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"https://www.academia.edu/login?post_login_redirect_url=https%3A%2F%2Fwww.academia.edu%2F11134235%2FA_new_species_of_Psittacosaurus_Dinosauria_Ceratopsia_from_the_Early_Cretaceous_Yixian_Formation_Liaoning_China%3Fauto%3Ddownload"; window.loswp.translateUrl = "https://www.academia.edu/login?post_login_redirect_url=https%3A%2F%2Fwww.academia.edu%2F11134235%2FA_new_species_of_Psittacosaurus_Dinosauria_Ceratopsia_from_the_Early_Cretaceous_Yixian_Formation_Liaoning_China%3Fshow_translation%3Dtrue"; window.loswp.previewableAttachments = [{"id":36791620,"identifier":"Attachment_36791620","shouldShowBulkDownload":false}]; window.loswp.shouldDetectTimezone = true; window.loswp.shouldShowBulkDownload = true; window.loswp.showSignupCaptcha = false window.loswp.willEdgeCache = false; window.loswp.work = {"work":{"id":11134235,"created_at":"2015-02-27T01:19:55.438-08:00","from_world_paper_id":132291196,"updated_at":"2024-11-16T16:50:33.448-08:00","_data":{"grobid_abstract":"Newly discovered, well-preserved skulls and mandibles from the lowest part of the Early Cretaceous Yixian Formation, western Liaoning Province, China, document the earliest known record of psittacosaurs and provide the basis for recognition of a new species, Psittacosaurus lujiatunensis. This discovery increases the taxonomic diversity of psittacosaurs to eight valid species and extends the stratigraphic range of the family Psittacosauridae from late Barremian-Albian to Hauterivian. Comparative study of the new species with other well-known psittacosaurs indicates that the new taxon is probably the most basal member of the psittacosaur clade now known. This interpretation is in agreement with the temporal distribution of the clade and supports the hypothesis that the Psittacosauridae originated from the basal ceratopsian stem no later than the earliest part of the Cretaceous.","publication_date":"2006,,","publication_name":"Palaeoworld","grobid_abstract_attachment_id":"36791620"},"document_type":"paper","pre_hit_view_count_baseline":null,"quality":"high","language":"en","title":"A new species of Psittacosaurus (Dinosauria: Ceratopsia) from the Early Cretaceous Yixian Formation, Liaoning, China","broadcastable":true,"draft":null,"has_indexable_attachment":true,"indexable":true}}["work"]; window.loswp.workCoauthors = [26889494]; window.loswp.locale = "en"; window.loswp.countryCode = "SG"; window.loswp.cwvAbTestBucket = ""; window.loswp.designVariant = "ds_vanilla"; window.loswp.fullPageMobileSutdModalVariant = "control"; window.loswp.useOptimizedScribd4genScript = false; window.loswp.appleClientId = 'edu.academia.applesignon';</script><script defer="" src="https://accounts.google.com/gsi/client"></script><div class="ds-loswp-container"><div class="ds-work-card--grid-container"><div class="ds-work-card--container js-loswp-work-card"><div class="ds-work-card--cover"><div class="ds-work-cover--wrapper"><div class="ds-work-cover--container"><button class="ds-work-cover--clickable js-swp-download-button" data-signup-modal="{"location":"swp-splash-paper-cover","attachmentId":36791620,"attachmentType":"pdf"}"><img alt="First page of “A new species of Psittacosaurus (Dinosauria: Ceratopsia) from the Early Cretaceous Yixian Formation, Liaoning, China”" class="ds-work-cover--cover-thumbnail" src="https://0.academia-photos.com/attachment_thumbnails/36791620/mini_magick20190307-26410-d2llh4.png?1551953443" /><img alt="PDF Icon" class="ds-work-cover--file-icon" src="//a.academia-assets.com/assets/single_work_splash/adobe.icon-574afd46eb6b03a77a153a647fb47e30546f9215c0ee6a25df597a779717f9ef.svg" /><div class="ds-work-cover--hover-container"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">download</span><p>Download Free PDF</p></div><div class="ds-work-cover--ribbon-container">Download Free PDF</div><div class="ds-work-cover--ribbon-triangle"></div></button></div></div></div><div class="ds-work-card--work-information"><h1 class="ds-work-card--work-title">A new species of Psittacosaurus (Dinosauria: Ceratopsia) from the Early Cretaceous Yixian Formation, Liaoning, China</h1><div class="ds-work-card--work-authors ds-work-card--detail"><a class="ds-work-card--author js-wsj-grid-card-author ds2-5-body-md ds2-5-body-link" data-author-id="26889494" href="https://214.academia.edu/ChangFuZhou"><img alt="Profile image of Chang-Fu Zhou" class="ds-work-card--author-avatar" src="//a.academia-assets.com/images/s65_no_pic.png" />Chang-Fu Zhou</a></div><div