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Beach vole - Wikipedia
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Available in 14 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-14" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">14 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Catalan" lang="ca" hreflang="ca" data-title="Microtus breweri" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Cebuano" lang="ceb" hreflang="ceb" data-title="Microtus breweri" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Strandw%C3%BChlmaus" title="Strandwühlmaus – German" lang="de" hreflang="de" data-title="Strandwühlmaus" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Spanish" lang="es" hreflang="es" data-title="Microtus breweri" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Basque" lang="eu" hreflang="eu" data-title="Microtus breweri" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%95%B4%EB%B3%80%EB%B0%AD%EC%A5%90" title="해변밭쥐 – Korean" lang="ko" hreflang="ko" data-title="해변밭쥐" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-avk mw-list-item"><a href="https://avk.wikipedia.org/wiki/Tovukol_(Microtus_breweri)" title="Tovukol (Microtus breweri) – Kotava" lang="avk" hreflang="avk" data-title="Tovukol (Microtus breweri)" data-language-autonym="Kotava" data-language-local-name="Kotava" class="interlanguage-link-target"><span>Kotava</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%81%D9%88%D9%84%D9%89_%D8%A7%D9%84%D8%B4%D8%A7%D8%B7%D8%A6" title="فولى الشاطئ – Egyptian Arabic" lang="arz" hreflang="arz" data-title="فولى الشاطئ" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Strandwoelmuis" title="Strandwoelmuis – Dutch" lang="nl" hreflang="nl" data-title="Strandwoelmuis" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Serbian" lang="sr" hreflang="sr" data-title="Microtus breweri" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Swedish" lang="sv" hreflang="sv" data-title="Microtus breweri" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Ukrainian" lang="uk" hreflang="uk" data-title="Microtus breweri" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Vietnamese" lang="vi" hreflang="vi" data-title="Microtus breweri" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Microtus_breweri" title="Microtus breweri – Waray" lang="war" hreflang="war" data-title="Microtus breweri" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1765454#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul 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</th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Microtus_breweri.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Microtus_breweri.jpg/220px-Microtus_breweri.jpg" decoding="async" width="220" height="219" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Microtus_breweri.jpg/330px-Microtus_breweri.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/57/Microtus_breweri.jpg/440px-Microtus_breweri.jpg 2x" data-file-width="1170" data-file-height="1166" /></a></span> </td></tr> <tr style="color:inherit; text-align: center; background-color: rgb(235,235,210)"> <th colspan="2"><div style="text-align: center"><a href="/wiki/Conservation_status" title="Conservation status">Conservation status</a></div> </th></tr> <tr> <td colspan="2"><div style="text-align: center"><span class="mw-default-size skin-invert-image" typeof="mw:File/Frameless"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Status_iucn3.1_VU.svg/220px-Status_iucn3.1_VU.svg.png" decoding="async" width="220" height="59" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Status_iucn3.1_VU.svg/330px-Status_iucn3.1_VU.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Status_iucn3.1_VU.svg/440px-Status_iucn3.1_VU.svg.png 2x" data-file-width="512" data-file-height="137" /></span></span><br /><a href="/wiki/Vulnerable_species" title="Vulnerable species">Vulnerable</a> <small> (<a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN 3.1</a>)<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></small></div> </td></tr> <tr> <td colspan="2"><div style="text-align: center"><span class="mw-default-size skin-invert-image" typeof="mw:File/Frameless"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/39/Status_TNC_G1.svg/220px-Status_TNC_G1.svg.png" decoding="async" width="220" height="59" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/39/Status_TNC_G1.svg/330px-Status_TNC_G1.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/39/Status_TNC_G1.svg/440px-Status_TNC_G1.svg.png 2x" data-file-width="240" data-file-height="64" /></span></span><br />Critically Imperiled <small> (<a href="/wiki/NatureServe_conservation_status" title="NatureServe conservation status">NatureServe</a>)<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></small></div> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Microtus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Mammal" title="Mammal">Mammalia</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Rodent" title="Rodent">Rodentia</a> </td></tr> <tr> <td>Family: </td> <td><a href="/wiki/Cricetidae" title="Cricetidae">Cricetidae</a> </td></tr> <tr> <td>Subfamily: </td> <td><a href="/wiki/Arvicolinae" title="Arvicolinae">Arvicolinae</a> </td></tr> <tr> <td>Genus: </td> <td><a href="/wiki/Microtus" title="Microtus"><i>Microtus</i></a> </td></tr> <tr> <td>Species: </td> <td><div style="display:inline" class="species"><i><b>M. breweri</b></i></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">Binomial name</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><b><span class="binomial"><span style="font-weight:normal;"></span><i>Microtus breweri</i></span></b><br /><div style="font-size: 85%;"><a href="/wiki/Spencer_Fullerton_Baird" title="Spencer Fullerton Baird">Baird</a>, 1858</div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i>Microtus pennsylvanicus breweri</i></li> <li><i>Arvicola breweri</i></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p>The <b>beach vole</b> (<i><b>Microtus breweri</b></i>) or <b>Muskeget vole</b> is a <a href="/wiki/Rodent" title="Rodent">rodent</a> in the family <a href="/wiki/Cricetidae" title="Cricetidae">Cricetidae</a>. This close relative of the <a href="/wiki/Eastern_meadow_vole" title="Eastern meadow vole">eastern meadow vole</a> (<i>Microtus pennsylvanicus</i>) is <a href="/wiki/Endemic" class="mw-redirect" title="Endemic">endemic</a> to the 0.87 km<sup>2</sup> <a href="/wiki/Muskeget_Island" title="Muskeget Island">Muskeget Island</a>, <a href="/wiki/Massachusetts" title="Massachusetts">Massachusetts</a>. Due to its relatively short period of <a href="/wiki/Reproductive_isolation" title="Reproductive isolation">reproductive isolation</a>, there is debate over the beach vole's designation as a <a href="/wiki/Subspecies" title="Subspecies">subspecies</a> of <i>M. pennsylvanicus</i>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=1" title="Edit section: Taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Microtus breweri</i> is currently recognized by the IUCN, NatureServe and ITIS<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> as a distinct species, due to its larger size, geographic isolation, and contrasting mating habits.