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A nearly complete juvenile specimen of Gladocephaloideus" /> <title>(PDF) New Material of the Pterosaur Gladocephaloideus Lü et al., 2012 from the Early Cretaceous of Liaoning Province, China, with Comments on Its Systematic Position | Martin Kundrát - Academia.edu</title> <link rel="canonical" href="https://www.academia.edu/106057126/New_Material_of_the_Pterosaur_Gladocephaloideus_L%C3%BC_et_al_2012_from_the_Early_Cretaceous_of_Liaoning_Province_China_with_Comments_on_Its_Systematic_Position" /> <script async src="https://www.googletagmanager.com/gtag/js?id=G-5VKX33P2DS"></script> <script> window.dataLayer = window.dataLayer || []; function gtag(){dataLayer.push(arguments);} gtag('js', new Date()); gtag('config', 'G-5VKX33P2DS', { cookie_domain: 'academia.edu', send_page_view: false, }); gtag('event', 'page_view', { 'controller': "single_work", 'action': "show", 'controller_action': 'single_work#show', 'logged_in': 'false', 'edge': 'unknown', // Send nil if there is no A/B test bucket, in case some records get logged // with missing data - that way we can distinguish between the two cases. // ab_test_bucket should be of the form <ab_test_name>:<bucket> 'ab_test_bucket': null, }) </script> <script> var $controller_name = 'single_work'; var $action_name = "show"; var $rails_env = 'production'; var $app_rev = '9387f500ddcbb8d05c67bef28a2fe0334f1aafb8'; var $domain = 'academia.edu'; var $app_host = "academia.edu"; var $asset_host = "academia-assets.com"; var $start_time = new Date().getTime(); var $recaptcha_key = "6LdxlRMTAAAAADnu_zyLhLg0YF9uACwz78shpjJB"; var $recaptcha_invisible_key = "6Lf3KHUUAAAAACggoMpmGJdQDtiyrjVlvGJ6BbAj"; var $disableClientRecordHit = false; </script> <script> window.require = { config: function() { return function() {} } } </script> <script> window.Aedu = window.Aedu || {}; window.Aedu.hit_data = null; window.Aedu.serverRenderTime = new Date(1733114478000); window.Aedu.timeDifference = new Date().getTime() - 1733114478000; </script> <script type="application/ld+json">{"@context":"https://schema.org","@type":"ScholarlyArticle","abstract":"Although there are nine genera of ctenochasmatoids reported from the Jehol Biota, at present each is known from a specimen that has either a skull or a relatively complete postcranial skeleton. A nearly complete juvenile specimen of Gladocephaloideus from the Lower Cretaceous Jiufotang Formation of Sihedang, Lingyuan of Liaoning Province is the most complete ctenochasmatoid preserved to date with a skull and postcranial skeleton. Based on the holotype (IG-CAGS 08-07) and the nearly complete new specimen (JPM 2014-004), the diagnosis of Gladocephaloideus is amended: approximately 50 teeth in total with sharp tips; small nasoantorbital opening, occupying approximately 13% of total skull length; ratio of prenarial rostrum length to skull length approximately 0.63; deep groove along the mid-line of the mandibular symphysis; length to width ratio of the longest cervical vertebra = 4.1; ratio of femur length to tibia length = 0.61; tibia as long as the wing-phalange 1. Phylogenetic analys...","author":[{"@context":"https://schema.org","@type":"Person","name":"Martin Kundrát"}],"contributor":[],"dateCreated":"2023-08-28","dateModified":null,"datePublished":"2016-01-01","headline":"New Material of the Pterosaur Gladocephaloideus Lü et al., 2012 from the Early Cretaceous of Liaoning Province, China, with Comments on Its Systematic Position","inLanguage":"en","keywords":["Evolutionary Biology","Biology","Ecology","China","Medicine","Multidisciplinary","Phylogeny","Anatomy","Dinosaurs","Animals","Skull","PLoS one","Holotype","Postcranial Skeleton","Ratio","Specimen"],"locationCreated":null,"publication":"PloS one","publisher":{"@context":"https://schema.org","@type":"Organization","name":null},"image":null,"thumbnailUrl":null,"url":"https://www.academia.edu/106057126/New_Material_of_the_Pterosaur_Gladocephaloideus_L%C3%BC_et_al_2012_from_the_Early_Cretaceous_of_Liaoning_Province_China_with_Comments_on_Its_Systematic_Position","sourceOrganization":[{"@context":"https://schema.org","@type":"EducationalOrganization","name":null}]}</script><link