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mw-first-heading"><i>Diandongosuchus</i></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. 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href="/wiki/Middle_Triassic" title="Middle Triassic">Middle Triassic</a> <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">242–237 <a href="/wiki/Megaannum" class="mw-redirect" title="Megaannum">Ma</a></span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"> <div style="position:absolute; left:138.09230769231px; font-size:50%"><div style="position:relative; left:-0.42em">↓</div></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Diandongosuchus_fuyuanensis_334.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Diandongosuchus_fuyuanensis_334.jpg/240px-Diandongosuchus_fuyuanensis_334.jpg" decoding="async" width="240" height="136" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Diandongosuchus_fuyuanensis_334.jpg/360px-Diandongosuchus_fuyuanensis_334.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Diandongosuchus_fuyuanensis_334.jpg/480px-Diandongosuchus_fuyuanensis_334.jpg 2x" data-file-width="700" data-file-height="397" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Skull in <a href="/wiki/Beijing_Museum_of_Natural_History" class="mw-redirect" title="Beijing Museum of Natural History">Beijing Museum of Natural History</a> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Diandongosuchus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Reptile" title="Reptile">Reptilia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Archosauromorpha" title="Archosauromorpha">Archosauromorpha</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Archosauriformes" title="Archosauriformes">Archosauriformes</a> </td></tr> <tr> <td>Order: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Phytosaur" title="Phytosaur">Phytosauria</a> </td></tr> <tr> <td>Genus: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a class="mw-selflink selflink"><i>Diandongosuchus</i></a><br /><small>Li <i>et al.</i>, <a href="/wiki/2012_in_paleontology" title="2012 in paleontology">2012</a></small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Type_species" title="Type species">Type species</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><b>Diandongosuchus fuyuanensis</b></i><br /><div style="font-size: 85%;">Li <i>et al.</i>, 2012</div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Diandongosuchus</b></i> is an extinct genus of <a href="/wiki/Archosauriformes" title="Archosauriformes">archosauriform</a> reptile, possibly a member of the <a href="/wiki/Phytosaur" title="Phytosaur">Phytosauria</a>, known from the <a href="/wiki/Middle_Triassic" title="Middle Triassic">Middle Triassic</a> of China. The <a href="/wiki/Type_species" title="Type species">type species</a> <i><b>Diandongosuchus fuyuanensis</b></i> was named in 2012 from the <a href="/wiki/Zhuganpo_Formation" title="Zhuganpo Formation">Zhuganpo Formation</a> of <a href="/wiki/Yunnan_Province" class="mw-redirect" title="Yunnan Province">Yunnan Province</a>. It is a marine species that shows similarities with another Chinese Triassic species called <i><a href="/wiki/Qianosuchus_mixtus" class="mw-redirect" title="Qianosuchus mixtus">Qianosuchus mixtus</a></i>, although it has fewer adaptations toward marine life. It was originally classified as the <a href="/wiki/Basal_(phylogenetics)" title="Basal (phylogenetics)">basal</a>-most member of the <a href="/wiki/Pseudosuchia" title="Pseudosuchia">pseudosuchian</a> clade <a href="/wiki/Poposauroidea" title="Poposauroidea">Poposauroidea</a>.<sup id="cite_ref-Letal12_1-0" class="reference"><a href="#cite_note-Letal12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> However, a subsequent study conducted by Stocker <i>et al.</i> (2016, 2017) indicated it to be the basalmost known phytosaur instead.<sup id="cite_ref-:0_2-0" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_3-0" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Diandongosuchus&action=edit&section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Diandongosuchus_fuyuanensis.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/21/Diandongosuchus_fuyuanensis.jpg/220px-Diandongosuchus_fuyuanensis.jpg" decoding="async" width="220" height="176" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/21/Diandongosuchus_fuyuanensis.jpg/330px-Diandongosuchus_fuyuanensis.