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class="mw-page-title-main">Йошинори Осуми</span></h1> <div id="bodyContent" class="vector-body"> <div id="siteSub" class="noprint">Уикипедия — ашық энциклопедиясынан алынған мәлімет</div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="contentSub2"></div> <div id="jump-to-nav"></div> <a class="mw-jump-link" href="#mw-head">Навигацияға өту</a> <a class="mw-jump-link" href="#searchInput">Іздеуге өту</a> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="kk" dir="ltr"><table class="infobox vcard" cellspacing="2" style="width: 22em; text-align: left; font-size: 88%; line-height: 1.5em; width: 23em;"><tbody><tr><td colspan="2" class="fn" style="text-align:center; font-size: 125%; font-weight: bold; background:#eee; padding: 0.3em;">Йошинори Осуми</td></tr><tr><td colspan="2" class="" style="text-align:center;"> <span typeof="mw:File"><a href="/wiki/%D0%A1%D1%83%D1%80%D0%B5%D1%82:Nobel_Laureates_1042_(30647248184).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/67/Nobel_Laureates_1042_%2830647248184%29.jpg/200px-Nobel_Laureates_1042_%2830647248184%29.jpg" decoding="async" width="200" height="267" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/67/Nobel_Laureates_1042_%2830647248184%29.jpg/300px-Nobel_Laureates_1042_%2830647248184%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/67/Nobel_Laureates_1042_%2830647248184%29.jpg/400px-Nobel_Laureates_1042_%2830647248184%29.jpg 2x" data-file-width="3456" data-file-height="4608" /></a></span> </td></tr><tr><th style="">Туған күні</th><td class="" style=""> <p><span class="no-wikidata" data-wikidata-property-id="P569"><a href="/wiki/9_%D0%B0%D2%9B%D0%BF%D0%B0%D0%BD" title="9 ақпан">9 ақпан</a> <a href="/wiki/1945_%D0%B6%D1%8B%D0%BB" title="1945 жыл">1945 </a></span><span style="display:none">(<span class="bday">1945-02-09</span>)</span> <span class="nowrap">(79 жас)</span> </p> </td></tr><tr><th style="">Туған жері</th><td class="" style=""> <p><a href="/w/index.php?title=%D0%A4%D1%83%D0%BA%D1%83%D0%BE%D0%BA%D0%B0&action=edit&redlink=1" class="new" title="Фукуока (мұндай бет жоқ)">Фукуока</a>, <a href="/wiki/%D0%96%D0%B0%D0%BF%D0%BE%D0%BD%D0%B8%D1%8F" title="Жапония">Жапония</a> </p> </td></tr><tr><th style="">Ғылыми аясы</th><td class="" style=""> <p><a href="/wiki/%D0%9C%D0%BE%D0%BB%D0%B5%D0%BA%D1%83%D0%BB%D0%B0%D0%BB%D1%8B%D2%9B_%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Молекулалық биология">молекулалық биология</a> </p> </td></tr><tr><th style="">Жұмыс орны</th><td class="org" style=""> <p><a href="/w/index.php?title=%D0%A2%D0%BE%D0%BA%D0%B8%D0%BE_%D1%82%D0%B5%D1%85%D0%BD%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F%D0%BB%D1%8B%D2%9B_%D0%B8%D0%BD%D1%81%D1%82%D0%B8%D1%82%D1%83%D1%82%D1%8B&action=edit&redlink=1" class="new" title="Токио технологиялық институты (мұндай бет жоқ)">Токио технологиялық институты</a> </p> </td></tr><tr><th style="">Ғылыми атағы</th><td class="" style=""> <p><a href="/w/index.php?title=%D2%AE%D0%BB%D0%B3%D1%96:%D2%92%D1%8B%D0%BB%D1%8B%D0%BC%D0%B8_%D0%B0%D1%82%D0%B0%D2%9B&action=edit&redlink=1" class="new" title="Үлгі:Ғылыми атақ (мұндай бет жоқ)">Үлгі:Ғылыми атақ</a> (1974) </p> </td></tr><tr><th style="">Альма-матер</th><td class="" style=""> <p><a href="/wiki/%D0%A2%D0%BE%D0%BA%D0%B8%D0%BE_%D1%83%D0%BD%D0%B8%D0%B2%D0%B5%D1%80%D1%81%D0%B8%D1%82%D0%B5%D1%82%D1%96" title="Токио университеті">Токио университеті</a> </p> </td></tr><tr><th style="">Марапаттары</th><td class="" style=""> <p><br style="line-height:0;" /> </p> <table style="background:transparent"> <tbody><tr><td></td></tr></tbody></table> <p><span typeof="mw:File"><a href="/wiki/%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B" title="Нобель сыйлығы — 2016"><img alt="Нобель сыйлығы — 2016" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/20px-Nobel_prize_medal.svg.png" decoding="async" width="20" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/30px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/40px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span> <a href="/wiki/%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B" title="Физиология немесе медицина саласындағы Нобель сыйлығы">Медицина саласындағы Нобель сыйлығы</a> (<a href="/wiki/2016_%D0%B6%D1%8B%D0%BB" title="2016 жыл">2016</a>) <br /> <a href="/w/index.php?title=%D0%A0%D0%BE%D0%B7%D0%B5%D0%BD%D1%81%D1%82%D0%B8%D0%BB%D0%B0_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Розенстила жүлдесі (мұндай бет жоқ)">Розенстила жүлдесі</a> (2015) <br /> <a href="/w/index.php?title=%D0%A5%D0%B0%D0%BB%D1%8B%D2%9B%D0%B0%D1%80%D0%B0%D0%BB%D1%8B%D2%9B_%D0%93%D0%B0%D0%B9%D1%80%D0%B4%D0%BD%D0%B5%D1%80_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Халықаралық Гайрднер жүлдесі (мұндай бет жоқ)">Халықаралық Гайрднер жүлдесі</a> (2015) <br /> <a href="/w/index.php?title=%D0%A5%D0%B0%D0%BB%D1%8B%D2%9B%D0%B0%D1%80%D0%B0%D0%BB%D1%8B%D2%9B_%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Халықаралық биология жүлдесі (мұндай бет жоқ)">Халықаралық биология жүлдесі</a> (2015) <br /> <a href="/w/index.php?title=Thomson_Reuters_Citation_Laureates&action=edit&redlink=1" class="new" title="Thomson Reuters Citation Laureates (мұндай бет жоқ)">Thomson Reuters Citation Laureates</a> (2013)<br /> <a href="/w/index.php?title=%D0%9A%D0%B8%D0%BE%D1%82%D0%BE_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Киото жүлдесі (мұндай бет жоқ)">Киото жүлдесі</a> (2012) <br /> <a href="/w/index.php?title=%D0%90%D1%81%D0%B0%D1%85%D0%B8_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Асахи жүлдесі (мұндай бет жоқ)">Асахи жүлдесі</a> (2008) </p> </td></tr> </tbody></table> <p><b>Йошинори Осуми</b> (Фукуока, <a href="/wiki/%D0%96%D0%B0%D0%BF%D0%BE%D0%BD%D0%B8%D1%8F" title="Жапония">Жапония</a>, <a href="/wiki/1945_%D0%B6%D1%8B%D0%BB" title="1945 жыл">1945 жылы</a> 9-ақпанда т.) <a href="/wiki/%D0%90%D0%B2%D1%82%D0%BE%D1%84%D0%B0%D0%B3%D0%B8%D1%8F" title="Автофагия">аутофагияға</a> маманданған <a href="/wiki/%D0%96%D0%B0%D0%BF%D0%BE%D0%BD%D0%B8%D1%8F" title="Жапония">жапон</a> <a href="/wiki/%D0%A6%D0%B8%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Цитология">молелекулалық биолог</a>. Токио Технологиялық Институтының профессоры (құрметті).<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Ол <a href="/wiki/%D0%90%D0%B2%D1%82%D0%BE%D1%84%D0%B0%D0%B3%D0%B8%D1%8F" title="Автофагия">аутофагия</a> механизмдеріне қатысты ашылулары үшін 2012 жылы Киото сыйлығын,<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> , және 2016 жылы <a href="/wiki/%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B_%D0%BB%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B%D0%BD%D1%8B%D2%A3_%D1%82%D1%96%D0%B7%D1%96%D0%BC%D1%96" title="Физиология немесе медицина саласында Нобель сыйлығы лауреаттарының тізімі">Медицина бойынша Нобель сыйлығын</a> алған.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div id="toc" class="toc" role="navigation" aria-labelledby="mw-toc-heading"><input type="checkbox" role="button" id="toctogglecheckbox" class="toctogglecheckbox" style="display:none" /><div class="toctitle" lang="kk" dir="ltr"><h2 id="mw-toc-heading">Мазмұны</h2><span class="toctogglespan"><label class="toctogglelabel" for="toctogglecheckbox"></label></span></div> <ul> <li class="toclevel-1 tocsection-1"><a href="#Марапаттар"><span class="tocnumber">1</span> <span class="toctext">Марапаттар</span></a></li> <li class="toclevel-1 tocsection-2"><a href="#Жұмыс"><span class="tocnumber">2</span> <span class="toctext">Жұмыс</span></a> <ul> <li class="toclevel-2 tocsection-3"><a href="#Жарияланымдар"><span class="tocnumber">2.1</span> <span class="toctext">Жарияланымдар</span></a></li> </ul> </li> <li class="toclevel-1 tocsection-4"><a href="#Дереккөздер"><span class="tocnumber">3</span> <span class="toctext">Дереккөздер</span></a></li> </ul> </div> <div class="mw-heading mw-heading2"><h2 id="Марапаттар"><span id=".D0.9C.D0.B0.D1.80.D0.B0.D0.BF.D0.B0.D1.82.D1.82.D0.B0.D1.80"></span>Марапаттар</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&veaction=edit&section=1" title="Бөлімді өңдеу: Марапаттар" class="mw-editsection-visualeditor"><span>өңдеу</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&action=edit&section=1" title="Edit section's source code: Марапаттар"><span>қайнарын өңдеу</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="https://ja.wikipedia.org/wiki/%E8%97%A4%E5%8E%9F%E8%B3%9E" class="extiw" title="ja:藤原賞">Фудзихара жүлдесі</a><span style="white-space: nowrap"><span class="noprint" style="font-size:95%; position: relative; top: .4em;"> <span class="ref-info" title="жапон тілінде" style="font-size:85%; cursor:help; color:#888;">(жап.)