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id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Ecosystem event</div> <p><b>Trophic cascades</b> are powerful indirect interactions that can control entire <a href="/wiki/Ecosystems" class="mw-redirect" title="Ecosystems">ecosystems</a>, occurring when a <a href="/wiki/Trophic_level" title="Trophic level">trophic level</a> in a <a href="/wiki/Food_web" title="Food web">food web</a> is suppressed. For example, a top-down cascade will occur if predators are effective enough in predation to reduce the abundance, or alter the behavior of their <a href="/wiki/Prey" class="mw-redirect" title="Prey">prey</a>, thereby releasing the next lower trophic level from <a href="/wiki/Predation" title="Predation">predation</a> (or <a href="/wiki/Herbivory" class="mw-redirect" title="Herbivory">herbivory</a> if the intermediate trophic level is a herbivore). </p><p>The trophic cascade is an ecological concept which has stimulated new research in many areas of <a href="/wiki/Ecology" title="Ecology">ecology</a>. For example, it can be important for understanding the knock-on effects of removing <a href="/wiki/Apex_predator" title="Apex predator">top predators</a> from <a href="/wiki/Food_web" title="Food web">food webs</a>, as humans have done in many places through <a href="/wiki/Hunting" title="Hunting">hunting</a> and <a href="/wiki/Fishing" title="Fishing">fishing</a>. </p><p>A <b>top-down cascade</b> is a trophic cascade where the top consumer/predator controls the <a href="/wiki/Primary_consumer" class="mw-redirect" title="Primary consumer">primary consumer</a> population. In turn, the primary producer population thrives. The removal of the top predator can alter the food web dynamics. In this case, the primary consumers would overpopulate and exploit the primary producers. Eventually there would not be enough primary producers to sustain the consumer population. Top-down food web stability depends on competition and predation in the higher trophic levels. Invasive species can also alter this cascade by removing or becoming a top predator. This interaction may not always be negative. Studies have shown that certain invasive species have begun to shift cascades; and as a consequence, ecosystem degradation has been repaired.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>For example, if the abundance of large <a href="/wiki/Piscivore" title="Piscivore">piscivorous</a> fish is increased in a <a href="/wiki/Lake" title="Lake">lake</a>, the abundance of their prey, smaller fish that eat <a href="/wiki/Zooplankton" title="Zooplankton">zooplankton</a>, should decrease. The resulting increase in zooplankton should, in turn, cause the <a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">biomass</a> of its prey, <a href="/wiki/Phytoplankton" title="Phytoplankton">phytoplankton</a>, to decrease. </p><p>In a <b>bottom-up cascade</b>, the population of primary producers will always control the increase/decrease of the energy in the higher trophic levels. Primary producers are plants and phytoplankton that require photosynthesis. Although light is important, primary producer populations are altered by the amount of nutrients in the system. This food web relies on the availability and limitation of resources. All populations will experience growth if there is initially a large amount of nutrients.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>In a <b>subsidy cascade</b>, species populations at one trophic level can be supplemented by external food. For example, native animals can forage on resources that don't originate in their same habitat, such as native predators eating livestock. This may increase their local abundances thereby affecting other species in the ecosystem and causing an ecological cascade. For example, Luskin et al. (2017) found that native animals living in protected primary rainforest in Malaysia found food subsidies in neighboring oil palm plantations.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> This subsidy allowed native animal populations to increase, which then triggered powerful secondary ‘cascading’ effects on forest tree community. Specifically, crop-raiding <a href="/wiki/Wild_boar" title="Wild boar">wild boar</a> (<i>Sus scrofa</i>) built thousands of nests from the forest understory vegetation and this caused a 62% decline in forest tree sapling density over a 24-year study period. Such cross-boundary subsidy cascades may be widespread in both terrestrial and marine ecosystems and present significant conservation challenges. </p><p>These trophic interactions shape patterns of biodiversity globally. Humans and <a href="/wiki/Climate_change" title="Climate change">climate change</a> have affected these cascades drastically. One example can be seen with <a href="/wiki/Sea_otter" title="Sea otter">sea otters</a> (<i>Enhydra lutris</i>) on the Pacific coast of the United States of America. Over time, human interactions caused a removal of sea otters. One of their main prey, the Pacific purple sea urchin (<i><a href="/wiki/Strongylocentrotus_purpuratus" title="Strongylocentrotus purpuratus">Strongylocentrotus purpuratus</a></i>) eventually began to overpopulate. The overpopulation caused increased predation of <a href="/wiki/Giant_kelp" class="mw-redirect" title="Giant kelp">giant kelp</a> (<i>Macrocystis pyrifera</i>). As a result, there was extreme deterioration of the kelp forests along the California coast. This is why it is important for countries to regulate marine and terrestrial ecosystems.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>Predator-induced interactions could heavily influence the flux of atmospheric carbon if managed on a global scale. For example, a study was conducted to determine the cost of potential stored carbon in living kelp biomass in sea otter (<i>Enhydra lutris</i>) enhanced ecosystems. The study valued the potential storage between $205 million and $408 million dollars (US) on the European Carbon Exchange (2012).<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Origins_and_theory">Origins and theory</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_cascade&action=edit&section=1" title="Edit section: Origins and theory"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Aldo_Leopold" title="Aldo Leopold">Aldo Leopold</a> is generally credited with first describing the mechanism of a trophic cascade, based on his observations of <a href="/wiki/Overgrazing" title="Overgrazing">overgrazing</a> of mountain slopes by deer after human extermination of wolves.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Nelson_Hairston" title="Nelson Hairston">Nelson Hairston</a>, Frederick E. Smith and <a href="/wiki/Lawrence_B._Slobodkin" title="Lawrence B. Slobodkin">Lawrence B. Slobodkin</a> are generally credited with introducing the concept into scientific discourse, although they did not use the term either. Hairston, Smith and Slobodkin argued that predators reduce the abundance of herbivores, allowing <a href="/wiki/Plant" title="Plant">plants</a> to flourish.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> This is often referred to as the <a href="/wiki/Green_world_hypothesis" title="Green world hypothesis">green world hypothesis</a>. The green world hypothesis is credited with bringing attention to the role of top-down forces (e.g. predation) and indirect effects in shaping <a href="/wiki/Biocoenosis" title="Biocoenosis">ecological communities</a>. The prevailing view of communities prior to Hairston, Smith and Slobodkin was trophodynamics, which attempted to explain the structure of communities using only bottom-up forces (e.g. resource limitation). Smith may have been inspired by the experiments of a Czech ecologist, <a href="/wiki/Jaroslav_Hrb%C3%A1%C4%8Dek" title="Jaroslav Hrbáček">Hrbáček</a>, whom he met on a <a href="/wiki/United_States_State_Department" class="mw-redirect" title="United States State Department">United States State Department</a> cultural exchange. Hrbáček had shown that fish in artificial <a href="/wiki/Pond" title="Pond">ponds</a> reduced the abundance of <a href="/wiki/Zooplankton" title="Zooplankton">zooplankton</a>, leading to an increase in the abundance of <a href="/wiki/Phytoplankton" title="Phytoplankton">phytoplankton</a>.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p><p>Hairston, Smith and Slobodkin feuded that the ecological communities acted as <a href="/wiki/Food_chains" class="mw-redirect" title="Food chains">food chains</a> with three trophic levels. Subsequent models expanded the argument to food chains with more than or fewer than three trophic levels.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> Lauri Oksanen argued that the top trophic level in a food chain increases the abundance of <a href="/wiki/Primary_productivity" class="mw-redirect" title="Primary productivity">producers</a> in food chains with an odd number of trophic levels (such as in Hairston, Smith and Slobodkin's three trophic level model), but decreases the abundance of the producers in food chains with an even number of trophic levels. Additionally, he argued that the number of trophic levels in a food chain increases as the productivity of the <a href="/wiki/Ecosystem" title="Ecosystem">ecosystem</a> increases. </p> <div class="mw-heading mw-heading2"><h2 id="Classic_examples">Classic examples</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_cascade&action=edit&section=2" title="Edit section: Classic examples"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Kelp_forest_and_sardines,_San_Clemente_Island,_Channel_Islands,_California.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/60/Kelp_forest_and_sardines%2C_San_Clemente_Island%2C_Channel_Islands%2C_California.jpg/220px-Kelp_forest_and_sardines%2C_San_Clemente_Island%2C_Channel_Islands%2C_California.jpg" decoding="async" width="220" height="293" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/60/Kelp_forest_and_sardines%2C_San_Clemente_Island%2C_Channel_Islands%2C_California.jpg/330px-Kelp_forest_and_sardines%2C_San_Clemente_Island%2C_Channel_Islands%2C_California.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/60/Kelp_forest_and_sardines%2C_San_Clemente_Island%2C_Channel_Islands%2C_California.jpg/440px-Kelp_forest_and_sardines%2C_San_Clemente_Island%2C_Channel_Islands%2C_California.jpg 2x" data-file-width="450" data-file-height="600" /></a><figcaption>Healthy Pacific kelp forests, like this one at <a href="/wiki/San_Clemente_Island" title="San Clemente Island">San Clemente Island</a> of California's <a href="/wiki/Channel_Islands_of_California" class="mw-redirect" title="Channel Islands of California">Channel Islands</a>, have been shown to flourish when sea otters are present. When otters are absent, sea urchin populations can <a href="/wiki/Irruptive_growth" title="Irruptive growth">irrupt</a> and severely degrade the kelp forest ecosystem.</figcaption></figure> <p>Although Hairston, Smith and Slobodkin formulated their argument in terms of terrestrial food chains, the earliest empirical demonstrations of trophic cascades came from <a href="/wiki/Marine_(ocean)" class="mw-redirect" title="Marine (ocean)">marine</a> and, especially, <a href="/wiki/Aquatic_ecosystem" title="Aquatic ecosystem">aquatic ecosystems</a>. Some of the most famous examples are: </p> <ul><li>In <a href="/wiki/North_America" title="North America">North American</a> lakes, <a href="/wiki/Piscivore" title="Piscivore">piscivorous</a> fish can dramatically reduce populations of zooplanktivorous fish; zooplanktivorous fish can dramatically alter <a href="/wiki/Freshwater" class="mw-redirect" title="Freshwater">freshwater</a> <a href="/wiki/Zooplankton" title="Zooplankton">zooplankton</a> communities, and zooplankton grazing can in turn have large impacts on <a href="/wiki/Phytoplankton" title="Phytoplankton">phytoplankton</a> communities. Removal of piscivorous fish can change lake water from clear to green by allowing phytoplankton to flourish.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup></li> <li>In the <a href="/wiki/Eel_River_(California)" title="Eel River (California)">Eel River</a>, in Northern <a href="/wiki/California" title="California">California</a>, fish (<a href="/wiki/Rainbow_trout" title="Rainbow trout">steelhead</a> and <a href="/wiki/California_roach" title="California roach">roach</a>) consume fish larvae and predatory <a href="/wiki/Insect" title="Insect">insects</a>. These smaller predators prey on <a href="/wiki/Midge" title="Midge">midge</a> larvae, which feed on <a href="/wiki/Algae" title="Algae">algae</a>. Removal of the larger fish increases the abundance of algae.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup></li> <li>In <a href="/wiki/Pacific_Ocean" title="Pacific Ocean">Pacific</a> <a href="/wiki/Kelp_forest" title="Kelp forest">kelp forests</a>, <a href="/wiki/Sea_otter" title="Sea otter">sea otters</a> feed on <a href="/wiki/Sea_urchin" title="Sea urchin">sea urchins</a>. In areas where sea otters have been <a href="/wiki/Hunting" title="Hunting">hunted</a> to <a href="/wiki/Local_extinction" title="Local extinction">extinction</a>, sea urchins increase in abundance and <a href="/wiki/Kelp" title="Kelp">kelp</a> populations are reduced.<sup id="cite_ref-uwo.academia_15-0" class="reference"><a href="#cite_note-uwo.academia-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup></li> <li>A classic example of a terrestrial trophic cascade is the reintroduction of <a href="/wiki/Gray_wolf" class="mw-redirect" title="Gray wolf">gray wolves</a> (<i>Canis lupus</i>) to <a href="/wiki/Yellowstone_National_Park" title="Yellowstone National Park">Yellowstone National Park</a>, which reduced the number, and changed the behavior, of <a href="/wiki/Elk" title="Elk">elk</a> (<i>Cervus canadensis</i>). This in turn released several plant species from grazing pressure and subsequently led to the transformation of riparian ecosystems.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Terrestrial_trophic_cascades">Terrestrial trophic cascades</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_cascade&action=edit&section=3" title="Edit section: Terrestrial trophic cascades"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The fact that the earliest documented trophic cascades all occurred in lakes and <a href="/wiki/Stream" title="Stream">streams</a> led a scientist to speculate that fundamental differences between aquatic and terrestrial <a href="/wiki/Food_web" title="Food web">food webs</a> made trophic cascades primarily an aquatic phenomenon. Trophic cascades were restricted to communities with relatively low <a href="/wiki/Biodiversity" title="Biodiversity">species diversity</a>, in which a small number of species could have overwhelming influence and the food web could operate as a linear food chain. Additionally, well documented trophic cascades at that point in time all occurred in food chains with algae as the <a href="/wiki/Primary_producer" class="mw-redirect" title="Primary producer">primary producer</a>. Trophic cascades, Strong argued, may only occur in communities with fast-growing producers which lack <a href="/wiki/Plant_defense_against_herbivory" title="Plant defense against herbivory">defenses against herbivory</a>.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p><p>Subsequent research has documented trophic cascades in terrestrial ecosystems, including: </p> <ul><li>In the <a href="/wiki/California_coastal_prairie" title="California coastal prairie">coastal prairie</a> of Northern California, <a href="/wiki/Yellow_bush_lupine" class="mw-redirect" title="Yellow bush lupine">yellow bush lupines</a> are fed upon by a particularly destructive herbivore, the root-boring caterpillar of the lupine ghost moth <i><a href="/wiki/Phymatopus_californicus" title="Phymatopus californicus">Phymatopus californicus</a></i>. <a href="/wiki/Entomopathogenic_nematode" title="Entomopathogenic nematode">Entomopathogenic nematodes</a> kill the caterpillars, and can increase the survival and <a href="/wiki/Seed" title="Seed">seed</a> production of lupines.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup></li> <li>In <a href="/wiki/Costa_Rica" title="Costa Rica">Costa Rican</a> <a href="/wiki/Rain_forest" class="mw-redirect" title="Rain forest">rain forest</a>, a <a href="/wiki/Cleridae" title="Cleridae">clerid</a> <a href="/wiki/Beetle" title="Beetle">beetle</a> specializes in eating <a href="/wiki/Ant" title="Ant">ants</a>. The ant <i>Pheidole bicornis</i> has a <a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">mutualistic</a> association with <a href="/wiki/Piper_(genus)" class="mw-redirect" title="Piper (genus)"><i>Piper</i></a> plants: the ant lives on the <i>Piper</i> and removes <a href="/wiki/Caterpillar" title="Caterpillar">caterpillars</a> and other <a href="/wiki/Insect" title="Insect">insect</a> herbivores. The clerid beetle, by reducing the abundance of ants, increases the leaf area removed from <i>Piper</i> plants by insect herbivores.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup></li></ul> <p>Critics pointed out that published terrestrial trophic cascades generally involved smaller subsets of the food web (often only a single plant species). This was quite different from aquatic trophic cascades, in which the biomass of producers as a whole were reduced when predators were removed. Additionally, most terrestrial trophic cascades did not demonstrate reduced plant biomass when predators were removed, but only increased plant damage from herbivores.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> It was unclear if such damage would actually result in reduced plant biomass or abundance. In 2002 a <a href="/wiki/Meta-analysis" title="Meta-analysis">meta-analysis</a> found trophic cascades to be generally <i>weaker</i> in terrestrial ecosystems, meaning that changes in predator biomass resulted in smaller changes in plant biomass.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> In contrast, a study published in 2009 demonstrated that multiple species of trees with highly varying <a href="/wiki/Autoecology" class="mw-redirect" title="Autoecology">autecologies</a> are in fact heavily impacted by the loss of an apex predator.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> Another study, published in 2011, demonstrated that the loss of large terrestrial predators also significantly degrades the integrity of river and stream systems, impacting their <a href="/wiki/River_morphology" title="River morphology">morphology</a>, <a href="/wiki/Hydrology" title="Hydrology">hydrology</a>, and associated biological communities.