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Air</span> </div> </a> <ul id="toc-Dispersion_through_Air-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Deposition_to_Rest" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Deposition_to_Rest"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3</span> <span>Deposition to Rest</span> </div> </a> <ul id="toc-Deposition_to_Rest-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Experimental_methods" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Experimental_methods"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Experimental methods</span> </div> </a> <ul id="toc-Experimental_methods-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Effects_on_human_health" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Effects_on_human_health"> <div 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class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" title="Table of Contents" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Aerobiology</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 26 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-26" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">26 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B9%D9%84%D9%85_%D8%A7%D9%84%D8%A3%D8%AD%D9%8A%D8%A7%D8%A1_%D8%A7%D9%84%D9%87%D9%88%D8%A7%D8%A6%D9%8A%D8%A9" title="علم الأحياء الهوائية – Arabic" lang="ar" hreflang="ar" data-title="علم الأحياء الهوائية" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AC%E0%A6%BE%E0%A6%AF%E0%A6%BC%E0%A7%81%E0%A6%9C%E0%A7%80%E0%A6%AC%E0%A6%AC%E0%A6%BF%E0%A6%A6%E0%A7%8D%E0%A6%AF%E0%A6%BE" title="বায়ুজীববিদ্যা – Bangla" lang="bn" hreflang="bn" data-title="বায়ুজীববিদ্যা" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Aerobiologia" title="Aerobiologia – Catalan" lang="ca" hreflang="ca" data-title="Aerobiologia" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Aerobiologie" title="Aerobiologie – German" lang="de" hreflang="de" data-title="Aerobiologie" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Aerobioloogia" title="Aerobioloogia – Estonian" lang="et" hreflang="et" data-title="Aerobioloogia" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Aerobiolog%C3%ADa" title="Aerobiología – Spanish" lang="es" hreflang="es" data-title="Aerobiología" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/A%C3%A9robiologie" title="Aérobiologie – French" lang="fr" hreflang="fr" data-title="Aérobiologie" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Aerobiologija" title="Aerobiologija – Croatian" lang="hr" hreflang="hr" data-title="Aerobiologija" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Aerobiologi" title="Aerobiologi – Indonesian" lang="id" hreflang="id" data-title="Aerobiologi" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Aerobiologia" title="Aerobiologia – Italian" lang="it" hreflang="it" data-title="Aerobiologia" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Aerobiol%C3%B3gia" title="Aerobiológia – Hungarian" lang="hu" hreflang="hu" data-title="Aerobiológia" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%8E%E0%B4%AF%E0%B4%B1%E0%B5%8B_%E0%B4%AC%E0%B4%AF%E0%B5%8B%E0%B4%B3%E0%B4%9C%E0%B4%BF" title="എയറോ ബയോളജി – Malayalam" lang="ml" hreflang="ml" data-title="എയറോ ബയോളജി" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Aerobiologie" title="Aerobiologie – Dutch" lang="nl" hreflang="nl" data-title="Aerobiologie" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E7%A9%BA%E4%B8%AD%E7%94%9F%E7%89%A9%E5%AD%A6" title="空中生物学 – Japanese" lang="ja" hreflang="ja" data-title="空中生物学" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Aerobiologia" title="Aerobiologia – Polish" lang="pl" hreflang="pl" data-title="Aerobiologia" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Aerobiologia" title="Aerobiologia – Portuguese" lang="pt" hreflang="pt" data-title="Aerobiologia" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Aerobiologie" title="Aerobiologie – Romanian" lang="ro" hreflang="ro" data-title="Aerobiologie" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-si mw-list-item"><a href="https://si.wikipedia.org/wiki/%E0%B7%80%E0%B7%8F%E0%B6%BA%E0%B7%80_%E0%B6%A2%E0%B7%93%E0%B7%80_%E0%B7%80%E0%B7%92%E0%B6%AF%E0%B7%8A%E2%80%8D%E0%B6%BA%E0%B7%8F%E0%B7%80" title="වායව ජීව විද්‍යාව – Sinhala" lang="si" hreflang="si" data-title="වායව ජීව විද්‍යාව" data-language-autonym="සිංහල" data-language-local-name="Sinhala" class="interlanguage-link-target"><span>සිංහල</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%90%D0%B5%D1%80%D0%BE%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%98%D0%B0" title="Аеробиологија – Serbian" lang="sr" hreflang="sr" data-title="Аеробиологија" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Aerobiologia" title="Aerobiologia – Finnish" lang="fi" hreflang="fi" data-title="Aerobiologia" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-tl mw-list-item"><a href="https://tl.wikipedia.org/wiki/Aerobiyolohiya" title="Aerobiyolohiya – Tagalog" lang="tl" hreflang="tl" data-title="Aerobiyolohiya" data-language-autonym="Tagalog" data-language-local-name="Tagalog" class="interlanguage-link-target"><span>Tagalog</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%B5%E0%AE%B3%E0%AE%BF%E0%AE%AF%E0%AF%81%E0%AE%AF%E0%AE%BF%E0%AE%B0%E0%AE%BF%E0%AE%AF%E0%AE%B2%E0%AF%8D" title="வளியுயிரியல் – Tamil" lang="ta" hreflang="ta" data-title="வளியுயிரியல்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Aerobiyoloji" title="Aerobiyoloji – Turkish" lang="tr" hreflang="tr" data-title="Aerobiyoloji" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Sinh_h%E1%BB%8Dc_kh%C3%B4ng_kh%C3%AD" title="Sinh học không khí – Vietnamese" lang="vi" hreflang="vi" data-title="Sinh học không khí" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Aerobiyolohiya" title="Aerobiyolohiya – Waray" lang="war" hreflang="war" data-title="Aerobiyolohiya" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E7%A9%BA%E6%B0%A3%E7%94%9F%E7%89%A9%E5%AD%B8" title="空氣生物學 – Cantonese" lang="yue" hreflang="yue" data-title="空氣生物學" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Not to be confused with <a href="/wiki/Bioastronautics" title="Bioastronautics">bioastronautics</a>.</div> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Spores_Trapped_From_Ambient_Air.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Spores_Trapped_From_Ambient_Air.jpg/231px-Spores_Trapped_From_Ambient_Air.jpg" decoding="async" width="231" height="171" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Spores_Trapped_From_Ambient_Air.jpg/347px-Spores_Trapped_From_Ambient_Air.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Spores_Trapped_From_Ambient_Air.jpg/462px-Spores_Trapped_From_Ambient_Air.jpg 2x" data-file-width="2080" data-file-height="1536" /></a><figcaption>Some common air-borne spores</figcaption></figure> <p><b>Aerobiology</b> (from <a href="/wiki/Ancient_Greek" title="Ancient Greek">Greek</a> <a href="https://en.wiktionary.org/wiki/%E1%BC%80%CE%AE%CF%81" class="extiw" title="wiktionary:ἀήρ">ἀήρ</a>, <i>aēr</i>, "<a href="/wiki/Air" class="mw-redirect" title="Air">air</a>"; <a href="https://en.wiktionary.org/wiki/%CE%B2%CE%AF%CE%BF%CF%82" class="extiw" title="wiktionary:βίος">βίος</a>, <i>bios</i>, "<a href="/wiki/Life" title="Life">life</a>"; and <a href="https://en.wiktionary.org/wiki/-%CE%BB%CE%BF%CE%B3%CE%AF%CE%B1" class="extiw" title="wiktionary:-λογία">-λογία</a>, <i><a href="https://en.wiktionary.org/wiki/-logia" class="extiw" title="wiktionary:-logia">-logia</a></i>) is a branch of <a href="/wiki/Biology" title="Biology">biology</a> that studies the passive transport of organic particles, such as <a href="/wiki/Bacteria" title="Bacteria">bacteria</a>, fungal <a href="/wiki/Spore" title="Spore">spores</a>, very small <a href="/wiki/Insects" class="mw-redirect" title="Insects">insects</a>, <a href="/wiki/Pollen" title="Pollen">pollen</a> grains and <a href="/wiki/Viruses" class="mw-redirect" title="Viruses">viruses</a>.<sup id="cite_ref-RSB_1-0" class="reference"><a href="#cite_note-RSB-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Aerobiologists have traditionally been involved in the measurement and reporting of airborne pollen and fungal spores as a service to those with <a href="/wiki/Allergy" title="Allergy">allergies</a>.<sup id="cite_ref-RSB_1-1" class="reference"><a href="#cite_note-RSB-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> However, aerobiology is a varied field, relating to <a href="/wiki/Environmental_science" title="Environmental science">environmental science</a>, <a href="/wiki/Plant_Science" class="mw-redirect" title="Plant Science">plant science</a>, <a href="/wiki/Meteorology" title="Meteorology">meteorology</a>, <a href="/wiki/Phenology" title="Phenology">phenology</a>, and <a href="/wiki/Climate_change" title="Climate change">climate change</a>.<sup id="cite_ref-:0_2-0" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Overview">Overview</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=1" title="Edit section: Overview"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The first mention of "aerobiology" was made by Fred Campbell Meier in the 1930s.<sup id="cite_ref-:0_2-1" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The particles, which can be described as <a href="/wiki/Aeroplankton" title="Aeroplankton">Aeroplankton</a>, generally range in size from nanometers to micrometers which makes them challenging to detect.