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Evolution of emotion - Wikipedia

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.sidebar-title-with-pretitle{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .sidebar:not(.notheme) .sidebar-title-with-pretitle a{color:var(--color-progressive)!important}}@media print{body.ns-0 .mw-parser-output .sidebar{display:none!important}}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><table class="sidebar sidebar-collapse nomobile nowraplinks"><tbody><tr><td class="sidebar-pretitle">Part of <a href="/wiki/Category:Emotions" title="Category:Emotions">a series</a> on</td></tr><tr><th class="sidebar-title-with-pretitle"><a href="/wiki/Emotion" title="Emotion">Emotions</a></th></tr><tr><td class="sidebar-image"><span class="skin-invert-image" typeof="mw:File"><a href="/wiki/File:Plutchik-wheel.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Plutchik-wheel.svg/90px-Plutchik-wheel.svg.png" decoding="async" width="90" height="91" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Plutchik-wheel.svg/135px-Plutchik-wheel.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Plutchik-wheel.svg/180px-Plutchik-wheel.svg.png 2x" data-file-width="715" data-file-height="725" /></a></span><span class="skin-invert-image" typeof="mw:File"><a href="/wiki/File:Plutchik_Dyads.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Plutchik_Dyads.svg/90px-Plutchik_Dyads.svg.png" decoding="async" width="90" height="90" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Plutchik_Dyads.svg/135px-Plutchik_Dyads.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Plutchik_Dyads.svg/180px-Plutchik_Dyads.svg.png 2x" data-file-width="1357" data-file-height="1356" /></a></span></td></tr><tr><td class="sidebar-content"> <div class="hlist"> <ul><li><a href="/wiki/Affect_(psychology)" title="Affect (psychology)">Affect</a></li> <li><a href="/wiki/Emotion_classification" title="Emotion classification">Classification</a></li> <li><a href="/wiki/Emotion_in_animals" title="Emotion in animals">In animals</a></li> <li><a href="/wiki/Emotional_intelligence" title="Emotional intelligence">Emotional intelligence</a></li> <li><a href="/wiki/Mood_(psychology)" title="Mood (psychology)">Mood</a></li> <li><br /><a href="/wiki/Emotional_self-regulation" title="Emotional self-regulation">Self-regulation</a> <ul><li><a href="/wiki/Interpersonal_emotion_regulation" title="Interpersonal emotion regulation">Interpersonal</a></li> <li><a href="/wiki/Emotional_dysregulation" title="Emotional dysregulation">Dysregulation</a></li></ul></li> <li><a href="/wiki/Valence_(psychology)" title="Valence (psychology)">Valence</a></li></ul> </div></td> </tr><tr><td class="sidebar-content"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="color: var(--color-base)"><div class="sidebar-list-title-c">Emotions</div></div><div class="sidebar-list-content mw-collapsible-content"><div class="hlist"> <ul><li><a href="/wiki/Acceptance" title="Acceptance">Acceptance</a></li> <li><a href="/wiki/Admiration" title="Admiration">Admiration</a></li> <li><a href="/wiki/Affection" title="Affection">Affection</a></li> <li><a href="/wiki/Amusement" title="Amusement">Amusement</a></li> <li><a href="/wiki/Anger" title="Anger">Anger</a></li> <li><a href="/wiki/Angst" title="Angst">Angst</a></li> <li><a href="/wiki/Anguish" title="Anguish">Anguish</a></li> <li><a href="/wiki/Annoyance" title="Annoyance">Annoyance</a></li> <li><a href="/wiki/Anticipation" title="Anticipation">Anticipation</a></li> <li><a href="/wiki/Anxiety" title="Anxiety">Anxiety</a></li> <li><a href="/wiki/Apathy" title="Apathy">Apathy</a></li> <li><a href="/wiki/Arousal" title="Arousal">Arousal</a></li> <li><a href="/wiki/Awe" title="Awe">Awe</a></li> <li><a href="/wiki/Belongingness" title="Belongingness">Belongingness</a></li> <li><a href="/wiki/Boredom" title="Boredom">Boredom</a></li> <li><a href="/wiki/Confidence" title="Confidence">Confidence</a></li> <li><a href="/wiki/Confusion" title="Confusion">Confusion</a></li> <li><a href="/wiki/Contempt" title="Contempt">Contempt</a></li> <li><a href="/wiki/Contentment" title="Contentment">Contentment</a></li> <li><a href="/wiki/Courage" title="Courage">Courage</a></li> <li><a href="/wiki/Curiosity" title="Curiosity">Curiosity</a></li> <li><a href="/wiki/Depression_(mood)" title="Depression (mood)">Depression</a></li> <li><a href="/wiki/Desire" title="Desire">Desire</a></li> <li><a href="/wiki/Determination" title="Determination">Determination</a></li> <li><a href="/wiki/Disappointment" title="Disappointment">Disappointment</a></li> <li><a href="/wiki/Disgust" title="Disgust">Disgust</a></li> <li><a href="/wiki/Distrust" title="Distrust">Distrust</a></li> <li><a href="/wiki/Doubt" title="Doubt">Doubt</a></li> <li><a href="/wiki/Ecstasy_(emotion)" title="Ecstasy (emotion)">Ecstasy</a></li> <li><a href="/wiki/Elevation_(emotion)" title="Elevation (emotion)">Elevation</a></li> <li><a href="/wiki/Embarrassment" title="Embarrassment">Embarrassment</a></li> <li><a href="/wiki/Emotional_detachment" title="Emotional detachment">Emotional Detachment</a></li> <li><a href="/wiki/Empathy" title="Empathy">Empathy</a></li> <li><a href="/wiki/Enthusiasm" title="Enthusiasm">Enthusiasm</a></li> <li><a href="/wiki/Envy" title="Envy">Envy</a></li> <li><a href="/wiki/Euphoria" title="Euphoria">Euphoria</a></li> <li><a href="/wiki/Faith" title="Faith">Faith</a></li> <li><a href="/wiki/Fear" title="Fear">Fear</a></li> <li><a href="/wiki/Frustration" title="Frustration">Frustration</a></li> <li><a href="/wiki/Gratification" title="Gratification">Gratification</a></li> <li><a href="/wiki/Gratitude" title="Gratitude">Gratitude</a></li> <li><a href="/wiki/Greed" title="Greed">Greed</a></li> <li><a href="/wiki/Grief" title="Grief">Grief</a></li> <li><a href="/wiki/Guilt_(emotion)" title="Guilt (emotion)">Guilt</a></li> <li><a href="/wiki/Happiness" title="Happiness">Happiness</a></li> <li><a href="/wiki/Hatred" title="Hatred">Hatred</a></li> <li><a href="/wiki/Hope" title="Hope">Hope</a></li> <li><a href="/wiki/Horror_and_terror" title="Horror and terror">Horror</a></li> <li><a href="/wiki/Hostility" title="Hostility">Hostility</a></li> <li><a href="/wiki/Hubris" title="Hubris">Hubris</a></li> <li><a href="/wiki/Humiliation" title="Humiliation">Humiliation</a></li> <li><a href="/wiki/Interest_(emotion)" title="Interest (emotion)">Interest</a></li> <li><a href="/wiki/Jealousy" title="Jealousy">Jealousy</a></li> <li><a href="/wiki/Joy" title="Joy">Joy</a></li> <li><a href="/wiki/Kindness" title="Kindness">Kindness</a></li> <li><a href="/wiki/Limerence" title="Limerence">Limerence</a></li> <li><a href="/wiki/Loneliness" title="Loneliness">Loneliness</a></li> <li><a href="/wiki/Love" title="Love">Love</a></li> <li><a href="/wiki/Lust" title="Lust">Lust</a></li> <li><a href="/wiki/Nostalgia" title="Nostalgia">Nostalgia</a></li> <li><a href="/wiki/Outrage_(emotion)" title="Outrage (emotion)">Outrage</a></li> <li><a href="/wiki/Panic" title="Panic">Panic</a></li> <li><a href="/wiki/Passion_(emotion)" title="Passion (emotion)">Passion</a></li> <li><a href="/wiki/Pity" title="Pity">Pity</a></li> <li><a href="/wiki/Pleasure" title="Pleasure">Pleasure</a></li> <li><a href="/wiki/Pride" title="Pride">Pride</a></li> <li><a href="/wiki/Rage_(emotion)" title="Rage (emotion)">Rage</a></li> <li><a href="/wiki/Regret" title="Regret">Regret</a></li> <li><a href="/wiki/Social_rejection" title="Social rejection">Rejection</a></li> <li><a href="/wiki/Relief_(emotion)" title="Relief (emotion)">Relief</a></li> <li><a href="/wiki/Remorse" title="Remorse">Remorse</a></li> <li><a href="/wiki/Resentment" title="Resentment">Resentment</a></li> <li><a href="/wiki/Sadness" title="Sadness">Sadness</a></li> <li><a href="/wiki/Saudade" title="Saudade">Saudade</a></li> <li><a href="/wiki/Schadenfreude" title="Schadenfreude">Schadenfreude</a></li> <li><a href="/wiki/Self-pity" title="Self-pity">Self-pity</a></li> <li><a href="/wiki/Shame" title="Shame">Shame</a></li> <li><a href="/wiki/Acute_stress_reaction" title="Acute stress reaction">Shock</a></li> <li><a href="/wiki/Shyness" title="Shyness">Shyness</a></li> <li><a href="/wiki/Social_connection" title="Social connection">Social connection</a></li> <li><a href="/wiki/Sorrow_(emotion)" title="Sorrow (emotion)">Sorrow</a></li> <li><a href="/wiki/Suffering" title="Suffering">Suffering</a></li> <li><a href="/wiki/Surprise_(emotion)" title="Surprise (emotion)">Surprise</a></li> <li><a href="/wiki/Suspicion_(emotion)" title="Suspicion (emotion)">Suspicion</a></li> <li><a href="/wiki/Trust_(social_science)" title="Trust (social science)">Trust</a></li> <li><a href="/wiki/Wonder_(emotion)" title="Wonder (emotion)">Wonder</a></li> <li><a href="/wiki/Worry" title="Worry">Worry</a></li></ul> </div></div></div></td> </tr><tr><td class="sidebar-navbar"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Emotion_sidebar" title="Template:Emotion sidebar"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Emotion_sidebar" title="Template talk:Emotion sidebar"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Emotion_sidebar" title="Special:EditPage/Template:Emotion sidebar"><abbr title="Edit this template">e</abbr></a></li></ul></div></td></tr></tbody></table> <p>The study of the <b>evolution of emotions</b> dates back to the 19th century. <a href="/wiki/Evolution" title="Evolution">Evolution</a> and <a href="/wiki/Natural_selection" title="Natural selection">natural selection</a> has been applied to the study of <a href="/wiki/Human_communication" title="Human communication">human communication</a>, mainly by <a href="/wiki/Charles_Darwin" title="Charles Darwin">Charles Darwin</a> in his 1872 work, <i><a href="/wiki/The_Expression_of_the_Emotions_in_Man_and_Animals" title="The Expression of the Emotions in Man and Animals">The Expression of the Emotions in Man and Animals</a></i>.<sup id="cite_ref-Darwin2_1-0" class="reference"><a href="#cite_note-Darwin2-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> Darwin researched the expression of <a href="/wiki/Emotion" title="Emotion">emotions</a> in an effort to support his materialist theory of unguided evolution. He proposed that much like other <a href="/wiki/Phenotypic_trait" title="Phenotypic trait">traits</a> found in <a href="/wiki/Animal" title="Animal">animals</a>, emotions apparently also evolved and were adapted over time.<sup id="cite_ref-Hess_2-0" class="reference"><a href="#cite_note-Hess-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> His work looked at not only <a href="/wiki/Facial_expression" title="Facial expression">facial expressions</a> in <a href="/wiki/Animals" class="mw-redirect" title="Animals">animals</a> and specifically <a href="/wiki/Human" title="Human">humans</a>, but attempted to point out parallels between <a href="/wiki/Behavior" title="Behavior">behaviors</a> in humans and other animals. </p><p>According to evolutionary theory, different emotions evolved at different times. Primal emotions, such as <a href="/wiki/Love" title="Love">love</a> and <a href="/wiki/Fear" title="Fear">fear</a>, are associated with ancient parts of the psyche. Social emotions, such as <a href="/wiki/Guilt_(emotion)" title="Guilt (emotion)">guilt</a> and <a href="/wiki/Pride" title="Pride">pride</a>, evolved among social <a href="/wiki/Primate" title="Primate">primates</a>. <a href="/wiki/Evolutionary_psychology" title="Evolutionary psychology">Evolutionary psychologists</a> consider human emotions to be best adapted to the life our ancestors led in <a href="/wiki/Nomad" title="Nomad">nomadic</a> foraging bands. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Origins">Origins</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Evolution_of_emotion&amp;action=edit&amp;section=1" title="Edit section: Origins"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Darwin's original plan was to include his findings about expression of emotions in a chapter of his work, <i><a href="/wiki/The_Descent_of_Man" class="mw-redirect" title="The Descent of Man">The Descent of Man, and Selection in Relation to Sex</a></i> (Darwin, 1871) but found that he had enough material for a whole book. It was based on <a href="/wiki/Observations" class="mw-redirect" title="Observations">observations</a>, both those around him and of people in many parts of the world. One important observation he made was that even in individuals who were born <a href="/wiki/Blindness" class="mw-redirect" title="Blindness">blind</a>, body and facial expressions displayed are similar to those of anyone else. The ideas found in his book on <a href="/wiki/Universality_(philosophy)" title="Universality (philosophy)">universality</a> of emotions were intended to go against Sir <a href="/wiki/Charles_Bell" title="Charles Bell">Charles Bell's</a> 1844 claim<sup id="cite_ref-Bell_3-0" class="reference"><a href="#cite_note-Bell-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> that human facial <a href="/wiki/Muscles" class="mw-redirect" title="Muscles">muscles</a> were created to give them the unique ability to express emotions.<sup id="cite_ref-Hess_2-1" class="reference"><a href="#cite_note-Hess-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> The main purpose of Darwin's work was to support the theory of evolution by demonstrating that emotions in humans and other animals are similar. Most of the similarities he found were between <a href="/wiki/Species" title="Species">species</a> closely related, but he found some similarities between distantly related species as well. He proposed the idea that emotional states are adaptive, and therefore only those able to express certain emotions passed on their characteristics.<sup id="cite_ref-Darwin2_1-1" class="reference"><a href="#cite_note-Darwin2-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Darwin's_principles"><span id="Darwin.27s_principles"></span>Darwin's principles</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Evolution_of_emotion&amp;action=edit&amp;section=2" title="Edit section: Darwin&#039;s principles"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In the 1872 work, Darwin proposed three principles. The first of the three is the "principle of serviceable habits", which he defined as we have certain habits or we perform different actions in certain states of mind, which get associated when that state of mind is induced, even when its not needed then. He used as an example contracting of eyebrows (furrowing the brow), which he noted is serviceable to prevent too much light from entering the eyes. He also said that the raising of eyebrows serves to increase the <a href="/wiki/Field_of_vision" class="mw-redirect" title="Field of vision">field of vision</a>. He cited examples of people attempting to remember something and raising their brows, as though they could "see" what they were trying to remember. </p><p>The second of the principles is that of <a href="/wiki/Antithesis" title="Antithesis">antithesis</a>. While some habits are serviceable, Darwin proposed that some actions or habits are carried out merely because they are opposite in nature to a serviceable habit, but are not serviceable themselves. Shrugging of the shoulders is an example Darwin used of antithesis, because it has no service. Shoulder shrugging is a passive expression, and very opposite of a confident or aggressive expression.<sup id="cite_ref-Darwin2_1-2" class="reference"><a href="#cite_note-Darwin2-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> </p><p>The third of the principles is expressive habits, or nervous discharge from the nervous system. This principle proposes that some habits are performed because of a build-up to the <a href="/wiki/Nervous_system" title="Nervous system">nervous system</a>, which causes a <a href="https://en.wiktionary.org/wiki/discharge" class="extiw" title="wikt:discharge">discharge</a> of the <a href="/wiki/Psychomotor_agitation" title="Psychomotor agitation">excitement</a>. Examples include foot and finger tapping, as well as vocal expressions and expressions of <a href="/wiki/Anger" title="Anger">anger</a>. Darwin noted that many animals rarely make noises, even when in <a href="/wiki/Pain" title="Pain">pain</a>, but under extreme circumstances they vocalize in response to pain and <a href="/wiki/Fear" title="Fear">fear</a>.<sup id="cite_ref-Darwin2_1-3" class="reference"><a href="#cite_note-Darwin2-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Research">Research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Evolution_of_emotion&amp;action=edit&amp;section=3" title="Edit section: Research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Emotion_in_animals" title="Emotion in animals">Emotion in animals</a></div> <p><a href="/wiki/Paul_Ekman" title="Paul Ekman">Paul Ekman</a> is most noted in this field for conducting research involving facial expressions of emotions. His work provided data to back up Darwin's ideas about universality of facial expressions, even across cultures. He conducted research by showing photographs exhibiting expressions of basic emotion to people and asking them to identify what emotion was being expressed. In 1971, Ekman and Wallace Friesen presented to people in a preliterate culture a story involving a certain emotion, along with photographs of specific facial expressions. The photographs had been previously used in studies using subjects from <a href="/wiki/Western_cultures" class="mw-redirect" title="Western cultures">Western cultures</a>. When asked to choose, from two or three photographs, the emotion being expressed in the story, the preliterate subjects' choices matched those of the Western subjects most of the time. These results indicated that certain expressions are universally associated with particular emotions, even in instances in which the people had little or no exposure to Western culture. The only emotions the preliterate people found hard to distinguish between were fear and <a href="/wiki/Surprise_(emotion)" title="Surprise (emotion)">surprise</a>.<sup id="cite_ref-Friesen_4-0" class="reference"><a href="#cite_note-Friesen-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> Ekman noted that while universal expressions do not necessarily prove Darwin's theory that they evolved, they do provide strong evidence of the possibility.<sup id="cite_ref-Ekman_1993_5-0" class="reference"><a href="#cite_note-Ekman_1993-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> He mentioned the similarities between human expressions and those of other <a href="/wiki/Primates" class="mw-redirect" title="Primates">primates</a>, as well as an overall universality of certain expressions to back up Darwin's ideas. The expressions of emotion that Ekman noted as most universal based on research are: anger, fear, <a href="/wiki/Disgust" title="Disgust">disgust</a>, <a href="/wiki/Sadness" title="Sadness">sadness</a>, and <a href="/wiki/Enjoyment" class="mw-redirect" title="Enjoyment">enjoyment</a>.<sup id="cite_ref-Ekman_1993_5-1" class="reference"><a href="#cite_note-Ekman_1993-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> </p><p>A common view is that facial expressions initially served a non-communicative <a href="/wiki/Adaptation" title="Adaptation">adaptive</a> function. Thus, the widened eyes in the facial expression of <a href="/wiki/Fear" title="Fear">fear</a> have been shown to increase the visual field and the speed of moving the eyes which helps finding and following threats. The wrinkled nose and mouth of the facial expression of <a href="/wiki/Disgust" title="Disgust">disgust</a> limit the intake of foul-smelling and possibly dangerous air and particles. Later, such reactions, which could be observed by other members of the group, increasingly become more distinctive and exaggerated in order to fulfill a primarily socially communicative function. This communicative function can dramatically or subtly influence the behavior of other members in the group. Thus, rhesus monkeys or human infants can learn to fear potential dangers based on only the facial expressions of fear of other group members or parents. Seeing fear expressions increases the tendency for flight responses while seeing anger expressions increases the tendency for fight responses. <a href="/wiki/Classical_conditioning" title="Classical conditioning">Classical conditioning</a> studies have found that it is easier to create a pairing between a negative stimulant and anger/fear expressions than between a negative stimulant and a happiness expression. Cross-cultural studies and studies on the congenitally <a href="/wiki/Blindness" class="mw-redirect" title="Blindness">blind</a> have found that these groups display the same expressions of shame and pride in situations related to social status. These expressions have clear similarities to displays of submission and dominance by other primates. Humans viewing expression of pride automatically assign a higher social status to such individuals than to those expressing other emotions.<sup id="cite_ref-Shariff2011_6-0" class="reference"><a href="#cite_note-Shariff2011-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> </p> <table class="wikitable"> <caption>Expressed emotions and adaptive functions<sup id="cite_ref-Shariff2011_6-1" class="reference"><a href="#cite_note-Shariff2011-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> </caption> <tbody><tr> <th>Expressed emotion </th> <th>Initial physiological function </th> <th>Evolved communicative function </th></tr> <tr> <td><a href="/wiki/Fear" title="Fear">Fear</a> </td> <td>Increased visual field and speed of eye movement from widened eyes. Sensory hypersensitization (e.g. hearing and touch). Substantial adrenal response (e.g. Vomiting and bodily fluid evacuation). </td> <td>Fight-or-flight (posturing, thanatosis, hostility, etc). Eusocial alerting to potential threats. Appeasement to peer aggressor. Hindered or shutdown learning postures (loops close in favor of survival). </td></tr> <tr> <td><a href="/wiki/Surprise_(emotion)" title="Surprise (emotion)">Surprise</a> </td> <td>Increased visual field from widened eyes, decreased stimulus sharpness. Hormonal shifts (e.g. noradrenaline). </td> <td>Improved (receptive) learning postures and modified perception filtering. </td></tr> <tr> <td><a href="/wiki/Disgust" title="Disgust">Disgust</a> </td> <td>Constriction of head openings reduce dangerous inhalations and ingestions, reduced blood flow and digestion. Nausea and dizziness, vomiting and bodily fluid evacuation. </td> <td>Warning of dangerous foods (in particular poisons) and stimulus changes. Psychogenic vomiting. Unable to assume learning postures. </td></tr> <tr> <td><a href="/wiki/Happiness" title="Happiness">Happiness</a> </td> <td>Vascularization and hormonal rebalancing (e.g. Oxytocin, dopamine, and serotonin shifts). Wide range of physiological signaling (e.g. innarticulate grunts such as laughter). </td> <td>Absence of threat, Reinforced learning postures. Nurturing. </td></tr> <tr> <td><a href="/wiki/Sadness" title="Sadness">Sadness</a> </td> <td>Tears, disphoria, mirror behavior desensitization, avoidance behavior. </td> <td>Vision handicapped by tears to show appeasement and evoke juvenile care taking behaviors in vertebates (engage peer sympathy). Downregulated or shutdown learning postures. </td></tr> <tr> <td><a href="/wiki/Anger" title="Anger">Anger</a> </td> <td>Primed adrenal responses (pupil dilation, pain desensitization, sweat, gastro-intestinal compartization, hostility) </td> <td>Warning of impending threats. Dominant posturing. Adversarial learning postures. </td></tr> <tr> <td><a href="/wiki/Pride" title="Pride">Pride</a> </td> <td>Increased lung volume in preparation for encountering challengers. Arousal. </td> <td>Increased social status posturing and mating attempts. </td></tr> <tr> <td><a href="/wiki/Shame" title="Shame">Shame</a>/<a href="/wiki/Embarrassment" title="Embarrassment">Embarrassment</a> </td> <td>Reduces and hides vulnerable body areas from potential attacks, arousal downregulation. </td> <td>Decreased social status posturing and mating attempts. Wish for appeasement. </td></tr></tbody></table> <p><a href="/wiki/Robert_Zajonc" title="Robert Zajonc">Robert Zajonc</a> published two reviews in 1989 of the "facial efference theory of emotion", also known as <a href="/wiki/Facial_feedback_hypothesis" title="Facial feedback hypothesis">facial feedback theory</a>,<sup id="cite_ref-Adelmann_7-0" class="reference"><a href="#cite_note-Adelmann-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Zajonc2_8-0" class="reference"><a href="#cite_note-Zajonc2-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> which he had first introduced to the scientific literature in an article published in <i><a href="/wiki/Science_(journal)" title="Science (journal)">Science</a></i> in 1985.