class="ds-work-card--detail"><p class="ds-work-card--detail ds2-5-body-sm">2006, Palaeoworld</p><div class="ds-work-card--work-metadata"><div class="ds-work-card--work-metadata__stat"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">visibility</span><p class="ds2-5-body-sm" id="work-metadata-view-count">…</p></div><div class="ds-work-card--work-metadata__stat"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">description</span><p class="ds2-5-body-sm">15 pages</p></div><div class="ds-work-card--work-metadata__stat"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">link</span><p class="ds2-5-body-sm">1 file</p></div></div><script>(async () => { const workId = 11134235; 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The pathological fibula is significantly different from a normal fibula, being shorter and much stouter, and with a distinct swelling on the shaft. The swollen portion (representing a mass of necrotic bone) of the fibula indicates that the animal survived for a relatively long time after becoming</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A new Psittacosaurus (Dinosauria: Ceratopsia) specimen from the Yixian Formation of western Liaoning, China: the first pathological psittacosaurid","attachmentId":99095952,"attachmentType":"pdf","work_url":"https://www.academia.edu/97487972/A_new_Psittacosaurus_Dinosauria_Ceratopsia_specimen_from_the_Yixian_Formation_of_western_Liaoning_China_the_first_pathological_psittacosaurid","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/97487972/A_new_Psittacosaurus_Dinosauria_Ceratopsia_specimen_from_the_Yixian_Formation_of_western_Liaoning_China_the_first_pathological_psittacosaurid"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="1" data-entity-id="11134116" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/11134116/Endocranial_morphology_of_psittacosaurs_Dinosauria_Ceratopsia_based_on_CT_scans_of_new_fossils_from_the_Lower_Cretaceous_China">Endocranial morphology of psittacosaurs (Dinosauria: Ceratopsia) based on CT scans of new fossils from the Lower Cretaceous, China</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="26889494" href="https://214.academia.edu/ChangFuZhou">Chang-Fu Zhou</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Palaeoworld, 2007</p><p class="ds-related-work--abstract ds2-5-body-sm">Psittacosaurs, small basal ceratopsians with a parrot-like beak, are among the most abundant dinosaurs, but occur only in the Early Cretaceous of East Asia. Although the general morphology of psittacosaurs is fairly well understood, the endocranial anatomy of the group has never been described. New discoveries of well-preserved skulls from the celebrated Liaoning beds in northeastern China provide the material for conducting research on psittacosaur endocranial morphology. Using computed tomography scans of three-dimensionally preserved skulls, this study reveals basic endocranial anatomy of psittacosaurs and provides the first palaeoneurological evidence of psittacosaurs in relation to their behaviour. Although commonly believed to have had a small brain and small eyes, psittacosaurs had relatively high brain/body size ratios that are comparable to those in the large theropod Tyrannosaurus, and probably had a keen sense of smell and acute vision, as evidenced by their enlarged olfactory lobes and bulbous optic lobes. The configuration of the semicircular canals agrees with limb proportions to suggest that psittacosaurs were agile animals, perhaps better able to escape predation by carnivorous dinosaurs on that account. The behavioural adaptations implied by this study may have been crucial for the successful radiation of psittacosaurs during the Early Cretaceous of East Asia.</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Endocranial morphology of psittacosaurs (Dinosauria: Ceratopsia) based on CT scans of new fossils from the Lower Cretaceous, China","attachmentId":46871021,"attachmentType":"pdf","work_url":"https://www.academia.edu/11134116/Endocranial_morphology_of_psittacosaurs_Dinosauria_Ceratopsia_based_on_CT_scans_of_new_fossils_from_the_Lower_Cretaceous_China","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/11134116/Endocranial_morphology_of_psittacosaurs_Dinosauria_Ceratopsia_based_on_CT_scans_of_new_fossils_from_the_Lower_Cretaceous_China"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="2" data-entity-id="11134224" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/11134224/A_possible_azhdarchid_pterosaur_from_the_Lower_Cretaceous_Qingshan_Group_of_Laiyang_Shandong_China">A