<sup id="cite_ref-:4_3-0" class="reference"><a href="#cite_note-:4-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p><a href="/wiki/Aerosolization" title="Aerosolization">Aerosolization</a> is the process of a small and light particles becoming suspended in moving air. Now <a href="/wiki/Bioaerosol" title="Bioaerosol">bioaerosols</a>, these pollen and fungal spores can be transported across an ocean, or even travel around the globe.<sup id="cite_ref-:2_4-0" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Due to the high quantities of microbes and the ease of dispersion, <a href="/wiki/Martinus_Beijerinck" title="Martinus Beijerinck">Martinus Beijerinck</a> once said "Everything is everywhere, the environment selects".<sup id="cite_ref-:3_5-0" class="reference"><a href="#cite_note-:3-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> This means that aeroplankton are everywhere and have been everywhere, and it solely depends on environmental factors to determine which remain. Aeroplankton are found in significant quantities even in the <a href="/wiki/Atmospheric_boundary_layer" class="mw-redirect" title="Atmospheric boundary layer">Atmospheric boundary layer (ABL)</a>.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> The effects on climate and cloud chemistry of these atmospheric populations is still under review. </p><p><a href="/wiki/NASA" title="NASA">NASA</a> and other research agencies are studying how long these bioaerosols can remain afloat and how they can survive in such extreme climates. The conditions of the upper atmosphere are similar to the climate on Mars' surface, and the microbes found are helping redefine the conditions which can support life.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Dispersal_of_particles">Dispersal of particles</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=2" title="Edit section: Dispersal of particles"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The process of dispersal of aerobiological particles has 3 steps: removal from source, dispersion through air, and deposition to rest.<sup id="cite_ref-:1_8-0" class="reference"><a href="#cite_note-:1-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> The particle geometry and environment affect all three phases, however once it is aerosolized, its fate depends on the laws of physics governing the motion of the air. </p> <div class="mw-heading mw-heading3"><h3 id="Removal_from_Source">Removal from Source</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=3" title="Edit section: Removal from Source"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Puffballs_emitting_spores.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d1/Puffballs_emitting_spores.jpg/220px-Puffballs_emitting_spores.jpg" decoding="async" width="220" height="272" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d1/Puffballs_emitting_spores.jpg/330px-Puffballs_emitting_spores.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d1/Puffballs_emitting_spores.jpg/440px-Puffballs_emitting_spores.jpg 2x" data-file-width="2126" data-file-height="2630" /></a><figcaption>A puffball mushroom ejecting its spores.</figcaption></figure> <p>Pollen and spores can be blown from their surface or shaken loose. Generally the wind speed required for release is higher than average wind speed.<sup id="cite_ref-:1_8-1" class="reference"><a href="#cite_note-:1-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Rain splatter can also dislodge spores. Some fungi can even be triggered by environmental factors to actively eject spores.<sup id="cite_ref-:1_8-2" class="reference"><a href="#cite_note-:1-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Dispersion_through_Air">Dispersion through Air</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=4" title="Edit section: Dispersion through Air"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Once released from rest, the <a href="/wiki/Aeroplankton" title="Aeroplankton">aeroplankton</a> is at the mercy of the wind and physics. The settling speed of spores and pollen vary and is a major factor in dispersion; the longer the particle is floating, the longer it can be caught by a turbulent wind gust. Wind speed and direction fluctuate with time and height, so the specific path of once neighboring particles can vary significantly.<sup id="cite_ref-:1_8-3" class="reference"><a href="#cite_note-:1-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> The concentration of particles in the air decreases with distance from source, and the dispersion distance is most accurately modeled as a <a href="/wiki/Power_Function" class="mw-redirect" title="Power Function">power function</a>.