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Max Delbrück - Wikipedia

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id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Max Delbrück</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 57 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-57" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">57 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%85%D8%A7%D9%83%D8%B3_%D8%AF%D9%84%D8%A8%D8%B1%D9%88%D9%83" title="ماكس دلبروك – Arabic" lang="ar" hreflang="ar" data-title="ماكس دلبروك" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%85%DA%A9%D8%B3_%D8%AF%D9%84%D8%A8%D8%B1%D9%88%DA%A9" title="مکس دلبروک – South Azerbaijani" lang="azb" hreflang="azb" data-title="مکس دلبروک" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AE%E0%A6%BE%E0%A6%95%E0%A7%8D%E0%A6%B8_%E0%A6%A1%E0%A7%87%E0%A6%B2%E0%A6%AC%E0%A7%8D%E0%A6%B0%E0%A7%8D%E0%A6%AF%E0%A7%81%E0%A6%95" title="মাক্স ডেলব্র্যুক – Bangla" lang="bn" hreflang="bn" data-title="মাক্স ডেলব্র্যুক" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Minnan" lang="nan" hreflang="nan" data-title="Max Delbrück" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%9C%D0%B0%D0%BA%D1%81_%D0%94%D1%8D%D0%BB%D1%8C%D0%B1%D1%80%D1%83%D0%BA" title="Макс Дэльбрук – Belarusian" lang="be" hreflang="be" data-title="Макс Дэльбрук" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9C%D0%B0%D0%BA%D1%81_%D0%94%D0%B5%D0%BB%D0%B1%D1%80%D1%8E%D0%BA" title="Макс Делбрюк – Bulgarian" lang="bg" hreflang="bg" data-title="Макс Делбрюк" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Catalan" lang="ca" hreflang="ca" data-title="Max Delbrück" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Czech" lang="cs" hreflang="cs" data-title="Max Delbrück" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Max_Delbr%C3%BCck_(Biophysiker)" title="Max Delbrück (Biophysiker) – German" lang="de" hreflang="de" data-title="Max Delbrück (Biophysiker)" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Estonian" lang="et" hreflang="et" data-title="Max Delbrück" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%9C%CE%B1%CE%BE_%CE%9D%CF%84%CE%B5%CE%BB%CE%BC%CF%80%CF%81%CE%BF%CF%8D%CE%BA" title="Μαξ Ντελμπρούκ – Greek" lang="el" hreflang="el" data-title="Μαξ Ντελμπρούκ" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Spanish" lang="es" hreflang="es" data-title="Max Delbrück" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Max_Delbruck" title="Max Delbruck – Basque" lang="eu" hreflang="eu" data-title="Max Delbruck" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%85%DA%A9%D8%B3_%D8%AF%D9%84%D8%A8%D8%B1%D9%88%DA%A9" title="مکس دلبروک – Persian" lang="fa" hreflang="fa" data-title="مکس دلبروک" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – French" lang="fr" hreflang="fr" data-title="Max Delbrück" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Irish" lang="ga" hreflang="ga" data-title="Max Delbrück" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%A7%89%EC%8A%A4_%EB%8D%B8%EB%B8%8C%EB%A4%BC%ED%81%AC" title="막스 델브뤼크 – Korean" lang="ko" hreflang="ko" data-title="막스 델브뤼크" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Croatian" lang="hr" hreflang="hr" data-title="Max Delbrück" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Ido" lang="io" hreflang="io" data-title="Max Delbrück" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Max_Ludwig_Henning_Delbr%C3%BCck" title="Max Ludwig Henning Delbrück – Indonesian" lang="id" hreflang="id" data-title="Max Ludwig Henning Delbrück" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Icelandic" lang="is" hreflang="is" data-title="Max Delbrück" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Italian" lang="it" hreflang="it" data-title="Max Delbrück" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%9E%D7%A7%D7%A1_%D7%93%D7%9C%D7%91%D7%A8%D7%99%D7%A7" title="מקס דלבריק – Hebrew" lang="he" hreflang="he" data-title="מקס דלבריק" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%9C%D0%B0%D0%BA%D1%81_%D0%94%D0%B5%D0%BB%D1%8C%D0%B1%D1%80%D1%8E%D0%BA" title="Макс Дельбрюк – Kazakh" lang="kk" hreflang="kk" data-title="Макс Дельбрюк" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Max_Delbruck" title="Max Delbruck – Swahili" lang="sw" hreflang="sw" data-title="Max Delbruck" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ht mw-list-item"><a href="https://ht.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Haitian Creole" lang="ht" hreflang="ht" data-title="Max Delbrück" data-language-autonym="Kreyòl ayisyen" data-language-local-name="Haitian Creole" class="interlanguage-link-target"><span>Kreyòl ayisyen</span></a></li><li class="interlanguage-link interwiki-lb mw-list-item"><a href="https://lb.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Luxembourgish" lang="lb" hreflang="lb" data-title="Max Delbrück" data-language-autonym="Lëtzebuergesch" data-language-local-name="Luxembourgish" class="interlanguage-link-target"><span>Lëtzebuergesch</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Lithuanian" lang="lt" hreflang="lt" data-title="Max Delbrück" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Hungarian" lang="hu" hreflang="hu" data-title="Max Delbrück" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%9B%E1%83%90%E1%83%A5%E1%83%A1_%E1%83%93%E1%83%94%E1%83%9A%E1%83%91%E1%83%A0%E1%83%98%E1%83%A3%E1%83%99%E1%83%98" title="მაქს დელბრიუკი – Mingrelian" lang="xmf" hreflang="xmf" data-title="მაქს დელბრიუკი" data-language-autonym="მარგალური" data-language-local-name="Mingrelian" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%85%D8%A7%D9%83%D8%B3_%D8%AF%D9%84%D8%A8%D8%B1%D9%88%D9%83" title="ماكس دلبروك – Egyptian Arabic" lang="arz" hreflang="arz" data-title="ماكس دلبروك" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%85%DA%A9%D8%B3_%D8%AF%D9%84%D8%A8%D8%B1%D9%88%DA%A9" title="مکس دلبروک – Mazanderani" lang="mzn" hreflang="mzn" data-title="مکس دلبروک" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Malay" lang="ms" hreflang="ms" data-title="Max Delbrück" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Dutch" lang="nl" hreflang="nl" data-title="Max Delbrück" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%9E%E3%83%83%E3%82%AF%E3%82%B9%E3%83%BB%E3%83%87%E3%83%AB%E3%83%96%E3%83%AA%E3%83%A5%E3%83%83%E3%82%AF" title="マックス・デルブリュック – Japanese" lang="ja" hreflang="ja" data-title="マックス・デルブリュック" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Max Delbrück" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Occitan" lang="oc" hreflang="oc" data-title="Max Delbrück" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Uzbek" lang="uz" hreflang="uz" data-title="Max Delbrück" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%85%DB%8C%DA%A9%D8%B3_%DA%88%DB%8C%D9%84%D8%A8%D8%B1%D9%88%DA%A9" title="میکس ڈیلبروک – Western Punjabi" lang="pnb" hreflang="pnb" data-title="میکس ڈیلبروک" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Polish" lang="pl" hreflang="pl" data-title="Max Delbrück" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Portuguese" lang="pt" hreflang="pt" data-title="Max Delbrück" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Romanian" lang="ro" hreflang="ro" data-title="Max Delbrück" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%94%D0%B5%D0%BB%D1%8C%D0%B1%D1%80%D1%8E%D0%BA,_%D0%9C%D0%B0%D0%BA%D1%81" title="Дельбрюк, Макс – Russian" lang="ru" hreflang="ru" data-title="Дельбрюк, Макс" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Simple English" lang="en-simple" hreflang="en-simple" data-title="Max Delbrück" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Slovenian" lang="sl" hreflang="sl" data-title="Max Delbrück" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D9%85%D8%A7%DA%A9%D8%B3_%D8%AF%DB%8E%D9%84%D8%A8%D8%B1%DB%86%DA%A9" title="ماکس دێلبرۆک – Central Kurdish" lang="ckb" hreflang="ckb" data-title="ماکس دێلبرۆک" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%9C%D0%B0%D0%BA%D1%81_%D0%94%D0%B5%D0%BB%D0%B1%D1%80%D0%B8%D0%BA" title="Макс Делбрик – Serbian" lang="sr" hreflang="sr" data-title="Макс Делбрик" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Max_Delbr%C3%BCck" title="Max Delbrück – Finnish" lang="fi" hreflang="fi" data-title="Max Delbrück" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Max_Delbr%C3%BCck_(biofysiker)" title="Max Delbrück (biofysiker) – Swedish" lang="sv" hreflang="sv" data-title="Max Delbrück (biofysiker)" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%9C%D0%B0%D0%BA%D1%81_%D0%94%D0%B5%D0%BB%D0%B1%D1%80%D2%AF%D0%BA" title="Макс Делбрүк – Tatar" lang="tt" hreflang="tt" data-title="Макс Делбрүк" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a 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Yoruba" lang="yo" hreflang="yo" data-title="Max Delbrück" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E9%A6%AC%E5%85%8B%E6%96%AF%C2%B7%E5%BE%B7%E7%88%BE%E5%B8%83%E5%91%82%E5%85%8B" title="馬克斯·德爾布呂克 – Chinese" lang="zh" hreflang="zh" data-title="馬克斯·德爾布呂克" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q76807#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div 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class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Biophysicist</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Not to be confused with his uncle, <a href="/wiki/Max_Delbr%C3%BCck_(chemist)" title="Max Delbrück (chemist)">Max Delbrück (chemist)</a>, an agricultural chemist.