authors are often abbreviated: the abbreviation <i>L.</i>, for <i>Linnaeus,</i> is commonly used. In botany, there is, in fact, a regulated list of standard abbreviations (see <a href="/wiki/List_of_botanists_by_author_abbreviation" class="mw-redirect" title="List of botanists by author abbreviation">list of botanists by author abbreviation</a>).<sup id="cite_ref-97" class="reference"><a href="#cite_note-97"><span class="cite-bracket">[</span>96<span class="cite-bracket">]</span></a></sup> The system for assigning authorities differs slightly between <a href="/wiki/Author_citation_(botany)" title="Author citation (botany)">botany</a> and <a href="/wiki/Author_citation_(zoology)" title="Author citation (zoology)">zoology</a>.<sup id="cite_ref-Herbarium_8-2" class="reference"><a href="#cite_note-Herbarium-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> However, it is standard that if the genus of a species has been changed since the original description, the original authority's name is placed in parentheses.<sup id="cite_ref-98" class="reference"><a href="#cite_note-98"><span class="cite-bracket">[</span>97<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Phenetics">Phenetics</h2></div> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Phylogenetics.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/69/Phylogenetics.svg/300px-Phylogenetics.svg.png" decoding="async" width="300" height="200" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/69/Phylogenetics.svg/450px-Phylogenetics.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/69/Phylogenetics.svg/600px-Phylogenetics.svg.png 2x" data-file-width="1500" data-file-height="1000" /></a><figcaption>A comparison of phylogenetic and phenetic (character-based) concepts</figcaption></figure> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Phenetics" title="Phenetics">Phenetics</a></div> <p>In phenetics, also known as taximetrics, or numerical taxonomy, organisms are classified based on overall similarity, regardless of their phylogeny or evolutionary relationships.<sup id="cite_ref-BiologyDiscussion_20-7" class="reference"><a href="#cite_note-BiologyDiscussion-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> It results in a measure of hypergeometric "distance" between taxa. Phenetic methods have become relatively rare in modern times, largely superseded by <a href="/wiki/Cladistics" title="Cladistics">cladistic</a> analyses, as phenetic methods do not distinguish shared ancestral (or <a href="/wiki/Plesiomorph" class="mw-redirect" title="Plesiomorph">plesiomorphic</a>) traits from shared derived (or <a href="/wiki/Apomorphic" class="mw-redirect" title="Apomorphic">apomorphic</a>) traits.<sup id="cite_ref-99" class="reference"><a href="#cite_note-99"><span class="cite-bracket">[</span>98<span class="cite-bracket">]</span></a></sup> However, certain phenetic methods, such as <a href="/wiki/Neighbor_joining" title="Neighbor joining">neighbor joining</a>, have persisted, as rapid estimators of relationships when more advanced methods (such as <a href="/wiki/Bayesian_inference" title="Bayesian inference">Bayesian inference</a>) are too computationally expensive.<sup id="cite_ref-100" class="reference"><a href="#cite_note-100"><span class="cite-bracket">[</span>99<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Databases">Databases</h2></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Taxonomic_database" title="Taxonomic database">Taxonomic database</a></div> <p>Modern taxonomy uses <a href="/wiki/Database" title="Database">database</a> technologies to search and catalogue classifications and their documentation.<sup id="cite_ref-101" class="reference"><a href="#cite_note-101"><span class="cite-bracket">[</span>100<span class="cite-bracket">]</span></a></sup> While there is no commonly used database, there are comprehensive databases such as the <i><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">Catalogue of Life</a></i>, which attempts to list every documented species.<sup id="cite_ref-102" class="reference"><a href="#cite_note-102"><span class="cite-bracket">[</span>101<span class="cite-bracket">]</span></a></sup> The catalogue listed 1.64 million species for all kingdoms as of April 2016<sup class="plainlinks noexcerpt noprint asof-tag update" style="display:none;"><a class="external text" href="https://en.wikipedia.org/w/index.php?title=Taxonomy_(biology)&action=edit">[update]</a></sup>, claiming coverage of more than three-quarters of the estimated species known to modern science.<sup id="cite_ref-About_the_Catalogue_of_Life_103-0" class="reference"><a href="#cite_note-About_the_Catalogue_of_Life-103"><span class="cite-bracket">[</span>102<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1184024115"><div class="div-col div-col-small"> <ul><li><a href="/wiki/Automated_species_identification" title="Automated species identification">Automated species identification</a></li> <li><a href="/wiki/Bacterial_taxonomy" title="Bacterial taxonomy">Bacterial taxonomy</a></li> <li><a href="/wiki/Cluster_analysis" title="Cluster analysis">Cluster analysis</a></li> <li><a href="/wiki/Consortium_for_the_Barcode_of_Life" title="Consortium for the Barcode of Life">Consortium for the Barcode of Life</a></li> <li><a href="/wiki/Consortium_of_European_Taxonomic_Facilities" title="Consortium of European Taxonomic Facilities">Consortium of European Taxonomic Facilities</a></li> <li><a href="/wiki/Genetypes" title="Genetypes">Genetypes</a></li> <li><a href="/wiki/Glossary_of_scientific_naming" title="Glossary