<sup id="cite_ref-Explore_the_Database_4-0" class="reference"><a href="#cite_note-Explore_the_Database-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> A recent genetic study indicates that it may be a <a href="/wiki/Basal_(phylogenetics)" title="Basal (phylogenetics)">basal</a> subspecies of <i>M. pennsylvanicus</i>.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=2" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Beach voles are typically much larger than meadow voles. On average, male beach voles are 192.9 mm long, ranging between 169 and 215 mm; with the tail making up approximately one-fourth of the total length.<sup id="cite_ref-Explore_the_Database_4-1" class="reference"><a href="#cite_note-Explore_the_Database-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Generally, females are smaller, with an average length of 181.2 mm, ranging between 165 and 201 mm.<sup id="cite_ref-Explore_the_Database_4-2" class="reference"><a href="#cite_note-Explore_the_Database-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> The weight of a beach vole ranges between 29 and 79 g, proportional to age, with females weighing less than males.<sup id="cite_ref-Explore_the_Database_4-3" class="reference"><a href="#cite_note-Explore_the_Database-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Compared to meadow voles, beach voles exhibit a lower <a href="/wiki/Metabolism" title="Metabolism">metabolism</a>, and their <a href="/wiki/Fur" title="Fur">coat</a> is made up of more guard hairs and fewer underhairs.<sup id="cite_ref-Explore_the_Database_4-4" class="reference"><a href="#cite_note-Explore_the_Database-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> A white blaze can often be found on the chin or forehead. </p><p>The <a href="/wiki/Skulls" class="mw-redirect" title="Skulls">skulls</a> of <i>M. breweri</i> can be easily distinguished from those of meadow voles. The average cranial length of beach voles is 13.1 mm, while the cranial breadth is generally 11.2 mm.<sup id="cite_ref-Explore_the_Database_4-5" class="reference"><a href="#cite_note-Explore_the_Database-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Their skull is larger, and their <a href="/wiki/Brain_case" class="mw-redirect" title="Brain case">brain case</a> is longer and narrower than meadow voles. Recent unpublished observations include a distinctly shaped <a href="/wiki/Interparietal_bone" title="Interparietal bone">interparietal bone</a> and a narrow bridge between the <a href="/wiki/Orbits" class="mw-redirect" title="Orbits">orbits</a>. </p><p>Beach voles tend closer towards <a href="/wiki/R/K_selection_theory" title="R/K selection theory">K selection</a> than meadow voles, and some of their distinctive features are indicative of <a href="/wiki/Island_gigantism" title="Island gigantism">island gigantism</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Distribution">Distribution</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=3" title="Edit section: Distribution"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Microtus breweri</i> can only be found on Muskeget Island, off the west coast of Nantucket, Massachusetts, in the <a href="/wiki/United_States" title="United States">United States</a>. However, historically it was also found on nearby South Point and Adams Islands, which are sometimes considered as a part of Muskeget Island, but are no longer above <a href="/wiki/Sea_level" title="Sea level">sea level</a>. The vole was <a href="/wiki/Extirpated" class="mw-redirect" title="Extirpated">extirpated</a> from Muskeget proper by 1891 as a result of <a href="/wiki/Predation" title="Predation">predation</a> by stray <a href="/wiki/Cats" class="mw-redirect" title="Cats">cats</a> (<i>Felis catus</i>), but was reintroduced by <a href="/wiki/Gerrit_S._Miller" class="mw-redirect" title="Gerrit S. Miller">Gerrit S. Miller</a>, <a href="/wiki/Outram_Bangs" title="Outram Bangs">Outram Bangs</a>, and Chas F. Batchelder in 1893 from a colony captured on South Point Island. Moreover, Muskeget Island has moved about 1,000 feet eastwardly over the last about 200 years, along with its changing process of shape, size and position due to <a href="/wiki/Erosion" title="Erosion">erosion</a> and tidal buildup.<sup id="cite_ref-Wetherbee_6-0" class="reference"><a href="#cite_note-Wetherbee-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Habitat_and_diet">Habitat and diet</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=4" title="Edit section: Habitat and diet"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>These voles dominate an open habitat of <a href="/wiki/Poison_ivy" title="Poison ivy">poison ivy</a> (<i>Toxicodendron radicans</i>) and <a href="/wiki/Ammophila_(plant)" title="Ammophila (plant)">beach grass</a> (<i>Ammophilia breviligulata</i>). Swaths of bare sand and <a href="/wiki/Salt_marsh" title="Salt marsh">saltwater marsh</a> can be found on the island, but the voles tend to avoid these areas. They may burrow in loose sand or loose soil under or near any of their shelters.<sup id="cite_ref-Wetherbee_6-1" class="reference"><a href="#cite_note-Wetherbee-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> <i> M. breweri</i> feeds on beach grass stalks, leaves, seeds, and insect adults and <a href="/wiki/Larvae" class="mw-redirect" title="Larvae">larvae</a>. Their predators include birds of prey such as the <a href="/wiki/Short-eared_owl" title="Short-eared owl">short-eared owl</a> (<i>Asio flammeus</i>) and the <a href="/wiki/Northern_harrier" title="Northern harrier">northern harrier</a> (<i>Circus hudsonius</i>). Cats were introduced onto Muskeget in the late 1800s, decimating the vole's population, but are no longer present. Their home range is usually less than one acre.