</div> <p class="mw-empty-elt"> </p> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Max Delbrück</div><div class="honorific-suffix" style="font-size: 77%; font-weight: normal;"><span class="nobold noexcerpt nowraplinks" style="font-size:;"><span style="font-size: 100%;"><a href="/wiki/Foreign_Member_of_the_Royal_Society" class="mw-redirect" title="Foreign Member of the Royal Society">ForMemRS</a></span></span></div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Maxdelbr%C3%BCck-cr.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/93/Maxdelbr%C3%BCck-cr.jpg/220px-Maxdelbr%C3%BCck-cr.jpg" decoding="async" width="220" height="229" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/93/Maxdelbr%C3%BCck-cr.jpg/330px-Maxdelbr%C3%BCck-cr.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/93/Maxdelbr%C3%BCck-cr.jpg/440px-Maxdelbr%C3%BCck-cr.jpg 2x" data-file-width="1248" data-file-height="1300" /></a></span></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">Max Ludwig Henning Delbrück</div><br /><span style="display:none">(<span class="bday">1906-09-04</span>)</span>September 4, 1906<br /><div style="display:inline" class="birthplace"><a href="/wiki/Berlin" title="Berlin">Berlin</a>, <a href="/wiki/German_Empire" title="German Empire">German Empire</a></div></td></tr><tr><th scope="row" class="infobox-label">Died</th><td class="infobox-data">March 9, 1981<span style="display:none">(1981-03-09)</span> (aged&#160;74)<br /><div style="display:inline" class="deathplace"><a href="/wiki/Pasadena,_California" title="Pasadena, California">Pasadena, California</a>, U.S.</div></td></tr><tr><th scope="row" class="infobox-label">Citizenship</th><td class="infobox-data category"><a href="/wiki/Citizenship_in_the_United_States" class="mw-redirect" title="Citizenship in the United States">United States</a><sup id="cite_ref-brittanica-x1_1-0" class="reference"><a href="#cite_note-brittanica-x1-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup></td></tr><tr><th scope="row" class="infobox-label">Alma&#160;mater</th><td class="infobox-data"><a href="/wiki/University_of_G%C3%B6ttingen" title="University of Göttingen">University of Göttingen</a></td></tr><tr><th scope="row" class="infobox-label">Known&#160;for</th><td class="infobox-data"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style><div class="plainlist"> <ul><li>"discoveries concerning the replication mechanism and the genetic structure of viruses" <small>(Nobel Committee)<sup id="cite_ref-nobel1969rationale_2-0" class="reference"><a href="#cite_note-nobel1969rationale-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup></small></li> <li><a href="/wiki/Delbr%C3%BCck_scattering" title="Delbrück scattering">Delbrück scattering</a></li> <li><a href="/wiki/Luria%E2%80%93Delbr%C3%BCck_experiment" title="Luria–Delbrück experiment">Luria–Delbrück experiment</a></li> <li><a href="/wiki/Saffman%E2%80%93Delbr%C3%BCck_model" title="Saffman–Delbrück model">Saffman–Delbrück model</a></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label">Spouse</th><td class="infobox-data">Mary Bruce</td></tr><tr><th scope="row" class="infobox-label">Children</th><td class="infobox-data">4</td></tr><tr><th scope="row" class="infobox-label">Father</th><td class="infobox-data"><a href="/wiki/Hans_Delbr%C3%BCck" title="Hans Delbrück">Hans Delbrück</a></td></tr><tr><th scope="row" class="infobox-label">Relatives</th><td class="infobox-data"><a href="/wiki/Emmi_Bonhoeffer" title="Emmi Bonhoeffer">Emmi Bonhoeffer</a> (sister)</td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li><a href="/wiki/Mendel_Medal_(genetics)" class="mw-redirect" title="Mendel Medal (genetics)">Mendel Medal</a> (1968)</li> <li><a href="/wiki/Louisa_Gross_Horwitz_Prize" title="Louisa Gross Horwitz Prize">Louisa Gross Horwitz Prize</a> (1969)</li> <li><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> (1969)</li></ul> </div></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Biophysics" title="Biophysics">Biophysics</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/Kaiser_Wilhelm_Society" title="Kaiser Wilhelm Society">Kaiser Wilhelm Institute for Chemistry</a><br /><a href="/wiki/Vanderbilt_University" title="Vanderbilt University">Vanderbilt University</a><br /><a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">Caltech</a></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Doctoral_advisor" title="Doctoral advisor">Doctoral advisor</a></th><td class="infobox-data"><a href="/wiki/Lise_Meitner" title="Lise Meitner">Lise Meitner</a></td></tr><tr><th scope="row" class="infobox-label">Doctoral students</th><td class="infobox-data"><a href="/wiki/Lily_Jan" title="Lily Jan">Lily Jan</a>, <a href="/wiki/Yuh_Nung_Jan" title="Yuh Nung Jan">Yuh Nung Jan</a>, <a href="/wiki/Ernst_Peter_Fischer" title="Ernst Peter Fischer">Ernst Peter Fischer</a>, <a href="/wiki/Charles_M._Steinberg" title="Charles M. Steinberg">Charles M. Steinberg</a></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>Max Ludwig Henning Delbrück</b> (<style data-mw-deduplicate="TemplateStyles:r1177148991">.mw-parser-output .IPA-label-small{font-size:85%}.mw-parser-output .references .IPA-label-small,.mw-parser-output .infobox .IPA-label-small,.mw-parser-output .navbox .IPA-label-small{font-size:100%}</style><span class="IPA-label IPA-label-small">German:</span> <span class="IPA nowrap" lang="de-Latn-fonipa"><a href="/wiki/Help:IPA/Standard_German" title="Help:IPA/Standard German">&#91;maks<span class="wrap"> </span>ˈdɛl.bʁʏk&#93;</a></span> <span class="noprint"><span class="ext-phonos"><span data-nosnippet="" id="ooui-php-1" class="ext-phonos-PhonosButton noexcerpt ext-phonos-PhonosButton-emptylabel oo-ui-widget oo-ui-widget-enabled oo-ui-buttonElement oo-ui-buttonElement-frameless oo-ui-iconElement oo-ui-buttonWidget" data-ooui="{&quot;_&quot;:&quot;mw.Phonos.PhonosButton&quot;,&quot;href&quot;:&quot;\/\/upload.wikimedia.org\/wikipedia\/commons\/transcoded\/c\/cc\/De-Max_Delbr%C3%BCck.ogg\/De-Max_Delbr%C3%BCck.ogg.mp3&quot;,&quot;rel&quot;:[&quot;nofollow&quot;],&quot;framed&quot;:false,&quot;icon&quot;:&quot;volumeUp&quot;,&quot;data&quot;:{&quot;ipa&quot;:&quot;&quot;,&quot;text&quot;:&quot;&quot;,&quot;lang&quot;:&quot;en&quot;,&quot;wikibase&quot;:&quot;&quot;,&quot;file&quot;:&quot;De-Max Delbr\u00fcck.ogg&quot;},&quot;classes&quot;:[&quot;ext-phonos-PhonosButton&quot;,&quot;noexcerpt&quot;,&quot;ext-phonos-PhonosButton-emptylabel&quot;]}"><a role="button" tabindex="0" href="//upload.wikimedia.org/wikipedia/commons/transcoded/c/cc/De-Max_Delbr%C3%BCck.ogg/De-Max_Delbr%C3%BCck.ogg.mp3" rel="nofollow" aria-label="Play audio" title="Play audio" class="oo-ui-buttonElement-button"><span class="oo-ui-iconElement-icon oo-ui-icon-volumeUp"></span><span class="oo-ui-labelElement-label"></span><span class="oo-ui-indicatorElement-indicator oo-ui-indicatorElement-noIndicator"></span></a></span><sup class="ext-phonos-attribution noexcerpt navigation-not-searchable"><a href="/wiki/File:De-Max_Delbr%C3%BCck.ogg" title="File:De-Max Delbrück.ogg">ⓘ</a></sup></span></span>; September 4, 1906 – March 9, 1981) was a <a href="/wiki/German%E2%80%93American" class="mw-redirect" title="German–American">German–American</a> <a href="/wiki/Biophysicist" class="mw-redirect" title="Biophysicist">biophysicist</a> who participated in launching the <a href="/wiki/Molecular_biology" title="Molecular biology">molecular biology</a> <a href="/wiki/Research_program" title="Research program">research program</a> in the late 1930s. He stimulated <a href="/wiki/Physical_science" class="mw-redirect" title="Physical science">physical scientists</a>' interest into <a href="/wiki/Biology" title="Biology">biology</a>, especially as to <a href="/wiki/Basic_research" title="Basic research">basic research</a> to physically explain <a href="/wiki/Genes" class="mw-redirect" title="Genes">genes</a>, mysterious at the time. Formed in 1945 and led by Delbrück along with <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a> and <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a>, the <a href="/wiki/Phage_Group" class="mw-redirect" title="Phage Group">Phage Group</a> made substantial headway unraveling important aspects of <a href="/wiki/Genetics" title="Genetics">genetics</a>. The three shared the 1969 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> "for their discoveries concerning the replication mechanism and the genetic structure of viruses".