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class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/71/Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg/220px-Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg" decoding="async" width="220" height="301" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/71/Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg/330px-Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/71/Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg/440px-Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg 2x" data-file-width="3418" data-file-height="4684" /></a><figcaption>Yoshinori Ōsumi (2016)</figcaption></figure> <p><b>Yoshinori Ōsumi</b> (<a href="/wiki/Japanische_Schrift" title="Japanische Schrift">japanisch</a> <span lang="ja-Jpan" class="Jpan">大隅良典</span>, <i>Ōsumi Yoshinori</i>; * <a href="/wiki/9._Februar" title="9. Februar">9. Februar</a> <a href="/wiki/1945" title="1945">1945</a> in <a href="/wiki/Fukuoka" title="Fukuoka">Fukuoka</a>) ist ein japanischer <a href="/wiki/Zellbiologie" title="Zellbiologie">Zellbiologe</a>. Er ist bekannt für Forschungen und Entdeckungen zur <a href="/wiki/Autophagozytose" title="Autophagozytose">Autophagozytose</a> (Autophagie). 2016 erhielt er dafür den <a href="/wiki/Nobelpreis_f%C3%BCr_Physiologie_oder_Medizin" title="Nobelpreis für Physiologie oder Medizin">Nobelpreis für Physiologie oder Medizin</a>.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div id="toc" class="toc" role="navigation" aria-labelledby="mw-toc-heading"><input type="checkbox" role="button" id="toctogglecheckbox" class="toctogglecheckbox" style="display:none" /><div class="toctitle" lang="de" dir="ltr"><h2 id="mw-toc-heading">Inhaltsverzeichnis</h2><span class="toctogglespan"><label class="toctogglelabel" for="toctogglecheckbox"></label></span></div> <ul> <li class="toclevel-1 tocsection-1"><a href="#Leben"><span class="tocnumber">1</span> <span class="toctext">Leben</span></a></li> <li class="toclevel-1 tocsection-2"><a href="#Werk"><span class="tocnumber">2</span> <span class="toctext">Werk</span></a></li> <li class="toclevel-1 tocsection-3"><a href="#Auszeichnungen_(Auswahl)"><span class="tocnumber">3</span> <span class="toctext">Auszeichnungen (Auswahl)</span></a></li> <li class="toclevel-1 tocsection-4"><a href="#Schriften_(Auswahl)"><span class="tocnumber">4</span> <span class="toctext">Schriften (Auswahl)</span></a></li> <li class="toclevel-1 tocsection-5"><a href="#Weblinks"><span class="tocnumber">5</span> <span class="toctext">Weblinks</span></a></li> <li class="toclevel-1 tocsection-6"><a href="#Einzelnachweise"><span class="tocnumber">6</span> <span class="toctext">Einzelnachweise</span></a></li> </ul> </div> <div class="mw-heading mw-heading2"><h2 id="Leben">Leben</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&veaction=edit&section=1" title="Abschnitt bearbeiten: Leben" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&action=edit&section=1" title="Quellcode des Abschnitts bearbeiten: Leben"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Ōsumi wurde 1945 in Fukuoka geboren. Sein Vater war Professor an der <a href="/wiki/Universit%C3%A4t_Ky%C5%ABsh%C5%AB" title="Universität Kyūshū">Universität Kyūshū</a> und Ingenieur. Ōsumi erhielt 1967 den <a href="/wiki/Bachelor" title="Bachelor">Bachelor</a>-Abschluss an der <a href="/wiki/Universit%C3%A4t_Tokio" title="Universität Tokio">Universität Tokio</a> und wurde dort 1974 bei Kazutomi Imahori <a href="/wiki/Promotion_(Doktor)" title="Promotion (Doktor)">promoviert</a>. Während des Studiums wechselte er von der Chemie zur damals im Aufbruch befindlichen <a href="/wiki/Molekularbiologie" title="Molekularbiologie">Molekularbiologie</a>. Bis 1977 war er als <a href="/wiki/Postdoktorand" class="mw-redirect" title="Postdoktorand">Postdoktorand</a> an der <a href="/wiki/Rockefeller_University" title="Rockefeller University">Rockefeller University</a> bei <a href="/wiki/Gerald_Edelman" class="mw-redirect" title="Gerald Edelman">Gerald Edelman</a>, wo seine Beschäftigung mit Weizenzellen begann (DNA-Duplizierung), und danach forschte er an der Universität Tokio (bei Yasuhiro Anraku), an der er 1986 Dozent (<a href="/wiki/Lecturer" title="Lecturer">Lecturer</a>) und 1988 Assistenzprofessor (Associate Professor) wurde. Zu der Zeit befasste er sich schon mit Membrantransport in <a href="/wiki/Vakuole" title="Vakuole">Vakuolen</a> in der Zelle, die Teil des Systems der Autophagie sind. 1996 ging er als Professor an das <a href="/wiki/National_Institute_for_Basic_Biology" class="mw-redirect" title="National Institute for Basic Biology">National Institute for Basic Biology</a> in <a href="/wiki/Okazaki" title="Okazaki">Okazaki</a>. Außerdem war er 2004 bis 2009 Professor an der <a href="/wiki/Graduate_University_for_Advanced_Studies" title="Graduate University for Advanced Studies">Graduate University for Advanced Studies</a> in <a href="/wiki/Hayama_(Kanagawa)" title="Hayama (Kanagawa)">Hayama</a>. 2009 <a href="/wiki/Emeritierung" title="Emeritierung">emeritierte</a> er und war danach Professor an der <a href="/wiki/T%C5%8Dky%C5%8D_K%C5%8Dgy%C5%8D_Daigaku" title="Tōkyō Kōgyō Daigaku">Technischen Hochschule Tokio</a>.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&veaction=edit&section=2" title="Abschnitt bearbeiten: Werk" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&action=edit&section=2" title="Quellcode des Abschnitts bearbeiten: Werk"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/Datei:Yoshinori_Osumi_20161003.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Yoshinori_Osumi_20161003.jpg/220px-Yoshinori_Osumi_20161003.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Yoshinori_Osumi_20161003.jpg/330px-Yoshinori_Osumi_20161003.