of scientific naming">Glossary of scientific naming</a></li> <li><a href="/wiki/Identification_(biology)" title="Identification (biology)">Identification (biology)</a></li> <li><i><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></i></li> <li><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a></li> <li><a href="/wiki/Parataxonomy" title="Parataxonomy">Parataxonomy</a></li> <li><a href="/wiki/Phenogram" title="Phenogram">Phenogram</a></li> <li><a href="/wiki/Set_theory" title="Set theory">Set theory</a></li> <li><a href="/wiki/Taxonomy" title="Taxonomy">Taxonomy</a> (general)</li> <li><a href="/wiki/Virus_classification" title="Virus classification">Virus classification</a></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="Notes">Notes</h2></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-89"><span class="mw-cite-backlink"><b><a href="#cite_ref-89">^</a></b></span> <span class="reference-text">This ranking system, except for "Strain", can be remembered by the mnemonic "Do Kings Play Chess On Fine Glass Sets?"</span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239543626"><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-Wilkins2011-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Wilkins2011_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Wilkins2011_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFWilkins2011" class="citation web cs1">Wilkins, J. S. (5 February 2011). <a rel="nofollow" class="external text" href="https://evolvingthoughts.net/2011/02/what-is-systematics-and-what-is-taxonomy/">"What is systematics and what is taxonomy?"</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20160827124330/http://evolvingthoughts.net/2011/02/what-is-systematics-and-what-is-taxonomy/">Archived</a> from the original on 27 August 2016<span class="reference-accessdate">. Retrieved <span class="nowrap">21 August</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=What+is+systematics+and+what+is+taxonomy%3F&rft.date=2011-02-05&rft.aulast=Wilkins&rft.aufirst=J.+S.&rft_id=http%3A%2F%2Fevolvingthoughts.net%2F2011%2F02%2Fwhat-is-systematics-and-what-is-taxonomy%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Judd-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-Judd_2-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFJuddCampbellKelloggStevens2007" class="citation book cs1">Judd, W. S.; Campbell, C. S.; Kellogg, E. A.; Stevens, P. F.; Donoghue, M. J. (2007). "Taxonomy". <i>Plant Systematics: A Phylogenetic Approach</i> (3rd ed.). Sunderland: Sinauer Associates.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Taxonomy&rft.btitle=Plant+Systematics%3A+A+Phylogenetic+Approach&rft.place=Sunderland&rft.edition=3rd&rft.pub=Sinauer+Associates&rft.date=2007&rft.aulast=Judd&rft.aufirst=W.+S.&rft.au=Campbell%2C+C.+S.&rft.au=Kellogg%2C+E.+A.&rft.au=Stevens%2C+P.+F.&rft.au=Donoghue%2C+M.+J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Simpson-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-Simpson_3-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSimpson2010" class="citation book cs1">Simpson, Michael G. (2010). "Chapter 1 Plant Systematics: an Overview". <i>Plant Systematics</i> (2nd ed.). Academic Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780123743800" title="Special:BookSources/9780123743800"><bdi>9780123743800</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Chapter+1+Plant+Systematics%3A+an+Overview&rft.btitle=Plant+Systematics&rft.edition=2nd&rft.pub=Academic+Press&rft.date=2010&rft.isbn=9780123743800&rft.aulast=Simpson&rft.aufirst=Michael+G.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKirkCannonMinterStalpers2008" class="citation book cs1">Kirk, P. M.; Cannon, P. F.; Minter, D. W.; Stalpers, J. A., eds. (2008). "Taxonomy". <i>Dictionary of the Fungi</i> (10th ed.). CABI.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Taxonomy&rft.btitle=Dictionary+of+the+Fungi&rft.edition=10th&rft.pub=CABI&rft.date=2008&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWalker1988" class="citation book cs1">Walker, P. M. B., ed. (1988). <i>The Wordsworth Dictionary of Science and Technology</i>. W. R. Chambers Ltd. and Cambridge University Press.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=The+Wordsworth+Dictionary+of+Science+and+Technology&rft.pub=W.+R.+Chambers+Ltd.+and+Cambridge+University+Press&rft.date=1988&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Henderson-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-Henderson_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Henderson_6-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLawrence2005" class="citation book cs1">Lawrence, E. (2005). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=-PLgy6DWe0wC"><i>Henderson's Dictionary Of Biology</i></a>. Pearson/Prentice Hall. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780131273849" title="Special:BookSources/9780131273849"><bdi>9780131273849</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Henderson%27s+Dictionary+Of+Biology&rft.pub=Pearson%2FPrentice+Hall&rft.date=2005&rft.isbn=9780131273849&rft.aulast=Lawrence&rft.aufirst=E.&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3D-PLgy6DWe0wC&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWheeler2004" class="citation journal cs1 cs1-prop-long-vol"><a href="/wiki/Quentin_D._Wheeler" title="Quentin D. 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(eds.). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1693342">"Taxonomic triage and the poverty of phylogeny"</a>. <i><a href="/wiki/Philosophical_Transactions_of_the_Royal_Society" title="Philosophical Transactions of the Royal Society">Philosophical Transactions of the Royal Society</a></i>. 359: Taxonomy for the twenty-first century (1444): 571–583. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frstb.2003.1452">10.1098/rstb.2003.1452</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0962-8436">0962-8436</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1693342">1693342</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15253345">15253345</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Philosophical+Transactions+of+the+Royal+Society&rft.atitle=Taxonomic+triage+and+the+poverty+of+phylogeny&rft.volume=359%3A+Taxonomy+for+the+twenty-first+century&rft.issue=1444&rft.pages=571-583&rft.date=2004&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1693342%23id-name%3DPMC&rft.issn=0962-8436&rft_id=info%3Apmid%2F15253345&rft_id=info%3Adoi%2F10.1098%2Frstb.2003.1452&rft.aulast=Wheeler&rft.aufirst=Quentin+D.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1693342&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Herbarium-8"><span class="mw-cite-backlink">^ <a href="#cite_ref-Herbarium_8-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Herbarium_8-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Herbarium_8-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20161123030604/http://herbarium.usu.edu/teaching/4420/botnom.htm">"Nomenclature, Names, and Taxonomy"</a>. <i>Intermountain Herbarium</i>. Utah State University. 2005. Archived from <a rel="nofollow" class="external text" href="http://herbarium.usu.edu:80/teaching/4420/botnom.htm">the original</a> on 23 November 2016.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Intermountain+Herbarium&rft.atitle=Nomenclature%2C+Names%2C+and+Taxonomy&rft.date=2005&rft_id=http%3A%2F%2Fherbarium.usu.edu%3A80%2Fteaching%2F4420%2Fbotnom.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Laurin_2023_Advent-9"><span class="mw-cite-backlink">^ <a href="#cite_ref-Laurin_2023_Advent_9-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Laurin_2023_Advent_9-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Laurin_2023_Advent_9-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLaurin2023" class="citation book cs1">Laurin, Michel (3 August 2023). <a rel="nofollow" class="external text" href="https://www.taylorfrancis.com/books/mono/10.1201/9781003092827/advent-phylocode-michel-laurin"><i>The Advent of PhyloCode: The Continuing Evolution of Biological Nomenclature</i></a>. Boca Raton, Florida: CRC Press. pp. xv + 209. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1201%2F9781003092827">10.1201/9781003092827</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9781003092827" title="Special:BookSources/9781003092827"><bdi>9781003092827</bdi></a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230905140719/https://www.taylorfrancis.com/books/mono/10.1201/9781003092827/advent-phylocode-michel-laurin">Archived</a> from the original on 5 September 2023<span class="reference-accessdate">. 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Retrieved <span class="nowrap">15 September</span> 2017</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Chapter+6%3A+Microtaxonomy%2C+the+science+of+species&rft.btitle=The+Growth+of+Biological+Thought%3A+Diversity%2C+Evolution%2C+and+Inheritance&rft.pub=Belknap+Press+of+Harvard+University+Press&rft.date=1982&rft.isbn=9780674364462&rft.aulast=Mayr&rft.aufirst=Ernst&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3DpHThtE2R0UQC&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-27"><span class="mw-cite-backlink"><b><a href="#cite_ref-27">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.biological-concepts.com/views/search.php?term=1508">"Result of Your Query"</a>. <i>biological-concepts.com</i>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20170405170414/http://www.biological-concepts.com/views/search.php?term=1508">Archived</a> from the original on 5 April 2017.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=biological-concepts.com&rft.atitle=Result+of+Your+Query&rft_id=http%3A%2F%2Fwww.biological-concepts.com%2Fviews%2Fsearch.php%3Fterm%3D1508&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Cantino_&_de_Queiroz_2020-28"><span class="mw-cite-backlink">^ <a href="#cite_ref-Cantino_&_de_Queiroz_2020_28-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Cantino_&_de_Queiroz_2020_28-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Cantino_&_de_Queiroz_2020_28-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCantinode_Queiroz2020" class="citation book cs1">Cantino, Philip D.; de Queiroz, Kevin (29 April 2020). <a rel="nofollow" class="external text" href="https://www.routledge.com/International-Code-of-Phylogenetic-Nomenclature-PhyloCode/Queiroz-Cantino/p/book/9781138332829"><i>International Code of Phylogenetic Nomenclature (PhyloCode): A Phylogenetic Code of Biological Nomenclature</i></a>. 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Retrieved <span class="nowrap">19 September</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=International+Code+of+Phylogenetic+Nomenclature+%28PhyloCode%29%3A+A+Phylogenetic+Code+of+Biological+Nomenclature&rft.place=Boca+Raton%2C+Florida&rft.pages=xl+%2B+149&rft.pub=CRC+Press&rft.date=2020-04-29&rft.isbn=978-0429821356&rft.aulast=Cantino&rft.aufirst=Philip+D.