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Plant pathology - Wikipedia
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href="#Physiological_plant_disorders"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Physiological plant disorders</span> </div> </a> <ul id="toc-Physiological_plant_disorders-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Epidemiology" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Epidemiology"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Epidemiology</span> </div> </a> <ul id="toc-Epidemiology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Disease_resistance" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Disease_resistance"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Disease resistance</span> </div> </a> <ul id="toc-Disease_resistance-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Management" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Management"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Management</span> </div> </a> <button aria-controls="toc-Management-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Management subsection</span> </button> <ul id="toc-Management-sublist" class="vector-toc-list"> <li id="toc-Detection" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Detection"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.1</span> <span>Detection</span> </div> </a> <ul id="toc-Detection-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Biological" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Biological"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2</span> <span>Biological</span> </div> </a> <ul id="toc-Biological-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-History" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#History"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>History</span> </div> </a> <ul id="toc-History-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notable_People_in_Plant_Pathology" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notable_People_in_Plant_Pathology"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Notable People in Plant Pathology</span> </div> </a> <ul id="toc-Notable_People_in_Plant_Pathology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">9</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">10</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Plant pathology</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 55 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-55" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">55 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Plantsiektes" title="Plantsiektes – Afrikaans" lang="af" hreflang="af" data-title="Plantsiektes" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B9%D9%84%D9%85_%D8%A3%D9%85%D8%B1%D8%A7%D8%B6_%D8%A7%D9%84%D9%86%D8%A8%D8%A7%D8%AA" title="علم أمراض النبات – Arabic" lang="ar" hreflang="ar" data-title="علم أمراض النبات" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Fitopatologiya" title="Fitopatologiya – Azerbaijani" lang="az" hreflang="az" data-title="Fitopatologiya" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%89%E0%A6%A6%E0%A7%8D%E0%A6%AD%E0%A6%BF%E0%A6%A6_%E0%A6%B0%E0%A7%8B%E0%A6%97%E0%A6%A4%E0%A6%A4%E0%A7%8D%E0%A6%A4%E0%A7%8D%E0%A6%AC" title="উদ্ভিদ রোগতত্ত্ব – Bangla" lang="bn" hreflang="bn" data-title="উদ্ভিদ রোগতত্ত্ব" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Si%CC%8Dt-bu%CC%8Dt_p%C4%93%E2%81%BF-l%C3%AD-ha%CC%8Dk" title="Si̍t-bu̍t pēⁿ-lí-ha̍k – Minnan" lang="nan" hreflang="nan" data-title="Si̍t-bu̍t pēⁿ-lí-ha̍k" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A4%D1%96%D1%82%D0%B0%D0%BF%D0%B0%D1%82%D0%B0%D0%BB%D0%BE%D0%B3%D1%96%D1%8F" title="Фітапаталогія – Belarusian" lang="be" hreflang="be" data-title="Фітапаталогія" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A4%D0%B8%D1%82%D0%BE%D0%BF%D0%B0%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Фитопатология – Bulgarian" lang="bg" hreflang="bg" data-title="Фитопатология" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Patologija_biljaka" title="Patologija biljaka – Bosnian" lang="bs" hreflang="bs" data-title="Patologija biljaka" data-language-autonym="Bosanski" data-language-local-name="Bosnian" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Fitopatologia" title="Fitopatologia – Catalan" lang="ca" hreflang="ca" data-title="Fitopatologia" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Rostlinol%C3%A9ka%C5%99stv%C3%AD" title="Rostlinolékařství – Czech" lang="cs" hreflang="cs" data-title="Rostlinolékařství" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Plantesygdomme" title="Plantesygdomme – Danish" lang="da" hreflang="da" data-title="Plantesygdomme" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Phytopathologie" title="Phytopathologie – German" lang="de" hreflang="de" data-title="Phytopathologie" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/F%C3%BCtopatoloogia" title="Fütopatoloogia – Estonian" lang="et" hreflang="et" data-title="Fütopatoloogia" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A6%CF%85%CF%84%CE%BF%CF%80%CE%B1%CE%B8%CE%BF%CE%BB%CE%BF%CE%B3%CE%AF%CE%B1" title="Φυτοπαθολογία – Greek" lang="el" hreflang="el" data-title="Φυτοπαθολογία" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Fitopatolog%C3%ADa" title="Fitopatología – Spanish" lang="es" hreflang="es" data-title="Fitopatología" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Fitopatologio" title="Fitopatologio – Esperanto" lang="eo" hreflang="eo" data-title="Fitopatologio" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A8%DB%8C%D9%85%D8%A7%D8%B1%DB%8C%E2%80%8C%D8%B4%D9%86%D8%A7%D8%B3%DB%8C_%DA%AF%DB%8C%D8%A7%D9%87%DB%8C" title="بیماریشناسی گیاهی – Persian" lang="fa" hreflang="fa" data-title="بیماریشناسی گیاهی" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Pathologie_v%C3%A9g%C3%A9tale" title="Pathologie végétale – French" lang="fr" hreflang="fr" data-title="Pathologie végétale" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Fitopatolox%C3%ADa" title="Fitopatoloxía – Galician" lang="gl" hreflang="gl" data-title="Fitopatoloxía" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%8B%9D%EB%AC%BC%EB%B3%91%EB%A6%AC%ED%95%99" title="식물병리학 – Korean" lang="ko" hreflang="ko" data-title="식물병리학" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%AA%E0%A4%BE%E0%A4%A6%E0%A4%AA_%E0%A4%B0%E0%A5%8B%E0%A4%97%E0%A4%B5%E0%A4%BF%E0%A4%9C%E0%A5%8D%E0%A4%9E%E0%A4%BE%E0%A4%A8" title="पादप रोगविज्ञान – Hindi" lang="hi" hreflang="hi" data-title="पादप रोगविज्ञान" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Fitopatologija" title="Fitopatologija – Croatian" lang="hr" hreflang="hr" data-title="Fitopatologija" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Fitopatologi" title="Fitopatologi – Indonesian" lang="id" hreflang="id" data-title="Fitopatologi" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Patologia_vegetale" title="Patologia vegetale – Italian" lang="it" hreflang="it" data-title="Patologia vegetale" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A4%D7%99%D7%98%D7%95%D7%A4%D7%AA%D7%95%D7%9C%D7%95%D7%92%D7%99%D7%94" title="פיטופתולוגיה – Hebrew" lang="he" hreflang="he" data-title="פיטופתולוגיה" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%A4%E1%83%98%E1%83%A2%E1%83%9D%E1%83%9E%E1%83%90%E1%83%97%E1%83%9D%E1%83%9A%E1%83%9D%E1%83%92%E1%83%98%E1%83%90" title="ფიტოპათოლოგია – Georgian" lang="ka" hreflang="ka" data-title="ფიტოპათოლოგია" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A4%D0%B8%D1%82%D0%BE%D0%BF%D0%B0%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Фитопатология – Kazakh" lang="kk" hreflang="kk" data-title="Фитопатология" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%A4%D0%B8%D1%82%D0%BE%D0%BF%D0%B0%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Фитопатология – Kyrgyz" lang="ky" hreflang="ky" data-title="Фитопатология" data-language-autonym="Кыргызча" data-language-local-name="Kyrgyz" class="interlanguage-link-target"><span>Кыргызча</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Fitopatolo%C4%A3ija" title="Fitopatoloģija – Latvian" lang="lv" hreflang="lv" data-title="Fitopatoloģija" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Fitopatologija" title="Fitopatologija – Lithuanian" lang="lt" hreflang="lt" data-title="Fitopatologija" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/N%C3%B6v%C3%A9nyk%C3%B3rtan" title="Növénykórtan – Hungarian" lang="hu" hreflang="hu" data-title="Növénykórtan" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%A4%D0%B8%D1%82%D0%BE%D0%BF%D0%B0%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%98%D0%B0" title="Фитопатологија – Macedonian" lang="mk" hreflang="mk" data-title="Фитопатологија" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%B8%E0%B4%B8%E0%B5%8D%E0%B4%AF%E0%B4%B0%E0%B5%8B%E0%B4%97%E0%B4%B6%E0%B4%BE%E0%B4%B8%E0%B5%8D%E0%B4%A4%E0%B5%8D%E0%B4%B0%E0%B4%82" title="സസ്യരോഗശാസ്ത്രം – Malayalam" lang="ml" hreflang="ml" data-title="സസ്യരോഗശാസ്ത്രം" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Patologi_tumbuhan" title="Patologi tumbuhan – Malay" lang="ms" hreflang="ms" data-title="Patologi tumbuhan" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Fytopathologie" title="Fytopathologie – Dutch" lang="nl" hreflang="nl" data-title="Fytopathologie" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E6%A4%8D%E7%89%A9%E7%97%85%E7%90%86%E5%AD%A6" title="植物病理学 – Japanese" lang="ja" hreflang="ja" data-title="植物病理学" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Fitopatologiya" title="Fitopatologiya – Uzbek" lang="uz" hreflang="uz" data-title="Fitopatologiya" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Fitopatologia" title="Fitopatologia – Polish" lang="pl" hreflang="pl" data-title="Fitopatologia" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Fitopatologia" title="Fitopatologia – Portuguese" lang="pt" hreflang="pt" data-title="Fitopatologia" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Fitopatologie" title="Fitopatologie – Romanian" lang="ro" hreflang="ro" data-title="Fitopatologie" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A4%D0%B8%D1%82%D0%BE%D0%BF%D0%B0%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Фитопатология – Russian" lang="ru" hreflang="ru" data-title="Фитопатология" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Fitopatologjia" title="Fitopatologjia – Albanian" lang="sq" hreflang="sq" data-title="Fitopatologjia" data-language-autonym="Shqip" data-language-local-name="Albanian" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-si mw-list-item"><a href="https://si.wikipedia.org/wiki/%E0%B7%81%E0%B7%8F%E0%B6%9A_%E0%B7%80%E0%B7%8A%E2%80%8D%E0%B6%BA%E0%B7%8F%E0%B6%B0%E0%B7%92_%E0%B7%80%E0%B7%92%E0%B6%AF%E0%B7%8A%E2%80%8D%E0%B6%BA%E0%B7%8F%E0%B7%80" title="ශාක ව්යාධි විද්යාව – Sinhala" lang="si" hreflang="si" data-title="ශාක ව්යාධි විද්යාව" data-language-autonym="සිංහල" data-language-local-name="Sinhala" class="interlanguage-link-target"><span>සිංහල</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Plant_pathology" title="Plant pathology – Simple English" lang="en-simple" hreflang="en-simple" data-title="Plant pathology" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Fytopatol%C3%B3gia" title="Fytopatológia – Slovak" lang="sk" hreflang="sk" data-title="Fytopatológia" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D8%B2%D8%A7%D9%86%D8%B3%D8%AA%DB%8C_%D9%86%DB%95%D8%AE%DB%86%D8%B4%DB%8C%DB%8C%DB%95%DA%A9%D8%A7%D9%86%DB%8C_%DA%95%D9%88%DB%95%DA%A9" title="زانستی نەخۆشییەکانی ڕوەک – Central Kurdish" lang="ckb" hreflang="ckb" data-title="زانستی نەخۆشییەکانی ڕوەک" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-su mw-list-item"><a href="https://su.wikipedia.org/wiki/Pitopatologi" title="Pitopatologi – Sundanese" lang="su" hreflang="su" data-title="Pitopatologi" data-language-autonym="Sunda" data-language-local-name="Sundanese" class="interlanguage-link-target"><span>Sunda</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Kasvipatologia" title="Kasvipatologia – Finnish" lang="fi" hreflang="fi" data-title="Kasvipatologia" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/V%C3%A4xtpatologi" title="Växtpatologi – Swedish" lang="sv" hreflang="sv" data-title="Växtpatologi" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%A4%E0%AE%BE%E0%AE%B5%E0%AE%B0_%E0%AE%A8%E0%AF%8B%E0%AE%AF%E0%AE%BF%E0%AE%AF%E0%AE%B2%E0%AF%8D" title="தாவர நோயியல் – Tamil" lang="ta" hreflang="ta" data-title="தாவர நோயியல்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Fitopatoloji" title="Fitopatoloji – Turkish" lang="tr" hreflang="tr" data-title="Fitopatoloji" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A4%D1%96%D1%82%D0%BE%D0%BF%D0%B0%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D1%96%D1%8F" title="Фітопатологія – Ukrainian" lang="uk" hreflang="uk" data-title="Фітопатологія" data-language-autonym="Українська" data-language-local-name="Ukrainian" 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id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Scientific study of plant diseases</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">For the journal, see <a href="/wiki/Plant_Pathology_(journal)" title="Plant Pathology (journal)"><i>Plant Pathology</i> (journal)</a>. For organisms that damage crops and forestry, see <a href="/wiki/Pest_(organism)" title="Pest (organism)">Pest (organism)</a>.</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">"Phytopathology" redirects here. For the journal, see <a href="/wiki/Phytopathology_(journal)" class="mw-redirect" title="Phytopathology (journal)"><i>Phytopathology</i> (journal)</a>.</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Black_rot_lifecycle.tif" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Black_rot_lifecycle.tif/lossy-page1-390px-Black_rot_lifecycle.tif.jpg" decoding="async" width="390" height="313" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Black_rot_lifecycle.tif/lossy-page1-585px-Black_rot_lifecycle.tif.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Black_rot_lifecycle.tif/lossy-page1-780px-Black_rot_lifecycle.tif.jpg 2x" data-file-width="1270" data-file-height="1020" /></a><figcaption>Life cycle of the black rot pathogen, the <a href="/wiki/Gram-negative_bacteria" title="Gram-negative bacteria">gram negative</a> bacterium <a href="/wiki/Xanthomonas_campestris_pv._campestris" title="Xanthomonas campestris pv. campestris"><i>Xanthomonas campestris</i> pathovar <i>campestris</i></a></figcaption></figure> <p><b>Plant pathology</b> or <b>phytopathology</b> is the scientific study of <a href="/wiki/Plant_disease" title="Plant disease">plant diseases</a> caused by <a href="/wiki/Pathogen" title="Pathogen">pathogens</a> (infectious organisms) and environmental conditions (physiological factors).<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Plant pathology involves the study of pathogen identification, disease <a href="/wiki/Etiology" title="Etiology">etiology</a>, disease cycles, economic impact, <a href="/wiki/Plant_disease_epidemiology" title="Plant disease epidemiology">plant disease epidemiology</a>, <a href="/wiki/Plant_disease_resistance" title="Plant disease resistance">plant disease resistance</a>, how plant diseases affect humans and animals, <a href="/wiki/Pathosystem" title="Pathosystem">pathosystem</a> genetics, and management of plant diseases. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Plant_pathogenicity">Plant pathogenicity</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=1" title="Edit section: Plant pathogenicity"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Plant_disease" title="Plant disease">Plant disease</a></div> <p>Plant pathogens, organisms that cause infectious <a href="/wiki/Plant_disease" title="Plant disease">plant diseases</a>, include <a href="/wiki/Fungus" title="Fungus">fungi</a>, <a href="/wiki/Oomycetes" class="mw-redirect" title="Oomycetes">oomycetes</a>, <a href="/wiki/Bacterium" class="mw-redirect" title="Bacterium">bacteria</a>, <a href="/wiki/Plant_virus" title="Plant virus">viruses</a>, <a href="/wiki/Viroid" title="Viroid">viroids</a>, <a href="/wiki/Virus" title="Virus">virus</a>-like organisms, <a href="/wiki/Phytoplasmas" class="mw-redirect" title="Phytoplasmas">phytoplasmas</a>, <a href="/wiki/Protozoa" title="Protozoa">protozoa</a>, <a href="/wiki/Nematode" title="Nematode">nematodes</a> and <a href="/wiki/Parasitic_plant" title="Parasitic plant">parasitic plants</a>.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> In most plant <a href="/wiki/Pathosystem" title="Pathosystem">pathosystems</a>, <a href="/wiki/Virulence" title="Virulence">virulence</a> depends on <a href="/wiki/Hydrolase" title="Hydrolase">hydrolases</a> and enzymes that degrade the <a href="/wiki/Cell_wall" title="Cell wall">cell wall</a>. The vast majority of these act on <a href="/wiki/Pectin" title="Pectin">pectins</a> (for example, <a href="/wiki/Pectinesterase" title="Pectinesterase">pectinesterase</a>, <a href="/wiki/Pectate_lyase" title="Pectate lyase">pectate lyase</a>, and <a href="/wiki/Pectinase" title="Pectinase">pectinases</a>). For microbes, the cell wall <a href="/wiki/Polysaccharide" title="Polysaccharide">polysaccharides</a> are both a food source and a barrier to be overcome. Many pathogens grow opportunistically when the host breaks down its own cell walls, most often during <a href="/wiki/Fruit_ripening" class="mw-redirect" title="Fruit ripening">fruit ripening</a>.