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PhD Degree Program in Biophysics (BP) · UCSF
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PhD</a><ul><li class="menu-880056 menu-path-node-17191 first odd"><a href="/degree-program/path-to-phd" tabindex="0">Path to PhD Overview</a></li> <li class="menu-880176 menu-path-node-16971 even"><a href="/degree-program/path-to-phd/guidelines" tabindex="0">Path to PhD Guidelines</a></li> <li class="menu-956986 menu-path-node-18776 odd"><a href="/degree-program/path-to-phd/rotations-research-advisor" tabindex="0">Rotations and Advisor Selection</a></li> <li class="menu-880106 menu-path-node-16536 even"><a href="/degree-program/path-to-phd/qbc-journal-club-requirements" tabindex="0">QBC Journal Club Requirements</a></li> <li class="menu-956991 menuparent menu-path-node-18796 odd"><a tabindex="0">Qualifying Examination</a><ul><li class="menu-956991 menu-path-node-18796 first odd"><a href="/degree-program/path-to-phd/qualifying-exam" tabindex="0">Qualifying Examination Overview</a></li> <li class="menu-956996 menu-path-node-18791 even"><a href="/degree-program/path-to-phd/qualifying-exam/about" tabindex="0">About the Examination</a></li> <li class="menu-957001 menu-path-node-18786 odd"><a href="/degree-program/path-to-phd/qualifying-exam/preparation" tabindex="0">Preparation</a></li> <li class="menu-957006 menu-path-node-18781 even last"><a href="/degree-program/path-to-phd/qualifying-exam/advice" tabindex="0">Student and Faculty Advice</a></li> </ul></li> <li class="menu-880141 menu-path-node-16671 even"><a href="/degree-program/path-to-phd/advancement-thesis" tabindex="0">Advancement to Candidacy and Thesis Progression</a></li> <li class="menu-961261 menu-path-node-19686 odd"><a href="/degree-program/path-to-phd/how-to-navigate-difficult-situation" tabindex="0">How to Navigate a Difficult Situation</a></li> <li class="menu-880136 menu-path-node-16566 even last"><a href="/degree-program/path-to-phd/six-year-policy" tabindex="0">Six-year Policy</a></li> </ul></li> <li class="menu-857736 menu-path-node-16051 odd last"><a href="/degree-program/career-development" tabindex="0">Career Development</a></li> </ul></li> <li class="menu-857791 menuparent menu-path-node-16656 even"><a tabindex="0">Admissions</a><ul><li class="menu-857791 menu-path-node-16656 first odd"><a href="/admissions" tabindex="0">Admissions Overview</a></li> <li class="menu-956496 menuparent menu-path-node-18366 even last"><a tabindex="0">Life After Biophysics</a><ul><li class="menu-956496 menu-path-node-18366 first odd"><a href="/admissions/life-after-biophysics" tabindex="0">Life After Biophysics Overview</a></li> <li class="menu-956546 menu-path-node-18586 even"><a href="/admissions/life-after-biophysics/lillian-chong" tabindex="0">Lillian Chong, PhD</a></li> <li class="menu-956501 menu-path-node-18376 odd"><a href="/admissions/life-after-biophysics/stephen-floor" tabindex="0">Stephen Floor, PhD</a></li> <li class="menu-956506 menu-path-node-18371 even last"><a href="/admissions/life-after-biophysics/jeremy-wilbur" tabindex="0">Jeremy Wilbur, PhD</a></li> </ul></li> </ul></li> <li class="menu-819541 menuparent menu-path-node-16036 odd"><a tabindex="0">People</a><ul><li class="menu-819541 menu-path-node-16036 first odd"><a href="/people" tabindex="0">People Overview</a></li> <li class="menu-961981 menu-path-node-21561 even"><a href="/people/manglik" tabindex="0">Director Aashish Manglik, MD, PhD</a></li> <li class="menu-857906 menu-path-node-16691 odd"><a href="/people/flowers" tabindex="0">Manager Nicole Flowers</a></li> <li class="menu-857981 menu-path-node-16706 even"><a href="/people/faculty" tabindex="0">Faculty</a></li> <li class="menu-857986 menu-path-node-16711 odd"><a href="/people/students" tabindex="0">Students</a></li> <li class="menu-857991 menu-path-node-16716 even"><a href="/people/alumni" tabindex="0">Alumni</a></li> <li class="menu-857891 menu-path-node-16681 