<sup id="cite_ref-:4_3-1" class="reference"><a href="#cite_note-:4-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Deposition_to_Rest">Deposition to Rest</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=5" title="Edit section: Deposition to Rest"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Deposition is a combination of gravity and inertia. The fall speed for small particles can be calculated by mass and geometry, but the complex shapes of pollen and spores often fall slower than their estimated speed modeled with simple shapes.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Spores can also be removed from the air from impact; the inertia of the particles will cause them to hit surfaces along their path, instead of flowing around them like air.<sup id="cite_ref-:1_8-4" class="reference"><a href="#cite_note-:1-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Experimental_methods">Experimental methods</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=6" title="Edit section: Experimental methods"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>There have been many studies performed to understand real-life dispersal patterns of pollen and spores. To collect samples, studies often use a volumetric spore trap such as a Hirst-type sampler. Particles stick to a sampling strip and then can be inspected under a microscope.<sup id="cite_ref-:0_2-2" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Scientists have to count the particles under magnification, and then analyze sample DNA by <a href="/wiki/Amplicon_sequence_variant" title="Amplicon sequence variant">Amplicon sequence variant</a> (ASV) or another common method.<sup id="cite_ref-:5_10-0" class="reference"><a href="#cite_note-:5-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>A challenge repeatedly cited in literature is that because of differing testing or analysis methodologies, results are not always comparable across studies.<sup id="cite_ref-:3_5-1" class="reference"><a href="#cite_note-:3-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> Therefore, extensive data collection must be performed in each study to get an accurate model. Unfortunately there is no database of aerobiological particle distribution to compare results to.<sup id="cite_ref-:3_5-2" class="reference"><a href="#cite_note-:3-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Effects_on_human_health">Effects on human health</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=7" title="Edit section: Effects on human health"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Blausen_0015_AllergicRhinitis.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/55/Blausen_0015_AllergicRhinitis.png/220px-Blausen_0015_AllergicRhinitis.png" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/55/Blausen_0015_AllergicRhinitis.png/330px-Blausen_0015_AllergicRhinitis.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/55/Blausen_0015_AllergicRhinitis.png/440px-Blausen_0015_AllergicRhinitis.png 2x" data-file-width="1024" data-file-height="768" /></a><figcaption>Illustration depicting inflammation associated with allergic rhinitis.</figcaption></figure> <p><a href="/wiki/Allergic_rhinitis" title="Allergic rhinitis">Allergic rhinitis</a> is a type of inflammation in the nose that occurs when the immune system overreacts to allergens in the air.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> It is typically triggered in humans by pollen and other bioaerosols. Between 10% and 30% of people in Western countries are affected.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> Symptoms are usually worse during pollination periods, when there is significantly more pollen aerosolized in the air.<sup id="cite_ref-:6_13-0" class="reference"><a href="#cite_note-:6-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> In these peak periods, staying indoors is one way to limit exposure. However, studies have shown that there are still significant levels of pollen indoors. In the winter, pollen levels indoors actually exceed outdoor levels.<sup id="cite_ref-:6_13-1" class="reference"><a href="#cite_note-:6-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>Up to date data on pollen levels is critical for humans that have allergies. A current limitation is that many spore traps require scientists to identify and count individual pollen grains under magnification.<sup id="cite_ref-:5_10-1" class="reference"><a href="#cite_note-:5-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> This causes data to be delayed, sometimes by over a week. There are currently a number of fully automatic spore traps in development, and once they are fully functional they will improve the lives of people with allergies.<sup id="cite_ref-:5_10-2" class="reference"><a href="#cite_note-:5-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Effects_of_climate_change">Effects of climate change</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=8" title="Edit section: Effects of climate change"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Precipitation_and_climate_change.