&rft.au=de+Queiroz%2C+Kevin&rft_id=https%3A%2F%2Fwww.routledge.com%2FInternational-Code-of-Phylogenetic-Nomenclature-PhyloCode%2FQueiroz-Cantino%2Fp%2Fbook%2F9781138332829&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Voultsiadou_&_Vafidis_2007-29"><span class="mw-cite-backlink"><b><a href="#cite_ref-Voultsiadou_&_Vafidis_2007_29-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFVoultsiadouVafidis2007" class="citation journal cs1">Voultsiadou, Eleni; Vafidis, Dimitris (1 January 2007). <a rel="nofollow" class="external text" href="https://doi.org/10.1163/18759866-07602004">"Marine invertebrate diversity in Aristotle's zoology"</a>. <i>Contributions to Zoology</i>. <b>76</b> (2): 103–120. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1163%2F18759866-07602004">10.1163/18759866-07602004</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1875-9866">1875-9866</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230425063507/https://brill.com/view/journals/ctoz/76/2/article-p103_4.xml">Archived</a> from the original on 25 April 2023<span class="reference-accessdate">. Retrieved <span class="nowrap">19 September</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Contributions+to+Zoology&rft.atitle=Marine+invertebrate+diversity+in+Aristotle%27s+zoology&rft.volume=76&rft.issue=2&rft.pages=103-120&rft.date=2007-01-01&rft_id=info%3Adoi%2F10.1163%2F18759866-07602004&rft.issn=1875-9866&rft.aulast=Voultsiadou&rft.aufirst=Eleni&rft.au=Vafidis%2C+Dimitris&rft_id=https%3A%2F%2Fdoi.org%2F10.1163%2F18759866-07602004&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-Voultsiadou_et_al._2017-30"><span class="mw-cite-backlink"><b><a href="#cite_ref-Voultsiadou_et_al._2017_30-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFVoultsiadouGerovasileiouVandepitteGanias2017" class="citation journal cs1">Voultsiadou, Eleni; Gerovasileiou, Vasilis; Vandepitte, Leen; Ganias, Kostas; Arvanitidis, Christos (2017). <a rel="nofollow" class="external text" href="https://doi.org/10.12681%2Fmms.13874">"Aristotle's scientific contributions to the classification, nomenclature and distribution of marine organisms"</a>. <i>Mediterranean Marine Science</i>. <b>18</b> (3): 468–478. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.12681%2Fmms.13874">10.12681/mms.13874</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1791-6763">1791-6763</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Mediterranean+Marine+Science&rft.atitle=Aristotle%27s+scientific+contributions+to+the+classification%2C+nomenclature+and+distribution+of+marine+organisms&rft.volume=18&rft.issue=3&rft.pages=468-478&rft.date=2017&rft_id=info%3Adoi%2F10.12681%2Fmms.13874&rft.issn=1791-6763&rft.aulast=Voultsiadou&rft.aufirst=Eleni&rft.au=Gerovasileiou%2C+Vasilis&rft.au=Vandepitte%2C+Leen&rft.au=Ganias%2C+Kostas&rft.au=Arvanitidis%2C+Christos&rft_id=https%3A%2F%2Fdoi.org%2F10.12681%252Fmms.13874&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-FOOTNOTEDatta1988-31"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEDatta1988_31-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFDatta1988">Datta 1988</a>.</span> </li> <li id="cite_note-FOOTNOTEStace1989-32"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEStace1989_32-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFStace1989">Stace 1989</a>.</span> </li> <li id="cite_note-FOOTNOTEStuessy2009-33"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEStuessy2009_33-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFStuessy2009">Stuessy 2009</a>.</span> </li> <li id="cite_note-Manktelow-34"><span class="mw-cite-backlink">^ <a href="#cite_ref-Manktelow_34-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Manktelow_34-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Manktelow_34-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFManktelow2010" class="citation web cs1">Manktelow, M. (2010). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150529020546/http://atbi.eu/summerschool/files/summerschool/Manktelow_Syllabus.pdf">"History of Taxonomy"</a> <span class="cs1-format">(PDF)</span>. Dept. of Systematic Biology, <a href="/wiki/Uppsala_University" title="Uppsala University">Uppsala University</a>. 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Retrieved <span class="nowrap">8 April</span> 2017</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=theplantlist.org&rft.atitle=About+%E2%80%93+The+Plant+List&rft_id=http%3A%2F%2Fwww.theplantlist.org%2Fabout%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> <li id="cite_note-About_the_Catalogue_of_Life-103"><span class="mw-cite-backlink"><b><a href="#cite_ref-About_the_Catalogue_of_Life_103-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.catalogueoflife.org/annual-checklist/2016/info/about">"About the Catalogue of Life: 2016 Annual Checklist"</a>. <i>Catalogue of Life</i>. <a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">Integrated Taxonomic Information System</a> (ITIS). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20160515032942/http://www.catalogueoflife.org/annual-checklist/2016/info/about">Archived</a> from the original on 15 May 2016<span class="reference-accessdate">. Retrieved <span class="nowrap">22 May</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Catalogue+of+Life&rft.atitle=About+the+Catalogue+of+Life%3A+2016+Annual+Checklist&rft_id=http%3A%2F%2Fwww.catalogueoflife.org%2Fannual-checklist%2F2016%2Finfo%2Fabout&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Bibliography">Bibliography</h2></div> <style data-mw-deduplicate="TemplateStyles:r1239549316">.mw-parser-output .refbegin{margin-bottom:0.5em}.mw-parser-output .refbegin-hanging-indents>ul{margin-left:0}.mw-parser-output .refbegin-hanging-indents>ul>li{margin-left:0;padding-left:3.2em;text-indent:-3.2em}.mw-parser-output .refbegin-hanging-indents ul,.mw-parser-output .refbegin-hanging-indents ul li{list-style:none}@media(max-width:720px){.mw-parser-output .refbegin-hanging-indents>ul>li{padding-left:1.6em;text-indent:-1.6em}}.mw-parser-output .refbegin-columns{margin-top:0.3em}.mw-parser-output .refbegin-columns ul{margin-top:0}.mw-parser-output .refbegin-columns li{page-break-inside:avoid;break-inside:avoid-column}@media screen{.mw-parser-output .refbegin{font-size:90%}}</style><div class="refbegin" style=""> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDatta1988" class="citation book cs1">Datta, Subhash Chandra (1988). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=X7lfMACvjs4C"><i>Systematic Botany</i></a> (4 ed.). New Delhi: New Age International. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9788122400137" title="Special:BookSources/9788122400137"><bdi>9788122400137</bdi></a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20231113101351/https://books.google.com/books?id=X7lfMACvjs4C">Archived</a> from the original on 13 November 2023<span class="reference-accessdate">. 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Cambridge: <a href="/wiki/Cambridge_University_Press" title="Cambridge University Press">Cambridge University Press</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780521427852" title="Special:BookSources/9780521427852"><bdi>9780521427852</bdi></a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230113180249/https://books.google.com/books?id=VfQnuwh3bw8C">Archived</a> from the original on 13 January 2023<span class="reference-accessdate">. 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Columbia University Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780231147125" title="Special:BookSources/9780231147125"><bdi>9780231147125</bdi></a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20231113101356/https://books.google.com/books?id=0bYs8F0Mb9gC">Archived</a> from the original on 13 November 2023<span class="reference-accessdate">. Retrieved <span class="nowrap">6 February</span> 2014</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Plant+Taxonomy%3A+The+Systematic+Evaluation+of+Comparative+Data&rft.pub=Columbia+University+Press&rft.date=2009&rft.isbn=9780231147125&rft.aulast=Stuessy&rft.aufirst=Tod+F.&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3D0bYs8F0Mb9gC&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFTurrill1938" class="citation journal cs1">Turrill, W. B. (1938). "The Expansion Of Taxonomy With Special Reference To Spermatophyta". <i>Biological Reviews</i>. <b>13</b> (4): 342–373. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.1469-185X.1938.tb00522.x">10.1111/j.1469-185X.1938.tb00522.x</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:84905335">84905335</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biological+Reviews&rft.atitle=The+Expansion+Of+Taxonomy+With+Special+Reference+To+Spermatophyta&rft.volume=13&rft.issue=4&rft.pages=342-373&rft.date=1938&rft_id=info%3Adoi%2F10.1111%2Fj.1469-185X.1938.tb00522.x&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A84905335%23id-name%3DS2CID&rft.aulast=Turrill&rft.aufirst=W.+B.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWileyLieberman2011" class="citation book cs1">Wiley, Edward O.; Lieberman, Bruce S. (2011). <i>Phylogenetics: Theory and Practice of Phylogenetic Systematics</i> (2nd ed.). John Wiley & Sons. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780470905968" title="Special:BookSources/9780470905968"><bdi>9780470905968</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Phylogenetics%3A+Theory+and+Practice+of+Phylogenetic+Systematics&rft.edition=2nd&rft.pub=John+Wiley+%26+Sons&rft.date=2011&rft.isbn=9780470905968&rft.aulast=Wiley&rft.aufirst=Edward+O.&rft.au=Lieberman%2C+Bruce+S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATaxonomy+%28biology%29" class="Z3988"></span></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2></div> <ul><li><a rel="nofollow" class="external text" href="http://www.nhm.ac.uk/nature-online/science-of-natural-history/taxonomy-systematics/what-is-taxonomy/index.html">What is taxonomy?</a> at the <a rel="nofollow" class="external text" href="http://www.nhm.ac.uk/"><i>Natural History Museum London</i></a></li> <li><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/">Taxonomy</a> at <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/">NCBI</a> the <i>National Center for Biotechnology Information</i></li> <li><a rel="nofollow" class="external text" href="https://www.uniprot.org/taxonomy/">Taxonomy</a> at <a rel="nofollow" class="external text" href="https://www.uniprot.org/">UniProt</a> the <i>Universal Protein Resource</i></li> <li><a rel="nofollow" class="external text" href="https://www.itis.gov/">ITIS</a> the <i>Integrated Taxonomic Information System</i></li> <li><a rel="nofollow" class="external text" href="http://www.cetaf.org/">CETaF</a> the <i>Consortium of European Taxonomic Facilities</i></li> <li><a class="external text" href="https://species.wikimedia.org/">Wikispecies</a> <i>free species directory</i></li> <li><a rel="nofollow" class="external text" href="https://www.thescienceclass.online/2019/10/biological-classification.html">Biological classification.