<sup id="cite_ref-Wetherbee_6-2" class="reference"><a href="#cite_note-Wetherbee-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Tamarin_7-0" class="reference"><a href="#cite_note-Tamarin-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Reproduction_and_behavior">Reproduction and behavior</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=5" title="Edit section: Reproduction and behavior"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Young beach voles are born in the <a href="/wiki/Burrow" title="Burrow">burrow</a> <a href="/wiki/Nests" class="mw-redirect" title="Nests">nests</a>, underneath fragments of <a href="/wiki/Debris" title="Debris">wreckage</a>, or at the base of <a href="/wiki/Goldenrod" title="Goldenrod">goldenrod</a> (<i>Solidago</i> sp.). They do not demonstrate a typical vole breeding cycle, and show many of the attributes of a <a href="/wiki/R/K_selection_theory" title="R/K selection theory">K-selected organism</a>, such as large size, later age at maturation, <a href="/wiki/Sex_ratio" title="Sex ratio">sex ratio</a> weighted towards the males, and low reproductive output.<sup id="cite_ref-Robert_Tamarin_8-0" class="reference"><a href="#cite_note-Robert_Tamarin-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> The adults can breed from the spring to the fall, and their <a href="/wiki/Gestation" title="Gestation">gestation</a> lasts for about one month. Every year, an individual female beach vole typically produces two litters of three to five offspring, but most will live for less than one year.<sup id="cite_ref-Wetherbee_6-3" class="reference"><a href="#cite_note-Wetherbee-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>The beach voles have a habit of building runways above or in the ground under the beach grass, in order to better stay hidden from aerial predators. These runways may contain cut grass. During winter, underground tunnels become more common due to the cold weather.<sup id="cite_ref-Robert_Tamarin_8-1" class="reference"><a href="#cite_note-Robert_Tamarin-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=6" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFRoach,_N.2020" class="citation journal cs1">Roach, N. (2020). <a rel="nofollow" class="external text" href="https://www.iucnredlist.org/species/13417/22349291">"<i>Microtus breweri </i>"</a>. <i><a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN Red List of Threatened Species</a></i>. <b>2020</b>: e.T13417A22349291. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.2305%2FIUCN.UK.2020-2.RLTS.T13417A22349291.en">10.2305/IUCN.UK.2020-2.RLTS.T13417A22349291.en</a></span><span class="reference-accessdate">. Retrieved <span class="nowrap">19 January</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=IUCN+Red+List+of+Threatened+Species&rft.atitle=Microtus+breweri+&rft.volume=2020&rft.pages=e.T13417A22349291&rft.date=2020&rft_id=info%3Adoi%2F10.2305%2FIUCN.UK.2020-2.RLTS.T13417A22349291.en&rft.au=Roach%2C+N.&rft_id=https%3A%2F%2Fwww.iucnredlist.org%2Fspecies%2F13417%2F22349291&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://explorer.natureserve.org/Taxon/ELEMENT_GLOBAL.2.102648/Microtus_breweri">"<i>Microtus breweri</i>. NatureServe Explorer 2.0"</a>. <i>explorer.natureserve.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">19 January</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=explorer.natureserve.org&rft.atitle=Microtus+breweri.+NatureServe+Explorer+2.0&rft_id=https%3A%2F%2Fexplorer.natureserve.org%2FTaxon%2FELEMENT_GLOBAL.2.102648%2FMicrotus_breweri&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=180304#null">"Microtus breweri (Baird, 1857)"</a>. <i>ITIS</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2 February</span> 2022</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=ITIS&rft.atitle=Microtus+breweri+%28Baird%2C+1857%29&rft_id=https%3A%2F%2Fwww.itis.gov%2Fservlet%2FSingleRpt%2FSingleRpt%3Fsearch_topic%3DTSN%26search_value%3D180304%23null&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> <li id="cite_note-Explore_the_Database-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-Explore_the_Database_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Explore_the_Database_4-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Explore_the_Database_4-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Explore_the_Database_4-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Explore_the_Database_4-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-Explore_the_Database_4-5"><sup><i><b>f</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.mammaldiversity.org/explore.html#species-id=1002096">"Explore the Database"</a>. <i>www.mammaldiversity.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2021-07-09</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.mammaldiversity.org&rft.atitle=Explore+the+Database&rft_id=https%3A%2F%2Fwww.mammaldiversity.org%2Fexplore.html%23species-id%3D1002096&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFJacksonCook2020" class="citation journal cs1">Jackson, Donavan J; Cook, Joseph A (2020-02-21). <a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fjmammal%2Fgyz196">"A precarious future for distinctive peripheral populations of meadow voles (Microtus pennsylvanicus)"</a>. <i>Journal of Mammalogy</i>. <b>101</b> (1): 36–51. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fjmammal%2Fgyz196">10.1093/jmammal/gyz196</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0022-2372">0022-2372</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Mammalogy&rft.atitle=A+precarious+future+for+distinctive+peripheral+populations+of+meadow+voles+%28Microtus+pennsylvanicus%29&rft.volume=101&rft.issue=1&rft.pages=36-51&rft.date=2020-02-21&rft_id=info%3Adoi%2F10.1093%2Fjmammal%2Fgyz196&rft.issn=0022-2372&rft.aulast=Jackson&rft.aufirst=Donavan+J&rft.au=Cook%2C+Joseph+A&rft_id=https%3A%2F%2Fdoi.org%2F10.1093%252Fjmammal%252Fgyz196&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> <li id="cite_note-Wetherbee-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-Wetherbee_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Wetherbee_6-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Wetherbee_6-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Wetherbee_6-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDavid_Kenneth_WetherbeeRaymond_Parke_CoppingerRichard_E._Walsh1972" class="citation book cs1">David Kenneth Wetherbee; Raymond Parke Coppinger; Richard E. Walsh (1972). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=9ag1AAAAMAAJ"><i>Time Lapse Ecology, Muskeget Island, Nantucket, Massachusetts</i></a>. MSS Educational Pub. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780842201858" title="Special:BookSources/9780842201858"><bdi>9780842201858</bdi></a><span class="reference-accessdate">. Retrieved <span class="nowrap">28 December</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Time+Lapse+Ecology%2C+Muskeget+Island%2C+Nantucket%2C+Massachusetts&rft.pub=MSS+Educational+Pub.&rft.date=1972&rft.isbn=9780842201858&rft.au=David+Kenneth+Wetherbee&rft.au=Raymond+Parke+Coppinger&rft.au=Richard+E.+Walsh&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3D9ag1AAAAMAAJ&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> <li id="cite_note-Tamarin-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-Tamarin_7-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFTamarin1977" class="citation journal cs1">Tamarin, Robert H. (1977). "demography of the Beach Vole (<i>Microtus breweri</i>) and the Meadow Vole (<i>Microtus pennsylvanicus</i>) in Southeastern Massachusetts". <i>Ecology</i>. <b>58</b> (6): 1310–1321. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1977Ecol...58.1310T">1977Ecol...58.1310T</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F1935083">10.2307/1935083</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/1935083">1935083</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecology&rft.atitle=demography+of+the+Beach+Vole+%28Microtus+breweri%29+and+the+Meadow+Vole+%28Microtus+pennsylvanicus%29+in+Southeastern+Massachusetts&rft.volume=58&rft.issue=6&rft.pages=1310-1321&rft.date=1977&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F1935083%23id-name%3DJSTOR&rft_id=info%3Adoi%2F10.2307%2F1935083&rft_id=info%3Abibcode%2F1977Ecol...58.1310T&rft.aulast=Tamarin&rft.aufirst=Robert+H.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> <li id="cite_note-Robert_Tamarin-8"><span class="mw-cite-backlink">^ <a href="#cite_ref-Robert_Tamarin_8-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Robert_Tamarin_8-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFTamarin1977" class="citation journal cs1">Tamarin, Robert H. (1977). "Reproduction in the Island Beach Vole, <i>Microtus breweri</i>, and the Mainland Meadow Vole, <i>Microtus pennsylvanicus</i>, in Southeastern Massachusetts". <i>Journal of Mammalogy</i>. <b>58</b> (American Society of Mammalogists): 536–548. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F1380002">10.2307/1380002</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/1380002">1380002</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/336824">336824</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Mammalogy&rft.atitle=Reproduction+in+the+Island+Beach+Vole%2C+Microtus+breweri%2C+and+the+Mainland+Meadow+Vole%2C+Microtus+pennsylvanicus%2C+in+Southeastern+Massachusetts&rft.volume=58&rft.issue=American+Society+of+Mammalogists&rft.pages=536-548&rft.date=1977&rft_id=info%3Apmid%2F336824&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F1380002%23id-name%3DJSTOR&rft_id=info%3Adoi%2F10.2307%2F1380002&rft.aulast=Tamarin&rft.aufirst=Robert+H.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABeach+vole" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=7" title="Edit section: Further reading"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Musser, G. G. and M. D. Carleton. 2005. Superfamily Muroidea. pp. 894–1531 <i>in</i> Mammal Species of the World a Taxonomic and Geographic Reference. D. E. Wilson and D. M. Reeder eds. Johns Hopkins University Press, Baltimore.</li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Beach_vole&action=edit&section=8" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external free" href="https://training.fws.gov/library/pubs5/necas/web_link/39_muskeget.htm">http://training.fws.gov/library/pubs5/necas/web_link/39_muskeget.htm</a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline 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.navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r886047488"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r886047488"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r886047488"></div><div role="navigation" class="navbox" aria-labelledby="Extant_species_of_subfamily_Arvicolinae" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Arvicolinae" title="Template:Arvicolinae"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Arvicolinae" title="Template talk:Arvicolinae"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Arvicolinae" title="Special:EditPage/Template:Arvicolinae"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Extant_species_of_subfamily_Arvicolinae" style="font-size:114%;margin:0 4em">Extant species of subfamily <a href="/wiki/Arvicolinae" title="Arvicolinae">Arvicolinae</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Mammal" title="Mammal">Mammalia</a></li> <li>Infraclass: <a href="/wiki/Eutheria" title="Eutheria">Eutheria</a></li> <li>Superorder: <a href="/wiki/Euarchontoglires" title="Euarchontoglires">Euarchontoglires</a></li> <li>Order: <a href="/wiki/Rodent" title="Rodent">Rodentia</a></li> <li>Family: <a href="/wiki/Cricetidae" title="Cricetidae">Cricetidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Arvicolini" title="Arvicolini">Arvicolini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Arvicola(Water_voles)" scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Arvicola" title="Arvicola">Arvicola</a></i><br /><small><span style="color:#696969"><span class="nobold">(Water voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/European_water_vole" title="European water vole">European water vole (<i>A. amphibius</i>)</a></li> <li><a href="/wiki/Southwestern_water_vole" title="Southwestern water vole">Southwestern water vole (<i>A. sapidus</i>)</a></li> <li><a href="/wiki/Montane_water_vole" title="Montane water vole">Montane water vole (<i>A. scherman</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Microtini" title="Microtini">Microtini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Alexandromys" title="Alexandromys">Alexandromys</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Clarke%27s_vole" title="Clarke's vole">Clarke's vole (<i>M. clarkei</i>)</a></li> <li><a href="/wiki/Evorsk_vole" title="Evorsk vole">Evorsk vole (<i>M. evoronensis</i>)</a></li> <li><a href="/wiki/Reed_vole" title="Reed vole">Reed vole (<i>M. fortis</i>)</a></li> <li><a href="/wiki/Gerbe%27s_vole" title="Gerbe's