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id="toc-Structure_and_function-sublist" class="vector-toc-list"> <li id="toc-Structure_of_eukaryotic_cells" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Structure_of_eukaryotic_cells"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.1</span> <span>Structure of eukaryotic cells</span> </div> </a> <ul id="toc-Structure_of_eukaryotic_cells-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cell_metabolism" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cell_metabolism"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.2</span> <span>Cell metabolism</span> </div> </a> <ul id="toc-Cell_metabolism-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cell_signaling" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cell_signaling"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.3</span> <span>Cell signaling</span> </div> </a> <ul id="toc-Cell_signaling-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Growth_and_development" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Growth_and_development"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Growth and development</span> </div> </a> <button aria-controls="toc-Growth_and_development-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Growth and development subsection</span> </button> <ul id="toc-Growth_and_development-sublist" class="vector-toc-list"> <li id="toc-Eukaryotic_cell_cycle" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Eukaryotic_cell_cycle"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.1</span> <span>Eukaryotic cell cycle</span> </div> </a> <ul id="toc-Eukaryotic_cell_cycle-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cell_mortality,_cell_lineage_immortality" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cell_mortality,_cell_lineage_immortality"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2</span> <span>Cell mortality, cell lineage immortality</span> </div> </a> <ul id="toc-Cell_mortality,_cell_lineage_immortality-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cell_cycle_phases" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cell_cycle_phases"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.3</span> <span>Cell cycle phases</span> </div> </a> <ul id="toc-Cell_cycle_phases-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Pathology" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Pathology"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Pathology</span> </div> </a> <ul id="toc-Pathology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cell_cycle_checkpoints_and_DNA_damage_repair_system" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Cell_cycle_checkpoints_and_DNA_damage_repair_system"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Cell cycle checkpoints and DNA damage repair system</span> </div> </a> <ul id="toc-Cell_cycle_checkpoints_and_DNA_damage_repair_system-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Mitochondrial_membrane_dynamics" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Mitochondrial_membrane_dynamics"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>Mitochondrial membrane dynamics</span> </div> </a> <ul id="toc-Mitochondrial_membrane_dynamics-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Autophagy" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Autophagy"> <div class="vector-toc-text"> <span class="vector-toc-numb">9</span> <span>Autophagy</span> </div> </a> <ul id="toc-Autophagy-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notable_cell_biologists" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notable_cell_biologists"> <div class="vector-toc-text"> <span class="vector-toc-numb">10</span> <span>Notable cell biologists</span> </div> </a> <ul id="toc-Notable_cell_biologists-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">11</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notes" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notes"> <div class="vector-toc-text"> <span class="vector-toc-numb">12</span> <span>Notes</span> </div> </a> <ul id="toc-Notes-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">13</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">14</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span 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Available in 83 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-83" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">83 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B9%D9%84%D9%85_%D8%A7%D9%84%D8%AE%D9%84%D9%8A%D8%A9" title="علم الخلية – Arabic" lang="ar" hreflang="ar" data-title="علم الخلية" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Biolox%C3%ADa_celular" title="Bioloxía celular – Asturian" lang="ast" hreflang="ast" data-title="Bioloxía celular" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Sitologiya" title="Sitologiya – Azerbaijani" lang="az" hreflang="az" data-title="Sitologiya" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%95%E0%A7%8B%E0%A6%B7%E0%A6%AC%E0%A6%BF%E0%A6%A6%E0%A7%8D%E0%A6%AF%E0%A6%BE" title="কোষবিদ্যা – Bangla" lang="bn" hreflang="bn" data-title="কোষবিদ্যা" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/S%C3%A8-pau_seng-bu%CC%8Dt-ha%CC%8Dk" title="Sè-pau seng-bu̍t-ha̍k – Minnan" lang="nan" hreflang="nan" data-title="Sè-pau seng-bu̍t-ha̍k" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9A%D0%BB%D0%B5%D1%82%D1%8A%D1%87%D0%BD%D0%B0_%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Клетъчна биология – Bulgarian" lang="bg" hreflang="bg" data-title="Клетъчна биология" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Citologija" title="Citologija – Bosnian" lang="bs" hreflang="bs" data-title="Citologija" data-language-autonym="Bosanski" data-language-local-name="Bosnian" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Biologia_cel%C2%B7lular" title="Biologia cel·lular – Catalan" lang="ca" hreflang="ca" data-title="Biologia cel·lular" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Bun%C4%9B%C4%8Dn%C3%A1_biologie" title="Buněčná biologie – Czech" lang="cs" hreflang="cs" data-title="Buněčná biologie" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/Bioleg_cell" title="Bioleg cell – Welsh" lang="cy" hreflang="cy" data-title="Bioleg cell" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Zellbiologie" title="Zellbiologie – German" lang="de" hreflang="de" data-title="Zellbiologie" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Rakubioloogia" title="Rakubioloogia – Estonian" lang="et" hreflang="et" data-title="Rakubioloogia" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%9A%CF%85%CF%84%CF%84%CE%B1%CF%81%CE%B9%CE%BA%CE%AE_%CE%B2%CE%B9%CE%BF%CE%BB%CE%BF%CE%B3%CE%AF%CE%B1" title="Κυτταρική βιολογία – Greek" lang="el" hreflang="el" data-title="Κυτταρική βιολογία" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Biolog%C3%ADa_celular" title="Biología celular – Spanish" lang="es" hreflang="es" data-title="Biología celular" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/%C4%88elbiologio" title="Ĉelbiologio – Esperanto" lang="eo" hreflang="eo" data-title="Ĉelbiologio" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Biologia_zelular" title="Biologia zelular – Basque" lang="eu" hreflang="eu" data-title="Biologia zelular" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%D9%84%D9%88%D9%84%E2%80%8C%D8%B4%D9%86%D8%A7%D8%B3%DB%8C" title="سلول‌شناسی – Persian" lang="fa" hreflang="fa" data-title="سلول‌شناسی" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Biologie_cellulaire" title="Biologie cellulaire – French" lang="fr" hreflang="fr" data-title="Biologie cellulaire" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-fy mw-list-item"><a href="https://fy.wikipedia.org/wiki/Selbiology" title="Selbiology – Western Frisian" lang="fy" hreflang="fy" data-title="Selbiology" data-language-autonym="Frysk" data-language-local-name="Western Frisian" class="interlanguage-link-target"><span>Frysk</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Bitheola%C3%ADocht_na_gceall" title="Bitheolaíocht na gceall – Irish" lang="ga" hreflang="ga" data-title="Bitheolaíocht na gceall" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Biolox%C3%ADa_celular" title="Bioloxía celular – Galician" lang="gl" hreflang="gl" data-title="Bioloxía celular" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-hak mw-list-item"><a href="https://hak.wikipedia.org/wiki/Se-p%C3%A2u_s%C3%A2ng-vu%CC%8Dt-ho%CC%8Dk" title="Se-pâu sâng-vu̍t-ho̍k – Hakka Chinese" lang="hak" hreflang="hak" data-title="Se-pâu sâng-vu̍t-ho̍k" data-language-autonym="客家語 / Hak-kâ-ngî" data-language-local-name="Hakka Chinese" class="interlanguage-link-target"><span>客家語 / Hak-kâ-ngî</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%84%B8%ED%8F%AC%EC%83%9D%EB%AC%BC%ED%95%99" title="세포생물학 – Korean" lang="ko" hreflang="ko" data-title="세포생물학" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%95%E0%A5%8B%E0%A4%B6%E0%A4%BF%E0%A4%95%E0%A4%BE_%E0%A4%B5%E0%A4%BF%E0%A4%9C%E0%A5%8D%E0%A4%9E%E0%A4%BE%E0%A4%A8" title="कोशिका विज्ञान – Hindi" lang="hi" hreflang="hi" data-title="कोशिका विज्ञान" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Stani%C4%8Dna_biologija" title="Stanična biologija – Croatian" lang="hr" hreflang="hr" data-title="Stanična biologija" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Celulala_biologio" title="Celulala biologio – Ido" lang="io" hreflang="io" data-title="Celulala biologio" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Biologi_sel" title="Biologi sel – Indonesian" lang="id" hreflang="id" data-title="Biologi sel" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-ia mw-list-item"><a href="https://ia.wikipedia.org/wiki/Biologia_cellular" title="Biologia cellular – Interlingua" lang="ia" hreflang="ia" data-title="Biologia cellular" data-language-autonym="Interlingua" data-language-local-name="Interlingua" class="interlanguage-link-target"><span>Interlingua</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Frumul%C3%ADffr%C3%A6%C3%B0i" title="Frumulíffræði – Icelandic" lang="is" hreflang="is" data-title="Frumulíffræði" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Biologia_cellulare" title="Biologia cellulare – Italian" lang="it" hreflang="it" data-title="Biologia cellulare" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%91%D7%99%D7%95%D7%9C%D7%95%D7%92%D7%99%D7%94_%D7%A9%D7%9C_%D7%94%D7%AA%D7%90" title="ביולוגיה של התא – Hebrew" lang="he" hreflang="he" data-title="ביולוגיה של התא" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Sitologi" title="Sitologi – Javanese" lang="jv" hreflang="jv" data-title="Sitologi" data-language-autonym="Jawa" data-language-local-name="Javanese" class="interlanguage-link-target"><span>Jawa</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%AA%E1%83%98%E1%83%A2%E1%83%9D%E1%83%9A%E1%83%9D%E1%83%92%E1%83%98%E1%83%90" title="ციტოლოგია – Georgian" lang="ka" hreflang="ka" data-title="ციტოლოგია" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A6%D0%B8%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Цитология – Kazakh" lang="kk" hreflang="kk" data-title="Цитология" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Cytologia" title="Cytologia – Latin" lang="la" hreflang="la" data-title="Cytologia" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Citolo%C4%A3ija" title="Citoloģija – Latvian" lang="lv" hreflang="lv" data-title="Citoloģija" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lb mw-list-item"><a href="https://lb.wikipedia.org/wiki/Zytologie" title="Zytologie – Luxembourgish" lang="lb" hreflang="lb" data-title="Zytologie" data-language-autonym="Lëtzebuergesch" data-language-local-name="Luxembourgish" class="interlanguage-link-target"><span>Lëtzebuergesch</span></a></li><li class="interlanguage-link interwiki-lfn mw-list-item"><a href="https://lfn.wikipedia.org/wiki/Biolojia_selulal" title="Biolojia selulal – Lingua Franca Nova" lang="lfn" hreflang="lfn" data-title="Biolojia selulal" data-language-autonym="Lingua Franca Nova" data-language-local-name="Lingua Franca Nova" class="interlanguage-link-target"><span>Lingua Franca Nova</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Sejtbiol%C3%B3gia" title="Sejtbiológia – Hungarian" lang="hu" hreflang="hu" data-title="Sejtbiológia" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%A6%D0%B8%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%98%D0%B0" title="Цитологија – Macedonian" lang="mk" hreflang="mk" data-title="Цитологија" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%95%E0%B5%8B%E0%B4%B6%E0%B4%9C%E0%B5%80%E0%B4%B5%E0%B4%B6%E0%B4%BE%E0%B4%B8%E0%B5%8D%E0%B4%A4%E0%B5%8D%E0%B4%B0%E0%B4%82" title="കോശജീവശാസ്ത്രം – Malayalam" lang="ml" hreflang="ml" data-title="കോശജീവശാസ്ത്രം" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%AA%E1%83%98%E1%83%A2%E1%83%9D%E1%83%9A%E1%83%9D%E1%83%92%E1%83%98%E1%83%90" title="ციტოლოგია – Mingrelian" lang="xmf" hreflang="xmf" data-title="ციტოლოგია" data-language-autonym="მარგალური" data-language-local-name="Mingrelian" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Biologi_sel" title="Biologi