<sup id="cite_ref-Zajonc1_9-0" class="reference"><a href="#cite_note-Zajonc1-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> This theory proposes that the facial musculature of mammals can control the temperature of the base of the brain (in particular the <a href="/wiki/Hypothalamus" title="Hypothalamus">hypothalamus</a>) by varying the degree of forward and backward flow through a vascular network (a so-called <i><a href="/wiki/Rete_mirabile" title="Rete mirabile">rete mirabile</a></i>). The theory is based on the idea that increasing the temperature of portions of the hypothalamus can produce aggressive behavior, whereas cooling can produce relaxation. Our emotional language has comparable descriptors, such as "hot-head" and "cool-breezy". The theory offers an explanation for the evolution of common facial expressions of emotion in mammals. Little experimental work has been done to extend the theory, however. </p><p><a href="/wiki/Carroll_Izard" title="Carroll Izard">Carroll Izard</a> discussed gains and losses associated with the evolution of emotions. He said that discrete emotion experiences emerge in ontogeny before language or conceptual structures that frame the qualia known as discrete emotion feelings are acquired. He noted that in evolution, when humans gained the <a href="https://en.wiktionary.org/wiki/capability" class="extiw" title="wikt:capability">capability</a> of expressing themselves with <a href="/wiki/Language" title="Language">language</a>, this contributed greatly to emotional evolution. Not only can humans <a href="https://en.wiktionary.org/wiki/articulate" class="extiw" title="wikt:articulate">articulate</a> and share their emotions, they can use their experiences to foresee and take appropriate action in future experiences. He did, however, raise the question of whether or not humans have lost some of their <a href="/wiki/Empathy" title="Empathy">empathy</a> for one another, citing things such as murder and crime against one another as destructive.<sup id="cite_ref-Izard_10-0" class="reference"><a href="#cite_note-Izard-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> </p><p><a href="/wiki/Joseph_LeDoux" class="mw-redirect" title="Joseph LeDoux">Joseph LeDoux</a> focuses much of his research on the emotion fear. Fear can be evoked by two systems in the <a href="/wiki/Brain" title="Brain">brain</a>, both involving the <a href="/wiki/Thalamus" title="Thalamus">thalamus</a> and the <a href="/wiki/Amygdala" title="Amygdala">amygdala</a>: one old, short and fast, the other more recently evolved, more circuitous and slower. In the older system, sensory information travels directly and quickly from the thalamus to the amygdala where it elicits the autonomic and motor responses we call fear. In the younger system, sensory information travels from the thalamus to the relevant cortical sensory areas (touch to the somatosensory cortex, vision to the visual cortex, etc.) and on to frontal association areas, where appraisal occurs. These frontal areas communicate directly with the amygdala and, in light of appraisal, may reduce or magnify the amygdala's fear response. If you glimpse what looks like a snake, long before your younger frontal areas have had time to determine it is a stick, the old thalamus-amygdala system will have evoked fear. LeDoux hypothesizes that the old fast system persists because a behavioral response at the first hint of danger is of little consequence when mistaken but may mean the difference between life and death when appropriate.<sup id="cite_ref-LeDoux1998_11-0" class="reference"><a href="#cite_note-LeDoux1998-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-LeDoux1996_12-0" class="reference"><a href="#cite_note-LeDoux1996-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Evolution_of_emotion&amp;action=edit&amp;section=4" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Animal_sexual_behaviour#Pleasure" title="Animal sexual behaviour">Animal sexual behaviour § Sex for pleasure</a></li> <li><a href="/wiki/Empathy#Other_animals_and_empathy_between_species" title="Empathy">Empathy § In animals</a></li> <li><a href="/wiki/Facial_expression" title="Facial expression">Facial expression</a></li> <li><a href="/wiki/Fear#Mechanism" title="Fear">Fear § In animals</a></li> <li><a href="/wiki/Reward_system#Other_species" title="Reward system">Reward system § Animals vs. humans</a></li> <li><a href="/wiki/Paul_Ekman" title="Paul Ekman">Paul Ekman</a></li> <li><a href="/wiki/Posture_(psychology)" title="Posture (psychology)">Posture (psychology)</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Evolution_of_emotion&amp;action=edit&amp;section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-Darwin2-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Darwin2_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Darwin2_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Darwin2_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Darwin2_1-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFDarwin2007" class="citation book cs1">Darwin, C. 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In Jenkins, JM; Oatley, K; Stein, NL (eds.). <i>Human Emotions: A reader</i>. Malden, Massachusetts: <a href="/wiki/Blackwell_Publishers" class="mw-redirect" title="Blackwell Publishers">Blackwell Publishers</a>. pp.&#160;<span class="nowrap">98–</span>111. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/0-631-20747-3" title="Special:BookSources/0-631-20747-3"><bdi>0-631-20747-3</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=bookitem&amp;rft.atitle=The+emotional+brain&amp;rft.btitle=Human+Emotions%3A+A+reader&amp;rft.place=Malden%2C+Massachusetts&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E98-%3C%2Fspan%3E111&amp;rft.pub=Blackwell+Publishers&amp;rft.date=1998&amp;rft.isbn=0-631-20747-3&amp;rft.aulast=LeDoux&amp;rft.aufirst=J&amp;rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3DkB19aJKJLoEC%26q%3Dthe%2Bemotional%2Bbrain%2Bthe%2Bmysterious%26pg%3DPA98&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AEvolution+of+emotion" class="Z3988"></span></span> </li> <li id="cite_note-LeDoux1996-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-LeDoux1996_12-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLedoux1996" class="citation journal cs1">Ledoux, J. (1996). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2723854">"Emotion Theory and Research: Highlights, Unanswered Questions, and Emerging Issues"</a>. <i><a href="/wiki/Annual_Review_of_Psychology" title="Annual Review of Psychology">Annual Review of Psychology</a></i>. <b>60</b> (1): <span class="nowrap">1–</span>25. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1146%2Fannurev.psych.60.110707.163539">10.1146/annurev.psych.60.110707.163539</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2723854">2723854</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18729725">18729725</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Annual+Review+of+Psychology&amp;rft.atitle=Emotion+Theory+and+Research%3A+Highlights%2C+Unanswered+Questions%2C+and+Emerging+Issues&amp;rft.volume=60&amp;rft.issue=1&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E1-%3C%2Fspan%3E25&amp;rft.date=1996&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2723854%23id-name%3DPMC&amp;rft_id=info%3Apmid%2F18729725&amp;rft_id=info%3Adoi%2F10.1146%2Fannurev.psych.60.110707.163539&amp;rft.aulast=Ledoux&amp;rft.aufirst=J.&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2723854&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AEvolution+of+emotion" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Evolution_of_emotion&amp;action=edit&amp;section=6" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="http://darwin-online.org.uk/content/frameset?itemID=F1142&amp;viewtype=text&amp;pageseq=1">The Expression of the Emotions in Man and Animals</a></li></ul> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output 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class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Evolutionary_psychology" title="Template:Evolutionary psychology"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Evolutionary_psychology" title="Template talk:Evolutionary psychology"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Evolutionary_psychology" title="Special:EditPage/Template:Evolutionary psychology"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Evolutionary_psychology1033" style="font-size:114%;margin:0 4em"><a href="/wiki/Evolutionary_psychology" title="Evolutionary psychology">Evolutionary psychology</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><a href="/wiki/History_of_evolutionary_psychology" title="History of evolutionary psychology">History</a> <ul><li><a href="/wiki/History_of_evolutionary_thought" title="History of evolutionary thought">Evolutionary thought</a></li></ul></li> <li><a href="/wiki/Theoretical_foundations_of_evolutionary_psychology" title="Theoretical foundations of evolutionary psychology">Theoretical foundations</a> <ul><li><a href="/wiki/Adaptationism" title="Adaptationism">Adaptationism</a></li> <li><a href="/wiki/Cognitive_revolution" title="Cognitive revolution">Cognitive revolution</a></li> <li><a href="/wiki/Cognitivism_(psychology)" title="Cognitivism (psychology)">Cognitivism</a></li> <li><a href="/wiki/Gene-centered_view_of_evolution" title="Gene-centered view of evolution">Gene selection theory</a></li> <li><a href="/wiki/Modern_synthesis_(20th_century)" title="Modern synthesis (20th century)">Modern synthesis</a></li></ul></li> <li><a href="/wiki/Criticism_of_evolutionary_psychology" title="Criticism of evolutionary psychology">Criticism</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align:left;"><a href="/wiki/Human_evolution" title="Human evolution">Evolutionary<br />processes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Psychological_adaptation" title="Psychological adaptation">Adaptations</a></li> <li><a href="/wiki/Altruism_(biology)" title="Altruism (biology)">Altruism</a> <ul><li><a href="/wiki/Cheating_(biology)" title="Cheating (biology)">Cheating</a></li> <li><a href="/wiki/Hamiltonian_spite" title="Hamiltonian spite">Hamiltonian spite</a></li> <li><a href="/wiki/Reciprocal_altruism_in_humans" title="Reciprocal altruism in humans">Reciprocal</a></li></ul></li> <li><a href="/wiki/Baldwin_effect" title="Baldwin effect">Baldwin effect</a></li> <li><a href="/wiki/Spandrel_(biology)" title="Spandrel (biology)">By-products</a></li> <li><a href="/wiki/Evolutionarily_stable_strategy" title="Evolutionarily stable strategy">Evolutionarily stable strategy</a></li> <li><a href="/wiki/Exaptation" title="Exaptation">Exaptation</a></li> <li><a href="/wiki/Fitness_(biology)" title="Fitness (biology)">Fitness</a> <ul><li><a href="/wiki/Inclusive_fitness_in_humans" title="Inclusive fitness in humans">Inclusive</a></li></ul></li> <li><a href="/wiki/Kin_selection" title="Kin selection">Kin selection</a></li> <li><a href="/wiki/Evolutionary_mismatch" title="Evolutionary mismatch">Mismatch</a></li> <li><a href="/wiki/Natural_selection" title="Natural selection">Natural selection</a></li> <li><a href="/wiki/Parental_investment" title="Parental investment">Parental investment</a> <ul><li><a href="/wiki/Parent%E2%80%93offspring_conflict" title="Parent–offspring conflict">Parent–offspring conflict</a></li></ul></li> <li><a href="/wiki/Sexual_selection_in_humans" title="Sexual selection in humans">Sexual selection</a> <ul><li><a href="/wiki/Costly_signaling_theory_in_evolutionary_psychology" title="Costly signaling theory in evolutionary psychology">Costly signaling</a></li> <li><a href="/wiki/Male_intrasexual_competition" title="Male intrasexual competition">Male</a>/<a href="/wiki/Female_intrasexual_competition" title="Female intrasexual competition">female intrasexual competition</a></li> <li><a href="/wiki/Mate_choice" title="Mate