<sup id="cite_ref-Cantu-et-al-2008_3-0" class="reference"><a href="#cite_note-Cantu-et-al-2008-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Unlike human and animal pathology, plant pathology usually focuses on a single causal organism; however, some plant diseases have been shown to be interactions between multiple pathogens.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>To colonize a plant, pathogens have specific <a href="/wiki/Pathogenicity_factor" class="mw-redirect" title="Pathogenicity factor">pathogenicity factors</a>, of five main types: uses of cell wall–degrading enzymes, <a href="/wiki/Toxins" class="mw-redirect" title="Toxins">toxins</a>, effector proteins, <a href="/wiki/Phytohormone" class="mw-redirect" title="Phytohormone">phytohormones</a>, and <a href="/wiki/Exopolysaccharide" class="mw-redirect" title="Exopolysaccharide">exopolysaccharides</a>. </p> <ul><li><b>Cell wall-degrading enzymes</b>: These are used to break down the plant <a href="/wiki/Cell_wall" title="Cell wall">cell wall</a> in order to release the nutrients inside and include esterases, glycosyl hydrolases, lyases and oxidoreductases.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li> <li><b><a href="/wiki/Toxins" class="mw-redirect" title="Toxins">Toxins</a></b>: These can be non-host-specific, which damage all plants, or host-specific, which cause damage only on a host plant.</li> <li><b>Effector proteins</b>: These can be secreted by pathogens such as bacteria, fungi, and oomycetes<sup id="cite_ref-watermold_6-0" class="reference"><a href="#cite_note-watermold-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Oomycetes_7-0" class="reference"><a href="#cite_note-Oomycetes-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> into the extracellular environment or directly into the host cell, often via the <a href="/wiki/Type_three_secretion_system" class="mw-redirect" title="Type three secretion system">Type three secretion system</a>. Some effectors are known to suppress host defense processes. This can include reducing the plant's internal signaling mechanisms or reduction of phytochemicals production.<sup id="cite_ref-effector_8-0" class="reference"><a href="#cite_note-effector-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li><b><a href="/wiki/Phytohormone" class="mw-redirect" title="Phytohormone">Phytohormones</a></b> are chemicals used by plants for signaling; pathogens can produce these to modify plant growth to their own advantage.</li> <li><b><a href="/wiki/Exopolysaccharide" class="mw-redirect" title="Exopolysaccharide">Exopolysaccharides</a></b> are mostly small chains of sugars that help pathogens to adhere to a plant's surface, enabling them to begin the process of infection.</li></ul> <div class="mw-heading mw-heading2"><h2 id="Physiological_plant_disorders">Physiological plant disorders</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=2" title="Edit section: Physiological plant disorders"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Physiological_plant_disorder" title="Physiological plant disorder">Physiological plant disorder</a></div> <p>Some abiotic disorders can be confused with pathogen-induced disorders. Abiotic causes include natural processes such as <a href="/wiki/Drought" title="Drought">drought</a>, <a href="/wiki/Frost" title="Frost">frost</a>, <a href="/wiki/Snow" title="Snow">snow</a> and <a href="/wiki/Hail" title="Hail">hail</a>; <a href="/wiki/Flood" title="Flood">flooding</a> and poor drainage; <a href="/wiki/Micronutrient_deficiency" title="Micronutrient deficiency">nutrient deficiency</a>; deposition of mineral salts such as <a href="/wiki/Sodium_chloride" title="Sodium chloride">sodium chloride</a> and <a href="/wiki/Gypsum" title="Gypsum">gypsum</a>; <a href="/wiki/Wind" title="Wind">windburn</a> and breakage by storms; and <a href="/wiki/Wildfire" title="Wildfire">wildfires</a>. <sup id="cite_ref-Schutzki2007_9-0" class="reference"><a href="#cite_note-Schutzki2007-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Epidemiology">Epidemiology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=3" title="Edit section: Epidemiology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Plant_disease_epidemiology" title="Plant disease epidemiology">Plant disease epidemiology</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Plant_Disease_Triangle.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/de/Plant_Disease_Triangle.svg/220px-Plant_Disease_Triangle.svg.png" decoding="async" width="220" height="220" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/de/Plant_Disease_Triangle.svg/330px-Plant_Disease_Triangle.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/de/Plant_Disease_Triangle.svg/440px-Plant_Disease_Triangle.svg.png 2x" data-file-width="500" data-file-height="500" /></a><figcaption>Plant disease triangle</figcaption></figure> <p>Epidemiology is the study of factors affecting the outbreak and spread of infectious diseases.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>A disease triangle describes the basic factors required for plant diseases. These are the host plant, the pathogen, and the environment. Any one of these can be modified to control a disease.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Disease_resistance">Disease resistance</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=4" title="Edit section: Disease resistance"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Plant_disease_resistance" title="Plant disease resistance">Plant disease resistance</a></div> <p>Plant disease resistance is the ability of a plant to prevent and terminate infections from plant pathogens. Structures that help plants prevent pathogens from entering are the cuticular layer, cell walls, and stomata guard cells. Once pathogens have overcome these barriers, plant receptors initiate signaling pathways to create molecules to compete against the foreign molecules. These pathways are influenced and triggered by genes within the host plant and can manipulated by genetic breeding to create resistant varieties.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Management">Management</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=5" title="Edit section: Management"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Antagonism_(phytopathology)" title="Antagonism (phytopathology)">Antagonism (phytopathology)</a></div> <div class="mw-heading mw-heading3"><h3 id="Detection">Detection</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=6" title="Edit section: Detection"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Ancient methods of leaf examination and breaking open plant material by hand are now augmented by newer technologies. These include <a href="/wiki/Molecular_pathology" title="Molecular pathology">molecular pathology</a> assays such as <a href="/wiki/Polymerase_chain_reaction" title="Polymerase chain reaction">polymerase chain reaction</a> (PCR), <a href="/wiki/RT-PCR" class="mw-redirect" title="RT-PCR">RT-PCR</a> and <a href="/wiki/Loop-mediated_isothermal_amplification" title="Loop-mediated isothermal amplification">loop-mediated isothermal amplification</a> (LAMP).<sup id="cite_ref-Mumford-et-al-2006_13-0" class="reference"><a href="#cite_note-Mumford-et-al-2006-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Although PCR can detect multiple molecular targets in a single solution there are limits.<sup id="cite_ref-Mumford-et-al-2006_13-1" class="reference"><a href="#cite_note-Mumford-et-al-2006-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Bertolini et al. 2001, Ito et al. 2002, and Ragozzino et al. 2004 developed PCR methods for multiplexing six or seven plant pathogen molecular products and Persson et al. 2005 for multiplexing four with RT-PCR.<sup id="cite_ref-Mumford-et-al-2006_13-2" class="reference"><a href="#cite_note-Mumford-et-al-2006-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> More extensive <a href="/wiki/Molecular_diagnosis" class="mw-redirect" title="Molecular diagnosis">molecular diagnosis</a> requires <a href="/wiki/PCR_array" class="mw-redirect" title="PCR array">PCR arrays</a>.