vole">Gerbe's vole (<i>M. gerbei</i>)</a></li> <li><a href="/wiki/Taiwan_vole" title="Taiwan vole">Taiwan vole (<i>M. kikuchii</i>)</a></li> <li><a href="/wiki/Lacustrine_vole" title="Lacustrine vole">Lacustrine vole (<i>M. limnophilus</i>)</a></li> <li><a href="/wiki/Maximowicz%27s_vole" title="Maximowicz's vole">Maximowicz's vole (<i>M. maximowiczii</i>)</a></li> <li><a href="/wiki/Middendorff%27s_vole" title="Middendorff's vole">Middendorff's vole (<i>M. middendorffi</i>)</a></li> <li><a href="/wiki/Mongolian_vole" title="Mongolian vole">Mongolian vole (<i>M. mongolicus</i>)</a></li> <li><a href="/wiki/Japanese_grass_vole" title="Japanese grass vole">Japanese grass vole (<i>M. montebelli</i>)</a></li> <li><a href="/wiki/Muisk_vole" class="mw-redirect" title="Muisk vole">Muisk vole (<i>M. mujanensis</i>)</a></li> <li><a href="/wiki/Tundra_vole" title="Tundra vole">Tundra vole (<i>M. oeconomus</i>)</a></li> <li><a href="/wiki/Sakhalin_vole" title="Sakhalin vole">Sakhalin vole (<i>M. sachalinensis</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Chionomys" title="Chionomys">Chionomys</a></i><br /><small><span style="color:#696969"><span class="nobold">(Snow voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Caucasian_snow_vole" title="Caucasian snow vole">Caucasian snow vole (<i>C. gud</i>)</a></li> <li><a href="/wiki/European_snow_vole" title="European snow vole">European snow vole (<i>C. nivalis</i>)</a></li> <li><a href="/wiki/Robert%27s_snow_vole" title="Robert's snow vole">Robert's snow vole (<i>C. roberti</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Hyperacrius" title="Hyperacrius">Hyperacrius</a></i><br /><small><span style="color:#696969"><span class="nobold">(Voles from Pakistan)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/True%27s_vole" title="True's vole">True's vole (<i>H. fertilis</i>)</a></li> <li><a href="/wiki/Murree_vole" title="Murree vole">Murree vole (<i>H. wynnei</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Lasiopodomys" title="Lasiopodomys">Lasiopodomys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Brandt%27s_vole" title="Brandt's vole">Brandt's vole (<i>L. brandtii</i>)</a></li> <li><a href="/wiki/Plateau_vole" title="Plateau vole">Plateau vole (<i>L. fuscus</i>)</a></li> <li><a href="/wiki/Mandarin_vole" title="Mandarin vole">Mandarin vole (<i>L. mandarinus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Lemmiscus" class="mw-redirect" title="Lemmiscus">Lemmiscus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Sagebrush_vole" title="Sagebrush vole">Sagebrush vole (<i>L. curtatus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Microtus" title="Microtus">Microtus</a></i><br /><small><span style="color:#696969"><span class="nobold">(Voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><u>Subgenus <i><a href="/w/index.php?title=Microtus_(subgenus)&action=edit&redlink=1" class="new" title="Microtus (subgenus) (page does not exist)">Microtus</a></i></u>: <a href="/wiki/Field_vole" class="mw-redirect" title="Field vole">Field vole (<i>M. agrestis</i>)</a></li> <li><a href="/wiki/Anatolian_vole" title="Anatolian vole">Anatolian vole (<i>M. anatolicus</i>)</a></li> <li><a href="/wiki/Common_vole" title="Common vole">Common vole (<i>M. arvalis</i>)</a></li> <li><a href="/wiki/Cabrera%27s_vole" title="Cabrera's vole">Cabrera's vole (<i>M. cabrerae</i>)</a></li> <li><a href="/wiki/Do%C4%9Framaci%27s_vole" title="Doğramaci's vole">Doğramaci's vole (<i>M. dogramacii</i>)</a></li> <li><a href="/wiki/G%C3%BCnther%27s_vole" title="Günther's vole">Günther's vole (<i>M. guentheri</i>)</a></li> <li><a href="/wiki/Tien_Shan_vole" title="Tien Shan vole">Tien Shan vole (<i>M. ilaeus</i>)</a></li> <li><a href="/wiki/Persian_vole" title="Persian vole">Persian vole (<i>M. irani</i>)</a></li> <li><a href="/wiki/Kerman_vole" title="Kerman vole">Kerman vole (<i>M. kermanensis</i>)</a></li> <li><a href="/wiki/Southern_vole" class="mw-redirect" title="Southern vole">Southern vole (<i>M. levis</i>)</a></li> <li><a href="/wiki/Paradox_vole" title="Paradox vole">Paradox vole (<i>M. paradoxus</i>)</a></li> <li><a href="/wiki/Qazvin_vole" title="Qazvin vole">Qazvin vole (<i>M. qazvinensis</i>)</a></li> <li><a href="/wiki/Schidlovsky%27s_vole" title="Schidlovsky's vole">Schidlovsky's vole (<i>M. schidlovskii</i>)</a></li> <li><a href="/wiki/Social_vole" title="Social vole">Social vole (<i>M. socialis</i>)</a></li> <li><a href="/wiki/European_pine_vole" title="European pine vole">European pine vole (<i>M. subterraneus</i>)</a></li> <li><a href="/wiki/Transcaspian_vole" title="Transcaspian vole">Transcaspian vole (<i>M. transcaspicus</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/wiki/Blanfordimys" title="Blanfordimys">Blanfordimys</a></i></u>: <a href="/wiki/Afghan_vole" title="Afghan vole">Afghan vole (<i>B. afghanus</i>)</a></li> <li><a href="/wiki/Bucharian_vole" title="Bucharian vole">Bucharian vole (<i>B. bucharicus</i>)</a></li> <li><a href="/wiki/Juniper_vole" title="Juniper vole">Juniper vole (<i>B. juldaschi</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/wiki/Terricola_(subgenus)" title="Terricola (subgenus)">Terricola</a></i></u>: <a href="/wiki/Bavarian_pine_vole" title="Bavarian pine vole">Bavarian pine vole (<i>M. bavaricus</i>)</a></li> <li><a href="/wiki/Calabria_pine_vole" title="Calabria pine vole">Calabria pine vole (<i>M. brachycercus</i>)</a></li> <li><a href="/wiki/Daghestan_pine_vole" title="Daghestan pine vole">Daghestan pine vole (<i>M. daghestanicus</i>)</a></li> <li><a href="/wiki/Mediterranean_pine_vole" title="Mediterranean pine vole">Mediterranean pine vole (<i>M. duodecimcostatus</i>)</a></li> <li><a href="/wiki/Felten%27s_vole" title="Felten's vole">Felten's vole (<i>M. felteni</i>)</a></li> <li><a href="/wiki/Liechtenstein%27s_pine_vole" title="Liechtenstein's pine vole">Liechtenstein's pine vole (<i>M. liechtensteini</i>)</a></li> <li><a href="/wiki/Lusitanian_pine_vole" title="Lusitanian pine vole">Lusitanian pine vole (<i>M. lusitanicus</i>)</a></li> <li><a href="/wiki/Major%27s_pine_vole" title="Major's pine vole">Major's pine vole (<i>M. majori</i>)</a></li> <li><a href="/wiki/Alpine_pine_vole" title="Alpine pine vole">Alpine pine vole (<i>M. multiplex</i>)</a></li> <li><a href="/wiki/Savi%27s_pine_vole" title="Savi's pine vole">Savi's pine vole (<i>M. savii</i>)</a></li> <li><a href="/wiki/Tatra_pine_vole" title="Tatra pine vole">Tatra pine vole (<i>M. tatricus</i>)</a></li> <li><a