sel – Malay" lang="ms" hreflang="ms" data-title="Biologi sel" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mn mw-list-item"><a href="https://mn.wikipedia.org/wiki/%D0%A6%D0%B8%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8" title="Цитологи – Mongolian" lang="mn" hreflang="mn" data-title="Цитологи" data-language-autonym="Монгол" data-language-local-name="Mongolian" class="interlanguage-link-target"><span>Монгол</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Celbiologie" title="Celbiologie – Dutch" lang="nl" hreflang="nl" data-title="Celbiologie" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-new mw-list-item"><a href="https://new.wikipedia.org/wiki/%E0%A4%95%E0%A5%8B%E0%A4%B7_%E0%A4%9C%E0%A5%80%E0%A4%B5%E0%A4%B6%E0%A4%BE%E0%A4%B8%E0%A5%8D%E0%A4%A4%E0%A5%8D%E0%A4%B0" title="कोष जीवशास्त्र – Newari" lang="new" hreflang="new" data-title="कोष जीवशास्त्र" data-language-autonym="नेपाल भाषा" data-language-local-name="Newari" class="interlanguage-link-target"><span>नेपाल भाषा</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E7%B4%B0%E8%83%9E%E7%94%9F%E7%89%A9%E5%AD%A6" title="細胞生物学 – Japanese" lang="ja" hreflang="ja" data-title="細胞生物学" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Cytologi" title="Cytologi – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Cytologi" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-nov mw-list-item"><a href="https://nov.wikipedia.org/wiki/Selulal_biologia" title="Selulal biologia – Novial" lang="nov" hreflang="nov" data-title="Selulal biologia" data-language-autonym="Novial" data-language-local-name="Novial" class="interlanguage-link-target"><span>Novial</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Biologia_cellulara" title="Biologia cellulara – Occitan" lang="oc" hreflang="oc" data-title="Biologia cellulara" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Sitologiya" title="Sitologiya – Uzbek" lang="uz" hreflang="uz" data-title="Sitologiya" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pa mw-list-item"><a href="https://pa.wikipedia.org/wiki/%E0%A8%95%E0%A9%8B%E0%A8%B8%E0%A8%BC%E0%A8%BE%E0%A8%A3%E0%A9%82_%E0%A8%B5%E0%A8%BF%E0%A8%97%E0%A8%BF%E0%A8%86%E0%A8%A8" title="ਕੋਸ਼ਾਣੂ ਵਿਗਿਆਨ – Punjabi" lang="pa" hreflang="pa" data-title="ਕੋਸ਼ਾਣੂ ਵਿਗਿਆਨ" data-language-autonym="ਪੰਜਾਬੀ" data-language-local-name="Punjabi" class="interlanguage-link-target"><span>ਪੰਜਾਬੀ</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AE%D9%84%DB%8C%D8%A7%D8%A6%DB%8C_%D8%AD%DB%8C%D8%A7%D8%AA%DB%8C%D8%A7%D8%AA" title="خلیائی حیاتیات – Western Punjabi" lang="pnb" hreflang="pnb" data-title="خلیائی حیاتیات" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-ps mw-list-item"><a href="https://ps.wikipedia.org/wiki/%D8%AF_%D8%AD%D8%AC%D8%B1%DB%90_%D8%A8%D9%8A%D9%88%D9%84%D9%88%DA%98%D9%8A" title="د حجرې بيولوژي – Pashto" lang="ps" hreflang="ps" data-title="د حجرې بيولوژي" data-language-autonym="پښتو" data-language-local-name="Pashto" class="interlanguage-link-target"><span>پښتو</span></a></li><li class="interlanguage-link interwiki-pl badge-Q70894304 mw-list-item" title=""><a href="https://pl.wikipedia.org/wiki/Biologia_kom%C3%B3rki" title="Biologia komórki – Polish" lang="pl" hreflang="pl" data-title="Biologia komórki" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Biologia_celular" title="Biologia celular – Portuguese" lang="pt" hreflang="pt" data-title="Biologia celular" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-kaa mw-list-item"><a href="https://kaa.wikipedia.org/wiki/Citologiya" title="Citologiya – Kara-Kalpak" lang="kaa" hreflang="kaa" data-title="Citologiya" data-language-autonym="Qaraqalpaqsha" data-language-local-name="Kara-Kalpak" class="interlanguage-link-target"><span>Qaraqalpaqsha</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Biologie_celular%C4%83" title="Biologie celulară – Romanian" lang="ro" hreflang="ro" data-title="Biologie celulară" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru badge-Q70894304 mw-list-item" title=""><a href="https://ru.wikipedia.org/wiki/%D0%9A%D0%BB%D0%B5%D1%82%D0%BE%D1%87%D0%BD%D0%B0%D1%8F_%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Клеточная биология – Russian" lang="ru" hreflang="ru" data-title="Клеточная биология" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sah mw-list-item"><a href="https://sah.wikipedia.org/wiki/%D0%A1%D0%B0%D0%B0%D0%BD%D1%8B%D0%BA_%D0%B1%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F%D1%82%D0%B0" title="Саанык биологията – Yakut" lang="sah" hreflang="sah" data-title="Саанык биологията" data-language-autonym="Саха тыла" data-language-local-name="Yakut" class="interlanguage-link-target"><span>Саха тыла</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Cell_biology" title="Cell biology – Scots" lang="sco" hreflang="sco" data-title="Cell biology" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Biologjia_qelizore" title="Biologjia qelizore – Albanian" lang="sq" hreflang="sq" data-title="Biologjia qelizore" data-language-autonym="Shqip" data-language-local-name="Albanian" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Cell_biology" title="Cell biology – Simple English" lang="en-simple" hreflang="en-simple" data-title="Cell biology" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Cytol%C3%B3gia" title="Cytológia – Slovak" lang="sk" hreflang="sk" data-title="Cytológia" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Celi%C4%8Dna_biologija" title="Celična biologija – Slovenian" lang="sl" hreflang="sl" data-title="Celična biologija" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%DA%98%DB%8C%D9%86%D9%86%D8%A7%D8%B3%DB%8C%DB%8C_%D8%AE%D8%A7%D9%86%DB%95%DB%8C%DB%8C" title="ژینناسیی خانەیی – Central Kurdish" lang="ckb" hreflang="ckb" data-title="ژینناسیی خانەیی" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-sr badge-Q70893996 mw-list-item" title=""><a href="https://sr.wikipedia.org/wiki/%D0%91%D0%B8%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%98%D0%B0_%D1%9B%D0%B5%D0%BB%D0%B8%D1%98%D0%B5" title="Биологија ћелије – Serbian" lang="sr" hreflang="sr" data-title="Биологија ћелије" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Celularna_biologija" title="Celularna biologija – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Celularna biologija" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Solubiologia" title="Solubiologia – Finnish" lang="fi" hreflang="fi" data-title="Solubiologia" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Cellbiologi" title="Cellbiologi – Swedish" lang="sv" hreflang="sv" data-title="Cellbiologi" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tl mw-list-item"><a href="https://tl.wikipedia.org/wiki/Biyolohiyang_pangselula" title="Biyolohiyang pangselula – Tagalog" lang="tl" hreflang="tl" data-title="Biyolohiyang pangselula" data-language-autonym="Tagalog" data-language-local-name="Tagalog" class="interlanguage-link-target"><span>Tagalog</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%89%E0%AE%AF%E0%AE%BF%E0%AE%B0%E0%AE%A3%E0%AF%81_%E0%AE%89%E0%AE%AF%E0%AE%BF%E0%AE%B0%E0%AE%BF%E0%AE%AF%E0%AE%B2%E0%AF%8D" title="உயிரணு உயிரியல் – Tamil" lang="ta" hreflang="ta" data-title="உயிரணு உயிரியல்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%9A%D2%AF%D0%B7%D3%99%D0%BD%D3%99%D0%BA_%D1%84%D3%99%D0%BD%D0%B5" title="Күзәнәк фәне – Tatar" lang="tt" hreflang="tt" data-title="Күзәнәк фәне" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B8%8A%E0%B8%B5%E0%B8%A7%E0%B8%A7%E0%B8%B4%E0%B8%97%E0%B8%A2%E0%B8%B2%E0%B8%82%E0%B8%AD%E0%B8%87%E0%B9%80%E0%B8%8B%E0%B8%A5%E0%B8%A5%E0%B9%8C" title="ชีววิทยาของเซลล์ – Thai" lang="th" hreflang="th" data-title="ชีววิทยาของเซลล์" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tg mw-list-item"><a href="https://tg.wikipedia.org/wiki/%D0%A1%D0%B8%D1%82%D0%BE%D0%BB%D0%BE%D0%B3%D0%B8%D1%8F" title="Ситология – Tajik" lang="tg" hreflang="tg" data-title="Ситология" data-language-autonym="Тоҷикӣ" data-language-local-name="Tajik" class="interlanguage-link-target"><span>Тоҷикӣ</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Sitoloji" title="Sitoloji – Turkish" lang="tr" hreflang="tr" data-title="Sitoloji" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9A%D0%BB%D1%96%D1%82%D0%B8%D0%BD%D0%BD%D0%B0_%D0%B1%D1%96%D0%BE%D0%BB%D0%BE%D0%B3%D1%96%D1%8F" title="Клітинна біологія – Ukrainian" lang="uk" hreflang="uk" data-title="Клітинна біологія" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%AE%D9%84%DB%8C%D8%A7%D8%A6%DB%8C_%D8%AD%DB%8C%D8%A7%D8%AA%DB%8C%D8%A7%D8%AA" title="خلیائی حیاتیات – Urdu" lang="ur" hreflang="ur" data-title="خلیائی حیاتیات" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Sinh_h%E1%BB%8Dc_t%E1%BA%BF_b%C3%A0o" title="Sinh học tế bào – Vietnamese" lang="vi" hreflang="vi" data-title="Sinh học tế bào" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Biyolohiya_hin_selula" title="Biyolohiya hin selula – Waray" lang="war" hreflang="war" data-title="Biyolohiya hin selula" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E7%BB%86%E8%83%9E%E7%94%9F%E7%89%A9%E5%AD%A6" title="细胞生物学 – Wu" lang="wuu" hreflang="wuu" data-title="细胞生物学" data-language-autonym="吴语" data-language-local-name="Wu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E7%B4%B0%E8%83%9E%E7%94%9F%E7%89%A9%E5%AD%B8" title="細胞生物學 – Cantonese" lang="yue" hreflang="yue" data-title="細胞生物學" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E7%BB%86%E8%83%9E%E7%94%9F%E7%89%A9%E5%AD%A6" title="细胞生物学 – Chinese" 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.mw-editsection-like{font-style:normal}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1066933788"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><table class="sidebar sidebar-collapse nomobile nowraplinks" style="border:2px solid #90db90"><tbody><tr><td class="sidebar-pretitle">Part of a series on</td></tr><tr><th class="sidebar-title-with-pretitle" style="padding:0.2em;font-size:175%;font-weight:bold"><a href="/wiki/Biology" title="Biology">Biology</a></th></tr><tr><td class="sidebar-image"><span typeof="mw:File"><a href="/wiki/File:DNA_simple.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/DNA_simple.svg/45px-DNA_simple.svg.png" decoding="async" width="45" height="107" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/DNA_simple.svg/68px-DNA_simple.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e7/DNA_simple.svg/90px-DNA_simple.svg.png 2x" data-file-width="114" data-file-height="270" /></a></span><div class="sidebar-caption"><b>Science of life</b></div></td></tr><tr><td class="sidebar-above" style="background:#ddddff;display:block; margin-bottom:0.55em; background-color: #e7f4e7;"> <div class="hlist"> <ul><li><a href="/wiki/Index_of_biology_articles" title="Index of biology articles">Index</a></li> <li><a href="/wiki/Outline_of_biology" title="Outline of biology">Outline</a></li> <li><a href="/wiki/Glossary_of_biology" title="Glossary of biology">Glossary</a></li></ul> <div class="hlist"> </div> <ul><li><a href="/wiki/History_of_biology" title="History of biology">History</a> (<a href="/wiki/Timeline_of_biology_and_organic_chemistry" title="Timeline of biology and organic chemistry">timeline</a>)</li></ul> </div></td></tr><tr><td class="sidebar-content" style="background:#e7f4e7;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90db90;font-size:100%;font-weight:bold;;color: var(--color-base)">Key components</div><div class="sidebar-list-content mw-collapsible-content hlist"> <ul><li><a href="/wiki/Cell_theory" title="Cell theory">Cell theory</a></li> <li><a href="/wiki/Ecosystem" title="Ecosystem">Ecosystem</a></li> <li><a href="/wiki/Evolution" title="Evolution">Evolution</a></li> <li><a href="/wiki/Phylogenetic_tree" title="Phylogenetic tree"> Phylogeny</a></li></ul> <hr /> <ul><li><b>Properties of <a href="/wiki/Life" title="Life">life</a></b></li></ul> <hr /> <ul><li><a href="/wiki/Adaptation" title="Adaptation">Adaptation</a></li> <li><a href="/wiki/Metabolism" title="Metabolism"> Energy processing</a></li> <li><a href="/wiki/Developmental_biology" title="Developmental biology">Growth</a></li> <li><a href="/wiki/Structure#Biological" title="Structure">Order</a></li> <li><a href="/wiki/Homeostasis" title="Homeostasis">Regulation</a></li> <li><a href="/wiki/Reproduction" title="Reproduction">Reproduction</a></li> <li><a href="/wiki/Stimulus_(physiology)" title="Stimulus (physiology)">Response to environment</a></li></ul> <hr /> <ul><li><b><a href="/wiki/Domain_(biology)" title="Domain (biology)"> Domains</a> and <a href="/wiki/Kingdom_(biology)" title="Kingdom (biology)">Kingdoms</a> of life</b></li></ul> <hr /> <ul><li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Bacteria" title="Bacteria">Bacteria</a></li> <li><a href="/wiki/Eukaryote" title="Eukaryote">Eukarya</a> (<a href="/wiki/Animals" class="mw-redirect" title="Animals">Animals</a>, <a href="/wiki/Fungi" class="mw-redirect" title="Fungi">Fungi</a>, <a href="/wiki/Plants" class="mw-redirect" title="Plants">Plants</a>, <a href="/wiki/Protists" class="mw-redirect" title="Protists">Protists</a>)</li></ul></div></div></td> </tr><tr><td