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p><p>The critics' model is challenged by studies accumulating since the reintroduction of <a href="/wiki/Gray_wolf" class="mw-redirect" title="Gray wolf">gray wolves</a> (<i>Canis lupus</i>) to <a href="/wiki/Yellowstone_National_Park" title="Yellowstone National Park">Yellowstone National Park</a>. The gray wolf, after being <a href="/wiki/Local_extinction" title="Local extinction">extirpated</a> in the 1920s and absent for 70 years, was <a href="/wiki/Wolf_reintroduction" title="Wolf reintroduction">reintroduced</a> to the park in 1995 and 1996. Since then a three-tiered trophic cascade has been reestablished involving wolves, <a href="/wiki/Elk" title="Elk">elk</a> (<i>Cervus elaphus</i>), and woody <a href="/wiki/Browsing_(herbivory)" title="Browsing (herbivory)">browse</a> species such as <a href="/wiki/Populus_tremuloides" title="Populus tremuloides">aspen</a> (<i>Populus tremuloides</i>), <a href="/wiki/Populus_sect._Aigeiros" title="Populus sect. Aigeiros">cottonwoods</a> (<i>Populus</i> spp.), and <a href="/wiki/Willows" class="mw-redirect" title="Willows">willows</a> (<i>Salix</i> spp.). Mechanisms likely include actual wolf predation of elk, which reduces their numbers, and the threat of predation, which alters elk behavior and feeding habits, resulting in these plant species being released from intensive browsing pressure. Subsequently, their survival and <a href="/wiki/Recruitment_(biology)" title="Recruitment (biology)">recruitment</a> rates have significantly increased in some places within Yellowstone's northern range. This effect is particularly noted among the range's <a href="/wiki/Riparian" class="mw-redirect" title="Riparian">riparian</a> plant communities, with upland communities only recently beginning to show similar signs of recovery.<sup id="cite_ref-rippbesch2012_26-0" class="reference"><a href="#cite_note-rippbesch2012-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> </p><p>Examples of this phenomenon include: </p> <ul><li>A 2–3 fold increase in <a href="/wiki/Deciduous" title="Deciduous">deciduous</a> woody vegetation cover, mostly of willow, in the <a href="/wiki/Soda_Butte_Creek" title="Soda Butte Creek">Soda Butte Creek</a> area between 1995 and 1999.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup></li> <li>Heights of the tallest willows in the <a href="/wiki/Gallatin_Range" title="Gallatin Range">Gallatin River valley</a> increasing from 75 cm to 200 cm between 1998 and 2002.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup></li> <li>Heights of the tallest willows in the Blacktail Creek area increased from less than 50 cm to more than 250 cm between 1997 and 2003. Additionally, canopy cover over streams increased significantly, from only 5% to a range of 14–73%.<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup></li> <li>In the northern range, tall deciduous woody vegetation cover increased by 170% between 1991 and 2006.<sup id="cite_ref-baril2009_30-0" class="reference"><a href="#cite_note-baril2009-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup></li> <li>In the <a href="/wiki/Lamar_River" title="Lamar River">Lamar</a> and Soda Butte Valleys the number of young cottonwood trees that had been successfully recruited went from 0 to 156 between 2001 and 2010.<sup id="cite_ref-rippbesch2012_26-1" class="reference"><a href="#cite_note-rippbesch2012-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup></li></ul> <p>Trophic cascades also impact the <a href="/wiki/Biodiversity" title="Biodiversity">biodiversity</a> of ecosystems, and when examined from that perspective wolves appear to be having multiple, positive cascading impacts on the biodiversity of Yellowstone National Park. These impacts include: </p> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Trophic_Cascade_1.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Trophic_Cascade_1.svg/200px-Trophic_Cascade_1.svg.png" decoding="async" width="200" height="283" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Trophic_Cascade_1.svg/300px-Trophic_Cascade_1.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d6/Trophic_Cascade_1.svg/400px-Trophic_Cascade_1.svg.png 2x" data-file-width="724" data-file-height="1024" /></a><figcaption>This diagram illustrates trophic cascade caused by removal of the top predator. When the top predator is removed the population of deer is able to grow unchecked and this causes over-consumption of the primary producers.</figcaption></figure> <ul><li><a href="/wiki/Scavenger" title="Scavenger">Scavengers</a>, such as <a href="/wiki/Common_raven" title="Common raven">ravens</a> (<i>Corvus corax</i>), <a href="/wiki/Bald_eagles" class="mw-redirect" title="Bald eagles">bald eagles</a> (<i>Haliaeetus leucocephalus</i>), and even <a href="/wiki/Grizzly_bears" class="mw-redirect" title="Grizzly bears">grizzly bears</a> (<i>Ursus arctos horribilis</i>), are likely subsidized by the carcasses of wolf kills.<sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup></li> <li>In the northern range, the relative abundance of six out of seven native songbirds which utilize willow was found to be greater in areas of willow recovery as opposed to those where willows remained suppressed.<sup id="cite_ref-baril2009_30-1" class="reference"><a href="#cite_note-baril2009-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/American_bison" title="American bison">Bison</a> (<i>Bison bison</i>) numbers in the northern range have been steadily increasing as elk numbers have declined, presumably due to a decrease in <a href="/wiki/Interspecific_competition" title="Interspecific competition">interspecific competition</a> between the two species.<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup></li> <li>Importantly, the number of <a href="/wiki/American_beaver" class="mw-redirect" title="American beaver">beaver</a> (<i>Castor canadensis</i>) colonies in the park has increased from one in 1996 to twelve in 2009. The recovery is likely due to the increase in willow availability, as they have been feeding almost exclusively on it. As <a href="/wiki/Keystone_species" title="Keystone species">keystone species</a>, the resurgence of beaver is a critical event for the region. The presence of beavers has been shown to positively impact streambank <a href="/wiki/Erosion" title="Erosion">erosion</a>, <a href="/wiki/Sediment" title="Sediment">sediment</a> retention, <a href="/wiki/Water_table" title="Water table">water tables</a>, <a href="/wiki/Nutrient_cycling" class="mw-redirect" title="Nutrient cycling">nutrient cycling</a>, and both the diversity and abundance of plant and animal life among riparian communities.<sup id="cite_ref-rippbesch2012_26-2" class="reference"><a href="#cite_note-rippbesch2012-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup></li></ul> <p>There are a number of other examples of trophic cascades involving large terrestrial mammals, including: </p> <ul><li>In both <a href="/wiki/Zion_National_Park" title="Zion National Park">Zion National Park</a> and <a href="/wiki/Yosemite_National_Park" title="Yosemite National Park">Yosemite National Park</a>, the increase in human visitation during the first half of the 20th century was found to correspond to the decline of native <a href="/wiki/Cougar" title="Cougar">cougar</a> (<i>Puma concolor</i>) populations in at least part of their range. Soon after, native populations of <a href="/wiki/Mule_deer" title="Mule deer">mule deer</a> (<i>Odocoileus hemionus</i>) erupted, subjecting resident communities of cottonwoods (<i>Populus fremontii</i>) in Zion and <a href="/wiki/California_black_oak" class="mw-redirect" title="California black oak">California black oak</a> (<i>Quercus kelloggii</i>) in Yosemite to intensified browsing. This halted successful recruitment of these species except in refugia inaccessible to the deer. In Zion the suppression of cottonwoods increased stream erosion and decreased the diversity and abundance of amphibians, reptiles, butterflies, and wildflowers. In parts of the park where cougars were still common these negative impacts were not expressed and riparian communities were significantly healthier.<sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup></li> <li>In <a href="/wiki/Sub-Saharan_Africa" title="Sub-Saharan Africa">sub-Saharan Africa</a>, the decline of <a href="/wiki/Lion" title="Lion">lion</a> (<i>Panthera leo</i>) and <a href="/wiki/Leopard" title="Leopard">leopard</a> (<i>Panthera pardus</i>) populations has led to a rising population of <a href="/wiki/Olive_baboon" title="Olive baboon">olive baboon</a> (<i>Papio anubis</i>). This case of <a href="/wiki/Mesopredator_release" class="mw-redirect" title="Mesopredator release">mesopredator release</a> negatively impacted already declining <a href="/wiki/Ungulate" title="Ungulate">ungulate</a> populations and is one of the reasons for increased conflict between baboons and humans, as the primates raid crops and spread <a href="/wiki/Intestinal_parasite" class="mw-redirect" title="Intestinal parasite">intestinal parasites</a>.<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup></li> <li>In the Australian states of <a href="/wiki/New_South_Wales" title="New South Wales">New South Wales</a> and <a href="/wiki/South_Australia" title="South Australia">South Australia</a>, the presence or absence of <a href="/wiki/Australian_dingo" class="mw-redirect" title="Australian dingo">dingoes</a> (<i>Canis lupus dingo</i>) was found to be inversely related to the abundance of invasive <a href="/wiki/Red_fox" title="Red fox">red foxes</a> (<i>Vulpes vulpes</i>). In other words, the foxes were most common where the dingoes were least common. Subsequently, populations of an endangered prey species, the <a href="/wiki/Dusky_hopping_mouse" title="Dusky hopping mouse">dusky hopping mouse</a> (<i>Notomys fuscus</i>) were also less abundant where dingoes were absent due to the foxes, which consume the mice, no longer being held in check by the top predator.<sup id="cite_ref-37" class="reference"><a href="#cite_note-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Marine_trophic_cascades">Marine trophic cascades</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_cascade&action=edit&section=4" title="Edit section: Marine trophic cascades"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In addition to the classic examples listed above, more recent examples of trophic cascades in <a href="/wiki/Marine_ecosystem" title="Marine ecosystem">marine ecosystems</a> have been identified: </p> <ul><li>An example of a cascade in a complex, open-ocean ecosystem occurred in the northwest <a href="/wiki/Atlantic_Ocean" title="Atlantic Ocean">Atlantic</a> during the 1980s and 1990s. The removal of <a href="/wiki/Atlantic_cod" title="Atlantic cod">Atlantic cod</a> (<i>Gadus morhua</i>) and other ground fishes by sustained <a href="/wiki/Overfishing" title="Overfishing">overfishing</a> resulted in increases in the abundance of the prey species for these ground fishes, particularly smaller <a href="/wiki/Forage_fish" title="Forage fish">forage fishes</a> and invertebrates such as the northern <a href="/wiki/Chionoecetes" title="Chionoecetes">snow crab</a> (<i>Chionoecetes opilio</i>) and northern <a href="/wiki/Shrimp" title="Shrimp">shrimp</a> (<i>Pandalus borealis</i>). The increased abundance of these prey species altered the community of <a href="/wiki/Zooplankton" title="Zooplankton">zooplankton</a> that serve as food for smaller fishes and invertebrates as an indirect effect.<sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup></li> <li>A similar cascade, also involving the Atlantic cod, occurred in the <a href="/wiki/Baltic_Sea" title="Baltic Sea">Baltic Sea</a> at the end of the 1980s. After a decline in Atlantic cod, the abundance of its main prey, the <a href="/wiki/Sprat" title="Sprat">sprat</a> (<i>Sprattus sprattus</i>), increased<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">[</span>39<span class="cite-bracket">]</span></a></sup> and the Baltic Sea ecosystem shifted from being dominated by cod into being dominated by sprat. The next level of trophic cascade was a decrease in the abundance of <i>Pseudocalanus acuspes</i>,<sup id="cite_ref-40" class="reference"><a href="#cite_note-40"><span class="cite-bracket">[</span>40<span class="cite-bracket">]</span></a></sup> a <a href="/wiki/Copepod" title="Copepod">copepod</a> which the sprat prey on.</li> <li>On <a href="/wiki/Caribbean_Sea" title="Caribbean Sea">Caribbean</a> <a href="/wiki/Coral_reef" title="Coral reef">coral reefs</a>, several species of angelfishes and <a href="/wiki/Parrotfish" title="Parrotfish">parrotfishes</a> eat species of <a href="/wiki/Sponge" title="Sponge">sponges</a> that lack <a href="/wiki/Chemical_defense" title="Chemical defense">chemical defenses</a>. Removal of these sponge-eating fish species from reefs by fish-trapping and netting has resulted in a shift in the sponge community toward fast-growing sponge species that lack chemical defenses.<sup id="cite_ref-41" class="reference"><a href="#cite_note-41"><span class="cite-bracket">[</span>41<span class="cite-bracket">]</span></a></sup> These fast-growing sponge species are superior competitors for space, and overgrow and smother reef-building corals to a greater extent on overfished reefs.<sup id="cite_ref-42" class="reference"><a href="#cite_note-42"><span class="cite-bracket">[</span>42<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Criticisms">Criticisms</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_cascade&action=edit&section=5" title="Edit section: Criticisms"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Although the existence of trophic cascades is not controversial, ecologists have long debated how ubiquitous they are. Hairston, Smith and Slobodkin argued that <a href="/wiki/Terrestrial_ecoregion" class="mw-redirect" title="Terrestrial ecoregion">terrestrial</a> <a href="/wiki/Ecosystems" class="mw-redirect" title="Ecosystems">ecosystems</a>, as a rule, behave as a three <a href="/wiki/Trophic_level" title="Trophic level">trophic level</a> trophic cascade, which provoked immediate controversy. Some of the criticisms, both of Hairston, Smith and Slobodkin's model and of Oksanen's later model, were: </p> <ul><li><a href="/wiki/Plant" title="Plant">Plants</a> possess numerous <a href="/wiki/Plant_defense_against_herbivory" title="Plant defense against herbivory">defenses</a> against herbivory, and these defenses also contribute to reducing the impact of herbivores on plant populations.<sup id="cite_ref-Murdoch_1966_43-0" class="reference"><a href="#cite_note-Murdoch_1966-43"><span class="cite-bracket">[</span>43<span class="cite-bracket">]</span></a></sup></li> <li>Herbivore populations may be limited by factors other than food or predation, such as nesting sites or available territory.<sup id="cite_ref-Murdoch_1966_43-1" class="reference"><a href="#cite_note-Murdoch_1966-43"><span class="cite-bracket">[</span>43<span class="cite-bracket">]</span></a></sup></li> <li>For trophic cascades to be ubiquitous, communities must generally act as food chains, with discrete trophic levels. Most communities, however, have complex <a href="/wiki/Food_webs" class="mw-redirect" title="Food webs">food webs</a>. In real food webs, consumers often feed at multiple trophic levels (<a href="/wiki/Omnivory" class="mw-redirect" title="Omnivory">omnivory</a>), organisms often change their <a href="/wiki/Diet_(nutrition)" title="Diet (nutrition)">diet</a> as they grow larger, <a href="/wiki/Cannibalism_(zoology)" class="mw-redirect" title="Cannibalism (zoology)">cannibalism</a> occurs, and consumers are subsidized by inputs of resources from outside the local community, all of which blur the distinctions between trophic levels.<sup id="cite_ref-44" class="reference"><a href="#cite_note-44"><span class="cite-bracket">[</span>44<span class="cite-bracket">]</span></a></sup></li></ul> <p>Antagonistically, this principle is sometimes called the "trophic trickle".<sup id="cite_ref-45" class="reference"><a href="#cite_note-45"><span class="cite-bracket">[</span>45<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-46" class="reference"><a href="#cite_note-46"><span class="cite-bracket">[</span>46<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_cascade&action=edit&section=6" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 30em;"> <ul><li><a href="/wiki/Cascade_effect_(ecology)" title="Cascade effect (ecology)">Cascade effect (ecology)</a></li> <li><a href="/wiki/Ecological_release" title="Ecological release">Ecological release</a></li> <li><a href="/wiki/Fishing_down_the_food_web" title="Fishing down the food web">Fishing down the food web</a></li> <li><a href="/wiki/Food_web" title="Food web">Food web</a></li> <li><a href="/wiki/Lake_ecology" class="mw-redirect" title="Lake ecology">Lake ecology</a></li> <li><a href="/wiki/Mesopredator_release_hypothesis" title="Mesopredator release hypothesis">Mesopredator release hypothesis</a></li> <li><a href="/wiki/Population_dynamics_of_fisheries" title="Population dynamics of fisheries">Population dynamics of fisheries</a></li> <li><a href="/wiki/Soil_food_web" title="Soil food web">Soil food web</a></li> <li><a href="/wiki/Stream_ecology" title="Stream ecology">Stream ecology</a></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Trophic_cascade&action=edit&section=7" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width reflist-columns-2"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFKottaWernbergJänesKotta2018" class="citation journal cs1">Kotta, J.; Wernberg, T.; Jänes, H.; Kotta, I.; Nurkse, K.; Pärnoja, M.; Orav-Kotta, H. (2018). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5897427">"Novel crab predator causes marine ecosystem regime shift"</a>. <i>Scientific Reports</i>. <b>8</b> (1): 4956. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2018NatSR...8.4956K">2018NatSR...8.4956K</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fs41598-018-23282-w">10.1038/s41598-018-23282-w</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5897427">5897427</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29651152">29651152</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Scientific+Reports&rft.atitle=Novel+crab+predator+causes+marine+ecosystem+regime+shift&rft.volume=8&rft.issue=1&rft.pages=4956&rft.date=2018&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5897427%23id-name%3DPMC&rft_id=info%3Apmid%2F29651152&rft_id=info%3Adoi%2F10.1038%2Fs41598-018-23282-w&rft_id=info%3Abibcode%2F2018NatSR...8.4956K&rft.aulast=Kotta&rft.aufirst=J.