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class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="Lloc"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="Menú principal" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">Menú principal</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">Menú principal</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">mou a la barra lateral</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">amaga</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> Navegació </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage-description" class="mw-list-item"><a href="/wiki/Portada" title="Visiteu la pàgina principal [z]" accesskey="z"><span>Portada</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/Especial:Article_aleatori" title="Carrega una pàgina a l’atzar [x]" accesskey="x"><span>Article a l'atzar</span></a></li><li id="n-Articles-de-qualitat" class="mw-list-item"><a href="/wiki/Viquip%C3%A8dia:Articles_de_qualitat"><span>Articles de qualitat</span></a></li> </ul> </div> </div> <div id="p-Comunitat" class="vector-menu mw-portlet mw-portlet-Comunitat" > <div class="vector-menu-heading"> Comunitat </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-portal" class="mw-list-item"><a href="/wiki/Viquip%C3%A8dia:Portal" title="Sobre el projecte, què podeu fer, on trobareu les coses"><span>Portal viquipedista</span></a></li><li id="n-Agenda-d'actes" class="mw-list-item"><a href="/wiki/Viquip%C3%A8dia:Trobades"><span>Agenda d'actes</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/Especial:Canvis_recents" title="Una llista dels canvis recents al wiki [r]" accesskey="r"><span>Canvis recents</span></a></li><li id="n-La-taverna" class="mw-list-item"><a href="/wiki/Viquip%C3%A8dia:La_taverna"><span>La taverna</span></a></li><li id="n-contactpage" class="mw-list-item"><a href="/wiki/Viquip%C3%A8dia:Contacte"><span>Contacte</span></a></li><li id="n-Xat" class="mw-list-item"><a href="/wiki/Viquip%C3%A8dia:Canals_IRC"><span>Xat</span></a></li><li id="n-help" class="mw-list-item"><a href="/wiki/Viquip%C3%A8dia:Ajuda" title="El lloc per a saber més coses"><span>Ajuda</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/Portada" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Viquipèdia" src="/static/images/mobile/copyright/wikipedia-wordmark-ca.svg" style="width: 7.5em; height: 1.4375em;"> <img class="mw-logo-tagline" alt="l'Enciclopèdia Lliure" src="/static/images/mobile/copyright/wikipedia-tagline-ca.svg" width="120" height="14" style="width: 7.5em; height: 0.875em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/Especial:Cerca" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Cerca a la Viquipèdia [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>Cerca</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Cerca a Viquipèdia" aria-label="Cerca a Viquipèdia" autocapitalize="sentences" title="Cerca a la Viquipèdia [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="Especial:Cerca"> </div> <button class="cdx-button cdx-search-input__end-button">Cerca</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="Eines personals"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="Aparença"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="Change the appearance of the page's font size, width, and color" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="Aparença" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">Aparença</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="//donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_ca.wikipedia.org&uselang=ca" class=""><span>Donatius</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Especial:Crea_compte&returnto=Termolisina" title="Us animem a crear un compte i iniciar una sessió, encara que no és obligatori" class=""><span>Crea un compte</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Especial:Registre_i_entrada&returnto=Termolisina" title="Us animem a registrar-vos, però no és obligatori [o]" accesskey="o" class=""><span>Inicia la sessió</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="Més opcions" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="Eines personals" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">Eines personals</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="Menú d'usuari" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="//donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_ca.wikipedia.org&uselang=ca"><span>Donatius</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Especial:Crea_compte&returnto=Termolisina" title="Us animem a crear un compte i iniciar una sessió, encara que no és obligatori"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>Crea un compte</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Especial:Registre_i_entrada&returnto=Termolisina" title="Us animem a