</span></span><span class="link-ru metadata noprint" style="font-size:80%; margin-left:-1.7em; position: relative; top: -.4em;"><a href="/w/index.php?title=%D0%A4%D1%83%D0%B4%D0%B7%D0%B8%D1%85%D0%B0%D1%80%D0%B0_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Фудзихара жүлдесі (мұндай бет жоқ)">қаз.</a></span></span> (2005)<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li> <li><a href="https://en.wikipedia.org/wiki/Japan_Academy_Prize_(academics)" class="extiw" title="en:Japan Academy Prize (academics)">Жапон ғылым академиясының жүлдесі</a><span style="white-space: nowrap"><span class="noprint" style="font-size:95%; position: relative; top: .4em;"> <span class="ref-info" title="ағылшын тілінде" style="font-size:85%; cursor:help; color:#888;">(ағыл.)</span></span><span class="link-ru metadata noprint" style="font-size:80%; margin-left:-1.7em; position: relative; top: -.4em;"><a href="/w/index.php?title=%D0%96%D0%B0%D0%BF%D0%BE%D0%BD_%D2%93%D1%8B%D0%BB%D1%8B%D0%BC_%D0%B0%D0%BA%D0%B0%D0%B4%D0%B5%D0%BC%D0%B8%D1%8F%D1%81%D1%8B%D0%BD%D1%8B%D2%A3_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Жапон ғылым академиясының жүлдесі (мұндай бет жоқ)">қаз.</a></span></span> (2006)<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></li> <li>Жапония Ботаникалық Қоғамының сыйлығы (2007)<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup></li> <li><a href="/w/index.php?title=%D0%90%D1%81%D0%B0%D1%85%D0%B8_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Асахи жүлдесі (мұндай бет жоқ)">Асахи жүлдесі</a> (2008)<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li><a href="/w/index.php?title=%D0%9A%D0%B8%D0%BE%D1%82%D0%BE_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Киото жүлдесі (мұндай бет жоқ)">Киото жүлдесі</a> (2012)<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup></li> <li><a href="/w/index.php?title=Thomson_Reuters_Citation_Laureates&action=edit&redlink=1" class="new" title="Thomson Reuters Citation Laureates (мұндай бет жоқ)">Thomson Reuters Citation Laureates</a> (2013)<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup></li> <li><a href="/w/index.php?title=%D0%A5%D0%B0%D0%BB%D1%8B%D2%9B%D0%B0%D1%80%D0%B0%D0%BB%D1%8B%D2%9B_%D0%93%D0%B0%D0%B9%D0%B4%D0%BD%D0%B5%D1%80_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B&action=edit&redlink=1" class="new" title="Халықаралық Гайднер сыйлығы (мұндай бет жоқ)">Халықаралық Гайднер сыйлығы</a> (2015)<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup></li> <li><a href="/w/index.php?title=%D0%91%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%B1%D0%BE%D0%B9%D1%8B%D0%BD%D1%88%D0%B0_%D1%85%D0%B0%D0%BB%D1%8B%D2%9B%D0%B0%D1%80%D0%B0%D0%BB%D1%8B%D2%9B_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%9B&action=edit&redlink=1" class="new" title="Биология бойынша халықаралық сыйлық (мұндай бет жоқ)">Биология бойынша халықаралық сыйлық</a> (2015)</li> <li><a href="/w/index.php?title=%D0%A0%D0%BE%D0%B7%D0%B5%D0%BD%D1%81%D1%82%D0%B8%D0%BB%D0%B0_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Розенстила жүлдесі (мұндай бет жоқ)">Розенстила жүлдесі</a> (2015)</li> <li><a href="https://en.wikipedia.org/wiki/Keio_Medical_Science_Prize" class="extiw" title="en:Keio Medical Science Prize">Кэйо жүлдесі</a><span style="white-space: nowrap"><span class="noprint" style="font-size:95%; position: relative; top: .4em;"> <span class="ref-info" title="ағылшын тілінде" style="font-size:85%; cursor:help; color:#888;">(ағыл.)</span></span><span class="link-ru metadata noprint" style="font-size:80%; margin-left:-1.7em; position: relative; top: -.4em;"><a href="/w/index.php?title=%D0%9A%D1%8D%D0%B9%D0%BE_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Кэйо жүлдесі (мұндай бет жоқ)">қаз.</a></span></span> (2015)<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup></li> <li><a href="https://en.wikipedia.org/wiki/Person_of_Cultural_Merit" class="extiw" title="en:Person of Cultural Merit">Еңбегі сіңген мәдениет қызметкері (Жапония)</a><span style="white-space: nowrap"><span class="noprint" style="font-size:95%; position: relative; top: .4em;"> <span class="ref-info" title="ағылшын тілінде" style="font-size:85%; cursor:help; color:#888;">(ағыл.)</span></span><span class="link-ru metadata noprint" style="font-size:80%; margin-left:-1.7em; position: relative; top: -.4em;"><a href="/w/index.php?title=%D0%95%D2%A3%D0%B1%D0%B5%D0%B3%D1%96_%D1%81%D1%96%D2%A3%D0%B3%D0%B5%D0%BD_%D0%BC%D3%99%D0%B4%D0%B5%D0%BD%D0%B8%D0%B5%D1%82_%D2%9B%D1%8B%D0%B7%D0%BC%D0%B5%D1%82%D0%BA%D0%B5%D1%80%D1%96_(%D0%96%D0%B0%D0%BF%D0%BE%D0%BD%D0%B8%D1%8F)&action=edit&redlink=1" class="new" title="Еңбегі сіңген мәдениет қызметкері (Жапония) (мұндай бет жоқ)">қаз.</a></span></span> (2015)<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup></li> <li><a href="https://en.wikipedia.org/wiki/Wiley_Prize" class="extiw" title="en:Wiley Prize">Уайли жүлдесі</a><span style="white-space: nowrap"><span class="noprint" style="font-size:95%; position: relative; top: .4em;"> <span class="ref-info" title="ағылшын тілінде" style="font-size:85%; cursor:help; color:#888;">(ағыл.)</span></span><span class="link-ru metadata noprint" style="font-size:80%; margin-left:-1.7em; position: relative; top: -.4em;"><a href="/w/index.php?title=%D0%A3%D0%B0%D0%B9%D0%BB%D0%B8_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Уайли жүлдесі (мұндай бет жоқ)">қаз.</a></span></span> (2016)<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup></li> <li><a href="https://en.wikipedia.org/wiki/Dr._Paul_Janssen_Award_for_Biomedical_Research" class="extiw" title="en:Dr. Paul Janssen Award for Biomedical Research">Поль Джанссен жүлдесі</a><span style="white-space: nowrap"><span class="noprint" style="font-size:95%; position: relative; top: .4em;"> <span class="ref-info" title="ағылшын тілінде" style="font-size:85%; cursor:help; color:#888;">(ағыл.)</span></span><span class="link-ru metadata noprint" style="font-size:80%; margin-left:-1.7em; position: relative; top: -.4em;"><a href="/w/index.php?title=%D0%9F%D0%BE%D0%BB%D1%8C_%D0%94%D0%B6%D0%B0%D0%BD%D1%81%D1%81%D0%B5%D0%BD_%D0%B6%D2%AF%D0%BB%D0%B4%D0%B5%D1%81%D1%96&action=edit&redlink=1" class="new" title="Поль Джанссен жүлдесі (мұндай бет жоқ)">қаз.