&rft.au=Wernberg%2C+T.&rft.au=J%C3%A4nes%2C+H.&rft.au=Kotta%2C+I.&rft.au=Nurkse%2C+K.&rft.au=P%C3%A4rnoja%2C+M.&rft.au=Orav-Kotta%2C+H.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5897427&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMegreyWerner" class="citation web cs1">Megrey, Bernard; Werner, Francisco. <a rel="nofollow" class="external text" href="https://pices.int/publications/presentations/PICES_13/PICES_13_S2/Megrey_S2.pdf">"Evaluating the Role of Topdown vs. Bottom-up Ecosystem Regulation from a Modeling Perspective"</a> <span class="cs1-format">(PDF)</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Evaluating+the+Role+of+Topdown+vs.+Bottom-up+Ecosystem+Regulation+from+a+Modeling+Perspective&rft.aulast=Megrey&rft.aufirst=Bernard&rft.au=Werner%2C+Francisco&rft_id=https%3A%2F%2Fpices.int%2Fpublications%2Fpresentations%2FPICES_13%2FPICES_13_S2%2FMegrey_S2.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMatsuzakiSuzukiKadoyaNakagawa2018" class="citation journal cs1">Matsuzaki, Shin-Ichiro S.; Suzuki, Kenta; Kadoya, Taku; Nakagawa, Megumi; Takamura, Noriko (2018). <a rel="nofollow" class="external text" href="https://doi.org/10.1002%2Fecy.2414">"Bottom-up linkages between primary production, zooplankton, and fish in a shallow, hypereutrophic lake"</a>. <i>Ecology</i>. <b>99</b> (9): <span class="nowrap">2025–</span>2036. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2018Ecol...99.2025M">2018Ecol...99.2025M</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1002%2Fecy.2414">10.1002/ecy.2414</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29884987">29884987</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:46996957">46996957</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecology&rft.atitle=Bottom-up+linkages+between+primary+production%2C+zooplankton%2C+and+fish+in+a+shallow%2C+hypereutrophic+lake&rft.volume=99&rft.issue=9&rft.pages=%3Cspan+class%3D%22nowrap%22%3E2025-%3C%2Fspan%3E2036&rft.date=2018&rft_id=info%3Adoi%2F10.1002%2Fecy.2414&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A46996957%23id-name%3DS2CID&rft_id=info%3Apmid%2F29884987&rft_id=info%3Abibcode%2F2018Ecol...99.2025M&rft.aulast=Matsuzaki&rft.aufirst=Shin-Ichiro+S.&rft.au=Suzuki%2C+Kenta&rft.au=Kadoya%2C+Taku&rft.au=Nakagawa%2C+Megumi&rft.au=Takamura%2C+Noriko&rft_id=https%3A%2F%2Fdoi.org%2F10.1002%252Fecy.2414&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLynamLlopeMöllmannHelaouët2017" class="citation journal cs1">Lynam, Christopher Philip; Llope, Marcos; Möllmann, Christian; Helaouët, Pierre; Bayliss-Brown, Georgia Anne; Stenseth, Nils C. (Feb 2017). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5338359">"Trophic and environmental control in the North Sea"</a>. <i>Proceedings of the National Academy of Sciences</i>. <b>114</b> (8): <span class="nowrap">1952–</span>1957. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.1621037114">10.1073/pnas.1621037114</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5338359">5338359</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/28167770">28167770</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences&rft.atitle=Trophic+and+environmental+control+in+the+North+Sea&rft.volume=114&rft.issue=8&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1952-%3C%2Fspan%3E1957&rft.date=2017-02&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5338359%23id-name%3DPMC&rft_id=info%3Apmid%2F28167770&rft_id=info%3Adoi%2F10.1073%2Fpnas.1621037114&rft.aulast=Lynam&rft.aufirst=Christopher+Philip&rft.au=Llope%2C+Marcos&rft.au=M%C3%B6llmann%2C+Christian&rft.au=Helaou%C3%ABt%2C+Pierre&rft.au=Bayliss-Brown%2C+Georgia+Anne&rft.au=Stenseth%2C+Nils+C.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5338359&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLuskin2017" class="citation journal cs1">Luskin, M. (2017). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5738359">"Cross-boundary subsidy cascades from oil palm degrade distant tropical forests"</a>. <i>Nature Communications</i>. <b>8</b> (8): 2231. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2017NatCo...8.2231L">2017NatCo...8.2231L</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fs41467-017-01920-7">10.1038/s41467-017-01920-7</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5738359">5738359</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29263381">29263381</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature+Communications&rft.atitle=Cross-boundary+subsidy+cascades+from+oil+palm+degrade+distant+tropical+forests&rft.volume=8&rft.issue=8&rft.pages=2231&rft.date=2017&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5738359%23id-name%3DPMC&rft_id=info%3Apmid%2F29263381&rft_id=info%3Adoi%2F10.1038%2Fs41467-017-01920-7&rft_id=info%3Abibcode%2F2017NatCo...8.2231L&rft.aulast=Luskin&rft.aufirst=M.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5738359&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFZhangQianGirardelloPellissier2018" class="citation journal cs1">Zhang, J.; Qian, H.; Girardello, M.; Pellissier, V.; Nielsen, S. E.; Svenning, J.-C. (2018). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6083253">"Trophic interactions among vertebrate guilds and plants shape global patterns in species diversity"</a>. <i>Proceedings of the Royal Society B: Biological Sciences</i>. <b>285</b> (1883): 20180949. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frspb.2018.0949">10.1098/rspb.2018.0949</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6083253">6083253</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/30051871">30051871</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+Royal+Society+B%3A+Biological+Sciences&rft.atitle=Trophic+interactions+among+vertebrate+guilds+and+plants+shape+global+patterns+in+species+diversity&rft.volume=285&rft.issue=1883&rft.pages=20180949&rft.date=2018&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6083253%23id-name%3DPMC&rft_id=info%3Apmid%2F30051871&rft_id=info%3Adoi%2F10.1098%2Frspb.2018.0949&rft.aulast=Zhang&rft.aufirst=J.&rft.au=Qian%2C+H.&rft.au=Girardello%2C+M.&rft.au=Pellissier%2C+V.&rft.au=Nielsen%2C+S.+E.&rft.au=Svenning%2C+J.-C.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6083253&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.uky.edu/Classes/ENT/530/Lecture_Notes/feb11/feb11_files/frame.htm">"University of Kentucky Lecture Notes"</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=University+of+Kentucky+Lecture+Notes&rft_id=http%3A%2F%2Fwww.uky.edu%2FClasses%2FENT%2F530%2FLecture_Notes%2Ffeb11%2Ffeb11_files%2Fframe.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilmersEstesEdwardsLaidre2012" class="citation journal cs1">Wilmers, C. C.; Estes, J. A.; Edwards, M.; Laidre, K. L.; Konar, B. (2012). <a rel="nofollow" class="external text" href="https://doi.org/10.1890%2F110176">"Do trophic cascades affect the storage and flux of atmospheric carbon? An analysis of sea otters and kelp forests"</a>. <i>Frontiers in Ecology and the Environment</i>. <b>10</b> (8): <span class="nowrap">409–</span>415. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012FrEE...10..409W">2012FrEE...10..409W</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1890%2F110176">10.1890/110176</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1540-9309">1540-9309</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:51684842">51684842</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Frontiers+in+Ecology+and+the+Environment&rft.atitle=Do+trophic+cascades+affect+the+storage+and+flux+of+atmospheric+carbon%3F+An+analysis+of+sea+otters+and+kelp+forests&rft.volume=10&rft.issue=8&rft.pages=%3Cspan+class%3D%22nowrap%22%3E409-%3C%2Fspan%3E415&rft.date=2012&rft_id=info%3Adoi%2F10.1890%2F110176&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A51684842%23id-name%3DS2CID&rft.issn=1540-9309&rft_id=info%3Abibcode%2F2012FrEE...10..409W&rft.aulast=Wilmers&rft.aufirst=C.+C.&rft.au=Estes%2C+J.+A.&rft.au=Edwards%2C+M.&rft.au=Laidre%2C+K.+L.&rft.au=Konar%2C+B.&rft_id=https%3A%2F%2Fdoi.org%2F10.1890%252F110176&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text">Leopold, A. (1949) <a rel="nofollow" class="external text" href="http://www.missionwolf.org/page/thinking-leopold/">"Thinking like a mountain"</a> in "Sand county almanac"</span> </li> <li id="cite_note-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-10">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHairstonSmithSlobodkin1960" class="citation journal cs1">Hairston, NG; Smith, FE; Slobodkin, LB (1960). "Community structure, population control and competition". <i>American Naturalist</i>. <b>94</b> (879): <span class="nowrap">421–</span>425. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1086%2F282146">10.1086/282146</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:84548124">84548124</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Naturalist&rft.atitle=Community+structure%2C+population+control+and+competition&rft.volume=94&rft.issue=879&rft.pages=%3Cspan+class%3D%22nowrap%22%3E421-%3C%2Fspan%3E425&rft.date=1960&rft_id=info%3Adoi%2F10.1086%2F282146&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A84548124%23id-name%3DS2CID&rft.aulast=Hairston&rft.aufirst=NG&rft.au=Smith%2C+FE&rft.au=Slobodkin%2C+LB&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHrbáčekDvořakovaKořínekProcházkóva1961" class="citation journal cs1">Hrbáček, J; Dvořakova, M; Kořínek, V; Procházkóva, L (1961). "Demonstration of the effect of the fish stock on the species composition of zooplankton and the intensity of metabolism of the whole plankton association". <i>Verh. Internat. Verein. Limnol</i>. <b>14</b> (1): <span class="nowrap">192–</span>195. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1961SILP...14..192H">1961SILP...14..192H</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1080%2F03680770.1959.11899269">10.1080/03680770.1959.11899269</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Verh.+Internat.+Verein.+Limnol&rft.atitle=Demonstration+of+the+effect+of+the+fish+stock+on+the+species+composition+of+zooplankton+and+the+intensity+of+metabolism+of+the+whole+plankton+association&rft.volume=14&rft.issue=1&rft.pages=%3Cspan+class%3D%22nowrap%22%3E192-%3C%2Fspan%3E195&rft.date=1961&rft_id=info%3Adoi%2F10.1080%2F03680770.1959.11899269&rft_id=info%3Abibcode%2F1961SILP...14..192H&rft.aulast=Hrb%C3%A1%C4%8Dek&rft.aufirst=J&rft.au=Dvo%C5%99akova%2C+M&rft.au=Ko%C5%99%C3%ADnek%2C+V&rft.au=Proch%C3%A1zk%C3%B3va%2C+L&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFOksanenFretwellArrudaNiemala1981" class="citation journal cs1">Oksanen, L; Fretwell, SD; Arruda, J; Niemala, P (1981). "Exploitation ecosystems in gradients of primary productivity". <i>American Naturalist</i>. <b>118</b> (2): <span class="nowrap">240–</span>261. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1086%2F283817">10.1086/283817</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:84215344">84215344</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Naturalist&rft.atitle=Exploitation+ecosystems+in+gradients+of+primary+productivity&rft.volume=118&rft.issue=2&rft.pages=%3Cspan+class%3D%22nowrap%22%3E240-%3C%2Fspan%3E261&rft.date=1981&rft_id=info%3Adoi%2F10.1086%2F283817&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A84215344%23id-name%3DS2CID&rft.aulast=Oksanen&rft.aufirst=L&rft.au=Fretwell%2C+SD&rft.au=Arruda%2C+J&rft.au=Niemala%2C+P&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCarpenterKitchellHodgson1985" class="citation journal cs1">Carpenter, SR; Kitchell, JF; Hodgson, JR (1985). "Cascading trophic interactions and lake productivity". <i>BioScience</i>. <b>35</b> (10): <span class="nowrap">634–</span>639. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F1309989">10.2307/1309989</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/1309989">1309989</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=BioScience&rft.atitle=Cascading+trophic+interactions+and+lake+productivity&rft.volume=35&rft.issue=10&rft.pages=%3Cspan+class%3D%22nowrap%22%3E634-%3C%2Fspan%3E639&rft.date=1985&rft_id=info%3Adoi%2F10.2307%2F1309989&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F1309989%23id-name%3DJSTOR&rft.aulast=Carpenter&rft.aufirst=SR&rft.au=Kitchell%2C+JF&rft.au=Hodgson%2C+JR&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPower1990" class="citation journal cs1">Power, ME (1990). "Effects of fish in river food webs". <i>Science</i>. <b>250</b> (4982): <span class="nowrap">811–</span>814. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1990Sci...250..811P">1990Sci...250..811P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.250.4982.811">10.1126/science.250.4982.811</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/17759974">17759974</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:24780727">24780727</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Effects+of+fish+in+river+food+webs&rft.volume=250&rft.issue=4982&rft.pages=%3Cspan+class%3D%22nowrap%22%3E811-%3C%2Fspan%3E814&rft.date=1990&rft_id=info%3Adoi%2F10.1126%2Fscience.250.4982.811&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A24780727%23id-name%3DS2CID&rft_id=info%3Apmid%2F17759974&rft_id=info%3Abibcode%2F1990Sci...250..811P&rft.aulast=Power&rft.aufirst=ME&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-uwo.academia-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-uwo.academia_15-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSzpakOrchardSalomonGröcke2013" class="citation journal cs1">Szpak, Paul; Orchard, Trevor J.; Salomon, Anne K.; Gröcke, Darren R. (2013). <a rel="nofollow" class="external text" href="https://www.academia.edu/2568296">"Regional ecological variability and impact of the maritime fur trade on nearshore ecosystems in southern Haida Gwaii (British Columbia, Canada): evidence from stable isotope analysis of rockfish (Sebastes spp.) bone collagen"</a>. <i><a href="/w/index.php?title=Archaeological_and_Anthropological_Sciences_(journal)&action=edit&redlink=1" class="new" title="Archaeological and Anthropological Sciences (journal) (page does not exist)">Archaeological and Anthropological Sciences</a></i>. <b>5</b> (X): XX. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2013ArAnS...5..159S">2013ArAnS...5..159S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2Fs12520-013-0122-y">10.1007/s12520-013-0122-y</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:84866250">84866250</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Archaeological+and+Anthropological+Sciences&rft.atitle=Regional+ecological+variability+and+impact+of+the+maritime+fur+trade+on+nearshore+ecosystems+in+southern+Haida+Gwaii+%28British+Columbia%2C+Canada%29%3A+evidence+from+stable+isotope+analysis+of+rockfish+%28Sebastes+spp.%29+bone+collagen&rft.volume=5&rft.issue=X&rft.pages=XX&rft.date=2013&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A84866250%23id-name%3DS2CID&rft_id=info%3Adoi%2F10.1007%2Fs12520-013-0122-y&rft_id=info%3Abibcode%2F2013ArAnS...5..159S&rft.aulast=Szpak&rft.aufirst=Paul&rft.au=Orchard%2C+Trevor+J.&rft.au=Salomon%2C+Anne+K.&rft.au=Gr%C3%B6cke%2C+Darren+R.&rft_id=https%3A%2F%2Fwww.academia.edu%2F2568296&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-16">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFEstesPalmisano1974" class="citation journal cs1">Estes, JA; Palmisano, JF (1974). "Sea otters: their role in structuring nearshore communities". <i>Science</i>. <b>185</b> (4156): <span class="nowrap">1058–</span>1060. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1974Sci...185.1058E">1974Sci...185.1058E</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.185.4156.1058">10.1126/science.185.4156.1058</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/17738247">17738247</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:35892592">35892592</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Sea+otters%3A+their+role+in+structuring+nearshore+communities&rft.volume=185&rft.issue=4156&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1058-%3C%2Fspan%3E1060&rft.date=1974&rft_id=info%3Adoi%2F10.1126%2Fscience.185.4156.1058&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A35892592%23id-name%3DS2CID&rft_id=info%3Apmid%2F17738247&rft_id=info%3Abibcode%2F1974Sci...185.1058E&rft.aulast=Estes&rft.aufirst=JA&rft.au=Palmisano%2C+JF&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWeston2022" class="citation news cs1">Weston, Phoebe (2022-06-23). <a rel="nofollow" class="external text" href="https://www.theguardian.com/environment/2022/jun/23/rebalancing-act-bringing-back-wolf-fix-broken-ecosystem-aoe">"<span class="cs1-kern-left"></span>'People may be overselling the myth': should we bring back the wolf?"</a>. <i>The Guardian</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2022-06-24</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Guardian&rft.atitle=%27People+may+be+overselling+the+myth%27%3A+should+we+bring+back+the+wolf%3F&rft.date=2022-06-23&rft.aulast=Weston&rft.aufirst=Phoebe&rft_id=https%3A%2F%2Fwww.theguardian.com%2Fenvironment%2F2022%2Fjun%2F23%2Frebalancing-act-bringing-back-wolf-fix-broken-ecosystem-aoe&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-18"><span class="mw-cite-backlink"><b><a href="#cite_ref-18">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFStrong1992" class="citation journal cs1">Strong, D. R. (1992). "Are trophic cascades all wet? Differentiation and donor-control in speciose ecosystems". <i>Ecology</i>. <b>73</b> (3): <span class="nowrap">747–</span>754. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1992Ecol...73..747S">1992Ecol...73..747S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F1940154">10.2307/1940154</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/1940154">1940154</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecology&rft.atitle=Are+trophic+cascades+all+wet%3F+Differentiation+and+donor-control+in+speciose+ecosystems&rft.volume=73&rft.issue=3&rft.pages=%3Cspan+class%3D%22nowrap%22%3E747-%3C%2Fspan%3E754&rft.date=1992&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F1940154%23id-name%3DJSTOR&rft_id=info%3Adoi%2F10.2307%2F1940154&rft_id=info%3Abibcode%2F1992Ecol...73..747S&rft.aulast=Strong&rft.aufirst=D.+R.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-19"><span class="mw-cite-backlink"><b><a href="#cite_ref-19">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFStrongWhippleChildDennis1999" class="citation journal cs1">Strong, D. R.; Whipple, A. V.; Child, A. L.; Dennis, B. (1999). "Model selection for a subterranean trophic cascade: Root-feeding caterpillars and entomopathogenic nematodes". <i>Ecology</i>. <b>80</b> (8): <span class="nowrap">2750–</span>2761. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F177255">10.2307/177255</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/177255">177255</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecology&rft.atitle=Model+selection+for+a+subterranean+trophic+cascade%3A+Root-feeding+caterpillars+and+entomopathogenic+nematodes&rft.volume=80&rft.issue=8&rft.pages=%3Cspan+class%3D%22nowrap%22%3E2750-%3C%2Fspan%3E2761&rft.date=1999&rft_id=info%3Adoi%2F10.2307%2F177255&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F177255%23id-name%3DJSTOR&rft.aulast=Strong&rft.aufirst=D.+R.&rft.au=Whipple%2C+A.+V.&rft.au=Child%2C+A.+L.&rft.au=Dennis%2C+B.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-20"><span class="mw-cite-backlink"><b><a href="#cite_ref-20">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPreisser2003" class="citation journal cs1">Preisser, E. L. (2003). "Field evidence for a rapidly cascading underground food web". <i>Ecology</i>. <b>84</b> (4): <span class="nowrap">869–</span>874. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1890%2F0012-9658%282003%29084%5B0869%3Afefarc%5D2.0.co%3B2">10.1890/0012-9658(2003)084[0869:fefarc]2.0.co;2</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecology&rft.atitle=Field+evidence+for+a+rapidly+cascading+underground+food+web&rft.volume=84&rft.issue=4&rft.pages=%3Cspan+class%3D%22nowrap%22%3E869-%3C%2Fspan%3E874&rft.date=2003&rft_id=info%3Adoi%2F10.1890%2F0012-9658%282003%29084%5B0869%3Afefarc%5D2.0.co%3B2&rft.aulast=Preisser&rft.aufirst=E.+L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-21"><span class="mw-cite-backlink"><b><a href="#cite_ref-21">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLetourneauDyer1998" class="citation journal cs1">Letourneau, D. K.; Dyer, L. A. (1998). "Experimental test in lowland tropical forest shows top-down effects through four trophic levels". <i>Ecology</i>. <b>79</b> (5): <span class="nowrap">1678–</span>1687. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F176787">10.2307/176787</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/176787">176787</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecology&rft.atitle=Experimental+test+in+lowland+tropical+forest+shows+top-down+effects+through+four+trophic+levels&rft.volume=79&rft.issue=5&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1678-%3C%2Fspan%3E1687&rft.date=1998&rft_id=info%3Adoi%2F10.2307%2F176787&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F176787%23id-name%3DJSTOR&rft.aulast=Letourneau&rft.aufirst=D.+K.&rft.au=Dyer%2C+L.+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-22"><span class="mw-cite-backlink"><b><a href="#cite_ref-22">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPolisSearsHuxel2000" class="citation journal cs1">Polis, G. A.; Sears, A. L. W.; Huxel, G. R.; et al. (2000). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fs0169-5347%2800%2901971-6">"When is a trophic cascade a trophic cascade?"</a>. <i>Trends in Ecology & Evolution</i>. <b>15</b> (11): <span class="nowrap">473–</span>475. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2000TEcoE..15..473P">2000TEcoE..15..473P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fs0169-5347%2800%2901971-6">10.1016/s0169-5347(00)01971-6</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11050351">11050351</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Trends+in+Ecology+%26+Evolution&rft.atitle=When+is+a+trophic+cascade+a+trophic+cascade%3F&rft.volume=15&rft.issue=11&rft.pages=%3Cspan+class%3D%22nowrap%22%3E473-%3C%2Fspan%3E475&rft.date=2000&rft_id=info%3Apmid%2F11050351&rft_id=info%3Adoi%2F10.1016%2Fs0169-5347%2800%2901971-6&rft_id=info%3Abibcode%2F2000TEcoE..15..473P&rft.aulast=Polis&rft.aufirst=G.+A.&rft.au=Sears%2C+A.+L.+W.&rft.au=Huxel%2C+G.+R.&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252Fs0169-5347%252800%252901971-6&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-23"><span class="mw-cite-backlink"><b><a href="#cite_ref-23">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFShurinBorerSeabloomAnderson2002" class="citation journal cs1">Shurin, J. B.; Borer, E. T.; Seabloom, E. W.; Anderson, K.; Blanchette, C. A.; Broitman, B; Cooper, S. D.; Halpern, B. S. (2002). "A cross-ecosystem comparison of the strength of trophic cascades". <i>Ecology Letters</i>. <b>5</b> (6): <span class="nowrap">785–</span>791. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2002EcolL...5..785S">2002EcolL...5..785S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1046%2Fj.1461-0248.2002.00381.x">10.1046/j.1461-0248.2002.00381.x</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecology+Letters&rft.atitle=A+cross-ecosystem+comparison+of+the+strength+of+trophic+cascades&rft.volume=5&rft.issue=6&rft.pages=%3Cspan+class%3D%22nowrap%22%3E785-%3C%2Fspan%3E791&rft.date=2002&rft_id=info%3Adoi%2F10.1046%2Fj.1461-0248.2002.00381.x&rft_id=info%3Abibcode%2F2002EcolL...5..785S&rft.aulast=Shurin&rft.aufirst=J.+B.&rft.au=Borer%2C+E.+T.&rft.au=Seabloom%2C+E.+W.&rft.au=Anderson%2C+K.&rft.au=Blanchette%2C+C.+A.&rft.au=Broitman%2C+B&rft.au=Cooper%2C+S.+D.&rft.au=Halpern%2C+B.+S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-24"><span class="mw-cite-backlink"><b><a href="#cite_ref-24">^</a></b></span> <span class="reference-text">Beschta, R.L., and W.J. Ripple. 2009. Large predators and trophic cascades in terrestrial ecosystems of the western United States Biological Conservation. 142, 2009: 2401–2414.</span> </li> <li id="cite_note-25"><span class="mw-cite-backlink"><b><a href="#cite_ref-25">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBeschtaRipple2011" class="citation journal cs1 cs1-prop-long-vol">Beschta, R. L.; Ripple, W. J. (2011). "The role of large predators in maintaining riparian plant communities and river morphology". <i>Geomorphology</i>. <span class="nowrap">157–</span>158: <span class="nowrap">88–</span>98. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.geomorph.2011.04.042">10.1016/j.geomorph.2011.04.042</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Geomorphology&rft.atitle=The+role+of+large+predators+in+maintaining+riparian+plant+communities+and+river+morphology&rft.volume=157%E2%80%93158&rft.pages=%3Cspan+class%3D%22nowrap%22%3E88-%3C%2Fspan%3E98&rft.date=2011&rft_id=info%3Adoi%2F10.1016%2Fj.geomorph.2011.04.042&rft.aulast=Beschta&rft.aufirst=R.+L.&rft.au=Ripple%2C+W.+J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-rippbesch2012-26"><span class="mw-cite-backlink">^ <a href="#cite_ref-rippbesch2012_26-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-rippbesch2012_26-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-rippbesch2012_26-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRippleBeschta2012" class="citation journal cs1">Ripple, W. J.; Beschta, R. L. (2012). "Trophic cascades in Yellowstone: The first 15 years after wolf reintroduction". <i>Biological Conservation</i>. <b>145</b> (1): <span class="nowrap">205–</span>213. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012BCons.145..205R">2012BCons.145..205R</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.biocon.2011.11.005">10.1016/j.biocon.2011.11.005</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:9750513">9750513</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biological+Conservation&rft.atitle=Trophic+cascades+in+Yellowstone%3A+The+first+15+years+after+wolf+reintroduction&rft.volume=145&rft.issue=1&rft.pages=%3Cspan+class%3D%22nowrap%22%3E205-%3C%2Fspan%3E213&rft.date=2012&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A9750513%23id-name%3DS2CID&rft_id=info%3Adoi%2F10.1016%2Fj.biocon.2011.11.005&rft_id=info%3Abibcode%2F2012BCons.145..205R&rft.aulast=Ripple&rft.aufirst=W.+J.&rft.au=Beschta%2C+R.+L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-27"><span class="mw-cite-backlink"><b><a href="#cite_ref-27">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGroshong2004" class="citation thesis cs1">Groshong, L. C. (2004). <i>Mapping Riparian Vegetation Change in Yellowstone's Northern Range using High Spatial Resolution Imagery</i> (MA Thesis). Eugene, Oregon, USA: University of Oregon.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adissertation&rft.title=Mapping+Riparian+Vegetation+Change+in+Yellowstone%27s+Northern+Range+using+High+Spatial+Resolution+Imagery&rft.inst=University+of+Oregon&rft.date=2004&rft.aulast=Groshong&rft.aufirst=L.+C.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-28"><span class="mw-cite-backlink"><b><a href="#cite_ref-28">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRippleBeschta2004" class="citation journal cs1">Ripple, W.J.; Beschta, R.L. (2004). "Wolves, elk, willows, and trophic cascades in the upper Gallatin Range of Southwestern Montana, USA". <i>Forest Ecology and Management</i>. <b>200</b> (<span class="nowrap">1–</span>3): <span class="nowrap">161–</span>181. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2004ForEM.200..161R">2004ForEM.200..161R</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.foreco.2004.06.017">10.1016/j.foreco.2004.06.017</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Forest+Ecology+and+Management&rft.atitle=Wolves%2C+elk%2C+willows%2C+and+trophic+cascades+in+the+upper+Gallatin+Range+of+Southwestern+Montana%2C+USA&rft.volume=200&rft.issue=%3Cspan+class%3D%22nowrap%22%3E1%E2%80%93%3C%2Fspan%3E3&rft.pages=%3Cspan+class%3D%22nowrap%22%3E161-%3C%2Fspan%3E181&rft.date=2004&rft_id=info%3Adoi%2F10.1016%2Fj.foreco.2004.06.017&rft_id=info%3Abibcode%2F2004ForEM.200..161R&rft.aulast=Ripple&rft.aufirst=W.J.&rft.au=Beschta%2C+R.L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-29"><span class="mw-cite-backlink"><b><a href="#cite_ref-29">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBeschtaRipple2007" class="citation journal cs1">Beschta, R.L.; Ripple, W.J. (2007). <a rel="nofollow" class="external text" href="https://scholarsarchive.byu.edu/cgi/viewcontent.cgi?article=2114&context=wnan">"Increased Willow Heights along northern Yellowstone's Blacktail Deer Creek following wolf reintroduction"</a>. <i>Western North American Naturalist</i>. <b>67</b> (4): <span class="nowrap">613–</span>617. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.3398%2F1527-0904%282007%2967%5B613%3Aiwhany%5D2.0.co%3B2">10.3398/1527-0904(2007)67[613:iwhany]2.0.co;2</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:85023589">85023589</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Western+North+American+Naturalist&rft.atitle=Increased+Willow+Heights+along+northern+Yellowstone%27s+Blacktail+Deer+Creek+following+wolf+reintroduction&rft.volume=67&rft.issue=4&rft.pages=%3Cspan+class%3D%22nowrap%22%3E613-%3C%2Fspan%3E617&rft.date=2007&rft_id=info%3Adoi%2F10.3398%2F1527-0904%282007%2967%5B613%3Aiwhany%5D2.0.co%3B2&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A85023589%23id-name%3DS2CID&rft.aulast=Beschta&rft.aufirst=R.L.&rft.au=Ripple%2C+W.J.&rft_id=https%3A%2F%2Fscholarsarchive.byu.edu%2Fcgi%2Fviewcontent.cgi%3Farticle%3D2114%26context%3Dwnan&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-baril2009-30"><span class="mw-cite-backlink">^ <a href="#cite_ref-baril2009_30-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-baril2009_30-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBaril2009" class="citation thesis cs1">Baril, L. M. (2009). <i>Change in Deciduous Woody Vegetation, Implications of Increased Willow (</i>Salix<i> spp.) Growth for Bird Species Diversity and Willow Species Composition in and around Yellowstone National Park's Northern Range</i> (MS). Bozeman, USA: Montana State University.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adissertation&rft.title=Change+in+Deciduous+Woody+Vegetation%2C+Implications+of+Increased+Willow+%28Salix+spp.%29+Growth+for+Bird+Species+Diversity+and+Willow+Species+Composition+in+and+around+Yellowstone+National+Park%27s+Northern+Range&rft.inst=Montana+State+University&rft.date=2009&rft.aulast=Baril&rft.aufirst=L.+M.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-31"><span class="mw-cite-backlink"><b><a href="#cite_ref-31">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilmersCrabtreeSmithMurphy2003" class="citation journal cs1">Wilmers, C.C.; Crabtree, R. L.; Smith, D. W.; Murphy, K. M.; Getz, W. M. (2003). <a rel="nofollow" class="external text" href="https://escholarship.org/uc/item/0h09s21c">"Trophic facilitation by introduced top predators: grey wolf subsidies to scavengers in Yellowstone National Park"</a>. <i>Journal of Animal Ecology</i>. <b>72</b> (6): <span class="nowrap">909–</span>916. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2003JAnEc..72..909W">2003JAnEc..72..909W</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1046%2Fj.1365-2656.2003.00766.x">10.1046/j.1365-2656.2003.00766.x</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Animal+Ecology&rft.atitle=Trophic+facilitation+by+introduced+top+predators%3A+grey+wolf+subsidies+to+scavengers+in+Yellowstone+National+Park&rft.volume=72&rft.issue=6&rft.pages=%3Cspan+class%3D%22nowrap%22%3E909-%3C%2Fspan%3E916&rft.date=2003&rft_id=info%3Adoi%2F10.1046%2Fj.1365-2656.2003.00766.x&rft_id=info%3Abibcode%2F2003JAnEc..72..909W&rft.aulast=Wilmers&rft.aufirst=C.C.&rft.au=Crabtree%2C+R.+L.&rft.au=Smith%2C+D.+W.&rft.au=Murphy%2C+K.+M.&rft.au=Getz%2C+W.+M.&rft_id=https%3A%2F%2Fescholarship.org%2Fuc%2Fitem%2F0h09s21c&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-32"><span class="mw-cite-backlink"><b><a href="#cite_ref-32">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPainterRipple2012" class="citation journal cs1">Painter, L. E.; Ripple, W. J. (2012). "Effects of bison on willow and cottonwood in northern Yellowstone National Park". <i>Forest Ecology and Management</i>. <b>264</b>: <span class="nowrap">150–</span>158. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012ForEM.264..150P">2012ForEM.264..150P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.foreco.2011.10.010">10.1016/j.foreco.2011.10.010</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Forest+Ecology+and+Management&rft.atitle=Effects+of+bison+on+willow+and+cottonwood+in+northern+Yellowstone+National+Park&rft.volume=264&rft.pages=%3Cspan+class%3D%22nowrap%22%3E150-%3C%2Fspan%3E158&rft.date=2012&rft_id=info%3Adoi%2F10.1016%2Fj.foreco.2011.10.010&rft_id=info%3Abibcode%2F2012ForEM.264..150P&rft.aulast=Painter&rft.aufirst=L.+E.&rft.au=Ripple%2C+W.+J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-33"><span class="mw-cite-backlink"><b><a href="#cite_ref-33">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRippleBeschta2006" class="citation journal cs1">Ripple, W.J.; Beschta, R.L. (2006). "Linking a cougar decline, trophic cascade, and catastrophic regime shift in Zion National Park". <i>Biological Conservation</i>. <b>133</b> (4): <span class="nowrap">397–</span>408. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2006BCons.133..397R">2006BCons.133..397R</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.biocon.2006.07.002">10.1016/j.biocon.2006.07.002</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biological+Conservation&rft.atitle=Linking+a+cougar+decline%2C+trophic+cascade%2C+and+catastrophic+regime+shift+in+Zion+National+Park&rft.volume=133&rft.issue=4&rft.pages=%3Cspan+class%3D%22nowrap%22%3E397-%3C%2Fspan%3E408&rft.date=2006&rft_id=info%3Adoi%2F10.1016%2Fj.biocon.2006.07.002&rft_id=info%3Abibcode%2F2006BCons.133..397R&rft.aulast=Ripple&rft.aufirst=W.J.&rft.au=Beschta%2C+R.L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-34"><span class="mw-cite-backlink"><b><a href="#cite_ref-34">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRippleBeschta2008" class="citation journal cs1">Ripple, W.J.; Beschta, R.L. (2008). "Trophic cascades involving cougar, mule deer, and black oaks in Yosemite National Park". <i>Biological Conservation</i>. <b>141</b> (5): <span class="nowrap">1249–</span>1256. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2008BCons.141.1249R">2008BCons.141.1249R</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.biocon.2008.02.028">10.1016/j.biocon.2008.02.028</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biological+Conservation&rft.atitle=Trophic+cascades+involving+cougar%2C+mule+deer%2C+and+black+oaks+in+Yosemite+National+Park&rft.volume=141&rft.issue=5&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1249-%3C%2Fspan%3E1256&rft.date=2008&rft_id=info%3Adoi%2F10.1016%2Fj.biocon.2008.02.028&rft_id=info%3Abibcode%2F2008BCons.141.1249R&rft.aulast=Ripple&rft.aufirst=W.J.&rft.au=Beschta%2C+R.L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-35"><span class="mw-cite-backlink"><b><a href="#cite_ref-35">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFEstes2011" class="citation journal cs1">Estes, James A.; et al. (2011). "2011. Trophic Downgrading of Planet Earth". <i>Science</i>. <b>333</b> (6040): <span class="nowrap">301–</span>306. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2011Sci...333..301E">2011Sci...333..301E</a>. <a href="/wiki/CiteSeerX_(identifier)" class="mw-redirect" title="CiteSeerX (identifier)">CiteSeerX</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.701.8043">10.1.1.701.8043</a></span>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1205106">10.1126/science.1205106</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21764740">21764740</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:7752940">7752940</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=2011.+Trophic+Downgrading+of+Planet+Earth&rft.volume=333&rft.issue=6040&rft.pages=%3Cspan+class%3D%22nowrap%22%3E301-%3C%2Fspan%3E306&rft.date=2011&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A7752940%23id-name%3DS2CID&rft_id=info%3Abibcode%2F2011Sci...333..301E&rft_id=https%3A%2F%2Fciteseerx.ist.psu.edu%2Fviewdoc%2Fsummary%3Fdoi%3D10.1.1.701.8043%23id-name%3DCiteSeerX&rft_id=info%3Apmid%2F21764740&rft_id=info%3Adoi%2F10.1126%2Fscience.1205106&rft.aulast=Estes&rft.aufirst=James+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-36"><span class="mw-cite-backlink"><b><a href="#cite_ref-36">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPrugh2009" class="citation journal cs1">Prugh, Laura R.; et al. (2009). "2009. The Rise of the Mesopredator". <i>BioScience</i>. <b>59</b> (9): <span class="nowrap">779–</span>791. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1525%2Fbio.2009.59.9.9">10.1525/bio.2009.59.9.9</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:40484905">40484905</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=BioScience&rft.atitle=2009.