CINXE.COM
Neuroscience | The Canadian Encyclopedia
<!DOCTYPE html> <html lang="en"> <head> <meta name="title" content="Neuroscience"> <meta property="og:title" content="Neuroscience"> <meta property="og:description" content="Neuroscience is the study of the structure and function of the nervous system. It includes a number of subfields such as neuroanatomy, neurochemistry, neurophar..."> <meta property="og:image" content="https://d3d0lqu00lnqvz.cloudfront.net/TCE_placeholder.png"> <meta name="twitter:card" content="summary_large_image"> <meta name="twitter:title" content="Neuroscience"> <meta name="twitter:description" content="Neuroscience is the study of the structure and function of the nervous system. It includes a number of subfields such as neuroanatomy, neurochemistry, neurophar..."> <meta name="twitter:image" content="https://d3d0lqu00lnqvz.cloudfront.net/TCE_placeholder.png"> <!-- Google Tag Manager --> <script async> (function(w,d,s,l,i){w[l]=w[l]||[];w[l].push({'gtm.start': new Date().getTime(),event:'gtm.js'});var f=d.getElementsByTagName(s)[0], j=d.createElement(s),dl=l!='dataLayer'?'&l='+l:'';j.async=true;j.src= 'https://www.googletagmanager.com/gtm.js?id='+i+dl;f.parentNode.insertBefore(j,f); })(window,document,'script','dataLayer','GTM-NT3GRKD'); </script> <noscript> <iframe src="https://www.googletagmanager.com/ns.html?id=GTM-NT3GRKD" height="0" width="0" style="display:none;visibility:hidden" ></iframe> </noscript> <!-- End Google Tag Manager --> <!-- Push user ID --> <script> if (document.querySelector('.mainUserSubMenu') !== null) { dataLayer.push({ 'userID': '' }); } </script> <script> const Laravel = { csrfToken: "TBjMRcqZMlBKqq8FfQbZoiVreBacEvyIeMS7AQMI", locale: "en" }; </script> <title>Neuroscience | The Canadian Encyclopedia</title> <meta charset="utf-8"> <meta http-equiv="X-UA-Compatible" content="IE=edge,chrome=1"> <meta name="viewport" content="width=device-width, initial-scale=1"> <link rel="manifest" href="https://www.thecanadianencyclopedia.ca/manifest.json"> <meta name="description" content="Neuroscience is the study of the structure and function of the nervous system. It includes a number of subfields such as neuroanatomy, neurochemistry, neurophar..."> <link rel="apple-touch-icon" sizes="57x57" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-57x57.png?v=f22552fe571739798798bb695565903f"> <link rel="apple-touch-icon" sizes="60x60" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-60x60.png?v=f417df6dda9ce41f908afc590778f94b"> <link rel="apple-touch-icon" sizes="72x72" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-72x72.png?v=3af0c2e4ca8c973438f91f44e7a58d6c"> <link rel="apple-touch-icon" sizes="76x76" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-76x76.png?v=41438835e152256cf9b311eb7eb116c0"> <link rel="apple-touch-icon" sizes="192x192" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-precomposed.png?v=6d047b2f33e6b90d2af0b41ac01b0566"> <link rel="apple-touch-icon" sizes="192x192" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon.png?v=6d047b2f33e6b90d2af0b41ac01b0566"> <link rel="apple-touch-icon" sizes="114x114" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-114x114.png?v=91501eda47ce85d33e0feae1630a4e99"> <link rel="apple-touch-icon" sizes="120x120" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-120x120.png?v=43adb2769c1827f1dda68db736544ec4"> <link rel="apple-touch-icon" sizes="144x144" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-144x144.png?v=670a8df278b4101e8bf081d7b9ebb258"> <link rel="apple-touch-icon" sizes="152x152" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-152x152.png?v=db581da4dde2d406e52a132691b14680"> <link rel="apple-touch-icon" sizes="180x180" href="https://www.thecanadianencyclopedia.ca/images/favicon/apple-icon-180x180.png?v=8d4920caa59ad54fb454f2cd80ebfc1c"> <link rel="icon" type="image/png" sizes="192x192" href="https://www.thecanadianencyclopedia.ca/images/favicon/android-icon-192x192.png?v=6d047b2f33e6b90d2af0b41ac01b0566"> <link rel="icon" type="image/png" sizes="36x36" href="https://www.thecanadianencyclopedia.ca/images/favicon/android-icon-36x36.png?v=3e9842d534281da81a0f4b4c81a17592"> <link rel="icon" type="image/png" sizes="48x48" href="https://www.thecanadianencyclopedia.ca/images/favicon/android-icon-48x48.png?v=7eb74dc9409ece0737d81a8ba5841bf7"> <link rel="icon" type="image/png" sizes="72x72" href="https://www.thecanadianencyclopedia.ca/images/favicon/android-icon-72x72.png?v=3af0c2e4ca8c973438f91f44e7a58d6c"> <link rel="icon" type="image/png" sizes="36x36" href="https://www.thecanadianencyclopedia.ca/images/favicon/android-icon-36x36.png?v=3e9842d534281da81a0f4b4c81a17592"> <link rel="icon" type="image/png" sizes="44x44" href="https://www.thecanadianencyclopedia.ca/images/favicon/android-icon-44x44.png"> <link rel="icon" type="image/png" sizes="96x96" href="https://www.thecanadianencyclopedia.ca/images/favicon/favicon-96x96.png?v=a46206812744ada382bc23a1bc4f1357"> <link rel="icon" type="image/png" sizes="96x96" href="https://www.thecanadianencyclopedia.ca/images/favicon/favicon-96x96.png?v=a46206812744ada382bc23a1bc4f1357"> <link rel="icon" type="image/png" sizes="16x16" href="https://www.thecanadianencyclopedia.ca/images/favicon/favicon-16x16.png?v=ff43d9c7c67bb75a3d9a471a65e6724f"> <link rel="manifest" href="https://www.thecanadianencyclopedia.ca/images/favicon/manifest.json?v=de0c530598b698ec2dbe765b440b7479"> <meta name="msapplication-TileColor" content="#ffffff"> <meta name="msapplication-TileImage" content="/ms-icon-144x144.png')}}"> <meta name="theme-color" content="#ffffff"> <link rel="shortcut icon" href="https://www.thecanadianencyclopedia.ca/images/favicon/favicon.ico?v=45c5ae5e358dcde8acbbf3140859fb18" type="image/x-icon"> <link rel="icon" href="https://www.thecanadianencyclopedia.ca/images/favicon/favicon.ico?v=45c5ae5e358dcde8acbbf3140859fb18" type="image/x-icon"> <link rel="canonical" href="https://www.thecanadianencyclopedia.ca/en/article/neuroscience"> <link rel="alternate" hreflang="fr" href="https://www.thecanadianencyclopedia.ca/fr/article/neuroscience"> <!