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Breviatea - Wikipedia

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dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Group of protists</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(245,215,255)">Breviatea </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Mastigamoeba_invertens.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/db/Mastigamoeba_invertens.jpg/220px-Mastigamoeba_invertens.jpg" decoding="async" width="220" height="90" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/db/Mastigamoeba_invertens.jpg/330px-Mastigamoeba_invertens.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/db/Mastigamoeba_invertens.jpg/440px-Mastigamoeba_invertens.jpg 2x" data-file-width="832" data-file-height="341" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i><a href="/wiki/Breviata_anathema" class="mw-redirect" title="Breviata anathema">Breviata anathema</a></i> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(245,215,255)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Breviatidae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Amorphea" title="Amorphea">Amorphea</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Obazoa" title="Obazoa">Obazoa</a> </td></tr> <tr> <td>Class: </td> <td><a class="mw-selflink selflink">Breviatea</a><br /><small><a href="/wiki/Cavalier-Smith" class="mw-redirect" title="Cavalier-Smith">Cavalier-Smith</a>, 2004<sup id="cite_ref-CavSmith_et_al_2004_2-1" class="reference"><a href="#cite_note-CavSmith_et_al_2004-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup></small> </td></tr> <tr> <td>Order: </td> <td><a class="mw-selflink selflink">Breviatida</a><br /><small>Cavalier-Smith, 2004<sup id="cite_ref-CavSmith_et_al_2004_2-0" class="reference"><a href="#cite_note-CavSmith_et_al_2004-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup></small> </td></tr> <tr> <td>Family: </td> <td><a class="mw-selflink selflink">Breviatidae</a><br /><small>Cavalier-Smith, 2013<sup id="cite_ref-Cavalier-Smith_et_al._2013_1-0" class="reference"><a href="#cite_note-Cavalier-Smith_et_al._2013-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup></small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(245,215,255)">Genera </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i><a href="/wiki/Breviata" title="Breviata">Breviata</a></i></li> <li><i><a href="/w/index.php?title=Subulatomonas&amp;action=edit&amp;redlink=1" class="new" title="Subulatomonas (page does not exist)">Subulatomonas</a></i></li> <li><i><a href="/w/index.php?title=Lenisia&amp;action=edit&amp;redlink=1" class="new" title="Lenisia (page does not exist)">Lenisia</a></i></li> <li><i><a href="/w/index.php?title=Pygsuia&amp;action=edit&amp;redlink=1" class="new" title="Pygsuia (page does not exist)">Pygsuia</a></i></li></ul> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(245,215,255)"><a href="/wiki/Biodiversity" title="Biodiversity">Diversity</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><a href="#Classification">4 species</a> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Breviatea</b>, commonly known as <b>breviate amoebae</b>,<sup id="cite_ref-Brown_2013_3-0" class="reference"><a href="#cite_note-Brown_2013-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> are a group of free-living, <a href="/wiki/Amitochondriate" class="mw-redirect" title="Amitochondriate">amitochondriate</a> <a href="/wiki/Protist" title="Protist">protists</a> with uncertain phylogenetic position.<sup id="cite_ref-Hamann_2016_4-0" class="reference"><a href="#cite_note-Hamann_2016-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> They are <a href="/wiki/Flagellate" title="Flagellate">biflagellate</a>, and can live in anaerobic (oxygen-free) environments.<sup id="cite_ref-Hamann_2016_4-1" class="reference"><a href="#cite_note-Hamann_2016-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Brown_2013_3-1" class="reference"><a href="#cite_note-Brown_2013-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Modern_Soil_Biology_3rd_5-0" class="reference"><a href="#cite_note-Modern_Soil_Biology_3rd-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> They are currently placed in the <a href="/wiki/Obazoa" title="Obazoa">Obazoa</a> clade.<sup id="cite_ref-Kang_2021_6-0" class="reference"><a href="#cite_note-Kang_2021-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> They likely do not possess <a href="/wiki/Vinculin_family" title="Vinculin family">vinculin</a> proteins.