</a></span></span> (2016)<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B" title="Физиология немесе медицина саласындағы Нобель сыйлығы">Медицина саласындағы Нобель сыйлығы</a> (2016)</li></ul> <div class="mw-heading mw-heading2"><h2 id="Жұмыс"><span id=".D0.96.D2.B1.D0.BC.D1.8B.D1.81"></span>Жұмыс</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&veaction=edit&section=2" title="Бөлімді өңдеу: Жұмыс" class="mw-editsection-visualeditor"><span>өңдеу</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&action=edit&section=2" title="Edit section's source code: Жұмыс"><span>қайнарын өңдеу</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Жарияланымдар"><span id=".D0.96.D0.B0.D1.80.D0.B8.D1.8F.D0.BB.D0.B0.D0.BD.D1.8B.D0.BC.D0.B4.D0.B0.D1.80"></span>Жарияланымдар</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&veaction=edit&section=3" title="Бөлімді өңдеу: Жарияланымдар" class="mw-editsection-visualeditor"><span>өңдеу</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&action=edit&section=3" title="Edit section's source code: Жарияланымдар"><span>қайнарын өңдеу</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><span class="citation Journal">Tsukada, M; Ōsumi, Y (25 de octubre de 1993). "Isolation and characterization of autophagy-defective mutants of Saccharomyces cerevisiae". <i>FEBS Letters</i> <b>333</b> (1-2): 169-74. <a href="/w/index.php?title=Digital_object_identifier&action=edit&redlink=1" class="new" title="Digital object identifier (мұндай бет жоқ)">doi</a>:<a href="//doi.org/10.1016/0014-5793(93)80398-E" class="extiw" title="doi:10.1016/0014-5793(93)80398-E">10.1016/0014-5793(93)80398-E</a>. <a href="/w/index.php?title=International_Standard_Serial_Number&action=edit&redlink=1" class="new" title="International Standard Serial Number (мұндай бет жоқ)">ISSN</a> <a rel="nofollow" class="external text" href="http://www.worldcat.org/issn/0014-5793">0014-5793</a>. <a href="/w/index.php?title=PubMed_Identifier&action=edit&redlink=1" class="new" title="PubMed Identifier (мұндай бет жоқ)">PMID</a> <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/pubmed/8224160">8224160</a>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Isolation+and+characterization+of+autophagy-defective+mutants+of+Saccharomyces+cerevisiae&rft.jtitle=FEBS+Letters&rft.aulast=Tsukada&rft.aufirst=M&rft.au=Tsukada%2C%26%2332%3BM&rft.au=%C5%8Csumi%2C%26%2332%3BY&rft.date=25+de+octubre+de+1993&rft.volume=333&rft.issue=1-2&rft.pages=169-74&rft_id=info:doi/10.1016%2F0014-5793%2893%2980398-E&rft.issn=0014-5793&rft_id=info:pmid/8224160&rfr_id=info:sid/en.wikipedia.org:%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8"><span style="display: none;"> </span></span></li> <li><span class="citation Journal">Mizushima, N; Noda, T; Yoshimori, T; Tanaka, Y; Ishii, T; George, MD; Klionsky, DJ; Ohsumi, M et al. (24 de septiembre de 1998). "A protein conjugation system essential for autophagy". <i>Nature</i> <b>395</b> (6700): 395-8. <a href="/w/index.php?title=Digital_object_identifier&action=edit&redlink=1" class="new" title="Digital object identifier (мұндай бет жоқ)">doi</a>:<a href="//doi.org/10.1038/26506" class="extiw" title="doi:10.1038/26506">10.1038/26506</a>. <a href="/w/index.php?title=PubMed_Identifier&action=edit&redlink=1" class="new" title="PubMed Identifier (мұндай бет жоқ)">PMID</a> <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/pubmed/9759731">9759731</a>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+protein+conjugation+system+essential+for+autophagy&rft.jtitle=Nature&rft.aulast=Mizushima&rft.aufirst=N&rft.au=Mizushima%2C%26%2332%3BN&rft.au=Noda%2C%26%2332%3BT&rft.au=Yoshimori%2C%26%2332%3BT&rft.au=Tanaka%2C%26%2332%3BY&rft.au=Ishii%2C%26%2332%3BT&rft.au=George%2C%26%2332%3BMD&rft.au=Klionsky%2C%26%2332%3BDJ&rft.au=Ohsumi%2C%26%2332%3BM&rft.au=%C5%8Csumi%2C%26%2332%3BY&rft.date=24+de+septiembre+de+1998&rft.volume=395&rft.issue=6700&rft.pages=395-8&rft_id=info:doi/10.1038%2F26506&rft_id=info:pmid/9759731&rfr_id=info:sid/en.wikipedia.org:%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8"><span style="display: none;"> </span></span>Se sugiere usar <code>|número-autores=</code> (ayuda) <a href="/w/index.php?title=Categor%C3%ADa:Wikipedia:P%C3%A1ginas_con_referencias_con_et_al._impl%C3%ADcito_en_los_autores&action=edit&redlink=1" class="new" title="Categoría:Wikipedia:Páginas con referencias con et al. implícito en los autores (мұндай бет жоқ)">Categoría:Wikipedia:Páginas con referencias con et al. implícito en los autores</a></li> <li><span class="citation Journal">Kabeya, Y.; Mizushima, N.; Ueno, T.; Yamamoto, A.; Kirisako, T.; Noda, T.; Kominami, E.; Ōsumi, Y. et al. (2000). "LC3, a mammalian homologue of yeast Apg8p, is localized in autophagosome membranes after processing". <i>The EMBO Journal</i> <b>19</b> (21): 5720-5728. <a href="/w/index.php?title=Digital_object_identifier&action=edit&redlink=1" class="new" title="Digital object identifier (мұндай бет жоқ)">doi</a>:<a href="//doi.org/10.1093/emboj/19.21.5720" class="extiw" title="doi:10.1093/emboj/19.21.5720">10.1093/emboj/19.21.5720</a>. <a href="/w/index.php?title=PubMed_Central&action=edit&redlink=1" class="new" title="PubMed Central (мұндай бет жоқ)">PMC</a> <a rel="nofollow" class="external text" href="http://www.pubmedcentral.gov/articlerender.fcgi?tool=pmcentrez&artid=305793">305793</a>. <a href="/w/index.php?title=PubMed_Identifier&action=edit&redlink=1" class="new" title="PubMed Identifier (мұндай бет жоқ)">PMID</a> <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/pubmed/11060023">11060023</a>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=LC3%2C+a+mammalian+homologue+of+yeast+Apg8p%2C+is+localized+in+autophagosome+membranes+after+processing&rft.jtitle=The+EMBO+Journal&rft.aulast=Kabeya&rft.aufirst=Y.&rft.au=Kabeya%2C%26%2332%3BY.&rft.au=Mizushima%2C%26%2332%3BN.&rft.au=Ueno%2C%26%2332%3BT.&rft.au=Yamamoto%2C%26%2332%3BA.&rft.au=Kirisako%2C%26%2332%3BT.&rft.au=Noda%2C%26%2332%3BT.&rft.au=Kominami%2C%26%2332%3BE.&rft.au=%C5%8Csumi%2C%26%2332%3BY.&rft.au=Yoshimori%2C%26%2332%3BT.&rft.date=2000&rft.volume=19&rft.issue=21&rft.pages=5720-5728&rft_id=info:doi/10.1093%2Femboj%2F19.21.5720&rft_id=info:pmc/305793&rft_id=info:pmid/11060023&rfr_id=info:sid/en.wikipedia.org:%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8"><span style="display: none;"> </span></span>Se sugiere usar <code>|número-autores=</code> (ayuda) <a href="/w/index.php?title=Categor%C3%ADa:Wikipedia:P%C3%A1ginas_con_referencias_con_et_al._impl%C3%ADcito_en_los_autores&action=edit&redlink=1" class="new" title="Categoría:Wikipedia:Páginas con referencias con et al. implícito en los autores (мұндай бет жоқ)">Categoría:Wikipedia:Páginas con referencias con et al. implícito en los autores</a></li> <li><span class="citation Journal">Ichimura, Y; Kirisako, T; Takao, T; Satomi, Y; Shimonishi, Y; Ishihara, N; Mizushima, N; Tanida, I et al. (23 de noviembre de 2000). "A ubiquitin-like system mediates protein lipidation". <i>Nature</i> <b>408</b> (6811): 488-92. <a href="/w/index.php?title=Digital_object_identifier&action=edit&redlink=1" class="new" title="Digital object identifier (мұндай бет жоқ)">doi</a>:<a href="//doi.org/10.1038/35044114" class="extiw" title="doi:10.1038/35044114">10.1038/35044114</a>. <a href="/w/index.php?title=PubMed_Identifier&action=edit&redlink=1" class="new" title="PubMed Identifier (мұндай бет жоқ)">PMID</a> <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/pubmed/11100732">11100732</a>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+ubiquitin-like+system+mediates+protein+lipidation&rft.jtitle=Nature&rft.aulast=Ichimura&rft.aufirst=Y&rft.au=Ichimura%2C%26%2332%3BY&rft.au=Kirisako%2C%26%2332%3BT&rft.au=Takao%2C%26%2332%3BT&rft.au=Satomi%2C%26%2332%3BY&rft.au=Shimonishi%2C%26%2332%3BY&rft.au=Ishihara%2C%26%2332%3BN&rft.au=Mizushima%2C%26%2332%3BN&rft.au=Tanida%2C%26%2332%3BI&rft.au=Kominami%2C%26%2332%3BE&rft.date=23+de+noviembre+de+2000&rft.volume=408&rft.issue=6811&rft.pages=488-92&rft_id=info:doi/10.1038%2F35044114&rft_id=info:pmid/11100732&rfr_id=info:sid/en.wikipedia.org:%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8"><span style="display: none;"> </span></span></li> <li><span class="citation Journal">Ōsumi, Y (marzo de 2001). "Molecular dissection of autophagy: two ubiquitin-like systems". <i>Nature</i> <b>2</b> (3): 211-6. <a href="/w/index.php?title=Digital_object_identifier&action=edit&redlink=1" class="new" title="Digital object identifier (мұндай бет жоқ)">doi</a>:<a href="//doi.org/10.1038/35044114" class="extiw" title="doi:10.1038/35044114">10.1038/35044114</a>. <a href="/w/index.php?title=PubMed_Identifier&action=edit&redlink=1" class="new" title="PubMed Identifier (мұндай бет жоқ)">PMID</a> <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/pubmed/11265251">11265251</a>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Molecular+dissection+of+autophagy%3A+two+ubiquitin-like+systems&rft.jtitle=Nature&rft.aulast=%C5%8Csumi&rft.aufirst=Y&rft.au=%C5%8Csumi%2C%26%2332%3BY&rft.date=marzo+de+2001&rft.volume=2&rft.issue=3&rft.pages=211-6&rft_id=info:doi/10.1038%2F35044114&rft_id=info:pmid/11265251&rfr_id=info:sid/en.wikipedia.org:%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8"><span