-- preload fonts --> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/publico/Publico-Roman.ttf?v=da5cf4834925266e3fe9373d7c20987a" as="font" type="font/ttf" crossorigin> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/publico/Publico-Bold.ttf?v=3ad7881260304548a929a8add091efad" as="font" type="font/ttf" crossorigin> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/publico/Publico-Italic.ttf?v=a670bc7591f940131b96c1427a8bfcf9" as="font" type="font/ttf" crossorigin> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/publico/Publico-BoldItalic.ttf?v=1e6ee343e8f0d8d4784e7adaf55be5ed" as="font" type="font/ttf" crossorigin> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/avenir/Avenir-Roman.ttf?v=c4ed2390b8880049113ca6b174e5393e" as="font" type="font/ttf" crossorigin> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/avenir/Avenir-Heavy.ttf?v=6880b7f440fee3d3d23778d295fda4fc" as="font" type="font/ttf" crossorigin> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/avenir/Avenir-Oblique.ttf?v=b934234a7cf41ebc3b0ed646c856d94a" as="font" type="font/ttf" crossorigin> <link rel="preload" href="https://www.thecanadianencyclopedia.ca/fonts/avenir/Avenir-HeavyOblique.ttf?v=86f5c8cc80dbc9fdac444dacf0798808" as="font" type="font/ttf" crossorigin> <!-- end preload fonts --> <link rel="stylesheet" type="text/css" href="https://www.thecanadianencyclopedia.ca/css/styles.css?v=d49542238e3b3e929d3830648a711e8e"> <link rel="stylesheet" type="text/css" href="https://cdnjs.cloudflare.com/ajax/libs/Trumbowyg/2.20.0/ui/trumbowyg.min.css"> <script> // Polyfill if IE11 var isIE11 = !!window.MSInputMethodContext && !!document.documentMode; if (isIE11) { document.write('<script src="https://cdnjs.cloudflare.com/polyfill/v3/polyfill.min.js"><\/script>'); } if ('serviceWorker' in navigator ) { window.addEventListener('load', function() { navigator.serviceWorker.register('/service-worker.js').then(function(registration) { // Registration was successful console.log('ServiceWorker registration successful with scope: ', registration.scope); }, function(err) { // registration failed :( console.log('ServiceWorker registration failed: ', err); }); }); } </script> <script src="https://code.jquery.com/jquery-2.2.4.min.js" integrity="sha256-BbhdlvQf/xTY9gja0Dq3HiwQF8LaCRTXxZKRutelT44=" crossorigin="anonymous"></script> <script src="https://code.jquery.com/ui/1.12.1/jquery-ui.min.js"></script> </head> <body id="body" data-lang="en" class="haveSidebar en "> <script> const Laravel = { csrfToken: "TBjMRcqZMlBKqq8FfQbZoiVreBacEvyIeMS7AQMI", locale: "en" }; </script> <header id="header" class="header"> <nav class="nav" role="navigation"> <div class="nav-menu--top"> <div class="nav-logo--en"> <a aria-label="The Canadian Encyclopedia homepage" class="nav-logo__link" href="https://www.thecanadianencyclopedia.ca/en"> </a> </div> <button class="nav-icon--search js-search-tab" aria-label="Search in Canadian Encyclopedia"></button> <button id="nav-menu-toggle" class="nav-icon--hamburger-menu" aria-label="open main navigation"> <div class="icon-container"> <span class="top"></span> <span class="middle"></span> <span class="bottom"></span> </div> </button> </div> <div id="nav-mid" class="nav-menu--mid"> <section class="nav-menu--mid__item dropdown-group"> <button class="nav-menu--mid__item__heading dropdown-toggler" aria-haspopup="true" > <span class="top-level-link"> topics </span> <span class="toggle-icon"></span> </button> <div class="sub-menu--topics dropdown-body" aria-expanded="false"> <div class="sub-menu__container"> <section id="People" class="sub-menu__wrapper nav-menu--mid__item dropdown-group"> <div class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/browse/people" aria-label="Browse topic: People"> People </a> <button class="sub-menu__button dropdown-toggler accordion-toggler" aria-label="expand section People"> <span class="toggle-icon"></span> </button> </div> <ul class="sub-menu nested dropdown-body" aria-expanded="false"> </ul> </section> <section id="Places" class="sub-menu__wrapper nav-menu--mid__item dropdown-group"> <div class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/browse/places" aria-label="Browse topic: Places"> Places </a> <button class="sub-menu__button dropdown-toggler accordion-toggler" aria-label="expand section Places"> <span class="toggle-icon"></span> </button> </div> <ul class="sub-menu nested dropdown-body" aria-expanded="false"> </ul> </section> <section id="Things" class="sub-menu__wrapper nav-menu--mid__item dropdown-group"> <div class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/browse/things" aria-label="Browse topic: Things"> Things </a> <button class="sub-menu__button dropdown-toggler accordion-toggler" aria-label="expand section Things"> <span class="toggle-icon"></span> </button> </div> <ul class="sub-menu nested dropdown-body" aria-expanded="false"> </ul> </section> </div> </div> </section> <section class="nav-menu--mid__item dropdown-group"> <button class="nav-menu--mid__item__heading dropdown-toggler" aria-haspopup="true" > <span class="top-level-link"> Collections </span> <span class="toggle-icon"></span> </button> <div class="sub-menu--cta dropdown-body" aria-expanded="false"> <div class="sub-menu__container"> <ul class="sub-menu__wrapper nav-menu--mid__item dropdown-group"> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/collections"> All Collections </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/the-memory-project"> The Memory Project </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/cdc"> Canada During Covid </a> </li> </ul> <div class="sub-menu__image"> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/950x479/MemoryProject/TCECollectionArchiveCollage-2.png" data-src="https://d14fiu1i7ba797.cloudfront.net/950x479/MemoryProject/TCECollectionArchiveCollage-2.png" alt="" /> </div> <a href="https://www.thecanadianencyclopedia.ca/en/collection/the-memory-project-archive-collection" class="sub-menu__image__link"> Explore the The Memory Project Archive collection <span class="link-arrow"></span> </a> </div> </div> </div> </section> <section class="nav-menu--mid__item dropdown-group"> <button class="nav-menu--mid__item__heading dropdown-toggler" aria-haspopup="true" > <span class="top-level-link"> Educators </span> <span class="toggle-icon"></span> </button> <div class="sub-menu--cta dropdown-body" aria-expanded="false"> <div class="sub-menu__container"> <ul class="sub-menu__wrapper nav-menu--mid__item dropdown-group"> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/educators#education-guides"> Education Guides </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/educators#primary-sources"> Primary Sources </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/educators#interactive-resources"> Interactive Resources </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/quiz/cc-quiz-citizenship"> The Challenge </a> </li> </ul> <div class="sub-menu__image"> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/950x579/cc_43.png" data-src="https://d14fiu1i7ba797.cloudfront.net/950x579/cc_43.png" alt="" /> </div> <a href="https://www.thecanadianencyclopedia.ca/en/collection/citizenship" class="sub-menu__image__link"> Explore the Citizenship Challenge collection <span class="link-arrow"></span> </a> </div> </div> </div> </section> <section class="nav-menu--mid__item dropdown-group"> <button class="nav-menu--mid__item__heading dropdown-toggler" aria-haspopup="true" > <span class="top-level-link"> Timelines </span> <span class="toggle-icon"></span> </button> <div class="sub-menu--cta dropdown-body" aria-expanded="false"> <div class="sub-menu__container"> <ul class="sub-menu__wrapper nav-menu--mid__item dropdown-group"> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/timelines"> All Timelines </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/timelines/people"> People </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/timelines/places"> Places </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/timelines/things"> Things </a> </li> </ul> <div class="sub-menu__image"> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/950x579/media/media/49936680-e046-4501-8593-85b8bc94d071.jpg" data-src="https://d14fiu1i7ba797.cloudfront.net/950x579/media/media/49936680-e046-4501-8593-85b8bc94d071.jpg" alt=""/> </div> <a href="https://www.thecanadianencyclopedia.ca/en/timeline/lgbtq2" class="sub-menu__image__link">Explore the 2SLGBTQ+ History timeline <span class="link-arrow" aria-hidden="true"></span> </a> </div> </div> </div> </section> <section class="nav-menu--mid__item dropdown-group"> <button class="nav-menu--mid__item__heading dropdown-toggler" aria-haspopup="true" > <span class="top-level-link"> Quizzes </span> <span class="toggle-icon"></span> </button> <div class="sub-menu--cta dropdown-body" aria-expanded="false"> <div class="sub-menu__container"> <ul class="sub-menu__wrapper nav-menu--mid__item dropdown-group"> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/quizzes"> Quizzes </a> </li> <li class="sub-menu__heading"> <a href="https://www.thecanadianencyclopedia.ca/en/quiz/cc-quiz-citizenship"> The Challenge </a> </li> </ul> <div class="sub-menu__image"> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/950x579/can_quiz.png" data-src="https://d14fiu1i7ba797.cloudfront.net/950x579/can_quiz.png" alt=""/> </div> <a href="https://www.thecanadianencyclopedia.ca/en/quiz/timeline" class="sub-menu__image__link">Timeline <span class="link-arrow"></span> </a> </div> </div> </div> </section> <button class="nav-icon--search js-search-tab" aria-label="Search in Canadian Encyclopedia"></button> </div> <ul id="nav-low" class="nav-menu--low"> <li class="nav-menu--low__item"> <a class="articleNavPopupEvent btn-login nav-link" href="#" data-target="popup-login" data-toggle="modal" aria-haspopup="dialog"> Sign in </a> </li> <li class="nav-menu--low__item"> <a class="nav-link help-btn" href="/en/about/faq" id="nav-help"> <i class="i-help"></i>Help </a> </li> <li class="nav-menu--low__item--btn"> <a class="b b-sm b-monochrome b-icon-heart" href="https://www.thecanadianencyclopedia.ca/en/donate" target="_blank"> Donate </a> </li> <li class="nav-menu--low__item--lang-toggle"> <a class="nav-link" href="https://www.thecanadianencyclopedia.ca/fr/article/neuroscience" aria-label="Toggle site language">fr</a> </li> </ul> </nav> </header> <div id="search" class="searchContainer" > <div class="searchPanel"> <div class="searchPanel-head"> <img src="https://www.thecanadianencyclopedia.ca/images/logo-tce.svg?v=96550f8de20f4537bc4ad37b01dff7f1" alt="The Canadian Encyclopedia"> <h2 class="searchPanel-title">Search The Canadian Encyclopedia</h2> </div> <div class="searchPanel-input-wrap"> <form id="form-search" name="form-search" method="get" data-action="https://www.thecanadianencyclopedia.ca/en/search"> <input data-getsearchroute="https://www.thecanadianencyclopedia.ca/en/json/search" name="query" type="text" class="input-search input-xl input-fullSpan searchPanel-input" id="inputSearch" /> <button class="searchPanel-searchBtn"> <img src="https://www.thecanadianencyclopedia.ca/images/i-search.svg?v=a1e99ea3dd07831352832b022428584e" alt="Search"> </button> <button class="searchPanel-exit"> <img src="https://www.thecanadianencyclopedia.ca/images/i-exit.svg?v=2601c4155c28e62e1eb2ebc7de9bd1ee" alt="Exit search"> </button> </form> <ul class="search-results"> </ul> </div> <p aria-hidden="true" role="alert" class="searchPanel-errorMsg">Enter your search term</p> <div></div> </div> </div> <div id="popup-login" class="modal"> <div class="popup-wrapper"> <div class="popup-content"> <div class="popup-connect"> <h2 class="popup-title">Login</h2> <form method="POST" action="https://www.thecanadianencyclopedia.ca/en/members" accept-charset="UTF-8" id="tceLogin" class="form"><input name="_token" type="hidden" value="TBjMRcqZMlBKqq8FfQbZoiVreBacEvyIeMS7AQMI"> <div class="input-group"> <div class="input-group"> <input class="form-login-email form-control input input-fullSpan" placeholder="Email" name="email" type="email" value=""> <span class="glyphicon glyphicon-envelope form-control-feedback"></span> </div> <div class="input-group"> <input class="form-login-password form-control form-password input input-fullSpan" placeholder="Password" name="password" type="password" value=""> <span class="glyphicon glyphicon-envelope form-control-feedback"></span> </div> </div> <div class="js-login-error error"></div> <div class="popup-login-option"> <div class="popup-helpers-container"> <div class="form-remember"> <input name="remember_me" type="checkbox" value="1"> <label for="remember_me">Remember me</label> </div> <div class="form-forgot-pass"> <a href="https://www.thecanadianencyclopedia.ca/auth/password/reset" class="b-link">I forgot my password</a><br> </div> </div> <input class="b b-md b-invert btn-login-submit" type="submit" value="Submit"> <p class="text-line">or</p> <a href="https://www.thecanadianencyclopedia.ca/auth/facebook" class="btn btn-facebook"><img src="https://www.thecanadianencyclopedia.ca/images/i-facebook-white.svg?v=3decbb62f20c49bf2a86d220cbf213d8" alt="Share on Facebook" title="Share on Facebook"> Login With Facebook</a> <a href="https://www.thecanadianencyclopedia.ca/auth/twitter" class="btn btn-twitter"><img src="https://www.thecanadianencyclopedia.ca/images/i-twitter-x-white.svg?v=762dd4e2da316070386a19085588add2" alt="Share on X" title="Share on X"> Login With X</a> </div> </form> </div> <div class="popup-body login-popup-body"> <a class="popup-close" data-dismiss="modal"></a> <h3>Why sign up?</h3> <p>Signing up enhances your TCE experience with the ability to save items to your personal reading list, and access the interactive map.