choice">Mate choice</a></li> <li><a href="/wiki/Sexual_dimorphism" title="Sexual dimorphism">Sexual dimorphism</a></li></ul></li> <li><a href="/wiki/Social_selection" title="Social selection">Social selection</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align:left;">Areas</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Evolution_of_cognition" title="Evolution of cognition">Cognition</a> /<br /><a class="mw-selflink selflink">Emotion</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Affect_(psychology)" title="Affect (psychology)">Affect</a> <ul><li><a href="/wiki/Affect_display" title="Affect display">Display</a></li> <li><a href="/wiki/Display_rules" title="Display rules">Display rules</a></li> <li><a href="/wiki/Facial_expression" title="Facial expression">Facial expression</a></li></ul></li> <li><a href="/wiki/Behavioral_modernity" title="Behavioral modernity">Behavioral modernity</a></li> <li><a href="/wiki/Cognitive_module" title="Cognitive module">Cognitive module</a>/<a href="/wiki/Modularity_of_mind" title="Modularity of mind">modularity of mind</a> <ul><li><a href="/wiki/Automatic_and_controlled_processes" title="Automatic and controlled processes">Automatic and controlled processes</a></li> <li><a href="/wiki/Computational_theory_of_mind" title="Computational theory of mind">Computational theory of mind</a></li> <li><a href="/wiki/Domain-general_learning" title="Domain-general learning">Domain generality</a></li> <li><a href="/wiki/Domain_specificity" title="Domain specificity">Domain specificity</a></li> <li><a href="/wiki/Dual_process_theory" title="Dual process theory">Dual process theory</a></li></ul></li> <li><a href="/wiki/Cognitive_tradeoff_hypothesis" title="Cognitive tradeoff hypothesis">Cognitive tradeoff hypothesis</a></li> <li><a href="/wiki/Evolution_of_the_brain" title="Evolution of the brain">Evolution of the brain</a></li> <li><a href="/wiki/Evolution_of_nervous_systems" title="Evolution of nervous systems">Evolution of nervous systems</a></li> <li><a href="/wiki/Fight-or-flight_response" title="Fight-or-flight response">Fight-or-flight response</a> <ul><li><a href="/wiki/Arachnophobia" title="Arachnophobia">Arachnophobia</a></li> <li><a href="/wiki/Fear_of_falling" title="Fear of falling">Basophobia</a></li> <li><a href="/wiki/Ophidiophobia" title="Ophidiophobia">Ophidiophobia</a></li></ul></li> <li><a href="/wiki/Folk_biology" title="Folk biology">Folk biology</a>/<a href="/wiki/Folk_taxonomy" title="Folk taxonomy">taxonomy</a></li> <li><a href="/wiki/Folk_psychology" title="Folk psychology">Folk psychology</a>/<a href="/wiki/Theory_of_mind" title="Theory of mind">theory of mind</a></li> <li><a href="/wiki/Evolution_of_human_intelligence" title="Evolution of human intelligence">Intelligence</a> <ul><li><a href="/wiki/Flynn_effect" title="Flynn effect">Flynn effect</a></li> <li><a href="/wiki/Wason_selection_task" title="Wason selection task">Wason selection task</a></li></ul></li> <li><a href="/wiki/Motor_control" title="Motor control">Motor control</a>/<a href="/wiki/Motor_skill" title="Motor skill">skill</a></li> <li><a href="/wiki/Human_multitasking" title="Human multitasking">Multitasking</a></li> <li><a href="/wiki/Neuroscience_of_sleep" title="Neuroscience of sleep">Sleep</a></li> <li><a href="/wiki/Visual_perception" title="Visual perception">Visual perception</a> <ul><li><a href="/wiki/Evolution_of_color_vision_in_primates" title="Evolution of color vision in primates">Color vision</a></li> <li><a href="/wiki/Evolution_of_the_eye" title="Evolution of the eye">Eye</a></li> <li><a href="/wiki/Na%C3%AFve_physics" title="Naïve physics">Naïve physics</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Evolutionary_psychology_and_culture" title="Evolutionary psychology and culture">Culture</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolutionary_aesthetics" title="Evolutionary aesthetics">Aesthetics</a> <ul><li><a href="/wiki/Darwinian_literary_studies" title="Darwinian literary studies">Literary criticism</a></li> <li><a href="/wiki/Evolutionary_musicology" title="Evolutionary musicology">Musicology</a></li></ul></li> <li><a href="/wiki/Evolutionary_anthropology" title="Evolutionary anthropology">Anthropology</a> <ul><li><a href="/wiki/Biological_anthropology" title="Biological anthropology">Biological</a></li></ul></li> <li><a href="/wiki/Biosocial_criminology" title="Biosocial criminology">Crime</a></li> <li><a href="/wiki/Evolutionary_linguistics" title="Evolutionary linguistics">Language</a> <ul><li><a href="/wiki/Origin_of_language" title="Origin of language">Origin</a></li> <li><a href="/wiki/Evolutionary_psychology_of_language" title="Evolutionary psychology of language">Psychology</a></li> <li><a href="/wiki/Origin_of_speech" title="Origin of speech">Speech</a></li></ul></li> <li><a href="/wiki/Evolution_of_morality" title="Evolution of morality">Morality</a> <ul><li><a href="/wiki/Moral_foundations_theory" title="Moral foundations theory">Moral foundations</a></li></ul></li> <li><a href="/wiki/Evolutionary_psychology_of_religion" title="Evolutionary psychology of religion">Religion</a> <ul><li><a href="/wiki/Evolutionary_origin_of_religions" class="mw-redirect" title="Evolutionary origin of religions">Origin</a></li></ul></li> <li><a href="/wiki/Cultural_universal" title="Cultural universal">Universals</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Evolutionary_developmental_psychology" title="Evolutionary developmental psychology">Development</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Attachment_theory" title="Attachment theory">Attachment</a></li> <li><a href="/wiki/Human_bonding" title="Human bonding">Bonding</a></li> <li><a href="/wiki/Affectional_bond" title="Affectional bond">Affectional</a>/<a href="/wiki/Maternal_bond" title="Maternal bond">maternal</a>/<a href="/wiki/Paternal_bond" title="Paternal bond">paternal bond</a></li> <li><a href="/wiki/Maternal_deprivation" title="Maternal deprivation">Caregiver deprivation</a></li> <li><a href="/wiki/Attachment_in_children" title="Attachment in children">Childhood attachment</a></li> <li><a href="/wiki/Cinderella_effect" title="Cinderella effect">Cinderella effect</a></li> <li><a href="/wiki/Cognitive_development" title="Cognitive development">Cognitive development</a></li> <li><a href="/wiki/Evolutionary_educational_psychology" title="Evolutionary educational psychology">Education</a></li> <li><a href="/wiki/Language_acquisition" title="Language acquisition">Language acquisition</a></li> <li><a href="/wiki/Personality_development" title="Personality development">Personality development</a></li> <li><a href="/wiki/Socialization" title="Socialization">Socialization</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ergonomics" title="Ergonomics">Human factors</a> /<br /><a href="/wiki/Evolutionary_psychiatry" title="Evolutionary psychiatry">Mental health</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cognitive_ergonomics" title="Cognitive ergonomics">Cognitive ergonomics</a> <ul><li><a href="/wiki/Computer-mediated_communication" title="Computer-mediated communication">Computer-mediated communication</a></li> <li><a href="/wiki/Engineering_psychology" title="Engineering psychology">Engineering psychology</a></li> <li><a href="/wiki/Human%E2%80%93computer_interaction" title="Human–computer interaction">Human–computer interaction</a></li> <li><a href="/wiki/Media_naturalness_theory" title="Media naturalness theory">Media naturalness theory</a></li> <li><a href="/wiki/Neuroergonomics" title="Neuroergonomics">Neuroergonomics</a></li></ul></li> <li><a href="/wiki/Evolutionary_approaches_to_depression" title="Evolutionary approaches to depression">Depression</a></li> <li><a href="/wiki/Digital_media_use_and_mental_health" title="Digital media use and mental health">Digital media use and mental health</a></li> <li><a href="/wiki/Accident-proneness" title="Accident-proneness">Hypophobia</a></li> <li><a href="/wiki/Imprinted_brain_hypothesis" title="Imprinted brain hypothesis">Imprinted brain hypothesis</a></li> <li><a href="/wiki/Mind-blindness" title="Mind-blindness">Mind-blindness</a></li> <li><a href="/wiki/Psychological_effects_of_Internet_use" title="Psychological effects of Internet use">Psychological effects of Internet use</a></li> <li><a href="/wiki/Rank_theory_of_depression" title="Rank theory of depression">Rank theory of depression</a></li> <li><a href="/wiki/Evolution_of_schizophrenia" title="Evolution of schizophrenia">Schizophrenia</a></li> <li><a href="/wiki/Screen_time" title="Screen time">Screen time</a></li> <li><a href="/wiki/Smartphones_and_pedestrian_safety" title="Smartphones and pedestrian safety">Smartphones and pedestrian safety</a></li> <li><a href="/wiki/Social_aspects_of_television" title="Social aspects of television">Social aspects of television</a></li> <li><a href="/wiki/Societal_effects_of_cars" title="Societal effects of cars">Societal effects of cars</a> <ul><li><a href="/wiki/Distracted_driving" title="Distracted driving">Distracted driving</a></li> <li><a href="/wiki/Lead%E2%80%93crime_hypothesis" title="Lead–crime hypothesis">Lead–crime hypothesis</a></li> <li><a href="/wiki/Mobile_phones_and_driving_safety" title="Mobile phones and driving safety">Mobile phones and driving safety</a></li> <li><a href="/wiki/Texting_while_driving" title="Texting while driving">Texting while driving</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sexology" title="Sexology">Sex</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Human_sexual_activity" title="Human sexual activity">Activity</a></li> <li><a href="/wiki/Attachment_in_adults" title="Attachment in adults">Adult attachment</a></li> <li><a href="/wiki/Age_disparity_in_sexual_relationships" title="Age disparity in sexual relationships">Age disparity</a></li> <li><a href="/wiki/Sexual_arousal" title="Sexual arousal">Arousal</a></li> <li><a href="/wiki/Concealed_ovulation" title="Concealed ovulation">Concealed ovulation</a></li> <li><a href="/wiki/Coolidge_effect" title="Coolidge effect">Coolidge effect</a></li> <li><a href="/wiki/Sexual_desire" title="Sexual desire">Desire</a></li> <li><a href="/wiki/Sexual_fantasy" title="Sexual fantasy">Fantasy</a></li> <li><a href="/wiki/Effects_of_hormones_on_sexual_motivation" title="Effects of hormones on sexual motivation">Hormonal motivation</a></li> <li><a href="/wiki/Sexual_jealousy" title="Sexual jealousy">Jealousy</a></li> <li><a href="/wiki/Mate_guarding_in_humans" title="Mate guarding in humans">Mate guarding</a></li> <li><a href="/wiki/Mating_preferences" title="Mating preferences">Mating preferences</a></li> <li><a href="/wiki/Human_mating_strategies" title="Human mating strategies">Mating strategies</a></li> <li><a href="/wiki/Biology_and_sexual_orientation" title="Biology and sexual orientation">Orientation</a></li> <li><a href="/wiki/Ovulatory_shift_hypothesis" title="Ovulatory shift hypothesis">Ovulatory shift hypothesis</a></li> <li><a href="/wiki/Pair_bond" title="Pair bond">Pair bond</a></li> <li><a href="/wiki/Physical_attractiveness" title="Physical attractiveness">Physical</a>/<a href="/wiki/Sexual_attraction" title="Sexual attraction">Sexual attraction</a></li> <li><a href="/wiki/Human_sexuality" title="Human sexuality">Sexuality</a>/<a href="/wiki/Human_male_sexuality" title="Human male sexuality">male</a>/<a href="/wiki/Human_female_sexuality" title="Human female sexuality">female</a></li> <li><a href="/wiki/Sexy_son_hypothesis" title="Sexy son hypothesis">Sexy son hypothesis</a></li> <li><a href="/wiki/Westermarck_effect" title="Westermarck effect">Westermarck effect</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sex_differences_in_psychology" title="Sex differences in psychology">Sex differences</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aggression" title="Aggression">Aggression</a></li> <li><a href="/wiki/Sex_and_gender_differences_in_autism" title="Sex and gender differences in autism">Autism</a></li> <li><a href="/wiki/Sex_differences_in_cognition" title="Sex differences in cognition">Cognition</a></li> <li><a href="/wiki/Sex_differences_in_crime" title="Sex differences in crime">Crime</a></li> <li><a href="/wiki/Sexual_division_of_labour" title="Sexual division of labour">Division of