odd"><a href="/people/honors-awards-faculty" tabindex="0">Faculty Honors and Awards</a></li> <li class="menu-857896 menu-path-node-16686 even"><a href="/people/honors-awards-student" tabindex="0">Student Honors and Awards</a></li> <li class="menu-962891 menu-path-node-22866 odd last"><a href="/people/honors-awards-student/bp-awards" tabindex="0">Biophysics Graduate Program Awards</a></li> </ul></li> <li class="menu-857996 menuparent menu-path-node-16721 even"><a tabindex="0">Seminars and Events</a><ul><li class="menu-857996 menu-path-node-16721 first odd"><a href="/seminars-events" tabindex="0">Seminars and Events Overview</a></li> <li class="menu-858006 menu-path-node-16586 even"><a href="/seminars-events/bbc-seminar-series" tabindex="0">BBC Seminar Series</a></li> <li class="menu-858016 menu-path-node-16596 odd"><a href="/seminars-events/bp223" tabindex="0">BP 223: Research in Progress Talks</a></li> <li class="menu-858001 menu-path-node-16581 even"><a href="/seminars-events/bp-ccb-pizza-talks" tabindex="0">BP/CCB Pizza Talks</a></li> <li class="menu-957086 menuparent menu-path-node-18811 odd"><a tabindex="0">Outreach Activities</a><ul><li class="menu-957086 menu-path-node-18811 first odd"><a href="/seminars-events/outreach-activities" tabindex="0">Outreach Activities Overview</a></li> <li class="menu-962446 menu-path-node-22276 even last"><a href="/seminars-events/outreach-activities/community-college-outreach" tabindex="0">Community College Outreach Program</a></li> </ul></li> <li class="menu-962451 menu-path-node-22271 even"><a href="/seminars-events/retreat" tabindex="0">QBC Graduate Program Retreat</a></li> <li class="menu-858011 menu-path-node-16601 odd"><a href="/seminars-events/qbc-journal-club" tabindex="0">QBC Journal Club</a></li> <li class="menu-961331 menu-path-node-19846 even"><a href="/seminars-events/student-run-initiatives" tabindex="0">Student-Run Initiatives</a></li> <li class="menu-857756 menu-path-node-16651 odd last"><a href="/seminars-events/undergrad-summer-internships" tabindex="0">Undergraduate Summer Internships</a></li> </ul></li> <li class="menu-854606 menu-path-news odd"><a href="/news" tabindex="0">News</a></li> <li class="menu-837141 menu-path-node-15826 even"><a href="/tour" tabindex="0">Mission Bay Tour</a></li> <li class="menu-837121 menu-path-node-15831 odd last"><a href="/support" tabindex="0">Support the Program</a></li> </ul> </div> </div> <div id="content" class="lab-title-group"> <h1 class="lab-title">PhD Degree Program in Biophysics (BP)</h1> <div class="subtitle"><span>Preparing scientists to reveal the intricacies of living systems through the application of physics and chemistry</span></div> </div> </div> <!-- /header --> <div id="main" class="clearfix" role="main"> <div> <div id="content-inner" class="inner column center"> <div id="content-header"> <div class="tabs"></div> </div> <!-- /#content-header --> <div id="content-area" class="clearfix"> <div class="region region-content"> <div id="block-system-main" class="block block-system"> <div class="content"> <div id="node-16026" class="node node-page clearfix" about="/" 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/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/biophysics_-_thor_swift_photography_0.jpg?itok=e-Hz6fjN 3x" media="(min-width: 0px) and (max-width: 430px)" /> <!--[if IE 9]></video><![endif]--> <!--[if lt IE 9]> <img src="/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/biophysics_-_thor_swift_photography_0.jpg?itok=e-Hz6fjN" alt="Woman handling scientific equipment." title="" /> <![endif]--> <!--[if !lt IE 9]><!--> <img srcset="" alt="Woman handling scientific equipment." title="" /> <!-- <![endif]--> </picture></li> <li data-thumb="https://biophysics.ucsf.edu/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/genentech-hall_0.jpg?itok=wkpB5k2o"><picture > <!