<sup id="cite_ref-Mumford-et-al-2006_13-3" class="reference"><a href="#cite_note-Mumford-et-al-2006-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> The primary detection method used worldwide is <a href="/wiki/Enzyme_linked_immunosorbent_assay" class="mw-redirect" title="Enzyme linked immunosorbent assay">enzyme linked immunosorbent assay</a>.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Biological">Biological</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=7" title="Edit section: Biological"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Crop_rotation" title="Crop rotation">Crop rotation</a> is a traditional and sometimes effective means of preventing a parasitic population from becoming well-established. For example, protection against infection by <i><a href="/wiki/Agrobacterium_tumefaciens" title="Agrobacterium tumefaciens">Agrobacterium tumefaciens</a></i>, which causes gall diseases in many plants, by dipping cuttings in suspensions of <i><a href="/wiki/Agrobacterium_radiobacter" title="Agrobacterium radiobacter">Agrobacterium radiobacter</a></i> before inserting them in the ground to take root.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="History">History</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=8" title="Edit section: History"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Timeline_of_plant_pathology" title="Timeline of plant pathology">Timeline of plant pathology</a></div> <p>Plant pathology has developed from antiquity, starting with <a href="/wiki/Theophrastus" title="Theophrastus">Theophrastus</a> in the ancient era, but scientific study began in the <a href="/wiki/Early_Modern" class="mw-redirect" title="Early Modern">Early Modern</a> period with the invention of the <a href="/wiki/Microscope" title="Microscope">microscope</a>, and developed in the 19th century.<sup id="cite_ref-Ainsworth_16-0" class="reference"><a href="#cite_note-Ainsworth-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Notable_People_in_Plant_Pathology">Notable People in Plant Pathology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=9" title="Edit section: Notable People in Plant Pathology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/George_Washington_Carver" title="George Washington Carver">George Washington Carver</a></li> <li><a href="/wiki/Heinrich_Anton_de_Bary" title="Heinrich Anton de Bary">Anton de Bary</a></li> <li><a href="/wiki/Erwin_Frink_Smith" title="Erwin Frink Smith">Erwin Frink Smith</a></li> <li><a href="/wiki/Agnes_Arber" title="Agnes Arber">Agnes Robertson Arber</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=10" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col"> <ul><li><a href="/wiki/American_Phytopathological_Society" title="American Phytopathological Society">American Phytopathological Society</a></li> <li><a href="/wiki/Australasian_Plant_Pathology_Society" title="Australasian Plant Pathology Society">Australasian Plant Pathology Society</a></li> <li><a href="/wiki/British_Society_for_Plant_Pathology" title="British Society for Plant Pathology">British Society for Plant Pathology</a></li> <li><a href="/wiki/Forest_pathology" title="Forest pathology">Forest pathology</a></li> <li><a href="/wiki/Gene-for-gene_relationship" title="Gene-for-gene relationship">Gene-for-gene relationship</a></li> <li><a href="/wiki/Global_Plant_Clinic" title="Global Plant Clinic">Global Plant Clinic</a></li> <li><a href="/wiki/Glossary_of_phytopathology" title="Glossary of phytopathology">Glossary of phytopathology</a></li> <li><a href="/wiki/Horsfall-Barratt_scale" class="mw-redirect" title="Horsfall-Barratt scale">Horsfall-Barratt scale</a></li> <li><a href="/wiki/List_of_phytopathology_journals" class="mw-redirect" title="List of phytopathology journals">List of phytopathology journals</a></li> <li><a href="/wiki/Microbial_inoculant" title="Microbial inoculant">Microbial inoculant</a></li> <li><a href="/wiki/Phytopharmacology" title="Phytopharmacology">Phytopharmacology</a></li> <li><a href="/wiki/Plant_disease_forecasting" title="Plant disease forecasting">Plant disease forecasting</a></li> <li><a href="/wiki/Stunt_(botany)" title="Stunt (botany)">Stunting</a></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=11" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFOliver2024" class="citation book cs1">Oliver, Richard, ed. 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Boonham, Neil; Tomlinson, Jenny; Barker, Ian (September 2006). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7087944">"Advances in molecular phytodiagnostics - new solutions for old problems"</a>. <i>European Journal of Plant Pathology</i>. <b>116</b> (1): 1–19. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2006EJPP..116....1M">2006EJPP..116....1M</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2Fs10658-006-9037-0">10.1007/s10658-006-9037-0</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7087944">7087944</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/32214677">32214677</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=European+Journal+of+Plant+Pathology&rft.atitle=Advances+in+molecular+phytodiagnostics+-+new+solutions+for+old+problems&rft.volume=116&rft.issue=1&rft.pages=1-19&rft.date=2006-09&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7087944%23id-name%3DPMC&rft_id=info%3Apmid%2F32214677&rft_id=info%3Adoi%2F10.1007%2Fs10658-006-9037-0&rft_id=info%3Abibcode%2F2006EJPP..116....1M&rft.aulast=Mumford&rft.aufirst=Rick&rft.au=Boonham%2C+Neil&rft.au=Tomlinson%2C+Jenny&rft.au=Barker%2C+Ian&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7087944&rfr_id=info%3Asid%2Fen.wikipedia.org%3APlant+pathology" class="Z3988"></span></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFVenbruxCrauwelsRediers2023" class="citation journal cs1">Venbrux, Marc; Crauwels, Sam; Rediers, Hans (8 May 2023). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10200959">"Current and emerging trends in techniques for plant pathogen detection"</a>. <i>Frontiers in Plant Science</i>. <b>14</b>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3389%2Ffpls.2023.1120968">10.3389/fpls.2023.1120968</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC10200959">10200959</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/37223788">37223788</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Frontiers+in+Plant+Science&rft.atitle=Current+and+emerging+trends+in+techniques+for+plant+pathogen+detection&rft.volume=14&rft.date=2023-05-08&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC10200959%23id-name%3DPMC&rft_id=info%3Apmid%2F37223788&rft_id=info%3Adoi%2F10.3389%2Ffpls.2023.1120968&rft.aulast=Venbrux&rft.aufirst=Marc&rft.au=Crauwels%2C+Sam&rft.au=Rediers%2C+Hans&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC10200959&rfr_id=info%3Asid%2Fen.wikipedia.org%3APlant+pathology" class="Z3988"></span></span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-15">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRyderJones1991" class="citation journal cs1">Ryder, Mh; Jones, Da (1991). "Biological Control of Crown Gall Using Using Agrobacterium Strains K84 and K1026". <i><a href="/wiki/Functional_Plant_Biology" title="Functional Plant Biology">Functional Plant Biology</a></i>. <b>18</b> (5): 571. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1071%2Fpp9910571">10.1071/pp9910571</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Functional+Plant+Biology&rft.atitle=Biological+Control+of+Crown+Gall+Using+Using+Agrobacterium+Strains+K84+and+K1026&rft.volume=18&rft.issue=5&rft.pages=571&rft.date=1991&rft_id=info%3Adoi%2F10.1071%2Fpp9910571&rft.aulast=Ryder&rft.aufirst=Mh&rft.au=Jones%2C+Da&rfr_id=info%3Asid%2Fen.wikipedia.org%3APlant+pathology" class="Z3988"></span></span> </li> <li id="cite_note-Ainsworth-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-Ainsworth_16-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAisnworth1981" class="citation book cs1">Aisnworth, Geoffrey Clough (1981). <i>Introduction to the History of Plant Pathology</i>. Cambridge University Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-521-23032-2" title="Special:BookSources/978-0-521-23032-2"><bdi>978-0-521-23032-2</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Introduction+to+the+History+of+Plant+Pathology&rft.pub=Cambridge+University+Press&rft.date=1981&rft.isbn=978-0-521-23032-2&rft.aulast=Aisnworth&rft.aufirst=Geoffrey+Clough&rfr_id=info%3Asid%2Fen.wikipedia.org%3APlant+pathology" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plant_pathology&action=edit&section=12" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239549316">.mw-parser-output .refbegin{margin-bottom:0.5em}.mw-parser-output .refbegin-hanging-indents>ul{margin-left:0}.mw-parser-output .refbegin-hanging-indents>ul>li{margin-left:0;padding-left:3.2em;text-indent:-3.2em}.mw-parser-output .refbegin-hanging-indents ul,.mw-parser-output .refbegin-hanging-indents ul li{list-style:none}@media(max-width:720px){.mw-parser-output .refbegin-hanging-indents>ul>li{padding-left:1.6em;text-indent:-1.6em}}.mw-parser-output .refbegin-columns{margin-top:0.3em}.mw-parser-output .refbegin-columns ul{margin-top:0}.mw-parser-output .refbegin-columns li{page-break-inside:avoid;break-inside:avoid-column}@media screen{.mw-parser-output .refbegin{font-size:90%}}</style><div class="refbegin refbegin-columns references-column-width" style="column-width: 30em"> <ul><li><a rel="nofollow" class="external text" href="http://www.isppweb.org/">International Society for Plant Pathology</a></li> <li><a rel="nofollow" class="external text" href="http://www.apsnet.org/">American Phytopathological Society</a></li> <li><a rel="nofollow" class="external text" href="http://www.bspp.org.uk/">British Society for Plant Pathology</a></li> <li><a rel="nofollow" class="external text" href="http://www.plantmanagementnetwork.org/php/">Plant Health Progress, Online journal of applied plant pathology</a></li> <li><a rel="nofollow" class="external text" href="http://www.pnwfungi.org/">Pacific Northwest Fungi, online mycology journal with papers on fungal plant pathogens</a></li> <li><a rel="nofollow" class="external text" href="http://eppws.nmsu.edu/">New Mexico State University Department of Entomology Plant Pathology and Weed Science</a></li> <li><a rel="nofollow" class="external text" href="http://www.phibase.org/">Pathogen Host Interactions Database (PHI-base)</a></li></ul> </div> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output 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style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Agriculture_footer" title="Template:Agriculture footer"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Agriculture_footer" title="Template talk:Agriculture footer"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Agriculture_footer" title="Special:EditPage/Template:Agriculture footer"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Agriculture" style="font-size:114%;margin:0 4em"><a href="/wiki/Agriculture" title="Agriculture">Agriculture</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li><a href="/wiki/Outline_of_agriculture" title="Outline of agriculture">Outline</a></li> <li><a href="/wiki/History_of_agriculture" title="History of agriculture">History</a></li> <li><a href="/wiki/Index_of_agriculture_articles" title="Index of agriculture articles">Index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Occupations</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agriculturist" title="Agriculturist">Agriculturist</a></li> <li><a href="/wiki/Agricultural_engineering" title="Agricultural engineering">Agricultural Engineer</a></li> <li><a href="/wiki/Farmer" title="Farmer">Farmer</a></li> <li><a href="/wiki/Farmworker" title="Farmworker">Farm worker</a></li> <li><a href="/wiki/Herder" title="Herder">Herder</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="7" style="width:1px;padding:0 0 0 2px"><div><br /></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Food_system" title="Food system">General</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"><div class="excerpt-block"><div class="excerpt"> <ul><li><a href="/wiki/Agribusiness" title="Agribusiness">Agribusiness</a></li> <li><a href="/wiki/Agricultural_cooperative" title="Agricultural cooperative">Agricultural cooperative</a></li> <li><a href="/wiki/Agricultural_supply_store" class="mw-redirect" title="Agricultural supply store">Agricultural supplies</a></li> <li><a href="/wiki/Agricultural_science" title="Agricultural science">Agricultural science</a></li> <li><a href="/wiki/Agricultural_engineering" title="Agricultural engineering">Agricultural engineering</a></li> <li><a href="/wiki/Agricultural_technology" title="Agricultural technology">Agricultural technology</a> <ul><li><a href="/wiki/Digital_Agriculture" class="mw-redirect" title="Digital Agriculture">Digital</a></li> <li><a href="/wiki/Agricultural_biotechnology" title="Agricultural biotechnology">Biotechnology</a></li></ul></li> <li><a href="/wiki/Agroforestry" title="Agroforestry">Agroforestry</a></li> <li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Animal_husbandry" title="Animal husbandry">Animal husbandry</a></li> <li><a href="/wiki/Animal-free_agriculture" title="Animal-free agriculture">Animal-free agriculture</a></li> <li><a href="/wiki/Cash_crop" title="Cash crop">Cash crop</a></li> <li><a href="/wiki/Cellular_agriculture" title="Cellular agriculture">Cellular agriculture</a></li> <li><a href="/wiki/Cultural_methods" title="Cultural methods">Cultural methods</a></li> <li><a href="/wiki/Contract_farming" title="Contract farming">Contract farming</a></li> <li><a href="/wiki/Extensive_farming" title="Extensive farming">Extensive farming</a></li> <li><a href="/wiki/Farm" title="Farm">Farm</a></li> <li><a href="/wiki/Farmhouse" title="Farmhouse">Farmhouse</a></li> <li><a href="/wiki/Feed_ratio" title="Feed ratio">Feed ratio</a></li> <li><a href="/wiki/Free_range" title="Free range">Free range</a></li> <li><a href="/wiki/Horticulture" title="Horticulture">Horticulture</a></li> <li><a href="/wiki/Intensive_farming" title="Intensive farming">Intensive farming</a> <ul><li><a href="/wiki/Intensive_animal_farming" title="Intensive animal farming">animals</a></li> <li><a href="/wiki/Intensive_pig_farming" title="Intensive pig farming">pigs</a></li> <li><a href="/wiki/Intensive_crop_farming" title="Intensive crop farming">crops</a></li></ul></li> <li><a href="/wiki/Mechanised_agriculture" title="Mechanised agriculture">Mechanised agriculture</a></li> <li><a href="/wiki/Organic_farming" title="Organic farming">Organic farming</a></li> <li><a href="/wiki/Paludiculture" title="Paludiculture">Paludiculture</a></li> <li><a href="/wiki/Permaculture" title="Permaculture">Permaculture</a></li> <li><a href="/wiki/Plantation" title="Plantation">Plantation</a></li> <li><a href="/wiki/Polyculture" title="Polyculture">Polyculture</a> <ul><li><a href="/wiki/Rice-duck_farming" title="Rice-duck farming">Rice-duck farming</a></li> <li><a href="/wiki/Rice-fish_system" title="Rice-fish system">Rice-fish system</a></li></ul></li> <li><a href="/wiki/Sustainable_agriculture" title="Sustainable agriculture">Sustainable agriculture</a></li> <li><a href="/wiki/Sustainable_food_system" title="Sustainable food system">Sustainable food system</a></li> <li><a href="/wiki/List_of_agricultural_universities_and_colleges" title="List of agricultural universities and colleges">Universities and colleges</a></li> <li><a href="/wiki/Urban_agriculture" title="Urban agriculture">Urban agriculture</a></li></ul></div></div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/History_of_agriculture" title="History of agriculture">History</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <li><a href="/wiki/Ancient_Egyptian_agriculture" title="Ancient Egyptian agriculture">Ancient Egypt</a></li> <li><a href="/wiki/Agriculture_in_ancient_Greece" title="Agriculture in ancient Greece">Ancient Greece</a></li> <li><a href="/wiki/Agriculture_in_ancient_Rome" title="Agriculture in ancient Rome">Ancient Rome</a></li> <li><a href="/wiki/Domesticated_plants_and_animals_of_Austronesia" title="Domesticated plants and animals of Austronesia">Austronesia</a></li> <li><a href="/wiki/History_of_agricultural_science" title="History of agricultural science">Agricultural science</a></li> <li><a href="/wiki/Agricultural_History_Review" title="Agricultural History Review">Agricultural History Review</a></li> <li><a href="/wiki/Arab_Agricultural_Revolution" title="Arab Agricultural Revolution">Arab Agricultural Revolution</a></li> <li><a href="/wiki/History_of_agriculture_in_Argentina" title="History of agriculture in Argentina">Argentina</a></li> <li><a href="/wiki/History_of_agriculture_in_Canada" title="History of agriculture in Canada">Canada</a></li> <li><a href="/wiki/History_of_agriculture_in_China" title="History of agriculture in China">China</a></li> <li><a href="/wiki/History_of_agriculture_in_Chile" title="History of agriculture in