<sup id="cite_ref-Kang_2021_6-1" class="reference"><a href="#cite_note-Kang_2021-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> Their metabolism relies on <a href="/wiki/Fermentation" title="Fermentation">fermentative</a> production of <a href="/wiki/Adenosine_triphosphate" title="Adenosine triphosphate">ATP</a> as an adaptation to their low-oxygen environment.<sup id="cite_ref-Hamann_2016_4-2" class="reference"><a href="#cite_note-Hamann_2016-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p><p>The lineage emerged roughly one billion years ago, at a time when the oxygen content of the Earth's oceans was low, and they thus developed anaerobic lifestyles. Together with <a href="/wiki/Apusomonads" class="mw-redirect" title="Apusomonads">Apusomonads</a>, they are the closest relatives of the <a href="/wiki/Opisthokont" title="Opisthokont">Opisthokonts</a>, a group that includes animals and fungi.<sup id="cite_ref-Brown_2013_3-2" class="reference"><a href="#cite_note-Brown_2013-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Hydrogenosomes_and_Mitosomes_7-0" class="reference"><a href="#cite_note-Hydrogenosomes_and_Mitosomes-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Characteristics">Characteristics</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Breviatea&amp;action=edit&amp;section=1" title="Edit section: Characteristics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Mitochondrion-related_organelles">Mitochondrion-related organelles</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Breviatea&amp;action=edit&amp;section=2" title="Edit section: Mitochondrion-related organelles"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Mitochondrion-related organelles (MROs) are organelles that evolved from a degradation of ancestral, fully functional <a href="/wiki/Mitochondria" class="mw-redirect" title="Mitochondria">mitochondria</a>. Among Breviatea, MROs are present in <i><a href="/w/index.php?title=Pygsuia&amp;action=edit&amp;redlink=1" class="new" title="Pygsuia (page does not exist)">Pygsuia</a></i>, <i><a href="/wiki/Breviata" title="Breviata">Breviata</a></i> and <i><a href="/w/index.php?title=Subulatomonas&amp;action=edit&amp;redlink=1" class="new" title="Subulatomonas (page does not exist)">Subulatomonas</a></i>. In the cells of <i>Pygsuia</i>, for which the complete <a href="/wiki/Transcriptome" title="Transcriptome">transcriptome</a> is known, there is a single smooth MRO that lacks a <a href="/wiki/Mitochondrial_genome" class="mw-redirect" title="Mitochondrial genome">mitochondrial genome</a> and most components of the <a href="/wiki/Electron_transport_chain" title="Electron transport chain">electron transport chain</a>. Of the <a href="/wiki/Citric_acid_cycle" title="Citric acid cycle">citric acid cycle</a> enzymes, which are present in the mitochondria in other organisms, only two are present in <i>Pygsuia</i>: <a href="/wiki/Fumarase" title="Fumarase">fumarase</a> and <a href="/wiki/Succinate_dehydrogenase" title="Succinate dehydrogenase">succinate dehydrogenase</a>. In contrast, <i>Lenisia</i> cells contain multiple MROs with <a href="/wiki/Crista" title="Crista">cristae</a>.<sup id="cite_ref-Hydrogenosomes_and_Mitosomes_7-1" class="reference"><a href="#cite_note-Hydrogenosomes_and_Mitosomes-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Evolution">Evolution</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Breviatea&amp;action=edit&amp;section=3" title="Edit section: Evolution"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Breviatea is a <a href="/wiki/Clade" title="Clade">clade</a> of <a href="/wiki/Basal_group" class="mw-redirect" title="Basal group">basal</a> <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotes</a>. They are closely related to the <a href="/wiki/Apusomonad" class="mw-redirect" title="Apusomonad">apusomonads</a> and the <a href="/wiki/Opisthokonta" class="mw-redirect" title="Opisthokonta">Opisthokonta</a> supergroup, and together they compose the larger clade <a href="/wiki/Obazoa" title="Obazoa">Obazoa</a>, which is the sister group to <a href="/wiki/Amoebozoa" title="Amoebozoa">Amoebozoa</a>.