+The+Rise+of+the+Mesopredator&rft.volume=59&rft.issue=9&rft.pages=%3Cspan+class%3D%22nowrap%22%3E779-%3C%2Fspan%3E791&rft.date=2009&rft_id=info%3Adoi%2F10.1525%2Fbio.2009.59.9.9&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A40484905%23id-name%3DS2CID&rft.aulast=Prugh&rft.aufirst=Laura+R.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-37"><span class="mw-cite-backlink"><b><a href="#cite_ref-37">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLetnicDworjanyn2011" class="citation journal cs1">Letnic, M.; Dworjanyn, S.A. (2011). "Does a top predator reduce the predatory impact of an invasive mesopredator on an endangered rodent?". <i>Ecography</i>. <b>34</b> (5): <span class="nowrap">827–</span>835. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2011Ecogr..34..827L">2011Ecogr..34..827L</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.1600-0587.2010.06516.x">10.1111/j.1600-0587.2010.06516.x</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecography&rft.atitle=Does+a+top+predator+reduce+the+predatory+impact+of+an+invasive+mesopredator+on+an+endangered+rodent%3F&rft.volume=34&rft.issue=5&rft.pages=%3Cspan+class%3D%22nowrap%22%3E827-%3C%2Fspan%3E835&rft.date=2011&rft_id=info%3Adoi%2F10.1111%2Fj.1600-0587.2010.06516.x&rft_id=info%3Abibcode%2F2011Ecogr..34..827L&rft.aulast=Letnic&rft.aufirst=M.&rft.au=Dworjanyn%2C+S.A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-38"><span class="mw-cite-backlink"><b><a href="#cite_ref-38">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFrankPetrieChoiLeggett2005" class="citation journal cs1">Frank, K. T.; Petrie, B.; Choi, J. S.; Leggett, W. C. (2005). "Trophic Cascades in a Formerly Cod-Dominated Ecosystem". <i>Science</i>. <b>308</b> (5728): <span class="nowrap">1621–</span>1623. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2005Sci...308.1621F">2005Sci...308.1621F</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1113075">10.1126/science.1113075</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0036-8075">0036-8075</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15947186">15947186</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:45088691">45088691</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Trophic+Cascades+in+a+Formerly+Cod-Dominated+Ecosystem&rft.volume=308&rft.issue=5728&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1621-%3C%2Fspan%3E1623&rft.date=2005&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A45088691%23id-name%3DS2CID&rft_id=info%3Abibcode%2F2005Sci...308.1621F&rft.issn=0036-8075&rft_id=info%3Adoi%2F10.1126%2Fscience.1113075&rft_id=info%3Apmid%2F15947186&rft.aulast=Frank&rft.aufirst=K.+T.&rft.au=Petrie%2C+B.&rft.au=Choi%2C+J.+S.&rft.au=Leggett%2C+W.+C.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-39"><span class="mw-cite-backlink"><b><a href="#cite_ref-39">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAlheitMöllmannDutzKornilovs2005" class="citation journal cs1">Alheit, J; Möllmann, C; Dutz, J; Kornilovs, G; Loewe, P; Mohrholz, V; Wasmund, N (2005). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.icesjms.2005.04.024">"Synchronous ecological regime shifts in the central Baltic and the North Sea in the late 1980s"</a>. <i>ICES Journal of Marine Science</i>. <b>62</b> (7): <span class="nowrap">1205–</span>1215. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2005ICJMS..62.1205A">2005ICJMS..62.1205A</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.icesjms.2005.04.024">10.1016/j.icesjms.2005.04.024</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=ICES+Journal+of+Marine+Science&rft.atitle=Synchronous+ecological+regime+shifts+in+the+central+Baltic+and+the+North+Sea+in+the+late+1980s&rft.volume=62&rft.issue=7&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1205-%3C%2Fspan%3E1215&rft.date=2005&rft_id=info%3Adoi%2F10.1016%2Fj.icesjms.2005.04.024&rft_id=info%3Abibcode%2F2005ICJMS..62.1205A&rft.aulast=Alheit&rft.aufirst=J&rft.au=M%C3%B6llmann%2C+C&rft.au=Dutz%2C+J&rft.au=Kornilovs%2C+G&rft.au=Loewe%2C+P&rft.au=Mohrholz%2C+V&rft.au=Wasmund%2C+N&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252Fj.icesjms.2005.04.024&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-40"><span class="mw-cite-backlink"><b><a href="#cite_ref-40">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMollmannMuller-KarulisKornilovsSt_John2008" class="citation journal cs1">Mollmann, C.; Muller-Karulis, B.; Kornilovs, G.; St John, M. A. (2008). <a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Ficesjms%2Ffsm197">"Effects of climate and overfishing on zooplankton dynamics and ecosystem structure: regime shifts, trophic cascade, and feedback loops in a simple ecosystem"</a>. <i>ICES Journal of Marine Science</i>. <b>65</b> (3): <span class="nowrap">302–</span>310. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Ficesjms%2Ffsm197">10.1093/icesjms/fsm197</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=ICES+Journal+of+Marine+Science&rft.atitle=Effects+of+climate+and+overfishing+on+zooplankton+dynamics+and+ecosystem+structure%3A+regime+shifts%2C+trophic+cascade%2C+and+feedback+loops+in+a+simple+ecosystem&rft.volume=65&rft.issue=3&rft.pages=%3Cspan+class%3D%22nowrap%22%3E302-%3C%2Fspan%3E310&rft.date=2008&rft_id=info%3Adoi%2F10.1093%2Ficesjms%2Ffsm197&rft.aulast=Mollmann&rft.aufirst=C.&rft.au=Muller-Karulis%2C+B.&rft.au=Kornilovs%2C+G.&rft.au=St+John%2C+M.+A.&rft_id=https%3A%2F%2Fdoi.org%2F10.1093%252Ficesjms%252Ffsm197&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-41"><span class="mw-cite-backlink"><b><a href="#cite_ref-41">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLohPawlik2014" class="citation journal cs1">Loh, T.-L.; Pawlik, J. R. (2014). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3964098">"Chemical defenses and resource trade-offs structure sponge communities on Caribbean coral reefs"</a>. <i>Proceedings of the National Academy of Sciences</i>. <b>111</b> (11): <span class="nowrap">4151–</span>4156. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2014PNAS..111.4151L">2014PNAS..111.4151L</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.1321626111">10.1073/pnas.1321626111</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0027-8424">0027-8424</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3964098">3964098</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/24567392">24567392</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences&rft.atitle=Chemical+defenses+and+resource+trade-offs+structure+sponge+communities+on+Caribbean+coral+reefs&rft.volume=111&rft.issue=11&rft.pages=%3Cspan+class%3D%22nowrap%22%3E4151-%3C%2Fspan%3E4156&rft.date=2014&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3964098%23id-name%3DPMC&rft_id=info%3Abibcode%2F2014PNAS..111.4151L&rft_id=info%3Apmid%2F24567392&rft_id=info%3Adoi%2F10.1073%2Fpnas.1321626111&rft.issn=0027-8424&rft.aulast=Loh&rft.aufirst=T.-L.&rft.au=Pawlik%2C+J.+R.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3964098&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-42"><span class="mw-cite-backlink"><b><a href="#cite_ref-42">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLoh2015" class="citation journal cs1">Loh, T.-L.; et al. (2015). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4419544">"Indirect effects of overfishing on Caribbean reefs: sponges overgrow reef-building corals"</a>. <i>PeerJ</i>. <b>3</b>: e901. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.7717%2Fpeerj.901">10.7717/peerj.901</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4419544">4419544</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/25945305">25945305</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=PeerJ&rft.atitle=Indirect+effects+of+overfishing+on+Caribbean+reefs%3A+sponges+overgrow+reef-building+corals&rft.volume=3&rft.pages=e901&rft.date=2015&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC4419544%23id-name%3DPMC&rft_id=info%3Apmid%2F25945305&rft_id=info%3Adoi%2F10.7717%2Fpeerj.901&rft.aulast=Loh&rft.aufirst=T.-L.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC4419544&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-Murdoch_1966-43"><span class="mw-cite-backlink">^ <a href="#cite_ref-Murdoch_1966_43-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Murdoch_1966_43-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMurdoch1966" class="citation journal cs1">Murdoch, WM (1966). "Community structure, population control, and competition – a critique". <i>American Naturalist</i>. <b>100</b> (912): <span class="nowrap">219–</span>226. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1086%2F282415">10.1086/282415</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:84354616">84354616</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Naturalist&rft.atitle=Community+structure%2C+population+control%2C+and+competition+%E2%80%93+a+critique&rft.volume=100&rft.issue=912&rft.pages=%3Cspan+class%3D%22nowrap%22%3E219-%3C%2Fspan%3E226&rft.date=1966&rft_id=info%3Adoi%2F10.1086%2F282415&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A84354616%23id-name%3DS2CID&rft.aulast=Murdoch&rft.aufirst=WM&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-44"><span class="mw-cite-backlink"><b><a href="#cite_ref-44">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPolisStrong1996" class="citation journal cs1">Polis, GA; Strong, DR (1996). "Food web complexity and community dynamics". <i>American Naturalist</i>. <b>147</b> (5): <span class="nowrap">813–</span>846. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1086%2F285880">10.1086/285880</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:85155900">85155900</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Naturalist&rft.atitle=Food+web+complexity+and+community+dynamics&rft.volume=147&rft.issue=5&rft.pages=%3Cspan+class%3D%22nowrap%22%3E813-%3C%2Fspan%3E846&rft.date=1996&rft_id=info%3Adoi%2F10.1086%2F285880&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A85155900%23id-name%3DS2CID&rft.aulast=Polis&rft.aufirst=GA&rft.au=Strong%2C+DR&rfr_id=info%3Asid%2Fen.wikipedia.org%3ATrophic+cascade" class="Z3988"></span></span> </li> <li id="cite_note-45"><span class="mw-cite-backlink"><b><a href="#cite_ref-45">^</a></b></span> <span class="reference-text">Eisenberg, Cristina (2011) <a rel="nofollow" class="external text" href="https://books.google.com/books?id=LCyZI6Vbp0oC&q=The+Wolf%27s+Tooth%3A+Keystone+Predators%2C+Trophic+Cascades%2C+and+Biodiversity%22">"The Wolf's Tooth: Keystone Predators, Trophic Cascades, and Biodiversity</a> pp. 15. Island Press. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1-59726-398-6" title="Special:BookSources/978-1-59726-398-6">978-1-59726-398-6</a>.</span> </li> <li id="cite_note-46"><span class="mw-cite-backlink"><b><a href="#cite_ref-46">^</a></b></span> <span class="reference-text">Barbosa P and Castellanos I (Eds) (2005) <a rel="nofollow" class="external text" href="https://books.google.com/books?id=251KKrnZNEEC&q=Ecology+of+Predator-Prey+Interactions"><i>Ecology of Predator-Prey Interactions</i></a> pp. 306, Oxford University Press. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780199883677" title="Special:BookSources/9780199883677">9780199883677</a>.</span> </li> 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aria-labelledby="Ecology:_Modelling_ecosystems:_Trophic_components279" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar 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href="/wiki/Special:EditPage/Template:Modelling_ecosystems" title="Special:EditPage/Template:Modelling ecosystems"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Ecology:_Modelling_ecosystems:_Trophic_components279" style="font-size:114%;margin:0 4em"><a href="/wiki/Ecology" title="Ecology">Ecology</a>: <a href="/wiki/Ecosystem_model" title="Ecosystem model">Modelling ecosystems</a>: <a href="/wiki/Trophic_level" title="Trophic level">Trophic</a> components</div></th></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abiotic_component" title="Abiotic component">Abiotic component</a></li> <li><a href="/wiki/Abiotic_stress" title="Abiotic stress">Abiotic stress</a></li> <li><a href="/wiki/Behavioral_ecology" title="Behavioral ecology">Behaviour</a></li> <li><a href="/wiki/Biogeochemical_cycle" title="Biogeochemical cycle">Biogeochemical cycle</a></li> <li><a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">Biomass</a></li> <li><a href="/wiki/Biotic_component" class="mw-redirect" title="Biotic component">Biotic component</a></li> <li><a href="/wiki/Biotic_stress" title="Biotic stress">Biotic stress</a></li> <li><a href="/wiki/Carrying_capacity" title="Carrying capacity">Carrying capacity</a></li> <li><a href="/wiki/Competition_(biology)" title="Competition (biology)">Competition</a></li> <li><a href="/wiki/Ecosystem" title="Ecosystem">Ecosystem</a></li> <li><a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem ecology</a></li> <li><a href="/wiki/Ecosystem_model" title="Ecosystem model">Ecosystem model</a></li> <li><a href="/wiki/Green_world_hypothesis" title="Green world hypothesis">Green world hypothesis</a></li> <li><a href="/wiki/Keystone_species" title="Keystone species">Keystone species</a></li> <li><a href="/wiki/List_of_feeding_behaviours" title="List of feeding behaviours">List of feeding behaviours</a></li> <li><a href="/wiki/Metabolic_theory_of_ecology" title="Metabolic theory of ecology">Metabolic theory of ecology</a></li> <li><a href="/wiki/Productivity_(ecology)" title="Productivity (ecology)">Productivity</a></li> <li><a href="/wiki/Resource_(biology)" title="Resource (biology)">Resource</a></li> <li><a href="/wiki/Restoration_ecology" class="mw-redirect" title="Restoration ecology">Restoration</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Autotroph" title="Autotroph">Producers</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Autotroph" title="Autotroph">Autotrophs</a></li> <li><a href="/wiki/Chemosynthesis" title="Chemosynthesis">Chemosynthesis</a></li> <li><a href="/wiki/Chemotroph" title="Chemotroph">Chemotrophs</a></li> <li><a href="/wiki/Foundation_species" title="Foundation species">Foundation species</a></li> <li><a href="/wiki/Kinetotroph" title="Kinetotroph">Kinetotrophs</a></li> <li><a href="/wiki/Mixotroph" title="Mixotroph">Mixotrophs</a></li> <li><a href="/wiki/Myco-heterotrophy" title="Myco-heterotrophy">Myco-heterotrophy</a></li> <li><a href="/wiki/Mycotroph" title="Mycotroph">Mycotroph</a></li> <li><a href="/wiki/Organotroph" title="Organotroph">Organotrophs</a></li> <li><a href="/wiki/Photoheterotroph" title="Photoheterotroph">Photoheterotrophs</a></li> <li><a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a></li> <li><a href="/wiki/Photosynthetic_efficiency" title="Photosynthetic efficiency">Photosynthetic efficiency</a></li> <li><a href="/wiki/Phototroph" title="Phototroph">Phototrophs</a></li> <li><a href="/wiki/Primary_nutritional_groups" title="Primary nutritional groups">Primary nutritional groups</a></li> <li><a href="/wiki/Primary_production" title="Primary production">Primary production</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Consumer_(food_chain)" title="Consumer (food chain)">Consumers</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Apex_predator" title="Apex predator">Apex predator</a></li> <li><a href="/wiki/Bacterivore" title="Bacterivore">Bacterivore</a></li> <li><a href="/wiki/Carnivore" title="Carnivore">Carnivores</a></li> <li><a href="/wiki/Chemoorganotroph" class="mw-redirect" title="Chemoorganotroph">Chemoorganotroph</a></li> <li><a href="/wiki/Foraging" title="Foraging">Foraging</a></li> <li><a href="/wiki/Generalist_and_specialist_species" title="Generalist and specialist species">Generalist and specialist species</a></li> <li><a href="/wiki/Intraguild_predation" title="Intraguild predation">Intraguild predation</a></li> <li><a href="/wiki/Herbivore" title="Herbivore">Herbivores</a></li> <li><a href="/wiki/Heterotroph" title="Heterotroph">Heterotroph</a></li> <li><a href="/wiki/Heterotrophic_nutrition" title="Heterotrophic nutrition">Heterotrophic nutrition</a></li> <li><a href="/wiki/Insectivore" title="Insectivore">Insectivore</a></li> <li><a href="/wiki/Mesopredator" title="Mesopredator">Mesopredators</a></li> <li><a href="/wiki/Mesopredator_release_hypothesis" title="Mesopredator release hypothesis">Mesopredator release hypothesis</a></li> <li><a href="/wiki/Omnivore" title="Omnivore">Omnivores</a></li> <li><a href="/wiki/Optimal_foraging_theory" title="Optimal foraging theory">Optimal foraging theory</a></li> <li><a href="/wiki/Planktivore" title="Planktivore">Planktivore</a></li> <li><a href="/wiki/Predation" title="Predation">Predation</a></li> <li><a href="/wiki/Prey_switching" title="Prey switching">Prey switching</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Decomposer" title="Decomposer">Decomposers</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chemoorganoheterotrophy" class="mw-redirect" title="Chemoorganoheterotrophy">Chemoorganoheterotrophy</a></li> <li><a href="/wiki/Decomposition" title="Decomposition">Decomposition</a></li> <li><a href="/wiki/Detritivore" title="Detritivore">Detritivores</a></li> <li><a href="/wiki/Detritus" title="Detritus">Detritus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Microorganism#Habitats_and_ecology" title="Microorganism">Microorganisms</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Bacteriophage" title="Bacteriophage">Bacteriophage</a></li> <li><a href="/wiki/Lithoautotroph" title="Lithoautotroph">Lithoautotroph</a></li> <li><a href="/wiki/Lithotroph" title="Lithotroph">Lithotrophy</a></li> <li><a href="/wiki/Marine_microorganisms" title="Marine microorganisms">Marine</a></li> <li><a href="/wiki/Microbial_cooperation" title="Microbial cooperation">Microbial cooperation</a></li> <li><a href="/wiki/Microbial_ecology" title="Microbial ecology">Microbial ecology</a></li> <li><a href="/wiki/Microbial_food_web" title="Microbial food web">Microbial food web</a></li> <li><a href="/wiki/Microbial_intelligence" title="Microbial intelligence">Microbial intelligence</a></li> <li><a href="/wiki/Microbial_loop" title="Microbial loop">Microbial loop</a></li> <li><a href="/wiki/Microbial_mat" title="Microbial mat">Microbial mat</a></li> <li><a href="/wiki/Microbial_metabolism" title="Microbial metabolism">Microbial metabolism</a></li> <li><a href="/wiki/Phage_ecology" title="Phage ecology">Phage ecology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Food_web" title="Food web">Food webs</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biomagnification" title="Biomagnification">Biomagnification</a></li> <li><a href="/wiki/Ecological_efficiency" title="Ecological efficiency">Ecological