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Yoshinori_Osumi_20161003.jpg/440px-Yoshinori_Osumi_20161003.jpg 2x" data-file-width="480" data-file-height="360" /></a><figcaption>Yoshinori Ōsumi in einem Labor der Technischen Hochschule Tokio</figcaption></figure> <p>Er klärte die molekularen Mechanismen von Autophagie am Beispiel von <a href="/wiki/Hefen" title="Hefen">Hefezellen</a>, später auch bei Säugerzellen, die sich durch Abbau körpereigener <a href="/wiki/Proteine" class="mw-redirect" title="Proteine">Proteine</a> auf gewandelte Umweltbedingungen einstellen. Dabei identifizierte er mit seiner Gruppe die beteiligten Enzyme, deren Regulierung je nach Stoffwechselzustand und den Mechanismus der Bildung von Autophagosomen. In der Hefezelle bilden sich bei Stickstoffmangel Autophagosomen (Membrane, die die abzubauenden Proteine umschließen und an Vakuolen oder <a href="/wiki/Lysosom" title="Lysosom">Lysosomen</a> abgeben), es bilden sich <a href="/wiki/Vesikel_(Biologie)" title="Vesikel (Biologie)">Vesikel</a> in den Vakuolen und Vakuolen fusionieren. Ōsumi untersuchte verschiedene genetische Varianten von Hefezellen, denen für diese Prozesse des Zellabbaus wichtige <a href="/wiki/Proteinase" class="mw-redirect" title="Proteinase">Proteinasen</a> in den Vakuolen fehlten. 1991 fand seine Gruppe einen ersten Autophagie-defekten Mutanten (apg 1-1, später ATG 1 genannt) in Hefe und später fanden sie 13 weitere (ATG). Mit der bald darauf erfolgten Entschlüsselung des Erbguts der Hefezelle konnten die entsprechenden Gene geklont werden. ATG 1 entsprach einer <a href="/wiki/Proteinkinase" class="mw-redirect" title="Proteinkinase">Proteinkinase</a>, die anderen hatten kompliziertere Funktionen und entsprachen regulatorischen <a href="/wiki/Ubiquitin" title="Ubiquitin">Ubiquitin</a>-ähnlichen Enzymen. </p> <div class="mw-heading mw-heading2"><h2 id="Auszeichnungen_(Auswahl)"><span id="Auszeichnungen_.28Auswahl.29"></span>Auszeichnungen (Auswahl)</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&veaction=edit&section=3" title="Abschnitt bearbeiten: Auszeichnungen (Auswahl)" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&action=edit&section=3" title="Quellcode des Abschnitts bearbeiten: Auszeichnungen (Auswahl)"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>2005 erhielt er den Fujihara Award, 2006 den Preis der Japan Academy, 2008 den <a href="/wiki/Asahi-Preis" title="Asahi-Preis">Asahi-Preis</a>, 2012 den <a href="/wiki/Kyoto-Preis" title="Kyoto-Preis">Kyoto-Preis</a> und 2015 den <a href="/wiki/Canada_Gairdner_International_Award" title="Canada Gairdner International Award">Canada Gairdner International Award</a>. 2015 erhielt er den <a href="/wiki/Internationaler_Preis_f%C3%BCr_Biologie" title="Internationaler Preis für Biologie">Internationalen Preis für Biologie</a>, den <a href="/wiki/Keio_Medical_Science_Prize" title="Keio Medical Science Prize">Keio Medical Science Prize</a> und den <a href="/wiki/Rosenstiel_Award" title="Rosenstiel Award">Rosenstiel Award</a>. 2016 erhielt er den <a href="/wiki/Wiley_Prize_in_Biomedical_Sciences" title="Wiley Prize in Biomedical Sciences">Wiley Prize in Biomedical Sciences</a>, den <a href="/wiki/Dr._Paul_Janssen_Award_for_Biomedical_Research" title="Dr. Paul Janssen Award for Biomedical Research">Dr. Paul Janssen Award for Biomedical Research</a> und den <a href="/wiki/Nobelpreis_f%C3%BCr_Physiologie_oder_Medizin" title="Nobelpreis für Physiologie oder Medizin">Nobelpreis für Physiologie oder Medizin</a>. Bereits seit 2013 hatte ihn <a href="/wiki/Thomson_Reuters" title="Thomson Reuters">Thomson Reuters</a> aufgrund der Zahl seiner <a href="/wiki/Zitat" title="Zitat">Zitationen</a> zu den Favoriten auf einen Nobelpreis gehandelt (<a href="/wiki/Thomson_Reuters_Citation_Laureates" class="mw-redirect" title="Thomson Reuters Citation Laureates">Thomson Reuters Citation Laureates</a>).<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> 2016 wurde er auch mit dem japanischen <a href="/wiki/Kulturorden_(Japan)" title="Kulturorden (Japan)">Kulturorden</a> ausgezeichnet. Für 2017 wurde ihm für die Aufklärung des Mechanismus der Autophagie der <a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Breakthrough Prize in Life Sciences</a> zugesprochen, der mit drei Millionen US-Dollar dotiert ist. 2017 wurde er zum Fellow der <a href="/wiki/American_Association_for_the_Advancement_of_Science" title="American Association for the Advancement of Science">American Association for the Advancement of Science</a> gewählt.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Schriften_(Auswahl)"><span id="Schriften_.28Auswahl.29"></span>Schriften (Auswahl)</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&veaction=edit&section=4" title="Abschnitt bearbeiten: Schriften (Auswahl)" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&action=edit&section=4" title="Quellcode des Abschnitts bearbeiten: Schriften (Auswahl)"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>mit Tsukada: <i>Isolation and characterization of autophagy-defective mutants of Saccharomyces cerevisiae</i>. In: <i><a href="/wiki/FEBS_Letters" title="FEBS Letters">FEBS Letters</a>.</i> Band 333, 1993, S. 169–174, <a rel="nofollow" class="external text" href="http://www.sciencedirect.com/science/article/pii/001457939380398E">Abstract</a></li> <li>mit K. Takeshige, M. Baba, S. Tsuboi, T. Noda: <i>Autophagy in yeast demonstrated with proteinase-deficient mutants and conditions for its induction</i>. In: <i><a href="/wiki/Journal_of_Cell_Biology" title="Journal of Cell Biology">Journal of Cell Biology</a>.</i> Band 119, 1992, S. 301–311, <a rel="nofollow" class="external text" href="http://jcb.rupress.org/content/119/2/301.abstract?ijkey=30b8f86312f81144777be1eb4009c5d8b1726ae7&keytype2=tf_ipsecsha">Abstract</a></li> <li>mit Takeshige, M. Baba, N. Baba: <i>Ultrastructural analysis of the autophagic process in yeast: detection of autophagosomes and their characterization</i>. In: <i><a href="/wiki/Journal_of_Cell_Biology" title="Journal of Cell Biology">Journal of Cell Biology</a>.