registrar-vos, però no és obligatori [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Inicia la sessió</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Pàgines per a editors no registrats <a href="/wiki/Ajuda:Introducci%C3%B3" aria-label="Vegeu més informació sobre l'edició"><span>més informació</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Especial:Contribucions_pr%C3%B2pies" title="Una llista de les modificacions fetes des d'aquesta adreça IP [y]" accesskey="y"><span>Contribucions</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Especial:Discussi%C3%B3_personal" title="Discussió sobre les edicions per aquesta adreça ip. [n]" accesskey="n"><span>Discussió per aquest IP</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Lloc"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Contingut" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Contingut</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">mou a la barra lateral</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">amaga</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">Inici</div> </a> </li> <li id="toc-Síntesi" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Síntesi"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Síntesi</span> </div> </a> <ul id="toc-Síntesi-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Estructura" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Estructura"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Estructura</span> </div> </a> <ul id="toc-Estructura-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Referències" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Referències"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Referències</span> </div> </a> <ul id="toc-Referències-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Enllaços_externs" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Enllaços_externs"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Enllaços externs</span> </div> </a> <ul id="toc-Enllaços_externs-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contingut" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Commuta la taula de continguts." > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Commuta la taula de continguts.</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Termolisina</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Vés a un article en una altra llengua. Disponible en 7 llengües" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-7" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">7 llengües</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Thermolysin" title="Thermolysin - alemany" lang="de" hreflang="de" data-title="Thermolysin" data-language-autonym="Deutsch" data-language-local-name="alemany" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Thermolysin" title="Thermolysin - anglès" lang="en" hreflang="en" data-title="Thermolysin" data-language-autonym="English" data-language-local-name="anglès" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%86%E3%83%AB%E3%83%A2%E3%83%AA%E3%82%B7%E3%83%B3" title="テルモリシン - japonès" lang="ja" hreflang="ja" data-title="テルモリシン" data-language-autonym="日本語" data-language-local-name="japonès" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Termolisin" title="Termolisin - malai" lang="ms" hreflang="ms" data-title="Termolisin" data-language-autonym="Bahasa Melayu" data-language-local-name="malai" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Termolizin" title="Termolizin - serbocroat" lang="sh" hreflang="sh" data-title="Termolizin" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="serbocroat" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Termolizin" title="Termolizin - serbi" lang="sr" hreflang="sr" data-title="Termolizin" data-language-autonym="Српски / srpski" data-language-local-name="serbi" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Termolysin" title="Termolysin - suec" lang="sv" hreflang="sv" data-title="Termolysin" data-language-autonym="Svenska" data-language-local-name="suec" class="interlanguage-link-target"><span>Svenska</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q409163#sitelinks-wikipedia" title="Modifica enllaços interlingües" class="wbc-editpage">Modifica els enllaços</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Espais de noms"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Termolisina" title="Vegeu el contingut de la pàgina [c]" accesskey="c"><span>Pàgina</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Discussi%C3%B3:Termolisina" rel="discussion" title="Discussió sobre el contingut d'aquesta pàgina [t]" accesskey="t"><span>Discussió</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Canvia la variant de llengua" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">català</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Vistes"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Termolisina"><span>Mostra</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Termolisina&action=edit" title="Modifica