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/21/Diandongosuchus_fuyuanensis.jpg/440px-Diandongosuchus_fuyuanensis.jpg 2x" data-file-width="2000" data-file-height="1600" /></a><figcaption>Restoration</figcaption></figure> <p><i>Diandongosuchus</i> is known from a nearly complete articulated skeleton (<a href="/wiki/Zhejiang_Museum_of_Natural_History" title="Zhejiang Museum of Natural History">ZMNH</a> M8770) missing most of the tail. The total length of ZMNH M8770 is 97 centimetres (3.18 ft) and the estimated body length of the animal in life is around 155 centimetres (5.09 ft). The specimen is preserved on its right side, with the underside of the lower jaws and the trunk showing. It was prepared out of a limestone slab to reveal details on the left side of the skeleton, many of which are better preserved. The skull of <i>Diandongosuchus</i> is pointed, with oval-shaped eye sockets, <a href="/wiki/Antorbital_fenestra" title="Antorbital fenestra">antorbital</a> and <a href="/wiki/Temporal_fenestra" title="Temporal fenestra">temporal</a> openings. Distinctive features include a long <a href="/wiki/Premaxilla" title="Premaxilla">premaxilla</a> bone at the tip of the snout that extends backward past the nostril openings, a large ridge on the <a href="/wiki/Jugal_bone" title="Jugal bone">jugal bone</a> that runs beneath the eye socket, and two supratemporal openings on the <a href="/wiki/Skull_table" class="mw-redirect" title="Skull table">skull table</a> that have prominent ridges surrounding them. The skull has similar proportions to that of <i>Qianosuchus</i>, and has the same number of teeth in the premaxilla. Like the terrestrial poposauroid <i>Poposaurus</i>, <i>Diandongosuchus</i> has a <a href="/wiki/Maxilla" title="Maxilla">maxilla</a> (upper jaw) bone that does not reach the border of the nostril opening.<sup id="cite_ref-Letal12_1-1" class="reference"><a href="#cite_note-Letal12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p><p>ZMNH M8770 has 25 vertebrae in the back and neck, two <a href="/wiki/Sacral_vertebra" class="mw-redirect" title="Sacral vertebra">sacral vertebrae</a> (as in most Triassic pseudosuchians), and seven of the forward-most tail vertebrae. The neck vertebrae are taller and narrower than they are in <i>Qianosuchus</i>. Most of the back vertebrae are obscured by overlying ribs. At the back of the trunk near the hips are bones belonging to small vertebrates such as fish - likely the stomach contents of the individual. Small overlapping <a href="/wiki/Osteoderm" title="Osteoderm">osteoderms</a> (bony scutes) overlay many of the vertebrae. Two rows run along the neck, back, and tail with about two osteoderms overlaying each vertebra. Small osteoderms also cover the limb bones.<sup id="cite_ref-Letal12_1-2" class="reference"><a href="#cite_note-Letal12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p><p>Some features of the limbs, <a href="/wiki/Pelvic_girdle" class="mw-redirect" title="Pelvic girdle">pelvic</a> and <a href="/wiki/Pectoral_girdle" class="mw-redirect" title="Pectoral girdle">pectoral girdles</a> are also diagnostic in <i>Diandongosuchus</i>, including a thick <a href="/wiki/Ischium" title="Ischium">ischium</a> bone in the hip, an opening of the <a href="/wiki/Coracoid" title="Coracoid">coracoid</a> bone in the pectoral girdle that is much larger than those of other archosaurs and is closed by the end of the <a href="/wiki/Scapula" title="Scapula">scapula</a>, and a fourth <a href="/wiki/Metatarsal" class="mw-redirect" title="Metatarsal">metatarsal</a> bone in the foot that is longer than the other metatarsals. The scapula of <i>Diandongosuchus</i> is longer and narrower than that of <i>Qianosuchus</i>. The <a href="/wiki/Iliac_crest" title="Iliac crest">iliac blade</a> of the hip is unusual in that it is narrow and projects far back from the rest of the hip. As in <i>Qianosuchus</i>, the femur of <i>Diandongosuchus</i> is slightly twisted, but the fibula is thinner and more curved. The <a href="/wiki/Talus_bone" title="Talus bone">astragalus</a> and <a href="/wiki/Calcaneum" class="mw-redirect" title="Calcaneum">calcaneum</a> bones of the ankle fit together like a ball-and-socket, a feature that confirms <i>Diandongosuchus</i> as a pseudosuchian. Some of the <a href="/wiki/Phalanges" class="mw-redirect" title="Phalanges">phalanges</a> or toe bones are missing in ZMNH M8770, but the metatarsals are present and have unique proportions among Triassic archosaurs in which the fourth is longer than the third.