</i> Band 124, 1994, S. 903–913, <a rel="nofollow" class="external text" href="http://jcb.rupress.org/content/124/6/903.abstract?ijkey=393ed7ef6e9da0c321e574b67bede9d9ec8a1826&keytype2=tf_ipsecsha">Abstract</a></li> <li>mit Noboru Mizushima u. a.: <i>A protein conjugation system essential for autophagy</i>. In: <i><a href="/wiki/Nature" title="Nature">Nature</a>.</i> Band 395, 1998, S. 395, <a rel="nofollow" class="external text" href="http://www.nature.com/nature/journal/v395/n6700/full/395395a0.html">Abstract</a></li> <li>mit Kabeya u. a.: <i>LC3, a mammalian homologue of yeast Apg8p, is localized in autophagosome membranes after processing</i>. In: <i>EMBO Journal.</i> Band 19, 2000, S. 5270–5278, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC305793/">PMC 305793</a> (freier Volltext)</li> <li>mit K. Suzuki u. a.: <i>The Pre-Autophagosomal Structure Organized by Concerted Functions of APG Genes Is Essential for Autophagosome Formation</i>. In: <i><a href="/wiki/The_EMBO_Journal" title="The EMBO Journal">The EMBO Journal</a>.</i> Band 20, 2001, S. 5971–5981, <a href="//doi.org/10.1093/emboj/20.21.5971" class="extiw" title="doi:10.1093/emboj/20.21.5971">doi:10.1093/emboj/20.21.5971</a>.</li> <li>mit Mizushima, Yoshimori: <i>Autophagosome formation in mammalian cells</i>. In: <i>Cell Structure and Function.</i> Band 27, 2002, S. 421–429, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/12576635?dopt=Abstract">PMID 12576635</a></li> <li>mit Nakatogawa, Ichimura: <i>Atg8, a Ubiquitin-like Protein Required for Autophagosome Formation, Mediates Membrane Tethering and Hemifusion</i>. In: <i><a href="/wiki/Cell_(Zeitschrift)" title="Cell (Zeitschrift)">Cell</a>.</i> Band 130, 2007, S. 165–178, <a href="//doi.org/10.1016/j.cell.2007.05.021" class="extiw" title="doi:10.1016/j.cell.2007.05.021">doi:10.1016/j.cell.2007.05.021</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="Weblinks">Weblinks</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&veaction=edit&section=5" title="Abschnitt bearbeiten: Weblinks" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&action=edit&section=5" title="Quellcode des Abschnitts bearbeiten: Weblinks"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="sisterproject" style="margin:0.1em 0 0 0;"><div class="noresize noviewer" style="display:inline-block; line-height:10px; min-width:1.6em; text-align:center;" aria-hidden="true" role="presentation"><span class="mw-default-size" typeof="mw:File"><span title="Commons"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div><b><span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Category:Yoshinori_%C5%8Csumi?uselang=de"><span lang="en">Commons</span>: Yoshinori Ōsumi</a></span></b> – Sammlung von Bildern, Videos und Audiodateien</div> <ul><li><a rel="nofollow" class="external text" href="http://www.inamori-f.or.jp/laureates/k28_b_yoshinori/prf_e.html">Imamori Foundation</a></li> <li><a rel="nofollow" class="external text" href="http://jcb.rupress.org/content/197/2/164.full"><i>Yoshinori Ohsumi: Autophagy from beginning to end</i>.</a> In: <i>Journal of Cell Biology</i>, 2012; Interview</li></ul> <div class="mw-heading mw-heading2"><h2 id="Einzelnachweise">Einzelnachweise</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&veaction=edit&section=6" title="Abschnitt bearbeiten: Einzelnachweise" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Yoshinori_%C5%8Csumi&action=edit&section=6" title="Quellcode des Abschnitts bearbeiten: Einzelnachweise"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.nobelprize.org/nobel_prizes/medicine/laureates/2016/press.html">Pressemeldung</a> bei nobelprize.org, 3. Oktober 2016.</span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><span class="cite">SPIEGEL ONLINE, Hamburg Germany: <a rel="nofollow" class="external text" href="http://www.spiegel.de/gesundheit/diagnose/nobelpreis-medizin-nobelpreis-geht-an-den-japaner-yoshinori-ohsumi-a-1114972.html"><i>Auszeichnung 2016: Medizinnobelpreis geht an den Japaner Yoshinori Ohsumi.</i></a> In: <i>SPIEGEL ONLINE.</i><span class="Abrufdatum"> Abgerufen am 3. Oktober 2016</span>.</span><span style="display: none;" class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adc&rfr_id=info%3Asid%2Fde.wikipedia.org%3AYoshinori+%C5%8Csumi&rft.title=Auszeichnung+2016%3A+Medizinnobelpreis+geht+an+den+Japaner+Yoshinori+Ohsumi&rft.description=Auszeichnung+2016%3A+Medizinnobelpreis+geht+an+den+Japaner+Yoshinori+Ohsumi&rft.identifier=http%3A%2F%2Fwww.spiegel.de%2Fgesundheit%2Fdiagnose%2Fnobelpreis-medizin-nobelpreis-geht-an-den-japaner-yoshinori-ohsumi-a-1114972.html&rft.creator=SPIEGEL+ONLINE%2C+Hamburg+Germany"> </span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"> <a rel="nofollow" class="external text" href="http://sciencewatch.com/nobel/2013-predictions">2013 Predictions</a> bei <a href="/wiki/Thomson_Reuters" title="Thomson Reuters">Thomson Reuters</a> (sciencewatch.com); abgerufen am 25. September 2013</span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><span class="cite"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20180130205436/https://www.aaas.org/fellow/ohsumi-yoshinori"><i>Fellows der AAAS: Yoshinori Ohsumi.</i></a> American Association for the Advancement of Science, archiviert vom <style data-mw-deduplicate="TemplateStyles:r235239667">.mw-parser-output .dewiki-iconexternal>a{background-position:center right;background-repeat:no-repeat}body.skin-minerva .mw-parser-output .dewiki-iconexternal>a{background-image:url("https://upload.wikimedia.org/wikipedia/commons/a/a4/OOjs_UI_icon_external-link-ltr-progressive.svg")!important;background-size:10px;padding-right:13px!important}body.skin-timeless .mw-parser-output .dewiki-iconexternal>a,body.skin-monobook .mw-parser-output .dewiki-iconexternal>a{background-image:url("https://upload.wikimedia.org/wikipedia/commons/3/30/MediaWiki_external_link_icon.svg")!important;padding-right:13px!important}body.skin-vector .mw-parser-output .dewiki-iconexternal>a{background-image:url("https://upload.wikimedia.org/wikipedia/commons/9/96/Link-external-small-ltr-progressive.svg")!important;background-size:0.857em;padding-right:1em!important}</style><span class="dewiki-iconexternal"><a class="external text" href="https://redirecter.toolforge.org/?url=https%3A%2F%2Fwww.aaas.org%2Ffellow%2Fohsumi-yoshinori">Original</a></span> (nicht mehr online verfügbar) am <span style="white-space:nowrap;">30. Januar 2018</span><span>;</span><span class="Abrufdatum"> abgerufen am 30. Januar 2018</span>.