--[if IE 9]><video style="display: none;"><![endif]--> <source srcset="/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/genentech-hall_0.jpg?itok=wkpB5k2o 1x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop_2x/public/genentech-hall_0.jpg?itok=YMJoXFNc 2x, 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/sites/pharm.ucsf.edu/files/styles/slideshow_crop_1_5x/public/rutter-center_0.jpg?itok=a2SCMPoC 2x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop_3x/public/rutter-center_0.jpg?itok=xfseQ_rz 3x" media="(min-width: 801px) and (max-width: 1024px)" /> <source srcset="/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 1x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop_1_5x/public/rutter-center_0.jpg?itok=a2SCMPoC 2x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop_2x/public/rutter-center_0.jpg?itok=d6Io1zLj 3x" media="(min-width: 641px) and (max-width: 800px)" /> <source srcset="/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 1x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 2x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop_1_5x/public/rutter-center_0.jpg?itok=a2SCMPoC 3x" media="(min-width: 481px) and (max-width: 640px)" /> <source srcset="/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 1x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 2x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop_1_5x/public/rutter-center_0.jpg?itok=a2SCMPoC 3x" media="(min-width: 431px) and (max-width: 480px)" /> <source srcset="/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 1x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 2x, /sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE 3x" media="(min-width: 0px) and (max-width: 430px)" /> <!--[if IE 9]></video><![endif]--> <!--[if lt IE 9]> <img src="/sites/pharm.ucsf.edu/files/styles/slideshow_crop/public/rutter-center_0.jpg?itok=6zrV3PUE" alt="Dramatic night sky with modern buildings lit behind a grove." title="" /> <![endif]--> <!--[if !lt IE 9]><!--> <img srcset="" alt="Dramatic night sky with modern buildings lit behind a grove." title="" /> <!-- <![endif]--> </picture></li> </ul></div> </div></div></div><div class="field field-name-body field-type-text-with-summary field-label-hidden"><div class="field-items"><div class="field-item even" property="content:encoded"><h2>About the program</h2> <p>The Biophysics Graduate Program (BP) at the University of California, San Francisco (UCSF) prepares scientists to leverage new and emerging research tools—from nanotechnology to nuclear magnetic resonance spectroscopy—to quantify biological processes in living biological systems using physics and chemistry. The program is set in an atmosphere of intense research collaboration and within a university known for its research accomplishments and interdisciplinary approach to discovery and education.</p> <p>BP welcomes scientists of any race, religion, national origin, gender identity, caregiver and family commitments, political affiliation, sexual orientation, and eligible age or ability. We believe Black Lives Matter and are committed to sustained action to reduce racism and inequity in science.</p> <h2>Application window</h2> <table> <tbody> <tr> <th scope="row"> <p>Opens</p> </th> <td> <p>September 1, 2024</p> </td> </tr> <tr> <th scope="row"> <p>Closes</p> </th> <td> <p>December 1, 2024</p> </td> </tr> <tr> <th scope="row"> <p>How to apply</p> </th> <td> <p>Apply through the Molecular Cellular and Computational Biophysics application</p> </td> </tr> </tbody> </table> <p><a href="https://gradapp.ucsf.edu/register/apply"><div style="max-width: 284px;" class="media media-element-container media-media-full _none"><picture class="media-element file-media-full"> <!