<sup id="cite_ref-Letal12_1-3" class="reference"><a href="#cite_note-Letal12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Diandongosuchus&action=edit&section=2" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Diandongosuchus_skeleton.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/5e/Diandongosuchus_skeleton.jpg/220px-Diandongosuchus_skeleton.jpg" decoding="async" width="220" height="159" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/5e/Diandongosuchus_skeleton.jpg/330px-Diandongosuchus_skeleton.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/5e/Diandongosuchus_skeleton.jpg/440px-Diandongosuchus_skeleton.jpg 2x" data-file-width="926" data-file-height="671" /></a><figcaption>Diagram of the holotype specimen with insert photos showing traits shared with <a href="/wiki/Phytosaurs" class="mw-redirect" title="Phytosaurs">phytosaurs</a></figcaption></figure> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-Expand_section plainlinks metadata ambox mbox-small-left ambox-content" role="presentation"><tbody><tr><td class="mbox-image"><span typeof="mw:File"><a href="/wiki/File:Wiki_letter_w_cropped.svg" class="mw-file-description"><img alt="[icon]" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/20px-Wiki_letter_w_cropped.svg.png" decoding="async" width="20" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/30px-Wiki_letter_w_cropped.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/40px-Wiki_letter_w_cropped.svg.png 2x" data-file-width="44" data-file-height="31" /></a></span></td><td class="mbox-text"><div class="mbox-text-span">This section <b>needs expansion</b>. You can help by <a class="external text" href="https://en.wikipedia.org/w/index.php?title=Diandongosuchus&action=edit&section=">adding to it</a>. <span class="date-container"><i>(<span class="date">June 2017</span>)</i></span></div></td></tr></tbody></table> <p>A phylogenetic analysis conducted by Li <i>et al.</i> (2012) in the original description of <i>Diandongosuchus</i> showed that it was the most basal member of a clade called Poposauroidea, which includes mostly terrestrial pseudosuchians such as the bipedal <i><a href="/wiki/Poposaurus" title="Poposaurus">Poposaurus</a></i> and the sail-backed <i><a href="/wiki/Arizonasaurus" title="Arizonasaurus">Arizonasaurus</a></i>. It was found to be closely related to <i>Qianosuchus</i>, an aquatic pseudosuchian that was the second most basal member of Poposauroidea. The data matrix of Li <i>et al.</i>, a list of characteristics that was used in the analysis, was based on that of Nesbitt (2011), one of the most extensive on archosaurs. Because of this, many of the relationships found by Li <i>et al.</i> are the same as those found by Nesbitt. Below is a cladogram from the analysis:<sup id="cite_ref-Letal12_1-4" class="reference"><a href="#cite_note-Letal12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1261134923">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto;display:table}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:85%;line-height:85%"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Archosauria" class="mw-redirect" title="Archosauria">Archosauria</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Pseudosuchia" title="Pseudosuchia">Pseudosuchia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Ornithosuchidae" title="Ornithosuchidae">Ornithosuchidae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Ornithosuchus" title="Ornithosuchus">Ornithosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Riojasuchus" title="Riojasuchus">Riojasuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Suchia" title="Suchia">Suchia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Gracilisuchus" title="Gracilisuchus">Gracilisuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Turfanosuchus" title="Turfanosuchus">Turfanosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Revueltosaurus" title="Revueltosaurus">Revueltosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Aetosauria" class="mw-redirect" title="Aetosauria">Aetosauria</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Longosuchus" title="Longosuchus">Longosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Stagonolepis" title="Stagonolepis">Stagonolepis</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Aetosaurus" title="Aetosaurus">Aetosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Paracrocodylomorpha" title="Paracrocodylomorpha">Paracrocodylomorpha</