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<span>Behavior</span> </div> </a> <ul id="toc-Behavior-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Phylogeny_and_systematics" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Phylogeny_and_systematics"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Phylogeny and systematics</span> </div> </a> <ul id="toc-Phylogeny_and_systematics-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Curculionidae</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" 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Available in 59 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-59" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">59 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B3%D9%88%D8%B3%D9%8A%D8%A9_%D8%AD%D9%82%D9%8A%D9%82%D9%8A%D8%A9" title="سوسية حقيقية – Arabic" lang="ar" hreflang="ar" data-title="سوسية حقيقية" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Asturian" lang="ast" hreflang="ast" data-title="Curculionidae" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Azerbaijani" lang="az" hreflang="az" data-title="Curculionidae" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D0%BB%D0%BE%D0%BD%D1%96%D0%BA%D1%96" title="Слонікі – Belarusian" lang="be" hreflang="be" data-title="Слонікі" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bcl mw-list-item"><a href="https://bcl.wikipedia.org/wiki/Ulalo" title="Ulalo – Central Bikol" lang="bcl" hreflang="bcl" data-title="Ulalo" data-language-autonym="Bikol Central" data-language-local-name="Central Bikol" class="interlanguage-link-target"><span>Bikol Central</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A5%D0%BE%D0%B1%D0%BE%D1%82%D0%BD%D0%B8%D0%BA%D0%BE%D0%B2%D0%B8" title="Хоботникови – Bulgarian" lang="bg" hreflang="bg" data-title="Хоботникови" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Curculi%C3%B2nids" title="Curculiònids – Catalan" lang="ca" hreflang="ca" data-title="Curculiònids" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Cebuano" lang="ceb" hreflang="ceb" data-title="Curculionidae" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Nosatcovit%C3%AD" title="Nosatcovití – Czech" lang="cs" hreflang="cs" data-title="Nosatcovití" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Snudebiller" title="Snudebiller – Danish" lang="da" hreflang="da" data-title="Snudebiller" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/R%C3%BCsselk%C3%A4fer" title="Rüsselkäfer – German" lang="de" hreflang="de" data-title="Rüsselkäfer" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/Ts%C3%A9yoo%CA%BC%C3%A1a%C5%82ii" title="Tséyooʼáałii – Navajo" lang="nv" hreflang="nv" data-title="Tséyooʼáałii" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/K%C3%A4rsaklased" title="Kärsaklased – Estonian" lang="et" hreflang="et" data-title="Kärsaklased" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%9A%CE%BF%CF%85%CF%81%CE%BA%CE%BF%CF%85%CE%BB%CE%B9%CE%BF%CE%BD%CE%AF%CE%B4%CE%B5%CF%82" title="Κουρκουλιονίδες – Greek" lang="el" hreflang="el" data-title="Κουρκουλιονίδες" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Spanish" lang="es" hreflang="es" data-title="Curculionidae" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Kurkuliedoj" title="Kurkuliedoj – Esperanto" lang="eo" hreflang="eo" data-title="Kurkuliedoj" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Basque" lang="eu" hreflang="eu" data-title="Curculionidae" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%D9%88%D8%B3%DA%A9%DA%86%D9%87%E2%80%8C%D9%87%D8%A7%DB%8C_%D8%AE%D8%B1%D8%B7%D9%88%D9%85%E2%80%8C%D8%AF%D8%A7%D8%B1" title="سوسکچه‌های خرطوم‌دار – Persian" lang="fa" hreflang="fa" data-title="سوسکچه‌های خرطوم‌دار" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Curculionidae" title="Curculionidae – French" lang="fr" hreflang="fr" data-title="Curculionidae" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Gurgullo_(insecto)" title="Gurgullo (insecto) – Galician" lang="gl" hreflang="gl" data-title="Gurgullo (insecto)" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%B0%94%EA%B5%AC%EB%AF%B8%EA%B3%BC" title="바구미과 – Korean" lang="ko" hreflang="ko" data-title="바구미과" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D4%B5%D6%80%D5%AF%D5%A1%D6%80%D5%A1%D5%AF%D5%B6%D5%B3%D5%AB%D5%A9%D5%B6%D5%A5%D6%80" title="Երկարակնճիթներ – Armenian" lang="hy" hreflang="hy" data-title="Երկարակնճիթներ" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Pipe" title="Pipe – Croatian" lang="hr" hreflang="hr" data-title="Pipe" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Kurkuliono" title="Kurkuliono – Ido" lang="io" hreflang="io" data-title="Kurkuliono" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-os mw-list-item"><a href="https://os.wikipedia.org/wiki/%D0%94%D0%B0%D1%80%D0%B3%D1%8A%D0%B1%D1%8B%D1%80%D1%8B%D0%BD%D1%87%D1%8A%D1%8B%D1%82%C3%A6" title="Даргъбырынчъытæ – Ossetic" lang="os" hreflang="os" data-title="Даргъбырынчъытæ" data-language-autonym="Ирон" data-language-local-name="Ossetic" class="interlanguage-link-target"><span>Ирон</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Ranabj%C3%B6llur" title="Ranabjöllur – Icelandic" lang="is" hreflang="is" data-title="Ranabjöllur" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Italian" lang="it" hreflang="it" data-title="Curculionidae" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%97%D7%93%D7%A7%D7%95%D7%A0%D7%99%D7%AA%D7%99%D7%99%D7%9D" title="חדקוניתיים – Hebrew" lang="he" hreflang="he" data-title="חדקוניתיים" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%95%E0%B2%B0%E0%B3%8D%E0%B2%95%E0%B3%8D%E0%B2%AF%E0%B3%82%E0%B2%B2%E0%B2%BF%E0%B2%AF%E0%B2%BE%E0%B2%A8%E0%B2%BF%E0%B2%A1%E0%B3%80" title="ಕರ್ಕ್ಯೂಲಿಯಾನಿಡೀ – Kannada" lang="kn" hreflang="kn" data-title="ಕರ್ಕ್ಯೂಲಿಯಾನಿಡೀ" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%AA%E1%83%AE%E1%83%95%E1%83%98%E1%83%A0%E1%83%92%E1%83%A0%E1%83%AB%E1%83%94%E1%83%9A%E1%83%90%E1%83%A1%E1%83%94%E1%83%91%E1%83%A0%E1%83%9C%E1%83%98" title="ცხვირგრძელასებრნი – Georgian" lang="ka" hreflang="ka" data-title="ცხვირგრძელასებრნი" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D2%B0%D0%B7%D1%8B%D0%BD%D1%82%D2%B1%D0%BC%D1%81%D1%8B%D2%9B%D1%82%D1%8B%D0%BB%D0%B0%D1%80" title="Ұзынтұмсықтылар – Kazakh" lang="kk" hreflang="kk" data-title="Ұзынтұмсықтылар" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%A8%D0%B8%D1%88%D1%82%D1%83%D0%BC%D1%88%D1%83%D0%BA%D1%82%D0%B0%D1%80" title="Шиштумшуктар – Kyrgyz" lang="ky" hreflang="ky" data-title="Шиштумшуктар" data-language-autonym="Кыргызча" data-language-local-name="Kyrgyz" class="interlanguage-link-target"><span>Кыргызча</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Latin" lang="la" hreflang="la" data-title="Curculionidae" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Smecernieku_dzimta" title="Smecernieku dzimta – Latvian" lang="lv" hreflang="lv" data-title="Smecernieku dzimta" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Straubliukai" title="Straubliukai – Lithuanian" lang="lt" hreflang="lt" data-title="Straubliukai" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Orm%C3%A1nyosbog%C3%A1r-f%C3%A9l%C3%A9k" title="Ormányosbogár-félék – Hungarian" lang="hu" hreflang="hu" data-title="Ormányosbogár-félék" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%A7%D1%83%D1%80%D1%83%D0%BB%D0%BA%D0%B0%D1%80%D0%B8" title="Чурулкари – Macedonian" lang="mk" hreflang="mk" data-title="Чурулкари" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Malay" lang="ms" hreflang="ms" data-title="Curculionidae" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Snuitkevers_(familie)" title="Snuitkevers (familie) – Dutch" lang="nl" hreflang="nl" data-title="Snuitkevers (familie)" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja badge-Q70893996 mw-list-item" title=""><a href="https://ja.wikipedia.org/wiki/%E3%82%BE%E3%82%A6%E3%83%A0%E3%82%B7%E7%A7%91" title="ゾウムシ科 – Japanese" lang="ja" hreflang="ja" data-title="ゾウムシ科" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Snutebiller" title="Snutebiller – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Snutebiller" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Filchalar" title="Filchalar – Uzbek" lang="uz" hreflang="uz" data-title="Filchalar" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Snutenk%C3%A4vers" title="Snutenkävers – Low German" lang="nds" hreflang="nds" data-title="Snutenkävers" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Ryjkowcowate" title="Ryjkowcowate – Polish" lang="pl" hreflang="pl" data-title="Ryjkowcowate" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Curculion%C3%ADdeos" title="Curculionídeos – Portuguese" lang="pt" hreflang="pt" data-title="Curculionídeos" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Curculionide" title="Curculionide – Romanian" lang="ro" hreflang="ro" data-title="Curculionide" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-qu mw-list-item"><a href="https://qu.wikipedia.org/wiki/Sinqa_suntu" title="Sinqa suntu – Quechua" lang="qu" hreflang="qu" data-title="Sinqa suntu" data-language-autonym="Runa Simi" data-language-local-name="Quechua" class="interlanguage-link-target"><span>Runa Simi</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%94%D0%BE%D0%BB%D0%B3%D0%BE%D0%BD%D0%BE%D1%81%D0%B8%D0%BA%D0%B8" title="Долгоносики – Russian" lang="ru" hreflang="ru" data-title="Долгоносики" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Pravi_ril%C4%8Dkarji" title="Pravi rilčkarji – Slovenian" lang="sl" hreflang="sl" data-title="Pravi rilčkarji" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Pipe" title="Pipe – Serbian" lang="sr" hreflang="sr" data-title="Pipe" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/K%C3%A4rs%C3%A4kk%C3%A4%C3%A4t" title="Kärsäkkäät – Finnish" lang="fi" hreflang="fi" data-title="Kärsäkkäät" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/%C3%84kta_vivlar" title="Äkta vivlar – Swedish" lang="sv" hreflang="sv" data-title="Äkta vivlar" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Curculionidae" title="Curculionidae – Turkish" lang="tr" hreflang="tr" data-title="Curculionidae" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%94%D0%BE%D0%B2%D0%B3%D0%BE%D0%BD%D0%BE%D1%81%D0%B8%D0%BA%D0%BE%D0%B2%D1%96" title="Довгоносикові – Ukrainian" lang="uk" hreflang="uk" data-title="Довгоносикові" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/H%E1%BB%8D_B%E1%BB%8D_v%C3%B2i_voi" title="Họ Bọ vòi voi – Vietnamese" lang="vi" 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searchaux" style="display:none">Family of beetles</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">True weevils and bark beetles </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Weevil_(Cholus_cinctus)_Anton_Valley.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Weevil_%28Cholus_cinctus%29_Anton_Valley.jpg/220px-Weevil_%28Cholus_cinctus%29_Anton_Valley.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Weevil_%28Cholus_cinctus%29_Anton_Valley.jpg/330px-Weevil_%28Cholus_cinctus%29_Anton_Valley.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Weevil_%28Cholus_cinctus%29_Anton_Valley.jpg/440px-Weevil_%28Cholus_cinctus%29_Anton_Valley.jpg 2x" data-file-width="3701" data-file-height="2467" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i><a href="/w/index.php?title=Cholus_cinctus&action=edit&redlink=1" class="new" title="Cholus cinctus (page does not exist)">Cholus cinctus</a></i>, Panama </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Curculionidae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Insect" title="Insect">Insecta</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Beetle" title="Beetle">Coleoptera</a> </td></tr> <tr> <td>Infraorder: </td> <td><a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Phytophaga" title="Phytophaga">Phytophaga</a> </td></tr> <tr> <td>Superfamily: </td> <td><a href="/wiki/Weevil" title="Weevil">Curculionoidea</a> </td></tr> <tr> <td>Family: </td> <td><a class="mw-selflink selflink">Curculionidae</a><br /><small><a href="/wiki/Pierre_Andr%C3%A9_Latreille" title="Pierre André Latreille">Latreille</a>, 1802</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Biodiversity" title="Biodiversity">Diversity</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><a href="#Phylogeny_and_systematics">Some 20 subfamilies (see text)</a> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Flickr_-_Lukjonis_-_Bug-Cionus_hortulanus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Flickr_-_Lukjonis_-_Bug-Cionus_hortulanus.jpg/220px-Flickr_-_Lukjonis_-_Bug-Cionus_hortulanus.jpg" decoding="async" width="220" height="169" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Flickr_-_Lukjonis_-_Bug-Cionus_hortulanus.jpg/330px-Flickr_-_Lukjonis_-_Bug-Cionus_hortulanus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Flickr_-_Lukjonis_-_Bug-Cionus_hortulanus.jpg/440px-Flickr_-_Lukjonis_-_Bug-Cionus_hortulanus.jpg 2x" data-file-width="1024" data-file-height="787" /></a><figcaption>A true weevil</figcaption></figure> <p>The <b>Curculionidae</b> are a family of <a href="/wiki/Weevil" title="Weevil">weevils</a>, commonly called <b>snout beetles</b> or <b>true weevils</b>. They are one of the largest animal families with 6,800 genera and 83,000<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> species described worldwide. They are the sister group to the family <a href="/wiki/Brentidae" title="Brentidae">Brentidae</a>.<sup id="cite_ref-nlg_2-0" class="reference"><a href="#cite_note-nlg-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>They include the <a href="/wiki/Bark_beetle" title="Bark beetle">bark beetles</a> as the <a href="/wiki/Subfamily" title="Subfamily">subfamily</a> <a href="/wiki/Scolytinae" class="mw-redirect" title="Scolytinae">Scolytinae</a>, which are modified <sup class="noprint Inline-Template" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Reliable_sources" title="Wikipedia:Reliable sources"><span title="The material near this tag may rely on an unreliable source. (December 2013)">unreliable source?</span></a></i>]</sup> in shape in accordance with their wood-boring lifestyle. They do not much resemble other weevils, so they were traditionally considered a distinct family, Scolytidae. The family also includes the <a href="/wiki/Ambrosia_beetle" title="Ambrosia beetle">ambrosia beetles</a>, of which the present-day <a href="/wiki/Subfamily" title="Subfamily">subfamily</a> <a href="/wiki/Platypodinae" title="Platypodinae">Platypodinae</a> was formerly considered the distinct family Platypodidae. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Curculionidae&action=edit&section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Adult Curculionidae can be recognised by the well-developed, downwards-curved snout (<a href="/wiki/Rostrum_(anatomy)" title="Rostrum (anatomy)">rostrum</a>) possessed by many species, though the rostrum is sometimes short (e.g. Entiminae).<sup id="cite_ref-:0_3-0" class="reference"><a href="#cite_note-:0-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_4-0" class="reference"><a href="#cite_note-:1-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> They have elbowed <a href="/wiki/Antenna_(biology)" title="Antenna (biology)">antennae</a> that end in clubs, and the first antennal segment often fits into a groove in the side of the rostrum.<sup id="cite_ref-:0_3-1" class="reference"><a href="#cite_note-:0-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_4-1" class="reference"><a href="#cite_note-:1-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> The body tends to be robust, convex, heavily sclerotised and covered in scales or bristles.<sup id="cite_ref-:0_3-2" class="reference"><a href="#cite_note-:0-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Curculionidae range in size from 1–35 mm long, usually being 5–15 mm long.