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/18/Precipitation_and_climate_change.svg/215px-Precipitation_and_climate_change.svg.png" decoding="async" width="215" height="168" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/18/Precipitation_and_climate_change.svg/323px-Precipitation_and_climate_change.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/18/Precipitation_and_climate_change.svg/430px-Precipitation_and_climate_change.svg.png 2x" data-file-width="960" data-file-height="749" /></a><figcaption>Map of precipitation change at +2 °C of global warming</figcaption></figure> <p>Scientists have predicted that the meteorological results of <a href="/wiki/Climate_change" title="Climate change">climate change</a> will weaken pollen and spore dispersal barriers, and lead to less biological uniqueness in different regions.<sup id="cite_ref-:2_4-1" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Precipitation increases richness (number of species) of biodiversity in regions because clouds formulate in the upper atmosphere where there is more varied biodiversity.<sup id="cite_ref-:2_4-2" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Specifically in the Arctic, climate change has dramatically increased precipitation, and scientists have seen new microbes in the area because of it.<sup id="cite_ref-:2_4-3" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Rising summer temperatures and CO<sub>2</sub> levels have shown to increase total amounts of pollen released by certain trees, as well as delay the start of pollen season.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> However, more studies are needed to see long term effects of climate change. </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Aerobiology&action=edit&section=9" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output 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.navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Biology_nav" title="Template:Biology nav"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Biology_nav" title="Template talk:Biology nav"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Biology_nav" title="Special:EditPage/Template:Biology nav"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Biology858" style="font-size:114%;margin:0 4em"><a href="/wiki/Biology" title="Biology">Biology</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li>Introduction (<a href="/wiki/Introduction_to_genetics" title="Introduction to genetics">Genetics</a>, <a href="/wiki/Introduction_to_evolution" title="Introduction to evolution">Evolution</a>)</li> <li><a href="/wiki/Outline_of_biology" title="Outline of biology">Outline</a></li> <li><a href="/wiki/History_of_biology" title="History of biology">History</a> <ul><li><a href="/wiki/Timeline_of_biology_and_organic_chemistry" title="Timeline of biology and organic chemistry">Timeline</a></li></ul></li> <li><a href="/wiki/Index_of_biology_articles" title="Index of biology articles">Index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Biology" title="Biology">Biology</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Overview</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Science" title="Science">Science</a></li> <li><a href="/wiki/Life" title="Life">Life</a></li> <li>Properties (<a href="/wiki/Adaptation" title="Adaptation">Adaptation</a>, <a href="/wiki/Metabolism" title="Metabolism"> Energy processing</a>, <a href="/wiki/Developmental_biology" title="Developmental biology">Growth</a>, <a href="/wiki/Structure#Biological" title="Structure">Order</a>, <a href="/wiki/Homeostasis" title="Homeostasis">Regulation</a>, <a href="/wiki/Reproduction" title="Reproduction">Reproduction</a> (<a href="/wiki/Self-replication" title="Self-replication">Self-replication</a>), <a href="/wiki/Stimulus_(physiology)" title="Stimulus (physiology)">Response to environment</a>)</li> <li><a href="/wiki/Biological_organisation" title="Biological organisation">Hierarchy of life</a> (<a href="/wiki/Atom" title="Atom">Atom</a> > <a href="/wiki/Molecule" title="Molecule">Molecule</a> > <a href="/wiki/Organelle" title="Organelle">Organelle</a> > <a href="/wiki/Cell_(biology)" title="Cell (biology)">Cell</a> > <a href="/wiki/Tissue_(biology)" title="Tissue (biology)">Tissue</a> > <a href="/wiki/Organ_(biology)" title="Organ (biology)">Organ</a> > <a href="/wiki/Biological_system" title="Biological system">Organ system</a> > <a href="/wiki/Organism" title="Organism">Organism</a> > <a href="/wiki/Population" title="Population">Population</a> > <a href="/wiki/Biocoenosis" title="Biocoenosis">Community</a> > <a href="/wiki/Ecosystem" title="Ecosystem">Ecosystem</a> > <a