<sup id="cite_ref-Nobel_3-0" class="reference"><a href="#cite_note-Nobel-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> He was the first physicist to predict what is now called <a href="/wiki/Delbr%C3%BCck_scattering" title="Delbrück scattering">Delbrück scattering</a>.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Early_and_personal_life">Early and personal life</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Max_Delbr%C3%BCck&amp;action=edit&amp;section=1" title="Edit section: Early and personal life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Max_Delbruck.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/b/bb/Max_Delbruck.jpg/170px-Max_Delbruck.jpg" decoding="async" width="170" height="273" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/bb/Max_Delbruck.jpg/255px-Max_Delbruck.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/bb/Max_Delbruck.jpg/340px-Max_Delbruck.jpg 2x" data-file-width="1344" data-file-height="2162" /></a><figcaption>Delbrück in the early 1940s</figcaption></figure> <p>Delbrück was born in <a href="/wiki/Berlin" title="Berlin">Berlin</a>, <a href="/wiki/German_Empire" title="German Empire">German Empire</a>. His mother was granddaughter of <a href="/wiki/Justus_von_Liebig" title="Justus von Liebig">Justus von Liebig</a>, an eminent <a href="/wiki/Chemist" title="Chemist">chemist</a>, while his father <a href="/wiki/Hans_Delbr%C3%BCck" title="Hans Delbrück">Hans Delbrück</a> was a <a href="/wiki/History" title="History">history</a> professor at the <a href="/wiki/University_of_Berlin" class="mw-redirect" title="University of Berlin">University of Berlin</a>. In 1937, Delbrück left Nazi Germany for America—first California, then Tennessee—becoming a <a href="/wiki/Citizenship_in_the_United_States" class="mw-redirect" title="Citizenship in the United States">US citizen</a> in 1945.<sup id="cite_ref-brittanica-x1_1-1" class="reference"><a href="#cite_note-brittanica-x1-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> In 1941, he married Mary Bruce. They had four children. </p><p>Delbrück's brother Justus, a lawyer, as well as his sister <a href="/wiki/Emmi_Bonhoeffer" title="Emmi Bonhoeffer">Emmi Bonhoeffer</a> were active along with his brothers-in-law <a href="/wiki/Klaus_Bonhoeffer" title="Klaus Bonhoeffer">Klaus Bonhoeffer</a> and <a href="/wiki/Dietrich_Bonhoeffer" title="Dietrich Bonhoeffer">Dietrich Bonhoeffer</a> in <a href="/wiki/German_Resistance_to_Nazism" class="mw-redirect" title="German Resistance to Nazism">resistance to Nazism</a>. Found guilty by the <a href="/wiki/People%27s_Court_(Germany)" title="People&#39;s Court (Germany)">People's Court</a> for roles in the <a href="/wiki/20_July_plot" title="20 July plot">July 20, 1944 plot to assassinate Hitler</a>, Dietrich and Klaus were executed in 1945 by the <a href="/wiki/SS-Reichssicherheitshauptamt" class="mw-redirect" title="SS-Reichssicherheitshauptamt">RSHA</a>. Justus died in Soviet custody that same year. His son, Tobias Delbruck is a professor at the Institute of Neuroinformatics at the University of Zurich and ETH Zurich.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> Professor Tobias Delbruck is also one of the pioneers in the domain of event cameras,<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> now increasingly being deployed in dynamic vision systems. </p> <div class="mw-heading mw-heading2"><h2 id="Education">Education</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Max_Delbr%C3%BCck&amp;action=edit&amp;section=2" title="Edit section: Education"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Delbrück studied <a href="/wiki/Astrophysics" title="Astrophysics">astrophysics</a>, shifting towards <a href="/wiki/Theoretical_physics" title="Theoretical physics">theoretical physics</a>, at the <a href="/wiki/University_of_G%C3%B6ttingen" title="University of Göttingen">University of Göttingen</a>. After completing his <a href="/wiki/Doctor_of_Philosophy" title="Doctor of Philosophy">Ph.D.</a> there in 1930,<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> he traveled through <a href="/wiki/England" title="England">England</a>, <a href="/wiki/Denmark" title="Denmark">Denmark</a>, and <a href="/wiki/Switzerland" title="Switzerland">Switzerland</a>. He met <a href="/wiki/Wolfgang_Pauli" title="Wolfgang Pauli">Wolfgang Pauli</a> and <a href="/wiki/Niels_Bohr" title="Niels Bohr">Niels Bohr</a>, who interested him in <a href="/wiki/Biology" title="Biology">biology</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Career_and_research">Career and research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Max_Delbr%C3%BCck&amp;action=edit&amp;section=3" title="Edit section: Career and research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Freie_Universitaet_Berlin_Otto-Hahn-Bau_im_Winter_01-2005.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Freie_Universitaet_Berlin_Otto-Hahn-Bau_im_Winter_01-2005.jpg/220px-Freie_Universitaet_Berlin_Otto-Hahn-Bau_im_Winter_01-2005.jpg" decoding="async" width="220" height="152" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Freie_Universitaet_Berlin_Otto-Hahn-Bau_im_Winter_01-2005.jpg/330px-Freie_Universitaet_Berlin_Otto-Hahn-Bau_im_Winter_01-2005.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Freie_Universitaet_Berlin_Otto-Hahn-Bau_im_Winter_01-2005.jpg/440px-Freie_Universitaet_Berlin_Otto-Hahn-Bau_im_Winter_01-2005.jpg 2x" data-file-width="1279" data-file-height="885" /></a><figcaption>Delbrück's workplace in Berlin: Kaiser Wilhelm Institute for Chemistry, now the <a href="/wiki/Free_University_of_Berlin" title="Free University of Berlin">Free University of Berlin</a>.</figcaption></figure> <p>Delbrück returned to Berlin in 1932 as an assistant to <a href="/wiki/Lise_Meitner" title="Lise Meitner">Lise Meitner</a>, who was collaborating with <a href="/wiki/Otto_Hahn" title="Otto Hahn">Otto Hahn</a> on irradiation of <a href="/wiki/Uranium" title="Uranium">uranium</a> with <a href="/wiki/Neutrons" class="mw-redirect" title="Neutrons">neutrons</a>. Delbrück wrote a few papers, including one in 1933 on <a href="/wiki/Gamma_rays" class="mw-redirect" title="Gamma rays">gamma rays</a>' scattering by a <a href="/wiki/Coulomb" title="Coulomb">Coulomb</a> field's polarization of a vacuum. Though theoretically tenable, his conclusion was misplaced, whereas <a href="/wiki/Hans_Bethe" title="Hans Bethe">Hans Bethe</a> some 20 years later confirmed the phenomenon and named it "<a href="/wiki/Delbr%C3%BCck_scattering" title="Delbrück scattering">Delbrück scattering</a>".<sup id="cite_ref-NAS_10-0" class="reference"><a href="#cite_note-NAS-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> </p><p>In 1935, Delbrück published a collaboration with <a href="/wiki/Nikolay_Timofeev-Ressovsky" title="Nikolay Timofeev-Ressovsky">Nikolay Timofeev-Ressovsky</a> and <a href="/wiki/Karl_Zimmer" title="Karl Zimmer">Karl Zimmer</a> the major work, <i>Über die Natur der Genmutation und der Genstruktur</i>. It was considered to be a major advance in understanding the nature of gene mutation and gene structure.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> The work was a keystone in the formation of molecular genetics.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> It was also an inspirational starting point for <a href="/wiki/Erwin_Schr%C3%B6dinger" title="Erwin Schrödinger"> Erwin Schrödinger's</a> thinking, a course of lectures in 1943, and the eventual writing of the book <i>What Is Life? The Physical Aspect of the Living Cell</i>.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> </p><p>In 1937, he attained a fellowship from <a href="/wiki/Rockefeller_Foundation" title="Rockefeller Foundation">Rockefeller Foundation</a>—which was launching the <a href="/wiki/Molecular_biology" title="Molecular biology">molecular biology</a> <a href="/wiki/Research_program" title="Research program">research program</a>—to research <a href="/wiki/Genetics" title="Genetics">genetics</a> of the fruit fly, <i><a href="/wiki/Drosophila_melanogaster" title="Drosophila melanogaster">Drosophila melanogaster</a></i>, in <a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">California Institute of Technology</a>'s biology department,<sup id="cite_ref-MDC_14-0" class="reference"><a href="#cite_note-MDC-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> where Delbrück could blend interests in biochemistry and genetics.