<sup id="cite_ref-:1_4-2" class="reference"><a href="#cite_note-:1-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Most Curculionidae are <a href="/wiki/Sexual_dimorphism" title="Sexual dimorphism">sexually dimorphic</a> with females (compared to males) having antennae positioned more basally and a longer, thinner rostrum.<sup id="cite_ref-:1_4-3" class="reference"><a href="#cite_note-:1-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Larval Curculionidae are C-shaped and lightly sclerotised, with minute antennae, robust mandibles<sup id="cite_ref-:0_3-3" class="reference"><a href="#cite_note-:0-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> and no legs.<sup id="cite_ref-:2_5-0" class="reference"><a href="#cite_note-:2-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p> Most weevils feed on plants as larvae and adults, and they include important pests of cultivated plants that chew holes in fruits, nuts and other parts.<sup id="cite_ref-:1_4-4" class="reference"><a href="#cite_note-:1-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> The long rostrum possessed by most adult weevils is used by females to help lay eggs (oviposit) inside plant tissue.<sup id="cite_ref-:2_5-1" class="reference"><a href="#cite_note-:2-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> Some feed on rotten wood or bark (e.g. Cossoninae and Cryptorhynchinae), and some are <a href="/wiki/Woodboring_beetle" title="Woodboring beetle">wood-borers</a> that feed on <a href="/wiki/Ambrosia_fungi" title="Ambrosia fungi">ambrosia fungi</a> (Platypodinae and some Scolytinae).<sup id="cite_ref-:0_3-4" class="reference"><a href="#cite_note-:0-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></p><figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Curculionidae_._Curculio_elephas_(33105941022).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Curculionidae_._Curculio_elephas_%2833105941022%29.jpg/220px-Curculionidae_._Curculio_elephas_%2833105941022%29.jpg" decoding="async" width="220" height="167" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Curculionidae_._Curculio_elephas_%2833105941022%29.jpg/330px-Curculionidae_._Curculio_elephas_%2833105941022%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Curculionidae_._Curculio_elephas_%2833105941022%29.jpg/440px-Curculionidae_._Curculio_elephas_%2833105941022%29.jpg 2x" data-file-width="1554" data-file-height="1181" /></a><figcaption><i><a href="/wiki/Curculio_elephas" title="Curculio elephas">Curculio elephas</a></i></figcaption></figure> <p>Although pesticide resistance hasn't historically been an issue with these insects, recently a mutation was discovered in association with the voltage-gated sodium channel in the species <i><a href="/wiki/Maize_weevil" title="Maize weevil">Sitophilus zeamais</a></i>, indicating there is a lot to learn about how these insects adapt to changing environments.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Behavior">Behavior</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Curculionidae&action=edit&section=2" title="Edit section: Behavior"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>When disturbed, adult curculionids often <a href="/wiki/Apparent_death" title="Apparent death">play dead</a> by lying motionless on their backs.<sup id="cite_ref-:1_4-5" class="reference"><a href="#cite_note-:1-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Many species of weevils are common household and garden pests, but don't harm people, pets, or buildings. Their presence is more of a temporary nuisance, usually to plants and their fruits in their larval stage. In tropical areas they have larger effects, specifically several species in the genera <i><a href="/wiki/Conotrachelus" title="Conotrachelus">Conotrachelus</a></i> and <i><a href="/wiki/Copturus" title="Copturus">Copturus</a></i>.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Phylogeny_and_systematics">Phylogeny and systematics</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Curculionidae&action=edit&section=3" title="Edit section: Phylogeny and systematics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The <a href="/wiki/Phylogeny" class="mw-redirect" title="Phylogeny">phylogeny</a> of the group is complex; with so many species, a spirited debate exists about the relationships between subfamilies and genera. A 1997 analysis attempted to construct a phylogeny based mainly on larval characteristics.<sup id="cite_ref-:2_5-2" class="reference"><a href="#cite_note-:2-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>Recent work on the phylogenetic relationships in weevils mentions the two subfamily groups <b>Adelognatha</b> (<i>short-nosed weevils</i>, subfamily <a href="/wiki/Entiminae" title="Entiminae">Entiminae</a>) and <b>Phanerognatha</b> (<i>long-nosed weevils</i>, subfamilies of Curculionidae other than Entiminae) for the species of Curculionidae.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>Almost two dozen <a href="/wiki/Subfamilies" class="mw-redirect" title="Subfamilies">subfamilies</a> are recognized by some authors even when merging those that are certainly invalid. Others, however, recognize a lesser number – the only subfamilies that are almost universally considered valid are the <a href="/wiki/Baridinae" title="Baridinae">Baridinae</a>, <a href="/wiki/Cossoninae" title="Cossoninae">Cossoninae</a>, <a href="/wiki/Curculioninae" title="Curculioninae">Curculioninae</a>, <a href="/wiki/Cyclominae" title="Cyclominae">Cyclominae</a>, <a href="/wiki/Entiminae" title="Entiminae">Entiminae</a>, <a href="/wiki/Molytinae" title="Molytinae">Molytinae</a>, <a href="/wiki/Platypodinae" title="Platypodinae">Platypodinae</a>, and <a href="/wiki/Scolytinae" class="mw-redirect" title="Scolytinae">Scolytinae</a>. The various proposed <a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">taxonomic</a> schemes typically recognize as many additional subfamilies again, but little agreement is seen between authorities about which. In particular, the delimitation of the <a href="/wiki/Molytinae" title="Molytinae">Molytinae</a> has proven difficult.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (December 2022)">citation needed</span></a></i>]</sup> </p><p>The timeline for current and extant weevil speciation and diversification is consistent with the radiation of <a href="/wiki/Gymnosperm" title="Gymnosperm">gymnosperms</a> during the Mesozoic period.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Curculio_larva.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/59/Curculio_larva.jpg/160px-Curculio_larva.jpg" decoding="async" width="160" height="123" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/59/Curculio_larva.jpg/240px-Curculio_larva.