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Your current IP address is 8.222.208.146. </p> <div class="paragraphbreak" style="margin-top:0.5em"></div><div style="font-size: 16px;"> <p>Even while globally blocked, you will <i>usually</i> still be able to edit pages on <a href="https://meta.wikimedia.org/wiki/" class="extiw" title="m:">Meta-Wiki</a>. </p> </div> <div class="paragraphbreak" style="margin-top:0.5em"></div><div style="font-size: 16px;"> <p>If you believe you were blocked by mistake, you can find additional information and instructions in the <a href="https://meta.wikimedia.org/wiki/Special:MyLanguage/No_open_proxies" class="extiw" title="m:Special:MyLanguage/No open proxies">No open proxies</a> global policy. Otherwise, to discuss the block please <a href="https://meta.wikimedia.org/wiki/Steward_requests/Global" class="extiw" title="m:Steward requests/Global">post a request for review on Meta-Wiki</a>. You could also send an email to the <a href="https://meta.wikimedia.org/wiki/Special:MyLanguage/Stewards" class="extiw" title="m:Special:MyLanguage/Stewards">stewards</a> <a href="https://meta.wikimedia.org/wiki/Special:MyLanguage/VRT" class="extiw" title="m:Special:MyLanguage/VRT">VRT</a> queue at <kbd>stewards@wikimedia.org</kbd> including all above details. </p> </div> <p>Other useful links: <a href="https://meta.wikimedia.org/wiki/Global_blocks" class="extiw" title="m:Global blocks">Global blocks</a> · <a href="/wiki/Help:I_have_been_blocked" title="Help:I have been blocked">Help:I have been blocked</a> </p> </div></li></ul><hr /> <div id="viewsourcetext">You can view and copy the source of this page:</div><textarea readonly="" accesskey="," id="wpTextbox1" cols="80" rows="25" style="" class="mw-editfont-monospace" lang="en" dir="ltr" name="wpTextbox1">== Structure == [[File:TMV structure full.png|thumb|Schematic model of TMV: 1. nucleic acid ([[RNA]]), 2. capsomer protein ([[Protomer (structural biology)|protomer]]), 3. [[capsid]]]]''Tobacco mosaic virus'' has a rod-like appearance. Its [[capsid]] is made from 2130 [[molecules]] of coat protein and one molecule of genomic single strand RNA, 6400 bases long. The coat protein self-assembles into the rod-like helical structure (16.3 proteins per helix turn) around the RNA, which forms a hairpin loop structure (see the [[Electron Microscope|electron micrograph]] above). The structural organization of the virus gives stability.<ref>{{cite journal | vauthors = Caspar DL |title=Assembly and Stability of the Tobacco Mosaic Virus Particle |date= January 1964 | doi = 10.1016/S0065-3233(08)60268-5 | journal = Advances in Protein Chemistry |volume=18 |pages=37–121 | veditors = Anfinsen CB, Anson ML, Edsall JT |publisher=Academic Press |pmid=14151998 |isbn=9780120342181 |language=en }}</ref> The protein monomer consists of 158 [[amino acids]] which are assembled into four main alpha-helices, which are joined by a prominent loop proximal to the axis of the virion. Virions are ~300&nbsp;nm in length and ~18&nbsp;nm in diameter.<ref name="isbn0-7167-1843-X">{{cite book | vauthors = Stryer L | title = Biochemistry | publisher = W.H. Freeman | location = San Francisco | year = 1988 | isbn = 978-0-7167-1843-7 | url-access = registration | url = https://archive.org/details/biochemistry3rdedi00stry }}</ref> Negatively stained electron microphotographs show a distinct inner channel of radius ~2&nbsp;nm. The RNA is located at a radius of ~4&nbsp;nm and is protected from the action of cellular enzymes by the coat protein.<ref name="pmid10212932">{{cite journal | vauthors = Klug A | title = The tobacco mosaic virus particle: structure and assembly | journal = Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences | volume = 354 | issue = 1383 | pages = 531–5 | date = March 1999 | pmid = 10212932 | pmc = 1692534 | doi = 10.1098/rstb.1999.0404 }}</ref> X-ray [[fiber diffraction]] structure of the intact virus was studied based on an [[electron density]] map at 3.6 Å resolution.<ref name="pmid3952490">{{PDB|1VTM}}; {{cite journal | vauthors = Namba K, Stubbs G | title = Structure of tobacco mosaic virus at 3.6 A resolution: implications for assembly | journal = Science | volume = 231 | issue = 4744 | pages = 1401–6 | date = March 1986 | pmid = 3952490 | doi = 10.1126/science.3952490 }}</ref> Inside the capsid helix, near the core, is the coiled RNA molecule, which is made up of 6,395 ±10 nucleotides.