style="display: none;"> </span></span></li> <li><span class="citation Journal">Kuma, A; Hatano, M; Matsui, M; Yamamoto, A; Nakaya, H; Yoshimori, T; Ōsumi, Y; Tokuhisa, T et al. (23 de diciembre de 2004). "The role of autophagy during the early neonatal starvation period". <i>Nature</i> <b>432</b> (7020): 1032-6. <a href="/w/index.php?title=Digital_object_identifier&action=edit&redlink=1" class="new" title="Digital object identifier (мұндай бет жоқ)">doi</a>:<a href="//doi.org/10.1038/nature03029" class="extiw" title="doi:10.1038/nature03029">10.1038/nature03029</a>. <a href="/w/index.php?title=PubMed_Identifier&action=edit&redlink=1" class="new" title="PubMed Identifier (мұндай бет жоқ)">PMID</a> <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/pubmed/15525940">15525940</a>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=The+role+of+autophagy+during+the+early+neonatal+starvation+period&rft.jtitle=Nature&rft.aulast=Kuma&rft.aufirst=A&rft.au=Kuma%2C%26%2332%3BA&rft.au=Hatano%2C%26%2332%3BM&rft.au=Matsui%2C%26%2332%3BM&rft.au=Yamamoto%2C%26%2332%3BA&rft.au=Nakaya%2C%26%2332%3BH&rft.au=Yoshimori%2C%26%2332%3BT&rft.au=%C5%8Csumi%2C%26%2332%3BY&rft.au=Tokuhisa%2C%26%2332%3BT&rft.au=Mizushima%2C%26%2332%3BN&rft.date=23+de+diciembre+de+2004&rft.volume=432&rft.issue=7020&rft.pages=1032-6&rft_id=info:doi/10.1038%2Fnature03029&rft_id=info:pmid/15525940&rfr_id=info:sid/en.wikipedia.org:%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8"><span style="display: none;"> </span></span>Se sugiere usar <code>|número-autores=</code> (ayuda) <a href="/w/index.php?title=Categor%C3%ADa:Wikipedia:P%C3%A1ginas_con_referencias_con_et_al._impl%C3%ADcito_en_los_autores&action=edit&redlink=1" class="new" title="Categoría:Wikipedia:Páginas con referencias con et al. implícito en los autores (мұндай бет жоқ)">Categoría:Wikipedia:Páginas con referencias con et al. implícito en los autores</a></li> <li><span class="citation Journal">Hanada, T; Noda, NN; Satomi, Y; Ichimura, Y; Fujioka, Y; Takao, T; Inagaki, F; Ohsumi, Y (28 de diciembre de 2007). "The Atg12-Atg5 conjugate has a novel E3-like activity for protein lipidation in autophagy". <i>Journal of Biological Chemistry</i> <b>282</b> (52): 37298-302. <a href="/w/index.php?title=Digital_object_identifier&action=edit&redlink=1" class="new" title="Digital object identifier (мұндай бет жоқ)">doi</a>:<a href="//doi.org/10.1074/jbc.C700195200" class="extiw" title="doi:10.1074/jbc.C700195200">10.1074/jbc.C700195200</a>. <a href="/w/index.php?title=PubMed_Identifier&action=edit&redlink=1" class="new" title="PubMed Identifier (мұндай бет жоқ)">PMID</a> <a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/pubmed/17986448">17986448</a>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=The+Atg12-Atg5+conjugate+has+a+novel+E3-like+activity+for+protein+lipidation+in+autophagy&rft.jtitle=Journal+of+Biological+Chemistry&rft.aulast=Hanada&rft.aufirst=T&rft.au=Hanada%2C%26%2332%3BT&rft.au=Noda%2C%26%2332%3BNN&rft.au=Satomi%2C%26%2332%3BY&rft.au=Ichimura%2C%26%2332%3BY&rft.au=Fujioka%2C%26%2332%3BY&rft.au=Takao%2C%26%2332%3BT&rft.au=Inagaki%2C%26%2332%3BF&rft.au=Ohsumi%2C%26%2332%3BY&rft.date=28+de+diciembre+de+2007&rft.volume=282&rft.issue=52&rft.pages=37298-302&rft_id=info:doi/10.1074%2Fjbc.C700195200&rft_id=info:pmid/17986448&rfr_id=info:sid/en.wikipedia.org:%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8"><span style="display: none;"> </span></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="Дереккөздер"><span id=".D0.94.D0.B5.D1.80.D0.B5.D0.BA.D0.BA.D3.A9.D0.B7.D0.B4.D0.B5.D1.80"></span>Дереккөздер</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&veaction=edit&section=4" title="Бөлімді өңдеу: Дереккөздер" class="mw-editsection-visualeditor"><span>өңдеу</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8&action=edit&section=4" title="Edit section's source code: Дереккөздер"><span>қайнарын өңдеу</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="reflist columns" style="list-style-type: decimal;"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><span class="citation web" id="CITAREF3_de_octubre_de_2016"><a rel="nofollow" class="external text" href="http://www.titech.ac.jp/english/news/2016/036279.html">"Honorary Professor Йошинори Ohsumi wins Nobel Prize in Physiology or Medicine for 2016"</a>. <i>titech.ac.jp</i> (ағылшын тілінде). 3 қазан күні 2016 жылғы<span class="reference-accessdate">. </span></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><span class="citation noticia">Biemiller, Лоуренс (10 тамыз 2012). </span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.elmundo.es/salud/2016/10/03/57f2193e268e3edd358b457d.html">Нобель сыйлығы, Медицина қызметкерлері үшін Йошинори Ōsumi, олардың нәтижелері бойынша регенерация жасушалар." 3 қазан күні 2016 жылғы.</a></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><span class="citation web"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/2016/press.html">"The Nobel Prize in Physiology or Medicine 2016"</a>. <i>nobelprize.org</i> (ағылшын тілінде). 3 қазан күні 2016 жылғы<span class="reference-accessdate">. </span></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><a href="#cite_ref-5">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.fujizai.or.jp/e_prize_one46_50.htm">The Fujihara Foundation of Science</a> <small><a rel="nofollow" class="external text" href="https://web.archive.org/web/20161003201525/http://www.fujizai.or.jp/e_prize_one46_50.htm">Мұрағатталған</a> 3 қазанның 2016 жылы.</small></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><a href="#cite_ref-6">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.japan-acad.go.jp/en/activities/jyusho/091to100.html">The Imperial Prize,Japan Academy Prize,Duke of Edinburgh Prize Recipients</a> // Yhe Japan Academy</span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><a href="#cite_ref-7">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://bsj.or.jp/jpn/members/prize/awardee.php">日本植物学会賞受賞者一覧 (Список победителей премии)</a></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><a href="#cite_ref-8">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.asahi.com/shimbun/award/asahi/english.html#2008">The Asahi Prize</a> // The Asahi Shimbun Company</span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><a href="#cite_ref-9">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.kyotoprize.org/en/laureates/yoshinori-ohsumi">Yoshinori Ohsumi | 2012 Laureate</a> // Kyoto Prize</span> </li> <li id="cite_note-10"><span class="mw-cite-backlink"><a href="#cite_ref-10">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://sciencewatch.com/nobel/2013-predictions">2013 Predictions</a> // ScienceWatch | Thomson Reuters</span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><a href="#cite_ref-11">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.gairdner.org/content/yoshinori-ohsumi">Yoshinori Ohsumi</a> // Gairdner</span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><a href="#cite_ref-12">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.ms-fund.keio.ac.jp/prize/winner/prize2015.html">The 2015 Keio Medical Science Prize Awardees</a> <small><a rel="nofollow" class="external text" href="https://web.archive.org/web/20170630173138/http://www.ms-fund.keio.ac.jp/prize/winner/prize2015.html">Мұрағатталған</a> 30 маусымның 2017 жылы.</small></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><a href="#cite_ref-13">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.titech.ac.jp/english/news/2016/033374.html">Honorary Professor Yoshinori Ohsumi selected as 2015 Person of Cultural Merit</a></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><a href="#cite_ref-14">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://eu.wiley.com/WileyCDA/PressRelease/pressReleaseId-123822.html">The 15th Annual Wiley Prize in Biomedical Sciences Awarded for Studies of Autophagy</a></span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><a href="#cite_ref-15">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.jnj.com/news/all/Yoshinori-Ohsumi-PhD-Wins-2016-Dr-Paul-Janssen-Award-for-Biomedical-Research">Yoshinori Ohsumi, Ph.D., Wins 2016 Dr. Paul Janssen Award for Biomedical Research</a> <small><a rel="nofollow" class="external text" href="https://web.archive.org/web/20161003200138/http://www.jnj.com/news/all/Yoshinori-Ohsumi-PhD-Wins-2016-Dr-Paul-Janssen-Award-for-Biomedical-Research">Мұрағатталған</a> 3 қазанның 2016 жылы.