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> While at <a href="/wiki/Caltech" class="mw-redirect" title="Caltech">Caltech</a>, Delbrück researched <a href="/wiki/Bacteria" title="Bacteria">bacteria</a> and their viruses (<a href="/wiki/Bacteriophages" class="mw-redirect" title="Bacteriophages">bacteriophages</a> or <i>phages</i>). In 1939, with <a href="/wiki/Emory_Ellis" title="Emory Ellis">Emory L. Ellis</a>,<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">&#91;</span>17<span class="cite-bracket">&#93;</span></a></sup> he coauthored "The growth of bacteriophage", a paper reporting that the <a href="/wiki/Virus" title="Virus">viruses</a> reproduce in <i>one step</i>, not exponentially as do <a href="/wiki/Cell_(biology)" title="Cell (biology)">cellular</a> organisms. </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Plaque_in_Buttrick_Hall.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0a/Plaque_in_Buttrick_Hall.png/220px-Plaque_in_Buttrick_Hall.png" decoding="async" width="220" height="170" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0a/Plaque_in_Buttrick_Hall.png/330px-Plaque_in_Buttrick_Hall.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0a/Plaque_in_Buttrick_Hall.png/440px-Plaque_in_Buttrick_Hall.png 2x" data-file-width="1054" data-file-height="816" /></a><figcaption>Drawing of a plaque in Buttrick Hall, Vanderbilt University commemorating the work of Max Delbrück.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup></figcaption></figure> <p>Although Delbrück's <a href="/wiki/Rockefeller_Foundation" title="Rockefeller Foundation">Rockefeller Foundation</a> fellowship expired in 1939, the Foundation matched him up with <a href="/wiki/Vanderbilt_University" title="Vanderbilt University">Vanderbilt University</a> in <a href="/wiki/Nashville,_Tennessee" title="Nashville, Tennessee">Nashville, Tennessee</a>, where from 1940 to 1947 he taught <a href="/wiki/Physics" title="Physics">physics</a>, yet had his laboratory in the biology department.<sup id="cite_ref-Vanderbilt_19-0" class="reference"><a href="#cite_note-Vanderbilt-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> In 1941, Delbrück met <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a> of <a href="/wiki/Indiana_University" title="Indiana University">Indiana University</a> who began visiting Vanderbilt.<sup id="cite_ref-Vanderbilt_19-1" class="reference"><a href="#cite_note-Vanderbilt-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> In 1942, <a href="/wiki/Luria%E2%80%93Delbr%C3%BCck_experiment" title="Luria–Delbrück experiment">Delbrück and Luria published</a> on <a href="/wiki/Bacteria" title="Bacteria">bacterial</a> resistance to <a href="/wiki/Virus" title="Virus">virus</a> infection mediated by random <a href="/wiki/Mutation" title="Mutation">mutation</a>.<sup id="cite_ref-Vanderbilt_19-2" class="reference"><a href="#cite_note-Vanderbilt-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> of <a href="/wiki/Washington_University_in_St._Louis" title="Washington University in St. Louis">Washington University in St. Louis</a> began visiting in 1943.<sup id="cite_ref-Vanderbilt_19-3" class="reference"><a href="#cite_note-Vanderbilt-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> The Luria–Delbrück experiment, also called the Fluctuation Test, demonstrated that Darwin's theory of natural selection acting on random mutations applies to bacteria as well as to more complex organisms. The 1969 Nobel Prize in Physiology or Medicine was awarded to both scientists in part for this work. To put this work in its historical perspective, Lamarck in 1801 first presented his theory of Inheritance of Acquired Characteristics, which stated that if an organism changes during life in order to adapt to its environment (for example stretches its neck to reach for tall trees), those changes are passed on to its offspring. He also said that evolution happens according to a predetermined plan. Darwin published his theory of evolution in his 1859 book On the Origin of Species with compelling evidence contradicting Lamarck. Darwin said that evolution is not predetermined but that there are inherent variations in all organisms, and that those variations that confer increased fitness are selected by the environment and passed on to the offspring. In the feud between Lamarck and Darwin, Darwin talked of pre-existing changes, but the nature of these changes was not known and had to await the science of genetics by Gregor Mendel's experiments on pea plants published in 1866. Support for Darwin's theory was provided when Thomas Hunt Morgan discovered that a mutated white-eyed fruit fly among red-eyed flies was able to reproduce true white-eyed offspring. The most elegant and convincing support for Darwin's ideas, however, was provided by the Luria-Delbruck experiment,<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">&#91;</span>20<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup> which showed that mutations conferring resistance of the bacterium E. coli to T1 bacteriophage (virus) existed in the population prior to exposure to T1 and were not induced by adding T1. In other words, mutations are random events that occur whether or not they prove to be useful, while selection (for T1 resistance upon challenge with T1 in this case) provides the direction in evolution by retaining those mutations that are advantageous, discarding those that are harmful (T1 sensitivity in this case). This experiment dealt a blow to Lamarckian inheritance and set the stage for tremendous advances in genetics and molecular biology, launching a tsunami of research that eventually led to the discovery of DNA as the hereditary material and to cracking the genetic code. Of course, by then Avery, along with McCloud (and earlier, McCarty) was well on the way to showing the genetic capability of DNA. </p><p>In 1945, Delbrück, Luria, and Hershey set up a course in <a href="/wiki/Bacteriophage" title="Bacteriophage">bacteriophage</a> genetics at <a href="/wiki/Cold_Spring_Harbor_Laboratory" title="Cold Spring Harbor Laboratory">Cold Spring Harbor Laboratory</a> on <a href="/wiki/Long_Island,_New_York" class="mw-redirect" title="Long Island, New York">Long Island, New York</a>.<sup id="cite_ref-Vanderbilt_19-4" class="reference"><a href="#cite_note-Vanderbilt-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> This <a href="/wiki/Phage_Group" class="mw-redirect" title="Phage Group">Phage Group</a> spurred <a href="/wiki/Molecular_biology" title="Molecular biology">molecular biology</a>'s early development.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">&#91;</span>23<span class="cite-bracket">&#93;</span></a></sup> Delbrück received the 1969 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a>, shared with Luria and Hershey "for their discoveries concerning the replication mechanism and the genetic structure of viruses".<sup id="cite_ref-Nobel_3-1" class="reference"><a href="#cite_note-Nobel-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Vanderbilt_19-5" class="reference"><a href="#cite_note-Vanderbilt-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-FischerLipson_24-0" class="reference"><a href="#cite_note-FischerLipson-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup> The committee also noted that "The honour in the first place goes to Delbrück who transformed bacteriophage research from vague empiricism to an exact science. He analyzed and defined the conditions for precise measurement of the biological effects. Together with Luria he elaborated the quantitative methods and established the statistical criteria for evaluation which made the subsequent penetrating studies possible. Delbrück's and Luria's forte is perhaps mainly theoretical analysis, whereas Hershey above all is an eminently skillful experimenter. The three of them supplement each other well also in these respects." That year, Delbrück and <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> were also awarded by <a href="/wiki/Columbia_University" title="Columbia University">Columbia University</a> the <a href="/wiki/Louisa_Gross_Horwitz_Prize" title="Louisa Gross Horwitz Prize">Louisa Gross Horwitz Prize</a>. In late 1947, as Vanderbilt lacked the resources to keep him, Delbrück had returned to Caltech as a professor of biology, and remained there for the rest of his career.