rel="stylesheet" media="all" href="//a.academia-assets.com/assets/single_work_page/loswp-102fa537001ba4d8dcd921ad9bd56c474abc201906ea4843e7e7efe9dfbf561d.css" /><link rel="stylesheet" media="all" href="//a.academia-assets.com/assets/design_system/body-8d679e925718b5e8e4b18e9a4fab37f7eaa99e43386459376559080ac8f2856a.css" /><link rel="stylesheet" media="all" href="//a.academia-assets.com/assets/design_system/button-3cea6e0ad4715ed965c49bfb15dedfc632787b32ff6d8c3a474182b231146ab7.css" /><link rel="stylesheet" media="all" 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A nearly complete juvenile specimen of Gladocephaloideus from the Lower Cretaceous Jiufotang Formation of Sihedang, Lingyuan of Liaoning Province is the most complete ctenochasmatoid preserved to date with a skull and postcranial skeleton. Based on the holotype (IG-CAGS 08-07) and the nearly complete new specimen (JPM 2014-004), the diagnosis of Gladocephaloideus is amended: approximately 50 teeth in total with sharp tips; small nasoantorbital opening, occupying approximately 13% of total skull length; ratio of prenarial rostrum length to skull length approximately 0.63; deep groove along the mid-line of the mandibular symphysis; length to width ratio of the longest cervical vertebra = 4.1; ratio of femur length to tibia length = 0.61; tibia as long as the wing-phalange 1. Phylogenetic analys...","publication_date":"2016,,","publication_name":"PloS one"},"document_type":"paper","pre_hit_view_count_baseline":null,"quality":"high","language":"en","title":"New Material of the Pterosaur Gladocephaloideus Lü et al., 2012 from the Early Cretaceous of Liaoning Province, China, with Comments on Its Systematic Position","broadcastable":false,"draft":null,"has_indexable_attachment":true,"indexable":true}}["work"]; window.loswp.workCoauthors = [805759]; window.loswp.locale = "en"; window.loswp.countryCode = "SG"; window.loswp.cwvAbTestBucket = ""; window.loswp.designVariant = "ds_vanilla"; window.loswp.fullPageMobileSutdModalVariant = "full_page_mobile_sutd_modal"; window.loswp.useOptimizedScribd4genScript = false; window.loswp.appleClientId = 'edu.academia.applesignon';</script><script defer="" src="https://accounts.google.com/gsi/client"></script><div class="ds-loswp-container"><div class="ds-work-card--grid-container"><div class="ds-work-card--container js-loswp-work-card"><div class="ds-work-card--cover"><div class="ds-work-cover--wrapper"><div class="ds-work-cover--container"><button class="ds-work-cover--clickable js-swp-download-button" data-signup-modal="{"location":"swp-splash-paper-cover","attachmentId":105355191,"attachmentType":"pdf"}"><img alt="First page of “New Material of the Pterosaur Gladocephaloideus Lü et al., 2012 from the Early Cretaceous of Liaoning Province, China, with Comments on Its Systematic Position”" class="ds-work-cover--cover-thumbnail" src="https://0.academia-photos.com/attachment_thumbnails/105355191/mini_magick20230829-1-p25wse.png?1693286697" /><img alt="PDF Icon" class="ds-work-cover--file-icon" src="//a.academia-assets.com/assets/single_work_splash/adobe.icon-574afd46eb6b03a77a153a647fb47e30546f9215c0ee6a25df597a779717f9ef.svg" /><div class="ds-work-cover--hover-container"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">download</span><p>Download Free PDF</p></div><div class="ds-work-cover--ribbon-container">Download Free PDF</div><div class="ds-work-cover--ribbon-triangle"></div></button></div></div></div><div class="ds-work-card--work-information"><h1 class="ds-work-card--work-title">New Material of the Pterosaur Gladocephaloideus Lü et al., 2012 from the Early Cretaceous of Liaoning Province, China, with Comments on Its Systematic Position</h1><div class="ds-work-card--work-authors ds-work-card--detail"><a class="ds-work-card--author js-wsj-grid-card-author ds2-5-body-md ds2-5-body-link" data-author-id="805759" href="https://independent.academia.edu/MartinKundr%C3%A1t"><img alt="Profile image of Martin Kundrát" class="ds-work-card--author-avatar" src="https://0.academia-photos.com/805759/7383248/8300103/s65_martin.kundr_t.jpg" />Martin Kundrát</a></div><div class="ds-work-card--detail"><p class="ds-work-card--detail ds2-5-body-sm">2016, PloS one</p></div><p class="ds-work-card--work-abstract