<sup id="cite_ref-Brown_2013_3-3" class="reference"><a href="#cite_note-Brown_2013-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> Within Breviatea, the four known species are distributed into smaller clades of two species each: one uniting <i><a href="/wiki/Breviata" title="Breviata">Breviata</a></i> with <i><a href="/w/index.php?title=Subulatomonas&amp;action=edit&amp;redlink=1" class="new" title="Subulatomonas (page does not exist)">Subulatomonas</a></i>, and one uniting <i><a href="/w/index.php?title=Lenisia&amp;action=edit&amp;redlink=1" class="new" title="Lenisia (page does not exist)">Lenisia</a></i> with <i><a href="/w/index.php?title=Pygsuia&amp;action=edit&amp;redlink=1" class="new" title="Pygsuia (page does not exist)">Pygsuia</a></i>.<sup id="cite_ref-Hamann_2016_4-3" class="reference"><a href="#cite_note-Hamann_2016-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:90%;"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Podiata" title="Podiata">Podiata</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/CRuMs" title="CRuMs">CRuMs</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Amorphea" title="Amorphea">Amorphea</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Amoebozoa" title="Amoebozoa">Amoebozoa</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Obazoa" title="Obazoa">Obazoa</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><b>Breviatea</b>&#160;&#160; </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Breviata_anathema" class="mw-redirect" title="Breviata anathema">Breviata anathema</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Subulatomonas_tetraspora&amp;action=edit&amp;redlink=1" class="new" title="Subulatomonas tetraspora (page does not exist)">Subulatomonas tetraspora</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Lenisia_limosa&amp;action=edit&amp;redlink=1" class="new" title="Lenisia limosa (page does not exist)">Lenisia limosa</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Pygsuia_biforma&amp;action=edit&amp;redlink=1" class="new" title="Pygsuia biforma (page does not exist)">Pygsuia biforma</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Apusomonadida" class="mw-redirect" title="Apusomonadida">Apusomonadida</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Opisthokonta" class="mw-redirect" title="Opisthokonta">Opisthokonta</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Holozoa" title="Holozoa">Holozoa</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Holomycota" title="Holomycota">Holomycota</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Breviatea&amp;action=edit&amp;section=4" title="Edit section: Taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="History">History</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Breviatea&amp;action=edit&amp;section=5" title="Edit section: History"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The class Breviatea was created in 2004 by British protozoologist <a href="/wiki/Thomas_Cavalier-Smith" title="Thomas Cavalier-Smith">Thomas Cavalier-Smith</a> to group a problematic taxon previously known as <i>‘Mastigamoeba invertens’</i>. This organism, initially classified in the <a href="/wiki/Archamoebae" title="Archamoebae">Archamoebae</a> within phylum <a href="/wiki/Amoebozoa" title="Amoebozoa">Amoebozoa</a>, appeared to strongly diverge in <a href="/wiki/Phylogenetic" class="mw-redirect" title="Phylogenetic">phylogenetic</a> trees based on <a href="/wiki/Ribosomal_RNA" title="Ribosomal RNA">ribosomal RNA</a> and had a structure very different from other Archamoebae. Because of these results, <i>‘M. invertens’</i> was separated into the order <b>Breviatida</b>, contained in the monotypic class <b>Breviatea</b>.<sup id="cite_ref-CavSmith_et_al_2004_2-2" class="reference"><a href="#cite_note-CavSmith_et_al_2004-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> The organism was eventually renamed <i><a href="/wiki/Breviata_anathema" class="mw-redirect" title="Breviata anathema">Breviata anathema</a></i>.<sup id="cite_ref-Walker_2006_8-0" class="reference"><a href="#cite_note-Walker_2006-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> A family-level rank for these amoebae, <b>Breviatidae</b>, was <a href="/wiki/Taxonomy_(biology)#Taxonomic_descriptions" title="Taxonomy (biology)">formally described</a> by the same author in 2013.