href="/wiki/Biosphere" title="Biosphere">Biosphere</a>)</li> <li><a href="/wiki/Reductionism#In_science" title="Reductionism">Reductionistic</a></li> <li><a href="/wiki/Emergence" title="Emergence"> Emergent property</a></li> <li><a href="/wiki/Mechanical_philosophy" class="mw-redirect" title="Mechanical philosophy">Mechanistic</a></li> <li><a href="/wiki/Scientific_method" title="Scientific method">Scientific method</a></li> <li><a href="/wiki/Taxonomic_rank" title="Taxonomic rank">Taxonomic rank</a></li> <li><a href="/wiki/Scientific_theory" title="Scientific theory"> Theory</a></li> <li><a href="/wiki/Scientific_law" title="Scientific law"> Law</a></li> <li><a href="/wiki/Peer_review" title="Peer review">Peer review</a></li> <li><a href="/wiki/List_of_biology_journals" title="List of biology journals">Biology journals</a></li> <li><a href="/wiki/Common_name" title="Common name">Common name</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Chemical basis</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Atom" title="Atom">Atoms</a></li> <li><a href="/wiki/Amino_acid" title="Amino acid">Amino acids</a></li> <li><a href="/wiki/Carbohydrate" title="Carbohydrate">Carbohydrates</a></li> <li><a href="/wiki/Chemical_bond" title="Chemical bond">Chemical bond</a></li> <li><a href="/wiki/Chemical_element" title="Chemical element">Chemical element</a></li> <li><a href="/wiki/Lipid" title="Lipid">Lipids</a></li> <li><a href="/wiki/Matter" title="Matter">Matter</a> <ul><li><a href="/wiki/Quantum_biology" title="Quantum biology">Quantum</a></li></ul></li> <li><a href="/wiki/Molecule" title="Molecule">Molecules</a></li> <li><a href="/wiki/Monomer" title="Monomer">Monomer</a></li> <li><a href="/wiki/Nucleic_acid" title="Nucleic acid">Nucleic acids</a></li> <li><a href="/wiki/Organic_compound" title="Organic compound">Organic compounds</a></li> <li><a href="/wiki/PH" title="PH">pH</a></li> <li><a href="/wiki/Polymer" title="Polymer">Polymer</a></li> <li><a href="/wiki/Protein" title="Protein">Proteins</a></li> <li><a href="/wiki/Water" title="Water">Water</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cell_(biology)" title="Cell (biology)">Cells</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Adenosine_triphosphate" title="Adenosine triphosphate">ATP</a></li> <li><a href="/wiki/Cell_cycle" title="Cell cycle">Cell cycle</a></li> <li><a href="/wiki/Cell_theory" title="Cell theory">Cell theory</a></li> <li><a href="/wiki/Cell_signaling" title="Cell signaling">Cell signaling</a></li> <li><a href="/wiki/Cellular_respiration" title="Cellular respiration">Cellular respiration</a></li> <li><a href="/wiki/Energy" title="Energy">Energy transformation</a></li> <li><a href="/wiki/Enzyme" title="Enzyme">Enzyme</a></li> <li><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryote</a></li> <li><a href="/wiki/Fermentation" title="Fermentation">Fermentation</a></li> <li><a href="/wiki/Metabolism" title="Metabolism">Metabolism</a></li> <li><a href="/wiki/Meiosis" title="Meiosis">Meiosis</a></li> <li><a href="/wiki/Mitosis" title="Mitosis">Mitosis</a></li> <li><a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a></li> <li><a href="/wiki/Prokaryote" title="Prokaryote">Prokaryote</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Genetics" title="Genetics">Genetics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/DNA" title="DNA">DNA</a></li> <li><a href="/wiki/Epigenetics" title="Epigenetics">Epigenetics</a></li> <li><a href="/wiki/Evolutionary_developmental_biology" title="Evolutionary developmental biology">Evolutionary developmental biology</a></li> <li><a href="/wiki/Gene_expression" title="Gene expression">Gene expression</a></li> <li><a href="/wiki/Regulation_of_gene_expression" title="Regulation of gene expression">Gene regulation</a></li> <li><a href="/wiki/Genome" title="Genome">Genomes</a></li> <li><a href="/wiki/Mendelian_inheritance" title="Mendelian inheritance">Mendelian inheritance</a></li> <li><a href="/wiki/Post-transcriptional_modification" title="Post-transcriptional modification">Post-transcriptional modification</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Evolution" title="Evolution">Evolution</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Adaptation" title="Adaptation">Adaptation</a></li> <li><a href="/wiki/Earliest_known_life_forms" title="Earliest known life forms">Earliest known life forms</a></li> <li><a href="/wiki/Function_(biology)" title="Function (biology)">Function</a></li> <li><a href="/wiki/Genetic_drift" title="Genetic drift">Genetic drift</a></li> <li><a href="/wiki/Gene_flow" title="Gene flow">Gene flow</a></li> <li><a href="/wiki/History_of_life" title="History of life">History of life</a></li> <li><a href="/wiki/Macroevolution" title="Macroevolution">Macroevolution</a></li> <li><a