</small> // Johnson & Johnson</span> </li> </ol></div></div> <table class="navbox" style="border-spacing:0"><tbody><tr><td style="padding:2px"><table class="nowraplinks collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><span style="float:left;text-align:left;width:5em;margin-right:0.5em"><span typeof="mw:File"><a href="/wiki/%D2%AE%D0%BB%D0%B3%D1%96:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%9B%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B_2001-2025" title="Бұл үлгіні қарау"><img alt="Бұл үлгіні қарау" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/55/Magnifying_glass_icon.svg/14px-Magnifying_glass_icon.svg.png" decoding="async" width="14" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/55/Magnifying_glass_icon.svg/21px-Magnifying_glass_icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/55/Magnifying_glass_icon.svg/28px-Magnifying_glass_icon.svg.png 2x" data-file-width="490" data-file-height="490" /></a></span> <span typeof="mw:File"><a href="/wiki/%D2%AE%D0%BB%D0%B3%D1%96_%D1%82%D0%B0%D0%BB%D2%9B%D1%8B%D0%BB%D0%B0%D1%83%D1%8B:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%9B%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B_2001-2025" title="Бұл үлгіні талқылау"><img alt="Бұл үлгіні талқылау" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/29/Comment_alt_font_awesome.svg/14px-Comment_alt_font_awesome.svg.png" decoding="async" width="14" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/29/Comment_alt_font_awesome.svg/21px-Comment_alt_font_awesome.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/29/Comment_alt_font_awesome.svg/28px-Comment_alt_font_awesome.svg.png 2x" data-file-width="512" data-file-height="512" /></a></span> <span typeof="mw:File"><a href="//kk.wikipedia.org/w/index.php?title=%D2%AE%D0%BB%D0%B3%D1%96:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%9B%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B_2001-2025&action=edit" title="Бұл үлгіні өңдеу"><img alt="Бұл үлгіні өңдеу" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Template_alt_full_black_22.svg/16px-Template_alt_full_black_22.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Template_alt_full_black_22.svg/24px-Template_alt_full_black_22.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7e/Template_alt_full_black_22.svg/32px-Template_alt_full_black_22.svg.png 2x" data-file-width="160" data-file-height="160" /></a></span></span><div style="font-size:114%">2001—2025 жылдардағы <a href="/wiki/%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B_%D0%BB%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B%D0%BD%D1%8B%D2%A3_%D1%82%D1%96%D0%B7%D1%96%D0%BC%D1%96" title="Физиология немесе медицина саласында Нобель сыйлығы лауреаттарының тізімі">Физиология немесе медицина саласындағы Нобель сыйлығының лауреаттары</a></div></th></tr><tr style="height:2px"><td colspan="2"></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <div> <p><span style="white-space:nowrap;"><a href="/wiki/%D0%9B%D0%B5%D0%BB%D0%B0%D0%BD%D0%B4_%D0%A5%D0%B0%D1%80%D1%82%D0%B2%D0%B5%D0%BB%D0%BB" title="Леланд Хартвелл">Леланд Хартвелл</a> / <a href="/wiki/%D0%A2%D0%B8%D0%BC%D0%BE%D1%82%D0%B8_%D0%A5%D0%B0%D0%BD%D1%82" title="Тимоти Хант">Тимоти Хант</a> / <a href="/wiki/%D0%9F%D0%BE%D0%BB_%D0%9D%D0%B5%D1%80%D1%81" title="Пол Нерс">Пол Нерс</a> (2001) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%A1%D0%B8%D0%B4%D0%BD%D0%B5%D0%B9_%D0%91%D1%80%D0%B5%D0%BD%D0%BD%D0%B5%D1%80" title="Сидней Бреннер">Сидней Бреннер</a> / <a href="/wiki/%D0%A0%D0%BE%D0%B1%D0%B5%D1%80%D1%82_%D0%A5%D0%BE%D1%80%D0%B2%D0%B8%D1%86" title="Роберт Хорвиц">Роберт Хорвиц</a> / <a href="/wiki/%D0%94%D0%B6%D0%BE%D0%BD_%D0%A1%D0%B0%D0%BB%D1%81%D1%82%D0%BE%D0%BD" title="Джон Салстон">Джон Салстон</a> (2002) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%9F%D0%BE%D0%BB_%D0%9B%D0%BE%D1%82%D0%B5%D1%80%D0%B1%D1%83%D1%80" title="Пол Лотербур">Пол Лотербур</a> / <a href="/wiki/%D0%9F%D0%B8%D1%82%D0%B5%D1%80_%D0%9C%D1%8D%D0%BD%D1%81%D1%84%D0%B8%D0%BB%D0%B4" title="Питер Мэнсфилд">Питер Мэнсфилд</a> (2003) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%A0%D0%B8%D1%87%D0%B0%D1%80%D0%B4_%D0%AD%D0%BA%D1%81%D0%B5%D0%BB" title="Ричард Эксел">Ричард Эксел</a> / <a href="/wiki/%D0%9B%D0%B8%D0%BD%D0%B4%D0%B0_%D0%91%D0%B0%D0%BA" title="Линда Бак">Линда Бак</a> (2004) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%91%D0%B0%D1%80%D1%80%D0%B8_%D0%9C%D0%B0%D1%80%D1%88%D0%B0%D0%BB%D0%BB" title="Барри Маршалл">Барри Маршалл</a> / <a href="/wiki/%D0%A0%D0%BE%D0%B1%D0%B8%D0%BD_%D0%A3%D0%BE%D1%80%D1%80%D0%B5%D0%BD" title="Робин Уоррен">Робин Уоррен</a> (2005) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%AD%D0%BD%D0%B4%D1%80%D1%8E_%D0%A4%D0%B0%D0%B9%D0%B5%D1%80" title="Эндрю Файер">Эндрю Файер</a> / <a href="/wiki/%D0%9A%D1%80%D0%B5%D0%B9%D0%B3_%D0%9C%D0%B5%D0%BB%D0%BB%D0%BE" title="Крейг Мелло">Крейг Мелло</a> (2006) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%9C%D0%B0%D1%80%D0%B8%D0%BE_%D0%9A%D0%B0%D0%BF%D0%B5%D0%BA%D0%BA%D0%B8" title="Марио Капекки">Марио Капекки</a> / <a href="/wiki/%D0%9C%D0%B0%D1%80%D1%82%D0%B8%D0%BD_%D0%AD%D0%B2%D0%B0%D0%BD%D1%81" title="Мартин Эванс">Мартин Эванс</a> / <a href="/wiki/%D0%9E%D0%BB%D0%B8%D0%B2%D0%B5%D1%80_%D0%A1%D0%BC%D0%B8%D1%82%D0%B8%D1%81" title="Оливер Смитис">Оливер Смитис</a> (2007) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%A5%D0%B0%D1%80%D0%B0%D0%BB%D1%8C%D0%B4_%D1%86%D1%83%D1%80_%D0%A5%D0%B0%D1%83%D0%B7%D0%B5%D0%BD" title="Харальд цур Хаузен">Харальд цур Хаузен</a> / <a href="/wiki/%D0%9B%D1%8E%D0%BA_%D0%9C%D0%BE%D0%BD%D1%82%D0%B0%D0%BD%D1%8C%D0%B5" title="Люк Монтанье">Люк Монтанье</a> / <a href="/wiki/%D0%A4%D1%80%D0%B0%D0%BD%D1%81%D1%83%D0%B0%D0%B7%D0%B0_%D0%91%D0%B0%D1%80%D1%80%D0%B5-%D0%A1%D0%B8%D0%BD%D1%83%D1%81%D1%81%D0%B8" title="Франсуаза Барре-Синусси">Франсуаза Барре-Синусси</a> (2008) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%AD%D0%BB%D0%B8%D0%B7%D0%B0%D0%B1%D0%B5%D1%82_%D0%91%D0%BB%D1%8D%D0%BA%D0%B1%D0%B5%D1%80%D0%BD" title="Элизабет Блэкберн">Элизабет Блэкберн</a> / <a href="/wiki/%D0%9A%D1%8D%D1%80%D0%BE%D0%BB_%D0%93%D1%80%D0%B5%D0%B9%D0%B4%D0%B5%D1%80" title="Кэрол Грейдер">Кэрол Грейдер</a> / <a href="/wiki/%D0%94%D0%B6%D0%B5%D0%BA_%D0%A8%D0%BE%D1%81%D1%82%D0%B0%D0%BA" title="Джек Шостак">Джек Шостак</a> (2009) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%A0%D0%BE%D0%B1%D0%B5%D1%80%D1%82_%D0%94%D0%B6%D0%B5%D1%84%D1%84%D1%80%D0%B8_%D0%AD%D0%B4%D0%B2%D0%B0%D1%80%D0%B4%D1%81" title="Роберт Джеффри Эдвардс">Роберт Эдвардс</a> (2010) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%91%D1%80%D1%8E%D1%81_%D0%91%D0%B5%D1%82%D0%BB%D0%B5%D1%80" title="Брюс Бетлер">Брюс Бетлер</a> / <a href="/wiki/%D0%96%D1%8E%D0%BB%D1%8C_%D0%A5%D0%BE%D1%84%D1%84%D0%BC%D0%B0%D0%BD" title="Жюль Хоффман">Жюль Хоффман</a> / <a href="/wiki/%D0%A0%D0%B0%D0%BB%D1%8C%D1%84_%D0%A1%D1%82%D0%B0%D0%B9%D0%BD%D0%BC%D0%B0%D0%BD" title="Ральф Стайнман">Ральф Стайнман</a> (2011) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%A1%D0%B8%D0%BD%D1%8A%D1%8F_%D0%AF%D0%BC%D0%B0%D0%BD%D0%B0%D0%BA%D0%B0" title="Синъя Яманака">Синъя Яманака</a> / <a href="/wiki/%D0%94%D0%B6%D0%BE%D0%BD_%D0%93%D0%B5%D1%80%D0%B4%D0%BE%D0%BD" title="Джон Гердон">Джон Гердон</a> (2012) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%94%D0%B6%D0%B5%D0%B9%D0%BC%D1%81_%D0%A0%D0%BE%D1%82%D0%BC%D0%B0%D0%BD" title="Джеймс Ротман">Джеймс Ротман</a> / <a href="/wiki/%D0%A0%D1%8D%D0%BD%D0%B4%D0%B8_%D0%A8%D0%B5%D0%BA%D0%BC%D0%B0%D0%BD" title="Рэнди