</p> <a href="https://www.thecanadianencyclopedia.ca/en/members/register" class="b b-md b-primary">Create Account </a> </div> </div> </div> </div> <div class="sidebarGrid"> <div class="sidebarContainerDesktop"> <aside class="sb-anchor sb-articles"> <sidebar sb-header="Content" sb-subtitle-show="true"> <div class="sb-group-name sb-article-index">Index</div> <div class="sb-hidden-anchor"></div> <div class="sb-group-name">Resources</div> <div class="sb-subcat sb-group"> <div class="sb-tab sb-tab--separator"><a href="#recommended">Recommended</a></div> </div> </sidebar> </aside> </div> <div class="content-container-desktop"> <input class="js-picture-information" type="hidden" value="[]"> <main class="main-grid"> <article class="article main-grid__content"> <input type="hidden" id="isUserLogin" name="loginuser" value=""> <aside id="articlewidgets" class="article-widgets"> <input type="checkbox" class="menu-open" name="menu-open" id="menu-open"/> <label class="widget-open-button" for="menu-open"> <span class="dot--1"></span> <span class="dot--2"></span> <span class="dot--3"></span> </label> <button data-target="popup-citation" data-toggle="modal" class="widget-item articleNavPopupEvent"> <div class="widget-item__icon--citation"></div> <span class="widget-item__title">Citation</span> </button> <button data-toggle="modal" class="widget-item articleNavPopupEvent js-edit-submit"> <div class="widget-item__icon--edit"></div> <span class="widget-item__title">Suggest an Edit</span> </button> <button data-target="popup-translation" data-toggle="modal" class="widget-item articleNavPopupEvent js-translation"> <div class="widget-item__icon--translate"></div> <span class="widget-item__title">Request Translation</span> </button> <button id="popup-translation__original" class="widget-item hide-widget"> <div class="widget-item__icon--revert"></div> <span class="widget-item__title">Revert to original</span> </button> </aside> <div id="popup-edit" class="modal"> <div class="popup-wrapper"> <div class="popup-content"> <div class="popup-title"> <div>Suggest an Edit</div> <div> <a class="popup-close" data-dismiss="modal"></a> </div> </div> <div class="popup-body"> <div class="contact-form edit-form"> <div class="form-error-list"> <ul></ul> </div> <form method="POST" action="https://www.thecanadianencyclopedia.ca/en/submitArticle" accept-charset="UTF-8" class="form form-submit-edit form-main"><input name="_token" type="hidden" value="TBjMRcqZMlBKqq8FfQbZoiVreBacEvyIeMS7AQMI"> <input name="aid" type="hidden" value="78860"> <input name="edit" type="hidden" value="true"> <div class="input-group "> <label for="description">Enter your suggested edit(s) to this article in the form field below</label> <textarea id="text-editor" class="form-control input input-fullSpan" placeholder="Write your message here..." name="description" cols="50" rows="10"></textarea> </div> <div class="btn-container"> <div> <input class="disabled js-submitedit-submit b b-md b-invert" type="submit" value="Submit"> </div> <div> <a href="#" data-dismiss="modal" class="js-popup-close b-link">Cancel</a> </div> </div> </form> </div> </div> </div> </div> </div> <div id="popup-citation" class="modal"> <div class="popup-wrapper"> <div class="popup-content"> <div class="popup-title"> <div> Citation </div> <div> <a class="popup-close" data-dismiss="modal"></a> </div> </div> <div class="popup-body"> <ul class="list"> <li> <ul class="citation-reference"> <input id="copy1" type="text" value="MLA 8TH EDITION |聽Vanderwolf, C.H.. "Neuroscience". <em>The Canadian Encyclopedia</em>, 12 September 2024, <em>Historica Canada</em>. www.thecanadianencyclopedia.ca/en/article/neuroscience. Accessed 28 November 2024."> <li class="title">MLA 8TH EDITION</li> <li class="description">Vanderwolf, C.H.. "Neuroscience". <em>The Canadian Encyclopedia</em>, 12 September 2024, <em>Historica Canada</em>. www.thecanadianencyclopedia.ca/en/article/neuroscience. Accessed 28 November 2024.</li> <li class="link"><a data-text="Vanderwolf, C.H.. "Neuroscience". <em>The Canadian Encyclopedia</em>, 12 September 2024, <em>Historica Canada</em>. www.thecanadianencyclopedia.ca/en/article/neuroscience. Accessed 28 November 2024." href="#" class="js-copy-clipboard b b-md b-invert b-modal-copy">Copy</a></li> </ul> </li> <li> <ul class="citation-reference"> <input id="copy2" type="text" value="APA 6TH EDITION |聽Vanderwolf, C. (2024). Neuroscience. In <em>The Canadian Encyclopedia</em>. Retrieved from https://www.thecanadianencyclopedia.ca/en/article/neuroscience"> <li class="title">APA 6TH EDITION</li> <li class="description">Vanderwolf, C. (2024). Neuroscience. In <em>The Canadian Encyclopedia</em>. Retrieved from https://www.thecanadianencyclopedia.ca/en/article/neuroscience</li> <li class="link"><a data-text="Vanderwolf, C. (2024). Neuroscience. In <em>The Canadian Encyclopedia</em>. Retrieved from https://www.thecanadianencyclopedia.ca/en/article/neuroscience" href="#" class="js-copy-clipboard b b-md b-invert b-modal-copy">Copy</a></li> </ul> </li> <li> <ul class="citation-reference"> <input id="copy3" type="text" value="CHICAGO 17TH EDITION | Vanderwolf, C.H.. "Neuroscience." <em>The Canadian Encyclopedia</em>. Historica Canada. Article published February 07, 2006; Last Edited September 12, 2024."> <li class="title">CHICAGO 17TH EDITION</li> <li class="description">Vanderwolf, C.H.. "Neuroscience." <em>The Canadian Encyclopedia</em>. Historica Canada. Article published February 07, 2006; Last Edited September 12, 2024.</li> <li class="link"><a data-text="Vanderwolf, C.H.. "Neuroscience." <em>The Canadian Encyclopedia</em>. Historica Canada. Article published February 07, 2006; Last Edited September 12, 2024." href="#" class="js-copy-clipboard b b-md b-invert b-modal-copy">Copy</a></li> </ul> </li> <li> <ul class="citation-reference"> <input id="copy4" type="text" value="TURABIAN 8TH EDITION | <em>The Canadian Encyclopedia</em>, s.v. "Neuroscience," by C.H. Vanderwolf, Accessed November 28, 2024, https://www.thecanadianencyclopedia.ca/en/article/neuroscience"> <li class="title">TURABIAN 8TH EDITION</li> <li class="description"><em>The Canadian Encyclopedia</em>, s.v. "Neuroscience," by C.H. Vanderwolf, Accessed November 28, 2024, https://www.thecanadianencyclopedia.ca/en/article/neuroscience</li> <li class="link"><a data-text="<em>The Canadian Encyclopedia</em>, s.v. "Neuroscience," by C.H. Vanderwolf, Accessed November 28, 2024, https://www.thecanadianencyclopedia.ca/en/article/neuroscience" href="#" class="js-copy-clipboard b b-md b-invert b-modal-copy">Copy</a></li> </ul> </li> </ul> </div> </div> </div> </div> <div id="popup-share" class="modal"> <div class="popup-wrapper"> <div class="popup-content"> <div class="popup-title"> <div>Share</div> <div> <a class="popup-close" data-dismiss="modal"></a> </div> </div> <div class="popup-body"> <ul class="list"> <li><a