class="sidebar-content" style="background:#e7f4e7;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90db90;font-size:100%;font-weight:bold;;color: var(--color-base)">Branches</div><div class="sidebar-list-content mw-collapsible-content hlist"><div class="excerpt-block"><div class="excerpt"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></li> <li><a href="/wiki/Aerobiology" title="Aerobiology">Aerobiology</a></li> <li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Agrostology" title="Agrostology">Agrostology</a></li> <li><a href="/wiki/Anatomy" title="Anatomy">Anatomy</a></li> <li><a href="/wiki/Astrobiology" title="Astrobiology">Astrobiology</a></li> <li><a href="/wiki/Bacteriology" title="Bacteriology">Bacteriology</a></li> <li><a href="/wiki/Biochemistry" title="Biochemistry">Biochemistry</a></li> <li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a 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title="Lipidology">Lipidology</a></li> <li><a href="/wiki/Mammalogy" title="Mammalogy">Mammalogy</a></li> <li><a href="/wiki/Marine_biology" title="Marine biology">Marine biology</a></li> <li><a href="/wiki/Mathematical_and_theoretical_biology" title="Mathematical and theoretical biology">Mathematical biology</a></li> <li><a href="/wiki/Microbiology" title="Microbiology">Microbiology</a></li> <li><a href="/wiki/Molecular_biology" title="Molecular biology">Molecular biology</a></li> <li><a href="/wiki/Mycology" title="Mycology">Mycology</a></li> <li><a href="/wiki/Neontology" title="Neontology">Neontology</a></li> <li><a href="/wiki/Neuroscience" title="Neuroscience">Neuroscience</a></li> <li><a href="/wiki/Nutritional_science" title="Nutritional science">Nutrition</a></li> <li><a href="/wiki/Ornithology" title="Ornithology">Ornithology</a></li> <li><a href="/wiki/Osteology" title="Osteology">Osteology</a></li> <li><a href="/wiki/Paleontology" title="Paleontology">Paleontology</a></li> <li><a href="/wiki/Parasitology" title="Parasitology">Parasitology</a></li> <li><a href="/wiki/Pathology" title="Pathology">Pathology</a></li> <li><a href="/wiki/Pharmacology" title="Pharmacology">Pharmacology</a></li> <li><a href="/wiki/Photobiology" title="Photobiology">Photobiology</a></li> <li><a href="/wiki/Phycology" title="Phycology">Phycology</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a></li> <li><a href="/wiki/Physiology" title="Physiology">Physiology</a></li> <li><a href="/wiki/Pomology" title="Pomology">Pomology</a></li> <li><a href="/wiki/Primatology" title="Primatology">Primatology</a></li> <li><a href="/wiki/Proteomics" title="Proteomics">Proteomics</a></li> <li><a href="/wiki/Protistology" title="Protistology">Protistology</a></li> <li><a href="/wiki/Quantum_biology" title="Quantum biology">Quantum biology</a></li> <li><a href="/wiki/Relational_biology" class="mw-redirect" title="Relational biology">Relational biology</a></li> <li><a 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title="Zoology">Zoology</a></li></ul></div></div></div></div></td> </tr><tr><td class="sidebar-content" style="background:#e7f4e7;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90db90;font-size:100%;font-weight:bold;;color: var(--color-base)">Research</div><div class="sidebar-list-content mw-collapsible-content hlist"> <ul><li><a href="/wiki/Biologist" title="Biologist">Biologist</a> (<a href="/wiki/List_of_biologists" title="List of biologists">list)</a></li> <li><a href="/wiki/List_of_biology_awards" title="List of biology awards">List of biology awards</a></li> <li><a href="/wiki/List_of_biology_journals" title="List of biology journals">List of journals</a></li> <li><a href="/wiki/List_of_research_methods_in_biology" title="List of research methods in biology">List of research methods</a></li> <li><a href="/wiki/List_of_unsolved_problems_in_biology" title="List of unsolved problems in biology">List of unsolved problems</a></li></ul></div></div></td> </tr><tr><td class="sidebar-content" style="background:#e7f4e7;"> <div class="sidebar-list mw-collapsible mw-collapsed"><div class="sidebar-list-title" style="text-align:center;padding-bottom:0;background:#90db90;font-size:100%;font-weight:bold;;color: var(--color-base)">Applications</div><div class="sidebar-list-content mw-collapsible-content hlist"> <ul><li><a href="/wiki/Agricultural_science" title="Agricultural science">Agricultural science</a></li> <li><a href="/wiki/Biomedical_sciences" title="Biomedical sciences">Biomedical sciences</a></li> <li><a href="/wiki/Health_technology" title="Health technology">Health technology</a></li> <li><a href="/wiki/Pharming_(genetics)" title="Pharming (genetics)"> Pharming</a></li></ul></div></div></td> </tr><tr><td class="sidebar-below hlist"> <ul><li><span class="nowrap"><span class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Issoria_lathonia.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/16px-Issoria_lathonia.jpg" decoding="async" width="16" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/24px-Issoria_lathonia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/32px-Issoria_lathonia.jpg 2x" data-file-width="629" data-file-height="445" /></a></span> </span><a href="/wiki/Portal:Biology" title="Portal:Biology">Biology portal</a></span></li> <li><span class="nowrap"><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Biology" title="Category:Biology">Category</a></span></li></ul></td></tr><tr><td class="sidebar-navbar"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:TopicTOC-Biology" title="Template:TopicTOC-Biology"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:TopicTOC-Biology" title="Template talk:TopicTOC-Biology"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:TopicTOC-Biology" title="Special:EditPage/Template:TopicTOC-Biology"><abbr title="Edit this template">e</abbr></a></li></ul></div></td></tr></tbody></table> <p><b>Cell biology</b> (also <b>cellular biology</b> or <b>cytology</b>) is a branch of <a href="/wiki/Biology" title="Biology">biology</a> that studies the <a href="/wiki/Anatomy" title="Anatomy">structure</a>, <a href="/wiki/Physiology" title="Physiology">function</a>, and behavior of <a href="/wiki/Cell_(biology)" title="Cell (biology)">cells</a>.<sup id="cite_ref-alberts2017a_1-0" class="reference"><a href="#cite_note-alberts2017a-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> All living organisms are made of cells. A cell is the basic unit of life that is responsible for the living and functioning of organisms.