href="/wiki/Microevolution" title="Microevolution">Microevolution</a></li> <li><a href="/wiki/Mutation" title="Mutation">Mutation</a></li> <li><a href="/wiki/Natural_selection" title="Natural selection">Natural selection</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a></li> <li><a href="/wiki/Speciation" title="Speciation">Speciation</a></li> <li><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Taxonomy</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Biodiversity" title="Biodiversity">Diversity</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Bacteria" title="Bacteria">Bacteria</a></li> <li><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryote</a> <ul><li><a href="/wiki/Alga" class="mw-redirect" title="Alga">Alga</a></li> <li><a href="/wiki/Animal" title="Animal">Animal</a></li> <li><a href="/wiki/Fungus" title="Fungus">Fungus</a></li> <li><a href="/wiki/Plant" title="Plant">Plant</a></li> <li><a href="/wiki/Protist" title="Protist">Protist</a></li></ul></li> <li><a href="/wiki/Virus" title="Virus">Virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Plant form <br /> and function</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Epidermis_(botany)" title="Epidermis (botany)">Epidermis (botany)</a></li> <li><a href="/wiki/Flower" title="Flower">Flower</a></li> <li><a href="/wiki/Ground_tissue" title="Ground tissue">Ground tissue</a></li> <li><a href="/wiki/Leaf" title="Leaf">Leaf</a></li> <li><a href="/wiki/Phloem" title="Phloem">Phloem</a></li> <li><a href="/wiki/Plant_stem" title="Plant stem">Plant stem</a></li> <li><a href="/wiki/Root" title="Root">Root</a></li> <li><a href="/wiki/Shoot_(botany)" title="Shoot (botany)">Shoot</a></li> <li><a href="/wiki/Vascular_plant" title="Vascular plant">Vascular plant</a></li> <li><a href="/wiki/Vascular_tissue" title="Vascular tissue">Vascular tissue</a></li> <li><a href="/wiki/Xylem" title="Xylem">Xylem</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Animal form <br />and function</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Breathing" title="Breathing">Breathing</a></li> <li><a href="/wiki/Circulatory_system" title="Circulatory system">Circulatory system</a></li> <li><a href="/wiki/Endocrine_system" title="Endocrine system">Endocrine system</a></li> <li><a href="/wiki/Human_digestive_system" title="Human digestive system">Digestive system</a></li> <li><a href="/wiki/Homeostasis" title="Homeostasis">Homeostasis</a></li> <li><a href="/wiki/Immune_system" title="Immune system">Immune system</a></li> <li><a href="/wiki/Internal_environment" title="Internal environment">Internal environment</a></li> <li><a href="/wiki/Muscular_system" title="Muscular system">Muscular system</a></li> <li><a href="/wiki/Nervous_system" title="Nervous system">Nervous system</a></li> <li><a href="/wiki/Reproductive_system" title="Reproductive system">Reproductive system</a></li> <li><a href="/wiki/Respiratory_system" title="Respiratory system">Respiratory system</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ecology" title="Ecology">Ecology</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biogeochemical_cycle" title="Biogeochemical cycle">Biogeochemical cycle</a></li> <li><a href="/wiki/Biological_interaction" title="Biological interaction">Biological interaction</a></li> <li><a href="/wiki/Biomass" title="Biomass">Biomass</a></li> <li><a href="/wiki/Biome" title="Biome">Biomes</a></li> <li><a href="/wiki/Biosphere" title="Biosphere">Biosphere</a></li> <li><a href="/wiki/Climate" title="Climate">Climate</a></li> <li><a href="/wiki/Climate_change" title="Climate change">Climate change</a></li> <li><a href="/wiki/Community_(ecology)" title="Community (ecology)">Community</a></li> <li><a href="/wiki/Conservation_biology" title="Conservation biology">Conservation</a></li> <li><a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem</a></li> <li><a href="/wiki/Habitat" title="Habitat">Habitat</a> <ul><li><a href="/wiki/Ecological_niche" title="Ecological niche">niche</a></li></ul></li> <li><a href="/wiki/Microbiome" title="Microbiome">Microbiome</a></li> <li><a href="/wiki/Population_ecology" title="Population ecology">Population dynamics</a></li> <li><a href="/wiki/Resource_(biology)" title="Resource (biology)">Resources</a></li></ul> </div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="4" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Tree_of_life_by_Haeckel.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/de/Tree_of_life_by_Haeckel.jpg/140px-Tree_of_life_by_Haeckel.jpg" decoding="async" width="140" height="222" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/de/Tree_of_life_by_Haeckel.jpg/210px-Tree_of_life_by_Haeckel.