href="/wiki/Thomas%27s_pine_vole" title="Thomas's pine vole">Thomas's pine vole (<i>M. thomasi</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/wiki/Mynomes" title="Mynomes">Mynomes</a></i></u>: <a class="mw-selflink selflink">Beach Vole (<i>M. breweri</i>)</a></li> <li><a href="/wiki/Gray-tailed_vole" title="Gray-tailed vole">Gray-tailed vole (<i>M. canicaudus</i>)</a></li> <li><a href="/wiki/Montane_vole" title="Montane vole">Montane vole (<i>M. montanus</i>)</a></li> <li><a href="/wiki/Creeping_vole" title="Creeping vole">Creeping vole (<i>M. oregoni</i>)</a></li> <li><a href="/wiki/Meadow_vole" class="mw-redirect" title="Meadow vole">Meadow vole (<i>M. pennsylvanicus</i>)</a></li> <li><a href="/wiki/Townsend%27s_vole" title="Townsend's vole">Townsend's vole (<i>M. townsendii</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/wiki/Pitymys" title="Pitymys">Pitymys</a></i></u>: <a href="/wiki/Guatemalan_vole" title="Guatemalan vole">Guatemalan vole (<i>M. guatemalensis</i>)</a></li> <li><a href="/wiki/Tarabund%C3%AD_vole" title="Tarabundí vole">Tarabundí vole (<i>M. oaxacensis</i>)</a></li> <li><a href="/wiki/Woodland_vole" title="Woodland vole">Woodland Vole (<i>M. pinetorum</i>)</a></li> <li><a href="/wiki/Jalapan_pine_vole" title="Jalapan pine vole">Jalapan pine vole (<i>M. quasiater</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/wiki/Pedomys" class="mw-redirect" title="Pedomys">Pedomys</a></i></u>: <a href="/wiki/Prairie_vole" title="Prairie vole">Prairie vole (<i>M. ochrogaster</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/wiki/Hyrcanicola" class="mw-redirect" title="Hyrcanicola">Hyrcanicola</a></i></u>: <a href="/wiki/Schelkovnikov%27s_pine_vole" title="Schelkovnikov's pine vole">Schelkovnikov's pine vole (<i>M. schelkovnikovi</i>)</a></li></ul> <ul><li><u><i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i></u>: <a href="/wiki/Insular_vole" title="Insular vole">Insular vole (<i>M. abbreviatus</i>)</a></li> <li><a href="/wiki/California_vole" title="California vole">California vole (<i>M. californicus</i>)</a></li> <li><a href="/wiki/Rock_vole" title="Rock vole">Rock vole (<i>M. chrotorrhinus</i>)</a></li> <li><a href="/wiki/Long-tailed_vole" title="Long-tailed vole">Long-tailed vole (<i>M. longicaudus</i>)</a></li> <li><a href="/wiki/Mexican_vole" title="Mexican vole">Mexican vole (<i>M. mexicanus</i>)</a></li> <li><a href="/wiki/Singing_vole" title="Singing vole">Singing vole (<i>M. miurus</i>)</a></li> <li><a href="/wiki/Water_vole_(North_America)" class="mw-redirect" title="Water vole (North America)">Water vole (<i>M. richardsoni</i>)</a></li> <li><a href="/wiki/Zempoalt%C3%A9pec_vole" title="Zempoaltépec vole">Zempoaltépec vole (<i>M. umbrosus</i>)</a></li> <li><a href="/wiki/Taiga_vole" title="Taiga vole">Taiga vole (<i>M. xanthognathus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Neodon" title="Neodon">Neodon</a></i><br /><small><span style="color:#696969"><span class="nobold">(Mountain voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Forrest%27s_mountain_vole" title="Forrest's mountain vole">Forrest's mountain vole (<i>N. forresti</i>)</a></li> <li><a href="/wiki/Chinese_scrub_vole" title="Chinese scrub vole">Chinese scrub vole (<i>N. irene</i>)</a></li> <li><a href="/wiki/Linzhi_mountain_vole" title="Linzhi mountain vole">Linzhi mountain vole (<i>N. linzhiensis</i>)</a></li> <li><a href="/wiki/Sikkim_mountain_vole" title="Sikkim mountain vole">Sikkim mountain vole (<i>N. sikimensis</i>)</a></li> <li><a href="/wiki/Blyth%27s_vole" title="Blyth's vole">Blyth's vole (<i>N. leucurus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Proedromys" class="mw-redirect" title="Proedromys">Proedromys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Duke_of_Bedford%27s_vole" title="Duke of Bedford's vole">Duke of Bedford's vole (<i>P. bedfordi</i>)</a></li> <li><i><a href="/wiki/Proedromys_liangshanensis" class="mw-redirect" title="Proedromys liangshanensis">P. liangshanensis</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Stenocranius" title="Stenocranius">Stenocranius</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Narrow-headed_vole" title="Narrow-headed vole">Narrow-headed vole (<i>S. gregalis</i>)</a></li> <li><a href="/wiki/Radde%27s_vole" title="Radde's vole">Radde's vole (<i>S. raddei</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Volemys" title="Volemys">Volemys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Szechuan_vole" title="Szechuan vole">Szechuan vole (<i>V. millicens</i>)</a></li> <li><a href="/wiki/Marie%27s_vole" title="Marie's vole">Marie's vole (<i>V. musseri</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Dicrostonychini" title="Dicrostonychini">Dicrostonychini</a><br /><small><span style="color:#696969"><span class="nobold">(Collared <a href="/wiki/Lemming" title="Lemming">lemmings</a>)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dicrostonyx" scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Collared_lemming" title="Collared lemming">Dicrostonyx</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Northern_collared_lemming" title="Northern collared lemming">Northern collared lemming (<i>D. groenlandicus</i>)</a></li> <li><a href="/wiki/Ungava_collared_lemming" title="Ungava collared lemming">Ungava collared lemming (<i>D. hudsonius</i>)</a></li> <li><a href="/wiki/Nelson%27s_collared_lemming" title="Nelson's collared lemming">Nelson's collared lemming (<i>D. nelsoni</i>)</a></li> <li><a href="/wiki/Ogilvie_Mountains_collared_lemming" title="Ogilvie Mountains collared lemming">Ogilvie Mountains collared lemming (<i>D. nunatakensis</i>)</a></li> <li><a href="/wiki/Richardson%27s_collared_lemming" title="Richardson's collared lemming">Richardson's collared lemming (<i>D. richardsoni</i>)</a></li> <li><a href="/wiki/Arctic_lemming" title="Arctic lemming">Arctic lemming (<i>D. torquatus</i>)</a></li> <li><a href="/wiki/Unalaska_collared_lemming" title="Unalaska collared lemming">Unalaska collared lemming (<i>D. unalascensis</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ellobiusini" title="Ellobiusini">Ellobiusini</a><br /><small><span style="color:#696969"><span class="nobold">(mole voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ellobius(Mole_voles)" scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Ellobius" title="Ellobius">Ellobius</a></i><br /><small><span style="color:#696969"><span class="nobold">(Mole voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Alai_mole_vole" title="Alai