</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20200813180358/https://www.thescienceclass.online/2019/10/biological-classification.html">Archived</a> 13 August 2020 at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a></li></ul> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output .navbox-list-with-group{text-align:left;border-left-width:2px;border-left-style:solid}.mw-parser-output tr+tr>.navbox-abovebelow,.mw-parser-output tr+tr>.navbox-group,.mw-parser-output tr+tr>.navbox-image,.mw-parser-output tr+tr>.navbox-list{border-top:2px solid #fdfdfd}.mw-parser-output .navbox-title{background-color:#ccf}.mw-parser-output .navbox-abovebelow,.mw-parser-output .navbox-group,.mw-parser-output .navbox-subgroup .navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Branches_of_biology" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Branches_of_biology" title="Template:Branches of biology"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Branches_of_biology" title="Template talk:Branches of biology"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Branches_of_biology" title="Special:EditPage/Template:Branches of biology"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Branches_of_biology" style="font-size:114%;margin:0 4em"><a href="/wiki/Outline_of_biology#Branches" title="Outline of biology">Branches of biology</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></li> <li><a href="/wiki/Aerobiology" title="Aerobiology">Aerobiology</a></li> <li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Agrostology" title="Agrostology">Agrostology</a></li> <li><a href="/wiki/Anatomy" title="Anatomy">Anatomy</a></li> <li><a href="/wiki/Astrobiology" title="Astrobiology">Astrobiology</a></li> <li><a href="/wiki/Bacteriology" title="Bacteriology">Bacteriology</a></li> <li><a href="/wiki/Biochemistry" title="Biochemistry">Biochemistry</a></li> <li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Biogeology" title="Biogeology">Biogeology</a></li> <li><a href="/wiki/Bioinformatics" title="Bioinformatics">Bioinformatics</a></li> <li><a href="/wiki/Biological_engineering" title="Biological engineering">Biological engineering</a></li> <li><a href="/wiki/Biomechanics" title="Biomechanics">Biomechanics</a></li> <li><a href="/wiki/Biophysics" title="Biophysics">Biophysics</a></li> <li><a href="/wiki/Biosemiotics" title="Biosemiotics">Biosemiotics</a></li> <li><a href="/wiki/Biostatistics" title="Biostatistics">Biostatistics</a></li> <li><a href="/wiki/Biotechnology" title="Biotechnology">Biotechnology</a></li> <li><a href="/wiki/Botany" title="Botany">Botany</a></li> <li><a href="/wiki/Cell_biology" title="Cell biology">Cell biology</a></li> <li><a href="/wiki/Cellular_microbiology" title="Cellular microbiology">Cellular microbiology</a></li> <li><a href="/wiki/Chemical_biology" title="Chemical biology">Chemical biology</a></li> <li><a href="/wiki/Chronobiology" title="Chronobiology">Chronobiology</a></li> <li><a href="/wiki/Cognitive_biology" title="Cognitive biology">Cognitive biology</a></li> <li><a href="/wiki/Computational_biology" title="Computational biology">Computational biology</a></li> <li><a href="/wiki/Conservation_biology" title="Conservation biology">Conservation biology</a></li> <li><a href="/wiki/Cryobiology" title="Cryobiology">Cryobiology</a></li> <li><a href="/wiki/Cytogenetics" title="Cytogenetics">Cytogenetics</a></li> <li><a href="/wiki/Dendrology" title="Dendrology">Dendrology</a></li> <li><a href="/wiki/Developmental_biology" title="Developmental biology">Developmental biology</a></li> <li><a href="/wiki/Ecological_genetics" title="Ecological genetics">Ecological genetics</a></li> <li><a href="/wiki/Ecology" title="Ecology">Ecology</a></li> <li><a href="/wiki/Embryology" title="Embryology">Embryology</a></li> <li><a href="/wiki/Epidemiology" title="Epidemiology">Epidemiology</a></li> <li><a href="/wiki/Epigenetics" title="Epigenetics">Epigenetics</a></li> <li><a href="/wiki/Evolutionary_biology" title="Evolutionary biology">Evolutionary biology</a></li> <li><a href="/wiki/Freshwater_biology" title="Freshwater biology">Freshwater biology</a></li> <li><a href="/wiki/Generative_biology" class="mw-redirect" title="Generative biology">Generative biology</a></li> <li><a href="/wiki/Genetics" title="Genetics">Genetics</a></li> <li><a href="/wiki/Genomics" title="Genomics">Genomics</a></li> <li><a href="/wiki/Geobiology" title="Geobiology">Geobiology</a></li> <li><a href="/wiki/Gerontology" title="Gerontology">Gerontology</a></li> <li><a href="/wiki/Herpetology" title="Herpetology">Herpetology</a></li> <li><a href="/wiki/Histology" title="Histology">Histology</a></li> <li><a href="/wiki/Human_biology" title="Human biology">Human biology</a></li> <li><a href="/wiki/Ichthyology" title="Ichthyology">Ichthyology</a></li> <li><a href="/wiki/Immunology" title="Immunology">Immunology</a></li> <li><a href="/wiki/Lipidology" title="Lipidology">Lipidology</a></li> <li><a href="/wiki/Mammalogy" title="Mammalogy">Mammalogy</a></li> <li><a href="/wiki/Marine_biology" title="Marine biology">Marine biology</a></li> <li><a href="/wiki/Mathematical_and_theoretical_biology" title="Mathematical and theoretical biology">Mathematical biology</a></li> <li><a href="/wiki/Microbiology" title="Microbiology">Microbiology</a></li> <li><a href="/wiki/Molecular_biology" title="Molecular biology">Molecular biology</a></li> <li><a href="/wiki/Mycology" title="Mycology">Mycology</a></li> <li><a href="/wiki/Neontology" title="Neontology">Neontology</a></li> <li><a href="/wiki/Neuroscience" title="Neuroscience">Neuroscience</a></li> <li><a href="/wiki/Nutritional_science" title="Nutritional science">Nutrition</a></li> <li><a