efficiency</a></li> <li><a href="/wiki/Ecological_pyramid" title="Ecological pyramid">Ecological pyramid</a></li> <li><a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">Energy flow</a></li> <li><a href="/wiki/Food_chain" title="Food chain">Food chain</a></li> <li><a href="/wiki/Trophic_level" title="Trophic level">Trophic level</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Example webs</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lake_ecosystem#Trophic_relationships" title="Lake ecosystem">Lakes</a></li> <li><a href="/wiki/River_ecosystem#Trophic_relationships" title="River ecosystem">Rivers</a></li> <li><a href="/wiki/Soil_food_web" title="Soil food web">Soil</a></li> <li><a href="/wiki/Tritrophic_interactions_in_plant_defense" title="Tritrophic interactions in plant defense">Tritrophic interactions in plant defense</a></li> <li><a href="/wiki/Marine_food_web" title="Marine food web">Marine food webs</a> <ul><li><a href="/wiki/Cold_seep" title="Cold seep">cold seeps</a></li> <li><a href="/wiki/Hydrothermal_vent#Biological_communities" title="Hydrothermal vent">hydrothermal vents</a></li> <li><a href="/wiki/Intertidal_ecology" title="Intertidal ecology">intertidal</a></li> <li><a href="/wiki/Kelp_forest#Trophic_ecology" title="Kelp forest">kelp forests</a></li> <li><a href="/wiki/North_Pacific_Gyre" title="North Pacific Gyre">North Pacific Gyre</a></li> <li><a href="/wiki/Ecology_of_the_San_Francisco_Estuary#Food_web" title="Ecology of the San Francisco Estuary">San Francisco Estuary</a></li> <li><a href="/wiki/Tide_pool" title="Tide pool">tide pool</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Processes</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ascendency" title="Ascendency">Ascendency</a></li> <li><a href="/wiki/Bioaccumulation" title="Bioaccumulation">Bioaccumulation</a></li> <li><a href="/wiki/Cascade_effect_(ecology)" title="Cascade effect (ecology)">Cascade effect</a></li> <li><a href="/wiki/Climax_community" title="Climax community">Climax community</a></li> <li><a href="/wiki/Competitive_exclusion_principle" title="Competitive exclusion principle">Competitive exclusion principle</a></li> <li><a href="/wiki/Consumer%E2%80%93resource_interactions" title="Consumer–resource interactions">Consumer–resource interactions</a></li> <li><a href="/wiki/Copiotroph" title="Copiotroph">Copiotrophs</a></li> <li><a href="/wiki/Dominance_(ecology)" title="Dominance (ecology)">Dominance</a></li> <li><a href="/wiki/Ecological_network" title="Ecological network">Ecological network</a></li> <li><a href="/wiki/Ecological_succession" title="Ecological succession">Ecological succession</a></li> <li><a href="/wiki/Energy_quality" title="Energy quality">Energy quality</a></li> <li><a href="/wiki/Energy_systems_language" title="Energy systems language">Energy systems language</a></li> <li><a href="/wiki/F-ratio_(oceanography)" title="F-ratio (oceanography)">f-ratio</a></li> <li><a href="/wiki/Feed_conversion_ratio" title="Feed conversion ratio">Feed conversion ratio</a></li> <li><a href="/wiki/Feeding_frenzy" title="Feeding frenzy">Feeding frenzy</a></li> <li><a href="/wiki/Mesotrophic_soil" title="Mesotrophic soil">Mesotrophic soil</a></li> <li><a href="/wiki/Nutrient_cycle" title="Nutrient cycle">Nutrient cycle</a></li> <li><a href="/wiki/Oligotroph" title="Oligotroph">Oligotroph</a></li> <li><a href="/wiki/Paradox_of_the_plankton" title="Paradox of the plankton">Paradox of the plankton</a></li> <li><a class="mw-selflink selflink">Trophic cascade</a></li> <li><a href="/wiki/Trophic_mutualism" title="Trophic mutualism">Trophic mutualism</a></li> <li><a href="/wiki/Trophic_state_index" title="Trophic state index">Trophic state index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Defense,<br />counter</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Animal_coloration" title="Animal coloration">Animal coloration</a></li> <li><a href="/wiki/Anti-predator_adaptation" title="Anti-predator adaptation">Anti-predator adaptations</a></li> <li><a href="/wiki/Camouflage" title="Camouflage">Camouflage</a></li> <li><a href="/wiki/Deimatic_behaviour" title="Deimatic behaviour">Deimatic behaviour</a></li> <li><a href="/wiki/Herbivore_adaptations_to_plant_defense" title="Herbivore adaptations to plant defense">Herbivore adaptations to plant defense</a></li> <li><a href="/wiki/Mimicry" title="Mimicry">Mimicry</a></li> <li><a href="/wiki/Plant_defense_against_herbivory" title="Plant defense against herbivory">Plant defense against herbivory</a></li> <li><a href="/wiki/Shoaling_and_schooling#Predator_avoidance" title="Shoaling and schooling">Predator avoidance in schooling fish</a></li></ul> </div></td></tr></tbody></table></div><div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Ecology:_Modelling_ecosystems:_Other_components285" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Modelling_ecosystems" title="Template:Modelling ecosystems"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Modelling_ecosystems" title="Template talk:Modelling ecosystems"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Modelling_ecosystems" title="Special:EditPage/Template:Modelling ecosystems"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Ecology:_Modelling_ecosystems:_Other_components285" style="font-size:114%;margin:0 4em"><a href="/wiki/Ecology" title="Ecology">Ecology</a>: <a href="/wiki/Ecosystem_model" title="Ecosystem model">Modelling ecosystems</a>: Other components</div></th></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Population_ecology" title="Population ecology">Population<br />ecology</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abundance_(ecology)" title="Abundance (ecology)">Abundance</a></li> <li><a href="/wiki/Allee_effect" title="Allee effect">Allee effect</a></li> <li><a href="/wiki/Consumer-resource_model" title="Consumer-resource model">Consumer-resource model</a></li> <li><a href="/wiki/Depensation" title="Depensation">Depensation</a></li> <li><a href="/wiki/Ecological_yield" title="Ecological yield">Ecological yield</a></li> <li><a href="/wiki/Effective_population_size" title="Effective population size">Effective population size</a></li> <li><a href="/wiki/Intraspecific_competition" title="Intraspecific competition">Intraspecific competition</a></li> <li><a href="/wiki/Logistic_function" title="Logistic function">Logistic function</a></li> <li><a href="/wiki/Malthusian_growth_model" title="Malthusian growth model">Malthusian growth model</a></li> <li><a href="/wiki/Maximum_sustainable_yield" title="Maximum sustainable yield">Maximum sustainable yield</a></li> <li><a href="/wiki/Overpopulation" title="Overpopulation">Overpopulation</a></li> <li><a href="/wiki/Overexploitation" title="Overexploitation">Overexploitation</a></li> <li><a href="/wiki/Population_cycle" title="Population cycle">Population cycle</a></li> <li><a href="/wiki/Population_dynamics" title="Population dynamics">Population dynamics</a></li> <li><a href="/wiki/Population_model" title="Population model">Population modeling</a></li> <li><a href="/wiki/Population_size" title="Population size">Population size</a></li> <li><a href="/wiki/Lotka%E2%80%93Volterra_equations" title="Lotka–Volterra equations">Predator–prey (Lotka–Volterra) equations</a></li> <li><a href="/wiki/Recruitment_(biology)" title="Recruitment (biology)">Recruitment</a></li> <li><a href="/wiki/Small_population_size" title="Small population size">Small population size</a></li> <li><a href="/wiki/Ecological_stability" title="Ecological stability">Stability</a> <ul><li><a href="/wiki/Ecological_resilience" title="Ecological resilience">Resilience</a></li> <li><a href="/wiki/Resistance_(ecology)" title="Resistance (ecology)">Resistance</a></li></ul></li> <li><a href="/wiki/Random_generalized_Lotka%E2%80%93Volterra_model" title="Random generalized Lotka–Volterra model">Random generalized Lotka–Volterra model</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Species</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biodiversity" title="Biodiversity">Biodiversity</a></li> <li><a href="/wiki/Density_dependence" title="Density dependence">Density-dependent inhibition</a></li> <li><a href="/wiki/Ecological_effects_of_biodiversity" title="Ecological effects of biodiversity">Ecological effects of biodiversity</a></li> <li><a href="/wiki/Ecological_extinction" title="Ecological extinction">Ecological extinction</a></li> <li><a href="/wiki/Endemism" title="Endemism">Endemic species</a></li> <li><a href="/wiki/Flagship_species" title="Flagship species">Flagship species</a></li> <li><a href="/wiki/Ordination_(statistics)" title="Ordination (statistics)">Gradient analysis</a></li> <li><a href="/wiki/Bioindicator" title="Bioindicator">Indicator species</a></li> <li><a href="/wiki/Introduced_species" title="Introduced species">Introduced species</a></li> <li><a href="/wiki/Invasive_species" title="Invasive species">Invasive species</a> / <a href="/wiki/Native_species" title="Native species">Native species</a></li> <li><a href="/wiki/Latitudinal_gradients_in_species_diversity" title="Latitudinal gradients in species diversity">Latitudinal gradients in species diversity</a></li> <li><a href="/wiki/Minimum_viable_population" title="Minimum viable population">Minimum viable population</a></li> <li><a href="/wiki/Unified_neutral_theory_of_biodiversity" title="Unified neutral theory of biodiversity">Neutral theory</a></li> <li><a href="/wiki/Occupancy%E2%80%93abundance_relationship" title="Occupancy–abundance relationship">Occupancy–abundance relationship</a></li> <li><a href="/wiki/Population_viability_analysis" title="Population viability analysis">Population viability analysis</a></li> <li><a href="/wiki/Priority_effect" title="Priority effect">Priority effect</a></li> <li><a href="/wiki/Rapoport%27s_rule" title="Rapoport's rule">Rapoport's rule</a></li> <li><a href="/wiki/Relative_abundance_distribution" title="Relative abundance distribution">Relative abundance distribution</a></li> <li><a href="/wiki/Relative_species_abundance" title="Relative species abundance">Relative species abundance</a></li> <li><a href="/wiki/Species_diversity" title="Species diversity">Species diversity</a></li> <li><a href="/wiki/Species_homogeneity" title="Species homogeneity">Species homogeneity</a></li> <li><a href="/wiki/Species_richness" title="Species richness">Species richness</a></li> <li><a href="/wiki/Species_distribution" title="Species distribution">Species distribution</a></li> <li><a href="/wiki/Species%E2%80%93area_relationship" title="Species–area relationship">Species–area curve</a></li> <li><a href="/wiki/Umbrella_species" title="Umbrella species">Umbrella species</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Species<br />interaction</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Antibiosis" title="Antibiosis">Antibiosis</a></li> <li><a href="/wiki/Biological_interaction" title="Biological interaction">Biological interaction</a></li> <li><a href="/wiki/Commensalism" title="Commensalism">Commensalism</a></li> <li><a href="/wiki/Community_(ecology)" title="Community (ecology)">Community ecology</a></li> <li><a href="/wiki/Ecological_facilitation" title="Ecological facilitation">Ecological facilitation</a></li> <li><a href="/wiki/Interspecific_competition" title="Interspecific competition">Interspecific competition</a></li> <li><a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">Mutualism</a></li> <li><a href="/wiki/Parasitism" title="Parasitism">Parasitism</a></li> <li><a href="/wiki/Storage_effect" title="Storage effect">Storage effect</a></li> <li><a href="/wiki/Symbiosis" title="Symbiosis">Symbiosis</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Spatial_ecology" title="Spatial ecology">Spatial<br />ecology</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Cross-boundary_subsidy" title="Cross-boundary subsidy">Cross-boundary subsidy</a></li> <li><a href="/wiki/Cline_(biology)" title="Cline (biology)">Ecocline</a></li> <li><a href="/wiki/Ecotone" title="Ecotone">Ecotone</a></li> <li><a href="/wiki/Ecotype" title="Ecotype">Ecotype</a></li> <li><a href="/wiki/Disturbance_(ecology)" title="Disturbance (ecology)">Disturbance</a></li> <li><a href="/wiki/Edge_effects" title="Edge effects">Edge effects</a></li> <li><a href="/wiki/Foster%27s_rule" title="Foster's rule">Foster's rule</a></li> <li><a href="/wiki/Habitat_fragmentation" title="Habitat fragmentation">Habitat fragmentation</a></li> <li><a href="/wiki/Ideal_free_distribution" title="Ideal free distribution">Ideal free distribution</a></li> <li><a href="/wiki/Intermediate_disturbance_hypothesis" title="Intermediate disturbance hypothesis">Intermediate disturbance hypothesis</a></li> <li><a href="/wiki/Insular_biogeography" title="Insular biogeography">Insular biogeography</a></li> <li><a href="/wiki/Land_change_modeling" title="Land change modeling">Land change modeling</a></li> <li><a href="/wiki/Landscape_ecology" title="Landscape ecology">Landscape ecology</a></li> <li><a href="/wiki/Landscape_epidemiology" title="Landscape epidemiology">Landscape epidemiology</a></li> <li><a href="/wiki/Landscape_limnology" title="Landscape limnology">Landscape limnology</a></li> <li><a href="/wiki/Metapopulation" title="Metapopulation">Metapopulation</a></li> <li><a href="/wiki/Patch_dynamics" title="Patch dynamics">Patch dynamics</a></li> <li><a href="/wiki/R/K_selection_theory" title="R/K selection theory"><i>r</i>/<i>K</i> selection theory</a></li> <li><a href="/wiki/Resource_selection_function" title="Resource selection function">Resource selection function</a></li> <li><a href="/wiki/Source%E2%80%93sink_dynamics" title="Source–sink dynamics">Source–sink dynamics</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Ecological_niche" title="Ecological niche">Niche</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ecological_trap" title="Ecological trap">Ecological trap</a></li> <li><a href="/wiki/Ecosystem_engineer" title="Ecosystem engineer">Ecosystem engineer</a></li> <li><a href="/wiki/Species_distribution_modelling" title="Species distribution modelling">Environmental niche modelling</a></li> <li><a href="/wiki/Guild_(ecology)" title="Guild (ecology)">Guild</a></li> <li><a href="/wiki/Habitat" title="Habitat">Habitat</a> <ul><li><a href="/wiki/Marine_habitat" title="Marine habitat">Marine</a></li> <li><a href="/wiki/Semiaquatic" title="Semiaquatic">Semiaquatic</a></li> <li><a href="/wiki/Terrestrial_animal" title="Terrestrial animal">Terrestrial</a></li></ul></li> <li><a href="/wiki/Limiting_similarity" title="Limiting similarity">Limiting similarity</a></li> <li><a href="/wiki/Niche_apportionment_models" title="Niche apportionment models">Niche apportionment models</a></li> <li><a href="/wiki/Niche_construction" title="Niche construction">Niche construction</a></li> <li><a href="/wiki/Niche_differentiation" class="mw-redirect" title="Niche differentiation">Niche differentiation</a></li> <li><a href="/wiki/Ontogenetic_niche_shift" title="Ontogenetic niche shift">Ontogenetic niche shift</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em"><a href="/wiki/Non-trophic_networks" title="Non-trophic networks">Other<br />networks</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Assembly_rules" title="Assembly rules">Assembly rules</a></li> <li><a href="/wiki/Bateman%27s_principle" title="Bateman's principle">Bateman's principle</a></li> <li><a href="/wiki/Bioluminescence" title="Bioluminescence">Bioluminescence</a></li> <li><a href="/wiki/Ecological_collapse" class="mw-redirect" title="Ecological collapse">Ecological collapse</a></li> <li><a href="/wiki/Ecological_debt" title="Ecological debt">Ecological debt</a></li> <li><a href="/wiki/Ecological_debt" title="Ecological debt">Ecological deficit</a></li> <li><a href="/wiki/Energy_flow_(ecology)" title="Energy flow (ecology)">Ecological energetics</a></li> <li><a href="/wiki/Ecological_indicator" title="Ecological indicator">Ecological indicator</a></li> <li><a href="/wiki/Ecological_threshold" title="Ecological threshold">Ecological threshold</a></li> <li><a href="/wiki/Ecosystem_diversity" title="Ecosystem diversity">Ecosystem diversity</a></li> <li><a href="/wiki/Emergence" title="Emergence">Emergence</a></li> <li><a href="/wiki/Extinction_debt" title="Extinction debt">Extinction debt</a></li> <li><a href="/wiki/Kleiber%27s_law" title="Kleiber's law">Kleiber's law</a></li> <li><a href="/wiki/Liebig%27s_law_of_the_minimum" title="Liebig's law of the minimum">Liebig's law of the minimum</a></li> <li><a href="/wiki/Marginal_value_theorem" title="Marginal value theorem">Marginal value theorem</a></li> <li><a href="/wiki/Thorson%27s_rule" title="Thorson's rule">Thorson's rule</a></li> <li><a href="/wiki/Xerosere" title="Xerosere">Xerosere</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7.5em">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Allometry" title="Allometry">Allometry</a></li> <li><a href="/wiki/Alternative_stable_state" title="Alternative stable state">Alternative stable state</a></li> <li><a href="/wiki/Balance_of_nature" title="Balance of nature">Balance of nature</a></li> <li><a href="/wiki/Biological_data_visualization" title="Biological data visualization">Biological data visualization</a></li> <li><a href="/wiki/Ecological_economics" title="Ecological economics">Ecological economics</a></li> <li><a href="/wiki/Ecological_footprint" title="Ecological footprint">Ecological footprint</a></li> <li><a href="/wiki/Ecological_forecasting" title="Ecological forecasting">Ecological forecasting</a></li> <li><a href="/wiki/Environmental_humanities" title="Environmental humanities">Ecological humanities</a></li> <li><a href="/wiki/Ecological_stoichiometry" title="Ecological stoichiometry">Ecological stoichiometry</a></li> <li><a href="/wiki/Ecopath" title="Ecopath">Ecopath</a></li> <li><a href="/wiki/Ecosystem_based_fisheries" class="mw-redirect" title="Ecosystem based fisheries">Ecosystem based fisheries</a></li> <li><a href="/wiki/Endolith" title="Endolith">Endolith</a></li> <li><a href="/wiki/Evolutionary_ecology" title="Evolutionary ecology">Evolutionary ecology</a></li> <li><a href="/wiki/Functional_ecology" title="Functional ecology">Functional ecology</a></li> <li><a href="/wiki/Industrial_ecology" title="Industrial ecology">Industrial ecology</a></li> <li><a href="/wiki/Macroecology" title="Macroecology">Macroecology</a></li> <li><a href="/wiki/Microecosystem" title="Microecosystem">Microecosystem</a></li> <li><a href="/wiki/Natural_environment" title="Natural environment">Natural environment</a></li> <li><a href="/wiki/Regime_shift" title="Regime shift">Regime shift</a></li> <li><a