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Loricata" title="Loricata">Loricata</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Poposauroidea" title="Poposauroidea">Poposauroidea</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Diandongosuchus</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Qianosuchus" title="Qianosuchus">Qianosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Xilousuchus" title="Xilousuchus">Xilousuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Arizonasaurus" title="Arizonasaurus">Arizonasaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Poposaurus" title="Poposaurus">Poposaurus</a></i> (holotype) </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Poposaurus" title="Poposaurus">Poposaurus</a></i> (Yale specimen) </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Lotosaurus" title="Lotosaurus">Lotosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Sillosuchus" title="Sillosuchus">Sillosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261134923"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Effigia" title="Effigia">Effigia</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Shuvosaurus" title="Shuvosaurus">Shuvosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <p>However, more recent studies have found it to be a basal <a href="/wiki/Phytosaur" title="Phytosaur">phytosaur</a>.<sup id="cite_ref-:0_2-1" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_3-1" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Paleoecology">Paleoecology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Diandongosuchus&action=edit&section=3" title="Edit section: Paleoecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Diandongosuchus_skull.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Diandongosuchus_skull.jpg/220px-Diandongosuchus_skull.jpg" decoding="async" width="220" height="181" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Diandongosuchus_skull.jpg/330px-Diandongosuchus_skull.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Diandongosuchus_skull.jpg/440px-Diandongosuchus_skull.jpg 2x" data-file-width="926" data-file-height="760" /></a><figcaption>Skull features</figcaption></figure> <p><i>Diandongosuchus</i> was found in a <a href="/wiki/Ladinian" title="Ladinian">Ladinian</a>-age marine limestone formation that has preserved many marine reptiles including <a href="/wiki/Thallatosaur" class="mw-redirect" title="Thallatosaur">thallatosaurs</a>, <a href="/wiki/Nothosaur" title="Nothosaur">nothosaurs</a>, <a href="/wiki/Pistosaur" class="mw-redirect" title="Pistosaur">pistosaurs</a>, and some <a href="/wiki/Protorosaur" class="mw-redirect" title="Protorosaur">protorosaurs</a>. The closely related pseudosuchian <i>Qianosuchus</i> was found in a marine deposit about 50 kilometres (31 mi) northwest of the <i>Diandongosuchus</i> locality that is slightly older (<a href="/wiki/Anisian" title="Anisian">Anisian</a> in age) and possesses many features consistent with a marine lifestyle. However, <i>Diandongosuchus</i> shows no features that are clear adaptations to a marine lifestyle. Possible adaptations include nostrils that are positioned slightly farther back on the skull than most terrestrial pseudosuchians and a greater number of premaxillary teeth (a feature seen in possible semiaquatic archosaurs such as <i><a href="/wiki/Chanaresuchus" title="Chanaresuchus">Chanaresuchus</a></i> and <a href="/wiki/Spinosaurid" class="mw-redirect" title="Spinosaurid">spinosaurids</a>). Fish bones within its stomach contents are additional evidence that it was a marine archosaur. <i>Diandongosuchus</i> may have had a similar lifestyle to modern marine crocodylians like the <a href="/wiki/Saltwater_crocodile" title="Saltwater crocodile">saltwater crocodile</a> that live along coastlines yet are not fully marine.<sup id="cite_ref-Letal12_1-5" class="reference"><a href="#cite_note-Letal12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p><p>The fossil assemblage in which <i>Diandongosuchus</i> was found bears many similarities to that of European fossil localities such as <a href="/wiki/Monte_San_Giorgio" title="Monte San Giorgio">Monte San Giorgio</a>. Both include marine reptiles like thallatosaurs and nothosaurs and probably represented environments along the northern shorelines of the <a href="/wiki/Tethys_Ocean" title="Tethys Ocean">Tethys Ocean</a>. No marine archosaurs like <i>Diandongosuchus</i> and <i>Qianosuchus</i> are known from Europe, although the pseudosuchian <i><a href="/wiki/Ticinosuchus" title="Ticinosuchus">Ticinosuchus</a></i> from Monte San Giorgio was probably adapted to life along the shorelines of the Tethys. In the analyses of Li <i>et al.