--[if IE 9]><video style="display: none;"><![endif]--> <source srcset="/sites/pharm.ucsf.edu/files/styles/lab_full/public/apply-bp.png?itok=ywXI2sRg&timestamp=1583194049 1x" media="(min-width: 1025px)"></source> <source srcset="/sites/pharm.ucsf.edu/files/styles/lab_three_quarters/public/apply-bp.png?itok=x9AeabCn&timestamp=1583194049 1x" media="(min-width: 801px) and (max-width: 1024px)"></source> <source srcset="/sites/pharm.ucsf.edu/files/styles/lab_full/public/apply-bp.png?itok=ywXI2sRg&timestamp=1583194049 1x" media="(min-width: 641px) and (max-width: 800px)"></source> <source srcset="/sites/pharm.ucsf.edu/files/styles/lab_three_quarters/public/apply-bp.png?itok=x9AeabCn&timestamp=1583194049 1x" media="(min-width: 481px) and (max-width: 640px)"></source> <source srcset="/sites/pharm.ucsf.edu/files/styles/lab_half/public/apply-bp.png?itok=auETmCIl&timestamp=1583194049 1x" media="(min-width: 431px) and (max-width: 480px)"></source> <source srcset="/sites/pharm.ucsf.edu/files/styles/lab_half/public/apply-bp.png?itok=auETmCIl&timestamp=1583194049 1x" media="(min-width: 0px) and (max-width: 430px)"></source> <!--[if IE 9]></video><![endif]--> <!--[if lt IE 9]> <img class="media-element file-media-full" src="/sites/pharm.ucsf.edu/files/styles/lab_full/public/apply-bp.png?itok=ywXI2sRg&timestamp=1583194049" alt="Apply" title="" /> <![endif]--> <!--[if !lt IE 9]><!--> <img class="media-element file-media-full" srcset="" alt="Apply" title="" /> <!-- <![endif]--> </picture></div></a></p> <h2>BP is a member of</h2> <ul> <li><a href="https://qbc.ucsf.edu/">Quantitative Biosciences Consortium (QBC)</a></li> <li> <div><a href="https://pibs.ucsf.edu/">Program in Biological Sciences (PIBS)</a></div> </li> </ul> <h2>Optional designated emphasis in</h2> <p><a href="https://pharm.ucsf.edu/cbs">Complex Biological Systems (CBS)</a></p> <hr /> <p class="fine-print">Image credits: © Majed and Thor Swift Photography</p></div></div></div> </div> </div> </div> </div> <div id="block-views-news-news-block" class="block block-views"> <h2>News</h2> <div class="content"> <div class="view view-news view-id-news view-display-id-news_block view-dom-id-27ace2d587c290fdb7035d2c312c3075"> <div 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class="field-content"><a href="https://pharmacy.ucsf.edu/news/2024/04/school-pharmacy-shows-out-grad-slam-storytelling-competition"><img typeof="foaf:Image" src="/sites/pharm.ucsf.edu/files/styles/lab_square_thumbnail/public/featured/grad-slam-2024-reuben-hogan.jpg?itok=5DXpyZBB" alt="Ruben Hogan speaks in front of graphics on screen." title="Noah Berger" /></a></div> </div> <div class="views-field views-field-title"> <span class="field-content"><a href="https://pharmacy.ucsf.edu/news/2024/04/school-pharmacy-shows-out-grad-slam-storytelling-competition">School of Pharmacy shows out in Grad Slam storytelling competition</a></span> </div> </div> <div class="views-row views-row-3 views-row-odd views-row-last"> <div class="views-field views-field-field-featured-image views-field-field-image"> <div class="field-content"><a href="https://pharmacy.ucsf.edu/news/2023/09/qbi-director-krogans-covid-infectious-disease-efforts-continue-make-waves"><img typeof="foaf:Image" src="/sites/pharm.ucsf.edu/files/styles/lab_square_thumbnail/public/featured/national_health_research_forum_2023_-_dihprize_presentation_p1010262.jpg?itok=XnQfeeqR" alt="Nevan Krogan receives award." /></a></div> </div> <div class="views-field views-field-title"> <span class="field-content"><a href="https://pharmacy.ucsf.edu/news/2023/09/qbi-director-krogans-covid-infectious-disease-efforts-continue-make-waves">QBI and Director Krogan’s COVID and infectious disease efforts continue to make waves</a></span> </div> </div> </div> <div class="more-link"> <a href="/news"> See all </a> </div> </div> </div> </div> </div> </div> <div id="sidebar-first"> <div class="region region-sidebar-first"> <div id="block-domain-menu-block-main-menu" class="block block-domain-menu-block"> <div class="content"> <div class="skip primary"> <a href="#postnav">Skip primary navigation</a> </div> <div class="menu-block-wrapper menu-block-main-menu menu-name-95ab1d4d78cce734039268e7f3b0e0db parent-mlid-0 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