labour</a></li> <li><a href="/wiki/Sex_differences_in_emotional_intelligence" title="Sex differences in emotional intelligence">Emotional intelligence</a></li> <li><a href="/wiki/Empathising%E2%80%93systemising_theory" title="Empathising–systemising theory">Empathising–systemising theory</a></li> <li><a href="/wiki/Gender_role" title="Gender role">Gender role</a></li> <li><a href="/wiki/Sex_differences_in_intelligence" title="Sex differences in intelligence">Intelligence</a></li> <li><a href="/wiki/Sex_differences_in_memory" title="Sex differences in memory">Memory</a></li> <li><a href="/wiki/Mental_disorders_and_gender" title="Mental disorders and gender">Mental health</a></li> <li><a href="/wiki/Sex_differences_in_narcissism" title="Sex differences in narcissism">Narcissism</a></li> <li><a href="/wiki/Neuroscience_of_sex_differences" title="Neuroscience of sex differences">Neuroscience</a></li> <li><a href="/wiki/Sex_differences_in_schizophrenia" title="Sex differences in schizophrenia">Schizophrenia</a></li> <li><a href="/wiki/Substance_abuse" title="Substance abuse">Substance abuse</a></li> <li><a href="/wiki/Gender_differences_in_suicide" title="Gender differences in suicide">Suicide</a></li> <li><a href="/wiki/Variability_hypothesis" title="Variability hypothesis">Variability hypothesis</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align:left;">Related subjects</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Academic disciplines</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Behavioral_economics" title="Behavioral economics">Behavioral</a>/<a href="/wiki/Evolutionary_economics" title="Evolutionary economics">evolutionary economics</a></li> <li><a href="/wiki/Behavioral_epigenetics" title="Behavioral epigenetics">Behavioral epigenetics</a>/<a href="/wiki/Behavioural_genetics" title="Behavioural genetics">genetics</a></li> <li><a href="/wiki/Affective_neuroscience" title="Affective neuroscience">Affective</a>/<a href="/wiki/Behavioral_neuroscience" title="Behavioral neuroscience">behavioral</a>/<a href="/wiki/Cognitive_neuroscience" title="Cognitive neuroscience">cognitive</a>/<a href="/wiki/Evolutionary_neuroscience" title="Evolutionary neuroscience">evolutionary neuroscience</a></li> <li><a href="/wiki/Biocultural_anthropology" title="Biocultural anthropology">Biocultural anthropology</a></li> <li><a href="/wiki/Biological_psychiatry" title="Biological psychiatry">Biological psychiatry</a></li> <li><a href="/wiki/Cognitive_psychology" title="Cognitive psychology">Cognitive psychology</a></li> <li><a href="/wiki/Cognitive_science" title="Cognitive science">Cognitive science</a></li> <li><a href="/wiki/Cross-cultural_psychology" title="Cross-cultural psychology">Cross-cultural psychology</a></li> <li><a href="/wiki/Ethology" title="Ethology">Ethology</a></li> <li><a href="/wiki/Evolutionary_biology" title="Evolutionary biology">Evolutionary biology</a></li> <li><a href="/wiki/Evolutionary_medicine" title="Evolutionary medicine">Evolutionary medicine</a></li> <li><a href="/wiki/Functional_psychology" title="Functional psychology">Functional psychology</a></li> <li><a href="/wiki/Neuropsychology" title="Neuropsychology">Neuropsychology</a></li> <li><a href="/wiki/Philosophy_of_mind" title="Philosophy of mind">Philosophy of mind</a></li> <li><a href="/wiki/Population_genetics" title="Population genetics">Population genetics</a></li> <li><a href="/wiki/Primatology" title="Primatology">Primatology</a></li> <li><a href="/wiki/Sociobiology" title="Sociobiology">Sociobiology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Research topics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cultural_evolution" title="Cultural evolution">Cultural evolution</a></li> <li><a href="/wiki/Evolutionary_epistemology" title="Evolutionary epistemology">Evolutionary epistemology</a></li> <li><a href="/wiki/Great_ape_language" title="Great ape language">Great ape language</a></li> <li><a href="/wiki/Human%E2%80%93animal_communication" title="Human–animal communication">Human–animal communication</a></li> <li><a href="/wiki/Missing_heritability_problem" title="Missing heritability problem">Missing heritability problem</a></li> <li><a href="/wiki/Primate_cognition" title="Primate cognition">Primate cognition</a></li> <li><a href="/wiki/Unit_of_selection" title="Unit of selection">Unit of selection</a> <ul><li><a href="/wiki/Coevolution" title="Coevolution">Coevolution</a></li> <li><a href="/wiki/Cultural_group_selection" title="Cultural group selection">Cultural group selection</a></li> <li><a href="/wiki/Dual_inheritance_theory" title="Dual inheritance theory">Dual inheritance theory</a></li> <li><a href="/wiki/Fisher%27s_principle" title="Fisher&#39;s principle">Fisher's principle</a></li> <li><a href="/wiki/Group_selection" title="Group selection">Group selection</a></li> <li><a href="/wiki/Hologenome_theory_of_evolution" title="Hologenome theory of evolution">Hologenome theory</a></li> <li><a href="/wiki/Lamarckism" title="Lamarckism">Lamarckism</a></li> <li><a href="/wiki/Population" title="Population">Population</a></li> <li><a href="/wiki/Punctuated_equilibrium" title="Punctuated equilibrium">Punctuated equilibrium</a></li> <li><a href="/wiki/Recent_human_evolution" title="Recent human evolution">Recent human evolution</a></li> <li><a href="/wiki/Species" title="Species">Species</a></li> <li><a href="/wiki/Species_complex" title="Species complex">Species complex</a></li> <li><a href="/wiki/Transgenerational_epigenetic_inheritance" title="Transgenerational epigenetic inheritance">Transgenerational epigenetic inheritance</a></li> <li><a href="/wiki/Trivers%E2%80%93Willard_hypothesis" title="Trivers–Willard hypothesis">Trivers–Willard hypothesis</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Theoretical positions</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cultural_selection_theory" title="Cultural selection theory">Cultural selection theory</a></li> <li><a href="/wiki/Determinism" title="Determinism">Determinism</a>/<a href="/wiki/Indeterminism" title="Indeterminism">indeterminism</a> <ul><li><a href="/wiki/Biological_determinism" title="Biological determinism">Biological determinism</a></li> <li><a href="/wiki/Connectionism" title="Connectionism">Connectionism</a></li> <li><a href="/wiki/Cultural_determinism" title="Cultural determinism">Cultural determinism</a></li> <li><a href="/wiki/Environmental_determinism" title="Environmental determinism">Environmental determinism</a></li> <li><a href="/wiki/Nature_versus_nurture" title="Nature versus nurture">Nature versus nurture</a></li> <li><a href="/wiki/Psychological_nativism" title="Psychological nativism">Psychological nativism</a></li> <li><a href="/wiki/Social_constructionism" title="Social constructionism">Social constructionism</a></li> <li><a href="/wiki/Social_determinism" title="Social determinism">Social determinism</a></li> <li><a href="/wiki/Standard_social_science_model" title="Standard social science model">Standard social science model</a></li></ul></li> <li><a href="/wiki/Functionalism_(philosophy_of_mind)" title="Functionalism (philosophy of mind)">Functionalism</a></li> <li><a href="/wiki/Memetics" title="Memetics">Memetics</a></li> <li><a href="/wiki/Multilineal_evolution" title="Multilineal evolution">Multilineal evolution</a></li> <li><a href="/wiki/Neo-Darwinism" title="Neo-Darwinism">Neo-Darwinism</a></li> <li><a href="/wiki/Neoevolutionism" title="Neoevolutionism">Neoevolutionism</a></li> <li><a href="/wiki/Sociocultural_evolution" title="Sociocultural evolution">Sociocultural evolution</a></li> <li><a href="/wiki/Unilineal_evolution" title="Unilineal evolution">Unilineal evolution</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><b><a href="/wiki/List_of_evolutionary_psychologists" title="List of evolutionary psychologists">Evolutionary psychologists</a></b></li> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" 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life</a></li> <li><a href="/wiki/Index_of_evolutionary_biology_articles" title="Index of evolutionary biology articles">Index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Evolution" title="Evolution">Evolution</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></li> <li><a href="/wiki/Adaptation" title="Adaptation">Adaptation</a></li> <li><a href="/wiki/Adaptive_radiation" title="Adaptive radiation">Adaptive radiation</a></li> <li><a href="/wiki/Altruism_(biology)" title="Altruism (biology)">Altruism</a> <ul><li><a href="/wiki/Cheating_(biology)" title="Cheating (biology)">Cheating</a></li> <li><a href="/wiki/Reciprocal_altruism" title="Reciprocal altruism">Reciprocal</a></li></ul></li> <li><a href="/wiki/Baldwin_effect" title="Baldwin effect">Baldwin effect</a></li> <li><a href="/wiki/Cladistics" title="Cladistics">Cladistics</a></li> <li><a href="/wiki/Coevolution" title="Coevolution">Coevolution</a> <ul><li><a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">Mutualism</a></li></ul></li> <li><a href="/wiki/Common_descent" title="Common descent">Common descent</a></li> <li><a href="/wiki/Convergent_evolution" title="Convergent evolution">Convergence</a></li> <li><a href="/wiki/Divergent_evolution" title="Divergent evolution">Divergence</a></li> <li><a href="/wiki/Earliest_known_life_forms" title="Earliest known life forms">Earliest known life forms</a></li> <li><a href="/wiki/Evidence_of_common_descent" title="Evidence of common descent">Evidence of evolution</a></li> <li><a href="/wiki/Evolutionary_arms_race" title="Evolutionary arms race">Evolutionary arms race</a></li> <li><a href="/wiki/Evolutionary_pressure" title="Evolutionary pressure">Evolutionary pressure</a></li> <li><a href="/wiki/Exaptation" title="Exaptation">Exaptation</a></li> <li><a href="/wiki/Extinction" title="Extinction">Extinction</a> <ul><li><a href="/wiki/Extinction_event" title="Extinction event">Event</a></li></ul></li> <li><a href="/wiki/Homology_(biology)" title="Homology (biology)">Homology</a></li> <li><a href="/wiki/Last_universal_common_ancestor" title="Last universal common ancestor">Last universal common ancestor</a></li> <li><a href="/wiki/Macroevolution" title="Macroevolution">Macroevolution</a></li> <li><a href="/wiki/Microevolution" title="Microevolution">Microevolution</a></li> <li><a href="/wiki/Evolutionary_mismatch" title="Evolutionary mismatch">Mismatch</a></li> <li><a href="/wiki/Nonadaptive_radiation" title="Nonadaptive radiation">Non-adaptive radiation</a></li> <li><a href="/wiki/Abiogenesis" title="Abiogenesis">Origin of life</a></li> <li><a href="/wiki/Panspermia" title="Panspermia">Panspermia</a></li> <li><a href="/wiki/Parallel_evolution" title="Parallel evolution">Parallel evolution</a></li> <li><a href="/wiki/Signalling_theory" title="Signalling theory">Signalling theory</a> <ul><li><a href="/wiki/Handicap_principle" title="Handicap principle">Handicap principle</a></li></ul></li> <li><a href="/wiki/Speciation" title="Speciation">Speciation</a> <ul><li><a href="/wiki/Species" title="Species">Species</a></li> <li><a href="/wiki/Species_complex" title="Species complex">Species complex</a></li></ul></li> <li><a href="/wiki/Evolutionary_taxonomy" title="Evolutionary taxonomy">Taxonomy</a></li> <li><a href="/wiki/Unit_of_selection" title="Unit of selection">Unit of selection</a> <ul><li><a href="/wiki/Gene-centered_view_of_evolution" title="Gene-centered view of evolution">Gene-centered view of evolution</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Population_genetics" title="Population genetics">Population<br />genetics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Selective_breeding" title="Selective breeding">Artificial selection</a></li> <li><a href="/wiki/Biodiversity" title="Biodiversity">Biodiversity</a></li> <li><a href="/wiki/Evolutionarily_stable_strategy" title="Evolutionarily stable strategy">Evolutionarily stable strategy</a></li> <li><a href="/wiki/Fisher%27s_principle" title="Fisher&#39;s principle">Fisher's principle</a></li> <li><a href="/wiki/Fitness_(biology)" title="Fitness (biology)">Fitness</a> <ul><li><a href="/wiki/Inclusive_fitness" title="Inclusive fitness">Inclusive</a></li></ul></li> <li><a href="/wiki/Gene_flow" title="Gene flow">Gene flow</a></li> <li><a