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/de/Tree_of_life_by_Haeckel.jpg/280px-Tree_of_life_by_Haeckel.jpg 2x" data-file-width="1804" data-file-height="2856" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/List_of_research_methods_in_biology" title="List of research methods in biology">Research <br /> methods</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Laboratory <br /> techniques</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Genetic_engineering" title="Genetic engineering">Genetic engineering</a></li> <li><a href="/wiki/Transformation_(genetics)" class="mw-redirect" title="Transformation (genetics)">Transformation</a></li> <li><a href="/wiki/Gel_electrophoresis" title="Gel electrophoresis">Gel electrophoresis</a></li> <li><a href="/wiki/Chromatography" title="Chromatography">Chromatography</a></li> <li><a href="/wiki/Centrifugation" title="Centrifugation">Centrifugation</a></li> <li><a href="/wiki/Cell_culture" title="Cell culture">Cell culture</a></li> <li><a href="/wiki/DNA_sequencing" title="DNA sequencing">DNA sequencing</a></li> <li><a href="/wiki/DNA_microarray" title="DNA microarray">DNA microarray</a></li> <li><a href="/wiki/Green_fluorescent_protein" title="Green fluorescent protein">Green fluorescent protein</a></li> <li><a href="/wiki/Vector_(molecular_biology)" title="Vector (molecular biology)">vector</a></li> <li><a href="/wiki/Enzyme_assay" title="Enzyme assay">Enzyme assay</a></li> <li><a href="/wiki/Protein_purification" title="Protein purification">Protein purification</a></li> <li><a href="/wiki/Western_blot" title="Western blot">Western blot</a></li> <li><a href="/wiki/Northern_blot" title="Northern blot">Northern blot</a></li> <li><a href="/wiki/Southern_blot" title="Southern blot">Southern blot</a></li> <li><a href="/wiki/Restriction_enzyme" title="Restriction enzyme">Restriction enzyme</a></li> <li><a href="/wiki/Polymerase_chain_reaction" title="Polymerase chain reaction">Polymerase chain reaction</a></li> <li><a href="/wiki/Two-hybrid_screening" title="Two-hybrid screening">Two-hybrid screening</a></li> <li><i><a href="/wiki/In_vivo" title="In vivo">in vivo</a></i></li> <li><i><a href="/wiki/In_vitro" title="In vitro">in vitro</a></i></li> <li><i><a href="/wiki/In_silico" title="In silico">in silico</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Field techniques</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Belt_transect" title="Belt transect">Belt transect</a></li> <li><a href="/wiki/Mark_and_recapture" title="Mark and recapture">mark and recapture</a></li> <li><a href="/wiki/Species_discovery_curve" title="Species discovery curve">species discovery curve</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Branches</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><div class="excerpt-block"><div class="excerpt"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></li> <li>Aerobiology</li> <li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Agrostology" title="Agrostology">Agrostology</a></li> <li><a href="/wiki/Anatomy" title="Anatomy">Anatomy</a></li> <li><a href="/wiki/Astrobiology" title="Astrobiology">Astrobiology</a></li> <li><a href="/wiki/Bacteriology" title="Bacteriology">Bacteriology</a></li> <li><a href="/wiki/Biochemistry" title="Biochemistry">Biochemistry</a></li> <li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Biogeology" title="Biogeology">Biogeology</a></li> <li><a href="/wiki/Bioinformatics" title="Bioinformatics">Bioinformatics</a></li> <li><a href="/wiki/Biological_engineering" title="Biological engineering">Biological engineering</a></li> <li><a href="/wiki/Biomechanics" title="Biomechanics">Biomechanics</a></li> <li><a href="/wiki/Biophysics" title="Biophysics">Biophysics</a></li> <li><a href="/wiki/Biosemiotics" title="Biosemiotics">Biosemiotics</a></li> <li><a href="/wiki/Biostatistics" title="Biostatistics">Biostatistics</a></li> <li><a href="/wiki/Biotechnology" title="Biotechnology">Biotechnology</a></li> <li><a href="/wiki/Botany" title="Botany">Botany</a></li> <li><a href="/wiki/Cell_biology" title="Cell biology">Cell biology</a></li> <li><a href="/wiki/Cellular_microbiology" title="Cellular microbiology">Cellular microbiology</a></li> <li><a href="/wiki/Chemical_biology" title="Chemical biology">Chemical biology</a></li> <li><a href="/wiki/Chronobiology" title="Chronobiology">Chronobiology</a></li> <li><a href="/wiki/Cognitive_biology" title="Cognitive biology">Cognitive biology</a></li> <li><a href="/wiki/Computational_biology" title="Computational biology">Computational biology</a></li> <li><a href="/wiki/Conservation_biology" title="Conservation biology">Conservation biology</a></li> <li><a href="/wiki/Cryobiology" title="Cryobiology">Cryobiology</a></li> <li><a href="/wiki/Cytogenetics" title="Cytogenetics">Cytogenetics</a></li> <li><a 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