href="https://www.facebook.com/sharer/sharer.php?u=https://www.thecanadianencyclopedia.ca/en/article/neuroscience" target="_blank"><img class="popup-share-facebook" src="https://www.thecanadianencyclopedia.ca/images/i-facebook.svg?v=7fb8b09ec62e4de8848271514745bd2a" alt="Share on Facebook" title="Share on Facebook"> Share on Facebook</a></li> <li><a href="https://twitter.com/intent/tweet?url=https://www.thecanadianencyclopedia.ca/en/article/neuroscience" target="_blank"><img src="https://www.thecanadianencyclopedia.ca/images/i-twitter-x.svg?v=deacd0b3626a2053dbdc3843f1e13d44" alt="Share on X" title="Share on X"> Share on X</a></li> <li><a href="mailto:?body=https://www.thecanadianencyclopedia.ca/en/article/neuroscience"><img src="https://www.thecanadianencyclopedia.ca/images/i-email.svg?v=c763267e8d2e1fab14f745261b8a8c7a" alt="Share by Email" title="Share by Email"> Share by Email</a></li> <li class="share-classroom"><a target="_blank" href="https://classroom.google.com/share?url='https://www.thecanadianencyclopedia.ca/en/article/neuroscience'" > <img src="https://www.thecanadianencyclopedia.ca/images/i-google.svg?v=657db3c0413ddd5da4c7a6ee570546f7" alt="Share on Google Classroom" title="Share on Google Classroom" style="filter: grayscale(100%);"> Share on Google Classroom </a> <g:sharetoclassroom url="https://www.thecanadianencyclopedia.ca/en/article/neuroscience" size="20"></g:sharetoclassroom> </li> <div id="widget-div"></div> </ul> </div> </div> </div> </div> <div id="popup-thankyou" class="modal"> <div class="popup-wrapper"> <div class="popup-content"> <div class="popup-body"> <div class="contact-form feedback-form"> <h2>Thank you for your submission</h2> <p>Our team will be reviewing your submission <br/>and get back to you with any further questions.</p> <p>Thanks for contributing to The Canadian Encyclopedia.</p> <a href="#" data-dismiss="modal" class="js-popup-close b-link">Close</a> </div> </div> </div> </div> </div> <div class="article-heading"> <p class="article-heading__type">Article</p> <h1 id="article-title" class="article-heading__title">Neuroscience</h1> </div> <div class="article-info" > <div class="article-info__column"> <p class="info-text">Article by <span class="info-text__value"> <a href='https://www.thecanadianencyclopedia.ca/en/author/c-h-vanderwolf' class='b-link'>C.H. Vanderwolf</a> </span> </p> <p class="info-text">Updated by <span class="info-text__value"> The Canadian Encyclopedia </span> </p> </div> <div class="article-info__column"> <p class="info-text">Published Online <span class="info-text__value--date"> February 7, 2006 </span> </p> <p class="info-text">Last Edited <span class="info-text__value--date"> September 12, 2024 </span> </p> </div> </div> <div class="article-toolbar"> <button data-target="popup-share" data-toggle="modal" class="article-toolbar__item articleNavPopupEvent"> <div class="article-toolbar__item__icon--share"></div> <span class="article-toolbar__item__title">Share</span> </button> <button id="js-print" class="article-toolbar__item"> <div class="article-toolbar__item__icon--print"></div> <span class="article-toolbar__item__title">Print</span> </button> </div> <div class="article-content"> <div class="article-content__summary"> <p>Neuroscience is the study of the structure and function of the nervous system. It includes a number of subfields such as neuroanatomy, neurochemistry, neuropharmacology, neurophysiology and the study of brain mechanisms in behaviour (neuroethology, neuropsychology, psychobiology and psychopharmacology). In addition, the medical fields of neurology, neurosurgery and <a href="https://www.thecanadianencyclopedia.ca/article/psychiatry/">psychiatry</a> are closely related to that of neuroscience.</p> </div> <div class="article-content-body"> <div id="article-content-def"> <iframe width="640" height="360" src="https://www.youtube.com/embed/pUOG2g4hj8s?si=FCshvjwIY3ylUazi?rel=0" title="YouTube video player" frameborder="0" allow="accelerometer; autoplay; clipboard-write; encrypted-media; gyroscope; picture-in-picture; web-share" referrerpolicy="strict-origin-when-cross-origin" allowfullscreen=""></iframe> <br> <h3>Synaptic Transmission</h3> <p>Anatomical investigations of the nervous system have revealed the existence of specialized cells or neurons, which occur in vast numbers, in a great variety of shapes, sizes and functions, and form interconnected systems of enormous complexity. A British scientist, C.S. Sherrington, first understood that one neuron can influence the activity of a second neuron primarily via a specialized area of contact, which he referred to as a synapse. Subsequent work has revealed that synaptic transmission is usually the result of the release of a chemical substance (a neurohumor or neurotransmitter) which acts on specialized macromolecules (receptors) located in the membrane of the recipient neuron. It is of historical interest that some essential features of this concept were originally proposed in two papers published by a Canadian, F.H. Scott, in 1905 and 1906. However, Scott's ideas were too far in advance of the times and were ignored. The chemical nature of synaptic transmission did not become generally accepted until several decades later.</p> <p>Modern research has shown that there are many chemically distinctive neurotransmitters. As a result, it is possible to alter the operation of specific types of synapses by the administration of drugs that alter the release or the effect of specific neurotransmitters. Great practical importance attaches to this fact since it makes possible the pharmacological treatment of disorders of the nervous system. It also provides the basis for the widespread non-medical use of drugs that alter various aspects of the brain's function in a selective manner.</p> <h3>Development of Neuroscience in Canada</h3> <p>The establishment of the Montreal Neurological Institute (MNI) at <a href="https://www.thecanadianencyclopedia.ca/en/article/mcgill-university">McGill University</a> in 1934 with <a href="https://www.thecanadianencyclopedia.ca/en/article/wilder-penfield">Wilder Penfield</a> as its first director had a major effect on the development of neuroscience in Canada. The MNI, together with related departments at McGill and <a href="https://www.thecanadianencyclopedia.ca/en/article/universite-de-montreal">Universit茅 de Montr茅al</a> made <a href="https://www.thecanadianencyclopedia.ca/en/article/montreal">Montreal</a> known worldwide for research and teaching in neuroscience and for the study and treatment of disorders of the nervous system. Scientists and physicians trained in Montreal, and subsequently established eminent careers elsewhere in Canada and the world. For example, <a href="https://www.thecanadianencyclopedia.ca/en/article/david-h-hubel">David H. Hubel</a>, a Canadian-born US citizen who was educated at McGill University and subsequently took up a position at Harvard University in Boston, was awarded the Nobel Prize in 1981 for his studies on brain mechanisms in vision. (<em>See also</em> <a href="https://www.thecanadianencyclopedia.ca/en/article/nobel-prizes">Nobel Prizes and Canada</a>.)