</span> <small class="archiv-bot"><span class="wp_boppel noviewer" aria-hidden="true" role="presentation"><span typeof="mw:File"><span title="i"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Pictogram_voting_info.svg/15px-Pictogram_voting_info.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Pictogram_voting_info.svg/23px-Pictogram_voting_info.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Pictogram_voting_info.svg/30px-Pictogram_voting_info.svg.png 2x" data-file-width="250" data-file-height="250" /></span></span></span> <b>Info:</b> Der Archivlink wurde automatisch eingesetzt und noch nicht geprüft. 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div.NavHead{background-color:var(--dewiki-hintergrundfarbe5);font-weight:bold}.mw-parser-output div.NavFrame:after{clear:both;content:"";display:block}.mw-parser-output div.NavFrame+div.NavFrame,.mw-parser-output div.NavFrame+link+div.NavFrame,.mw-parser-output div.NavFrame+style+div.NavFrame{margin-top:-1px}.mw-parser-output .NavToggle{float:right;font-size:x-small}@media screen{html.skin-theme-clientpref-night .mw-parser-output .NavPic span[typeof="mw:File"] img{background-color:#c8ccd1}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .NavPic span[typeof="mw:File"] img{background-color:#c8ccd1}}</style><div class="NavFrame navigation-not-searchable" role="navigation"> <div class="NavHead"><a href="/wiki/Liste_der_Nobelpreistr%C3%A4ger_f%C3%BCr_Physiologie_oder_Medizin" title="Liste der Nobelpreisträger für Physiologie oder Medizin">Träger des Nobelpreises für Physiologie oder Medizin</a></div> <div class="NavContent"> <p>1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Behring</a> | 1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> | 1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> | 1904: <a href="/wiki/Iwan_Petrowitsch_Pawlow" title="Iwan Petrowitsch Pawlow">Pawlow</a> | 1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> | 1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a>, <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Cajal</a> | 1907: <a href="/wiki/Alphonse_Laveran" title="Alphonse Laveran">Laveran</a> | 1908: <a href="/wiki/Ilja_Iljitsch_Metschnikow" title="Ilja Iljitsch Metschnikow">Metschnikow</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> | 1909: <a href="/wiki/Theodor_Kocher" title="Theodor Kocher">Kocher</a> | 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> | 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> | 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> | 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> | 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> | 1915–1918: <i>nicht verliehen</i> | 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> | 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> | 1921: <i>nicht verliehen</i> | 1922: <a href="/wiki/Archibald_Vivian_Hill" title="Archibald Vivian Hill">Hill</a>, <a href="/wiki/Otto_Meyerhof" title="Otto Meyerhof">Meyerhof</a> | 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_James_Rickard_Macleod" title="John James Rickard Macleod">Macleod</a> | 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> | 1925: <i>nicht verliehen</i> | 1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Fibiger</a> | 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> | 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> | 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> | 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> | 1931: <a href="/wiki/Otto_Warburg_(Biochemiker)" title="Otto Warburg (Biochemiker)">Warburg</a> | 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Adrian,_1._Baron_Adrian" title="Edgar Adrian, 1. Baron Adrian">Adrian</a> | 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> | 1934: <a href="/wiki/George_Hoyt_Whipple" title="George Hoyt Whipple">Whipple</a>, <a href="/wiki/George_Richards_Minot" title="George Richards Minot">Minot</a>, <a href="/wiki/William_Parry_Murphy" title="William Parry Murphy">Murphy</a> | 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> | 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> | 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> | 1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Heymans</a> | 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> | 1940–1942: <i>nicht verliehen</i> | 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> | 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> | 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> | 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> | 1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a>, <a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a>, <a href="/wiki/Bernardo_Alberto_Houssay" title="Bernardo Alberto Houssay">Houssay</a> | 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> | 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> | 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> | 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> | 1952: <a href="/wiki/Selman_Abraham_Waksman" title="Selman Abraham Waksman">Waksman</a> | 1953: <a href="/wiki/Hans_Adolf_Krebs" title="Hans Adolf Krebs">H. A. Krebs</a>, <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> | 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> | 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> | 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_For%C3%9Fmann" title="Werner Forßmann">Forßmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> | 