Шекман">Рэнди Шекман</a> / <a href="/wiki/%D0%A2%D0%BE%D0%BC%D0%B0%D1%81_%D0%9A%D1%80%D0%B8%D1%81%D1%82%D0%B8%D0%B0%D0%BD_%D0%97%D1%8E%D0%B4%D1%85%D0%BE%D1%84" title="Томас Кристиан Зюдхоф">Томас Зюдхоф</a> (2013) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%94%D0%B6%D0%BE%D0%BD_%D0%9E%E2%80%99%D0%9A%D0%B8%D1%84" title="Джон О’Киф">Джон О’Киф</a> / <a href="/wiki/%D0%9C%D0%B5%D0%B9-%D0%91%D1%80%D0%B8%D1%82%D1%82_%D0%9C%D0%BE%D0%B7%D0%B5%D1%80" title="Мей-Бритт Мозер">Мей-Бритт Мозер</a> / <a href="/wiki/%D0%AD%D0%B4%D0%B2%D0%B0%D1%80%D0%B4_%D0%9C%D0%BE%D0%B7%D0%B5%D1%80" title="Эдвард Мозер">Эдвард Мозер</a> (2014) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%A3%D0%B8%D0%BB%D1%8C%D1%8F%D0%BC_%D0%A1%D0%B5%D1%81%D0%B8%D0%BB_%D0%9A%D1%8D%D0%BC%D0%BF%D0%B1%D0%B5%D0%BB%D0%BB" title="Уильям Сесил Кэмпбелл">Уильям Кэмпбелл</a> / <a href="/wiki/%D0%A1%D0%B0%D1%82%D0%BE%D1%88%D0%B8_%D0%9E%D0%BC%D1%83%D1%80%D0%B0" title="Сатоши Омура">Сатоши Омура</a> / <a href="/wiki/%D0%A2%D1%83_%D0%AE%D1%8E" title="Ту Юю">Ту Юю</a> (2015) •</span> <span style="white-space:nowrap;"><a class="mw-selflink selflink">Йошинори Осуми</a> (2016) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%94%D0%B6%D0%B5%D1%84%D1%84%D1%80%D0%B8_%D0%A5%D0%BE%D0%BB%D0%BB" title="Джеффри Холл">Джеффри Холл</a> / <a href="/wiki/%D0%9C%D0%B0%D0%B9%D0%BA%D0%BB_%D0%A0%D0%BE%D1%81%D0%B1%D0%B0%D1%88" title="Майкл Росбаш">Майкл Росбаш</a> / <a href="/wiki/%D0%9C%D0%B0%D0%B9%D0%BA%D0%BB_%D0%AF%D0%BD%D0%B3" title="Майкл Янг">Майкл Янг</a> (2017) •</span> <span style="white-space:nowrap;"><a href="/w/index.php?title=%D0%94%D0%B6%D0%B5%D0%B9%D0%BC%D1%81_%D0%AD%D0%BB%D0%BB%D0%B8%D1%81%D0%BE%D0%BD&action=edit&redlink=1" class="new" title="Джеймс Эллисон (мұндай бет жоқ)">Джеймс Эллисон</a> / <a href="/w/index.php?title=%D0%A2%D0%B0%D1%81%D1%83%D0%BA%D1%83_%D0%A5%D0%BE%D0%BD%D0%B4%D0%B7%D1%91&action=edit&redlink=1" class="new" title="Тасуку Хондзё (мұндай бет жоқ)">Тасуку Хондзё</a> (2018) •</span> <span style="white-space:nowrap;"><a href="/wiki/%D0%A3%D0%B8%D0%BB%D1%8C%D1%8F%D0%BC_%D0%9A%D1%8D%D0%BB%D0%B8%D0%BD" title="Уильям Кэлин">Уильям Кэлин</a> / <a href="/w/index.php?title=%D0%9F%D0%B8%D1%82%D0%B5%D1%80_%D0%A0%D1%8D%D1%82%D0%BA%D0%BB%D0%B8%D1%84%D1%84&action=edit&redlink=1" class="new" title="Питер Рэтклифф (мұндай бет жоқ)">Питер Рэтклифф</a> / <a href="/wiki/%D0%93%D1%80%D0%B5%D0%B3%D0%B3_%D0%A1%D0%B5%D0%BC%D0%B5%D0%BD%D0%B7%D0%B0" title="Грегг Семенза">Грегг Семенза</a> (2019) •</span> <span style="white-space:nowrap;"><a href="/w/index.php?title=%D0%A5%D0%B0%D1%80%D0%B2%D0%B8_%D0%94%D0%B6%D0%B5%D0%B9%D0%BC%D1%81_%D0%90%D0%BB%D1%82%D0%B5%D1%80&action=edit&redlink=1" class="new" title="Харви Джеймс Алтер (мұндай бет жоқ)">Харви Джеймс Алтер</a> / <a href="/w/index.php?title=%D0%9C%D0%B0%D0%B9%D0%BA%D0%BB_%D0%A5%D0%B0%D1%83%D1%82%D0%BE%D0%BD&action=edit&redlink=1" class="new" title="Майкл Хаутон (мұндай бет жоқ)">Майкл Хаутон</a> / <a href="/w/index.php?title=%D0%A7%D0%B0%D1%80%D0%BB%D1%8C%D0%B7_%D0%A0%D0%B0%D0%B9%D1%81&action=edit&redlink=1" class="new" title="Чарльз Райс (мұндай бет жоқ)">Чарльз Райс</a> (2020) •</span> <span style="white-space:nowrap;"><a href="/w/index.php?title=%D0%94%D1%8D%D0%B2%D0%B8%D0%B4_%D0%94%D0%B6%D1%83%D0%BB%D0%B8%D1%83%D1%81&action=edit&redlink=1" class="new" title="Дэвид Джулиус (мұндай бет жоқ)">Дэвид Джулиус</a> / <a href="/w/index.php?title=%D0%90%D1%80%D0%B4%D0%B5%D0%BC_%D0%9F%D0%B0%D1%82%D0%B0%D0%BF%D1%83%D1%82%D1%8F%D0%BD&action=edit&redlink=1" class="new" title="Ардем Патапутян (мұндай бет жоқ)">Ардем Патапутян</a> (2021) •</span> <span style="white-space:nowrap;"><a href="/w/index.php?title=%D0%A1%D0%B2%D0%B0%D0%BD%D1%82%D0%B5_%D0%9F%D0%B0%D0%B0%D0%B1%D0%BE&action=edit&redlink=1" class="new" title="Сванте Паабо (мұндай бет жоқ)">Сванте Паабо</a> (2022) •</span> <span style="white-space:nowrap;"><a href="/w/index.php?title=%D0%9A%D0%B0%D1%82%D0%B0%D0%BB%D0%B8%D0%BD_%D0%9A%D0%B0%D1%80%D0%B8%D0%BA%D0%BE&action=edit&redlink=1" class="new" title="Каталин Карико (мұндай бет жоқ)">Каталин Карико</a> / <a href="/w/index.php?title=%D0%94%D1%80%D1%8E_%D0%92%D0%B0%D0%B9%D1%81%D0%BC%D0%B0%D0%BD&action=edit&redlink=1" class="new" title="Дрю Вайсман (мұндай бет жоқ)">Дрю Вайсман</a> (2023)</span> </p> </div></div></td></tr><tr style="height:2px"><td colspan="2"></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><div> <hr /> <p><a href="/wiki/%D2%AE%D0%BB%D0%B3%D1%96:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B" title="Үлгі:Физиология немесе медицина саласындағы Нобель сыйлығы">Толық тізім</a> | <a href="/wiki/%D2%AE%D0%BB%D0%B3%D1%96:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%9B%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B_1901-1925" title="Үлгі:Физиология немесе медицина саласындағы Нобель сыйлығының Лауреаттары 1901-1925">1901—1925</a> | <a href="/wiki/%D2%AE%D0%BB%D0%B3%D1%96:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%9B%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B_1926-1950" title="Үлгі:Физиология немесе медицина саласындағы Нобель сыйлығының Лауреаттары 1926-1950">1926—1950</a> | <a href="/wiki/%D2%AE%D0%BB%D0%B3%D1%96:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%9B%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B_1951-1975" title="Үлгі:Физиология немесе медицина саласындағы Нобель сыйлығының Лауреаттары 1951-1975">1951—1975</a> | <a href="/wiki/%D2%AE%D0%BB%D0%B3%D1%96:%D0%A4%D0%B8%D0%B7%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F_%D0%BD%D0%B5%D0%BC%D0%B5%D1%81%D0%B5_%D0%BC%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%9B%D0%B0%D1%83%D1%80%D0%B5%D0%B0%D1%82%D1%82%D0%B0%D1%80%D1%8B_1976-2000" title="Үлгі:Физиология немесе медицина саласындағы Нобель сыйлығының Лауреаттары 1976-2000">1976—2000</a> | <b>2001-ден бері</b> </p> </div></div></td></tr></tbody></table></td></tr></tbody></table>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">«<a dir="ltr" href="https://kk.wikipedia.org/w/index.php?title=Йошинори_Осуми&oldid=3362455">https://kk.wikipedia.org/w/index.php?title=Йошинори_Осуми&oldid=3362455</a>» бетінен алынған</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/%D0%90%D1%80%D0%BD%D0%B0%D0%B9%D1%8B:%D0%A1%D0%B0%D0%BD%D0%B0%D1%82%D1%82%D0%B0%D1%80" title="Арнайы:Санаттар">Санаттар</a>: <ul><li><a href="/wiki/%D0%A1%D0%B0%D0%BD%D0%B0%D1%82:%D0%90%D0%BB%D1%84%D0%B0%D0%B2%D0%B8%D1%82_%D0%B1%D0%BE%D0%B9%D1%8B%D0%BD%D1%88%D0%B0_%D1%82%D2%B1%D0%BB%D2%93%D0%B0%D0%BB%D0%B0%D1%80" title="Санат:Алфавит бойынша тұлғалар">Алфавит бойынша тұлғалар</a></li><li><a href="/wiki/%D0%A1%D0%B0%D0%BD%D0%B0%D1%82:%D0%90%D0%BB%D1%84%D0%B0%D0%B2%D0%B8%D1%82_%D0%B1%D0%BE%D0%B9%D1%8B%D0%BD%D1%88%D0%B0_%D2%93%D0%B0%D0%BB%D1%8B%D0%BC%D0%B4%D0%B0%D1%80" title="Санат:Алфавит бойынша ғалымдар">Алфавит бойынша ғалымдар</a></li><li><a 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href="/wiki/%D0%A1%D0%B0%D0%BD%D0%B0%D1%82:%D0%9C%D0%B5%D0%B4%D0%B8%D1%86%D0%B8%D0%BD%D0%B0_%D1%81%D0%B0%D0%BB%D0%B0%D1%81%D1%8B%D0%BD%D0%B4%D0%B0%D2%93%D1%8B_%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%B8%D0%B5%D0%B3%D0%B5%D1%80%D0%BB%D0%B5%D1%80%D1%96" title="Санат:Медицина саласындағы Нобель сыйлығының иегерлері">Медицина саласындағы Нобель сыйлығының иегерлері</a></li><li><a href="/wiki/%D0%A1%D0%B0%D0%BD%D0%B0%D1%82:%D0%9D%D0%BE%D0%B1%D0%B5%D0%BB%D1%8C_%D1%81%D1%8B%D0%B9%D0%BB%D1%8B%D2%93%D1%8B%D0%BD%D1%8B%D2%A3_%D0%B8%D0%B5%D0%B3%D0%B5%D1%80%D0%BB%D0%B5%D1%80%D1%96_(%D0%96%D0%B0%D0%BF%D0%BE%D0%BD%D0%B8%D1%8F)" title="Санат:Нобель сыйлығының иегерлері (Жапония)">Нобель сыйлығының иегерлері (Жапония)</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Жасырын санат: <ul><li><a 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href="https://ar.wikipedia.org/wiki/%D9%8A%D9%88%D8%B4%D9%8A%D9%86%D9%88%D8%B1%D9%8A_%D8%A3%D9%88%D8%B3%D9%88%D9%85%D9%8A" title="يوشينوري أوسومي – араб тілі" lang="ar" hreflang="ar" data-title="يوشينوري أوسومي" data-language-autonym="العربية" data-language-local-name="араб