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/59/Curculio_larva.jpg/320px-Curculio_larva.jpg 2x" data-file-width="2016" data-file-height="1544" /></a><figcaption><i>Curculio</i>, scarabeiform larva</figcaption></figure> <p>The subfamilies considered valid by at least some authors today: </p> <ul><li><a href="/wiki/Bagoinae" title="Bagoinae">Bagoinae</a> (sometimes in Molytinae)</li> <li><a href="/wiki/Baridinae" title="Baridinae">Baridinae</a></li> <li><a href="/wiki/Brachycerinae" title="Brachycerinae">Brachycerinae</a> (disputed; sometimes placed at family level)</li> <li><a href="/wiki/Ceutorhynchinae" title="Ceutorhynchinae">Ceutorhynchinae</a> (sometimes in Baridinae)</li> <li><a href="/wiki/Conoderinae" title="Conoderinae">Conoderinae</a> (sometimes in Baridinae)</li> <li><a href="/wiki/Cossoninae" title="Cossoninae">Cossoninae</a></li> <li><a href="/wiki/Cryptorhynchinae" title="Cryptorhynchinae">Cryptorhynchinae</a> (sometimes in Curculioninae)<figure class="mw-default-size" typeof="mw:File/Thumb"><span><video id="mwe_player_0" poster="//upload.wikimedia.org/wikipedia/commons/thumb/f/f2/Cionus_tuberculosus_-_2012-06-29.ogv/220px--Cionus_tuberculosus_-_2012-06-29.ogv.jpg" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="220" height="124" data-durationhint="55" data-mwtitle="Cionus_tuberculosus_-_2012-06-29.ogv" data-mwprovider="wikimediacommons" resource="/wiki/File:Cionus_tuberculosus_-_2012-06-29.ogv"><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/f/f2/Cionus_tuberculosus_-_2012-06-29.ogv/Cionus_tuberculosus_-_2012-06-29.ogv.480p.vp9.webm" type="video/webm; codecs="vp9, opus"" data-transcodekey="480p.vp9.webm" data-width="854" data-height="480" /><source src="//upload.wikimedia.org/wikipedia/commons/f/f2/Cionus_tuberculosus_-_2012-06-29.ogv" type="video/ogg; codecs="theora"" data-width="1024" data-height="576" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/f/f2/Cionus_tuberculosus_-_2012-06-29.ogv/Cionus_tuberculosus_-_2012-06-29.ogv.144p.mjpeg.mov" type="video/quicktime" data-transcodekey="144p.mjpeg.mov" data-width="256" data-height="144" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/f/f2/Cionus_tuberculosus_-_2012-06-29.ogv/Cionus_tuberculosus_-_2012-06-29.ogv.240p.vp9.webm" type="video/webm; codecs="vp9, opus"" data-transcodekey="240p.vp9.webm" data-width="426" data-height="240" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/f/f2/Cionus_tuberculosus_-_2012-06-29.ogv/Cionus_tuberculosus_-_2012-06-29.ogv.360p.vp9.webm" type="video/webm; codecs="vp9, opus"" data-transcodekey="360p.vp9.webm" data-width="640" data-height="360" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/f/f2/Cionus_tuberculosus_-_2012-06-29.ogv/Cionus_tuberculosus_-_2012-06-29.ogv.360p.webm" type="video/webm; codecs="vp8, vorbis"" data-transcodekey="360p.webm" data-width="640" data-height="360" /></video></span><figcaption><i>Cionus tuberculosus</i> (<a href="/wiki/Curculioninae" title="Curculioninae">Curculioninae</a>)</figcaption></figure> <ul><li><i><a href="/wiki/Acalles" title="Acalles">Acalles</a></i></li></ul></li> <li><a href="/wiki/Curculioninae" title="Curculioninae">Curculioninae</a> – flower weevils, acorn and nut weevils</li></ul> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Weevil3.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/11/Weevil3.jpg/220px-Weevil3.jpg" decoding="async" width="220" height="181" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/11/Weevil3.jpg/330px-Weevil3.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/11/Weevil3.jpg/440px-Weevil3.jpg 2x" data-file-width="1052" data-file-height="867" /></a><figcaption><i>Curculio nucum</i>, a brown nut-weevil</figcaption></figure> <ul><li><a href="/wiki/Cyclominae" title="Cyclominae">Cyclominae</a></li> <li><a href="/wiki/Dryophthorinae" title="Dryophthorinae">Dryophthorinae</a> (sometimes placed at family level)</li> <li><a href="/wiki/Entiminae" title="Entiminae">Entiminae</a> – broad-nosed weevils</li> <li><a href="/wiki/Hyperinae" title="Hyperinae">Hyperinae</a> (sometimes in Molytinae)</li> <li><a href="/wiki/Lixinae" title="Lixinae">Lixinae</a> (sometimes in Molytinae)</li> <li><a href="/wiki/Mesoptiliinae" title="Mesoptiliinae">Mesoptiliinae</a> (sometimes in Molytinae)</li> <li><a href="/wiki/Molytinae" title="Molytinae">Molytinae</a></li> <li><a href="/w/index.php?title=Orobitidinae&action=edit&redlink=1" class="new" title="Orobitidinae (page does not exist)">Orobitidinae</a> (sometimes in Baridinae)</li> <li><a href="/wiki/Platypodinae" title="Platypodinae">Platypodinae</a> – typical <a href="/wiki/Ambrosia_beetle" title="Ambrosia beetle">ambrosia beetles</a>, "higher" Curculionidaes<sup id="cite_ref-nlg_2-1" class="reference"><a href="#cite_note-nlg-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Raymondionyminae" title="Raymondionyminae">Raymondionyminae</a> (sometimes in Brachycerinae)</li> <li><a href="/wiki/Scolytinae" class="mw-redirect" title="Scolytinae">Scolytinae</a> – bark beetles</li> <li><a href="/w/index.php?title=Xiphaspidinae&action=edit&redlink=1" class="new" title="Xiphaspidinae (page does not exist)">Xiphaspidinae</a> (sometimes in Baridinae)</li></ul> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Curculionidae&action=edit&section=4" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239009302">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{clear:left;float:left;margin:0.5em 1em 0.5em 0}.mw-parser-output .portalright{clear:right;float:right;margin:0.5em 0 0.5em 1em}}</style><ul role="navigation" aria-label="Portals" class="noprint portalbox portalborder portalright"> <li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Beetle_icon.svg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/25px-Beetle_icon.svg.png" decoding="async" width="25" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/38px-Beetle_icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/50px-Beetle_icon.svg.png 2x" data-file-width="143" data-file-height="159" /></a></span></span><span class="portalbox-link"><a href="/wiki/Portal:Insects" title="Portal:Insects">Insects portal</a></span></li><li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Cercophonius_squama.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/60/Cercophonius_squama.jpg/32px-Cercophonius_squama.jpg" decoding="async" width="32" height="26" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/60/Cercophonius_squama.jpg/48px-Cercophonius_squama.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/60/Cercophonius_squama.jpg/64px-Cercophonius_squama.jpg 2x" data-file-width="1974" data-file-height="1606" /></a></span></span><span class="portalbox-link"><a href="/wiki/Portal:Arthropods" title="Portal:Arthropods">Arthropods portal</a></span></li></ul> <ul><li><a href="/wiki/Black_vine_weevil" class="mw-redirect" title="Black vine weevil">Black vine weevil</a></li> <li><a href="/wiki/Boll_weevil" title="Boll weevil">Boll weevil</a></li> <li><a href="/wiki/Curculio_caryae" title="Curculio caryae">Pecan weevil</a></li> <li><a href="/wiki/Wheat_weevil" title="Wheat weevil">Wheat weevil</a></li> <li><i><a href="/wiki/Hylobius" title="Hylobius">Hylobius</a></i></li> <li><i><a href="/wiki/Orthorhinus_cylindrirostris" title="Orthorhinus cylindrirostris">Orthorhinus cylindrirostris</a></i></li> <li><i><a href="/wiki/Premnotrypes" title="Premnotrypes">Premnotrypes</a></i></li> <li><i><a href="/wiki/Scolytoplatypus" title="Scolytoplatypus">Scolytoplatypus</a></i></li> <li><a href="/wiki/Pests_and_diseases_of_roses" class="mw-redirect" title="Pests and diseases of roses">Pests and diseases of roses</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Curculionidae&action=edit&section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output 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.id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite class="citation web cs1"><a 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Archived from <a rel="nofollow" class="external text" href="http://www.cricyt.edu.ar/entomologia/Articulos/Cladistics13%281997%29.pdf">the original</a> <span class="cs1-format">(PDF)</span> on 2012-02-18<span class="reference-accessdate">. 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(2011). <a rel="nofollow" class="external text" href="https://onlinelibrary.wiley.com/doi/abs/10.1111/j.1365-2583.2011.01079.x">"Pyrethroid resistance in Sitophilus zeamais is associated with a mutation (T929I) in the voltage-gated sodium channel"</a>. <i>Insect Molecular Biology</i>. <b>20</b> (4): 437–445. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.1365-2583.2011.01079.x">10.1111/j.1365-2583.2011.01079.x</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1365-2583">1365-2583</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21496128">21496128</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:205307830">205307830</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Insect+Molecular+Biology&rft.atitle=Pyrethroid+resistance+in+Sitophilus+zeamais+is+associated+with+a+mutation+%28T929I%29+in+the+voltage-gated+sodium+channel&rft.volume=20&rft.issue=4&rft.pages=437-445&rft.date=2011&rft.issn=1365-2583&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A205307830%23id-name%3DS2CID&rft_id=info%3Apmid%2F21496128&rft_id=info%3Adoi%2F10.1111%2Fj.1365-2583.2011.01079.x&rft.aulast=Ara%C3%BAjo&rft.aufirst=R%C3%BAbia+A.