el codi font d'aquesta pàgina [e]" accesskey="e"><span>Modifica</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Termolisina&action=history" title="Versions antigues d'aquesta pàgina [h]" accesskey="h"><span>Mostra l'historial</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Eines de la pàgina"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Eines" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Eines</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Eines</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">mou a la barra lateral</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">amaga</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="Més opcions" > <div class="vector-menu-heading"> Accions </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Termolisina"><span>Mostra</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Termolisina&action=edit" title="Modifica el codi font d'aquesta pàgina [e]" accesskey="e"><span>Modifica</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Termolisina&action=history"><span>Mostra l'historial</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> General </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Especial:Enlla%C3%A7os/Termolisina" title="Una llista de totes les pàgines wiki que enllacen amb aquesta [j]" accesskey="j"><span>Què hi enllaça</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Especial:Seguiment/Termolisina" rel="nofollow" title="Canvis recents a pàgines enllaçades des d'aquesta pàgina [k]" accesskey="k"><span>Canvis relacionats</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Especial:P%C3%A0gines_especials" title="Llista totes les pàgines especials [q]" accesskey="q"><span>Pàgines especials</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Termolisina&oldid=34191220" title="Enllaç permanent a aquesta revisió de la pàgina"><span>Enllaç permanent</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Termolisina&action=info" title="Més informació sobre aquesta pàgina"><span>Informació de la pàgina</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Especial:Citau&page=Termolisina&id=34191220&wpFormIdentifier=titleform" title="Informació sobre com citar aquesta pàgina"><span>Citau aquest article</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Especial:UrlShortener&url=https%3A%2F%2Fca.wikipedia.org%2Fwiki%2FTermolisina"><span>Obtén una URL abreujada</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Especial:QrCode&url=https%3A%2F%2Fca.wikipedia.org%2Fwiki%2FTermolisina"><span>Descarrega el codi QR</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Imprimeix/exporta </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-create_a_book" class="mw-list-item"><a href="/w/index.php?title=Especial:Llibre&bookcmd=book_creator&referer=Termolisina"><span>Crea un llibre</span></a></li><li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Especial:DownloadAsPdf&page=Termolisina&action=show-download-screen"><span>Baixa com a PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Termolisina&printable=yes" title="Versió per a impressió d'aquesta pàgina [p]" accesskey="p"><span>Versió per a impressora</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> En altres projectes </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Thermolysin" hreflang="en"><span>Commons</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q409163" title="Enllaç a l'element del repositori de dades connectat [g]" accesskey="g"><span>Element a Wikidata</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Eines de la pàgina"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Aparença"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Aparença</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">mou a la barra lateral</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">amaga</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">De la Viquipèdia, l'enciclopèdia lliure</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="ca" dir="ltr"><table class="infobox" style="font-size:90%;width:25em"><caption style="font-weight:bold;background-color:#b0d1ad"><span style="float:left;"><span typeof="mw:File"><span title="Infotaula d'enzim"><img alt="Infotaula d'enzim" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d0/Noun_project_3067.svg/25px-Noun_project_3067.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d0/Noun_project_3067.svg/38px-Noun_project_3067.