<sup id="cite_ref-Cavalier-Smith_et_al._2013_1-1" class="reference"><a href="#cite_note-Cavalier-Smith_et_al._2013-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Classification">Classification</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Breviatea&amp;action=edit&amp;section=6" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>There are currently four accepted genera, each containing only one species. </p> <ul><li><i><a href="/wiki/Breviata" title="Breviata">Breviata</a></i> <span style="font-size:85%;">Walker, Dacks &amp; Embley, 2006</span><sup id="cite_ref-Walker_2006_8-1" class="reference"><a href="#cite_note-Walker_2006-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> <ul><li><i><a href="/wiki/Breviata_anathema" class="mw-redirect" title="Breviata anathema">Breviata anathema</a></i> <span style="font-size:85%;">Walker, Dacks &amp; Embley, 2006</span></li></ul></li> <li><i><a href="/w/index.php?title=Lenisia&amp;action=edit&amp;redlink=1" class="new" title="Lenisia (page does not exist)">Lenisia</a></i> <span style="font-size:85%;">Hamann et al., 2016</span><sup id="cite_ref-Hamann_2016_4-4" class="reference"><a href="#cite_note-Hamann_2016-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> <ul><li><i><a href="/w/index.php?title=Lenisia_limosa&amp;action=edit&amp;redlink=1" class="new" title="Lenisia limosa (page does not exist)">Lenisia limosa</a></i> <span style="font-size:85%;">Hamann et al., 2016</span></li></ul></li> <li><i><a href="/w/index.php?title=Pygsuia&amp;action=edit&amp;redlink=1" class="new" title="Pygsuia (page does not exist)">Pygsuia</a></i> <span style="font-size:85%;">Brown et al., 2013</span><sup id="cite_ref-Brown_2013_3-4" class="reference"><a href="#cite_note-Brown_2013-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> <ul><li><i><a href="/w/index.php?title=Pygsuia_biforma&amp;action=edit&amp;redlink=1" class="new" title="Pygsuia biforma (page does not exist)">Pygsuia biforma</a></i> <span style="font-size:85%;">Brown et al., 2013</span></li></ul></li> <li><i><a href="/w/index.php?title=Subulatomonas&amp;action=edit&amp;redlink=1" class="new" title="Subulatomonas (page does not exist)">Subulatomonas</a></i> <span style="font-size:85%;">Katz et al., 2011</span><sup id="cite_ref-Subulatomonas_tetraspora_9-0" class="reference"><a href="#cite_note-Subulatomonas_tetraspora-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> <ul><li><i><a href="/w/index.php?title=Subulatomonas_tetraspora&amp;action=edit&amp;redlink=1" class="new" title="Subulatomonas tetraspora (page does not exist)">Subulatomonas tetraspora</a></i> <span style="font-size:85%;">Katz et al., 2011</span></li></ul></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Breviatea&amp;action=edit&amp;section=7" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-Cavalier-Smith_et_al._2013-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Cavalier-Smith_et_al._2013_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Cavalier-Smith_et_al._2013_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output 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Sp., the Organism Previously Studied as "<i>Mastigamoeba invertens</i>"". <i>Journal of Eukaryotic Microbiology</i>. <b>53</b> (2): 65–78. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.1550-7408.2005.00087.x">10.1111/j.1550-7408.2005.00087.x</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16579808">16579808</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:31046569">31046569</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Journal+of+Eukaryotic+Microbiology&amp;rft.atitle=Ultrastructural+Description+of+Breviata+anathema%2C+N.+Gen.%2C+N.+Sp.%2C+the+Organism+Previously+Studied+as+%22Mastigamoeba+invertens%22&amp;rft.volume=53&amp;rft.issue=2&amp;rft.pages=65-78&amp;rft.date=2006&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A31046569%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F16579808&amp;rft_id=info%3Adoi%2F10.1111%2Fj.1550-7408.2005.00087.x&amp;rft.aulast=Walker&amp;rft.aufirst=Giselle&amp;rft.au=Dacks%2C+Joel+B.&amp;rft.au=Embley%2C+T.+Martin&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABreviatea" class="Z3988"></span></span> </li> <li id="cite_note-Subulatomonas_tetraspora-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-Subulatomonas_tetraspora_9-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKatzGrantParfreyGant2011" class="citation journal cs1">Katz, Laura A.; Grant, Jessica; Parfrey, Laura Wegener; Gant, Anastasia; O’Kelly, Charles J.; Anderson, O. Roger; Molestina, Robert E.; Nerad, Thomas (November 2011). "<i>Subulatomonas tetraspora</i> nov. gen. nov. sp. is a Member of a Previously Unrecognized Major Clade of Eukaryotes". <i>Protist</i>. <b>162</b> (5): 762–773. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.protis.2011.05.002">10.1016/j.protis.2011.05.002</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21723191">21723191</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Protist&amp;rft.atitle=Subulatomonas+tetraspora+nov.