&rft.au=Williamson%2C+Martin+S.&rft.au=Bass%2C+Christopher&rft.au=Field%2C+Linda+M.&rft.au=Duce%2C+Ian+R.&rft_id=https%3A%2F%2Fonlinelibrary.wiley.com%2Fdoi%2Fabs%2F10.1111%2Fj.1365-2583.2011.01079.x&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACurculionidae" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFuentesVildozolaUrias-Lopez2017" class="citation book cs1">Fuentes, Luis Martin Hernandez; Vildozola, Alvaro Castaneda; Urias-Lopez, Mario Alfonso (2017-04-12). Shields, Vonnie D.C. (ed.). <a rel="nofollow" class="external text" href="http://www.intechopen.com/books/insect-physiology-and-ecology/weevil-borers-in-tropical-fruit-crops-importance-biology-and-management"><i>Weevil Borers in Tropical Fruit Crops: Importance, Biology and Management</i></a>. InTech. p. 43. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.5772%2F66635">10.5772/66635</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-953-51-3033-8" title="Special:BookSources/978-953-51-3033-8"><bdi>978-953-51-3033-8</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Weevil+Borers+in+Tropical+Fruit+Crops%3A+Importance%2C+Biology+and+Management&rft.pages=43&rft.pub=InTech&rft.date=2017-04-12&rft_id=info%3Adoi%2F10.5772%2F66635&rft.isbn=978-953-51-3033-8&rft.aulast=Fuentes&rft.aufirst=Luis+Martin+Hernandez&rft.au=Vildozola%2C+Alvaro+Castaneda&rft.au=Urias-Lopez%2C+Mario+Alfonso&rft_id=http%3A%2F%2Fwww.intechopen.com%2Fbooks%2Finsect-physiology-and-ecology%2Fweevil-borers-in-tropical-fruit-crops-importance-biology-and-management&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACurculionidae" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLieutier2007" class="citation book cs1">Lieutier, François, ed. 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Dordrecht: Kluwer. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1-4020-2240-1" title="Special:BookSources/978-1-4020-2240-1"><bdi>978-1-4020-2240-1</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Bark+and+wood+boring+insects+in+living+trees+in+Europe%3A+a+synthesis&rft.place=Dordrecht&rft.edition=Repr&rft.pub=Kluwer&rft.date=2007&rft.isbn=978-1-4020-2240-1&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACurculionidae" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFShinClarkeLemmonMoriarty_Lemmon2018" class="citation journal cs1">Shin, Seunggwan; Clarke, Dave J; Lemmon, Alan R; Moriarty Lemmon, Emily; Aitken, Alexander L; Haddad, Stephanie; Farrell, Brian D; Marvaldi, Adriana E; Oberprieler, Rolf G; McKenna, Duane D (2018-04-01). <a rel="nofollow" class="external text" href="https://academic.oup.com/mbe/article/35/4/823/4765916">"Phylogenomic Data Yield New and Robust Insights into the Phylogeny and Evolution of Weevils"</a>. <i>Molecular Biology and Evolution</i>. <b>35</b> (4): 823–836. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fmolbev%2Fmsx324">10.1093/molbev/msx324</a></span>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/11336%2F57287">11336/57287</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0737-4038">0737-4038</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29294021">29294021</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Molecular+Biology+and+Evolution&rft.atitle=Phylogenomic+Data+Yield+New+and+Robust+Insights+into+the+Phylogeny+and+Evolution+of+Weevils&rft.volume=35&rft.issue=4&rft.pages=823-836&rft.date=2018-04-01&rft_id=info%3Ahdl%2F11336%2F57287&rft.issn=0737-4038&rft_id=info%3Apmid%2F29294021&rft_id=info%3Adoi%2F10.1093%2Fmolbev%2Fmsx324&rft.aulast=Shin&rft.aufirst=Seunggwan&rft.au=Clarke%2C+Dave+J&rft.au=Lemmon%2C+Alan+R&rft.au=Moriarty+Lemmon%2C+Emily&rft.au=Aitken%2C+Alexander+L&rft.au=Haddad%2C+Stephanie&rft.au=Farrell%2C+Brian+D&rft.au=Marvaldi%2C+Adriana+E&rft.au=Oberprieler%2C+Rolf+G&rft.au=McKenna%2C+Duane+D&rft_id=https%3A%2F%2Facademic.oup.com%2Fmbe%2Farticle%2F35%2F4%2F823%2F4765916&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACurculionidae" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Curculionidae&action=edit&section=6" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Commons-logo.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span> Media related to <a href="https://commons.wikimedia.org/wiki/Category:Curculionidae" class="extiw" title="commons:Category:Curculionidae">Curculionidae</a> at Wikimedia Commons</li> <li><span 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href="/wiki/Template:Coleoptera" title="Template:Coleoptera"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Coleoptera" title="Template talk:Coleoptera"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Coleoptera" title="Special:EditPage/Template:Coleoptera"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Extant_Coleoptera_families" style="font-size:114%;margin:0 4em">Extant <a href="/wiki/Beetle" title="Beetle">Coleoptera</a> families</div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a></li> <li>Class: <a href="/wiki/Insect" title="Insect">Insecta</a></li> <li>Subclass: <a href="/wiki/Pterygota" title="Pterygota">Pterygota</a></li> <li>Infraclass: <a href="/wiki/Neoptera" title="Neoptera">Neoptera</a></li> <li>Superorder: <a href="/wiki/Holometabola" title="Holometabola">Holometabola</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Archostemata" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Archostemata" title="Archostemata">Archostemata</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Crowsoniella" title="Crowsoniella">Crowsoniellidae <small>(<i>Crowsoniella relicta</i>)</small></a></li> <li><a href="/wiki/Cupedidae" title="Cupedidae">Cupedidae <small>(reticulated beetles)</small></a></li> <li><a href="/wiki/Jurodidae" title="Jurodidae">Jurodidae <small>(<i>Sikhotealinia zhiltzovae</i>)</small></a></li> <li><a href="/wiki/Micromalthidae" title="Micromalthidae">Micromalthidae</a></li> <li><a href="/wiki/Ommatidae" title="Ommatidae">Ommatidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Adephaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Adephaga" title="Adephaga">Adephaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Extant_families" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Extant_taxon" class="mw-redirect" title="Extant taxon">Extant</a> families</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amphizoa" title="Amphizoa">Amphizoidae <small>(trout-stream beetles)</small></a></li> <li><a href="/wiki/Aspidytidae" title="Aspidytidae">Aspidytidae</a></li> <li><a href="/wiki/Ground_beetle" title="Ground beetle">Carabidae <small>(ground beetles)</small></a></li> <li><a href="/wiki/Tiger_beetle" title="Tiger beetle">Cicindelidae <small>(tiger beetles)</small></a></li> <li><a href="/wiki/Dytiscidae" title="Dytiscidae">Dytiscidae <small>(predaceous diving beetles)</small></a></li> <li><a href="/wiki/Whirligig_beetle" title="Whirligig beetle">Gyrinidae <small>(whirligig beetles)</small></a></li> <li><a href="/wiki/Haliplidae" title="Haliplidae">Haliplidae <small>(crawling water beetles)</small></a></li> <li><a href="/wiki/Hygrobia" title="Hygrobia">Hygrobiidae</a></li> <li><a href="/wiki/Meru_phyllisae" title="Meru phyllisae">Meruidae <small>(<i>Meru phyllisae</i>)</small></a></li> <li><a href="/wiki/Noteridae" title="Noteridae">Noteridae <small>(burrowing water beetles)</small></a></li> <li><a href="/wiki/Trachypachidae" title="Trachypachidae">Trachypachidae <small>(false ground beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Myxophaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Myxophaga" title="Myxophaga">Myxophaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hydroscaphidae" title="Hydroscaphidae">Hydroscaphidae <small>(skiff beetles)</small></a></li> <li><a href="/wiki/Lepicerus" title="Lepicerus">Lepiceridae</a></li> <li><a href="/wiki/Sphaerius" title="Sphaerius">Sphaeriusidae</a></li> <li><a href="/wiki/Torridincolidae" title="Torridincolidae">Torridincolidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Polyphaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Polyphaga" title="Polyphaga">Polyphaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bostrichiformia" title="Bostrichiformia">Bostrichiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Bostrichoidea" title="Bostrichoidea">Bostrichoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bostrichidae" title="Bostrichidae">Bostrichidae <small>(auger beetles)</small></a></li> <li><a href="/wiki/Dermestidae" title="Dermestidae">Dermestidae <small>(skin beetles)</small></a></li> <li><a href="/wiki/Endecatomus" title="Endecatomus">Endecatomidae</a></li> <li><a href="/wiki/Jacobsoniidae" title="Jacobsoniidae">Jacobsoniidae <small>(Jacobson's beetles)</small></a></li> <li><a href="/wiki/Nosodendridae" title="Nosodendridae">Nosodendridae <small>(wounded-tree beetles)</small></a></li> <li><a href="/wiki/Ptinidae" title="Ptinidae">Ptiniidae <small>(furniture beetles, death watch beetles, spider beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Derodontidae" title="Derodontidae">Derodontoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Derodontidae" title="Derodontidae">Derodontidae <small>(tooth-necked fungus beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Chrysomeloidea" title="Chrysomeloidea">Chrysomeloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Longhorn_beetle" title="Longhorn beetle">Cerambycidae <small>(longhorn beetles)</small></a></li> <li><a href="/wiki/Leaf_beetle" title="Leaf beetle">Chrysomelidae <small>(leaf beetles)</small></a></li> <li><a href="/wiki/Disteniidae" title="Disteniidae">Disteniidae</a></li> <li><a href="/wiki/Megalopodidae" title="Megalopodidae">Megalopodidae</a></li> <li><a href="/wiki/Orsodacnidae" title="Orsodacnidae">Orsodacnidae</a></li> <li><a href="/wiki/Oxypeltidae" title="Oxypeltidae">Oxypeltidae</a></li> <li><a href="/wiki/Vesperidae" title="Vesperidae">Vesperidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cleroidea" title="Cleroidea">Cleroidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acanthocnemus" title="Acanthocnemus">Acanthocnemidae <small>(<i>Acanthocnemus nigricans</i>)</small></a></li> <li><a href="/wiki/Biphyllidae" title="Biphyllidae">Biphyllidae <small>(false skin beetles)</small></a></li> <li><a href="/wiki/Byturidae" title="Byturidae">Byturidae <small>(fruitworm beetles)</small></a></li> <li><a href="/wiki/Chaetosomatidae" title="Chaetosomatidae">Chaetosomatidae</a></li> <li><a href="/wiki/Cleridae" title="Cleridae">Cleridae <small>(checkered beetles)</small></a></li> <li><a href="/wiki/Lophocateridae" title="Lophocateridae">Lophocateridae</a></li> <li><a href="/wiki/Mauroniscidae" title="Mauroniscidae">Mauroniscidae</a></li> <li><a href="/wiki/Melyridae" title="Melyridae">Melyridae <small>(soft-wing flower beetles)</small></a></li> <li><a href="/wiki/Metaxina" title="Metaxina">Metaxinidae <small>(<i>Metaxina ornata</i>)</small></a></li> <li><a href="/wiki/Peltis" title="Peltis">Peltidae</a></li> <li><a href="/wiki/Phloiophilus" title="Phloiophilus">Phloiophilidae <small>(<i>Phloiophilus edwardsi</i>)</small></a></li> <li><a href="/wiki/Phycosecidae" title="Phycosecidae">Phycosecidae</a></li> <li><a href="/wiki/Prionoceridae" title="Prionoceridae">Prionoceridae</a></li> <li><a href="/wiki/Protopeltis" title="Protopeltis">Protopeltidae</a></li> <li><a href="/wiki/Rentoniidae" title="Rentoniidae">Rentoniidae</a></li> <li><a href="/wiki/Rhadalidae" title="Rhadalidae">Rhadalidae</a></li> <li><a href="/wiki/Thanerocleridae" title="Thanerocleridae">Thanerocleridae</a></li> <li><a href="/wiki/Thymalidae" title="Thymalidae">Thymalidae</a></li> <li><a href="/wiki/Trogossitidae" title="Trogossitidae">Trogossitidae <small>(bark-gnawing beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Coccinelloidea" title="Coccinelloidea">Coccinelloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Akalyptoischion" title="Akalyptoischion">Akalyptoischiidae</a></li> <li><a href="/wiki/Alexiidae" title="Alexiidae">Alexiidae</a></li> <li><a href="/wiki/Anamorphidae" title="Anamorphidae">Anamorphidae</a></li> <li><a href="/wiki/Bothrideridae" title="Bothrideridae">Bothrideridae <small>(dry bark beetles)</small></a></li> <li><a href="/wiki/Cerylonidae" title="Cerylonidae">Cerylonidae <small>(minute bark beetles)</small></a></li> <li><a href="/wiki/Coccinellidae" title="Coccinellidae">Coccinellidae <small>(lady beetles, or God's cows)</small></a></li> <li><a href="/wiki/Corylophidae" title="Corylophidae">Corylophidae <small>(minute fungus beetles)</small></a></li> <li><a href="/wiki/Discolomatidae" title="Discolomatidae">Discolomatidae</a></li> <li><a href="/wiki/Endomychidae" title="Endomychidae">Endomychidae <small>(handsome fungus beetles)</small></a></li> <li><a href="/wiki/Eupsilobiidae" title="Eupsilobiidae">Eupsilobiidae</a></li> <li><a href="/wiki/Euxestidae" title="Euxestidae">Euxestidae <small>(well polished beetles)</small></a></li> <li><a href="/wiki/Latridiidae" title="Latridiidae">Latridiidae <small>(minute brown scavenger beetles)</small></a></li> <li><a href="/wiki/Murmidiidae" title="Murmidiidae">Murmidiidae</a></li> <li><a href="/wiki/Mycetaeidae" title="Mycetaeidae">Mycetaeidae</a></li> <li><a href="/wiki/Teredidae" title="Teredidae">Teredidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cucujoidea" title="Cucujoidea">Cucujoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agapytho" title="Agapytho">Agapythidae <small>(<i>Agapytho foveicollis</i>)</small></a></li> <li><a href="/wiki/Boganiidae" title="Boganiidae">Boganiidae</a></li> <li><a href="/wiki/Cavognathidae" title="Cavognathidae">Cavognathidae</a></li> <li><a href="/wiki/Cryptophagidae" title="Cryptophagidae">Cryptophagidae <small>(silken fungus beetles)</small></a></li> <li><a href="/wiki/Cucujidae" title="Cucujidae">Cucujidae <small>(flat bark beetles)</small></a></li> <li><a href="/wiki/Cybocephalidae" title="Cybocephalidae">Cybocephalidae</a></li> <li><a href="/wiki/Cyclaxyridae" title="Cyclaxyridae">Cyclaxyridae</a></li> <li><a href="/wiki/Erotylidae" title="Erotylidae">Erotylidae <small>(pleasing fungus beetles)</small></a></li> <li><a href="/wiki/Helotidae" title="Helotidae">Helotidae</a></li> <li><a href="/wiki/Hobartiidae" title="Hobartiidae">Hobartiidae</a></li> <li><a href="/wiki/Kateretidae" title="Kateretidae">Kateretidae <small>(short-winged flower beetles)</small></a></li> <li><a href="/wiki/Laemophloeidae" title="Laemophloeidae">Laemophloeidae <small>(lined flat bark beetles)</small></a></li> <li><a href="/wiki/Lamingtonium" title="Lamingtonium">Lamingtoniidae <small>(<i>Lamingtonium binnaberrense</i>)</small></a></li> <li><a href="/wiki/Monotomidae" title="Monotomidae">Monotomidae <small>(root-eating beetles)</small></a></li> <li><a href="/wiki/Myraboliidae" class="mw-redirect" title="Myraboliidae">Myraboliidae</a></li> <li><a href="/wiki/Sap_beetle" title="Sap beetle">Nitidulidae <small>(sap beetles)</small></a></li> <li><a href="/wiki/Passandridae" title="Passandridae">Passandridae <small>(parasitic flat bark beetles)</small></a></li> <li><a href="/wiki/Phalacridae" title="Phalacridae">Phalacridae <small>(shining flower beetles)</small></a></li> <li><a href="/wiki/Phloeostichidae" title="Phloeostichidae">Phloeostichidae</a></li> <li><a href="/wiki/Priasilphidae" title="Priasilphidae">Priasilphidae</a></li> <li><a href="/wiki/Propalticidae" title="Propalticidae">Propalticidae</a></li> <li><a href="/wiki/Protocucujidae" title="Protocucujidae">Protocucujidae</a></li> <li><a