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> | 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> | 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> | 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> | 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> | 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> | 1963: <a href="/wiki/John_Carew_Eccles" title="John Carew Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Fielding_Huxley" title="Andrew Fielding Huxley">Huxley</a> | 1964: <a href="/wiki/Konrad_Bloch" title="Konrad Bloch">Bloch</a>, <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Lynen</a> | 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Lwoff" title="André Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod_(Biologe)" title="Jacques Monod (Biologe)">Monod</a> | 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> | 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> | 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> | 1969: <a href="/wiki/Max_Delbr%C3%BCck_(Biophysiker)" title="Max Delbrück (Biophysiker)">Delbrück</a>, <a href="/wiki/Alfred_Day_Hershey" title="Alfred Day Hershey">Hershey</a>, <a href="/wiki/Salvador_Edward_Luria" title="Salvador Edward Luria">Luria</a> | 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> | 1971: <a href="/wiki/Earl_Wilbur_Sutherland" title="Earl Wilbur Sutherland">Sutherland</a> | 1972: <a href="/wiki/Gerald_M._Edelman" title="Gerald M. Edelman">Edelman</a>, <a href="/wiki/Rodney_R._Porter" title="Rodney R. Porter">Porter</a> | 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Tinbergen</a> | 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">de Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> | 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_M._Temin" title="Howard M. Temin">Temin</a> | 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> | 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Victor_Schally" title="Andrew Victor Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> | 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> | 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> | 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> | 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_N._Wiesel" title="Torsten N. Wiesel">Wiesel</a> | 1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> | 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> | 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> | 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Goldstein</a> | 1986: <a href="/wiki/Stanley_Cohen_(Biochemiker)" title="Stanley Cohen (Biochemiker)">Cohen</a>, <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a> | 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> | 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Belle_Elion" title="Gertrude Belle Elion">Elion</a>, <a href="/wiki/George_Herbert_Hitchings" title="George Herbert Hitchings">Hitchings</a> | 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> | 1990: <a href="/wiki/Joseph_Edward_Murray" title="Joseph Edward Murray">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> | 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> | 1992: <a href="/wiki/Edmond_Henri_Fischer" title="Edmond Henri Fischer">Fischer</a>, <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">E. G. Krebs</a> | 1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> | 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> | 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a>, <a href="/wiki/Eric_Wieschaus" title="Eric Wieschaus">Wieschaus</a> | 1996: <a href="/wiki/Peter_Doherty_(Mediziner)" title="Peter Doherty (Mediziner)">Doherty</a>, <a href="/wiki/Rolf_Zinkernagel" title="Rolf Zinkernagel">Zinkernagel</a> | 1997: <a href="/wiki/Stanley_Prusiner" title="Stanley Prusiner">Prusiner</a> | 1998: <a href="/wiki/Robert_Francis_Furchgott" title="Robert Francis Furchgott">Furchgott</a>, <a href="/wiki/Louis_J._Ignarro" title="Louis J. Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> | 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> | 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> | 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> | 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_E._Sulston" title="John E. Sulston">Sulston</a> | 2003: <a href="/wiki/Paul_Christian_Lauterbur" title="Paul Christian Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> | 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> | 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> | 2006: <a href="/wiki/Andrew_Z._Fire" title="Andrew Z. Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> | 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a> | 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">zur Hausen</a>, <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré-Sinoussi</a>, <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> | 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> | 2010: <a href="/wiki/Robert_Edwards_(Genetiker)" title="Robert Edwards (Genetiker)">Edwards</a> | 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Steinman</a> | 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shin%E2%80%99ya_Yamanaka" title="Shin’ya Yamanaka">Yamanaka</a> | 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> | 2014: <a href="/wiki/John_O%E2%80%99Keefe_(Neurowissenschaftler)" title="John O’Keefe (Neurowissenschaftler)">O’Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">M. Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">E. Moser</a> | 2015: <a href="/wiki/William_C._Campbell" title="William C. Campbell">Campbell</a>, <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Ōmura</a>, <a href="/wiki/Tu_Youyou" title="Tu Youyou">Youyou</a> | 2016: <a class="mw-selflink selflink">Ōsumi</a> | 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> | 2018: <a href="/wiki/James_P._Allison" title="James P. Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> | 2019: <a href="/wiki/William_G._Kaelin" title="William G. Kaelin">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> | 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> | 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> | 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> | 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> | 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div> </div> <div class="hintergrundfarbe1 rahmenfarbe1 navigation-not-searchable normdaten-typ-p" style="border-style: solid; border-width: 1px; clear: left; margin-bottom:1em; margin-top:1em; padding: 0.25em; overflow: hidden; word-break: break-word; word-wrap: break-word;" id="normdaten"> <div style="display: table-cell; vertical-align: middle; width: 100%;"> <div> Normdaten (Person): <a href="/wiki/Gemeinsame_Normdatei" title="Gemeinsame Normdatei">GND</a>: <span class="plainlinks-print"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/1089644000">1089644000</a></span> <span class="noprint">(<a rel="nofollow" class="external text" href="https://lobid.org/gnd/1089644000">lobid</a>, <a rel="nofollow" class="external text" href="https://swb.bsz-bw.de/DB=2.104/SET=1/TTL=1/CMD?retrace=0&trm_old=&ACT=SRCHA&IKT=2999&SRT=RLV&TRM=1089644000">OGND</a><span class="metadata">, <a rel="nofollow" class="external text" href="https://prometheus.lmu.de/gnd/1089644000">AKS</a></span>)</span> | <a href="/wiki/Library_of_Congress_Control_Number" title="Library of Congress Control Number">LCCN</a>: <span class="plainlinks-print"><a rel="nofollow" class="external text" href="https://lccn.loc.gov/n2008083589">n2008083589</a></span> | <a href="/wiki/Web_NDL_Authorities" title="Web NDL Authorities">NDL</a>: <span class="plainlinks-print"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00204204">00204204</a></span> | <a href="/wiki/Virtual_International_Authority_File" title="Virtual International Authority File">VIAF</a>: <span class="plainlinks-print"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/252902927/">252902927</a></span> | <span class="noprint"><a class="external text" href="https://persondata.toolforge.org/p/Yoshinori_%C5%8Csumi">Wikipedia-Personensuche</a></span><span class="metadata"></span></div> </div></div> <table class="metadata rahmenfarbe1" id="Vorlage_Personendaten" style="border-style: solid; margin-top: 20px;"> <tbody><tr> <th colspan="2"><a href="/wiki/Hilfe:Personendaten" title="Hilfe:Personendaten">Personendaten</a> </th></tr> <tr> <td style="color: #aaa;">NAME </td> <td style="font-weight: bold;">Ōsumi, Yoshinori </td></tr> <tr> <td style="color: #aaa;">ALTERNATIVNAMEN </td> <td>大隅良典 (japanisch) </td></tr> <tr> <td style="color: #aaa;">KURZBESCHREIBUNG </td> <td>japanischer Zellbiologe </td></tr> <tr> <td style="color: #aaa;">GEBURTSDATUM </td> <td>9. 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data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-as mw-list-item"><a href="https://as.wikipedia.org/wiki/%E0%A6%AF%E0%A6%BC%E0%A6%B6%E0%A7%8D%E0%A6%AC%E0%A6%BF%E0%A6%A8%E0%A7%B0%E0%A7%80_%E0%A6%85%E0%A6%B9%E0%A6%9A%E0%A7%81%E0%A6%AE%E0%A6%BF" title="য়শ্বিনৰী অহচুমি – Assamesisch" lang="as" hreflang="as" data-title="য়শ্বিনৰী অহচুমি" data-language-autonym="অসমীয়া" data-language-local-name="Assamesisch" class="interlanguage-link-target"><span>অসমীয়া</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Yo%C5%9Finori_Osumi" title="Yoşinori Osumi – Aserbaidschanisch" lang="az" hreflang="az" data-title="Yoşinori Osumi" data-language-autonym="Azərbaycanca" data-language-local-name="Aserbaidschanisch" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اوسومی – Südaserbaidschanisch" lang="azb" hreflang="azb" data-title="یوشینوری اوسومی" data-language-autonym="تۆرکجه" data-language-local-name="Südaserbaidschanisch" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%81%D1%81%D1%96%D0%BD%D0%BE%D1%80%D1%8B_%D0%90%D1%81%D1%83%D0%BC%D1%96" title="Ёсіноры Асумі – Belarussisch" lang="be" hreflang="be" data-title="Ёсіноры Асумі" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми – Bulgarisch" lang="bg" hreflang="bg" data-title="Йошинори Осуми" data-language-autonym="Български" data-language-local-name="Bulgarisch" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%87%E0%A6%AF%E0%A6%BC%E0%A7%8B%E0%A6%B6%E0%A6%BF%E0%A6%A8%E0%A7%8B%E0%A6%B0%E0%A6%BF_%E0%A6%93%E0%A6%B8%E0%A7%81%E0%A6%AE%E0%A6%BF" title="ইয়োশিনোরি ওসুমি – Bengalisch" lang="bn" hreflang="bn" data-title="ইয়োশিনোরি ওসুমি" data-language-autonym="বাংলা" data-language-local-name="Bengalisch" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Bosnisch" lang="bs" hreflang="bs" data-title="Yoshinori Ohsumi" data-language-autonym="Bosanski" data-language-local-name="Bosnisch" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Katalanisch" lang="ca" hreflang="ca" data-title="Yoshinori Ohsumi" data-language-autonym="Català" data-language-local-name="Katalanisch" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Jo%C5%A1inori_%C3%93sumi" title="Jošinori Ósumi – Tschechisch" lang="cs" hreflang="cs" data-title="Jošinori Ósumi" data-language-autonym="Čeština" data-language-local-name="Tschechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Dänisch" lang="da" hreflang="da" data-title="Yoshinori Ohsumi" data-language-autonym="Dansk" data-language-local-name="Dänisch" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%93%CE%B9%CE%BF%CF%83%CE%B9%CE%BD%CF%8C%CF%81%CE%B9_%CE%9F%CF%87%CF%83%CE%BF%CF%8D%CE%BC%CE%B9" title="Γιοσινόρι Οχσούμι – Griechisch" lang="el" hreflang="el" data-title="Γιοσινόρι Οχσούμι" data-language-autonym="Ελληνικά" data-language-local-name="Griechisch" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Englisch" lang="en" hreflang="en" data-title="Yoshinori Ohsumi" data-language-autonym="English" data-language-local-name="Englisch" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Spanisch" lang="es" hreflang="es" data-title="Yoshinori