mole vole">Alai mole vole (<i>E. alaicus</i>)</a></li> <li><a href="/wiki/Southern_mole_vole" title="Southern mole vole">Southern mole vole (<i>E. fuscocapillus</i>)</a></li> <li><a href="/wiki/Transcaucasian_mole_vole" title="Transcaucasian mole vole">Transcaucasian mole vole (<i>E. lutescens</i>)</a></li> <li><a href="/wiki/Northern_mole_vole" title="Northern mole vole">Northern mole vole (<i>E. talpinus</i>)</a></li> <li><a href="/wiki/Zaisan_mole_vole" title="Zaisan mole vole">Zaisan mole vole (<i>E. tancrei</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lagurini" title="Lagurini">Lagurini</a><br /><small><span style="color:#696969"><span class="nobold">(Steppe <a href="/wiki/Lemming" title="Lemming">lemmings</a>)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Eolagurus" title="Eolagurus">Eolagurus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Yellow_steppe_lemming" title="Yellow steppe lemming">Yellow steppe lemming (<i>E. luteus</i>)</a></li> <li><a href="/wiki/Przewalski%27s_steppe_lemming" title="Przewalski's steppe lemming">Przewalski's steppe lemming (<i>E. przewalskii</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Lagurus_(rodent)" title="Lagurus (rodent)">Lagurus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Steppe_lemming" title="Steppe lemming">Steppe lemming (<i>L. lagurus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lemmini" title="Lemmini">Lemmini</a><br /><small><span style="color:#696969"><span class="nobold">(<a href="/wiki/Lemming" title="Lemming">Lemmings</a>)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/True_lemming" title="True lemming">Lemmus</a></i><br /><small><span style="color:#696969"><span class="nobold">(True lemmings)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amur_lemming" title="Amur lemming">Amur lemming (<i>L. amurensis</i>)</a></li> <li><a href="/wiki/Norway_lemming" title="Norway lemming">Norway lemming (<i>L. lemmus</i>)</a></li> <li><a href="/wiki/Beringian_lemming" title="Beringian lemming">Beringian lemming (<i>L. nigripes</i>)</a></li> <li><a href="/wiki/Wrangel_Island_lemming" class="mw-redirect" title="Wrangel Island lemming">Wrangel Island lemming (<i>L. paulus</i>)</a></li> <li><a href="/wiki/Siberian_brown_lemming" class="mw-redirect" title="Siberian brown lemming">Siberian brown lemming (<i>L. sibiricus</i>)</a></li> <li><a href="/wiki/North_American_brown_lemming" class="mw-redirect" title="North American brown lemming">North American brown lemming (<i>L. trimucronatus</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Myopus" class="mw-redirect" title="Myopus">Myopus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Wood_lemming" title="Wood lemming">Wood lemming (<i>M. schisticolor</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Bog_lemming" title="Bog lemming">Synaptomys</a></i><br /><small><span style="color:#696969"><span class="nobold">(Bog lemmings)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Northern_bog_lemming" title="Northern bog lemming">Northern bog lemming (<i>S. borealis</i>)</a></li> <li><a href="/wiki/Southern_bog_lemming" title="Southern bog lemming">Southern bog lemming (<i>S. cooperi</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Clethrionomyini" title="Clethrionomyini">Clethrionomyini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Alticola" title="Alticola">Alticola</a></i><br /><small><span style="color:#696969"><span class="nobold">(Voles from<br />Central Asia)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><u>Subgenus <i><a href="/w/index.php?title=Alticola_(subgenus)&action=edit&redlink=1" class="new" title="Alticola (subgenus) (page does not exist)">Alticola</a></i></u>: <a href="/wiki/White-tailed_mountain_vole" title="White-tailed mountain vole">White-tailed mountain vole (<i>A. albicauda</i>)</a></li> <li><a href="/wiki/Silver_mountain_vole" title="Silver mountain vole">Silver mountain vole (<i>A. argentatus</i>)</a></li> <li><a href="/wiki/Gobi_Altai_mountain_vole" title="Gobi Altai mountain vole">Gobi Altai mountain vole (<i>A. barakshin</i>)</a></li> <li><a href="/wiki/Central_Kashmir_vole" title="Central Kashmir vole">Central Kashmir vole (<i>A. montosa</i>)</a></li> <li><a href="/wiki/Royle%27s_mountain_vole" title="Royle's mountain vole">Royle's mountain vole (<i>A. roylei</i>)</a></li> <li><a href="/wiki/Mongolian_silver_vole" title="Mongolian silver vole">Mongolian silver vole (<i>A. semicanus</i>)</a></li> <li><a href="/wiki/Stoli%C4%8Dka%27s_mountain_vole" title="Stolička's mountain vole">Stolička's mountain vole (<i>A. stoliczkanus</i>)</a></li> <li><a href="/wiki/Tuva_silver_vole" title="Tuva silver vole">Tuva silver vole (<i>A. tuvinicus</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/w/index.php?title=Aschizomys&action=edit&redlink=1" class="new" title="Aschizomys (page does not exist)">Aschizomys</a></i></u>: <a href="/wiki/Lemming_vole" title="Lemming vole">Lemming vole (<i>A. lemminus</i>)</a></li> <li><a href="/wiki/Large-eared_vole" title="Large-eared vole">Large-eared vole (<i>A. macrotis</i>)</a></li> <li><a href="/wiki/Lake_Baikal_mountain_vole" title="Lake Baikal mountain vole">Lake Baikal mountain vole (<i>A. olchonensis</i>)</a></li></ul> <ul><li><u>Subgenus <i><a href="/w/index.php?title=Platycranius&action=edit&redlink=1" class="new" title="Platycranius (page does not exist)">Platycranius</a></i></u>: <a href="/wiki/Flat-headed_vole" title="Flat-headed vole">Flat-headed vole (<i>A. strelzowi</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Caryomys" title="Caryomys">Caryomys</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ganzu_vole" title="Ganzu vole">Ganzu vole (<i>C. eva</i>)</a></li> <li><a href="/wiki/Kolan_vole" title="Kolan vole">Kolan vole (<i>C. inez</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Eothenomys" title="Eothenomys">Eothenomys</a></i><br /><small><span style="color:#696969"><span class="nobold">(Voles from<br />East Asia)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Kachin_red-backed_vole" title="Kachin red-backed vole">Kachin red-backed vole (<i>E. cachinus</i>)</a></li> <li><a href="/wiki/Pratt%27s_vole" title="Pratt's vole">Pratt's vole (<i>E. chinensis</i>)</a></li> <li><a href="/wiki/Southwest_China_vole" title="Southwest China vole">Southwest China vole (<i>E. custos</i>)</a></li> <li><a href="/wiki/P%C3%A8re_David%27s_vole" title="Père David's vole">Père David's vole (<i>E. melanogaster</i>)</a></li> <li><a href="/wiki/Yunnan_red-backed_vole" title="Yunnan