</i> (2012) and Nesbitt (2011), <i>Ticinosuchus</i> is either the most basal member of a clade called <a href="/wiki/Loricata" title="Loricata">Loricata</a> which is the <a href="/wiki/Sister_taxon" class="mw-redirect" title="Sister taxon">sister taxon</a> of Poposauroidea, or the sister taxon of <a href="/wiki/Paracrocodylomorpha" title="Paracrocodylomorpha">Paracrocodylomorpha</a> which includes both Loricata and Poposauroidea.<sup id="cite_ref-Letal12_1-6" class="reference"><a href="#cite_note-Letal12-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Although <i>Ticinosuchus</i> and <i>Diandongosuchus</i> were initially believed to have been very closely related basal paracrocodylomorphs, this hypothesis is invalidated if <i>Diandongosuchus</i> is a phytosaur as other studies have shown.<sup id="cite_ref-:1_3-2" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Diandongosuchus&action=edit&section=4" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-Letal12-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Letal12_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Letal12_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Letal12_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Letal12_1-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Letal12_1-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-Letal12_1-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-Letal12_1-6"><sup><i><b>g</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFLiWuZhaoSato2012" class="citation journal cs1">Li, C.; Wu, X. C.; Zhao, L. J.; Sato, T.; Wang, L. T. (2012). <a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/254315203">"A new archosaur (Diapsida, Archosauriformes) from the marine Triassic of China"</a>. <i>Journal of Vertebrate Paleontology</i>. <b>32</b> (5): 1064. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1080%2F02724634.2012.694383">10.1080/02724634.2012.694383</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:86797826">86797826</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Vertebrate+Paleontology&rft.atitle=A+new+archosaur+%28Diapsida%2C+Archosauriformes%29+from+the+marine+Triassic+of+China&rft.volume=32&rft.issue=5&rft.pages=1064&rft.date=2012&rft_id=info%3Adoi%2F10.1080%2F02724634.2012.694383&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A86797826%23id-name%3DS2CID&rft.aulast=Li&rft.aufirst=C.&rft.au=Wu%2C+X.+C.&rft.au=Zhao%2C+L.+J.&rft.au=Sato%2C+T.&rft.au=Wang%2C+L.+T.&rft_id=https%3A%2F%2Fwww.researchgate.net%2Fpublication%2F254315203&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADiandongosuchus" class="Z3988"></span></span> </li> <li id="cite_note-:0-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-:0_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:0_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMichelle_R._StockerSterling_J._NesbittLi-Jun_ZhaoXiao-Chun_Wu2016" class="citation journal cs1">Michelle R. Stocker; Sterling J. Nesbitt; Li-Jun Zhao; Xiao-Chun Wu; Chun Li (2016). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20161018211820/http://vertpaleo.org/PDFS/2016/SVP-2016-Program-Book-v10-with-covers.aspx">"Mosaic evolution in Phytosauria: the origin of long-snouted morphologies based on a complete skeleton of a phytosaur from the Middle Triassic of China"</a>. <i>Society of Vertebrate Paleontology 76th Annual Meeting Program & Abstracts</i>: 232. Archived from <a rel="nofollow" class="external text" href="http://vertpaleo.org/PDFS/2016/SVP-2016-Program-Book-v10-with-covers.aspx">the original</a> on 2016-10-18<span class="reference-accessdate">. Retrieved <span class="nowrap">2016-10-30</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Society+of+Vertebrate+Paleontology+76th+Annual+Meeting+Program+%26+Abstracts&rft.atitle=Mosaic+evolution+in+Phytosauria%3A+the+origin+of+long-snouted+morphologies+based+on+a+complete+skeleton+of+a+phytosaur+from+the+Middle+Triassic+of+China&rft.pages=232&rft.date=2016&rft.au=Michelle+R.+Stocker&rft.au=Sterling+J.