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d0/Noun_project_3067.svg/50px-Noun_project_3067.svg.png 2x" data-file-width="100" data-file-height="100" /></span></span></span>Termolisina</caption><tbody><tr><th colspan="2" class="infobox-header" style="text-align:center;background-color:#b0d1ad">Identificadors</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left;background-color:#eeeeee"><a href="/wiki/N%C3%BAmero_EC" title="Número EC">Número EC</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://enzyme.expasy.org/EC/3.4.24.27">3.4.24.27</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th colspan="2" class="infobox-header" style="text-align:center;background-color:#b0d1ad">Bases de dades</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left;background-color:#eeeeee"><a href="/wiki/IntEnz" title="IntEnz">IntEnz</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://www.ebi.ac.uk/intenz/query?cmd=SearchEC&ec=3.4.24.27">IntEnz view</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left;background-color:#eeeeee"><a href="/wiki/BRENDA" title="BRENDA">BRENDA</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://www.brenda-enzymes.org/enzyme.php?ecno=3.4.24.27">BRENDA entry</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left;background-color:#eeeeee"><a href="/wiki/ExPASy" title="ExPASy">ExPASy</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://www.expasy.org/enzyme/3.4.24.27">NiceZyme view</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" 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PDB</a><br /><a rel="nofollow" class="external text" href="https://pdbj.org/mine/sql?query=SELECT+DISTINCT+pdbid+FROM+entity+WHERE+entity.pdbx_ec%3D%273.4.24.27%27">PDBj</a><br /><a rel="nofollow" class="external text" href="https://www.ebi.ac.uk/pdbe/entry/search/index?ec_number:3.4.24.27">PDBe</a><br /><a rel="nofollow" class="external text" href="https://www.ebi.ac.uk/thornton-srv/databases/cgi-bin/enzymes/GetPage.pl?ec_number=3.4.24.27">PDBsum</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></span></td></tr><tr><td colspan="2" class="infobox-full-data infobox-data" style="text-align:center"><table class="infobox collapsible collapsed" style="font-size:90%;width:25em;float:none; clear:none; margin:0; border-width:0; border-collapse:collapse; text-align:left; width:100%"><tbody><tr><th colspan="2" class="infobox-header" style="text-align:center;background-color: #ddd">Recerca</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left;background-color: #e7dcc3; border:#fafafa 2px solid; border-width:3px 2px 0 0;"><a href="/wiki/PubMed_Central" title="PubMed Central">PMC</a></th><td class="infobox-data" style="background-color: #eee; border:#fafafa 2px solid; border-width:3px 0 0 2px;"><a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?db=pubmed&term3.4.24.27%5BEC/RN%20Number%5D%20AND%20pubmed%20pmc%20local%5Bsb%5D">articles</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left;background-color: #e7dcc3; border:#fafafa 2px solid; border-width:3px 2px 0 0;"><a href="/wiki/PubMed" title="PubMed">PubMed</a></th><td class="infobox-data" style="background-color: #eee; border:#fafafa 2px solid; border-width:3px 0 0 2px;"><a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/entrez/query.fcgi?db=pubmed&term=3.4.24.27%5BEC/RN%20Number%5D">articles</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left;background-color: #e7dcc3; border:#fafafa 2px solid; border-width:3px 2px 0 0;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a></th><td class="infobox-data" style="background-color: #eee; border:#fafafa 2px solid; border-width:3px 0 0 2px;"><a rel="nofollow" class="external text" href="http://www.ncbi.nlm.nih.gov/protein?term=3.4.24.27%5BEC/RN%20Number%5D">proteins</a> <span class="penicon" data-bridge-edit-flow="single-best-value"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q409163?uselang=ca#P591" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr></tbody></table></td></tr></tbody></table> <p>La <b>termolisina</b> és un <a href="/wiki/Enzim" title="Enzim">enzim</a> metaloproteinasa termoestable neutral produït pel bacteri grampositiu <i>Bacillus thermoproteolyticus</i>.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Aquest necessita un ió zinc per a l'activitat de l'enzim i quatre ions calci per a l'estabilitat estructural.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> La termolisina catalitza específicament la <a href="/wiki/Hidr%C3%B2lisi" title="Hidròlisi">hidròlisi</a> de l'<a href="/wiki/Enlla%C3%A7_pept%C3%ADdic" title="Enllaç peptídic">enllaç peptídic</a> que conté <a href="/wiki/Amino%C3%A0cids" class="mw-redirect" title="Aminoàcids">aminoàcids</a> hidròfobs. Però la termolisina és utilitzada sovint per a la formació d'enllaços peptídics per la reacció inversa d'hidròlisi.