ds-work-card--detail ds2-5-body-md">Although there are nine genera of ctenochasmatoids reported from the Jehol Biota, at present each is known from a specimen that has either a skull or a relatively complete postcranial skeleton. A nearly complete juvenile specimen of Gladocephaloideus from the Lower Cretaceous Jiufotang Formation of Sihedang, Lingyuan of Liaoning Province is the most complete ctenochasmatoid preserved to date with a skull and postcranial skeleton. Based on the holotype (IG-CAGS 08-07) and the nearly complete new specimen (JPM 2014-004), the diagnosis of Gladocephaloideus is amended: approximately 50 teeth in total with sharp tips; small nasoantorbital opening, occupying approximately 13% of total skull length; ratio of prenarial rostrum length to skull length approximately 0.63; deep groove along the mid-line of the mandibular symphysis; length to width ratio of the longest cervical vertebra = 4.1; ratio of femur length to tibia length = 0.61; tibia as long as the wing-phalange 1. Phylogenetic analys...</p><div class="ds-work-card--button-container"><button class="ds2-5-button js-swp-download-button" data-signup-modal="{"location":"continue-reading-button--work-card","attachmentId":105355191,"attachmentType":"pdf","workUrl":"https://www.academia.edu/106057126/New_Material_of_the_Pterosaur_Gladocephaloideus_L%C3%BC_et_al_2012_from_the_Early_Cretaceous_of_Liaoning_Province_China_with_Comments_on_Its_Systematic_Position"}">See full PDF</button><button class="ds2-5-button ds2-5-button--secondary js-swp-download-button" data-signup-modal="{"location":"download-pdf-button--work-card","attachmentId":105355191,"attachmentType":"pdf","workUrl":"https://www.academia.edu/106057126/New_Material_of_the_Pterosaur_Gladocephaloideus_L%C3%BC_et_al_2012_from_the_Early_Cretaceous_of_Liaoning_Province_China_with_Comments_on_Its_Systematic_Position"}"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">download</span>Download 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href="https://www.academia.edu/125699395/A_new_toothless_pterosaur_from_the_Early_Cretaceous_Jehol_Biota_with_comments_on_the_Chaoyangopteridae"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="6" data-entity-id="120536768" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/120536768/A_new_specimen_ofSinopterus_dongi_Pterosauria_Tapejaridae_from_the_Jiufotang_Formation_Early_Cretaceous_China_">A new specimen ofSinopterus dongi(Pterosauria, Tapejaridae) from the Jiufotang Formation (Early Cretaceous, China)</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="805759" href="https://independent.academia.edu/MartinKundr%C3%A1t">Martin Kundrát</a></div><p class="ds-related-work--metadata ds2-5-body-xs">PeerJ, 2021</p><p class="ds-related-work--abstract ds2-5-body-sm">The Tapejarinae are edentulous pterosaurs that are relatively common in Cretaceous continental deposits in South America, North Africa, Europe, and China (mostly Early Cretaceous). The Chinese Jiufotang Formation is particularly rich in tapejarine specimens, having yielded over 10 described specimens and dozens of undescribed ones. For the Jiufotang Formation, a total of seven nominal tapejarid species and two genera have been proposed. Some debate exists over how many of those are valid or, alternatively, sexual or ontogenetic morphs of fewer (or even a single) species. Despite the abundance of specimens and the relevant taxonomic problems involved, detailed revisions of the matter are still lacking. This is partly due to the relatively scarce knowledge on the comparative osteology of theSinopteruscomplex, which is hampered by the fact that most specimens have been only preliminarily described. In this contribution, we present a new postcranial specimen, D3072, which we attribute t...