Chile">Chile</a></li> <li><a href="/wiki/Columbian_exchange" title="Columbian exchange">Columbian exchange</a></li> <li><a href="/wiki/Green_Revolution" title="Green Revolution">Green Revolution</a></li> <li><a href="/wiki/History_of_agriculture_in_the_Indian_subcontinent" title="History of agriculture in the Indian subcontinent">Indian subcontinent</a></li> <li><a href="/wiki/Agriculture_in_Mesoamerica" title="Agriculture in Mesoamerica">Mesoamerica</a></li> <li><a href="/wiki/Agriculture_in_the_Middle_Ages" title="Agriculture in the Middle Ages">Middle Ages</a></li> <li><a href="/wiki/Neolithic_Revolution" title="Neolithic Revolution">Neolithic Revolution</a></li> <li><a href="/wiki/History_of_organic_farming" title="History of organic farming">Organic farming</a></li> <li><a href="/wiki/History_of_agriculture_in_Palestine" title="History of agriculture in Palestine">Palestine</a></li> <li><a href="/wiki/Agricultural_history_of_Peru" title="Agricultural history of Peru">Peru</a></li> <li>United Kingdom <ul><li><a href="/wiki/British_Agricultural_Revolution" title="British Agricultural Revolution">British Agricultural Revolution</a></li> <li><a href="/wiki/History_of_agriculture_in_Cheshire" title="History of agriculture in Cheshire">Cheshire</a></li> <li><a href="/wiki/History_of_agriculture_in_Scotland" title="History of agriculture in Scotland">Scotland</a></li></ul></li> <li><a href="/wiki/History_of_agriculture_in_the_United_States" title="History of agriculture in the United States">United States</a> <ul><li><a href="/wiki/History_of_African-American_agriculture" title="History of African-American agriculture">African-American</a></li> <li><a href="/wiki/History_of_agriculture_in_California" title="History of agriculture in California">California</a></li></ul></li> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Farm" title="Farm">Farming Types</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agrivoltaics" title="Agrivoltaics">Agrivoltaic</a></li> <li><a href="/wiki/Aquaculture" title="Aquaculture">Aquaculture</a></li> <li><a href="/wiki/Cattle" title="Cattle">Cattle</a></li> <li><a href="/wiki/Dairy_farming" title="Dairy farming">Dairy farming</a></li> <li><a href="/wiki/Fur_farming" title="Fur farming">Fur farming</a></li> <li><a href="/wiki/Goat_farming" title="Goat farming">Goat farming</a></li> <li><a href="/wiki/Grazing" title="Grazing">Grazing</a> <ul><li><a href="/wiki/Convertible_husbandry" title="Convertible husbandry">Convertible husbandry</a></li> <li><a href="/wiki/Rotational_grazing" title="Rotational grazing">Rotational grazing</a></li></ul></li> <li><a href="/wiki/Hydroponics" title="Hydroponics">Hydroponics</a></li> <li><a href="/wiki/Insect_farming" title="Insect farming">Insect farming</a></li> <li><a href="/wiki/Livestock" title="Livestock">Livestock</a> <ul><li><a href="/wiki/Pasture" title="Pasture">Pasture</a></li></ul></li> <li><a href="/wiki/Mixed_farming" title="Mixed farming">Mixed</a></li> <li><a href="/wiki/Monoculture" title="Monoculture">Monoculture</a></li> <li><a href="/wiki/Paddy_field" title="Paddy field">Paddy field</a></li> <li><a href="/wiki/Pastoral_farming" class="mw-redirect" title="Pastoral farming">Pastoral</a> <ul><li><a href="/wiki/Bocage" title="Bocage">Bocage</a></li></ul></li> <li><a href="/wiki/Pig_farming" title="Pig farming">Pig farming</a></li> <li><a href="/wiki/Poultry_farming" title="Poultry farming">Poultry farming</a></li> <li><a href="/wiki/Ranch" title="Ranch">Ranch</a></li> <li><a href="/wiki/Orchard" title="Orchard">Orchards</a></li> <li><a href="/wiki/Subsistence_agriculture" title="Subsistence agriculture">Subsistence agriculture</a></li> <li><a href="/wiki/Sheep_farming" title="Sheep farming">Sheep farming</a></li> <li><a href="/wiki/Terrace_(earthworks)" title="Terrace (earthworks)">Terrace</a></li> <li><a href="/wiki/Wildlife_farming" title="Wildlife farming">Wildlife farming</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Environmental_impact_of_agriculture" title="Environmental impact of agriculture">Environmental<br />impact</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agricultural_expansion" title="Agricultural expansion">Agricultural expansion</a></li> <li><a href="/wiki/Agricultural_pollution" title="Agricultural pollution">Agricultural pollution</a></li> <li><a href="/wiki/Agricultural_wastewater" class="mw-redirect" title="Agricultural wastewater">Agricultural wastewater</a></li> <li><a href="/wiki/Overgrazing" title="Overgrazing">Overgrazing</a></li> <li><a href="/wiki/Environmental_impact_of_irrigation" title="Environmental impact of irrigation">Environmental impact of irrigation</a> <ul><li><a href="/wiki/Overdrafting" title="Overdrafting">Overdrafting</a></li></ul></li> <li><a href="/wiki/Effects_of_climate_change_on_agriculture" title="Effects of climate change on agriculture">Climate change and agriculture</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Categories</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Category:Agricultural_machinery" title="Category:Agricultural machinery">Agricultural machinery</a></li> <li><a href="/wiki/Category:Agriculture_by_country" title="Category:Agriculture by country">Agriculture by country</a></li> <li><a href="/wiki/Category:Agriculture_companies" title="Category:Agriculture companies">Agriculture companies</a></li> <li><a href="/wiki/Category:Biotechnology" title="Category:Biotechnology">Biotechnology</a></li> <li><a href="/wiki/Category:History_of_agriculture" title="Category:History of agriculture">History of agriculture</a></li> <li><a href="/wiki/Category:Livestock" title="Category:Livestock">Livestock</a></li> <li><a href="/wiki/Category:Meat_industry" title="Category:Meat industry">Meat industry</a></li> <li><a href="/wiki/Category:Poultry_farming" title="Category:Poultry farming">Poultry farming</a></li> <li><a href="/wiki/Category:Agriculture_and_the_environment" title="Category:Agriculture and the environment">Agriculture and the environment</a></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Lists</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"><div class="excerpt-block"><div class="excerpt"> <li><a href="/wiki/Agriculturist" title="Agriculturist">Agriculturist profession</a></li> <li><a href="/wiki/List_of_agricultural_machinery" title="List of agricultural machinery">Agricultural machinery</a></li> <li><a href="/wiki/List_of_food_origins" title="List of food origins">Food origins</a></li> <li><a href="/wiki/List_of_agriculture_ministries" title="List of agriculture ministries">Government ministries</a></li> <li><a href="/wiki/List_of_agricultural_universities_and_colleges" title="List of agricultural universities and colleges">Universities and colleges</a></li></div></div></div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <b><a href="/wiki/Category:Agriculture" title="Category:Agriculture">Category</a></b></li> <li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" class="mw-file-description" title="Portal"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/16px-Symbol_portal_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/23px-Symbol_portal_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/31px-Symbol_portal_class.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span> <b><a href="/wiki/Portal:Agriculture" title="Portal:Agriculture">Portal</a></b></li> <li><span class="noviewer" typeof="mw:File"><span title="Commons page"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span> <b><a href="https://commons.wikimedia.org/wiki/Agriculture" class="extiw" title="commons:Agriculture">Commons</a></b></li> <li><span class="noviewer" typeof="mw:File"><span title="WikiProject"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/People_icon.svg/16px-People_icon.