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> La termolisina és el membre més estable de la familia de les matelaoproteïnases produïdes per diverses espècies <i>Bacillus</i>. Aquests enzims són anomenats també proteinases “neutrals” o proteinases semblants a les termolisines (TLPs). </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Síntesi"><span id="S.C3.ADntesi"></span>Síntesi</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Termolisina&action=edit&section=1" title="Modifica la secció: Síntesi"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Com totes les proteases bacterianes extracel·lulars, la termolisina és sintetitzada primerament pel bacteri com a pre-proenzim.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> La termolisina és sintetitzada com un pre-proenzim que consisteix en un senyal peptídic de 28 aminoàcids, el propèptid té 204 aminoàcids i l'enzim madur té 316 aminoàcids de llargada. El senyal peptídic actua com un senyal de translocació de la pre-protermolisina cap a la <a href="/wiki/Membrana_citoplasm%C3%A0tica" class="mw-redirect" title="Membrana citoplasmàtica">membrana citoplasmàtica</a> del bacteri. En el periplasma, la pre-protermolisina és convertida a protermolisina per un senyal peptidasa. La proseqüència actua llavors com una xaperona molecular i condueix cap a l'autoescletxa de l'enllaç peptídic unint la seqüència pro i la madura. La proteïna madura és secretada al medi extracel·lular.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Estructura">Estructura</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Termolisina&action=edit&section=2" title="Modifica la secció: Estructura"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/Fitxer:Thermolysin_2A7G.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Thermolysin_2A7G.png/220px-Thermolysin_2A7G.png" decoding="async" width="220" height="184" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Thermolysin_2A7G.png/330px-Thermolysin_2A7G.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Thermolysin_2A7G.png/440px-Thermolysin_2A7G.png 2x" data-file-width="1362" data-file-height="1142" /></a><figcaption>Estructura de la termolisina</figcaption></figure> <p>La termolisina té un pes molecular de 34.600 Da. La seva estructura general consisteix en dos dominis aproximadament esfèrics amb una profunda escletxa al mig de la molècula, separant els dos dominis. L'estructura secundària de cada domini és força diferent; el domini N-terminal consisteix bàsicament d'una fulla plegada beta, mentre que el domini C-terminal és bàsicament una <a href="/wiki/H%C3%A8lix_alfa" title="Hèlix alfa">hèlix alfa</a>. Aquests dos dominis estan connectats per una hèlix alfa central, que atravessa els aminoàcids 137-151.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>En contrast amb altres proteïnes que pateixen canvis conformacionals per la temperatura i la desnaturalització, la termolisina no pateix cap canvi conformacional fins almenys els 70 °C.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> L'estabilitat tèrmica dels membres de la familia TLP és mesurada en termes de T50 temperatura. A aquesta temperatura d'incubació durant 30 minuts es redueix l'activitat dels enzims a la meitat. La termolisina té una T50 de 86,9 °C, fent d'ella el membre més termoestable de la familia de les TLP.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Estudis sobre la contribució del calci a l'estabilitat de la termolisina han mostrat que durant la inactivació tèrmica un ió de calci sol és alliberat de la molècula.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Evitar que aquest calci s'uneixi originàriament a la molècula per <a href="/wiki/Mutaci%C3%B3" title="Mutació">mutació</a> de la seva zona d'unió, redueix l'estabilitat de la termolisina per 7 °C. Tot i així, mentre que el calci realitza una significant contribució a l'estabilització de la termolisina, més crucial per a l'estabilitat és el petit cluster d'aminoàcids del domini N-terminal localitzat a les proteïnes de la superfície. En particular la <a href="/wiki/Fenilalanina" title="Fenilalanina">fenilalanina</a> (F) a l'aminoàcid de posició 63 i la <a href="/wiki/Prolina" title="Prolina">prolina</a> (P) a l'aminoàcid de posició 69 contribueixen de manera significativa a l'estabilitat de la termolisina. Canviant aquests aminoàcids a la treonina (T) i l'<a href="/wiki/Alanina" title="Alanina">alanina</a> (A) respectivament en una proteinasa com la termolisina menys estable produïda per Bacillus stearothermophillus (TLP-ste), desemboca en reduccions individuals en estabilitat de 7 °C (F63- T) i 6,3 °C (P69 – A) i quan combinen una reducció, en estabilitat de 12,3 °C. </p> <div class="mw-heading mw-heading2"><h2 id="Referències"><span id="Refer.C3.A8ncies"></span>Referències</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Termolisina&action=edit&section=3" title="Modifica la secció: Referències"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="reflist {{#if: | references-column-count references-column-count-{{{col}}}" style="list-style-type: decimal;"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text">Endo, S. (1962). "Studies on protease produced by thermophilic bacteria". J. Ferment. Technol. 