</p> <p>The MNI became especially well known for the advances made by Wilder Penfield and Herbert Jasper in the diagnosis, classification and treatment of epilepsy. The identification of the role of gamma-aminobutyric acid (GABA), now believed to be the major inhibitory transmitter in the brain with a role in conditions as diverse as epilepsy and in the effect of anti-anxiety drugs, was made at the MNI by A.W. Bazemore, <a href="https://www.thecanadianencyclopedia.ca/en/article/kenneth-allan-caldwell-elliott">Kenneth Allan Caldwell Elliott</a> and E. Florey in 1957. The discoveries that Parkinson's disease was due to a deficiency of dopamine, an important neurotransmitter, and that the symptoms of the disease could be dramatically reversed by treatment with L-dopa, a compound from which the brain can manufacture dopamine, were made almost simultaneously in 1960-62 by Andr茅 Barbeau, G.F. Murphy and <a href="https://www.thecanadianencyclopedia.ca/en/article/theodore-lionel-sourkes">Theodore Lionel Sourkes</a> in Montreal and by O. Hornykiewicz and his colleagues in Vienna.</p> <p>Work by H. Jasper, K. Krnjevic, H. MacIntosh and <a href="https://www.thecanadianencyclopedia.ca/article/judah-hirsch-quastel/">Juda Hirsch Quastel</a> of McGill University and J. Szerb of <a href="https://www.thecanadianencyclopedia.ca/en/article/dalhousie-university">Dalhousie University</a> provided much of the basis of our present understanding of the action of another neurotransmitter, acetylcholine, in the brain. This work has recently assumed considerable practical importance as a result of the demonstration by British and American investigators that Alzheimer's disease is associated with a severe loss of acetylcholine-containing neurons in the brain.</p> <h3>Brain Behaviour Research</h3> <p>The establishment of brain behaviour research in Canada was due, in large part, to <a href="https://www.thecanadianencyclopedia.ca/en/article/donald-olding-hebb">Donald Olding Hebb</a> of <a href="https://www.thecanadianencyclopedia.ca/en/article/mcgill-university">McGill University</a>. Work in Hebb's laboratory demonstrated the important role played by ordinary environmental stimulation in the development and maintenance of normal brain function. One of Hebb's students, <a href="https://www.thecanadianencyclopedia.ca/en/article/brenda-milner">Brenda Milner</a>, carried out a series of studies at the MNI which demonstrated that localized injuries in different parts of the cerebral cortex produced quite specific impairments in psychological functions. In 1953 James Olds and Peter M. Milner, working in Hebb's laboratory, discovered that animals would voluntarily deliver electrical stimulation to certain parts of their own brain when enabled to do so by means of an implanted electrode. It is widely believed that this phenomenon establishes that activity of specific brain circuits is related to motivation and pleasure.</p> <p>Research in neuroscience is undergoing an explosive development everywhere in the developed world. Every large Canadian university has programs in neuroscience fields in several academic departments. New technology such as computerized tomography, magnetic resonance imaging and positron-emission tomography now permit noninvasive investigation of the living human nervous system at a level of detail which was unimaginable a few years ago. It is reasonable to expect great improvements in our understanding of the nervous system and our ability to treat diseases of the nervous system in the coming decades.</p> </div> </div> <div id="def" class="article-content-body hide"></div> </div> <div id="article-definitions"> <definitions route="https://www.thecanadianencyclopedia.ca/en/json/define" id="neuroscience"></definitions> </div> <div class="article-widget widget-adsense"> <script async src="//pagead2.googlesyndication.com/pagead/js/adsbygoogle.js"></script> <ins class="adsbygoogle" style="display:inline-block;width:468px;height:60px" data-ad-client="ca-pub-3964766461879185" data-ad-slot="1946780682"></ins> <script> (adsbygoogle = window.adsbygoogle || []).push({}); </script> </div> </article> </main> <div id="modal-overlay" class="overlay"></div> <div id="article-image" class="article-img-modal" role="dialog" aria-labelledby="article-modal-title"> <button class="article-img-modal__close" aria-label="close image modal"></button> <div id="js-article-modal-image" class="article-img-modal__container"> <div class="image-container"> <img class="article-img-modal__img" src="" alt=""> </div> <div class="text-container"> <span role="heading" aria-level="3" id="article-modal-title" class="article-img-modal__title"></span> <span class="article-img-modal__description"></span> <span class="article-img-modal__creator"></span> </div> </div> </div> </div> </div> <div class="main-grid article-related"> <ul class="article-tags-list main-grid__content"> <li class="article-tags-list__item"> <a class="article-tags-list__item__link" href="https://www.thecanadianencyclopedia.ca/en/search?query=neurology&tag=neurology">neurology</a> </li> <li class="article-tags-list__item"> <a class="article-tags-list__item__link" href="https://www.thecanadianencyclopedia.ca/en/search?query=science&tag=science">science</a> </li> <li class="article-tags-list__item"> <a class="article-tags-list__item__link" href="https://www.thecanadianencyclopedia.ca/en/search?query=research&tag=research">research</a> </li> </ul> <div class="article-related-two-column main-grid__content--wide"> <h3 class="article-related-two-column__title">External Links</h3> <ul class="article-related-two-column__list"> <li class="article-related-two-column__list__item"><p><a href="http://www.epilepsy.ca/"><strong>Epilepsy Canada </strong>An extensive information source about epilepsy for patients and care givers.</a></p></li> <li class="article-related-two-column__list__item"><p><a href="http://www.michaeljfox.org/"><strong>The Michael J. Fox Foundation for Parkinson's Research </strong>This website describes the nature of Parkinson's disease and current research initiatives. Also provides a bio of Michael J. Fox.