href="/wiki/Sexecology" title="Sexecology">Sexecology</a></li> <li><a href="/wiki/Systems_ecology" title="Systems ecology">Systems ecology</a></li> <li><a href="/wiki/Urban_ecology" title="Urban ecology">Urban ecology</a></li> <li><a href="/wiki/Theoretical_ecology" title="Theoretical ecology">Theoretical ecology</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/Outline_of_ecology" title="Outline of ecology">Outline of ecology</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Aquatic_ecosystems487" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Aquatic_ecosystems" title="Template:Aquatic ecosystems"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Aquatic_ecosystems" title="Template talk:Aquatic ecosystems"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Aquatic_ecosystems" title="Special:EditPage/Template:Aquatic ecosystems"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Aquatic_ecosystems487" style="font-size:114%;margin:0 4em"><a href="/wiki/Aquatic_ecosystem" title="Aquatic ecosystem">Aquatic ecosystems</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="General_components_and_freshwater_ecosystems487" style="font-size:114%;margin:0 4em">General components and <a href="/wiki/Freshwater_ecosystem" title="Freshwater ecosystem">freshwater ecosystems</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acoustic_ecology" title="Acoustic ecology">Acoustic ecology</a></li> <li><a href="/wiki/Algal_bloom" title="Algal bloom">Algal bloom</a></li> <li><a href="/wiki/Anoxic_waters" title="Anoxic waters">Anoxic waters</a></li> <li><a href="/wiki/Aquatic_adaptation" class="mw-redirect" title="Aquatic adaptation">Aquatic adaptation</a></li> <li><a href="/wiki/Aquatic_animal" title="Aquatic animal">Aquatic animal</a> <ul><li><a href="/wiki/Aquatic_insect" title="Aquatic insect">Insect</a></li> <li><a href="/wiki/Aquatic_mammal" title="Aquatic mammal">Mammal</a></li> <li><a href="/wiki/Water_bird" title="Water bird">Water bird</a></li></ul></li> <li><a href="/wiki/Aquatic_biomonitoring" title="Aquatic biomonitoring">Aquatic biomonitoring</a></li> <li><a href="/wiki/Aquatic_plant" title="Aquatic plant">Aquatic plant</a></li> <li><a href="/wiki/Agent-based_model_in_biology#Aquatic_population_dynamics" title="Agent-based model in biology">Aquatic population dynamics</a></li> <li><a href="/wiki/Aquatic_predation" class="mw-redirect" title="Aquatic predation">Aquatic predation</a></li> <li><a href="/wiki/Aquatic_respiration" title="Aquatic respiration">Aquatic respiration</a></li> <li><a href="/wiki/Aquatic_science" title="Aquatic science">Aquatic science</a></li> <li><a href="/wiki/Aquatic_toxicology" title="Aquatic toxicology">Aquatic toxicology</a></li> <li><a href="/wiki/Benthos" title="Benthos">Benthos</a></li> <li><a href="/wiki/Bioluminescence" title="Bioluminescence">Bioluminescence</a></li> <li><a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">Biomass</a></li> <li><a href="/wiki/Cascade_effect_(ecology)" title="Cascade effect (ecology)">Cascade effect</a></li> <li><a href="/wiki/Colored_dissolved_organic_matter" title="Colored dissolved organic matter">Colored dissolved organic matter</a></li> <li><a href="/wiki/Dead_zone_(ecology)" title="Dead zone (ecology)">Dead zone</a></li> <li><a href="/wiki/Ecohydrology" title="Ecohydrology">Ecohydrology</a></li> <li><a href="/wiki/Eutrophication" title="Eutrophication">Eutrophication</a></li> <li><a href="/wiki/Fisheries_science" title="Fisheries science">Fisheries science</a></li> <li><a href="/wiki/Food_chain" title="Food chain">Food chain</a></li> <li><a href="/wiki/Food_web" title="Food web">Food web</a></li> <li><a href="/wiki/GIS_and_aquatic_science" title="GIS and aquatic science">GIS and aquatic science</a></li> <li><a href="/wiki/Hydrobiology" title="Hydrobiology">Hydrobiology</a></li> <li><a href="/wiki/Hypoxia_(environmental)" title="Hypoxia (environmental)">Hypoxia</a></li> <li><a href="/wiki/Macrobenthos" title="Macrobenthos">Macrobenthos</a></li> <li><a href="/wiki/Meiobenthos" title="Meiobenthos">Meiobenthos</a></li> <li><a href="/wiki/Microbial_ecology" title="Microbial ecology">Microbial ecology</a></li> <li><a href="/wiki/Microbial_food_web" title="Microbial food web">Microbial food web</a></li> <li><a href="/wiki/Microbial_loop" title="Microbial loop">Microbial loop</a></li> <li><a href="/wiki/Nekton" title="Nekton">Nekton</a></li> <li><a href="/wiki/Neuston" title="Neuston">Neuston</a></li> <li><a href="/wiki/Particle_(ecology)" title="Particle (ecology)">Particle</a></li> <li><a href="/wiki/Pelagic_zone" title="Pelagic zone">Pelagic zone</a></li> <li><a href="/wiki/Photic_zone" title="Photic zone">Photic zone</a></li> <li><a href="/wiki/Phytoplankton" title="Phytoplankton">Phytoplankton</a></li> <li><a href="/wiki/Plankton" title="Plankton">Plankton</a></li> <li><a href="/wiki/Pleuston" class="mw-redirect" title="Pleuston">Pleuston</a></li> <li><a href="/wiki/Productivity_(ecology)" title="Productivity (ecology)">Productivity</a></li> <li><a href="/wiki/Ramsar_Convention" title="Ramsar Convention">Ramsar Convention</a></li> <li><a href="/wiki/Sediment_trap" title="Sediment trap">Sediment trap</a></li> <li><a href="/wiki/Semiaquatic" title="Semiaquatic">Semiaquatic</a></li> <li><a href="/wiki/Shoaling_and_schooling" title="Shoaling and schooling">Shoaling and schooling</a></li> <li><a href="/wiki/Siltation" title="Siltation">Siltation</a></li> <li><a href="/wiki/Spawn_(biology)" class="mw-redirect" title="Spawn (biology)">Spawn</a></li> <li><a href="/wiki/Isotope_analysis#Stable_isotope_analysis_in_aquatic_ecosystems" title="Isotope analysis">Stable isotope analysis in aquatic ecosystems</a></li> <li><a href="/wiki/Stream_metabolism" title="Stream metabolism">Stream metabolism</a></li> <li><a href="/wiki/Substrate_(marine_biology)" class="mw-redirect" title="Substrate (marine biology)">Substrate</a></li> <li><a href="/wiki/Thermal_pollution" title="Thermal pollution">Thermal pollution</a></li> <li><a href="/wiki/Trophic_level" title="Trophic level">Trophic level</a></li> <li><a href="/wiki/Underwater_camouflage" title="Underwater camouflage">Underwater camouflage</a></li> <li><a href="/wiki/Water_column" title="Water column">Water column</a></li> <li><a href="/wiki/Zooplankton" title="Zooplankton">Zooplankton</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="3" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg/140px-Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg" decoding="async" width="140" height="202" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg/210px-Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg/280px-Aerial_view_of_Slapton_Sands_and_northern_part_of_Ley_-_geograph.org.uk_-_1434412.jpg 2x" data-file-width="382" data-file-height="551" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Freshwater</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Freshwater_biology" title="Freshwater biology">Freshwater biology</a></li> <li><a href="/wiki/Freshwater_biome" class="mw-redirect" title="Freshwater biome">Freshwater biome</a></li> <li><a href="/wiki/Freshwater_environmental_quality_parameters" title="Freshwater environmental quality parameters">Freshwater environmental quality parameters</a></li> <li><a href="/wiki/Freshwater_fish" title="Freshwater fish">Freshwater fish</a></li> <li><a href="/wiki/Hyporheic_zone" title="Hyporheic zone">Hyporheic zone</a></li> <li><a href="/wiki/Limnology" title="Limnology">Limnology</a></li> <li><a href="/wiki/Lake_ecosystem" title="Lake ecosystem">Lake ecosystem</a></li> <li><a href="/wiki/Lake_stratification" title="Lake stratification">Lake stratification</a></li> <li><a href="/wiki/Macrophyte" class="mw-redirect" title="Macrophyte">Macrophyte</a></li> <li><a href="/wiki/Pond" title="Pond">Pond</a> <ul><li><a href="/wiki/Fish_pond" title="Fish pond">Fish pond</a></li></ul></li> <li><a href="/wiki/Rheotaxis" title="Rheotaxis">Rheotaxis</a></li> <li><a href="/wiki/River_ecosystem" title="River ecosystem">River ecosystem</a></li> <li><a href="/wiki/Stream_bed" title="Stream bed">Stream bed</a></li> <li><a href="/wiki/Stream_pool" title="Stream pool">Stream pool</a></li> <li><a href="/wiki/Trophic_state_index" title="Trophic state index">Trophic state index</a></li> <li><a href="/wiki/Upland_and_lowland" title="Upland and lowland">Upland and lowland</a></li> <li><a href="/wiki/Water_garden" title="Water garden">Water garden</a></li> <li><a href="/wiki/Wetland" title="Wetland">Wetland</a> <ul><li><a href="/wiki/Bog" title="Bog">Bog</a></li> <li><a href="/wiki/Brackish_marsh" title="Brackish marsh">Brackish marsh</a></li> <li><a href="/wiki/Fen" title="Fen">Fen</a></li> <li><a href="/wiki/Freshwater_marsh" title="Freshwater marsh">Freshwater marsh</a></li> <li><a href="/wiki/Freshwater_swamp_forest" title="Freshwater swamp forest">Freshwater swamp forest</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Ecoregions</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_freshwater_ecoregions_(WWF)" title="List of freshwater ecoregions (WWF)">List of freshwater ecoregions (WWF)</a> <ul><li><a href="/wiki/List_of_freshwater_ecoregions_in_Africa_and_Madagascar" title="List of freshwater ecoregions in Africa and Madagascar">Africa and Madagascar</a></li> <li><a href="/wiki/List_of_freshwater_ecoregions_of_Latin_America_and_the_Caribbean" title="List of freshwater ecoregions of Latin America and the Caribbean">Latin America and the Caribbean</a></li></ul></li> <li><a href="/wiki/List_of_marine_ecoregions" title="List of marine ecoregions">List of marine ecoregions</a></li> <li>Specific examples <ul><li><a href="/wiki/Geography_and_ecology_of_the_Everglades" title="Geography and ecology of the Everglades">Everglades</a></li> <li><a href="/wiki/Freshwater_ecology_of_Maharashtra" title="Freshwater ecology of Maharashtra">Maharashtra</a></li> <li><a href="/wiki/Ecosystem_of_the_North_Pacific_Subtropical_Gyre" title="Ecosystem of the North Pacific Subtropical Gyre">North Pacific Subtropical Gyre</a></li> <li><a href="/wiki/Ecology_of_the_San_Francisco_Estuary" title="Ecology of the San Francisco Estuary">San Francisco Estuary</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Marine_ecosystems_(components)487" style="font-size:114%;margin:0 4em"><a href="/wiki/Marine_ecosystem" title="Marine ecosystem">Marine ecosystems</a> (<a href="/wiki/Template:Marine_aquatic_ecosystems" title="Template:Marine aquatic ecosystems">components</a>)</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Deep_scattering_layer" title="Deep scattering layer">Deep scattering layer</a></li> <li><a href="/wiki/Diel_vertical_migration" title="Diel vertical migration">Diel vertical migration</a></li> <li><a href="/wiki/F-ratio_(oceanography)" title="F-ratio (oceanography)">f-ratio</a></li> <li><a href="/wiki/Iron_fertilization" title="Iron fertilization">Iron fertilization</a></li> <li><a href="/wiki/Large_marine_ecosystem" class="mw-redirect" title="Large marine ecosystem">Large marine ecosystem</a></li> <li><a href="/wiki/Marine_biology" title="Marine biology">Marine biology</a></li> <li><a href="/wiki/Marine_chemistry" title="Marine chemistry">Marine chemistry</a></li> <li><a href="/wiki/Marine_food_web" title="Marine food web">Marine food web</a></li> <li><a href="/wiki/Marine_primary_production" title="Marine primary production">Marine primary production</a></li> <li><a href="/wiki/Marine_snow" title="Marine snow">Marine snow</a></li> <li><a href="/wiki/Ocean_fertilization" title="Ocean fertilization">Ocean fertilization</a></li> <li><a href="/wiki/Oceanic_physical-biological_process" title="Oceanic physical-biological process">Oceanic physical-biological process</a></li> <li><a href="/wiki/Ocean_turbidity" title="Ocean turbidity">Ocean turbidity</a></li> <li><a href="/wiki/Photophore" title="Photophore">Photophore</a></li> <li><a href="/wiki/Thorson%27s_rule" title="Thorson's rule">Thorson's rule</a></li> <li><a href="/wiki/Upwelling" title="Upwelling">Upwelling</a></li> <li><a href="/wiki/Viral_shunt" title="Viral shunt">Viral shunt</a></li> <li><a href="/wiki/Whale_fall" title="Whale fall">Whale fall</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="4" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Maldivesfish2.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/35/Maldivesfish2.jpg/140px-Maldivesfish2.jpg" decoding="async" width="140" height="95" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/35/Maldivesfish2.jpg/210px-Maldivesfish2.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/35/Maldivesfish2.jpg/280px-Maldivesfish2.jpg 2x" data-file-width="1818" data-file-height="1228" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_life" title="Marine life">Marine life</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Census_of_Marine_Life" title="Census of Marine Life">Census of Marine Life</a></li> <li><a href="/wiki/Deep-sea_community" title="Deep-sea community">Deep-sea community</a></li> <li><a href="/wiki/Deep-water_coral" title="Deep-water coral">Deep-water coral</a></li> <li><a href="/wiki/Marine_fungi" title="Marine fungi">Marine fungi</a></li> <li><a href="/wiki/Marine_invertebrates" title="Marine invertebrates">Marine invertebrates</a></li> <li><a href="/wiki/Marine_larval_ecology" title="Marine larval ecology">Marine larval ecology</a></li> <li><a href="/wiki/Seagrass" title="Seagrass">Seagrass</a></li> <li><a href="/wiki/Seashore_wildlife" title="Seashore wildlife">Seashore wildlife</a></li> <li><a href="/wiki/Wild_fisheries" title="Wild fisheries">Wild fisheries</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_microorganisms" title="Marine microorganisms">Microorganisms</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marine_bacteriophage" class="mw-redirect" title="Marine bacteriophage">Marine bacteriophage</a></li> <li><a href="/wiki/Marine_prokaryotes" title="Marine prokaryotes">Marine prokaryotes</a></li> <li><a href="/wiki/Marine_protists" title="Marine protists">Marine protists</a></li> <li><a href="/wiki/Marine_viruses" title="Marine viruses">Marine viruses</a></li> <li><a href="/wiki/Paradox_of_the_plankton" title="Paradox of the plankton">Paradox of the plankton</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_vertebrate" title="Marine vertebrate">Vertebrates</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marine_mammal" title="Marine mammal">Marine mammal</a></li> <li><a href="/wiki/Marine_reptile" title="Marine reptile">Marine reptile</a></li> <li><a href="/wiki/Saltwater_fish" title="Saltwater fish">Saltwater fish</a> <ul><li><a href="/wiki/Coastal_fish" title="Coastal fish">Coastal fish</a></li> <li><a href="/wiki/Coral_reef_fish" title="Coral reef fish">Coral reef fish</a></li> <li><a href="/wiki/Deep-sea_fish" title="Deep-sea fish">Deep-sea fish</a></li> <li><a href="/wiki/Demersal_fish" title="Demersal fish">Demersal fish</a></li> <li><a href="/wiki/Pelagic_fish" title="Pelagic fish">Pelagic fish</a></li></ul></li> <li><a href="/wiki/Seabird" title="Seabird">Seabird</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_habitat" title="Marine habitat">Marine habitats</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bay_mud" title="Bay mud">Bay mud</a></li> <li><a href="/wiki/Marine_coastal_ecosystem" title="Marine coastal ecosystem">Marine coastal ecosystem</a></li> <li><a href="/wiki/Coastal_biogeomorphology" title="Coastal biogeomorphology">Coastal biogeomorphology</a></li> <li><a href="/wiki/Cold_seep" title="Cold seep">Cold seep</a></li> <li><a href="/wiki/Coral_reef" title="Coral reef">Coral reef</a></li> <li><a href="/wiki/Davidson_Seamount#Ecology" title="Davidson Seamount">Davidson Seamount § Ecology</a></li> <li><a href="/wiki/Estuary" title="Estuary">Estuary</a></li> <li><a href="/wiki/Intertidal_ecology" title="Intertidal ecology">Intertidal ecology</a></li> <li><a href="/wiki/Intertidal_wetland" title="Intertidal wetland">Intertidal wetland</a></li> <li><a href="/wiki/Kelp_forest" title="Kelp forest">Kelp forest</a></li> <li><a href="/wiki/Hydrothermal_vent" title="Hydrothermal vent">Hydrothermal vent</a></li> <li><a href="/wiki/Lagoon" title="Lagoon">Lagoon</a></li> <li><a href="/wiki/Mangrove" title="Mangrove">Mangrove</a></li> <li><a href="/wiki/Marine_biomes" class="mw-redirect" title="Marine biomes">Marine biomes</a></li> <li><a href="/wiki/Mudflat" title="Mudflat">Mudflat</a></li> <li><a href="/wiki/Oyster_reef" title="Oyster reef">Oyster reef</a></li> <li><a href="/wiki/Rocky_shore" title="Rocky shore">Rocky shore</a></li> <li><a href="/wiki/Salt_marsh" title="Salt marsh">Salt marsh</a></li> <li><a href="/wiki/Salt_pannes_and_pools" title="Salt pannes and pools">Salt pannes and pools</a></li> <li><a href="/wiki/Seagrass_meadow" title="Seagrass meadow">Seagrass meadow</a></li> <li><a href="/wiki/Sponge_ground" title="Sponge ground">Sponge ground</a></li> <li><a href="/wiki/Sponge_reef" title="Sponge reef">Sponge reef</a></li> <li><a href="/wiki/Tide_pool" title="Tide pool">Tide pool</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marine_conservation" title="Marine conservation">Conservation</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Coral_bleaching" title="Coral bleaching">Coral bleaching</a></li> <li><a href="/wiki/Ecological_values_of_mangroves" title="Ecological values of mangroves">Ecological values of mangroves</a></li> <li><a href="/wiki/Fisheries_and_climate_change" class="mw-redirect" title="Fisheries and climate change">Fisheries and climate change</a></li> <li><a href="/wiki/Hotspot_Ecosystem_Research_and_Man%27s_Impact_On_European_Seas" title="Hotspot Ecosystem Research and Man's Impact On European Seas">HERMIONE</a></li> <li><a href="/wiki/Human_impact_on_marine_life" title="Human impact on marine life">Human impact on marine life</a></li> <li><a href="/wiki/Marine_conservation_activism" title="Marine conservation activism">Marine conservation activism</a></li> <li><a href="/wiki/Marine_pollution" title="Marine pollution">Marine pollution</a></li> <li><a href="/wiki/Marine_protected_area" title="Marine protected area">Marine protected area</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="nowrap"><span class="nowrap"><span class="noviewer" typeof="mw:File"><span><img alt="image" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png/16px-Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png/24px-Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png/32px-Walden_Pond%2C_Massachusetts_on_June_27%2C_2012.png 2x" data-file-width="1500" data-file-height="1500" /></span></span> </span><a href="/wiki/Portal:Lakes" title="Portal:Lakes">Lakes portal</a></span></li> <li><span class="nowrap"><span class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Waves_in_pacifica_1.