+gen.+nov.+sp.+is+a+Member+of+a+Previously+Unrecognized+Major+Clade+of+Eukaryotes&amp;rft.volume=162&amp;rft.issue=5&amp;rft.pages=762-773&amp;rft.date=2011-11&amp;rft_id=info%3Adoi%2F10.1016%2Fj.protis.2011.05.002&amp;rft_id=info%3Apmid%2F21723191&amp;rft.aulast=Katz&amp;rft.aufirst=Laura+A.&amp;rft.au=Grant%2C+Jessica&amp;rft.au=Parfrey%2C+Laura+Wegener&amp;rft.au=Gant%2C+Anastasia&amp;rft.au=O%E2%80%99Kelly%2C+Charles+J.&amp;rft.au=Anderson%2C+O.+Roger&amp;rft.au=Molestina%2C+Robert+E.&amp;rft.au=Nerad%2C+Thomas&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABreviatea" class="Z3988"></span></span> </li> </ol></div></div> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist 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class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Amoebozoa" scope="row" class="navbox-group" style="width:1%;line-height:1.2em;;background:rgb(245,215,255)"><a href="/wiki/Amoebozoa" title="Amoebozoa">Amoebozoa</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Discosea" title="Discosea">Discosea</a></li> <li><a href="/wiki/Tubulinea" title="Tubulinea">Tubulinea</a></li> <li><b><a href="/wiki/Eumycetozoa" title="Eumycetozoa">Eumycetozoa</a></b></li> <li><a href="/wiki/Variosea" class="mw-redirect" title="Variosea">Variosea</a></li> <li><a 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<li><a class="mw-selflink selflink">Breviatea</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(245,215,255)"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"><a href="/wiki/Opisthokonta" class="mw-redirect" title="Opisthokonta">Opisthokonta</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Holomycota" scope="row" class="navbox-group" style="width:1%;line-height:1.2em;;background:rgb(245,215,255)"><a href="/wiki/Holomycota" title="Holomycota">Holomycota</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cristidiscoidea" title="Cristidiscoidea">Rotosphaerida</a></li> <li><b><a href="/wiki/Fungus" title="Fungus">Fungi</a></b>¹</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(245,215,255)"><span style="margin:-32px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);">&#160;&#160;&#160;<a href="/wiki/Holozoa" title="Holozoa">Holozoa</a>&#160;&#160;&#160;</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tunicaraptor" title="Tunicaraptor">Tunicaraptor</a></i></li> <li>†<i><a href="/wiki/Bicellum" class="mw-redirect" title="Bicellum">Bicellum</a></i></li> <li><a href="/wiki/Ichthyosporea" title="Ichthyosporea">Ichthyosporea</a></li> <li><a href="/wiki/Pluriformea" title="Pluriformea">Pluriformea</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(245,215,255)"><a href="/wiki/Filozoa" title="Filozoa">Filozoa</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Filasterea" title="Filasterea">Filasterea</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(245,215,255)"><a href="/wiki/Choanozoa" title="Choanozoa">Choanozoa</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Choanoflagellate" title="Choanoflagellate">Choanoflagellata</a></li> <li><b><a href="/wiki/Animal" title="Animal">Animalia</a></b>¹</li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.2em;"><span style="margin:-34px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"><a href="/wiki/Diaphoretickes" title="Diaphoretickes">Diaphoretickes</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Provora" title="Provora">Provora</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(200,250,80)"><a href="/wiki/Haptista" title="Haptista">Haptista</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Centroplasthelida" class="mw-redirect" title="Centroplasthelida">Centroplasthelida</a></li> <li><a href="/wiki/Haptophyte" title="Haptophyte">Haptophyta</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.2em;;background:rgb(200,250,80)"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);">&#160;&#160;&#160;&#160;<a href="/wiki/TSAR" class="mw-redirect" title="TSAR">TSAR</a>&#160;&#160;&#160;&#160;</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Telonemia" title="Telonemia">Telonemia</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.2em;;background:rgb(200,250,80)"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);">&#160;&#160;&#160;&#160;&#160;<a href="/wiki/SAR_supergroup" title="SAR supergroup">SAR</a>&#160;&#160;&#160;&#160;&#160;</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(200,250,80)"><a