<ref name="PNA-Goelet">{{cite journal | vauthors = Goelet P, Lomonossoff GP, Butler PJ, Akam ME, Gait MJ, Karn J | title = Nucleotide sequence of tobacco mosaic virus RNA | journal = Proceedings of the National Academy of Sciences of the United States of America | volume = 79 | issue = 19 | pages = 5818–22 | date = October 1982 | pmid = 6964389 | pmc = 347001 | doi = 10.1073/pnas.79.19.5818 | bibcode = 1982PNAS...79.5818G | doi-access = free }}</ref><ref name="EMBL-EBI">{{cite web |title=Sequence: V01408.1 |url=https://www.ebi.ac.uk/ena/browser/view/V01408.1 |website=European Nucleotide Archive |publisher=[[EMBL]]- [[European Bioinformatics Institute|EBI]] |access-date=28 March 2020 |language=en |quote=International central site for archiving nucleic acid sequence. The reference standard in international science.}}</ref> The structure of the virus plays an important role in the recognition of the viral DNA. This happens due to the formation of an obligatory intermediate produced from a protein allows the virus to recognize a specific RNA hairpin structure.<ref>{{cite journal | vauthors = Klug A | title = The tobacco mosaic virus particle: structure and assembly | journal = Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences | volume = 354 | issue = 1383 | pages = 531–535 | date = March 1999 | pmid = 10212932 | pmc = 1692534 | doi = 10.1098/rstb.1999.0404 }}</ref> The intermediate induces the nucleation of TMV self-assembly by binding with the hairpin structure.<ref>{{cite journal | vauthors = Butler PJ | title = Self-assembly of tobacco mosaic virus: the role of an intermediate aggregate in generating both specificity and speed | journal = Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences | volume = 354 | issue = 1383 | pages = 537–550 | date = March 1999 | pmid = 10212933 | pmc = 1692540 | doi = 10.1098/rstb.1999.0405 }}</ref> </textarea><div class="templatesUsed"></div><p id="mw-returnto">Return to <a href="/wiki/Tobacco_mosaic_virus" title="Tobacco mosaic virus">Tobacco mosaic virus</a>.</p> <noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/wiki/Tobacco_mosaic_virus">https://en.wikipedia.org/wiki/Tobacco_mosaic_virus</a>"</div></div> <div id="catlinks" class="catlinks catlinks-allhidden" data-mw="interface"></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy">Privacy policy</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia:About">About Wikipedia</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:General_disclaimer">Disclaimers</a></li> <li id="footer-places-contact"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us">Contact Wikipedia</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">Code of Conduct</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">Developers</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/en.wikipedia.org">Statistics</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookie statement</a></li> <li id="footer-places-mobileview"><a href="//en.m.wikipedia.org/w/index.php?title=Tobacco_mosaic_virus&action=edit&section=2&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobile view</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-f69cdc8f6-znn46","wgBackendResponseTime":324,"wgPageParseReport":{"limitreport":{"cputime":"0.097","walltime":"0.118","ppvisitednodes":{"value":418,"limit":1000000},"postexpandincludesize":{"value":17740,"limit":2097152},"templateargumentsize":{"value":6556,"limit":2097152},"expansiondepth":{"value":9,"limit":100},"expensivefunctioncount":{"value":0,"limit":500},"unstrip-depth":{"value":0,"limit":20},"unstrip-size":{"value":469,"limit":5000000},"entityaccesscount":{"value":0,"limit":400},"timingprofile":["100.00% 85.195 1 -total"," 99.84% 85.060 2 Template:Blocked_text"," 40.41% 34.428 2 Template:Replace"," 37.83% 32.227 1 Template:Colocationwebhost"," 34.39% 29.295 1 Template:Hidden"," 15.87% 13.519 1 Template:Hidden_begin"," 12.88% 10.974 1 Template:Tlx"," 2.31% 1.970 1 Template:Hidden_end"," 2.04% 1.735 1 MediaWiki:Wikimedia-globalblocking-blockedtext-mistake"," 1.64% 1.394 1 MediaWiki:Wikimedia-globalblocking-blockedtext-mistake-email-steward"]},"scribunto":{"limitreport-timeusage":{"value":"0.020","limit":"10.000"},"limitreport-memusage":{"value":1043316,"limit":52428800}},"cachereport":{"origin":"mw-web.codfw.main-f69cdc8f6-znn46","timestamp":"20241124135631","ttl":2592000,"transientcontent":false}}});});</script> </body> </html>