red-backed vole">Yunnan red-backed vole (<i>E. miletus</i>)</a></li> <li><a href="/wiki/Chaotung_vole" title="Chaotung vole">Chaotung vole (<i>E. olitor</i>)</a></li> <li><a href="/wiki/Yulungshan_vole" title="Yulungshan vole">Yulungshan vole (<i>E. proditor</i>)</a></li> <li><a href="/wiki/Ward%27s_red-backed_vole" title="Ward's red-backed vole">Ward's red-backed vole (<i>E. wardi</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Clethrionomys" title="Clethrionomys">Clethrionomys</a></i><br /><small><span style="color:#696969"><span class="nobold">(some Red-backed<br />voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Western_red-backed_vole" title="Western red-backed vole">Western red-backed vole (<i>C. californicus</i>)</a></li> <li><a href="/wiki/Tien_Shan_red-backed_vole" title="Tien Shan red-backed vole">Tien Shan red-backed vole (<i>C. centralis</i>)</a></li> <li><a href="/wiki/Southern_red-backed_vole" title="Southern red-backed vole">Southern red-backed Vole (<i>C. gapperi</i>)</a></li> <li><a href="/wiki/Bank_vole" title="Bank vole">Bank vole (<i>C. glareolus</i>)</a></li> <li><a href="/wiki/Imaizumi%27s_red-backed_vole" title="Imaizumi's red-backed vole">Imaizumi's red-backed vole (<i>C. imaizumii</i>)</a></li> <li><a href="/wiki/Northern_red-backed_vole" title="Northern red-backed vole">Northern red-backed vole (<i>C. rutilus</i>)</a></li> <li><a href="/wiki/Shansei_vole" title="Shansei vole">Shansei vole (<i>C. shanseius</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Craseomys" title="Craseomys">Craseomys</a></i><br /><small><span style="color:#696969"><span class="nobold">(other Red-backed<br />voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Japanese_red-backed_vole" title="Japanese red-backed vole">Japanese red-backed vole (<i>C. andersoni</i>)</a></li> <li><a href="/wiki/Royal_vole" title="Royal vole">Royal vole (<i>C. regulus</i>)</a></li> <li><a href="/wiki/Hokkaido_red-backed_vole" title="Hokkaido red-backed vole">Hokkaido red-backed vole (<i>C. rex</i>)</a></li> <li><a href="/wiki/Grey_red-backed_vole" title="Grey red-backed vole">Grey red-backed vole (<i>C. rufocanus</i>)</a></li> <li><a href="/wiki/Smith%27s_vole" title="Smith's vole">Smith's vole (<i>C. smithii</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ondatrini" title="Ondatrini">Ondatrini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Neofiber" class="mw-redirect" title="Neofiber">Neofiber</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Round-tailed_muskrat" title="Round-tailed muskrat">Round-tailed muskrat (<i>N. alleni</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Muskrat" title="Muskrat">Ondatra</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Muskrat" title="Muskrat">Muskrat (<i>O. zibethicus</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pliomyini" class="mw-redirect" title="Pliomyini">Pliomyini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dinaromys" scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Dinaromys" class="mw-redirect" title="Dinaromys">Dinaromys</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Balkan_snow_vole" title="Balkan snow vole">Balkan snow vole (<i>D. bogdanovi</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pliophenacomyini" class="mw-redirect" title="Pliophenacomyini">Pliophenacomyini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Tree_vole" title="Tree vole">Arborimus</a></i><br /><small><span style="color:#696969"><span class="nobold">(Tree voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/White-footed_vole" title="White-footed vole">White-footed vole (<i>A. albipes</i>)</a></li> <li><a href="/wiki/Red_tree_vole" title="Red tree vole">Red tree vole (<i>A. longicaudus</i>)</a></li> <li><a href="/wiki/California_red_tree_mouse" class="mw-redirect" title="California red tree mouse">California red tree mouse (<i>A. pomo</i>)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Heather_vole" title="Heather vole">Phenacomys</a></i><br /><small><span style="color:#696969"><span class="nobold">(Heather voles)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Western_heather_vole" title="Western heather vole">Western heather vole (<i>P. intermedius</i>)</a></li> <li><a href="/wiki/Eastern_heather_vole" title="Eastern heather vole">Eastern heather vole (<i>P. ungava</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Prometheomyini" class="mw-redirect" title="Prometheomyini">Prometheomyini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Prometheomys" scope="row" class="navbox-group" style="width:9em"><i><a href="/wiki/Prometheomys" class="mw-redirect" title="Prometheomys">Prometheomys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Long-clawed_mole_vole" title="Long-clawed mole vole">Long-clawed mole vole (<i>P. schaposchnikowi</i>)</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Microtus breweri</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q1765454" class="extiw" title="wikidata:Q1765454">Q1765454</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Microtus_breweri" class="extiw" title="wikispecies:Microtus breweri">Microtus breweri</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/310283">310283</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/2438615">2438615</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/44325">44325</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=11270423">11270423</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=180304">180304</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://apiv3.iucnredlist.org/api/v3/taxonredirect/13417">13417</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Mammal_Species_of_the_World" title="Mammal Species of the World">MSW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?s=y&id=13000251">13000251</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/NatureServe" title="NatureServe">NatureServe</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://explorer.natureserve.org/Taxon/ELEMENT_GLOBAL.2.102648/">2.102648</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=663697">663697</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Observation.org" title="Observation.org">Observation.org</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://observation.org/species/84958/">84958</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=988006">988006</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a>: National <span 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