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/People_icon.svg/24px-People_icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/37/People_icon.svg/32px-People_icon.svg.png 2x" data-file-width="100" data-file-height="100" /></span></span> <b><a href="/wiki/Wikipedia:WikiProject_Agriculture" title="Wikipedia:WikiProject Agriculture">Wikiproject</a></b></li> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"></div><div role="navigation" class="navbox" aria-labelledby="Botany" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background: #C3EEC3;;background: #90EE90;"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Botany" title="Template:Botany"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Botany" title="Template talk:Botany"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Botany" title="Special:EditPage/Template:Botany"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Botany" style="font-size:114%;margin:0 4em"><a href="/wiki/Botany" title="Botany">Botany</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: #C3EEC3;"><div> <ul><li><a href="/wiki/History_of_botany" title="History of botany">History</a></li> <li><a href="/wiki/Outline_of_botany" title="Outline of botany">Outline</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Branches_of_botany" title="Branches of botany">Subdisciplines</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaeobotany" class="mw-redirect" title="Archaeobotany">Archaeobotany</a></li> <li><a href="/wiki/Astrobotany" title="Astrobotany">Astrobotany</a></li> <li><a href="/wiki/Bryology" title="Bryology">Bryology</a></li> <li><a href="/wiki/Dendrology" title="Dendrology">Dendrology</a></li> <li><a href="/wiki/Ethnobotany" title="Ethnobotany">Ethnobotany</a></li> <li><a href="/wiki/Paleobotany" title="Paleobotany">Paleobotany</a></li> <li><a href="/wiki/Phycology" title="Phycology">Phycology</a></li> <li><a href="/wiki/Phytochemistry" title="Phytochemistry">Phytochemistry</a></li> <li><a href="/wiki/Phytogeography" title="Phytogeography">Phytogeography</a> <ul><li><a href="/wiki/Geobotanical_prospecting" title="Geobotanical prospecting">Geobotany</a></li></ul></li> <li><a href="/wiki/Plant_anatomy" title="Plant anatomy">Plant anatomy</a></li> <li><a href="/wiki/Plant_ecology" title="Plant ecology">Plant ecology</a></li> <li><a class="mw-selflink selflink">Plant pathology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant" title="Plant">Plant</a> groups</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Algae" title="Algae">Algae</a></li> <li><a href="/wiki/Archaeplastida" title="Archaeplastida">Archaeplastida</a></li> <li><a href="/wiki/Bryophyte" title="Bryophyte">Bryophyte</a></li> <li><a href="/wiki/Non-vascular_plant" title="Non-vascular plant">Non-vascular plants</a></li> <li><a href="/wiki/Vascular_plant" title="Vascular plant">Vascular plants</a></li> <li><a href="/wiki/Fern" title="Fern">Fern</a></li> <li><a href="/wiki/Lycophyte" title="Lycophyte">Lycophyte</a></li> <li><a href="/wiki/Spermatophyte" class="mw-redirect" title="Spermatophyte">Spermatophytes</a></li> <li><a href="/wiki/Gymnosperm" title="Gymnosperm">Gymnosperm</a></li> <li><a href="/wiki/Flowering_plant" title="Flowering plant">Angiosperm</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_anatomy" title="Plant anatomy">Plant anatomy</a> <div class="hlist"><ul><li><a href="/wiki/Plant_morphology" title="Plant morphology">Plant morphology</a><br /><span class="nobold">(<a href="/wiki/Glossary_of_plant_morphology" title="Glossary of plant morphology">glossary</a>)</span></li></ul></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%"><a href="/wiki/Plant_cell" title="Plant cell">Plant cells</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cell_wall" title="Cell wall">Cell wall</a></li> <li><a href="/wiki/Phragmoplast" title="Phragmoplast">Phragmoplast</a></li> <li><a href="/wiki/Plastid" title="Plastid">Plastid</a></li> <li><a href="/wiki/Plasmodesma" title="Plasmodesma">Plasmodesma</a></li> <li><a href="/wiki/Vacuole" title="Vacuole">Vacuole</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%"><a href="/wiki/Tissue_(biology)" title="Tissue (biology)">Tissues</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cork_cambium" title="Cork cambium">Cork</a></li> <li><a href="/wiki/Ground_tissue" title="Ground tissue">Ground tissue</a> <ul><li><a href="/wiki/Leaf#Mesophyll" title="Leaf">Mesophyll</a></li></ul></li> <li><a href="/wiki/Meristem" title="Meristem">Meristem</a></li> <li><a href="/wiki/Storage_organ" title="Storage organ">Storage organs</a></li> <li><a href="/wiki/Vascular_tissue" title="Vascular tissue">Vascular tissue</a> <ul><li><a href="/wiki/Vascular_bundle" title="Vascular bundle">Vascular bundle</a></li></ul></li> <li><a href="/wiki/Wood" title="Wood">Wood</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%">Vegetative</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bulb" title="Bulb">Bulb</a></li> <li><a href="/wiki/Root" title="Root">Root</a></li> <li><a href="/wiki/Rhizoid" title="Rhizoid">Rhizoid</a></li> <li><a href="/wiki/Rhizome" title="Rhizome">Rhizome</a></li> <li><a href="/wiki/Shoot_(botany)" title="Shoot (botany)">Shoot</a> <ul><li><a href="/wiki/Bud" title="Bud">Bud</a></li> <li><a href="/wiki/Leaf" title="Leaf">Leaf</a> <ul><li><a href="/wiki/Cataphyll" title="Cataphyll">Cataphyll</a></li> <li><a href="/wiki/Petiole_(botany)" title="Petiole (botany)">Petiole</a></li></ul></li> <li><a href="/wiki/Sessility_(botany)" title="Sessility (botany)">Sessility</a></li> <li><a href="/wiki/Plant_stem" title="Plant stem">Stem</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%"><a href="/wiki/Plant_reproductive_morphology" title="Plant reproductive morphology">Reproductive</a><br />(incl. Flower)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archegonium" title="Archegonium">Archegonium</a></li> <li><a href="/wiki/Antheridium" title="Antheridium">Antheridium</a></li> <li><a href="/wiki/Stamen" title="Stamen">Androecium</a> <ul><li><a href="/wiki/Pollen" title="Pollen">Pollen</a></li> <li><a href="/wiki/Stamen" title="Stamen">Stamen</a> <ul><li><a href="/wiki/Anther" class="mw-redirect" title="Anther">Anther</a></li> <li><a href="/wiki/Stamen#Morphology_and_terminology" title="Stamen">Filament</a></li></ul></li> <li><a href="/wiki/Staminode" title="Staminode">Staminode</a></li> <li><a href="/wiki/Tapetum_(botany)" title="Tapetum (botany)">Tapetum</a></li></ul></li> <li><a href="/wiki/Flower" title="Flower">Flower</a> <ul><li><a href="/wiki/Aestivation_(botany)" title="Aestivation (botany)">Aestivation</a></li> <li><a href="/wiki/ABC_model_of_flower_development" title="ABC model of flower development">Flower development</a></li> <li><a href="/wiki/Floral_diagram" title="Floral diagram">Floral diagram</a></li> <li><a href="/wiki/Floral_formula" title="Floral formula">Floral formula</a></li> <li><a href="/wiki/Floral_symmetry" title="Floral symmetry">Floral symmetry</a></li> <li><a href="/wiki/Whorl_(botany)" title="Whorl (botany)">Whorl</a></li></ul></li> <li><a href="/wiki/Fruit" title="Fruit">Fruit</a> <ul><li><a href="/wiki/Fruit_anatomy" class="mw-redirect" title="Fruit anatomy">Anatomy</a></li> <li><a href="/wiki/Berry_(botany)" title="Berry (botany)">Berry</a></li> <li><a href="/wiki/Capsule_(fruit)" title="Capsule (fruit)">Capsule</a></li> <li><a href="/wiki/Nut_(fruit)" title="Nut (fruit)">Nut</a></li> <li><a href="/wiki/Pyrena" title="Pyrena">Pyrena</a></li> <li><a href="/wiki/Seed" title="Seed">Seed</a> <ul><li><a href="/wiki/Seed_dispersal" title="Seed dispersal">Dispersal</a></li> <li><a href="/wiki/Endosperm" title="Endosperm">Endosperm</a></li></ul></li></ul></li> <li><a href="/wiki/Gametophyte" title="Gametophyte">Gametophyte</a></li> <li><a href="/wiki/Column_(botany)" title="Column (botany)">Gynandrium</a></li> <li><a href="/wiki/Gynoecium" title="Gynoecium">Gynoecium</a> <ul><li><a href="/wiki/Carpel" class="mw-redirect" title="Carpel">Carpel</a> <ul><li><a href="/wiki/Ovary_(botany)" title="Ovary (botany)">Ovary</a> <ul><li><a href="/wiki/Locule" title="Locule">Locule</a></li> <li><a href="/wiki/Ovule" title="Ovule">Ovule</a></li></ul></li> <li><a href="/wiki/Stigma_(botany)" title="Stigma (botany)">Stigma</a></li> <li><a href="/wiki/Style_(botany)" title="Style (botany)">Style</a></li></ul></li></ul></li> <li><a href="/wiki/Hypanthium" title="Hypanthium">Hypanthium (Floral cup)</a></li> <li><a href="/wiki/Inflorescence" title="Inflorescence">Inflorescence</a> <ul><li><a href="/wiki/Bract" title="Bract">Bract</a></li> <li><a href="/wiki/Pedicel_(botany)" title="Pedicel (botany)">Pedicellate</a></li> <li><a href="/wiki/Raceme" title="Raceme">Raceme</a></li> <li><a href="/wiki/Umbel" title="Umbel">Umbel</a></li></ul></li> <li><a href="/wiki/Perianth" title="Perianth">Perianth</a> <ul><li><a href="/wiki/Tepal" title="Tepal">Tepal</a></li> <li><a href="/wiki/Petal" title="Petal">Petal</a></li> <li><a href="/wiki/Sepal" title="Sepal">Sepal</a></li></ul></li> <li><a href="/wiki/Embryo#Plant_embryos" title="Embryo">Plant embryo</a></li> <li><a href="/wiki/Receptacle_(botany)" title="Receptacle (botany)">Receptacle</a></li> <li><a href="/wiki/Sporophyll" title="Sporophyll">Sporophyll</a></li> <li><a