<sup id="cite_ref-Vanderbilt_19-6" class="reference"><a href="#cite_note-Vanderbilt-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> Meanwhile, he set up <a href="/wiki/University_of_Cologne" title="University of Cologne">University of Cologne</a>'s institute for <a href="/wiki/Molecular_genetics" title="Molecular genetics">molecular genetics</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Awards_and_honours">Awards and honours</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Max_Delbr%C3%BCck&amp;action=edit&amp;section=4" title="Edit section: Awards and honours"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In addition to the Nobel Prize, Delbrück was elected a <a href="/wiki/List_of_Fellows_of_the_Royal_Society_elected_in_1967" class="mw-redirect" title="List of Fellows of the Royal Society elected in 1967">Foreign Member of the Royal Society (ForMemRS) in 1967</a>.<sup id="cite_ref-frs_25-0" class="reference"><a href="#cite_note-frs-25"><span class="cite-bracket">&#91;</span>25<span class="cite-bracket">&#93;</span></a></sup> He was elected an <a href="/wiki/EMBO_Member" class="mw-redirect" title="EMBO Member">EMBO Member</a> in 1970.<sup id="cite_ref-membo_26-0" class="reference"><a href="#cite_note-membo-26"><span class="cite-bracket">&#91;</span>26<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Max_Delbruck_Prize" title="Max Delbruck Prize">Max Delbruck Prize</a>, formerly known as the biological physics prize, is awarded by the <a href="/wiki/American_Physical_Society" title="American Physical Society">American Physical Society</a> and named in his honour. The <a href="/wiki/Max_Delbr%C3%BCck_Center_for_Molecular_Medicine_in_the_Helmholtz_Association" title="Max Delbrück Center for Molecular Medicine in the Helmholtz Association">Max Delbrück Center</a>, in Berlin, Germany, national research center for molecular medicine of the <a href="/wiki/Helmholtz_Association" title="Helmholtz Association">Helmholtz Association</a> also bears his name. </p> <div class="mw-heading mw-heading2"><h2 id="Later_life_and_legacy">Later life and legacy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Max_Delbr%C3%BCck&amp;action=edit&amp;section=5" title="Edit section: Later life and legacy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Delbrück helped spur <a href="/wiki/Physical_science" class="mw-redirect" title="Physical science">physical scientists</a>' interest in biology. His inferences on genes' susceptibility to mutation was relied on by physicist <a href="/wiki/Erwin_Schr%C3%B6dinger" title="Erwin Schrödinger">Erwin Schrödinger</a> in his 1944 book <i><a href="/wiki/What_is_Life%3F_(Schr%C3%B6dinger)" class="mw-redirect" title="What is Life? (Schrödinger)">What Is Life?</a></i>,<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> which conjectured genes were an "aperiodic crystal" storing codescript and influenced crystallographer <a href="/wiki/Rosalind_Franklin" title="Rosalind Franklin">Rosalind Franklin</a> and biologists <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> and <a href="/wiki/James_D._Watson" class="mw-redirect" title="James D. Watson">James D. Watson</a> in their 1953 identification of cellular DNA's molecular structure as a double helix.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Judson_29-0" class="reference"><a href="#cite_note-Judson-29"><span class="cite-bracket">&#91;</span>29<span class="cite-bracket">&#93;</span></a></sup> In 1977, he retired from <a href="/wiki/Caltech" class="mw-redirect" title="Caltech">Caltech</a>, remaining a Professor of Biology emeritus. He became interested in the behavioral sciences and spent some unfruitful effort on mold behavior in the 1960s. </p><p>Max Delbrück died, at age 74, on the evening of Monday, March 9, 1981, at <a href="/wiki/Huntington_Memorial_Hospital" class="mw-redirect" title="Huntington Memorial Hospital">Huntington Memorial Hospital</a> in <a href="/wiki/Pasadena,_California" title="Pasadena, California">Pasadena, California</a>. On August 26 to 27, 2006—the year Delbrück would have turned 100—family and friends gathered at Cold Spring Harbor Laboratory to reminisce on his life and work.<sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">&#91;</span>30<span class="cite-bracket">&#93;</span></a></sup> Although Delbrück held some anti-<a href="/wiki/Reductionism" title="Reductionism">reductionist</a> views; he conjectured that ultimately a paradox—akin perhaps to the <a href="/wiki/Waveparticle_duality" class="mw-redirect" title="Waveparticle duality">waveparticle duality</a> of <a href="/wiki/Physics" title="Physics">physics</a>—would be revealed about life. His view however, was later refuted upon the discovery of the <a href="/wiki/Nucleic_acid_double_helix" title="Nucleic acid double helix">double helix structure of DNA</a>.<sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">&#91;</span>31<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Max_Delbr%C3%BCck&amp;action=edit&amp;section=6" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-brittanica-x1-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-brittanica-x1_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-brittanica-x1_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite class="citation encyclopaedia cs1"><a rel="nofollow" class="external text" href="http://www.britannica.com/EBchecked/topic/156446/Max-Delbruck">"Max Delbrück"</a>. <i>Encyclopædia Britannica</i>. Encyclopædia Britannica, Inc. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20181226090033/https://www.britannica.com/biography/Max-Delbruck">Archived</a> from the original on December 26, 2018<span class="reference-accessdate">. Retrieved <span class="nowrap">June 25,</span> 2013</span>. <q>A refugee from Nazi Germany, Delbrück went to the United States in 1937, serving as a faculty member of the California Institute of Technology (1937–39; 1947–81) and of Vanderbilt University (1940–47). He became a U.S. citizen in 1945.</q></cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=bookitem&amp;rft.atitle=Max+Delbr%C3%BCck&amp;rft.btitle=Encyclop%C3%A6dia+Britannica&amp;rft.pub=Encyclop%C3%A6dia+Britannica%2C+Inc.&amp;rft_id=http%3A%2F%2Fwww.britannica.com%2FEBchecked%2Ftopic%2F156446%2FMax-Delbruck&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-nobel1969rationale-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-nobel1969rationale_2-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20130526150123/http://www.nobelprize.org/nobel_prizes/medicine/laureates/1969/index.html">"The Nobel Prize in Physiology or Medicine 1969"</a>. Nobel Foundation. Archived from <a rel="nofollow" class="external text" href="http://nobelprize.org/nobel_prizes/medicine/laureates/1969/index.html">the original</a> on May 26, 2013<span class="reference-accessdate">. Retrieved <span class="nowrap">June 25,</span> 2013</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=unknown&amp;rft.btitle=The+Nobel+Prize+in+Physiology+or+Medicine+1969&amp;rft.pub=Nobel+Foundation&amp;rft_id=http%3A%2F%2Fnobelprize.org%2Fnobel_prizes%2Fmedicine%2Flaureates%2F1969%2Findex.html&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-Nobel-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-Nobel_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Nobel_3-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/1969">"The Nobel Prize in Physiology or Medicine 1969"</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20180627062510/https://www.nobelprize.org/nobel_prizes/medicine/laureates/1969/">Archived</a> June 27, 2018, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>, Nobel Media AB 2013, <i>Nobelprize.org</i>, Web access November 6, 2013.</span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFTon_van_Helvoort1992" class="citation journal cs1">Ton van Helvoort (1992). "The controversy between John H. Northrop and Max Delbrück on the formation of bacteriophage: Bacterial synthesis or autonomous multiplication?". <i>Annals of Science</i>. <b>49</b> (6): 545–575. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1080%2F00033799200200451">10.1080/00033799200200451</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11616207">11616207</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Annals+of+Science&amp;rft.atitle=The+controversy+between+John+H.+Northrop+and+Max+Delbr%C3%BCck+on+the+formation+of+bacteriophage%3A+Bacterial+synthesis+or+autonomous+multiplication%3F&amp;rft.volume=49&amp;rft.issue=6&amp;rft.pages=545-575&amp;rft.date=1992&amp;rft_id=info%3Adoi%2F10.1080%2F00033799200200451&amp;rft_id=info%3Apmid%2F11616207&amp;rft.au=Ton+van+Helvoort&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLily_E._Kay1985" class="citation journal cs1">Lily E. Kay (1985). "Conceptual models and analytical tools: The biology of physicist Max Delbrück". <i>Journal of the History of Biology</i>. <b>18</b> (2): 207–246. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2FBF00120110">10.1007/BF00120110</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11611706">11611706</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:13630670">13630670</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Journal+of+the+History+of+Biology&amp;rft.atitle=Conceptual+models+and+analytical+tools%3A+The+biology+of+physicist+Max+Delbr%C3%BCck&amp;rft.volume=18&amp;rft.issue=2&amp;rft.pages=207-246&amp;rft.date=1985&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A13630670%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F11611706&amp;rft_id=info%3Adoi%2F10.1007%2FBF00120110&amp;rft.au=Lily+E.+Kay&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDaniel_J._McKaughan2005" class="citation journal cs1">Daniel J. McKaughan (2005). "The Influence of Niels Bohr on Max Delbrück". <i>Isis</i>. <b>96</b> (4): 507–529. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1086%2F498591">10.1086/498591</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16536153">16536153</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:12282400">12282400</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Isis&amp;rft.atitle=The+Influence+of+Niels+Bohr+on+Max+Delbr%C3%BCck&amp;rft.volume=96&amp;rft.issue=4&amp;rft.pages=507-529&amp;rft.date=2005&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A12282400%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F16536153&amp;rft_id=info%3Adoi%2F10.1086%2F498591&amp;rft.au=Daniel+J.+McKaughan&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text">"<a rel="nofollow" class="external free" href="https://www.ini.uzh.ch/~tobi/">https://www.ini.uzh.ch/~tobi/</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230110002401/https://www.ini.uzh.ch/~tobi/">Archived</a> January 10, 2023, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>"</span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text">"<a rel="nofollow" class="external free" href="https://ieeexplore.ieee.org/document/8050295">https://ieeexplore.ieee.org/document/8050295</a> <a rel="nofollow" class="external autonumber" href="https://ieeexplore.ieee.org/document/8050295/">[1]</a>"</span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text">"<a rel="nofollow" class="external text" href="https://www.britannica.com/biography/Max-Delbruck">Max Delbrück</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20190127153607/https://www.britannica.com/biography/Max-Delbruck">Archived</a> January 27, 2019, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>". <i>Encyclopaedia Britannica</i>. britannica.com. Retrieved January 27, 2019.</span> </li> <li id="cite_note-NAS-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-NAS_10-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFW._Hayes1992" class="citation journal cs1">W. Hayes (1992). <a rel="nofollow" class="external text" href="http://books.nap.edu/openbook.php?record_id=2201&amp;page=67">"Max Ludwig Henning Delbrück – September 4, 1906 – March 10, 1981"</a>. <i>Biographical Memoirs of the National Academy of Sciences</i>. <b>62</b>: 67–117. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11639973">11639973</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20121015224202/http://books.nap.edu/openbook.php?record_id=2201&amp;page=67">Archived</a> from the original on October 15, 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">July 22,</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Biographical+Memoirs+of+the+National+Academy+of+Sciences&amp;rft.atitle=Max+Ludwig+Henning+Delbr%C3%BCck+%E2%80%93+September+4%2C+1906+%E2%80%93+March+10%2C+1981&amp;rft.volume=62&amp;rft.pages=67-117&amp;rft.date=1992&amp;rft_id=info%3Apmid%2F11639973&amp;rft.au=W.+Hayes&amp;rft_id=http%3A%2F%2Fbooks.nap.edu%2Fopenbook.php%3Frecord_id%3D2201%26page%3D67&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text">Timofeeff-Ressovky, N. W., K. G. Zimmer, and M. Delbrück "Über die Natur der Genmutation und der Genstruktur" (<a href="/wiki/Weidmannsche_Buchhandlung" title="Weidmannsche Buchhandlung">Weidmannsche Buchhandlung</a>, 1935). <a rel="nofollow" class="external text" href="http://www.ini.uzh.ch/~tobi/fun/max/timofeeffZimmerDelbruck1935.pdf">Nachrichten Göttingen</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20220303004033/http://www.ini.uzh.ch/~tobi/fun/max/timofeeffZimmerDelbruck1935.pdf">Archived</a> March 3, 2022, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a> - "Über die Natur der Genmutation und der Genstruktur" (1935).</span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text">Strauss BS. A Physicist's Quest in Biology: Max Delbrück and "Complementarity". Genetics. 2017 Jun;206(2):641-650. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1534%2Fgenetics.117.201517">10.1534/genetics.117.201517</a>. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/28592501">28592501</a>; <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5499177/">5499177</a></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text">Erwin Schrödinger <i>What Is Life? The Physical Aspect of the Living Cell</i> (Cambridge University Press, 1944).</span> </li> <li id="cite_note-MDC-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-MDC_14-0">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://archive.today/20120804081324/http://www.mdc-berlin.de/en/news/archive/2006/20060904-mdc_celebrates_centennial_of_max_delbr_ck/index.html">"MDC celebrates centennial of Max Delbrück"</a>. Max Delbrück Center for Molecular Medicine Berli-Buch. September 4, 2006.</span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-15">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFStefanie_Tapke" class="citation web cs1">Stefanie Tapke. <a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/1969/delbruck-bio.html">"Max Delbrück – Biographical"</a>. <i>Biographical article</i>. Nobel Media. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20180612144035/https://www.nobelprize.org/nobel_prizes/medicine/laureates/1969/delbruck-bio.html">Archived</a> from the original on June 12, 2018<span class="reference-accessdate">. Retrieved <span class="nowrap">September 13,</span> 2013</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=Biographical+article&amp;rft.atitle=Max+Delbr%C3%BCck+%E2%80%93+Biographical&amp;rft.au=Stefanie+Tapke&amp;rft_id=https%3A%2F%2Fwww.nobelprize.org%2Fnobel_prizes%2Fmedicine%2Flaureates%2F1969%2Fdelbruck-bio.html&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-16">^</a></b></span> <span class="reference-text">Ellis E. L., Delbrück M. The growth of bacteriophage. J Gen Physiol. 1939 Jan 20;22(3):365-84. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/19873108">19873108</a></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text">Ellis E.L. "Bacteriophage: One-step growth curve" in Phage and the Origins of Molecular Biology (2007) Edited by John Cairns, Gunther S. Stent, and James D. Watson, Cold Spring Harbor Laboratory of Quantitative Biology, Cold Spring Harbor, Long Island, New York <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-87969-800-3" title="Special:BookSources/978-0-87969-800-3">978-0-87969-800-3</a></span> </li> <li id="cite_note-18"><span class="mw-cite-backlink"><b><a href="#cite_ref-18">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.vanderbilt.edu/delbruck/Delbruck_Centenary_Celebration.html">Max Delbrück and the Next 100 Years of Biology: The Max Delbrück Vanderbilt Centenary Celebration, The Inaugural Vanderbilt Discovery Lecture</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20121119140031/http://www.vanderbilt.edu/delbruck/Delbruck_Centenary_Celebration.html">Archived</a> November 19, 2012, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>, Held September 14, 2006</span> </li> <li id="cite_note-Vanderbilt-19"><span class="mw-cite-backlink">^ <a href="#cite_ref-Vanderbilt_19-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Vanderbilt_19-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Vanderbilt_19-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Vanderbilt_19-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Vanderbilt_19-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-Vanderbilt_19-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-Vanderbilt_19-6"><sup><i><b>g</b></i></sup></a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.vanderbilt.edu/delbruck">"Max Delbrück at Vanderbilt, 1940–1947"</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20121011134613/http://www.vanderbilt.edu/delbruck/">Archived</a> October 11, 2012, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>, Vanderbilt University, Web access November 6, 2013.