href="/wiki/Silvanidae" title="Silvanidae">Silvanidae <small>(silvanid flat bark beetles)</small></a></li> <li><a href="/wiki/Smicripidae" title="Smicripidae">Smicripidae <small>(palmetto beetles)</small></a></li> <li><a href="/wiki/Sphindidae" title="Sphindidae">Sphindidae <small>(dry-fungus beetles)</small></a></li> <li><a href="/wiki/Tasmosalpingus" title="Tasmosalpingus">Tasmosalpingidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Weevil" title="Weevil">Curculionoidea</a><br /><small><span style="color:#696969"><span class="nobold">(weevils)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anthribidae" title="Anthribidae">Anthribidae <small>(fungus weevils)</small></a></li> <li><a href="/wiki/Attelabidae" title="Attelabidae">Attelabidae <small>(leaf-rolling weevils)</small></a></li> <li><a href="/wiki/Belidae" title="Belidae">Belidae <small>(primitive weevils)</small></a></li> <li><a href="/wiki/Brentidae" title="Brentidae">Brentidae <small>(straight snout weevils, New York weevil)</small></a></li> <li><a href="/wiki/Caridae" title="Caridae">Caridae</a></li> <li><a class="mw-selflink selflink">Curculionidae <small>(true weevils, bark beetles, ambrosia beetles)</small></a></li> <li><a href="/wiki/Nemonychidae" title="Nemonychidae">Nemonychidae <small>(pine flower weevils)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexyloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexylidae <small>(ship-timber beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Tenebrionoidea" title="Tenebrionoidea">Tenebrionoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aderidae" title="Aderidae">Aderidae <small>(ant-like leaf beetles)</small></a></li> <li><a href="/wiki/Anthicidae" title="Anthicidae">Anthicidae <small>(ant-like flower beetles)</small></a></li> <li><a href="/wiki/Archeocrypticidae" title="Archeocrypticidae">Archeocrypticidae <small>(cryptic fungus beetles)</small></a></li> <li><a href="/wiki/Boridae" title="Boridae">Boridae <small>(conifer bark beetles)</small></a></li> <li><a href="/wiki/Chalcodryidae" title="Chalcodryidae">Chalcodryidae</a></li> <li><a href="/wiki/Ciidae" title="Ciidae">Ciidae <small>(minute tree-fungus beetles)</small></a></li> <li><a href="/wiki/Melandryidae" title="Melandryidae">Melandryidae <small>(false darkling beetles)</small></a></li> <li><a href="/wiki/Blister_beetle" title="Blister beetle">Meloidae <small>(blister beetles)</small></a></li> <li><a href="/wiki/Mordellidae" title="Mordellidae">Mordellidae <small>(tumbling flower beetles)</small></a></li> <li><a href="/wiki/Mycetophagidae" title="Mycetophagidae">Mycetophagidae <small>(hairy fungus beetles)</small></a></li> <li><a href="/wiki/Mycteridae" title="Mycteridae">Mycteridae <small>(palm and flower beetles)</small></a></li> <li><a href="/wiki/Oedemeridae" title="Oedemeridae">Oedemeridae <small>(false blister beetle)</small></a></li> <li><a href="/wiki/Perimylopidae" class="mw-redirect" title="Perimylopidae">Perimylopidae, or Promecheilidae</a></li> <li><a href="/wiki/Prostomidae" title="Prostomidae">Prostomidae <small>(jugular-horned beetles)</small></a></li> <li><a href="/wiki/Pterogeniidae" title="Pterogeniidae">Pterogeniidae</a></li> <li><a href="/wiki/Fire-coloured_beetle" title="Fire-coloured beetle">Pyrochroidae <small>(fire-coloured beetles)</small></a></li> <li><a href="/wiki/Pythidae" title="Pythidae">Pythidae <small>(dead log bark beetles)</small></a></li> <li><a href="/wiki/Ripiphoridae" title="Ripiphoridae">Ripiphoridae <small>(wedge-shaped beetles)</small></a></li> <li><a href="/wiki/Salpingidae" title="Salpingidae">Salpingidae <small>(narrow-waisted bark beetles)</small></a></li> <li><a href="/wiki/Scraptiidae" title="Scraptiidae">Scraptiidae <small>(false flower beetles)</small></a></li> <li><a href="/wiki/Stenotrachelidae" title="Stenotrachelidae">Stenotrachelidae <small>(false longhorn beetles)</small></a></li> <li><a href="/wiki/Synchroidae" title="Synchroidae">Synchroidae <small>(synchroa bark beetles)</small></a></li> <li><a href="/wiki/Darkling_beetle" title="Darkling beetle">Tenebrionidae <small>(darkling beetles)</small></a></li> <li><a href="/wiki/Tetratomidae" title="Tetratomidae">Tetratomidae <small>(polypore fungus beetles)</small></a></li> <li><a href="/wiki/Trictenotomidae" title="Trictenotomidae">Trictenotomidae</a></li> <li><a href="/wiki/Ulodidae" title="Ulodidae">Ulodidae</a></li> <li><a href="/wiki/Zopheridae" title="Zopheridae">Zopheridae <small>(ironclad beetles, cylindrical bark beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Elateriformia" title="Elateriformia">Elateriformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Buprestoidea" title="Buprestoidea">Buprestoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Buprestidae" title="Buprestidae">Buprestidae <small>(jewel beetles, or metallic wood-boring beetles)</small></a></li> <li><a href="/wiki/Schizopodidae" title="Schizopodidae">Schizopodidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Byrrhoidea" title="Byrrhoidea">Byrrhoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Byrrhidae" title="Byrrhidae">Byrrhidae <small>(pill beetles)</small></a></li> <li><a href="/wiki/Callirhipidae" title="Callirhipidae">Callirhipidae <small>(cedar beetles)</small></a></li> <li><a href="/wiki/Chelonariidae" title="Chelonariidae">Chelonariidae <small>(turtle beetles)</small></a></li> <li><a href="/wiki/Cneoglossidae" title="Cneoglossidae">Cneoglossidae</a></li> <li><a href="/wiki/Dryopidae" title="Dryopidae">Dryopidae <small>(long-toed water beetles)</small></a></li> <li><a href="/wiki/Elmidae" title="Elmidae">Elmidae <small>(riffle beetles)</small></a></li> <li><a href="/wiki/Eulichadidae" title="Eulichadidae">Eulichadidae <small>(forest stream beetles)</small></a></li> <li><a href="/wiki/Heteroceridae" title="Heteroceridae">Heteroceridae <small>(variegated mud-loving beetles)</small></a></li> <li><a href="/wiki/Limnichidae" title="Limnichidae">Limnichidae <small>(minute mud beetles)</small></a></li> <li><a href="/wiki/Lutrochidae" class="mw-redirect" title="Lutrochidae">Lutrochidae <small>(travertine beetles)</small></a></li> <li><a href="/wiki/Water-penny_beetles" class="mw-redirect" title="Water-penny beetles">Psephenidae <small>(water-penny beetles)</small></a></li> <li><a href="/wiki/Ptilodactylidae" title="Ptilodactylidae">Ptilodactylidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Dascilloidea" title="Dascilloidea">Dascilloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dascillidae" title="Dascillidae">Dascillidae <small>(soft bodied plant beetles)</small></a></li> <li><a href="/wiki/Rhipiceridae" title="Rhipiceridae">Rhipiceridae <small>(cicada beetle, cicada parasite beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Elateroidea" title="Elateroidea">Elateroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Artematopodidae" title="Artematopodidae">Artematopodidae <small>(soft-bodied plant beetles)</small></a></li> <li><a href="/wiki/Brachypsectridae" title="Brachypsectridae">Brachypsectridae <small>(Texas beetles)</small></a></li> <li><a href="/wiki/Soldier_beetle" title="Soldier beetle">Cantharidae <small>(soldier beetles)</small></a></li> <li><a href="/wiki/Cerophytidae" title="Cerophytidae">Cerophytidae <small>(rare click beetles)</small></a></li> <li><a href="/wiki/Click_beetle" title="Click beetle">Elateridae <small>(click beetles)</small></a></li> <li><a href="/wiki/Eucnemidae" title="Eucnemidae">Eucnemidae <small>(false click beetles)</small></a></li> <li><a href="/wiki/Jurasaidae" title="Jurasaidae">Jurasaidae</a></li> <li><a href="/wiki/Firefly" title="Firefly">Lampyridae <small>(fireflies)</small></a></li> <li><a href="/wiki/Lycidae" title="Lycidae">Lycidae <small>(net-winged beetles)</small></a></li> <li><a href="/wiki/Omethidae" title="Omethidae">Omethidae <small>(false fireflies, long-lipped beetles)</small></a></li> <li><a