href="/wiki/Rhizaria" title="Rhizaria">Rhizaria</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cercozoa" title="Cercozoa">Cercozoa</a></li> <li><a href="/wiki/Endomyxa" title="Endomyxa">Endomyxa</a></li> <li><b><a href="/wiki/Foraminifera" title="Foraminifera">Foraminifera</a></b></li> <li><b><a href="/wiki/Radiolaria" title="Radiolaria">Radiolaria</a></b></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(200,250,80)"><a href="/wiki/Alveolata" class="mw-redirect" title="Alveolata">Alveolata</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Colponemida" class="mw-redirect" title="Colponemida">Colponemida</a>*</li> <li><b><a href="/wiki/Ciliate" title="Ciliate">Ciliophora</a></b></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(200,250,80); vertical-align:middle;"><a href="/wiki/Myzozoa" title="Myzozoa">Myzozoa</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b><a href="/wiki/Apicomplexa" title="Apicomplexa">Apicomplexa</a></b></li> <li><a href="/wiki/Chrompodellid" title="Chrompodellid">Chrompodellida</a></li> <li><b><a href="/wiki/Dinoflagellate" title="Dinoflagellate">Dinoflagellata</a></b></li> <li><a href="/wiki/Perkinsozoa" class="mw-redirect" title="Perkinsozoa">Perkinsozoa</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(200,250,80)"><a href="/wiki/Stramenopiles" class="mw-redirect" title="Stramenopiles">Stramenopiles</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Platysulcus" title="Platysulcus">Platysulcus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(200,250,80)"><a href="/wiki/Bigyra" title="Bigyra">Bigyra</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bicosoecea" class="mw-redirect" title="Bicosoecea">Bicosoecida</a></li> <li><a href="/wiki/Placidozoa" title="Placidozoa">Placidozoa</a></li> <li><a href="/wiki/Sagenista" title="Sagenista">Sagenista</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(200,250,80)"><a href="/wiki/Gyrista" title="Gyrista">Gyrista</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Developea" class="mw-redirect" title="Developea">Developea</a></li> <li><a href="/wiki/Pirsoniales" class="mw-redirect" title="Pirsoniales">Pirsoniales</a></li> <li><a href="/wiki/Hyphochytriomycetes" title="Hyphochytriomycetes">Hyphochytriomycetes</a></li> <li><b><a href="/wiki/Ochrophyta" class="mw-redirect" title="Ochrophyta">Ochrophyta</a></b></li> <li><b><a href="/wiki/Oomycetes" class="mw-redirect" title="Oomycetes">Oomycetes</a></b></li> <li><a href="/wiki/Actinophryida" class="mw-redirect" title="Actinophryida">Actinophryida</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.2em;background:rgb(180,250,180)"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);">&#160;&#160;&#160;&#160;&#160;<a href="/wiki/CAM_clade" class="mw-redirect" title="CAM clade">CAM</a>&#160;&#160;&#160;&#160;&#160;</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(180,250,180)"><a href="/wiki/Pancryptista" title="Pancryptista">Pancryptista</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Microheliella" title="Microheliella">Microheliella</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(180,250,180)"><a href="/wiki/Cryptista" title="Cryptista">Cryptista</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Palpitomonas" title="Palpitomonas">Palpitomonas</a></i></li> <li><a href="/wiki/Cryptophyta" class="mw-redirect" title="Cryptophyta">Cryptophyta</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:rgb(180,250,180)"><a href="/wiki/Archaeplastida" title="Archaeplastida">Archaeplastida</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glaucophyta" class="mw-redirect" title="Glaucophyta">Glaucophyta</a></li> <li><a href="/wiki/Picozoa" title="Picozoa">Picozoa</a></li> <li><a href="/wiki/Rhodelphidia" class="mw-redirect" title="Rhodelphidia">Rhodelphidia</a></li> <li><b><a href="/wiki/Rhodophyta" class="mw-redirect" title="Rhodophyta">Rhodophyta</a></b></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Viridiplantae(Plants_sensu_lato)" scope="row" class="navbox-group" style="width:1%;background:rgb(180,250,180)"><a href="/wiki/Viridiplantae" title="Viridiplantae">Viridiplantae</a><br /><span class="nobold">(<a href="/wiki/Plant#Alternative_concepts" title="Plant">Plants <i>sensu lato</i></a>)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Prasinodermophyta" title="Prasinodermophyta">Prasinodermophyta</a></li> <li><b><a href="/wiki/Chlorophyta" title="Chlorophyta">Chlorophyta</a></b></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Streptophyta" scope="row" class="navbox-group" style="width:1%;background:rgb(180,250,180)"><a