Ohsumi" data-language-autonym="Español" data-language-local-name="Spanisch" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Estnisch" lang="et" hreflang="et" data-title="Yoshinori Ōsumi" data-language-autonym="Eesti" data-language-local-name="Estnisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Baskisch" lang="eu" hreflang="eu" data-title="Yoshinori Ohsumi" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اسومی – Persisch" lang="fa" hreflang="fa" data-title="یوشینوری اسومی" data-language-autonym="فارسی" data-language-local-name="Persisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Finnisch" lang="fi" hreflang="fi" data-title="Yoshinori Ōsumi" data-language-autonym="Suomi" data-language-local-name="Finnisch" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Französisch" lang="fr" hreflang="fr" data-title="Yoshinori Ohsumi" data-language-autonym="Français" data-language-local-name="Französisch" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Galicisch" lang="gl" hreflang="gl" data-title="Yoshinori Ōsumi" data-language-autonym="Galego" data-language-local-name="Galicisch" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%99%D7%95%D7%A9%D7%99%D7%A0%D7%95%D7%A8%D7%99_%D7%90%D7%95%D7%A1%D7%95%D7%9E%D7%99" title="יושינורי אוסומי – Hebräisch" lang="he" hreflang="he" data-title="יושינורי אוסומי" data-language-autonym="עברית" data-language-local-name="Hebräisch" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/%C3%93szumi_Josinori" title="Ószumi Josinori – Ungarisch" lang="hu" hreflang="hu" data-title="Ószumi Josinori" data-language-autonym="Magyar" data-language-local-name="Ungarisch" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%85%D5%B8%D5%BD%D5%AB%D5%B6%D5%B8%D6%80%D5%AB_%D5%95%D5%BD%D5%B8%D6%82%D5%B4%D5%AB" title="Յոսինորի Օսումի – Armenisch" lang="hy" hreflang="hy" data-title="Յոսինորի Օսումի" data-language-autonym="Հայերեն" data-language-local-name="Armenisch" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Indonesisch" lang="id" hreflang="id" data-title="Yoshinori Ohsumi" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Ido" lang="io" hreflang="io" data-title="Yoshinori Ohsumi" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it badge-Q17437798 badge-goodarticle mw-list-item" title="lesenswerter Artikel"><a href="https://it.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Italienisch" lang="it" hreflang="it" data-title="Yoshinori Ōsumi" data-language-autonym="Italiano" data-language-local-name="Italienisch" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 – Japanisch" lang="ja" hreflang="ja" data-title="大隅良典" data-language-autonym="日本語" data-language-local-name="Japanisch" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми – Kasachisch" lang="kk" hreflang="kk" data-title="Йошинори Осуми" data-language-autonym="Қазақша" data-language-local-name="Kasachisch" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%AF%E0%B3%8A%E0%B2%B6%E0%B2%BF%E0%B2%A8%E0%B3%8B%E0%B2%B0%E0%B2%BF_%E0%B2%93%E0%B2%B6%E0%B3%81%E0%B2%AE%E0%B2%BF" title="ಯೊಶಿನೋರಿ ಓಶುಮಿ – Kannada" lang="kn" hreflang="kn" data-title="ಯೊಶಿನೋರಿ ಓಶುಮಿ" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%98%A4%EC%8A%A4%EB%AF%B8_%EC%9A%94%EC%8B%9C%EB%85%B8%EB%A6%AC" title="오스미 요시노리 – Koreanisch" lang="ko" hreflang="ko" data-title="오스미 요시노리" data-language-autonym="한국어" data-language-local-name="Koreanisch" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Latein" lang="la" hreflang="la" data-title="Yoshinori Ohsumi" data-language-autonym="Latina" data-language-local-name="Latein" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%88%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Јошинори Осуми – Mazedonisch" lang="mk" hreflang="mk" data-title="Јошинори Осуми" data-language-autonym="Македонски" data-language-local-name="Mazedonisch" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%AF%E0%B5%8B%E0%B4%B7%E0%B4%BF%E0%B4%A8%E0%B5%8B%E0%B4%B1%E0%B4%BF_%E0%B4%89%E0%B4%B7%E0%B5%81%E0%B4%AE%E0%B4%BF" title="യോഷിനോറി ഉഷുമി – Malayalam" lang="ml" hreflang="ml" data-title="യോഷിനോറി ഉഷുമി" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Malaiisch" lang="ms" hreflang="ms" data-title="Yoshinori Ohsumi" data-language-autonym="Bahasa Melayu" data-language-local-name="Malaiisch" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اسومی – Masanderanisch" lang="mzn" hreflang="mzn" data-title="یوشینوری اسومی" data-language-autonym="مازِرونی" data-language-local-name="Masanderanisch" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Niederdeutsch" lang="nds" hreflang="nds" data-title="Yoshinori Ōsumi" data-language-autonym="Plattdüütsch" data-language-local-name="Niederdeutsch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-ne mw-list-item"><a href="https://ne.wikipedia.org/wiki/%E0%A4%AF%E0%A5%8B%E0%A4%B8%E0%A4%BF%E0%A4%A8%E0%A5%8B%E0%A4%B0%E0%A5%80_%E0%A4%93%E0%A4%B8%E0%A5%81%E0%A4%AE%E0%A5%80" title="योसिनोरी ओसुमी – Nepalesisch" lang="ne" hreflang="ne" data-title="योसिनोरी ओसुमी" data-language-autonym="नेपाली" data-language-local-name="Nepalesisch" class="interlanguage-link-target"><span>नेपाली</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Yoshinori_Osumi" title="Yoshinori Osumi – Niederländisch" lang="nl" hreflang="nl" data-title="Yoshinori Osumi" data-language-autonym="Nederlands" data-language-local-name="Niederländisch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Norwegisch (Bokmål)" lang="nb" hreflang="nb" data-title="Yoshinori Ohsumi" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegisch (Bokmål)" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Okzitanisch" lang="oc" hreflang="oc" data-title="Yoshinori Ōsumi" data-language-autonym="Occitan" data-language-local-name="Okzitanisch" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pa mw-list-item"><a href="https://pa.wikipedia.org/wiki/%E0%A8%AF%E0%A9%8B%E0%A8%B8%E0%A8%BC%E0%A9%80%E0%A8%A8%E0%A9%8B%E0%A8%B0%E0%A9%80_%E0%A8%93%E0%A8%B8%E0%A9%81%E0%A8%AE%E0%A9%80" title="ਯੋਸ਼ੀਨੋਰੀ ਓਸੁਮੀ – Punjabi" lang="pa" hreflang="pa" data-title="ਯੋਸ਼ੀਨੋਰੀ ਓਸੁਮੀ" data-language-autonym="ਪੰਜਾਬੀ" data-language-local-name="Punjabi" class="interlanguage-link-target"><span>ਪੰਜਾਬੀ</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Polnisch" lang="pl" hreflang="pl" data-title="Yoshinori Ōsumi" data-language-autonym="Polski" data-language-local-name="Polnisch" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%DB%81%D8%B3%D9%85%DB%8C" title="یوشینوری اوہسمی – Westliches Panjabi" lang="pnb" hreflang="pnb" data-title="یوشینوری اوہسمی" data-language-autonym="پنجابی" data-language-local-name="Westliches Panjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Portugiesisch" lang="pt" hreflang="pt" data-title="Yoshinori Ohsumi" data-language-autonym="Português" data-language-local-name="Portugiesisch" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Rumänisch" lang="ro" hreflang="ro" data-title="Yoshinori Ohsumi" data-language-autonym="Română" data-language-local-name="Rumänisch" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9E%D1%81%D1%83%D0%BC%D0%B8,_%D0%81%D1%81%D0%B8%D0%BD%D0%BE%D1%80%D0%B8" title="Осуми, Ёсинори – Russisch" lang="ru" hreflang="ru" data-title="Осуми, Ёсинори" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Schottisch" lang="sco" hreflang="sco" data-title="Yoshinori Ohsumi" data-language-autonym="Scots" data-language-local-name="Schottisch" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – Serbokroatisch" lang="sh" hreflang="sh" data-title="Yoshinori Ōsumi" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbokroatisch" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-si mw-list-item"><a href="https://si.wikipedia.org/wiki/%E0%B6%BA%E0%B7%9D%E0%B7%82%E0%B7%92%E0%B6%B1%E0%B7%9C%E0%B6%BB%E0%B7%92_%E0%B6%94%E0%B7%83%E0%B7%94%E0%B6%B8%E0%B7%92" title="යෝෂිනොරි ඔසුමි – Singhalesisch" lang="si" hreflang="si" data-title="යෝෂිනොරි ඔසුමි" data-language-autonym="සිංහල" data-language-local-name="Singhalesisch" class="interlanguage-link-target"><span>සිංහල</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – einfaches Englisch" lang="en-simple" hreflang="en-simple" data-title="Yoshinori Ohsumi" data-language-autonym="Simple English" data-language-local-name="einfaches Englisch" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Jo%C5%A1inori_Ohsumi" title="Jošinori Ohsumi – Slowenisch" lang="sl" hreflang="sl" data-title="Jošinori Ohsumi" data-language-autonym="Slovenščina" data-language-local-name="Slowenisch" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Albanisch" lang="sq" hreflang="sq" data-title="Yoshinori Ohsumi" data-language-autonym="Shqip" data-language-local-name="Albanisch" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%88%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Јошинори Осуми – Serbisch" lang="sr" hreflang="sr" data-title="Јошинори Осуми" data-language-autonym="Српски / srpski" data-language-local-name="Serbisch" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Schwedisch" lang="sv" hreflang="sv" data-title="Yoshinori Ohsumi" data-language-autonym="Svenska" data-language-local-name="Schwedisch" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Suaheli" lang="sw" hreflang="sw" data-title="Yoshinori Ohsumi" data-language-autonym="Kiswahili" data-language-local-name="Suaheli" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%87%E0%AE%AF%E0%AF%8B%E0%AE%9A%E0%AE%BF%E0%AE%A9%E0%AF%8B%E0%AE%B0%E0%AE%BF_%E0%AE%93%E0%AE%9A%E0%AF%82%E0%AE%AE%E0%AE%BF" title="இயோசினோரி ஓசூமி – Tamil" lang="ta" hreflang="ta" data-title="இயோசினோரி ஓசூமி" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a href="https://te.wikipedia.org/wiki/%E0%B0%AF%E0%B1%8B%E0%B0%B7%E0%B0%BF%E0%B0%A8%E0%B1%8B%E0%B0%B0%E0%B0%BF_%E0%B0%93%E0%B0%B9%E0%B1%8D%E0%B0%B8%E0%B1%81%E0%B0%AE%E0%B1%80" title="యోషినోరి ఓహ్సుమీ – Telugu" lang="te" hreflang="te" data-title="యోషినోరి ఓహ్సుమీ" data-language-autonym="తెలుగు" data-language-local-name="Telugu" class="interlanguage-link-target"><span>తెలుగు</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B9%82%E0%B8%A2%E0%B8%8A%E0%B8%B4%E0%B9%82%E0%B8%99%E0%B8%A3%E0%B8%B4_%E0%B9%82%E0%B8%AD%E0%B8%8B%E0%B8%B9%E0%B8%A1%E0%B8%B4" title="โยชิโนริ โอซูมิ – Thailändisch" lang="th" hreflang="th" data-title="โยชิโนริ โอซูมิ" data-language-autonym="ไทย" data-language-local-name="Thailändisch" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Türkisch" lang="tr" hreflang="tr" data-title="Yoshinori Ohsumi" data-language-autonym="Türkçe" data-language-local-name="Türkisch" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми – Tatarisch" lang="tt" hreflang="tt" data-title="Йошинори Осуми" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatarisch" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9E%D1%81%D1%83%D0%BC%D1%96_%D0%99%D0%BE%D1%81%D1%96%D0%BD%D0%BE%D1%80%D1%96" title="Осумі Йосінорі – Ukrainisch" lang="uk" hreflang="uk" data-title="Осумі Йосінорі" data-language-autonym="Українська" data-language-local-name="Ukrainisch" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%D8%B4%D9%88%D9%85%DB%8C" title="یوشینوری اوشومی – Urdu" lang="ur" hreflang="ur" data-title="یوشینوری اوشومی" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/%C5%8Csumi_Yoshinori" title="Ōsumi Yoshinori – Vietnamesisch" lang="vi" hreflang="vi" data-title="Ōsumi Yoshinori" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamesisch" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 – Wu" lang="wuu" hreflang="wuu" data-title="大隅良典" data-language-autonym="吴语" data-language-local-name="Wu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 – Chinesisch" lang="zh" hreflang="zh" data-title="大隅良典" data-language-autonym="中文" data-language-local-name="Chinesisch" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/%C3%94sumi_Yosinori" title="Ôsumi Yosinori – Min Nan" lang="nan" hreflang="nan" data-title="Ôsumi Yosinori" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Min Nan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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