href="/wiki/Sporophyte" title="Sporophyte">Sporophyte</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #D4EED4;;width:1%">Surface structures</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Plant_cuticle" title="Plant cuticle">Cuticle</a></li> <li><a href="/wiki/Epicuticular_wax" title="Epicuticular wax">Epicuticular wax</a></li> <li><a href="/wiki/Epidermis_(botany)" title="Epidermis (botany)">Epidermis</a></li> <li><a href="/wiki/Nectar" title="Nectar">Nectar</a></li> <li><a href="/wiki/Stoma" title="Stoma">Stoma</a></li> <li><a href="/wiki/Thorns,_spines,_and_prickles" title="Thorns, spines, and prickles">Thorns, spines, and prickles</a></li> <li><a href="/wiki/Trichome" title="Trichome">Trichome</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_physiology" title="Plant physiology">Plant physiology</a><br />Materials</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aleurone" title="Aleurone">Aleurone</a></li> <li><a href="/wiki/Apical_dominance" title="Apical dominance">Apical dominance</a></li> <li><a href="/wiki/Bulk_movement" title="Bulk movement">Bulk flow</a></li> <li><a href="/wiki/Cellulose" title="Cellulose">Cellulose</a></li> <li><a href="/wiki/Plant_nutrition" title="Plant nutrition">Nutrition</a></li> <li><a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a> <ul><li><a href="/wiki/Chlorophyll" title="Chlorophyll">Chlorophyll</a></li></ul></li> <li><a href="/wiki/Phytomelanin" title="Phytomelanin">Phytomelanin</a></li> <li><a href="/wiki/Plant_hormone" title="Plant hormone">Plant hormones</a></li> <li><a href="/wiki/Respiratory_system#Plants" title="Respiratory system">Respiration</a> <ul><li><a href="/wiki/Gas_exchange#Plants" title="Gas exchange">Gas Exchange</a></li> <li><a href="/wiki/Cellular_respiration" title="Cellular respiration">Cellular respiration</a></li></ul></li> <li><a href="/wiki/Sap" title="Sap">Sap</a></li> <li><a href="/wiki/Starch" title="Starch">Starch</a></li> <li><a href="/wiki/Sugar" title="Sugar">Sugar</a></li> <li><a href="/wiki/Transpiration" title="Transpiration">Transpiration</a></li> <li><a href="/wiki/Turgor_pressure" title="Turgor pressure">Turgor pressure</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%">Plant growth<br />and habit</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Habit_(biology)#Structure" title="Habit (biology)">Habit</a> <ul><li><a href="/wiki/Cushion_plant" title="Cushion plant">Cushion plants</a></li> <li><a href="/wiki/Rosette_(botany)" title="Rosette (botany)">Rosettes</a></li> <li><a href="/wiki/Shrub" title="Shrub">Shrubs</a> <ul><li><a href="/wiki/Prostrate_shrub" title="Prostrate shrub">Prostrate shrubs</a></li> <li><a href="/wiki/Subshrub" title="Subshrub">Subshrubs</a></li></ul></li> <li><a href="/wiki/Succulent_plant" title="Succulent plant">Succulent plants</a></li> <li><a href="/wiki/Tree" title="Tree">Trees</a></li> <li><a href="/wiki/Vine" title="Vine">Vines</a> <ul><li><a href="/wiki/Liana" title="Liana">Lianas</a></li></ul></li></ul></li> <li><a href="/wiki/Herbaceous_plant" title="Herbaceous plant">Herbaceous plants</a></li> <li><a href="/wiki/Secondary_growth" title="Secondary growth">Secondary growth</a></li> <li><a href="/wiki/Woody_plant" title="Woody plant">Woody plants</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_reproduction" title="Plant reproduction">Reproduction</a> <div class="hlist"><ul><li><a href="/wiki/Plant_evolution" title="Plant evolution">Evolution</a></li><li><a href="/wiki/Plant_ecology" title="Plant ecology">Ecology</a></li></ul></div></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Alternation_of_generations" title="Alternation of generations">Alternation of generations</a></li> <li><a href="/wiki/Double_fertilization" title="Double fertilization">Double fertilization</a></li> <li><a href="/wiki/Plant_evolutionary_developmental_biology" title="Plant evolutionary developmental biology">Evolutionary development</a></li> <li><a href="/wiki/Evolutionary_history_of_plants" title="Evolutionary history of plants">Evolutionary history</a> <ul><li><a href="/wiki/Timeline_of_plant_evolution" title="Timeline of plant evolution">timeline</a></li></ul></li> <li><a href="/wiki/Flora" title="Flora">Flora</a></li> <li><a href="/wiki/Germination" title="Germination">Germination</a></li> <li><a href="/wiki/Pollination" title="Pollination">Pollination</a> <ul><li><a href="/wiki/Artificial_pollination" class="mw-redirect" title="Artificial pollination">Artificial</a></li> <li><a href="/wiki/Pollinator" title="Pollinator">Pollinators</a></li> <li><a href="/wiki/Pollen_tube" title="Pollen tube">Pollen tube</a></li> <li><a href="/wiki/Self-pollination" title="Self-pollination">Self</a></li></ul></li> <li><a href="/wiki/Sporangium" title="Sporangium">Sporangium</a> <ul><li><a href="/wiki/Microsporangia" class="mw-redirect" title="Microsporangia">Microsporangia</a> <ul><li><a href="/wiki/Microspore" title="Microspore">Microspore</a></li></ul></li> <li><a href="/wiki/Sporangium" title="Sporangium">Megasporangium</a> <ul><li><a href="/wiki/Megaspore" title="Megaspore">Megaspore</a></li></ul></li> <li><a href="/wiki/Spore" title="Spore">Spore</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><a href="/wiki/Plant_taxonomy" title="Plant taxonomy">Plant taxonomy</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Biological classification</a></li> <li><a href="/wiki/Botanical_nomenclature" title="Botanical nomenclature">Botanical nomenclature</a> <ul><li><a href="/wiki/Botanical_name" title="Botanical name">Botanical name</a></li> <li><a href="/wiki/Correct_name" title="Correct name">Correct name</a></li> <li><a href="/wiki/Author_citation_(botany)" title="Author citation (botany)">Author citation</a></li> <li><a href="/wiki/International_Code_of_Nomenclature_for_algae,_fungi,_and_plants" title="International Code of Nomenclature for algae, fungi, and plants">International Code of Nomenclature (ICN)</a></li> <li><a href="/wiki/International_Code_of_Nomenclature_for_Cultivated_Plants" title="International Code of Nomenclature for Cultivated Plants">ICN for Cultivated Plants (ICNCP)</a></li></ul></li> <li><a href="/wiki/Cultivated_plant_taxonomy" title="Cultivated plant taxonomy">Cultivated plant taxonomy</a> <ul><li><a href="/wiki/Citrus_taxonomy" title="Citrus taxonomy">Citrus taxonomy</a></li> <li><a href="/wiki/Cultigen" title="Cultigen">Cultigen</a> <ul><li><a href="/wiki/Cultivar" title="Cultivar">Cultivar</a></li> <li><a href="/wiki/Cultivar_group" title="Cultivar group">Group</a></li> <li><a href="/wiki/Grex_(horticulture)" title="Grex (horticulture)">Grex</a></li></ul></li></ul></li> <li><a href="/wiki/History_of_plant_systematics" title="History of plant systematics">History of plant systematics</a></li> <li><a href="/wiki/Herbarium" title="Herbarium">Herbarium</a></li> <li><a href="/wiki/International_Association_for_Plant_Taxonomy" title="International Association for Plant Taxonomy">International Association for Plant Taxonomy</a> (IAPT)</li> <li><a href="/wiki/List_of_systems_of_plant_taxonomy" title="List of systems of plant taxonomy">Plant taxonomy systems</a></li> <li><a href="/wiki/Taxonomic_rank" title="Taxonomic rank">Taxonomic rank</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%">Practice</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Floriculture" title="Floriculture">Floriculture</a></li> <li><a href="/wiki/Forestry" title="Forestry">Forestry</a></li> <li><a href="/wiki/Horticulture" title="Horticulture">Horticulture</a></li> <li><a href="/wiki/Phytochemical" title="Phytochemical">Phytochemical</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background: #C3EEC3;;width:1%"><div class="hlist"><ul><li>Lists</li><li>Related topics</li></ul></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glossary_of_botanical_terms" title="Glossary of botanical terms">Botanical terms</a></li> <li><a href="/wiki/List_of_botanists" title="List of botanists">Botanists</a> <ul><li><a href="/wiki/List_of_botanists_by_author_abbreviation_(W%E2%80%93Z)" title="List of botanists by author abbreviation (W–Z)">by author abbreviation</a></li></ul></li> <li><a href="/wiki/Botanical_expeditions" title="Botanical expeditions">Botanical expeditions</a></li> <li><a href="/wiki/List_of_individual_trees" title="List of individual trees">Individual trees</a></li> <li><a href="/wiki/Lists_of_plants" title="Lists of plants">Plants</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: #C3EEC3;"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" 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