href="/wiki/Ornithology" title="Ornithology">Ornithology</a></li> <li><a href="/wiki/Osteology" title="Osteology">Osteology</a></li> <li><a href="/wiki/Paleontology" title="Paleontology">Paleontology</a></li> <li><a href="/wiki/Parasitology" title="Parasitology">Parasitology</a></li> <li><a href="/wiki/Pathology" title="Pathology">Pathology</a></li> <li><a href="/wiki/Pharmacology" title="Pharmacology">Pharmacology</a></li> <li><a href="/wiki/Photobiology" title="Photobiology">Photobiology</a></li> <li><a href="/wiki/Phycology" title="Phycology">Phycology</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a></li> <li><a href="/wiki/Physiology" title="Physiology">Physiology</a></li> <li><a href="/wiki/Pomology" title="Pomology">Pomology</a></li> <li><a href="/wiki/Primatology" title="Primatology">Primatology</a></li> <li><a href="/wiki/Proteomics" title="Proteomics">Proteomics</a></li> <li><a href="/wiki/Protistology" title="Protistology">Protistology</a></li> <li><a href="/wiki/Quantum_biology" title="Quantum biology">Quantum biology</a></li> <li><a href="/wiki/Relational_biology" class="mw-redirect" title="Relational biology">Relational biology</a></li> <li><a href="/wiki/Reproductive_biology" title="Reproductive biology">Reproductive biology</a></li> <li><a href="/wiki/Sociobiology" title="Sociobiology">Sociobiology</a></li> <li><a href="/wiki/Structural_biology" title="Structural biology">Structural biology</a></li> <li><a href="/wiki/Synthetic_biology" title="Synthetic biology">Synthetic biology</a></li> <li><a href="/wiki/Systematics" title="Systematics">Systematics</a></li> <li><a href="/wiki/Systems_biology" title="Systems biology">Systems biology</a></li> <li><a class="mw-selflink selflink">Taxonomy</a></li> <li><a href="/wiki/Teratology" title="Teratology">Teratology</a></li> <li><a href="/wiki/Toxicology" title="Toxicology">Toxicology</a></li> <li><a href="/wiki/Virology" title="Virology">Virology</a></li> <li><a href="/wiki/Virophysics" title="Virophysics">Virophysics</a></li> <li><a href="/wiki/Xenobiology" title="Xenobiology">Xenobiology</a></li> <li><a href="/wiki/Zoology" title="Zoology">Zoology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">See also</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/History_of_biology" title="History of biology">History of biology</a></li> <li><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></li> <li><a href="/wiki/Timeline_of_biology_and_organic_chemistry" title="Timeline of biology and organic chemistry">Timeline of biology and organic chemistry</a></li></ul> </div></td></tr></tbody></table></div> <style data-mw-deduplicate="TemplateStyles:r1130092004">.mw-parser-output .portal-bar{font-size:88%;font-weight:bold;display:flex;justify-content:center;align-items:baseline}.mw-parser-output .portal-bar-bordered{padding:0 2em;background-color:#fdfdfd;border:1px solid #a2a9b1;clear:both;margin:1em auto 0}.mw-parser-output .portal-bar-related{font-size:100%;justify-content:flex-start}.mw-parser-output .portal-bar-unbordered{padding:0 1.7em;margin-left:0}.mw-parser-output .portal-bar-header{margin:0 1em 0 0.5em;flex:0 0 auto;min-height:24px}.mw-parser-output .portal-bar-content{display:flex;flex-flow:row wrap;flex:0 1 auto;padding:0.15em 0;column-gap:1em;align-items:baseline;margin:0;list-style:none}.mw-parser-output .portal-bar-content-related{margin:0;list-style:none}.mw-parser-output .portal-bar-item{display:inline-block;margin:0.15em 0.2em;min-height:24px;line-height:24px}@media screen and (max-width:768px){.mw-parser-output .portal-bar{font-size:88%;font-weight:bold;display:flex;flex-flow:column wrap;align-items:baseline}.mw-parser-output .portal-bar-header{text-align:center;flex:0;padding-left:0.5em;margin:0 auto}.mw-parser-output .portal-bar-related{font-size:100%;align-items:flex-start}.mw-parser-output .portal-bar-content{display:flex;flex-flow:row wrap;align-items:center;flex:0;column-gap:1em;border-top:1px solid #a2a9b1;margin:0 auto;list-style:none}.mw-parser-output .portal-bar-content-related{border-top:none;margin:0;list-style:none}}.mw-parser-output .navbox+link+.portal-bar,.mw-parser-output .navbox+style+.portal-bar,.mw-parser-output .navbox+link+.portal-bar-bordered,.mw-parser-output .navbox+style+.portal-bar-bordered,.mw-parser-output .sister-bar+link+.portal-bar,.mw-parser-output .sister-bar+style+.portal-bar,.mw-parser-output .portal-bar+.navbox-styles+.navbox,.mw-parser-output .portal-bar+.navbox-styles+.sister-bar{margin-top:-1px}</style><div class="portal-bar noprint metadata noviewer portal-bar-bordered" role="navigation" aria-label="Portals"><span class="portal-bar-header"><a href="/wiki/Wikipedia:Contents/Portals" title="Wikipedia:Contents/Portals">Portal</a>:</span><ul class="portal-bar-content"><li class="portal-bar-item"><span class="nowrap"><span typeof="mw:File"><a href="/wiki/File:Issoria_lathonia.