тілі" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-as mw-list-item"><a href="https://as.wikipedia.org/wiki/%E0%A6%AF%E0%A6%BC%E0%A6%B6%E0%A7%8D%E0%A6%AC%E0%A6%BF%E0%A6%A8%E0%A7%B0%E0%A7%80_%E0%A6%85%E0%A6%B9%E0%A6%9A%E0%A7%81%E0%A6%AE%E0%A6%BF" title="য়শ্বিনৰী অহচুমি – ассам тілі" lang="as" hreflang="as" data-title="য়শ্বিনৰী অহচুমি" data-language-autonym="অসমীয়া" data-language-local-name="ассам тілі" class="interlanguage-link-target"><span>অসমীয়া</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Yo%C5%9Finori_Osumi" title="Yoşinori Osumi – әзірбайжан тілі" lang="az" hreflang="az" data-title="Yoşinori Osumi" data-language-autonym="Azərbaycanca" data-language-local-name="әзірбайжан тілі" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اوسومی – South Azerbaijani" lang="azb" hreflang="azb" data-title="یوشینوری اوسومی" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%81%D1%81%D1%96%D0%BD%D0%BE%D1%80%D1%8B_%D0%90%D1%81%D1%83%D0%BC%D1%96" title="Ёсіноры Асумі – беларусь тілі" lang="be" hreflang="be" data-title="Ёсіноры Асумі" data-language-autonym="Беларуская" data-language-local-name="беларусь тілі" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми – болгар тілі" lang="bg" hreflang="bg" data-title="Йошинори Осуми" data-language-autonym="Български" data-language-local-name="болгар тілі" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%87%E0%A6%AF%E0%A6%BC%E0%A7%8B%E0%A6%B6%E0%A6%BF%E0%A6%A8%E0%A7%8B%E0%A6%B0%E0%A6%BF_%E0%A6%93%E0%A6%B8%E0%A7%81%E0%A6%AE%E0%A6%BF" title="ইয়োশিনোরি ওসুমি – бенгал тілі" lang="bn" hreflang="bn" data-title="ইয়োশিনোরি ওসুমি" data-language-autonym="বাংলা" data-language-local-name="бенгал тілі" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – босния тілі" lang="bs" hreflang="bs" data-title="Yoshinori Ohsumi" data-language-autonym="Bosanski" data-language-local-name="босния тілі" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – каталан тілі" lang="ca" hreflang="ca" data-title="Yoshinori Ohsumi" data-language-autonym="Català" data-language-local-name="каталан тілі" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Jo%C5%A1inori_%C3%93sumi" title="Jošinori Ósumi – чех тілі" lang="cs" hreflang="cs" data-title="Jošinori Ósumi" data-language-autonym="Čeština" data-language-local-name="чех тілі" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – дат тілі" lang="da" hreflang="da" data-title="Yoshinori Ohsumi" data-language-autonym="Dansk" data-language-local-name="дат тілі" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – неміс тілі" lang="de" hreflang="de" data-title="Yoshinori Ōsumi" data-language-autonym="Deutsch" data-language-local-name="неміс тілі" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%93%CE%B9%CE%BF%CF%83%CE%B9%CE%BD%CF%8C%CF%81%CE%B9_%CE%9F%CF%87%CF%83%CE%BF%CF%8D%CE%BC%CE%B9" title="Γιοσινόρι Οχσούμι – грек тілі" lang="el" hreflang="el" data-title="Γιοσινόρι Οχσούμι" data-language-autonym="Ελληνικά" data-language-local-name="грек тілі" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – ағылшын тілі" lang="en" hreflang="en" data-title="Yoshinori Ohsumi" data-language-autonym="English" data-language-local-name="ағылшын тілі" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – испан тілі" lang="es" hreflang="es" data-title="Yoshinori Ohsumi" data-language-autonym="Español" data-language-local-name="испан тілі" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – эстон тілі" lang="et" hreflang="et" data-title="Yoshinori Ōsumi" data-language-autonym="Eesti" data-language-local-name="эстон тілі" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – баск тілі" lang="eu" hreflang="eu" data-title="Yoshinori Ohsumi" data-language-autonym="Euskara" data-language-local-name="баск тілі" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اسومی – парсы тілі" lang="fa" hreflang="fa" data-title="یوشینوری اسومی" data-language-autonym="فارسی" data-language-local-name="парсы тілі" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – фин тілі" lang="fi" hreflang="fi" data-title="Yoshinori Ōsumi" data-language-autonym="Suomi" data-language-local-name="фин тілі" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – француз тілі" lang="fr" hreflang="fr" data-title="Yoshinori Ohsumi" data-language-autonym="Français" data-language-local-name="француз тілі" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – галисия тілі" lang="gl" hreflang="gl" data-title="Yoshinori Ōsumi" data-language-autonym="Galego" data-language-local-name="галисия тілі" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%99%D7%95%D7%A9%D7%99%D7%A0%D7%95%D7%A8%D7%99_%D7%90%D7%95%D7%A1%D7%95%D7%9E%D7%99" title="יושינורי אוסומי – иврит тілі" lang="he" hreflang="he" data-title="יושינורי אוסומי" data-language-autonym="עברית" data-language-local-name="иврит тілі" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/%C3%93szumi_Josinori" title="Ószumi Josinori – венгр тілі" lang="hu" hreflang="hu" data-title="Ószumi Josinori" data-language-autonym="Magyar" data-language-local-name="венгр тілі" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%85%D5%B8%D5%BD%D5%AB%D5%B6%D5%B8%D6%80%D5%AB_%D5%95%D5%BD%D5%B8%D6%82%D5%B4%D5%AB" title="Յոսինորի Օսումի – армян тілі" lang="hy" hreflang="hy" data-title="Յոսինորի Օսումի" data-language-autonym="Հայերեն" data-language-local-name="армян тілі" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – индонезия тілі" lang="id" hreflang="id" data-title="Yoshinori Ohsumi" data-language-autonym="Bahasa Indonesia" data-language-local-name="индонезия тілі" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – идо тілі" lang="io" hreflang="io" data-title="Yoshinori Ohsumi" data-language-autonym="Ido" data-language-local-name="идо тілі" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it badge-Q17437798 badge-goodarticle mw-list-item" title="жақсы мақалалар"><a href="https://it.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – итальян тілі" lang="it" hreflang="it" data-title="Yoshinori Ōsumi" data-language-autonym="Italiano" data-language-local-name="итальян тілі" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 – жапон тілі" lang="ja" hreflang="ja" data-title="大隅良典" data-language-autonym="日本語" data-language-local-name="жапон тілі" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%AF%E0%B3%8A%E0%B2%B6%E0%B2%BF%E0%B2%A8%E0%B3%8B%E0%B2%B0%E0%B2%BF_%E0%B2%93%E0%B2%B6%E0%B3%81%E0%B2%AE%E0%B2%BF" title="ಯೊಶಿನೋರಿ ಓಶುಮಿ – каннада тілі" lang="kn" hreflang="kn" data-title="ಯೊಶಿನೋರಿ ಓಶುಮಿ" data-language-autonym="ಕನ್ನಡ" data-language-local-name="каннада тілі" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%98%A4%EC%8A%A4%EB%AF%B8_%EC%9A%94%EC%8B%9C%EB%85%B8%EB%A6%AC" title="오스미 요시노리 – корей тілі" lang="ko" hreflang="ko" data-title="오스미 요시노리" data-language-autonym="한국어" data-language-local-name="корей тілі" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – латын тілі" lang="la" hreflang="la" data-title="Yoshinori Ohsumi" data-language-autonym="Latina" data-language-local-name="латын тілі" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%88%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Јошинори Осуми – македон тілі" lang="mk" hreflang="mk" data-title="Јошинори Осуми" data-language-autonym="Македонски" data-language-local-name="македон тілі" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%AF%E0%B5%8B%E0%B4%B7%E0%B4%BF%E0%B4%A8%E0%B5%8B%E0%B4%B1%E0%B4%BF_%E0%B4%89%E0%B4%B7%E0%B5%81%E0%B4%AE%E0%B4%BF" title="യോഷിനോറി ഉഷുമി – малаялам тілі" lang="ml" hreflang="ml" data-title="യോഷിനോറി ഉഷുമി" data-language-autonym="മലയാളം" data-language-local-name="малаялам тілі" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – малай тілі" lang="ms" hreflang="ms" data-title="Yoshinori Ohsumi" data-language-autonym="Bahasa Melayu" data-language-local-name="малай тілі" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اسومی – мазандеран тілі" lang="mzn" hreflang="mzn" data-title="یوشینوری اسومی" data-language-autonym="مازِرونی" data-language-local-name="мазандеран тілі" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – төменгі