<sup id="cite_ref-Cell_and_Molecular_Biology2_3-0" class="reference"><a href="#cite_note-Cell_and_Molecular_Biology2-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Cell biology is the study of the structural and functional units of cells. Cell biology encompasses both <a href="/wiki/Prokaryote" title="Prokaryote">prokaryotic</a> and <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotic cells</a> and has many subtopics which may include the study of <a href="/wiki/Metabolism" title="Metabolism">cell metabolism</a>, <a href="/wiki/Signaling_pathway" class="mw-redirect" title="Signaling pathway">cell communication</a>, <a href="/wiki/Cell_cycle" title="Cell cycle">cell cycle</a>, <a href="/wiki/Biochemistry" title="Biochemistry">biochemistry</a>, and <a href="/wiki/Cell_(biology)" title="Cell (biology)">cell composition</a>. The study of cells is performed using several <a href="/wiki/Microscopy" title="Microscopy">microscopy</a> techniques, <a href="/wiki/Cell_culture" title="Cell culture">cell culture</a>, and <a href="/wiki/Cell_fractionation" title="Cell fractionation">cell fractionation</a>. These have allowed for and are currently being used for discoveries and research pertaining to how cells function, ultimately giving insight into understanding larger organisms. Knowing the components of cells and how cells work is fundamental to all biological sciences while also being essential for research in <a href="/wiki/Bio-medical" class="mw-redirect" title="Bio-medical">biomedical</a> fields such as <a href="/wiki/Cancer" title="Cancer">cancer</a>, and other diseases. Research in cell biology is interconnected to other fields such as <a href="/wiki/Genetics" title="Genetics">genetics</a>, <a href="/wiki/Molecular_genetics" title="Molecular genetics">molecular genetics</a>, <a href="/wiki/Molecular_biology" title="Molecular biology">molecular biology</a>, <a href="/wiki/Medical_microbiology" title="Medical microbiology">medical microbiology</a>, <a href="/wiki/Immunology" title="Immunology">immunology</a>, and <a href="/wiki/Cytochemistry" title="Cytochemistry">cytochemistry</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="History">History</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=1" title="Edit section: History"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Cells were first seen in 17th-century <a href="/wiki/Europe" title="Europe">Europe</a> with the invention of the <a href="/wiki/Optical_microscope#Compound_microscope" title="Optical microscope">compound microscope</a>. In 1665, <a href="/wiki/Robert_Hooke" title="Robert Hooke">Robert Hooke</a> referred to the building blocks of all living organisms as "cells" (published in <i><a href="/wiki/Micrographia" title="Micrographia">Micrographia</a></i>) after looking at a piece of <a href="/wiki/Cork_(material)" title="Cork (material)">cork</a> and observing a structure reminiscent of a <a href="/wiki/Monastic_cell" title="Monastic cell">monastic cell</a>;<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-EB1911_5-0" class="reference"><a href="#cite_note-EB1911-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> however, the cells were dead. They gave no indication to the actual overall components of a cell. A few years later, in 1674, <a href="/wiki/Anton_Van_Leeuwenhoek" class="mw-redirect" title="Anton Van Leeuwenhoek">Anton Van Leeuwenhoek</a> was the first to analyze live cells in his examination of <a href="/wiki/Algae" title="Algae">algae</a>. Many years later, in 1831, <a href="/wiki/Robert_Brown_(botanist,_born_1773)" title="Robert Brown (botanist, born 1773)">Robert Brown</a> discovered the <a href="/wiki/Cell_nucleus" title="Cell nucleus">nucleus</a>. All of this preceded the <a href="/wiki/Cell_theory" title="Cell theory">cell theory</a> which states that all living things are made up of cells and that cells are organisms' functional and structural units. This was ultimately concluded by plant scientist <a href="/wiki/Matthias_Jakob_Schleiden" title="Matthias Jakob Schleiden">Matthias Schleiden</a><sup id="cite_ref-EB1911_5-1" class="reference"><a href="#cite_note-EB1911-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> and animal scientist <a href="/wiki/Theodor_Schwann" title="Theodor Schwann">Theodor Schwann</a> in 1838, who viewed live cells in plant and animal tissue, respectively.<sup id="cite_ref-Cell_and_Molecular_Biology2_3-1" class="reference"><a href="#cite_note-Cell_and_Molecular_Biology2-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> 19 years later, <a href="/wiki/Rudolf_Virchow" title="Rudolf Virchow">Rudolf Virchow</a> further contributed to the cell theory, adding that all cells come from the division of pre-existing cells.<sup id="cite_ref-Cell_and_Molecular_Biology2_3-2" class="reference"><a href="#cite_note-Cell_and_Molecular_Biology2-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Viruses" class="mw-redirect" title="Viruses">Viruses</a> are not considered in cell biology – they lack the characteristics of a living cell and instead are studied in the <a href="/wiki/Microbiology" title="Microbiology">microbiology</a> subclass of <a href="/wiki/Virology" title="Virology">virology</a>.<sup id="cite_ref-Nature_6-0" class="reference"><a href="#cite_note-Nature-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Techniques">Techniques</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=2" title="Edit section: Techniques"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Cell biology research looks at different ways to culture and manipulate cells outside of a living body to further research in <a href="/wiki/Human_anatomy" class="mw-redirect" title="Human anatomy">human anatomy</a> and <a href="/wiki/Physiology" title="Physiology">physiology</a>, and to derive medications. The techniques by which cells are studied have evolved. Due to advancements in microscopy, techniques and technology have allowed scientists to hold a better understanding of the structure and function of cells. Many techniques commonly used to study cell biology are listed below:<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <ul><li><a href="/wiki/Cell_culture" title="Cell culture">Cell culture</a>: Utilizes rapidly growing cells on media which allows for a large amount of a specific cell type and an efficient way to study cells.<sup id="cite_ref-:12_8-0" class="reference"><a href="#cite_note-:12-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Cell culture is one of the major tools used in cellular and molecular biology, providing excellent model systems for studying the normal physiology and biochemistry of cells (e.g., metabolic studies, aging), the effects of drugs and toxic compounds on the cells, and mutagenesis and carcinogenesis. It is also used in drug screening and development, and large scale manufacturing of biological compounds (e.g., vaccines, therapeutic proteins).</li> <li><a href="/wiki/Fluorescence_microscope" title="Fluorescence microscope">Fluorescence microscopy</a>: Fluorescent markers such as <a href="/wiki/Green_fluorescent_protein" title="Green fluorescent protein">GFP</a>, are used to label a specific component of the cell. Afterwards, a certain light wavelength is used to excite the fluorescent marker which can then be visualized.<sup id="cite_ref-:12_8-1" class="reference"><a href="#cite_note-:12-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Phase-contrast_microscopy" title="Phase-contrast microscopy">Phase-contrast microscopy</a>: Uses the optical aspect of light to represent the solid, liquid, and gas-phase changes as brightness differences.<sup id="cite_ref-:12_8-2" class="reference"><a href="#cite_note-:12-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Confocal_microscopy" title="Confocal microscopy">Confocal microscopy</a>: Combines fluorescence microscopy with imaging by focusing light and snap shooting instances to form a 3-D image.<sup id="cite_ref-:12_8-3" class="reference"><a href="#cite_note-:12-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Transmission_electron_microscopy" title="Transmission electron microscopy">Transmission electron microscopy</a>: Involves metal staining and the passing of electrons through the cells, which will be deflected upon interaction with metal. This ultimately forms an image of the components being studied.<sup id="cite_ref-:12_8-4" class="reference"><a href="#cite_note-:12-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Cytometry" title="Cytometry">Cytometry</a>: The cells are placed in the machine which uses a beam to scatter the cells based on different aspects and can therefore separate them based on size and content. Cells may also be tagged with GFP-fluorescence and can be separated that way as well.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Cell_fractionation" title="Cell fractionation">Cell fractionation</a>: This process requires breaking up the cell using high temperature or sonification followed by <a href="/wiki/Centrifugation" title="Centrifugation">centrifugation</a> to separate the parts of the cell allowing for them to be studied separately.<sup id="cite_ref-:12_8-5" class="reference"><a href="#cite_note-:12-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Cell_types">Cell types</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=3" title="Edit section: Cell types"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Cell_types" class="mw-redirect" title="Cell types">Cell types</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Prokaryote_cell.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c5/Prokaryote_cell.svg/220px-Prokaryote_cell.svg.png" decoding="async" width="220" height="247" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c5/Prokaryote_cell.svg/330px-Prokaryote_cell.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c5/Prokaryote_cell.svg/440px-Prokaryote_cell.svg.png 2x" data-file-width="910" data-file-height="1020" /></a><figcaption>A drawing of a prokaryotic cell</figcaption></figure> <p>There are two fundamental classifications of cells: <a href="/wiki/Prokaryote" title="Prokaryote">prokaryotic</a> and <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotic</a>. Prokaryotic cells are distinguished from eukaryotic cells by the absence of a <a href="/wiki/Cell_nucleus" title="Cell nucleus">cell nucleus</a> or other membrane-bound <a href="/wiki/Organelle" title="Organelle">organelle</a>.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> Prokaryotic cells are much smaller than eukaryotic cells, making them the smallest form of life.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Prokaryotic cells include <a href="/wiki/Bacteria" title="Bacteria">Bacteria</a> and <a href="/wiki/Archaea" title="Archaea">Archaea</a>, and lack an enclosed cell nucleus.  Eukaryotic cells are found in plants, animals, fungi, and protists. They range from 10 to 100 μm in diameter, and their DNA is contained within a membrane-bound nucleus. Eukaryotes are organisms containing eukaryotic cells. The four eukaryotic kingdoms are <b>Animalia, Plantae, Fungi, and Protista.</b><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p><p>They both reproduce through <a href="/wiki/Binary_fission" class="mw-redirect" title="Binary fission">binary fission</a>. Bacteria, the most prominent type, have several <a href="/wiki/Bacterial_cell_structure#Cell_morphology" title="Bacterial cell structure">different shapes</a>, although most are <a href="/wiki/Cocci" class="mw-redirect" title="Cocci">spherical</a> or <a href="/wiki/Bacillus_(shape)" class="mw-redirect" title="Bacillus (shape)">rod-shaped</a>. Bacteria can be classed as either <a href="/wiki/Gram-positive_bacteria" title="Gram-positive bacteria">gram-positive</a> or <a href="/wiki/Gram-negative_bacteria" title="Gram-negative bacteria">gram-negative</a> depending on the <a href="/wiki/Cell_wall#Bacterial_cell_wall" title="Cell wall">cell wall</a> composition. Gram-positive bacteria have a thicker <a href="/wiki/Peptidoglycan_layer" class="mw-redirect" title="Peptidoglycan layer">peptidoglycan layer</a> than gram-negative bacteria. Bacterial structural features include a <a href="/wiki/Flagellum" title="Flagellum">flagellum</a> that helps the cell to move,<sup id="cite_ref-:02_13-0" class="reference"><a href="#cite_note-:02-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Ribosome" title="Ribosome">ribosomes</a> for the translation of RNA to protein,<sup id="cite_ref-:02_13-1" class="reference"><a href="#cite_note-:02-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> and a <a href="/wiki/Nucleoid" title="Nucleoid">nucleoid</a> that holds all the genetic material in a circular structure.