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/21px-Issoria_lathonia.jpg" decoding="async" width="21" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/32px-Issoria_lathonia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/42px-Issoria_lathonia.jpg 2x" data-file-width="629" data-file-height="445" /></a></span> </span><a href="/wiki/Portal:Biology" title="Portal:Biology">Biology</a></li></ul></div><style data-mw-deduplicate="TemplateStyles:r1236088147">.mw-parser-output .sister-bar{display:flex;justify-content:center;align-items:baseline;font-size:88%;background-color:#fdfdfd;border:1px solid #a2a9b1;clear:both;margin:1em 0 0;padding:0 2em}.mw-parser-output .sister-bar-header{margin:0 1em 0 0.5em;padding:0.2em 0;flex:0 0 auto;min-height:24px;line-height:22px}.mw-parser-output .sister-bar-content{display:flex;flex-flow:row wrap;flex:0 1 auto;align-items:baseline;padding:0.2em 0;column-gap:1em;margin:0;list-style:none}.mw-parser-output .sister-bar-item{display:flex;align-items:baseline;margin:0.15em 0;min-height:24px;text-align:left}.mw-parser-output .sister-bar-logo{width:22px;line-height:22px;margin:0 0.2em;text-align:right}.mw-parser-output .sister-bar-link{margin:0 0.2em;text-align:left}@media screen and (max-width:960px){.mw-parser-output .sister-bar{flex-flow:column wrap;margin:1em auto 0}.mw-parser-output .sister-bar-header{flex:0 1}.mw-parser-output .sister-bar-content{flex:1;border-top:1px solid #a2a9b1;margin:0;list-style:none}.mw-parser-output .sister-bar-item{flex:0 0 20em;min-width:20em}}.mw-parser-output .navbox+link+.sister-bar,.mw-parser-output .navbox+style+.sister-bar,.mw-parser-output .portal-bar+link+.sister-bar,.mw-parser-output .portal-bar+style+.sister-bar,.mw-parser-output .sister-bar+.navbox-styles+.navbox,.mw-parser-output .sister-bar+.navbox-styles+.portal-bar{margin-top:-1px}@media print{body.ns-0 .mw-parser-output .sister-bar{display:none!important}}</style><div class="noprint metadata sister-bar" role="navigation" aria-label="sister-projects"><div class="sister-bar-header"><b>Taxonomy (biology)</b> at Wikipedia's <a href="/wiki/Wikipedia:Wikimedia_sister_projects" title="Wikipedia:Wikimedia sister projects"><span id="sister-projects" style="white-space:nowrap;">sister projects</span></a>:</div><ul class="sister-bar-content"><li class="sister-bar-item"><span class="sister-bar-logo"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/0/06/Wiktionary-logo-v2.svg/19px-Wiktionary-logo-v2.svg.png" decoding="async" width="19" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/0/06/Wiktionary-logo-v2.svg/29px-Wiktionary-logo-v2.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/0/06/Wiktionary-logo-v2.svg/38px-Wiktionary-logo-v2.svg.png 2x" data-file-width="391" data-file-height="391" /></span></span></span><span class="sister-bar-link"><b><a href="https://en.wiktionary.org/wiki/taxonomy" class="extiw" title="wikt:taxonomy">Definitions</a></b> from Wiktionary</span></li><li class="sister-bar-item"><span class="sister-bar-logo"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/14px-Commons-logo.svg.png" decoding="async" width="14" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/21px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/28px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></span><span class="sister-bar-link"><b><a href="https://commons.wikimedia.org/wiki/Category:Taxonomy_(biology)" class="extiw" title="c:Category:Taxonomy (biology)">Media</a></b> from Commons</span></li><li class="sister-bar-item"><span class="sister-bar-logo"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Wikiquote-logo.svg/16px-Wikiquote-logo.svg.png" decoding="async" width="16" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Wikiquote-logo.svg/24px-Wikiquote-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Wikiquote-logo.svg/32px-Wikiquote-logo.svg.png 2x" data-file-width="300" data-file-height="355" /></span></span></span><span class="sister-bar-link"><b><a href="https://en.wikiquote.org/wiki/Taxonomy_(biology)" class="extiw" title="q:Taxonomy (biology)">Quotations</a></b> from Wikiquote</span></li><li class="sister-bar-item"><span class="sister-bar-logo"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4c/Wikisource-logo.svg/18px-Wikisource-logo.svg.png" decoding="async" width="18" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4c/Wikisource-logo.svg/28px-Wikisource-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4c/Wikisource-logo.svg/36px-Wikisource-logo.svg.png 2x" data-file-width="410" data-file-height="430" /></span></span></span><span class="sister-bar-link"><b><a href="https://en.wikisource.org/wiki/Special:Search/Taxonomy_(biology)" class="extiw" title="s:Special:Search/Taxonomy (biology)">Texts</a></b> from Wikisource</span></li><li class="sister-bar-item"><span class="sister-bar-logo"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Wikiversity_logo_2017.svg/21px-Wikiversity_logo_2017.svg.png" decoding="async" width="21" height="17" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Wikiversity_logo_2017.svg/32px-Wikiversity_logo_2017.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Wikiversity_logo_2017.svg/42px-Wikiversity_logo_2017.svg.png 2x" data-file-width="626" data-file-height="512" /></span></span></span><span class="sister-bar-link"><b><a href="https://en.wikiversity.org/wiki/Taxonomy_(Biology)" class="extiw" title="v:Taxonomy (Biology)">Resources</a></b> from Wikiversity</span></li><li class="sister-bar-item"><span class="sister-bar-logo"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/21px-Wikidata-logo.svg.png" decoding="async" width="21" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/32px-Wikidata-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/42px-Wikidata-logo.svg.png 2x" data-file-width="1050" data-file-height="590" /></span></span></span><span class="sister-bar-link"><b><a href="https://www.wikidata.org/wiki/Q8269924" class="extiw" title="d:Q8269924">Data</a></b> from Wikidata</span></li></ul></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐f69cdc8f6‐542sp Cached time: 20241122140429 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 1.474 seconds Real time usage: 1.731 seconds Preprocessor 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