href="/wiki/Phengodidae" title="Phengodidae">Phengodidae <small>(glowworm beetles)</small></a></li> <li><a href="/wiki/Rhagophthalmidae" title="Rhagophthalmidae">Rhagophthalmidae</a></li> <li><a href="/wiki/Sinopyrophoridae" class="mw-redirect" title="Sinopyrophoridae">Sinopyrophoridae</a></li> <li><a href="/wiki/Throscidae" title="Throscidae">Throscidae <small>(false metallic wood-boring beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipidae <small>(<i>Rhinorhipus tamborinensis</i>)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scirtoidea" title="Scirtoidea">Scirtoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Clambidae" title="Clambidae">Clambidae</a></li> <li><a href="/wiki/Declinia" title="Declinia">Decliniidae <small>(<i>Declinia relicta</i>)</small></a></li> <li><a href="/wiki/Eucinetidae" title="Eucinetidae">Eucinetidae <small>(plate-thigh beetles)</small></a></li> <li><a href="/wiki/Scirtidae" title="Scirtidae">Scirtidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Belohina" title="Belohina">Belohinidae <small>(<i>Belohina inexpectata</i>)</small></a></li> <li><a href="/wiki/Bolboceratidae" title="Bolboceratidae">Bolboceratidae</a></li> <li><a href="/wiki/False_stag_beetle" title="False stag beetle">Diphyllostomatidae <small>(false stag beetles)</small></a></li> <li><a href="/wiki/Geotrupidae" title="Geotrupidae">Geotrupidae <small>(dor beetles)</small></a></li> <li><a href="/wiki/Glaphyridae" title="Glaphyridae">Glaphyridae <small>(bumble bee scarab beetles)</small></a></li> <li><a href="/wiki/Glaresis" title="Glaresis">Glaresidae <small>(enigmatic scarab beetles)</small></a></li> <li><a href="/wiki/Hybosoridae" title="Hybosoridae">Hybosoridae <small>(scavenger scarab beetles)</small></a></li> <li><a href="/wiki/Stag_beetle" title="Stag beetle">Lucanidae <small>(stag beetles)</small></a></li> <li><a href="/wiki/Ochodaeidae" title="Ochodaeidae">Ochodaeidae <small>(sand-loving scarab beetles)</small></a></li> <li><a href="/wiki/Passalidae" title="Passalidae">Passalidae <small>(betsy beetles)</small></a></li> <li><a href="/wiki/Rain_beetle" title="Rain beetle">Pleocomidae <small>(rain beetles)</small></a></li> <li><a href="/wiki/Scarabaeidae" title="Scarabaeidae">Scarabaeidae <small>(scarabs)</small></a></li> <li><a href="/wiki/Trogidae" title="Trogidae">Trogidae <small>(hide beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Staphyliniformia" title="Staphyliniformia">Staphyliniformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Histeroidea" title="Histeroidea">Histeroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Histeridae" title="Histeridae">Histeridae <small>(clown beetles)</small></a></li> <li><a href="/wiki/Sphaerites" title="Sphaerites">Sphaeritidae <small>(false clown beetles)</small></a></li> <li><a href="/wiki/Syntelia" title="Syntelia">Synteliidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Hydrophiloidea" title="Hydrophiloidea">Hydrophiloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Epimetopidae" title="Epimetopidae">Epimetopidae</a></li> <li><a href="/wiki/Georissus" title="Georissus">Georissidae <small>(minute mud-loving beetles)</small></a></li> <li><a href="/wiki/Helophorus" title="Helophorus">Helophoridae</a></li> <li><a href="/wiki/Hydrochidae" class="mw-redirect" title="Hydrochidae">Hydrochidae</a></li> <li><a href="/wiki/Hydrophilidae" title="Hydrophilidae">Hydrophilidae <small>(water scavenger beetles)</small></a></li> <li><a href="/wiki/Spercheidae" class="mw-redirect" title="Spercheidae">Spercheidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Staphylinoidea" title="Staphylinoidea">Staphylinoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agyrtidae" title="Agyrtidae">Agyrtidae <small>(primitive carrion beetles)</small></a></li> <li><a href="/wiki/Hydraenidae" title="Hydraenidae">Hydraenidae</a></li> <li><a href="/wiki/Leiodidae" title="Leiodidae">Leiodidae <small>(round fungus beetles)</small></a></li> <li><a href="/wiki/Ptiliidae" title="Ptiliidae">Ptiliidae <small>(feather-winged beetles)</small></a></li> <li><a href="/wiki/Silphidae" title="Silphidae">Silphidae <small>(carrion beetles)</small></a></li> <li><a href="/wiki/Rove_beetle" title="Rove beetle">Staphylinidae <small>(rove beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/List_of_subgroups_of_the_order_Coleoptera" title="List of subgroups of the order Coleoptera">List of subgroups of the order Coleoptera</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Curculionidae</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q7415384" class="extiw" title="wikidata:Q7415384">Q7415384</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Curculionidae" class="extiw" title="wikispecies:Curculionidae">Curculionidae</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Animal_Diversity_Web" title="Animal Diversity Web">ADW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://animaldiversity.org/accounts/Curculionidae/">Curculionidae</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Faunal_Directory" title="Australian Faunal Directory">AFD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://biodiversity.org.au/afd/taxa/Curculionidae">Curculionidae</a></span></span></li> <li><span style="white-space:nowrap;">BioLib: <span class="uid"><a rel="nofollow" class="external text" href="https://www.biolib.cz/en/taxon/id12366">12366</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=1752">1752</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/BugGuide" title="BugGuide">BugGuide</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://bugguide.net/node/view/139">139</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/8HBQ5">8HBQ5</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/5240">5240</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/EPPO_Code" title="EPPO Code">EPPO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://gd.eppo.int/taxon/1CURCF">1CURCF</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:faunaeur.org:taxname:11185">11185</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea (new)</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://fauna-eu.org/cdm_dataportal/taxon/30f5352c-4936-4191-8733-bc358913fa9c">30f5352c-4936-4191-8733-bc358913fa9c</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/4239">4239</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/48736">48736</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=104606">104606</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=114666">114666</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Biodiversity_Network" title="National Biodiversity Network">NBN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://data.nbn.org.uk/Taxa/NHMSYS0020151946">NHMSYS0020151946</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=7042">7042</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/76119f93-79f6-4c83-ae86-49ee3b25f76a">76119f93-79f6-4c83-ae86-49ee3b25f76a</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=787472">787472</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=69436">69436</a></span></span></li> <li><span style="white-space:nowrap;">PPE: <span class="uid"><a rel="nofollow" class="external text" href="https://bladmineerders.nl/parasites/animalia/arthropoda/insecta/coleoptera/polyphaga/cucujiformia/curculionoidea/curculionidae/">curculionidae</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Plazi" title="Plazi">Plazi</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://treatment.plazi.org/id/B718293D-FFEC-F228-FF0D-FDB6BB36F615">B718293D-FFEC-F228-FF0D-FDB6BB36F615</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=150716">150716</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1038841319">.mw-parser-output .tooltip-dotted{border-bottom:1px dotted;cursor:help}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319"><link 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