+Nesbitt&rft.au=Li-Jun+Zhao&rft.au=Xiao-Chun+Wu&rft.au=Chun+Li&rft_id=http%3A%2F%2Fvertpaleo.org%2FPDFS%2F2016%2FSVP-2016-Program-Book-v10-with-covers.aspx&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADiandongosuchus" class="Z3988"></span></span> </li> <li id="cite_note-:1-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-:1_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:1_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:1_3-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMichelle_R._StockerLi-Jun_ZhaoSterling_J._NesbittXiao-Chun_Wu2017" class="citation journal cs1">Michelle R. Stocker; Li-Jun Zhao; Sterling J. Nesbitt; Xiao-Chun Wu; Chun Li (2017). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5385495">"A Short-Snouted, Middle Triassic Phytosaur and its Implications for the Morphological Evolution and Biogeography of Phytosauria"</a>. <i>Scientific Reports</i>. <b>7</b>: Article number 46028. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2017NatSR...746028S">2017NatSR...746028S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fsrep46028">10.1038/srep46028</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5385495">5385495</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/28393843">28393843</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Scientific+Reports&rft.atitle=A+Short-Snouted%2C+Middle+Triassic+Phytosaur+and+its+Implications+for+the+Morphological+Evolution+and+Biogeography+of+Phytosauria&rft.volume=7&rft.pages=Article+number+46028&rft.date=2017&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5385495%23id-name%3DPMC&rft_id=info%3Apmid%2F28393843&rft_id=info%3Adoi%2F10.1038%2Fsrep46028&rft_id=info%3Abibcode%2F2017NatSR...746028S&rft.au=Michelle+R.+Stocker&rft.au=Li-Jun+Zhao&rft.au=Sterling+J.+Nesbitt&rft.au=Xiao-Chun+Wu&rft.au=Chun+Li&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5385495&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADiandongosuchus" class="Z3988"></span></span> </li> </ol></div></div> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist 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.navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Phytosauria" title="Template:Phytosauria"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Phytosauria" title="Template talk:Phytosauria"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Phytosauria" title="Special:EditPage/Template:Phytosauria"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Phytosauria" style="font-size:114%;margin:0 4em"><a href="/wiki/Phytosaur" title="Phytosaur">Phytosauria</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*_Kingdom:_Animalia_*_Phylum:_Chordata_*_Class:_Sauropsida_*_Clade:_Archosauromorpha_*_Clade:_Crurotarsi" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size:85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Sauropsida" title="Sauropsida">Sauropsida</a></li> <li><i>Clade</i>: <a href="/wiki/Archosauromorpha" title="Archosauromorpha">Archosauromorpha</a></li> <li><i>Clade</i>: <a href="/wiki/Crurotarsi" title="Crurotarsi">Crurotarsi</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Sauropsida" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sauropsida" title="Sauropsida">Sauropsida</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Sauropsida" title="Template:Sauropsida">Sauropsida</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Archosauromorpha" title="Archosauromorpha">Archosauromorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Archosauromorpha" title="Template:Archosauromorpha">Archosauromorpha</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Phytosaur" title="Phytosaur">Phytosauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background:#ddddff;font-size:115%"><div><b><a href="/wiki/Phytosaur" title="Phytosaur">Phytosauria</a></b></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Phytosauria" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Phytosaur" title="Phytosaur">Phytosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a class="mw-selflink selflink">Diandongosuchus</a></i></li> <li><i><a href="/wiki/Heteropelta" title="Heteropelta">Heteropelta</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Parasuchidae" title="Parasuchidae">Parasuchidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Arganarhinus" title="Arganarhinus">Arganarhinus</a></i></li> <li><i><a href="/wiki/Ebrachosuchus" title="Ebrachosuchus">Ebrachosuchus</a></i></li> <li><i><a href="/wiki/Paleorhinus" title="Paleorhinus">Paleorhinus</a></i></li> <li><i><a href="/wiki/Parasuchus" title="Parasuchus">Parasuchus</a></i></li> <li><i><a