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A new specimen ofSinopterus dongi(Pterosauria, Tapejaridae) from the Jiufotang Formation (Early Cretaceous, China)","attachmentId":115654023,"attachmentType":"pdf","work_url":"https://www.academia.edu/120536768/A_new_specimen_ofSinopterus_dongi_Pterosauria_Tapejaridae_from_the_Jiufotang_Formation_Early_Cretaceous_China_","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/120536768/A_new_specimen_ofSinopterus_dongi_Pterosauria_Tapejaridae_from_the_Jiufotang_Formation_Early_Cretaceous_China_"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="7" data-entity-id="112017444" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/112017444/A_new_specimen_of_i_Sinopterus_dongi_i_Pterosauria_Tapejaridae_from_the_Jiufotang_Formation_Early_Cretaceous_China_">A new specimen of<i>Sinopterus dongi</i>(Pterosauria, Tapejaridae) from the Jiufotang Formation (Early Cretaceous, China)</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="129681083" href="https://independent.academia.edu/MKundr%C3%A1t">Martin Kundrát</a></div><p class="ds-related-work--metadata ds2-5-body-xs">PeerJ, 2021</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A new specimen of\u003ci\u003eSinopterus dongi\u003c/i\u003e(Pterosauria, Tapejaridae) from the Jiufotang Formation (Early Cretaceous, China)","attachmentId":109379465,"attachmentType":"pdf","work_url":"https://www.academia.edu/112017444/A_new_specimen_of_i_Sinopterus_dongi_i_Pterosauria_Tapejaridae_from_the_Jiufotang_Formation_Early_Cretaceous_China_","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/112017444/A_new_specimen_of_i_Sinopterus_dongi_i_Pterosauria_Tapejaridae_from_the_Jiufotang_Formation_Early_Cretaceous_China_"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="8" data-entity-id="3139318" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/3139318/A_new_pterosaur_from_the_Liaoning_Province_of_China_the_phylogeny_of_the_Pterodactyloidea_and_convergence_in_their_cervical_vertebrae">A new pterosaur from the Liaoning Province of China, the phylogeny of the Pterodactyloidea, and convergence in their cervical vertebrae</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="1336790" href="https://usf.academia.edu/BrianAndres">Brian Andres</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Palaeontology, 2008</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A new pterosaur from the Liaoning Province of China, the phylogeny of the Pterodactyloidea, and convergence in their cervical vertebrae","attachmentId":31048712,"attachmentType":"pdf","work_url":"https://www.academia.edu/3139318/A_new_pterosaur_from_the_Liaoning_Province_of_China_the_phylogeny_of_the_Pterodactyloidea_and_convergence_in_their_cervical_vertebrae","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/3139318/A_new_pterosaur_from_the_Liaoning_Province_of_China_the_phylogeny_of_the_Pterodactyloidea_and_convergence_in_their_cervical_vertebrae"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="9" data-entity-id="11134224" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/11134224/A_possible_azhdarchid_pterosaur_from_the_Lower_Cretaceous_Qingshan_Group_of_Laiyang_Shandong_China">A possible azhdarchid pterosaur from the Lower Cretaceous Qingshan Group of Laiyang, Shandong, China</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="26889494" href="https://214.academia.edu/ChangFuZhou">Chang-Fu Zhou</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Journal of Vertebrate Paleontology, 2010</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A possible azhdarchid pterosaur from the Lower Cretaceous Qingshan Group of Laiyang, Shandong, China","attachmentId":46870959,"attachmentType":"pdf","work_url":"https://www.academia.edu/11134224/A_possible_azhdarchid_pterosaur_from_the_Lower_Cretaceous_Qingshan_Group_of_Laiyang_Shandong_China","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" 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class="ds-related-work--metadata ds2-5-body-xs">Proceedings of the Royal Society B: Biological Sciences, 2010</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A longirostrine tyrannosauroid from the Early Cretaceous of China","attachmentId":66966988,"attachmentType":"pdf","work_url":"https://www.academia.edu/48262291/A_longirostrine_tyrannosauroid_from_the_Early_Cretaceous_of_China","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-related-work-grid-card-view-pdf" href="https://www.academia.edu/48262291/A_longirostrine_tyrannosauroid_from_the_Early_Cretaceous_of_China"><span class="ds2-5-text-link__content">View PDF</span><span 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