href="/wiki/Genetic_drift" title="Genetic drift">Genetic drift</a></li> <li><a href="/wiki/Kin_selection" title="Kin selection">Kin selection</a> <ul><li><a href="/wiki/Parental_investment" title="Parental investment">Parental investment</a></li> <li><a href="/wiki/Parent%E2%80%93offspring_conflict" title="Parent–offspring conflict">Parent–offspring conflict</a></li></ul></li> <li><a href="/wiki/Mutation" title="Mutation">Mutation</a></li> <li><a href="/wiki/Population" title="Population">Population</a></li> <li><a href="/wiki/Natural_selection" title="Natural selection">Natural selection</a></li> <li><a href="/wiki/Sexual_dimorphism" title="Sexual dimorphism">Sexual dimorphism</a></li> <li><a href="/wiki/Sexual_selection" title="Sexual selection">Sexual selection</a> <ul><li><a href="/wiki/Sexual_selection_in_flowering_plants" title="Sexual selection in flowering plants">Flowering plants</a></li> <li><a href="/wiki/Sexual_selection_in_fungi" title="Sexual selection in fungi">Fungi</a></li> <li><a href="/wiki/Mate_choice" title="Mate choice">Mate choice</a></li></ul></li> <li><a href="/wiki/Social_selection" title="Social selection">Social selection</a></li> <li><a href="/wiki/Trivers%E2%80%93Willard_hypothesis" title="Trivers–Willard hypothesis">Trivers–Willard hypothesis</a></li> <li><a href="/wiki/Genetic_variation" title="Genetic variation">Variation</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Development</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Canalisation_(genetics)" title="Canalisation (genetics)">Canalisation</a></li> <li><a href="/wiki/Evolutionary_developmental_biology" title="Evolutionary developmental biology">Evolutionary developmental biology</a></li> <li><a href="/wiki/Genetic_assimilation" title="Genetic assimilation">Genetic assimilation</a></li> <li><a href="/wiki/Inversion_(evolutionary_biology)" title="Inversion (evolutionary biology)">Inversion</a></li> <li><a href="/wiki/Modularity_(biology)" title="Modularity (biology)">Modularity</a></li> <li><a href="/wiki/Phenotypic_plasticity" title="Phenotypic plasticity">Phenotypic plasticity</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Of <a href="/wiki/Taxon" title="Taxon">taxa</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolution_of_bacteria" title="Evolution of bacteria">Bacteria</a></li> <li><a href="/wiki/Evolution_of_birds" title="Evolution of birds">Birds</a> <ul><li><a href="/wiki/Origin_of_birds" title="Origin of birds">origin</a></li></ul></li> <li><a href="/wiki/Evolution_of_brachiopods" title="Evolution of brachiopods">Brachiopods</a></li> <li><a href="/wiki/Evolution_of_molluscs" title="Evolution of molluscs">Molluscs</a> <ul><li><a href="/wiki/Evolution_of_cephalopods" title="Evolution of cephalopods">Cephalopods</a></li></ul></li> <li><a href="/wiki/Dinosaur#Evolutionary_history" title="Dinosaur">Dinosaurs</a></li> <li><a href="/wiki/Evolution_of_fish" title="Evolution of fish">Fish</a></li> <li><a href="/wiki/Evolution_of_fungi" title="Evolution of fungi">Fungi</a></li> <li><a href="/wiki/Evolution_of_insects" title="Evolution of insects">Insects</a> <ul><li><a href="/wiki/Evolution_of_butterflies" title="Evolution of butterflies">butterflies</a></li></ul></li> <li><a href="/wiki/History_of_life" title="History of life">Life</a></li> <li><a href="/wiki/Evolution_of_mammals" title="Evolution of mammals">Mammals</a> <ul><li><a href="/wiki/Cat_gap" title="Cat gap">cats</a></li> <li><a href="/wiki/Canidae#Evolution" title="Canidae">canids</a> <ul><li><a href="/wiki/Evolution_of_the_wolf" title="Evolution of the wolf">wolves</a></li> <li><a href="/wiki/Domestication_of_the_dog" title="Domestication of the dog">dogs</a></li></ul></li> <li><a href="/wiki/Hyena#Evolution" title="Hyena">hyenas</a></li> <li><a href="/wiki/Evolution_of_cetaceans" title="Evolution of cetaceans">dolphins and whales</a></li> <li><a href="/wiki/Evolution_of_the_horse" title="Evolution of the horse">horses</a></li> <li><a href="/wiki/Evolution_of_Macropodidae" title="Evolution of Macropodidae">Kangaroos</a></li> <li><a href="/wiki/Evolution_of_primates" title="Evolution of primates">primates</a> <ul><li><a href="/wiki/Human_evolution" title="Human evolution">humans</a></li> <li><a href="/wiki/Evolution_of_lemurs" title="Evolution of lemurs">lemurs</a></li></ul></li> <li><a href="/wiki/Evolution_of_sirenians" title="Evolution of sirenians">sea cows</a></li></ul></li> <li><a href="/wiki/Evolutionary_history_of_plants" title="Evolutionary history of plants">Plants</a> <ul><li><a href="/wiki/Pollinator-mediated_selection" title="Pollinator-mediated selection">pollinator-mediated</a></li></ul></li> <li><a href="/wiki/Evolution_of_reptiles" title="Evolution of reptiles">Reptiles</a></li> <li><a href="/wiki/Evolution_of_spiders" title="Evolution of spiders">Spiders</a></li> <li><a href="/wiki/Evolution_of_tetrapods" title="Evolution of tetrapods">Tetrapods</a></li> <li><a href="/wiki/Viral_evolution" title="Viral evolution">Viruses</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Of <a href="/wiki/Organ_(anatomy)" class="mw-redirect" title="Organ (anatomy)">organs</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolution_of_cells" title="Evolution of cells">Cell</a></li> <li><a href="/wiki/Models_of_DNA_evolution" title="Models of DNA evolution">DNA</a></li> <li><a href="/wiki/Evolution_of_flagella" title="Evolution of flagella">Flagella</a></li> <li><a href="/wiki/Eukaryote#Origin_of_eukaryotes" title="Eukaryote">Eukaryotes</a> <ul><li><a href="/wiki/Symbiogenesis" title="Symbiogenesis">symbiogenesis</a></li> <li><a href="/wiki/Chromosome#Eukaryotes" title="Chromosome">chromosome</a></li> <li><a href="/wiki/Endomembrane_system#Evolution" title="Endomembrane system">endomembrane system</a></li> <li><a href="/wiki/Mitochondrion#Origin_and_evolution" title="Mitochondrion">mitochondria</a></li> <li><a href="/wiki/Cell_nucleus#Evolution" title="Cell nucleus">nucleus</a></li> <li><a href="/wiki/Plastid#Origin" title="Plastid">plastids</a></li></ul></li> <li>In animals <ul><li><a href="/wiki/Evolution_of_the_eye" title="Evolution of the eye">eye</a></li> <li><a href="/wiki/Hair#Evolution" title="Hair">hair</a></li> <li><a href="/wiki/Evolution_of_mammalian_auditory_ossicles" title="Evolution of mammalian auditory ossicles">auditory ossicle</a></li> <li><a href="/wiki/Evolution_of_nervous_systems" title="Evolution of nervous systems">nervous system</a></li> <li><a href="/wiki/Evolution_of_the_brain" title="Evolution of the brain">brain</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Of <a href="/wiki/Biological_process" title="Biological process">processes</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolution_of_ageing" title="Evolution of ageing">Aging</a> <ul><li><a href="/wiki/Death#Evolution_of_aging_and_mortality" title="Death">Death</a></li> <li><a href="/wiki/Programmed_cell_death#Evolutionary_origin_of_mitochondrial_apoptosis" title="Programmed cell death">Programmed cell death</a></li></ul></li> <li><a href="/wiki/Origin_of_avian_flight" title="Origin of avian flight">Avian flight</a></li> <li><a href="/wiki/Evolution_of_biological_complexity" title="Evolution of biological complexity">Biological complexity</a></li> <li><a href="/wiki/Cooperation_(evolution)" title="Cooperation (evolution)">Cooperation</a></li> <li><a href="/wiki/Evolution_of_color_vision" title="Evolution of color vision">Color vision</a> <ul><li><a href="/wiki/Evolution_of_color_vision_in_primates" title="Evolution of color vision in primates">in primates</a></li></ul></li> <li><a class="mw-selflink selflink">Emotion</a></li> <li><a href="/wiki/Empathy#Evolution_across_species" title="Empathy">Empathy</a></li> <li><a href="/wiki/Evolutionary_ethics" title="Evolutionary ethics">Ethics</a></li> <li><a href="/wiki/Evolution_of_eusociality" title="Evolution of eusociality">Eusociality</a></li> <li><a href="/wiki/Immune_system#Evolution_and_other_mechanisms" title="Immune system">Immune system</a></li> <li><a href="/wiki/Metabolism#Evolution" title="Metabolism">Metabolism</a></li> <li><a href="/wiki/Monogamy_in_animals" title="Monogamy in animals">Monogamy</a></li> <li><a href="/wiki/Evolution_of_morality" title="Evolution of morality">Morality</a></li> <li><a href="/wiki/Mosaic_evolution" title="Mosaic evolution">Mosaic evolution</a></li> <li><a href="/wiki/Multicellular_organism#Evolutionary_history" title="Multicellular organism">Multicellularity</a></li> <li><a href="/wiki/Evolution_of_sexual_reproduction" title="Evolution of sexual reproduction">Sexual reproduction</a> <ul><li><a href="/wiki/Anisogamy#Evolution" title="Anisogamy">Gamete differentiation/sexes</a></li> <li><a href="/wiki/Biological_life_cycle" title="Biological life cycle">Life cycles/nuclear phases</a></li> <li><a href="/wiki/Mating_type" title="Mating type">Mating types</a></li> <li><a href="/wiki/Origin_and_function_of_meiosis" title="Origin and function of meiosis">Meiosis</a></li> <li><a href="/wiki/Sex-determination_system#Evolution" title="Sex-determination system">Sex-determination</a></li></ul></li> <li><a href="/wiki/Evolution_of_snake_venom" title="Evolution of snake venom">Snake venom</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tempo_and_Mode_in_Evolution" title="Tempo and Mode in Evolution">Tempo and modes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Phyletic_gradualism" title="Phyletic gradualism">Gradualism</a>/<a href="/wiki/Punctuated_equilibrium" title="Punctuated equilibrium">Punctuated equilibrium</a>/<a href="/wiki/Saltation_(biology)" title="Saltation (biology)">Saltationism</a></li> <li><a href="/wiki/Point_mutation" title="Point mutation">Micromutation</a>/<a href="/wiki/Mutationism#Later_mutationist_theories" title="Mutationism">Macromutation</a></li> <li><a href="/wiki/Uniformitarianism" title="Uniformitarianism">Uniformitarianism</a>/<a href="/wiki/Catastrophism" title="Catastrophism">Catastrophism</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Speciation" title="Speciation">Speciation</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Allopatric_speciation" title="Allopatric speciation">Allopatric</a></li> <li><a href="/wiki/Anagenesis" title="Anagenesis">Anagenesis</a></li> <li><a href="/wiki/Catagenesis_(biology)" title="Catagenesis (biology)">Catagenesis</a></li> <li><a href="/wiki/Cladogenesis" title="Cladogenesis">Cladogenesis</a></li> <li><a href="/wiki/Cospeciation" title="Cospeciation">Cospeciation</a></li> <li><a href="/wiki/Ecological_speciation" title="Ecological speciation">Ecological</a></li> <li><a href="/wiki/Hybrid_speciation" title="Hybrid speciation">Hybrid</a></li> <li><a href="/wiki/Nonecological_speciation" title="Nonecological speciation">Non-ecological</a></li> <li><a href="/wiki/Parapatric_speciation" title="Parapatric speciation">Parapatric</a></li> <li><a href="/wiki/Peripatric_speciation" title="Peripatric speciation">Peripatric</a></li> <li><a href="/wiki/Reinforcement_(speciation)" title="Reinforcement (speciation)">Reinforcement</a></li> <li><a href="/wiki/Sympatric_speciation" title="Sympatric speciation">Sympatric</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/History_of_evolutionary_thought" title="History of evolutionary thought">History</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolutionary_ideas_of_the_Renaissance_and_Enlightenment" title="Evolutionary ideas of the Renaissance and Enlightenment">Renaissance and Enlightenment</a></li> <li><a href="/wiki/Transmutation_of_species" title="Transmutation of species">Transmutation of species</a></li> <li><a href="/wiki/David_Hume" title="David Hume">David Hume</a> <ul><li><i><a href="/wiki/Dialogues_Concerning_Natural_Religion" title="Dialogues Concerning Natural Religion">Dialogues Concerning Natural Religion</a></i></li></ul></li> <li><a href="/wiki/Charles_Darwin" title="Charles Darwin">Charles Darwin</a> <ul><li><i><a href="/wiki/On_the_Origin_of_Species" title="On the Origin of Species">On the Origin of Species</a></i></li></ul></li> <li><a href="/wiki/History_of_paleontology" title="History of paleontology">History of paleontology</a></li> <li><a href="/wiki/Transitional_fossil" title="Transitional fossil">Transitional fossil</a></li> <li><a href="/wiki/Blending_inheritance" title="Blending inheritance">Blending