</a></p></li> <li class="article-related-two-column__list__item"><p><a href="http://www.parkinson.ca/"><strong>Parkinson Canada </strong>The website for the Parkinson Society Canada. This website focuses on the nature of Parkinson's disease, treatment, care, and related research projects.</a></p></li> <li class="article-related-two-column__list__item"><p><a href="http://www.alzheimer.ca/"><strong>Alzheimer Society of Canada </strong>The Alzheimer Society provides support, information and education to people with Alzheimer Disease, families, physicians, and health-care providers.</a></p></li> <li class="article-related-two-column__list__item"><p><a href="https://can-acn.org/"><strong>Canadian Association of Neuroscience</strong> Access the official website of the Canadian Association of Neuroscience.</a></p></li> </ul> </div> <!-- Related Collections --> <div class="callouts-layout main-grid__content--full"> <h3 id="recommended" class="callouts-layout__title">Associated Collections</h3> <div class="callouts-layout__grid"> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/collection/health-and-healthcare-in-canada" > Health and Healthcare in Canada </a> </h6> <span class="callout-item__category">collection</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/media/350b6e59-1083-4f86-b24c-c0a1608e2fa0.jpg" class="image-container__image" alt="Banting and Best Laboratory" loading="lazy" /> </div> </div> </div> </div> <!-- Related Articles --> <div class="callouts-layout main-grid__content--full"> <h3 id="recommended" class="callouts-layout__title">Recommended</h3> <div class="callouts-layout__grid"> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/wilder-penfield" > Wilder Penfield </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/media/0a9f70e1-8b68-43d1-886b-006215336887.jpg" class="image-container__image" alt="Wilder Penfield" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/brenda-milner" > Brenda Milner </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/single_use_images/B_Milner_1956.jpg" class="image-container__image" alt="Photo of Brenda Milner in 1956" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/psychiatry" > Psychiatry </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img loading="lazy" src="https://www.thecanadianencyclopedia.ca/images/tce_placeholder.jpg?v=e9dca980c9bdb3aa11e832e7ea94f5d9" alt="" class="image-container__image" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/donald-olding-hebb" > Donald Olding Hebb </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/donaldoldinghebb/donaldhebb.jpg" class="image-container__image" alt="Donald Olding Hebb" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/david-h-hubel" > David H. Hubel </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/davidhubel/davidhubel.jpg" class="image-container__image" alt="David H. Hubel" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/theodore-lionel-sourkes" > Theodore Lionel Sourkes </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img loading="lazy" src="https://www.thecanadianencyclopedia.ca/images/tce_placeholder.jpg?v=e9dca980c9bdb3aa11e832e7ea94f5d9" alt="" class="image-container__image" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/mcgill-university" > McGill University </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/media/2b7f1364-6c18-4b8d-a208-0aac26b4f5fe.jpg" class="image-container__image" alt="McGill University" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/universite-de-montreal" > Universit茅 de Montr茅al </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/media/4ed313c7-50c9-4739-af34-b1f58675cf71.jpg" class="image-container__image" alt="Universit茅 de Montr茅al" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/article/nobel-prizes" > Nobel Prizes and Canada </a> </h6> <span class="callout-item__category">Article</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/media/352401f2-9e16-4265-9c53-189828385881.jpg" class="image-container__image" alt="Gerhard Herzberg, physicist, Nobel laureate " loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/collection/health-and-healthcare-in-canada" > Health and Healthcare in Canada </a> </h6> <span class="callout-item__category">collection</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/media/350b6e59-1083-4f86-b24c-c0a1608e2fa0.jpg" class="image-container__image" alt="Banting and Best Laboratory" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/quiz/cc-quiz-science-and-innovation" > Science and Innovation in Canada </a> </h6> <span class="callout-item__category">Quiz</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/quiz_images/Science-Innovation-1.jpg" class="image-container__image" alt="" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/quiz/canadian-fact-or-fiction-notable-figures-and-innovations" > Canadian Fact or Fiction: Notable Figures and Innovations </a> </h6> <span class="callout-item__category">Quiz</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/media/home-page-images/Fact-or-Fiction-quiz.jpg" class="image-container__image" alt="" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/quiz/cc-quiz-women-in-stem" > Women in STEM in Canada </a> </h6> <span class="callout-item__category">Quiz</span> </div> <div class="image-container"> <img src="https://d14fiu1i7ba797.cloudfront.net/340x207/stem_redo_textless2_crop.png" class="image-container__image" alt="" loading="lazy" /> </div> </div> <div class="callout-item"> <div class="callout-item__container"> <h6 class="callout-item__title"> <a class="callout-item__link" href="https://www.thecanadianencyclopedia.ca/en/timeline/history-of-health-and-healthcare-in-canada" > History of Health and Healthcare in Canada </a> </h6> <span class="callout-item__category">timeline</span> </div> <div class="image-container"> <img loading="lazy" src="" alt="" class="image-container__image" /> </div> </div> </div> </div> </div> </div> </div> <footer id="footer"> <div class="full-width footer-first-part"> <div class="wrap--narrow"> <div class="footer-information grid--col2"> <div class="divider"> <h6>Updates? Omissions? Article suggestions?</h6> <a href="https://www.thecanadianencyclopedia.ca/en/contact" class="b b-md b-primary">We want to hear from you</a> </div> <div> <h6>Did you like your experience?