неміс тілі" lang="nds" hreflang="nds" data-title="Yoshinori Ōsumi" data-language-autonym="Plattdüütsch" data-language-local-name="төменгі неміс тілі" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-ne mw-list-item"><a href="https://ne.wikipedia.org/wiki/%E0%A4%AF%E0%A5%8B%E0%A4%B8%E0%A4%BF%E0%A4%A8%E0%A5%8B%E0%A4%B0%E0%A5%80_%E0%A4%93%E0%A4%B8%E0%A5%81%E0%A4%AE%E0%A5%80" title="योसिनोरी ओसुमी – непал тілі" lang="ne" hreflang="ne" data-title="योसिनोरी ओसुमी" data-language-autonym="नेपाली" data-language-local-name="непал тілі" class="interlanguage-link-target"><span>नेपाली</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Yoshinori_Osumi" title="Yoshinori Osumi – нидерланд тілі" lang="nl" hreflang="nl" data-title="Yoshinori Osumi" data-language-autonym="Nederlands" data-language-local-name="нидерланд тілі" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – норвегиялық букмол тілі" lang="nb" hreflang="nb" data-title="Yoshinori Ohsumi" data-language-autonym="Norsk bokmål" data-language-local-name="норвегиялық букмол тілі" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – окситан тілі" lang="oc" hreflang="oc" data-title="Yoshinori Ōsumi" data-language-autonym="Occitan" data-language-local-name="окситан тілі" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pa mw-list-item"><a href="https://pa.wikipedia.org/wiki/%E0%A8%AF%E0%A9%8B%E0%A8%B8%E0%A8%BC%E0%A9%80%E0%A8%A8%E0%A9%8B%E0%A8%B0%E0%A9%80_%E0%A8%93%E0%A8%B8%E0%A9%81%E0%A8%AE%E0%A9%80" title="ਯੋਸ਼ੀਨੋਰੀ ਓਸੁਮੀ – пенджаб тілі" lang="pa" hreflang="pa" data-title="ਯੋਸ਼ੀਨੋਰੀ ਓਸੁਮੀ" data-language-autonym="ਪੰਜਾਬੀ" data-language-local-name="пенджаб тілі" class="interlanguage-link-target"><span>ਪੰਜਾਬੀ</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – поляк тілі" lang="pl" hreflang="pl" data-title="Yoshinori Ōsumi" data-language-autonym="Polski" data-language-local-name="поляк тілі" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%DB%81%D8%B3%D9%85%DB%8C" title="یوشینوری اوہسمی – Western Punjabi" lang="pnb" hreflang="pnb" data-title="یوشینوری اوہسمی" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – португал тілі" lang="pt" hreflang="pt" data-title="Yoshinori Ohsumi" data-language-autonym="Português" data-language-local-name="португал тілі" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – румын тілі" lang="ro" hreflang="ro" data-title="Yoshinori Ohsumi" data-language-autonym="Română" data-language-local-name="румын тілі" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9E%D1%81%D1%83%D0%BC%D0%B8,_%D0%81%D1%81%D0%B8%D0%BD%D0%BE%D1%80%D0%B8" title="Осуми, Ёсинори – орыс тілі" lang="ru" hreflang="ru" data-title="Осуми, Ёсинори" data-language-autonym="Русский" data-language-local-name="орыс тілі" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – шотланд тілі" lang="sco" hreflang="sco" data-title="Yoshinori Ohsumi" data-language-autonym="Scots" data-language-local-name="шотланд тілі" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi – серб-хорват тілі" lang="sh" hreflang="sh" data-title="Yoshinori Ōsumi" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="серб-хорват тілі" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-si mw-list-item"><a href="https://si.wikipedia.org/wiki/%E0%B6%BA%E0%B7%9D%E0%B7%82%E0%B7%92%E0%B6%B1%E0%B7%9C%E0%B6%BB%E0%B7%92_%E0%B6%94%E0%B7%83%E0%B7%94%E0%B6%B8%E0%B7%92" title="යෝෂිනොරි ඔසුමි – сингал тілі" lang="si" hreflang="si" data-title="යෝෂිනොරි ඔසුමි" data-language-autonym="සිංහල" data-language-local-name="сингал тілі" class="interlanguage-link-target"><span>සිංහල</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – Simple English" lang="en-simple" hreflang="en-simple" data-title="Yoshinori Ohsumi" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Jo%C5%A1inori_Ohsumi" title="Jošinori Ohsumi – словен тілі" lang="sl" hreflang="sl" data-title="Jošinori Ohsumi" data-language-autonym="Slovenščina" data-language-local-name="словен тілі" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – албан тілі" lang="sq" hreflang="sq" data-title="Yoshinori Ohsumi" data-language-autonym="Shqip" data-language-local-name="албан тілі" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%88%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Јошинори Осуми – серб тілі" lang="sr" hreflang="sr" data-title="Јошинори Осуми" data-language-autonym="Српски / srpski" data-language-local-name="серб тілі" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – швед тілі" lang="sv" hreflang="sv" data-title="Yoshinori Ohsumi" data-language-autonym="Svenska" data-language-local-name="швед тілі" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – суахили тілі" lang="sw" hreflang="sw" data-title="Yoshinori Ohsumi" data-language-autonym="Kiswahili" data-language-local-name="суахили тілі" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%87%E0%AE%AF%E0%AF%8B%E0%AE%9A%E0%AE%BF%E0%AE%A9%E0%AF%8B%E0%AE%B0%E0%AE%BF_%E0%AE%93%E0%AE%9A%E0%AF%82%E0%AE%AE%E0%AE%BF" title="இயோசினோரி ஓசூமி – тамил тілі" lang="ta" hreflang="ta" data-title="இயோசினோரி ஓசூமி" data-language-autonym="தமிழ்" data-language-local-name="тамил тілі" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a href="https://te.wikipedia.org/wiki/%E0%B0%AF%E0%B1%8B%E0%B0%B7%E0%B0%BF%E0%B0%A8%E0%B1%8B%E0%B0%B0%E0%B0%BF_%E0%B0%93%E0%B0%B9%E0%B1%8D%E0%B0%B8%E0%B1%81%E0%B0%AE%E0%B1%80" title="యోషినోరి ఓహ్సుమీ – телугу тілі" lang="te" hreflang="te" data-title="యోషినోరి ఓహ్సుమీ" data-language-autonym="తెలుగు" data-language-local-name="телугу тілі" class="interlanguage-link-target"><span>తెలుగు</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B9%82%E0%B8%A2%E0%B8%8A%E0%B8%B4%E0%B9%82%E0%B8%99%E0%B8%A3%E0%B8%B4_%E0%B9%82%E0%B8%AD%E0%B8%8B%E0%B8%B9%E0%B8%A1%E0%B8%B4" title="โยชิโนริ โอซูมิ – тай тілі" lang="th" hreflang="th" data-title="โยชิโนริ โอซูมิ" data-language-autonym="ไทย" data-language-local-name="тай тілі" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi – түрік тілі" lang="tr" hreflang="tr" data-title="Yoshinori Ohsumi" data-language-autonym="Türkçe" data-language-local-name="түрік тілі" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми – татар тілі" lang="tt" hreflang="tt" data-title="Йошинори Осуми" data-language-autonym="Татарча / tatarça" data-language-local-name="татар тілі" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9E%D1%81%D1%83%D0%BC%D1%96_%D0%99%D0%BE%D1%81%D1%96%D0%BD%D0%BE%D1%80%D1%96" title="Осумі Йосінорі – украин тілі" lang="uk" hreflang="uk" data-title="Осумі Йосінорі" data-language-autonym="Українська" data-language-local-name="украин тілі" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%D8%B4%D9%88%D9%85%DB%8C" title="یوشینوری اوشومی – урду тілі" lang="ur" hreflang="ur" data-title="یوشینوری اوشومی" data-language-autonym="اردو" data-language-local-name="урду тілі" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/%C5%8Csumi_Yoshinori" title="Ōsumi Yoshinori – вьетнам тілі" lang="vi" hreflang="vi" data-title="Ōsumi Yoshinori" data-language-autonym="Tiếng Việt" data-language-local-name="вьетнам тілі" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 – қытай тілі (У)" lang="wuu" hreflang="wuu" data-title="大隅良典" data-language-autonym="吴语" data-language-local-name="қытай тілі (У)" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 – қытай тілі" lang="zh" hreflang="zh" data-title="大隅良典" data-language-autonym="中文" data-language-local-name="қытай тілі" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/%C3%94sumi_Yosinori" title="Ôsumi Yosinori – Minnan" lang="nan" hreflang="nan" data-title="Ôsumi Yosinori" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q8056214#sitelinks-wikipedia" title="Тіларалық сілтемелерді өңдеу" class="wbc-editpage">Сілтемелерді өңдеу</a></span></div> </div> </nav> </div> </div> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> Бұл беттің соңғы рет өзгертілген кезі: 2024 ж. тамыздың 3, 08:10.</li> <li id="footer-info-copyright">Мәтін <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/">Creative Commons 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