</span> </li> <li id="cite_note-20"><span class="mw-cite-backlink"><b><a href="#cite_ref-20">^</a></b></span> <span class="reference-text">Luria SE, Delbrück M. Mutations of bacteria from virus sensitivity to virus resistance. Genetics. 1943 Nov;28(6):491-511. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fgenetics%2F28.6.491">10.1093/genetics/28.6.491</a>. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/17247100">17247100</a>; <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1209226/">1209226</a></span> </li> <li id="cite_note-21"><span class="mw-cite-backlink"><b><a href="#cite_ref-21">^</a></b></span> <span class="reference-text">Luria SE "Mutations of bacteria and bacteriophage" in Phage and the Origins of Molecular Biology (2007) Edited by John Cairns, Gunther S. Stent, and James D. Watson, Cold Spring Harbor Laboratory of Quantitative Biology, Cold Spring Harbor, Long Island, New York, pgs. 173-179. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-87969-800-3" title="Special:BookSources/978-0-87969-800-3">978-0-87969-800-3</a></span> </li> <li id="cite_note-22"><span class="mw-cite-backlink"><b><a href="#cite_ref-22">^</a></b></span> <span class="reference-text">Luria SE. A Slot Machine, a Broken Test Tube. An Autobiography. Harper and Row, New York, 1984. Alfred P. Sloan Foundation Series</span> </li> <li id="cite_note-23"><span class="mw-cite-backlink"><b><a href="#cite_ref-23">^</a></b></span> <span class="reference-text">J.D. Watson (2012). <a rel="nofollow" class="external text" href="http://www.cshl.edu/gradschool/Non-Research-Faculty/james-d-watson">"James D Watson: Chancellor emeritus"</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20131211150032/http://www.cshl.edu/gradschool/Non-Research-Faculty/james-d-watson">Archived</a> December 11, 2013, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>, Cold Spring Harbor Laboratory.</span> </li> <li id="cite_note-FischerLipson-24"><span class="mw-cite-backlink"><b><a href="#cite_ref-FischerLipson_24-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPeter_Fischer_Ernst_and_Carol_Lipson1988" class="citation book cs1">Peter Fischer Ernst and Carol Lipson (1988). <span class="id-lock-registration" title="Free registration required"><a rel="nofollow" class="external text" href="https://archive.org/details/thinkingaboutsci00fisc"><i>Thinking about science&#160;: Max Delbrück and the origins of molecular biology</i></a></span>. New York: Norton. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-393-02508-8" title="Special:BookSources/978-0-393-02508-8"><bdi>978-0-393-02508-8</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Thinking+about+science+%3A+Max+Delbr%C3%BCck+and+the+origins+of+molecular+biology&amp;rft.place=New+York&amp;rft.pub=Norton&amp;rft.date=1988&amp;rft.isbn=978-0-393-02508-8&amp;rft.au=Peter+Fischer+Ernst+and+Carol+Lipson&amp;rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fthinkingaboutsci00fisc&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-frs-25"><span class="mw-cite-backlink"><b><a href="#cite_ref-frs_25-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilliam_Hayes1982" class="citation journal cs1"><a href="/wiki/William_Hayes_(geneticist)" title="William Hayes (geneticist)">William Hayes</a> (1982). <a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frsbm.1982.0003">"Max Ludwig Henning Delbruck. 4 September 1906-10 March 1981"</a>. <i><a href="/wiki/Biographical_Memoirs_of_Fellows_of_the_Royal_Society" title="Biographical Memoirs of Fellows of the Royal Society">Biographical Memoirs of Fellows of the Royal Society</a></i>. <b>28</b>. 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Dronamraju (November 1999). <a rel="nofollow" class="external text" href="http://www.genetics.org/content/153/3/1071.full">"Erwin Schrödinger and the origins of molecular biology"</a>. <i>Genetics</i>. <b>153</b> (3): 1071–6. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fgenetics%2F153.3.1071">10.1093/genetics/153.3.1071</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1460808">1460808</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/10545442">10545442</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20120428155705/http://www.genetics.org/content/153/3/1071.full">Archived</a> from the original on April 28, 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">July 22,</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Genetics&amp;rft.atitle=Erwin+Schr%C3%B6dinger+and+the+origins+of+molecular+biology&amp;rft.volume=153&amp;rft.issue=3&amp;rft.pages=1071-6&amp;rft.date=1999-11&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1460808%23id-name%3DPMC&amp;rft_id=info%3Apmid%2F10545442&amp;rft_id=info%3Adoi%2F10.1093%2Fgenetics%2F153.3.1071&amp;rft.au=K.+R.+Dronamraju&amp;rft_id=http%3A%2F%2Fwww.genetics.org%2Fcontent%2F153%2F3%2F1071.full&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> <li id="cite_note-28"><span class="mw-cite-backlink"><b><a href="#cite_ref-28">^</a></b></span> <span class="reference-text">M. P. Murphy and L. A. J. O'Neill (1997). <i>What Is Life? the Next Fifty Years: Speculations on the Future of Biology</i>. Cambridge University Press. <a rel="nofollow" class="external text" href="https://books.google.com/books?id=B1Ejn6ravj8C&amp;pg=PA2">p 2</a>. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/0-521-59939-3" title="Special:BookSources/0-521-59939-3">0-521-59939-3</a></span> </li> <li id="cite_note-Judson-29"><span class="mw-cite-backlink"><b><a href="#cite_ref-Judson_29-0">^</a></b></span> <span class="reference-text"><a href="/wiki/Horace_Freeland_Judson" title="Horace Freeland Judson">Horace Freeland Judson</a> (1996) <i>The Eighth Day of Creation: Makers of the Revolution in Biology</i>. Cold Spring Harbor Laboratory Press. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/0-87969-478-5" title="Special:BookSources/0-87969-478-5">0-87969-478-5</a>.</span> </li> <li id="cite_note-30"><span class="mw-cite-backlink"><b><a href="#cite_ref-30">^</a></b></span> <span class="reference-text">Kiryn Haslinger. <a rel="nofollow" class="external text" href="http://www.cshl.edu/images/stories/about_us/harbor_transcript/2007/07_winter/07_winter_Delbruck100.pdf"><i>Max Delbruck 100.</i></a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150923211708/http://www.cshl.edu/images/stories/about_us/harbor_transcript/2007/07_winter/07_winter_Delbruck100.pdf">Archived</a> September 23, 2015, at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a> HT Winter 2007.</span> </li> <li id="cite_note-31"><span class="mw-cite-backlink"><b><a href="#cite_ref-31">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFN._H._Horowitz1994" class="citation journal cs1">N. H. Horowitz (1994). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1275794">"Review of kay, the Molecular Vision of Life: Caltech, the Rockefeller Foundation, and the Rise of the New Biology"</a>. <i>Biophysical Journal</i>. <b>66</b> (3 Pt 1): 929–930. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1994BpJ....66..929H">1994BpJ....66..929H</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0006-3495%2894%2980873-2">10.1016/S0006-3495(94)80873-2</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1275794">1275794</a></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Biophysical+Journal&amp;rft.atitle=Review+of+kay%2C+the+Molecular+Vision+of+Life%3A+Caltech%2C+the+Rockefeller+Foundation%2C+and+the+Rise+of+the+New+Biology&amp;rft.volume=66&amp;rft.issue=3+Pt+1&amp;rft.pages=929-930&amp;rft.date=1994&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1275794%23id-name%3DPMC&amp;rft_id=info%3Adoi%2F10.1016%2FS0006-3495%2894%2980873-2&amp;rft_id=info%3Abibcode%2F1994BpJ....66..929H&amp;rft.au=N.+H.+Horowitz&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1275794&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AMax+Delbr%C3%BCck" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Max_Delbr%C3%BCck&amp;action=edit&amp;section=7" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; 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This is a letter from Watson to Delbrück that describes the discovery.</li> <li><a rel="nofollow" class="external text" href="http://oralhistories.library.caltech.edu/16/">Interview with Max Delbrück</a> Oral History Project, California Institute of Technology Archives, Pasadena, California.