href="/wiki/Streptophyta" title="Streptophyta">Streptophyta</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chlorokybus" title="Chlorokybus">Chlorokybus</a></i></li> <li><i><a href="/wiki/Mesostigma" title="Mesostigma">Mesostigma</a></i></li> <li><a href="/wiki/Klebsormidiophyceae" class="mw-redirect" title="Klebsormidiophyceae">Klebsormidiophyceae</a></li> <li><a href="/wiki/Charophyceae" title="Charophyceae">Charophyceae</a></li> <li><a href="/wiki/Coleochaetophyceae" title="Coleochaetophyceae">Coleochaetophyceae</a></li> <li><b><a href="/wiki/Zygnematophyceae" title="Zygnematophyceae">Zygnematophyceae</a></b></li> <li><b><a href="/wiki/Embryophyta" class="mw-redirect" title="Embryophyta">Embryophyta</a></b>¹</li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.2em;"><span style="margin:-23px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"><a href="/wiki/Excavata" title="Excavata">Excavates</a>*</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Discoba" class="mw-redirect" title="Discoba">Discoba</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Jakobida" class="mw-redirect" title="Jakobida">Jakobida</a></li> <li><i><a href="/wiki/Tsukubamonas" title="Tsukubamonas">Tsukubamonas</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Discicristata" title="Discicristata">Discicristata</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Heterolobosea" title="Heterolobosea">Heterolobosea</a></li> <li><b><a href="/wiki/Euglenozoa" title="Euglenozoa">Euglenozoa</a></b></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Metamonada" class="mw-redirect" title="Metamonada">Metamonada</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Barthelona" title="Barthelona">Barthelona</a></i></li> <li><a href="/w/index.php?title=Anaeramoebae&amp;action=edit&amp;redlink=1" class="new" title="Anaeramoebae (page does not exist)">Anaeramoebae</a></li> <li><a href="/wiki/Anaeromonada" class="mw-redirect" title="Anaeromonada">Anaeromonada</a></li> <li><a href="/wiki/Fornicata" class="mw-redirect" title="Fornicata">Fornicata</a></li> <li><a href="/wiki/Parabasalia" class="mw-redirect" title="Parabasalia">Parabasalia</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Malawimonada" class="mw-redirect" title="Malawimonada">Malawimonada</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Malawimonadidae" title="Malawimonadidae">Malawimonadidae</a></li> <li><a href="/wiki/Imasidae" class="mw-redirect" title="Imasidae">Imasidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.0em;"><span style="margin:-31px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"><i><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></i></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Acritarch" title="Acritarch">Acritarchs</a></li> <li>†<i><a href="/wiki/Grypania" title="Grypania">Grypania</a></i></li> <li>†<a href="/wiki/Gunflint_chert" title="Gunflint chert">Gunflint microbiota</a></li> <li>†<a href="/wiki/Chitinozoan" title="Chitinozoan">Chitinozoan</a></li> <li><i><a href="/wiki/Meteora_sporadica" title="Meteora sporadica">Meteora sporadica</a></i></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Hemimastigophora" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Hemimastigophora" title="Hemimastigophora">Hemimastigophora</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Spironematellidae" title="Spironematellidae">Spironematellidae</a></li> <li><a href="/wiki/Paramastigidae" class="mw-redirect" title="Paramastigidae">Paramastigidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ancyromonadida" title="Ancyromonadida">Ancyromonadida</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Ancyromonadidae&amp;action=edit&amp;redlink=1" class="new" title="Ancyromonadidae (page does not exist)">Ancyromonadidae</a></li> <li><a href="/w/index.php?title=Planomonadidae&amp;action=edit&amp;redlink=1" class="new" title="Planomonadidae (page does not exist)">Planomonadidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/CRuMs" title="CRuMs">CRuMs</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Collodictyonidae" title="Collodictyonidae">Collodictyonidae</a></li> <li><a href="/wiki/Mantamonadida" class="mw-redirect" title="Mantamonadida">Mantamonadida</a></li> <li><a href="/wiki/Rigifilida" title="Rigifilida">Rigifilida</a></li></ul> </div></td></tr></tbody></table><div> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>¹traditional <a 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