possible azhdarchid pterosaur from the Lower Cretaceous Qingshan Group of Laiyang, Shandong, China</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="26889494" href="https://214.academia.edu/ChangFuZhou">Chang-Fu Zhou</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Journal of Vertebrate Paleontology, 2010</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A possible azhdarchid pterosaur from the Lower Cretaceous Qingshan Group of Laiyang, Shandong, China","attachmentId":46870959,"attachmentType":"pdf","work_url":"https://www.academia.edu/11134224/A_possible_azhdarchid_pterosaur_from_the_Lower_Cretaceous_Qingshan_Group_of_Laiyang_Shandong_China","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/11134224/A_possible_azhdarchid_pterosaur_from_the_Lower_Cretaceous_Qingshan_Group_of_Laiyang_Shandong_China"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="3" data-entity-id="106057126" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/106057126/New_Material_of_the_Pterosaur_Gladocephaloideus_L%C3%BC_et_al_2012_from_the_Early_Cretaceous_of_Liaoning_Province_China_with_Comments_on_Its_Systematic_Position">New Material of the Pterosaur Gladocephaloideus Lü et al., 2012 from the Early Cretaceous of Liaoning Province, China, with Comments on Its Systematic Position</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="805759" href="https://independent.academia.edu/MartinKundr%C3%A1t">Martin Kundrát</a></div><p class="ds-related-work--metadata ds2-5-body-xs">PloS one, 2016</p><p class="ds-related-work--abstract ds2-5-body-sm">Although there are nine genera of ctenochasmatoids reported from the Jehol Biota, at present each is known from a specimen that has either a skull or a relatively complete postcranial skeleton. A nearly complete juvenile specimen of Gladocephaloideus from the Lower Cretaceous Jiufotang Formation of Sihedang, Lingyuan of Liaoning Province is the most complete ctenochasmatoid preserved to date with a skull and postcranial skeleton. Based on the holotype (IG-CAGS 08-07) and the nearly complete new specimen (JPM 2014-004), the diagnosis of Gladocephaloideus is amended: approximately 50 teeth in total with sharp tips; small nasoantorbital opening, occupying approximately 13% of total skull length; ratio of prenarial rostrum length to skull length approximately 0.63; deep groove along the mid-line of the mandibular symphysis; length to width ratio of the longest cervical vertebra = 4.1; ratio of femur length to tibia length = 0.61; tibia as long as the wing-phalange 1. Phylogenetic analys...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"New Material of the Pterosaur Gladocephaloideus Lü et al., 2012 from the Early Cretaceous of Liaoning Province, China, with Comments on Its Systematic Position","attachmentId":105355191,"attachmentType":"pdf","work_url":"https://www.academia.edu/106057126/New_Material_of_the_Pterosaur_Gladocephaloideus_L%C3%BC_et_al_2012_from_the_Early_Cretaceous_of_Liaoning_Province_China_with_Comments_on_Its_Systematic_Position","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/106057126/New_Material_of_the_Pterosaur_Gladocephaloideus_L%C3%BC_et_al_2012_from_the_Early_Cretaceous_of_Liaoning_Province_China_with_Comments_on_Its_Systematic_Position"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="4" data-entity-id="124586170" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/124586170/Psittacosaurus_amitabha_a_New_Species_of_Ceratopsian_Dinosaur_from_the_Ondai_Sayr_Locality_Central_Mongolia">Psittacosaurus amitabha, a New Species of Ceratopsian Dinosaur from the Ondai Sayr Locality, Central Mongolia</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="207818290" 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class="ds-related-work--metadata ds2-5-body-xs">Nature, 2005</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Pterosaur diversity and faunal turnover in Cretaceous terrestrial ecosystems in China","attachmentId":69048440,"attachmentType":"pdf","work_url":"https://www.academia.edu/51244165/Pterosaur_diversity_and_faunal_turnover_in_Cretaceous_terrestrial_ecosystems_in_China","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-related-work-grid-card-view-pdf" href="https://www.academia.edu/51244165/Pterosaur_diversity_and_faunal_turnover_in_Cretaceous_terrestrial_ecosystems_in_China"><span class="ds2-5-text-link__content">View PDF</span><span 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