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Waves_in_pacifica_1.jpg/16px-Waves_in_pacifica_1.jpg" decoding="async" width="16" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Waves_in_pacifica_1.jpg/24px-Waves_in_pacifica_1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/45/Waves_in_pacifica_1.jpg/32px-Waves_in_pacifica_1.jpg 2x" data-file-width="2000" data-file-height="1358" /></a></span> </span><a href="/wiki/Portal:Oceans" title="Portal:Oceans">Oceans portal</a></span></li> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Aquatic_ecology" title="Category:Aquatic ecology">Category</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Fishery_science_topics26" style="padding:3px"><table class="nowraplinks mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Fishery_science_topics" title="Template:Fishery science topics"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Fishery_science_topics" title="Template talk:Fishery science topics"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Fishery_science_topics" title="Special:EditPage/Template:Fishery science topics"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Fishery_science_topics26" style="font-size:114%;margin:0 4em"><a href="/wiki/Fishery" title="Fishery">Fishery</a> science topics</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Fisheries_science_and_wild_fisheries26" style="font-size:114%;margin:0 4em"><a href="/wiki/Fisheries_science" title="Fisheries science">Fisheries science</a> and <a href="/wiki/Wild_fisheries" title="Wild fisheries">wild fisheries</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:7em"><a href="/wiki/Fisheries_science" title="Fisheries science">Fisheries<br />science</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Population_dynamics_of_fisheries" title="Population dynamics of fisheries">Population dynamics of fisheries</a></li> <li><a href="/wiki/Shifting_baseline" title="Shifting baseline">Shifting baseline</a></li> <li><a href="/wiki/Fish_stocks" title="Fish stocks">Fish stock</a></li> <li><a href="/wiki/Fish_mortality" title="Fish mortality">Fish mortality</a></li> <li><a href="/wiki/Condition_index_in_fish" title="Condition index in fish">Condition index</a></li> <li><a href="/wiki/Stock_assessment" title="Stock assessment">Stock assessment</a></li> <li><a href="/wiki/Fish_measurement" title="Fish measurement">Fish measurement</a></li> <li><a href="/wiki/Standard_weight_in_fish" title="Standard weight in fish">Standard weight in fish</a></li> <li><a href="/wiki/Fish_counter" title="Fish counter">Fish counter</a></li> <li><a href="/wiki/Data_storage_tag" title="Data storage tag">Data storage tag</a></li> <li><a href="/wiki/Catch_per_unit_effort" title="Catch per unit effort">Catch per unit effort</a></li> <li><a href="/wiki/Otolith_microchemical_analysis" title="Otolith microchemical analysis">Otolith microchemical analysis</a></li> <li><a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">Biomass</a></li> <li><a href="/wiki/Fisheries_acoustics" title="Fisheries acoustics">Fisheries acoustics</a></li> <li><a href="/wiki/Acoustic_tag" title="Acoustic tag">Acoustic tag</a></li> <li><a href="/wiki/Coded_wire_tag" title="Coded wire tag">Coded wire tag</a></li> <li><a href="/wiki/GIS_and_aquatic_science" title="GIS and aquatic science">GIS and aquatic science</a></li> <li><a href="/wiki/Pop-up_satellite_archival_tag" title="Pop-up satellite archival tag">Pop-up satellite archival tag</a></li> <li><a href="/wiki/EcoSCOPE" title="EcoSCOPE">EcoSCOPE</a></li> <li><a href="/wiki/Age_class_structure" title="Age class structure">Age class structure</a></li> <li><a href="/wiki/Trophic_level" title="Trophic level">Trophic level</a></li> <li><a class="mw-selflink selflink">Trophic cascades</a></li> <li><a href="/wiki/Hydrographic_containment" title="Hydrographic containment">Hydrographic containment</a></li> <li><a href="/wiki/Match/mismatch" title="Match/mismatch">Match/mismatch hypothesis</a></li> <li><a href="/wiki/Climate_change_and_fisheries" title="Climate change and fisheries">Climate change and fisheries</a></li> <li><a href="/wiki/Marine_biology" title="Marine biology">Marine biology</a></li> <li><a href="/wiki/Aquatic_ecosystem" title="Aquatic ecosystem">Aquatic ecosystems</a></li> <li><a href="/wiki/Bioeconomics_(fisheries)" title="Bioeconomics (fisheries)">Bioeconomics</a></li> <li><a href="/wiki/EconMult" title="EconMult">EconMult</a></li> <li><a href="/wiki/Ecopath" title="Ecopath">Ecopath</a></li> <li><a href="/wiki/FishBase" title="FishBase">FishBase</a></li> <li><a href="/wiki/Census_of_Marine_Life" title="Census of Marine Life">Census of Marine Life</a></li> <li><a href="/wiki/OSTM/Jason-2" title="OSTM/Jason-2">OSTM/Jason-2</a></li> <li><a href="/wiki/Category:Fisheries_databases" title="Category:Fisheries databases">Fisheries databases</a></li> <li><a href="/wiki/Category:Fisheries_and_aquaculture_research_institutes" title="Category:Fisheries and aquaculture research institutes">Institutes</a></li> <li><a href="/wiki/Fisheries_science#Notable_contributors" title="Fisheries science">Fisheries scientists</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="2" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Great_Barracuda_off_the_Netherland_Antilles.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/81/Great_Barracuda_off_the_Netherland_Antilles.jpg/140px-Great_Barracuda_off_the_Netherland_Antilles.jpg" decoding="async" width="140" height="105" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/81/Great_Barracuda_off_the_Netherland_Antilles.jpg/210px-Great_Barracuda_off_the_Netherland_Antilles.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/81/Great_Barracuda_off_the_Netherland_Antilles.jpg/280px-Great_Barracuda_off_the_Netherland_Antilles.jpg 2x" data-file-width="600" data-file-height="450" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em"><a href="/wiki/Wild_fisheries" title="Wild fisheries">Wild<br />fisheries</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ocean_fisheries" title="Ocean fisheries">Ocean fisheries</a></li> <li><a href="/wiki/Diversity_of_fish" title="Diversity of fish">Diversity of fish</a></li> <li><a href="/wiki/Coastal_fish" title="Coastal fish">Coastal fish</a></li> <li><a href="/wiki/Coral_reef_fish" title="Coral reef fish">Coral reef fish</a></li> <li><a href="/wiki/Demersal_fish" title="Demersal fish">Demersal fish</a></li> <li><a href="/wiki/Forage_fish" title="Forage fish">Forage fish</a></li> <li><a href="/wiki/Pelagic_fish" title="Pelagic fish">Pelagic fish</a></li> <li><a href="/wiki/Cod_fisheries" title="Cod fisheries">Cod fisheries</a></li> <li><a href="/wiki/Crab_fisheries" title="Crab fisheries">Crab fisheries</a></li> <li><a href="/wiki/Eel_life_history" title="Eel life history">Eel fisheries</a></li> <li><a href="/wiki/Krill_fishery" title="Krill fishery">Krill fisheries</a></li> <li><a href="/wiki/Kelp#Kelp_in_history_and_culture" title="Kelp">Kelp fisheries</a></li> <li><a href="/wiki/Lobster_fishing" title="Lobster fishing">Lobster fisheries</a></li> <li><a href="/wiki/Shrimp_fishery" title="Shrimp fishery">Shrimp fishery</a></li> <li><a href="/wiki/Eel_ladder" class="mw-redirect" title="Eel ladder">Eel ladder</a></li> <li><a href="/wiki/Fish_ladder" title="Fish ladder">Fish ladder</a></li> <li><a href="/wiki/Fish_screen" title="Fish screen">Fish screen</a></li> <li><a href="/wiki/Fish_migration" title="Fish migration">Migration</a></li> <li><a href="/wiki/Salmon_run" title="Salmon run">Salmon run</a></li> <li><a href="/wiki/Sardine_run" title="Sardine run">Sardine run</a></li> <li><a href="/wiki/Shoaling_and_schooling" title="Shoaling and schooling">Shoaling and schooling</a></li> <li><a href="/wiki/Marine_habitat" title="Marine habitat">Marine habitat</a></li> <li><a href="/wiki/Marine_snow" title="Marine snow">Marine snow</a></li> <li><a href="/wiki/Water_column" title="Water column">Water column</a></li> <li><a href="/wiki/Upwelling" title="Upwelling">Upwelling</a></li> <li><a href="/wiki/Humboldt_Current" title="Humboldt Current">Humboldt current</a></li> <li><a href="/wiki/Algal_bloom" title="Algal bloom">Algal blooms</a></li> <li><a href="/wiki/Dead_zone_(ecology)" title="Dead zone (ecology)">Dead zones</a></li> <li><a href="/wiki/Fish_kill" title="Fish kill">Fish kill</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Fisheries_law,_management,_sustainability_and_conservation26" style="font-size:114%;margin:0 4em"><a href="/wiki/Fisheries_law" title="Fisheries law">Fisheries law</a>, <a href="/wiki/Fisheries_management" title="Fisheries management">management</a>, <a href="/wiki/Sustainable_fishery" title="Sustainable fishery">sustainability</a> and <a href="/wiki/Marine_conservation" title="Marine conservation">conservation</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:7em">Law</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Fisheries_law" title="Fisheries law">Fisheries law</a></li> <li><a href="/wiki/Exclusive_economic_zone" title="Exclusive economic zone">Exclusive economic zone</a></li> <li><a href="/wiki/Illegal,_unreported_and_unregulated_fishing" title="Illegal, unreported and unregulated fishing">Illegal, unreported and unregulated fishing</a></li> <li><a href="/wiki/United_Nations_Convention_on_the_Law_of_the_Sea" title="United Nations Convention on the Law of the Sea">United Nations Convention on the Law of the Sea</a> <ul><li><a href="/wiki/United_Nations_Fish_Stocks_Agreement" title="United Nations Fish Stocks Agreement">United Nations Fish Stocks Agreement</a></li></ul></li> <li><a href="/wiki/Fisheries_Convention" title="Fisheries Convention">Fisheries Convention</a></li> <li><a href="/wiki/Magnuson%E2%80%93Stevens_Fishery_Conservation_and_Management_Act" title="Magnuson–Stevens Fishery Conservation and Management Act">Magnuson–Stevens Act</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="6" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Yellowfin_tuna_nurp.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f9/Yellowfin_tuna_nurp.jpg/140px-Yellowfin_tuna_nurp.jpg" decoding="async" width="140" height="99" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f9/Yellowfin_tuna_nurp.jpg/210px-Yellowfin_tuna_nurp.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f9/Yellowfin_tuna_nurp.jpg/280px-Yellowfin_tuna_nurp.jpg 2x" data-file-width="1558" data-file-height="1106" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em">Management</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Fisheries_management" title="Fisheries management">Fisheries management</a></li> <li><a href="/wiki/Fisheries_monitoring_control_and_surveillance" title="Fisheries monitoring control and surveillance">Monitoring control and surveillance</a></li> <li><a href="/wiki/Vessel_monitoring_system" title="Vessel monitoring system">Vessel monitoring system</a></li> <li><a href="/wiki/Fishery_Resources_Monitoring_System" title="Fishery Resources Monitoring System">Fishery Resources Monitoring System</a></li> <li><a href="/wiki/Catch_reporting" title="Catch reporting">Catch reporting</a></li> <li><a href="/wiki/Fisheries_observer" title="Fisheries observer">Fisheries observer</a></li> <li><a href="/wiki/Catch_share" title="Catch share">Catch share</a></li> <li><a href="/wiki/Individual_fishing_quota" title="Individual fishing quota">Individual fishing quota</a></li> <li><a href="/wiki/Minimum_landing_size" title="Minimum landing size">Minimum landing size</a></li> <li><a href="/wiki/Pulse_fishing" title="Pulse fishing">Pulse fishing</a></li> <li><a href="/wiki/Slot_limit" title="Slot limit">Slot limit</a></li> <li><a href="/wiki/Common_Fisheries_Policy" title="Common Fisheries Policy">EU Common Fisheries Policy</a></li> <li><a href="/wiki/European_Fishery_MLS" title="European Fishery MLS">EU MLS</a></li> <li><a href="/wiki/European_Fisheries_Control_Agency" title="European Fisheries Control Agency">European Fisheries Control Agency</a></li> <li><a href="/wiki/National_Marine_Fisheries_Service" title="National Marine Fisheries Service">National Marine Fisheries Service</a></li> <li><a href="/wiki/Regional_fishery_body" title="Regional fishery body">Regional fishery body</a> <ul><li><a href="/wiki/Regional_fishery_management_organisation" class="mw-redirect" title="Regional fishery management organisation">Regional fishery management organisation</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em">Sustainability</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Sustainable_fishery" title="Sustainable fishery">Sustainable fishery</a></li> <li><a href="/wiki/Maximum_sustainable_yield" title="Maximum sustainable yield">Maximum sustainable yield</a></li> <li><a href="/wiki/List_of_commercially_important_fish_species" title="List of commercially important fish species">List of commercially important fish species</a></li> <li><a href="/wiki/Sustainable_seafood" title="Sustainable seafood">Sustainable seafood</a></li> <li><a href="/wiki/Overfishing" title="Overfishing">Overfishing</a></li> <li><a href="/wiki/Environmental_impact_of_fishing" title="Environmental impact of fishing">Environmental impact of fishing</a></li> <li><a href="/wiki/Fishing_down_the_food_web" title="Fishing down the food web">Fishing down the food web</a></li> <li><a href="/wiki/Destructive_fishing_practices" title="Destructive fishing practices">Destructive fishing practices</a></li> <li><a href="/wiki/Future_of_Marine_Animal_Populations" title="Future of Marine Animal Populations">Future of Marine Animal Populations</a></li> <li><i><a href="/wiki/The_Sunken_Billions" title="The Sunken Billions">The Sunken Billions</a></i></li> <li><i><a href="/wiki/The_End_of_the_Line_(book)" title="The End of the Line (book)">The End of the Line</a></i></li> <li><a href="/wiki/Bycatch" title="Bycatch">Bycatch</a></li> <li><a href="/wiki/Discards" title="Discards">Discards</a></li> <li><a href="/wiki/Incidental_catch" title="Incidental catch">Incidental catch</a></li> <li><a href="/wiki/Cetacean_bycatch" title="Cetacean bycatch">Cetacean bycatch</a></li> <li><a href="/wiki/Turtle_excluder_device" title="Turtle excluder device">Turtle excluder device</a></li> <li><a href="/wiki/Shrimp-Turtle_Case" title="Shrimp-Turtle Case"><i>Shrimp-Turtle</i> case</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em">Conservation</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marine_protected_area" title="Marine protected area">Marine protected area</a></li> <li><a href="/wiki/Marine_reserve" title="Marine reserve">Marine reserve</a></li> <li><a href="/wiki/Marine_conservation" title="Marine conservation">Marine conservation</a></li> <li><a href="/wiki/Marine_conservation_activism" title="Marine conservation activism">Marine conservation activism</a></li> <li><a href="/wiki/Salmon_conservation" title="Salmon conservation">Salmon conservation</a></li> <li><a href="/wiki/Grey_nurse_shark_conservation" title="Grey nurse shark conservation">Grey nurse shark conservation</a></li> <li><a href="/wiki/Shark_sanctuary" title="Shark sanctuary">Shark sanctuary</a></li> <li><a href="/wiki/European_Environmental_Agency" class="mw-redirect" title="European Environmental Agency">European Environmental Agency</a></li> <li><a href="/wiki/United_States_Fish_and_Wildlife_Service" title="United States Fish and Wildlife Service">U.S. Fish and Wildlife Service</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em">Advocacy</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/FishAct" title="FishAct">FishAct</a></li> <li><a href="/wiki/Marine_Stewardship_Council" title="Marine Stewardship Council">Marine Stewardship Council</a></li> <li><a href="/wiki/Aquaculture_Stewardship_Council" title="Aquaculture Stewardship Council">Aquaculture Stewardship Council</a></li> <li><a href="/wiki/Friend_of_the_Sea" title="Friend of the Sea">Friend of the Sea</a></li> <li><a href="/wiki/SeaChoice" title="SeaChoice">SeaChoice</a></li> <li><a href="/wiki/Seafood_Watch" title="Seafood Watch">Seafood Watch</a></li> <li><a href="/wiki/Ocean_Conservancy" title="Ocean Conservancy">Ocean Conservancy</a></li> <li><a href="/wiki/Ocean_Outcomes" title="Ocean Outcomes">Ocean Outcomes</a></li> <li><a href="/wiki/Oceana_(non-profit_group)" title="Oceana (non-profit group)">Oceana</a></li> <li><a href="/wiki/Save_Our_Seas_Foundation" title="Save Our Seas Foundation">Save Our Seas Foundation</a></li> <li><a href="/wiki/Sea_Around_Us_(organization)" title="Sea Around Us (organization)">Sea Around Us</a></li> <li><a href="/wiki/WorldFish" title="WorldFish">WorldFish</a></li> <li><a href="/wiki/Defying_Ocean%27s_End" title="Defying Ocean's End">Defying Ocean's End</a></li> <li><a href="/wiki/Hotspot_Ecosystem_Research_and_Man%27s_Impact_On_European_Seas" title="Hotspot Ecosystem Research and Man's Impact On European Seas">HERMIONE</a></li> <li><a href="/wiki/PROFISH" title="PROFISH">PROFISH</a></li> <li><a href="/wiki/International_Seafood_Sustainability_Foundation" title="International Seafood Sustainability Foundation">International Seafood Sustainability Foundation</a></li> <li><a href="/wiki/Sea_Shepherd_Conservation_Society" title="Sea Shepherd Conservation Society">Sea Shepherd Conservation Society</a></li> <li><a href="/wiki/Greenpeace" title="Greenpeace">Greenpeace</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em">Related issues</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/CalCOFI" title="CalCOFI">CalCOFI</a></li> <li><a href="/wiki/Fish_slaughter" title="Fish slaughter">Fish slaughter</a></li> <li><a href="/wiki/Marine_pollution" title="Marine pollution">Marine pollution</a></li> <li><a href="/wiki/Mercury_in_fish" title="Mercury in fish">Mercury in fish</a></li> <li><a href="/wiki/Shark_culling" title="Shark culling">Shark culling</a></li> <li><a href="/wiki/Shark_finning" title="Shark finning">Shark finning</a></li> <li><a href="/wiki/List_of_threatened_sharks" title="List of threatened sharks">Threatened sharks</a></li> <li><a href="/wiki/List_of_threatened_rays" title="List of threatened rays">Threatened rays</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li><a href="/wiki/Outline_of_fishing" title="Outline of fishing">Outline of fishing</a></li> <li><a href="/wiki/Index_of_fishing_articles" title="Index of fishing articles">Index of fishing articles</a></li> <li><a href="/wiki/Glossary_of_fishery_terms" title="Glossary of fishery terms">Glossary of fishery terms</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?useformat=desktop&type=1x1&usesul3=0" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" 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