href="/wiki/Wannia" title="Wannia">Wannia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Mystriosuchinae" class="mw-redirect" title="Mystriosuchinae">Mystriosuchinae</a><br />/ <a href="/wiki/Phytosauridae" class="mw-redirect" title="Phytosauridae">Phytosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Angistorhinus" title="Angistorhinus">Angistorhinus</a></i></li> <li><i><a href="/wiki/Brachysuchus" title="Brachysuchus">Brachysuchus</a></i></li> <li><i><a href="/wiki/Colossosuchus" title="Colossosuchus">Colossosuchus</a></i></li> <li><i><a href="/wiki/Jupijkam" title="Jupijkam">Jupijkam</a></i></li> <li><i><a href="/wiki/Protome_batalaria" title="Protome batalaria">Protome</a></i></li> <li><i><a href="/wiki/Rutiodon" title="Rutiodon">Rutiodon</a></i></li> <li><i><a href="/wiki/Volcanosuchus" title="Volcanosuchus">Volcanosuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Leptosuchomorpha" title="Leptosuchomorpha">Leptosuchomorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Leptosuchus" title="Leptosuchus">Leptosuchus</a></i></li> <li><i><a href="/wiki/Pravusuchus" title="Pravusuchus">Pravusuchus</a></i></li> <li><i><a href="/wiki/Smilosuchus" title="Smilosuchus">Smilosuchus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Mystriosuchini" title="Mystriosuchini">Mystriosuchini</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Angistorhinopsis" title="Angistorhinopsis">Angistorhinopsis</a></i></li> <li><i><a href="/wiki/Machaeroprosopus" title="Machaeroprosopus">Machaeroprosopus</a></i></li> <li><i><a href="/wiki/Mystriosuchus" title="Mystriosuchus">Mystriosuchus</a></i></li> <li><i><a href="/wiki/Nicrosaurus" title="Nicrosaurus">Nicrosaurus</a></i></li> <li><i><a href="/wiki/Redondasaurus" title="Redondasaurus">Redondasaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nomen_dubium" title="Nomen dubium">Nomina dubia</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Belodon" title="Belodon">Belodon</a></i></li> <li><i><a href="/wiki/Centemodon" title="Centemodon">Centemodon</a></i></li> <li><i><a href="/wiki/Clepsysaurus" title="Clepsysaurus">Clepsysaurus</a></i>?</li> <li><i><a href="/wiki/Coburgosuchus" title="Coburgosuchus">Coburgosuchus</a></i></li> <li><i><a href="/wiki/Compsosaurus" title="Compsosaurus">Compsosaurus</a></i></li> <li><i><a href="/wiki/Francosuchus" title="Francosuchus">Francosuchus</a></i></li> <li><i><a href="/wiki/Heterodontosuchus" title="Heterodontosuchus">Heterodontosuchus</a></i></li> <li><i><a href="/wiki/Mesorhinosuchus" title="Mesorhinosuchus">Mesorhinosuchus</a></i></li> <li><i><a href="/wiki/Omosaurus" title="Omosaurus">Omosaurus</a></i>?</li> <li><i><a href="/wiki/Palaeoctonus" title="Palaeoctonus">Palaeoctonus</a></i>?</li> <li><i><a href="/wiki/Phytosaurus" title="Phytosaurus">Phytosaurus</a></i></li> <li><i><a href="/wiki/Rileyasuchus" title="Rileyasuchus">Rileyasuchus</a></i></li> <li><i><a href="/wiki/Suchoprion" title="Suchoprion">Suchoprion</a></i></li> <li><i><a href="/wiki/Termatosaurus" title="Termatosaurus">Termatosaurus</a></i>?</li></ul> </div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Smilosuchus_adamanensis_flipped.jpg" class="mw-file-description" title="Smilosuchus adamanensis"><img alt="Smilosuchus adamanensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Smilosuchus_adamanensis_flipped.jpg/140px-Smilosuchus_adamanensis_flipped.jpg" decoding="async" width="140" height="31" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Smilosuchus_adamanensis_flipped.jpg/210px-Smilosuchus_adamanensis_flipped.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/17/Smilosuchus_adamanensis_flipped.jpg/280px-Smilosuchus_adamanensis_flipped.jpg 2x" data-file-width="2894" data-file-height="648" /></a></span></span></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Phytosaurs" title="Category:Phytosaurs">Category</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Diandongosuchus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q5271376" class="extiw" title="wikidata:Q5271376">Q5271376</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=1480950">1480950</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=246047">246047</a></span></span></li></ul> </div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1&useformat=desktop" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Diandongosuchus&oldid=1201032966">https://en.wikipedia.org/w/index.php?title=Diandongosuchus&oldid=1201032966</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:Phytosaurs" 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