</h6> <a href="https://www.cognitoforms.com/HistoricaCanada1/TheCanadianEncyclopediaUserSurvey" target="_blank" class="b b-md b-primary">Take our survey</a> </div> </div> </div> </div> <div class="full-width full-width-grey footer-second-part"> <div class="wrap--narrow footer-sub-first"> <div class="footer-utilities grid--col3"> <div class="randomize-12"> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>62 375</b> images</div> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>25 520</b> articles</div> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>64</b> collections</div> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>4 172</b> primary sources</div> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>46</b> quizzes</div> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>73</b> timelines</div> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>83</b> studyguides</div> <div class="footer-articles-count randomBlock" style="display: none;">We have <b>5 169</b> authors</div> </div> <div class="divider uppercase text-xsmall footer-social "></div> <div class="uppercase footer-social"> <span>Follow us elsewhere</span> <ul> <li><a href="https://twitter.com/HistoricaCanada" class="footer-social-twitter" target="_blank"><i class="i i-invert i-twitter"></i>Twitter</a></li> <li><a href="https://www.facebook.com/TheCanadianEncyclopedia" class="footer-social-facebook" target="_blank"><i class="i i-invert i-facebook"></i>Facebook</a></li> <li><a href="https://www.youtube.com/channel/UCtagGjDO8X1rBROhmSQXZsA" class="footer-social-youtube" target="_blank"><i class="i i-invert i-youtube"></i>Youtube</a></li> </ul> </div> </div> </div> <div class="wrap--narrow container-grey top-divider-footerlinks footer-sub-second"> <div class="footer-link"> <ul class="list list-inline list-background-invert"> <li><a href="https://www.thecanadianencyclopedia.ca/en/about">About</a></li> <li><a href="https://www.thecanadianencyclopedia.ca/en/contributors">Contribute</a></li> <li><a href="https://www.thecanadianencyclopedia.ca/en/contact">Contact</a></li> <li><a href="https://www.thecanadianencyclopedia.ca/en/sitemap">Sitemap</a></li> <li><a href="https://www.thecanadianencyclopedia.ca/en/privacy-policy">privacy policy</a></li> <li><a href="https://www.thecanadianencyclopedia.ca/images/pdf/en/aoda_2023.pdf?v=77bccd03de4270ae1fbba4cea45a2197">accessibility</a></li> </ul> </div> </div> <div class="wrap--narrow container-grey top-divider footer-sub-third"> <div class="footer-sitemap grid--col12"> <div class="footer-copyright"> <a href="/en/copyright">The Canadian Encyclopedia 漏 2024 <span class="copyright-separator-spacing">|</span> Historica Canada</a> </div> <div class="footer-canada-logos"> <ul> <li tabindex="0" class="footer-image-submenu"> <img src="https://www.thecanadianencyclopedia.ca/images/historica-canada-logo.png?v=d8609094fb353837d01f458c7a6f905b" alt="Historica Canada" title="Historica Canada" loading="lazy">聽 <ul class="external_links"> <li><a href="https://www.historicacanada.ca/" target="_blank">Historica Canada</a></li> <li><a href="/en/quiz/cc-quiz-citizenship" target="_blank">Citizenship challenge</a></li> <li><a href="https://www.historicacanada.ca/heritageminutes" target="_blank">Heritage minutes</a></li> <li><a href="http://www.thememoryproject.com/" target="_blank">The memory project</a></li> </ul> </li> <li><a href="https://www.canada.ca/en.html" target="_blank"><img src="https://www.thecanadianencyclopedia.ca/images/gov-canada.svg?v=fdda8701f4d83ce99144e160774c2965" alt="Funding by the gouvernment of Canada" title="Funding by the gouvernment of Canada" loading="lazy"></a></li> </ul> </div> </div> </div> </div> </footer> <div id="highlight-menu" style="display:none;"> <a href="#">Suggest an Edit</a> <div class="caret"></div> </div> <div id="overlayPopup"></div> <div id="definitionOverlayPopup"></div> <div id="overlay"></div> <div class="fullpageLoader"> <svg class="circular-loader" viewBox="25 25 50 50"> <circle class="loader-path" cx="50" cy="50" r="20" fill="none" stroke="#FFFFFF" stroke-width="4" /> </svg> </div> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/manifest.js?v=069add80cc340895762d04c204a6279a"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/vendor.js?v=bad4c73f65ec1c7706543fa4a9ea4830"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/app.js?v=efc40233a3006944c927943e0b78a15c"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/layout.js?v=beaebd25c9a5059050695d1edaf766de"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/header.js?v=5244f0ce969d84d34f640d056c0fe0d9"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/event-bus.js?v=5e4c1d3e6f42f0c3e6c9410dd9164894"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/image-scale.js?v=9c719b6e3da705632a5f6afd99100d6d"></script> <script src="https://cdnjs.cloudflare.com/ajax/libs/Trumbowyg/2.20.0/trumbowyg.min.js"></script> <script src="https://apis.google.com/js/platform.js" async defer></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/article.js?v=5436bd4ddb2e9afd7ac819d7ed95d6e8"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/translation.js?v=57b5749d848b8c62b44d3c07f2b37a96"></script> <script type="text/javascript"> //聽Don't have choice to let this here to use blade helper //======== Send Edit form in a Ajax Call ======== $("form.form-submit-edit").submit(function(e){ var form = $(this); var url = form.attr('action'); var articleID = $('#articleID').val(); $('.form-error-list ul').empty(); $.ajax({ type: "POST", url: url, data: form.serialize(), // serializes the form's elements. success: function(data){ //聽Trigger Close Popup //聽Show Thank You Popup + Overlay $('#popup-edit .popup-close').trigger('click'); $('#popup-thankyou').addClass('open'); $('#overlayPopup').show(); // location.reload(); }, error: function(data) { var getErrors = data.responseJSON.errors; var errorObj = $.parseJSON(getErrors); $.each(errorObj, function () { var createErrorList = '<li>'+this+'</li>'; $('.form-error-list ul').append(createErrorList); }); } }); e.preventDefault(); // avoid to execute the actual submit of the form. }); </script> <script src="https://www.thecanadianencyclopedia.ca/js/media-player.js?v=5d8651067197abf119670259f6cf27da"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/definitions.js?v=b992214c955858530020661fed1a9b67"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/lazyload.min.js?v=c1238b309b71a0d2a1f46179635b599d"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/paginations.js?v=99f58d2977f14337f4359d5e79d26cbc"></script> <script src="https://www.thecanadianencyclopedia.ca/js/jquery.bxslider.min.js?v=9069c36e9a5887c056cc2ce20af7e3c7"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/sidebar.js?v=055046a71e350608040bcb584ac4ae68"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/article.js?v=5436bd4ddb2e9afd7ac819d7ed95d6e8"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/article-ads.js?v=00da714bab3cde87b6d8ce0fb07a472c"></script> <script type="text/javascript" src="https://www.thecanadianencyclopedia.ca/js/search.js?v=c7c70246f5f7881af57df6d9ed1d3e16"></script> </body> </html>