40: 346–353.</span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text">Tajima M, Urabe I. et al. (1976). "Role of calcium ions in the thermostability of thermolysin and Bacillus subtilis var. amylosacchariticus neutral protease". Eur. J. Biochem. 64 (1): 243–247. doi:10.1111/j.1432-1033.1976.tb10293.x. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/819262?dopt=Abstract" class="extiw" title="pmid:819262">819262</a></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text">Trusek-Holownia A. (2003). "Synthesis of ZAlaPheOMe, the precursor of bitter dipeptide in the two-phase ethyl acetate-water system catalysed by thermolysin". J. Biotechnol. 102 (2): 153–163. doi:10.1016/S0168-1656(03)00024-5. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/12697393?dopt=Abstract" class="extiw" title="pmid:12697393">12697393</a></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text">Yasukawa K, Kusano M, Inouye K. (2007). "A new method for the extracellular production of recombinant thermolysin by co-expressing the mature sequence and pro-sequence in Escherichia coli". Protein Eng. Des. Sel. 20 (8): 375–383. doi:10.1093/protein/gzm031. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/17616558?dopt=Abstract" class="extiw" title="pmid:17616558">17616558</a></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><a href="#cite_ref-5">↑</a></span> <span class="reference-text">Inouye K, Kusano M. et al. (2007). "Engineering, expression, purification, and production of recombinant thermolysin". Biotechnol. Annu. Rev. 13: 43–64. doi:10.1016/S1387-2656(07)13003-9. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/17875473?dopt=Abstract" class="extiw" title="pmid:17875473">17875473</a></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><a href="#cite_ref-6">↑</a></span> <span class="reference-text">Holmes MA and Matthews BW. (1982). "Structure of thermolysin refined at 1.6 A resolution". J. Mol. Biol. 160 (4): 623–639. doi:10.1016/0022-2836(82)90319-9. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/7175940?dopt=Abstract" class="extiw" title="pmid:7175940">7175940</a></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><a href="#cite_ref-7">↑</a></span> <span class="reference-text">Matthews BW, Weaver LH and Kester WR. (1974). "The conformation of thermolysin". J. Biol. Chem. 249 (24): 8030–8044. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/4214815?dopt=Abstract" class="extiw" title="pmid:4214815">4214815</a></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><a href="#cite_ref-8">↑</a></span> <span class="reference-text">Eijsink VG, Veltman OR, et al. (1995). "Structural determinants of the stability of thermolysin-like proteinases". Nat. Struct. Biol. 2 (5): 374–379. doi:10.1038/nsb0595-374. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/7664094?dopt=Abstract" class="extiw" title="pmid:7664094">7664094</a></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><a href="#cite_ref-9">↑</a></span> <span class="reference-text">Dahlquist FW, Long JW and Bigbee WL (1976). "Role of Calcium in the thermal stability of thermolysin". Biochemistry 15 (5): 1103–1111. doi:10.1021/bi00650a024. <a href="/wiki/PubMed" title="PubMed">PMID</a>: <a href="https://www.ncbi.nlm.nih.gov/pubmed/814920?dopt=Abstract" class="extiw" title="pmid:814920">814920</a></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Enllaços_externs"><span id="Enlla.C3.A7os_externs"></span>Enllaços externs</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Termolisina&action=edit&section=4" title="Modifica la secció: Enllaços externs"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>MeSH Thermolysin <a rel="nofollow" class="external autonumber" href="http://www.nlm.nih.gov/cgi/mesh/2009/MB_cgi?mode=&term=Thermolysin">[1]</a>.</li></ul>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Obtingut de «<a dir="ltr" href="https://ca.wikipedia.org/w/index.php?title=Termolisina&oldid=34191220">https://ca.wikipedia.org/w/index.php?title=Termolisina&oldid=34191220</a>»</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Especial:Categorias" title="Especial:Categorias">Categoria</a>: <ul><li><a href="/wiki/Categoria:Peptidases" title="Categoria:Peptidases">Peptidases</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> La pàgina va ser modificada per darrera vegada el 2 nov 2024 a les 16:16.</li> <li id="footer-info-copyright">El text està disponible sota la <a href="/wiki/Viquip%C3%A8dia:Text_de_la_llic%C3%A8ncia_de_Creative_Commons_Reconeixement-Compartir_Igual_4.0_No_adaptada" title="Viquipèdia:Text de la llicència de Creative Commons Reconeixement-Compartir Igual 4.0 No adaptada"> Llicència de Creative Commons Reconeixement i Compartir-Igual</a>; es poden aplicar termes addicionals. 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