inheritance</a></li> <li><a href="/wiki/Mendelian_inheritance" title="Mendelian inheritance">Mendelian inheritance</a></li> <li><a href="/wiki/The_eclipse_of_Darwinism" title="The eclipse of Darwinism">The eclipse of Darwinism</a></li> <li><a href="/wiki/Neo-Darwinism" title="Neo-Darwinism">Neo-Darwinism</a></li> <li><a href="/wiki/Modern_synthesis_(20th_century)" title="Modern synthesis (20th century)">Modern synthesis</a></li> <li><a href="/wiki/History_of_molecular_evolution" title="History of molecular evolution">History of molecular evolution</a></li> <li><a href="/wiki/Extended_evolutionary_synthesis" title="Extended evolutionary synthesis">Extended evolutionary synthesis</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Philosophy</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Darwinism" title="Darwinism">Darwinism</a></li> <li><a href="/wiki/Alternatives_to_Darwinian_evolution" title="Alternatives to Darwinian evolution">Alternatives</a> <ul><li><a href="/wiki/Catastrophism" title="Catastrophism">Catastrophism</a></li> <li><a href="/wiki/Lamarckism" title="Lamarckism">Lamarckism</a></li> <li><a href="/wiki/Orthogenesis" title="Orthogenesis">Orthogenesis</a></li> <li><a href="/wiki/Mutationism" title="Mutationism">Mutationism</a></li> <li><a href="/wiki/Saltation_(biology)" title="Saltation (biology)">Saltationism</a></li> <li><a href="/wiki/Structuralism_(biology)" title="Structuralism (biology)">Structuralism</a> <ul><li><a href="/wiki/Spandrel_(biology)" title="Spandrel (biology)">Spandrel</a></li></ul></li> <li><a href="/wiki/Theistic_evolution" title="Theistic evolution">Theistic</a></li> <li><a href="/wiki/Vitalism" title="Vitalism">Vitalism</a></li></ul></li> <li><a href="/wiki/Teleology_in_biology" title="Teleology in biology">Teleology in biology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Related</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Ecological_genetics" title="Ecological genetics">Ecological genetics</a></li> <li><a href="/wiki/Evolutionary_medicine" title="Evolutionary medicine">Evolutionary medicine</a></li> <li><a href="/wiki/Group_selection" title="Group selection">Group selection</a> <ul><li><a href="/wiki/Cultural_evolution" title="Cultural evolution">Cultural evolution</a></li> <li><a href="/wiki/Cultural_group_selection" title="Cultural group selection">Cultural group selection</a></li> <li><a href="/wiki/Dual_inheritance_theory" title="Dual inheritance theory">Dual inheritance theory</a></li></ul></li> <li><a href="/wiki/Hologenome_theory_of_evolution" title="Hologenome theory of evolution">Hologenome theory of evolution</a></li> <li><a href="/wiki/Missing_heritability_problem" title="Missing heritability problem">Missing heritability problem</a></li> <li><a href="/wiki/Molecular_evolution" title="Molecular evolution">Molecular evolution</a></li> <li><a href="/wiki/Astrobiology" title="Astrobiology">Astrobiology</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a> <ul><li><a href="/wiki/Phylogenetic_tree" title="Phylogenetic tree">Tree</a></li></ul></li> <li><a href="/wiki/Polymorphism_(biology)" title="Polymorphism (biology)">Polymorphism</a></li> <li><a href="/wiki/Protocell" title="Protocell">Protocell</a></li> <li><a href="/wiki/Systematics" title="Systematics">Systematics</a></li> <li><a href="/wiki/Transgenerational_epigenetic_inheritance" title="Transgenerational epigenetic inheritance">Transgenerational epigenetic inheritance</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Evolutionary_biology" title="Category:Evolutionary biology">Category</a></li> <li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" class="mw-file-description" title="Portal"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/16px-Symbol_portal_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/23px-Symbol_portal_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/31px-Symbol_portal_class.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span> <a href="/wiki/Portal:Evolutionary_biology" title="Portal:Evolutionary biology">Portal</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Emotions_(list)134" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Emotion_navbox" title="Template:Emotion navbox"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Emotion_navbox" title="Template talk:Emotion navbox"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Emotion_navbox" title="Special:EditPage/Template:Emotion navbox"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Emotions_(list)134" style="font-size:114%;margin:0 4em"><a href="/wiki/Emotion" title="Emotion">Emotions</a> (<a href="/wiki/Emotion_classification" title="Emotion classification">list</a>)</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Emotion" title="Emotion">Emotions</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acceptance" title="Acceptance">Acceptance</a></li> <li><a href="/wiki/Admiration" title="Admiration">Admiration</a></li> <li><a href="/wiki/Adoration" title="Adoration">Adoration</a></li> <li><a href="/wiki/Aesthetic_emotions" title="Aesthetic emotions">Aesthetic</a></li> <li><a href="/wiki/Affection" title="Affection">Affection</a></li> <li><a href="/wiki/Psychomotor_agitation" title="Psychomotor agitation">Agitation</a></li> <li><a href="/wiki/Pain" title="Pain">Agony</a></li> <li><a href="/wiki/Amusement" title="Amusement">Amusement</a></li> <li><a href="/wiki/Anger" title="Anger">Anger</a></li> <li><a href="/wiki/Angst" title="Angst">Angst</a></li> <li><a href="/wiki/Anguish" title="Anguish">Anguish</a></li> <li><a href="/wiki/Annoyance" title="Annoyance">Annoyance</a></li> <li><a href="/wiki/Anticipation" title="Anticipation">Anticipation</a></li> <li><a href="/wiki/Anxiety" title="Anxiety">Anxiety</a></li> <li><a href="/wiki/Apathy" title="Apathy">Apathy</a></li> <li><a href="/wiki/Arousal" title="Arousal">Arousal</a></li> <li><a href="/wiki/Interpersonal_attraction" title="Interpersonal attraction">Attraction</a></li> <li><a href="/wiki/Awe" title="Awe">Awe</a></li> <li><a href="/wiki/Belongingness" title="Belongingness">Belongingness</a></li> <li><a href="/wiki/Boredom" title="Boredom">Boredom</a></li> <li><a href="/wiki/Calmness" title="Calmness">Calmness</a></li> <li><a href="/wiki/Compassion" title="Compassion">Compassion</a></li> <li><a href="/wiki/Confidence" title="Confidence">Confidence</a></li> <li><a href="/wiki/Confusion" title="Confusion">Confusion</a></li> <li><a href="/wiki/Contempt" title="Contempt">Contempt</a></li> <li><a href="/wiki/Contentment" title="Contentment">Contentment</a></li> <li><a href="/wiki/Courage" title="Courage">Courage</a></li> <li><a href="/wiki/Cruelty" title="Cruelty">Cruelty</a></li> <li><a href="/wiki/Curiosity" title="Curiosity">Curiosity</a></li> <li><a href="/wiki/Defeatism" title="Defeatism">Defeat</a></li> <li><a href="/wiki/Depression_(mood)" title="Depression (mood)">Depression</a></li> <li><a href="/wiki/Desire" title="Desire">Desire</a></li> <li><a href="/wiki/Disappointment" title="Disappointment">Disappointment</a></li> <li><a href="/wiki/Disgust" title="Disgust">Disgust</a></li> <li><a href="/wiki/Distrust" title="Distrust">Distrust</a></li> <li><a href="/wiki/Doubt" title="Doubt">Doubt</a></li> <li><a href="/wiki/Ecstasy_(emotion)" title="Ecstasy (emotion)">Ecstasy</a></li> <li><a href="/wiki/Embarrassment" title="Embarrassment">Embarrassment</a> <ul><li><a href="/wiki/Vicarious_embarrassment" title="Vicarious embarrassment">vicarious</a></li></ul></li> <li><a href="/wiki/Emotion_work" title="Emotion work">Emotion work</a></li> <li><a href="/wiki/Empathy" title="Empathy">Empathy</a></li> <li><a href="/wiki/Emptiness" title="Emptiness">Emptiness</a></li> <li><a href="/wiki/Attention" title="Attention">Enthrallment</a></li> <li><a href="/wiki/Enthusiasm" title="Enthusiasm">Enthusiasm</a></li> <li><a href="/wiki/Envy" title="Envy">Envy</a></li> <li><a href="/wiki/Euphoria" title="Euphoria">Euphoria</a></li> <li><a href="/wiki/Stimulation" title="Stimulation">Excitement</a></li> <li><a href="/wiki/Faith" title="Faith">Faith</a></li> <li><a href="/wiki/Fear" title="Fear">Fear</a></li> <li><a href="/wiki/Flow_(psychology)" title="Flow (psychology)">Flow</a></li> <li><a href="/wiki/Frustration" title="Frustration">Frustration</a></li> <li><a href="/wiki/Gratification" title="Gratification">Gratification</a></li> <li><a href="/wiki/Gratitude" title="Gratitude">Gratitude</a></li> <li><a href="/wiki/Greed" title="Greed">Greed</a></li> <li><a href="/wiki/Grief" title="Grief">Grief</a></li> <li><a href="/wiki/Guilt_(emotion)" title="Guilt (emotion)">Guilt</a></li> <li><a href="/wiki/Happiness" title="Happiness">Happiness</a> <ul><li><i><a href="/wiki/Joie_de_vivre" title="Joie de vivre">Joie de vivre</a></i></li></ul></li> <li><a href="/wiki/Hatred" title="Hatred">Hatred</a></li> <li><span title="Welsh-language text"><i lang="cy"><a href="/wiki/Hiraeth" title="Hiraeth">Hiraeth</a></i></span></li> <li><a href="/wiki/Homesickness" title="Homesickness">Homesickness</a></li> <li><a href="/wiki/Hope" title="Hope">Hope</a></li> <li><a href="/wiki/Horror_and_terror" title="Horror and terror">Horror</a></li> <li><a href="/wiki/Hostility" title="Hostility">Hostility</a></li> <li><a href="/wiki/Humiliation" title="Humiliation">Humiliation</a></li> <li><span title="Danish-language text"><i lang="da"><a href="/wiki/Hygge" title="Hygge">Hygge</a></i></span></li> <li><a href="/wiki/Hysteria" title="Hysteria">Hysteria</a></li> <li><a href="/wiki/Ikigai" title="Ikigai"><i>Ikigai</i> (sense of purpose)</a></li> <li><a href="/wiki/Hedonism" title="Hedonism">Indulgence</a></li> <li><a href="/wiki/Infatuation" title="Infatuation">Infatuation</a></li> <li><a href="/wiki/Insecurity_(emotion)" title="Insecurity (emotion)">Insecurity</a></li> <li><a href="/wiki/Artistic_inspiration" title="Artistic inspiration">Inspiration</a></li> <li><a href="/wiki/Interest_(emotion)" title="Interest (emotion)">Interest</a></li> <li><a href="/wiki/Irritability" title="Irritability">Irritation</a></li> <li><a href="/wiki/Isolation_(psychology)" title="Isolation (psychology)">Isolation</a></li> <li><a href="/wiki/Jealousy" title="Jealousy">Jealousy</a></li> <li><a href="/wiki/Joy" title="Joy">Joy</a></li> <li><a href="/wiki/Kindness" title="Kindness">Kindness</a></li> <li><a href="/wiki/Loneliness" title="Loneliness">Loneliness</a></li> <li><a href="/wiki/Love" title="Love">Love</a> <ul><li><a href="/wiki/Limerence" title="Limerence">limerence</a></li> <li><a href="/wiki/Love_at_first_sight" title="Love at first sight">at first sight</a></li></ul></li> <li><a href="/wiki/Lust" title="Lust">Lust</a></li> <li><span title="Japanese-language text"><i lang="ja-Latn"><a href="/wiki/Mono_no_aware" title="Mono no aware">Mono no aware</a></i></span></li> <li><a href="/wiki/Neglect" title="Neglect">Neglect</a></li> <li><a href="/wiki/Nostalgia" title="Nostalgia">Nostalgia</a></li> <li><a href="/wiki/Outrage_(emotion)" title="Outrage (emotion)">Outrage</a></li> <li><a href="/wiki/Panic" title="Panic">Panic</a></li> <li><a href="/wiki/Passion_(emotion)" title="Passion (emotion)">Passion</a></li> <li><a href="/wiki/Pity" title="Pity">Pity</a> <ul><li><a href="/wiki/Self-pity" title="Self-pity">self-pity</a></li></ul></li> <li><a href="/wiki/Pleasure" title="Pleasure">Pleasure</a></li> <li><a href="/wiki/Pride" title="Pride">Pride</a> <ul><li><a href="/wiki/Grandiosity" title="Grandiosity">grandiosity</a></li> <li><a href="/wiki/Hubris" title="Hubris">hubris</a></li> <li><a href="/wiki/Insult" title="Insult">insult</a></li> <li><a href="/wiki/Vanity" title="Vanity">vanity</a></li></ul></li> <li><a href="/wiki/Rage_(emotion)" title="Rage (emotion)">Rage</a></li> <li><a href="/wiki/Regret" title="Regret">Regret</a></li> <li><a href="/wiki/Social_rejection" title="Social rejection">Rejection</a></li> <li><a href="/wiki/Relief_(emotion)" title="Relief (emotion)">Relief</a></li> <li><a href="/wiki/Remorse" title="Remorse">Remorse</a></li> <li><a href="/wiki/Resentment" title="Resentment">Resentment</a></li> <li><a href="/wiki/Revenge" title="Revenge">Revenge</a></li> <li><a href="/wiki/Sadness" title="Sadness">Sadness</a> <ul><li><a href="/wiki/Melancholia" title="Melancholia">melancholy</a></li></ul></li> <li><span title="Portuguese-language text"><i lang="pt"><a href="/wiki/Saudade" title="Saudade">Saudade</a></i></span></li> <li><span title="German-language text"><i lang="de"><a href="/wiki/Schadenfreude" title="Schadenfreude">Schadenfreude</a></i></span></li> <li><span title="German-language text"><i lang="de"><a href="/wiki/Sehnsucht" title="Sehnsucht">Sehnsucht</a></i></span></li> <li><a href="/wiki/Sentimentality" title="Sentimentality">Sentimentality</a></li> <li><a href="/wiki/Shame" title="Shame">Shame</a></li> <li><a href="/wiki/Acute_stress_reaction" title="Acute stress reaction">Shock</a></li> <li><a href="/wiki/Shyness" title="Shyness">Shyness</a></li> <li><a href="/wiki/Social_connection" title="Social connection">Social connection</a></li> <li><a href="/wiki/Sorrow_(emotion)" title="Sorrow (emotion)">Sorrow</a></li> <li><a href="/wiki/Spite_(sentiment)" title="Spite (sentiment)">Spite</a></li> <li><a href="/wiki/Psychological_stress" title="Psychological stress">Stress</a> <ul><li><a href="/wiki/Chronic_stress" title="Chronic stress">chronic</a></li></ul></li> <li><a href="/wiki/Suffering" title="Suffering">Suffering</a></li> <li><a href="/wiki/Surprise_(emotion)" title="Surprise (emotion)">Surprise</a></li> <li><a href="/wiki/Sympathy" title="Sympathy">Sympathy</a></li> <li><a href="/wiki/Trust_(social_science)" title="Trust (social science)">Trust</a></li> <li><a href="/wiki/Wonder_(emotion)" title="Wonder (emotion)">Wonder</a> <ul><li><a href="/wiki/Sense_of_wonder" title="Sense of wonder">sense of wonder</a></li></ul></li> <li><a href="/wiki/Worry" title="Worry">Worry</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="3" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Plutchik-wheel.