</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20060911044153/http://archives.caltech.edu/search_catalog.cfm?search_field=Delbruck&amp;entry_type=photo">Caltech Photo Archives of Max Delbrück</a></li> <li><a rel="nofollow" class="external text" href="http://www.cumc.columbia.edu/horwitz/">The Official Site of Louisa Gross Horwitz Prize</a></li> <li><a rel="nofollow" class="external text" href="http://osulibrary.oregonstate.edu/specialcollections/coll/pauling/dna/people/delbruck.html">Key Participants: Max Delbrück</a> – <i>Linus Pauling and the Race for DNA: A Documentary History</i></li> <li><a rel="nofollow" class="external text" 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title="Template talk:Nobel Prize in Physiology or Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Special:EditPage/Template:Nobel Prize in Physiology or Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Physiology_or_Medicine" title="List of Nobel laureates in Physiology or Medicine">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_Macleod_(physiologist)" title="John Macleod (physiologist)">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1925</span></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Krebs_(biochemist)" title="Hans Krebs (biochemist)">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Beadle" title="George Beadle">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a class="mw-selflink selflink">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_MacLeod_Cormack" title="Allan MacLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards_(physiologist)" title="Robert Edwards (physiologist)">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>posthumously</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe_(neuroscientist)" title="John O&#39;Keefe (neuroscientist)">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap"><a href="/wiki/2020_Nobel_Prize_in_Physiology_or_Medicine" title="2020 Nobel Prize in Physiology or Medicine">2020</a>: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton_(virologist)" class="mw-redirect" title="Michael Houghton (virologist)">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap"><a href="/wiki/2021_Nobel_Prize_in_Physiology_or_Medicine" title="2021 Nobel Prize in Physiology or Medicine">2021</a>: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap"><a href="/wiki/2022_Nobel_Prize_in_Physiology_or_Medicine" title="2022 Nobel Prize in Physiology or Medicine">2022</a>: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap"><a href="/wiki/2023_Nobel_Prize_in_Physiology_or_Medicine" title="2023 Nobel Prize in Physiology or Medicine">2023</a>: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="1969_Nobel_Prize_laureates" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:1969_Nobel_Prize_winners" title="Template:1969 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:1969_Nobel_Prize_winners" title="Template talk:1969 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:1969_Nobel_Prize_winners" title="Special:EditPage/Template:1969 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="1969_Nobel_Prize_laureates" style="font-size:114%;margin:0 4em">1969 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Derek_Barton" title="Derek Barton">Derek Barton</a> (Great Britain)</li> <li><a href="/wiki/Odd_Hassel" title="Odd Hassel">Odd Hassel</a> (Norway)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/1969_Nobel_Prize_in_Literature" title="1969 Nobel Prize in Literature">1969</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Samuel_Beckett" title="Samuel Beckett">Samuel Beckett</a> (Ireland)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a> (<a href="/wiki/1969_Nobel_Peace_Prize" title="1969 Nobel Peace Prize">1969</a>)</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/International_Labour_Organization" title="International Labour Organization">International Labour Organization</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Murray_Gell-Mann" title="Murray Gell-Mann">Murray Gell-Mann</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Max Delbrück</a> (United States)</li> <li><a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> (United States)</li> <li><a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economic Sciences</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ragnar_Frisch" title="Ragnar Frisch">Ragnar Frisch</a> (Norway)</li> <li><a href="/wiki/Jan_Tinbergen" title="Jan Tinbergen">Jan Tinbergen</a> (Netherlands)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/Template:1964_Nobel_Prize_winners" title="Template:1964 Nobel Prize winners">1964</a></dd> <dd><a href="/wiki/Template:1965_Nobel_Prize_winners" title="Template:1965 Nobel Prize winners">1965</a></dd> <dd><a href="/wiki/Template:1966_Nobel_Prize_winners" title="Template:1966 Nobel Prize winners">1966</a></dd> <dd><a href="/wiki/Template:1967_Nobel_Prize_winners" title="Template:1967 Nobel Prize winners">1967</a></dd> <dd><a href="/wiki/Template:1968_Nobel_Prize_winners" title="Template:1968 Nobel Prize winners">1968</a></dd> <dd><b>1969 </b></dd> <dd><a href="/wiki/Template:1970_Nobel_Prize_winners" title="Template:1970 Nobel Prize winners">1970</a></dd> <dd><a href="/wiki/Template:1971_Nobel_Prize_winners" title="Template:1971 Nobel Prize winners">1971</a></dd> <dd><a href="/wiki/Template:1972_Nobel_Prize_winners" title="Template:1972 Nobel Prize winners">1972</a></dd> <dd><a href="/wiki/Template:1973_Nobel_Prize_winners" title="Template:1973 Nobel Prize winners">1973</a></dd> <dd><a href="/wiki/Template:1974_Nobel_Prize_winners" title="Template:1974 Nobel Prize winners">1974</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&amp;#124;text-top&amp;#124;10px&amp;#124;alt=Edit_this_at_Wikidata&amp;#124;link=https&amp;#58;//www.wikidata.org/wiki/Q76807#identifiers&amp;#124;class=noprint&amp;#124;Edit_this_at_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Authority_control_databases_frameless&amp;#124;text-top&amp;#124;10px&amp;#124;alt=Edit_this_at_Wikidata&amp;#124;link=https&amp;#58;//www.wikidata.org/wiki/Q76807#identifiers&amp;#124;class=noprint&amp;#124;Edit_this_at_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q76807#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">International</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/0000000071501831">ISNI</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/100384850">VIAF</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://id.worldcat.org/fast/185042/">FAST</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.oclc.org/worldcat/entity/E39PBJjd9fGBFyxT8GhthbxkXd">WorldCat</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/120256568">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n85287457">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb12292744p">France</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb12292744p">BnF data</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00620570">Japan</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&amp;local_base=aut&amp;ccl_term=ica=nlk20000082229&amp;CON_LNG=ENG">Czech Republic</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p07523873X">Netherlands</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://libris.kb.se/97mppmtt0q6vxhg">Sweden</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://dbn.bn.org.pl/descriptor-details/9810606824705606">Poland</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&amp;local_base=NLX10&amp;find_code=UID&amp;request=987007276852705171">Israel</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://cantic.bnc.cat/registre/981058520237206706">Catalonia</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Academics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://ci.nii.ac.jp/author/DA01985393?l=en">CiNii</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.uri?authorId=6603037446">Scopus</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://zbmath.org/authors/?q=ai:delbruck.max">zbMATH</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.leopoldina.org/mitgliederverzeichnis/mitglieder/member/Member/show/max-delbrueck-1/">Leopoldina</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Artists</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://musicbrainz.org/artist/df880cb0-f6d5-45c0-acab-15073c345f2c">MusicBrainz</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">People</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.deutsche-biographie.de/pnd120256568.html?language=en">Deutsche Biographie</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.deutsche-digitale-bibliothek.de/person/gnd/120256568">DDB</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/072294426">IdRef</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://snaccooperative.org/ark:/99166/w66b9j96">SNAC</a></span></li></ul></div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐f69cdc8f6‐7j489 Cached time: 20241122175706 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 0.986 seconds Real time 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