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Plutchik-wheel.svg/90px-Plutchik-wheel.svg.png" decoding="async" width="90" height="91" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Plutchik-wheel.svg/135px-Plutchik-wheel.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/ce/Plutchik-wheel.svg/180px-Plutchik-wheel.svg.png 2x" data-file-width="715" data-file-height="725" /></a></span><br /><span typeof="mw:File"><a href="/wiki/File:Plutchik_Dyads.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Plutchik_Dyads.svg/90px-Plutchik_Dyads.svg.png" decoding="async" width="90" height="90" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Plutchik_Dyads.svg/135px-Plutchik_Dyads.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Plutchik_Dyads.svg/180px-Plutchik_Dyads.svg.png 2x" data-file-width="1357" data-file-height="1356" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Worldview" title="Worldview">Worldviews</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cynicism_(contemporary)" title="Cynicism (contemporary)">Cynicism</a></li> <li><a href="/wiki/Defeatism" title="Defeatism">Defeatism</a></li> <li><a href="/wiki/Fatalism" title="Fatalism">Fatalism</a></li> <li><a href="/wiki/Misanthropy" title="Misanthropy">Misanthropy</a></li> <li><a href="/wiki/Nihilism" title="Nihilism">Nihilism</a></li> <li><a href="/wiki/Optimism" title="Optimism">Optimism</a></li> <li><a href="/wiki/Pessimism" title="Pessimism">Pessimism</a></li> <li><a href="/wiki/Recluse" title="Recluse">Reclusion</a></li> <li><span title="German-language text"><i lang="de"><a href="/wiki/Weltschmerz" title="Weltschmerz">Weltschmerz</a></i></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Related</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Affect <ul><li><a href="/wiki/Affect_consciousness" title="Affect consciousness">consciousness</a></li> <li><a href="/wiki/Affect_(education)" title="Affect (education)">in education</a></li> <li><a href="/wiki/Affect_measures" title="Affect measures">measures</a></li> <li><a href="/wiki/Affect_(psychology)" title="Affect (psychology)">in psychology</a></li></ul></li> <li>Affective <ul><li><a href="/wiki/Affective_computing" title="Affective computing">computing</a></li> <li><a href="/wiki/Affective_forecasting" title="Affective forecasting">forecasting</a></li> <li><a href="/wiki/Affective_neuroscience" title="Affective neuroscience">neuroscience</a></li> <li><a href="/wiki/Affective_science" title="Affective science">science</a></li> <li><a href="/wiki/Affective_spectrum" title="Affective spectrum">spectrum</a></li></ul></li> <li>Affectivity <ul><li><a href="/wiki/Positive_affectivity" title="Positive affectivity">positive</a></li> <li><a href="/wiki/Negative_affectivity" title="Negative affectivity">negative</a></li></ul></li> <li><a href="/wiki/Appeal_to_emotion" title="Appeal to emotion">Appeal to emotion</a></li> <li><a href="/wiki/Amygdala_hijack" title="Amygdala hijack">Amygdala hijack</a></li> <li>Emotion <ul><li><a href="/wiki/Art_and_emotion" title="Art and emotion">and art</a></li> <li><a href="/wiki/Emotion_and_memory" title="Emotion and memory">and memory</a></li> <li><a href="/wiki/Music_and_emotion" title="Music and emotion">and music</a></li> <li><a href="/wiki/Sex_differences_in_psychology" title="Sex differences in psychology">and sex</a></li> <li><a href="/wiki/Sleep_and_emotions" title="Sleep and emotions">and sleep</a></li> <li><a href="/wiki/Emotion_classification" title="Emotion classification">classification</a></li> <li><a class="mw-selflink selflink">evolution</a></li> <li><a href="/wiki/Expressed_emotion" title="Expressed emotion">expressed</a></li> <li><a href="/wiki/Functional_accounts_of_emotion" title="Functional accounts of emotion">functional accounts</a></li> <li><a href="/wiki/Group_emotion" title="Group emotion">group</a></li> <li><a href="/wiki/Homeostatic_emotion" class="mw-redirect" title="Homeostatic emotion">homeostatic</a></li> <li><a href="/wiki/Emotion_in_animals" title="Emotion in animals">in animals</a></li> <li><a href="/wiki/Emotion_perception" title="Emotion perception">perception</a></li> <li><a href="/wiki/Emotion_recognition" title="Emotion recognition">recognition</a> <ul><li><a href="/wiki/Emotion_recognition_in_conversation" title="Emotion recognition in conversation">in conversation</a></li></ul></li> <li><a href="/wiki/Emotional_self-regulation" title="Emotional self-regulation">regulation</a> <ul><li><a href="/wiki/Interpersonal_emotion_regulation" title="Interpersonal emotion regulation">interpersonal</a></li></ul></li> <li><a href="/wiki/Emotion_work" title="Emotion work">work</a></li></ul></li> <li>Emotional <ul><li><a href="/wiki/Emotional_aperture" title="Emotional aperture">aperture</a></li> <li><a href="/wiki/Emotional_bias" title="Emotional bias">bias</a></li> <li><a href="/wiki/Emotional_blackmail" title="Emotional blackmail">blackmail</a></li> <li><a href="/wiki/Emotional_competence" title="Emotional competence">competence</a></li> <li><a href="/wiki/Emotional_conflict" title="Emotional conflict">conflict</a></li> <li><a href="/wiki/Emotional_contagion" title="Emotional contagion">contagion</a></li> <li><a href="/wiki/Emotional_detachment" title="Emotional detachment">detachment</a></li> <li><a href="/wiki/Emotional_dysregulation" title="Emotional dysregulation">dysregulation</a></li> <li><a href="/wiki/Emotional_eating" title="Emotional eating">eating</a></li> <li><a href="/wiki/Emotional_exhaustion" title="Emotional exhaustion">exhaustion</a></li> <li><a href="/wiki/Emotional_expression" title="Emotional expression">expression</a> <ul><li><a href="/wiki/Gender_and_emotional_expression" title="Gender and emotional expression">and gender</a></li></ul></li> <li><a href="/wiki/Emotional_intelligence" title="Emotional intelligence">intelligence</a> <ul><li><a href="/wiki/Bullying_and_emotional_intelligence" title="Bullying and emotional intelligence">and bullying</a></li> <li><a href="/wiki/Empathy_quotient" title="Empathy quotient">Empathy quotient</a></li></ul></li> <li><a href="/wiki/Emotional_intimacy" title="Emotional intimacy">intimacy</a></li> <li><a href="/wiki/Emotional_isolation" title="Emotional isolation">isolation</a></li> <li><a href="/wiki/Emotional_lability" title="Emotional lability">lability</a></li> <li><a href="/wiki/Emotional_labor" title="Emotional labor">labor</a></li> <li><a href="/wiki/Emotional_lateralization" title="Emotional lateralization">lateralization</a></li> <li><a href="/wiki/Emotional_literacy" title="Emotional literacy">literacy</a></li> <li><a href="/wiki/Emotional_prosody" title="Emotional prosody">prosody</a></li> <li><a href="/wiki/Emotional_reasoning" title="Emotional reasoning">reasoning</a></li> <li><a href="/wiki/Emotional_responsivity" title="Emotional responsivity">responsivity</a></li> <li><a href="/wiki/Emotional_security" class="mw-redirect" title="Emotional security">security</a></li> <li><a href="/wiki/Emotional_symbiosis" title="Emotional symbiosis">symbiosis</a></li> <li><a href="/wiki/Emotional_thought_method" title="Emotional thought method">thought method</a></li> <li><a href="/wiki/Emotional_well-being" class="mw-redirect" title="Emotional well-being">well-being</a></li></ul></li> <li><a href="/wiki/Emotionality" title="Emotionality">Emotionality</a> <ul><li><a href="/wiki/Bounded_emotionality" title="Bounded emotionality">bounded</a></li></ul></li> <li>Emotions <ul><li><a href="/wiki/Emotions_and_culture" title="Emotions and culture">and culture</a></li> <li><a href="/wiki/History_of_emotions" title="History of emotions">history</a></li> <li><a href="/wiki/Emotions_in_decision-making" title="Emotions in decision-making">in decision-making</a></li> <li><a href="/wiki/Emotions_in_the_workplace" title="Emotions in the workplace">in the workplace</a></li> <li><a href="/wiki/Emotions_in_virtual_communication" title="Emotions in virtual communication">in virtual communication</a></li> <li><a href="/wiki/Moral_emotions" title="Moral emotions">moral</a></li> <li><a href="/wiki/Self-conscious_emotions" title="Self-conscious emotions">self-conscious</a></li> <li><a href="/wiki/Social_emotions" title="Social emotions">social</a></li> <li><a href="/wiki/Social_sharing_of_emotions" title="Social sharing of emotions">social sharing</a></li> <li><a href="/wiki/Sociology_of_emotions" title="Sociology of emotions">sociology</a></li></ul></li> <li><a href="/wiki/Feeling" title="Feeling">Feeling</a></li> <li><a href="/wiki/Group_affective_tone" title="Group affective tone">Group affective tone</a></li> <li><a href="/wiki/Interactions_between_the_emotional_and_executive_brain_systems" title="Interactions between the emotional and executive brain systems">Interactions between the emotional and executive brain systems</a></li> <li><a href="/wiki/Jealousy_in_art" title="Jealousy in art">Jealousy in art</a></li> <li><a href="/wiki/Meta-emotion" title="Meta-emotion">Meta-emotion</a></li> <li><a href="/wiki/Pathognomy" title="Pathognomy">Pathognomy</a></li> <li><a href="/wiki/Pathos" title="Pathos">Pathos</a></li> <li><a href="/wiki/Social_emotional_development" title="Social emotional development">Social emotional development</a></li> <li><a href="/wiki/Stoic_passions" title="Stoic passions">Stoic passions</a></li> <li>Theory <ul><li><a href="/wiki/Affect_theory" title="Affect theory">affect</a></li> <li><a href="/wiki/Appraisal_theory" title="Appraisal theory">appraisal</a></li> <li><a href="/wiki/Theory_of_constructed_emotion" title="Theory of constructed emotion">constructed emotion</a></li> <li><a href="/wiki/Discrete_emotion_theory" title="Discrete emotion theory">discrete emotion</a></li> <li><a href="/wiki/Somatic_marker_hypothesis" title="Somatic marker hypothesis">somatic marker</a></li></ul></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div><i>Italics</i> indicate emotion names in foreign languages</div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐84749c7844‐55gsn Cached time: 20250207234445 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 0.489 seconds Real time usage: 0.623 seconds Preprocessor visited node count: 1415/1000000 Post‐expand include size: 157308/2097152 bytes Template argument size: 4364/2097152 bytes Highest expansion depth: 8/100 Expensive parser function count: 2/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 89416/5000000 bytes Lua time usage: 0.285/10.000 seconds Lua memory usage: 17586707/52428800 bytes Number of Wikibase entities loaded: 0/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 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