<sup id="cite_ref-:02_13-2" class="reference"><a href="#cite_note-:02-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> There are many processes that occur in prokaryotic cells that allow them to survive. In prokaryotes, <b>mRNA synthesis</b> is initiated at a promoter sequence on the DNA template comprising two consensus sequences that recruit RNA polymerase. The prokaryotic polymerase consists of a core enzyme of four protein subunits and a σ protein that assists only with initiation. For instance, in a process termed <a href="/wiki/Conjugation_pilus" class="mw-redirect" title="Conjugation pilus">conjugation</a>, the fertility factor allows the bacteria to possess a pilus which allows it to transmit DNA to another bacteria which lacks the F factor, permitting the transmittance of resistance allowing it to survive in certain environments.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Structure_and_function">Structure and function</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=4" title="Edit section: Structure and function"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Structure_of_eukaryotic_cells">Structure of eukaryotic cells</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=5" title="Edit section: Structure of eukaryotic cells"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Eukaryote" title="Eukaryote">Eukaryote</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Animal_cell_NIH.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Animal_cell_NIH.jpg/220px-Animal_cell_NIH.jpg" decoding="async" width="220" height="153" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Animal_cell_NIH.jpg/330px-Animal_cell_NIH.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Animal_cell_NIH.jpg/440px-Animal_cell_NIH.jpg 2x" data-file-width="600" data-file-height="418" /></a><figcaption>A diagram of an animal cell</figcaption></figure> <p><a href="/wiki/Eukaryotic_cell#Cell_features" class="mw-redirect" title="Eukaryotic cell">Eukaryotic cells</a> are composed of the following organelles: </p> <ul><li><a href="/wiki/Cell_nucleus" title="Cell nucleus">Nucleus</a>: The nucleus of the cell functions as the <a href="/wiki/Genome" title="Genome">genome</a> and genetic information storage for the cell, containing all the <a href="/wiki/DNA" title="DNA">DNA</a> organized in the form of <a href="/wiki/Chromosome" title="Chromosome">chromosomes</a>. It is surrounded by a <a href="/wiki/Nuclear_envelope" title="Nuclear envelope">nuclear envelope</a>, which includes nuclear pores allowing for the transportation of proteins between the inside and outside of the nucleus.<sup id="cite_ref-pmid29107331_15-0" class="reference"><a href="#cite_note-pmid29107331-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> This is also the site for replication of DNA as well as transcription of DNA to RNA. Afterwards, the RNA is modified and transported out to the cytosol to be translated to protein.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Nucleolus" title="Nucleolus">Nucleolus</a>: This structure is within the nucleus, usually dense and spherical. It is the site of ribosomal RNA (rRNA) synthesis, which is needed for ribosomal assembly.</li> <li><a href="/wiki/Endoplasmic_Reticulum" class="mw-redirect" title="Endoplasmic Reticulum">Endoplasmic reticulum (ER)</a>: This functions to synthesize, store, and secrete proteins to the Golgi apparatus.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> Structurally, the endoplasmic reticulum is a network of membranes found throughout the cell and connected to the nucleus. The membranes are slightly different from cell to cell and a cell's function determines the size and structure of the ER.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Mitochondria" class="mw-redirect" title="Mitochondria">Mitochondria</a>: Commonly known as the powerhouse of the cell is a double membrane bound cell organelle.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> This functions for the production of energy or ATP within the cell. Specifically, this is the place where the Krebs cycle or <a href="/wiki/Citric_acid_cycle" title="Citric acid cycle">TCA cycle</a> for the production of NADH and FADH occurs. Afterwards, these products are used within the electron transport chain (ETC) and oxidative phosphorylation for the final production of ATP.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Golgi_apparatus" title="Golgi apparatus">Golgi apparatus</a>: This functions to further process, package, and secrete the proteins to their destination. The proteins contain a signal sequence that allows the Golgi apparatus to recognize and direct it to the correct place. Golgi apparatus also produce <a href="/wiki/Glycoprotein" title="Glycoprotein">glycoproteins</a> and <a href="/wiki/Glycolipid" title="Glycolipid">glycolipids</a>.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Lysosome" title="Lysosome">Lysosome</a>: The lysosome functions to degrade material brought in from the outside of the cell or old organelles. This contains many acid hydrolases, proteases, nucleases, and lipases, which break down the various molecules. <a href="/wiki/Autophagy" title="Autophagy">Autophagy</a> is the process of degradation through lysosomes which occurs when a vesicle buds off from the ER and engulfs the material, then, attaches and fuses with the lysosome to allow the material to be degraded.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Ribosome" title="Ribosome">Ribosomes</a>: Functions to translate RNA to protein. it serves as a site of protein synthesis.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Cytoskeleton" title="Cytoskeleton">Cytoskeleton</a>: Cytoskeleton is a structure that helps to maintain the shape and general organization of the cytoplasm. It anchors organelles within the cells and makes up the structure and stability of the cell. The cytoskeleton is composed of three principal types of protein filaments: actin filaments, intermediate filaments, and microtubules, which are held together and linked to subcellular organelles and the plasma membrane by a variety of accessory proteins.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Cell_membrane" title="Cell membrane">Cell membrane</a>: The cell membrane can be described as a phospholipid bilayer and is also consisted of lipids and proteins.<sup id="cite_ref-:02_13-3" class="reference"><a href="#cite_note-:02-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Because the inside of the bilayer is hydrophobic and in order for molecules to participate in reactions within the cell, they need to be able to cross this membrane layer to get into the cell via <a href="/wiki/Osmotic_pressure" title="Osmotic pressure">osmotic pressure</a>, <a href="/wiki/Diffusion" title="Diffusion">diffusion</a>, concentration gradients, and membrane channels.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Centriole" title="Centriole">Centrioles</a>: Function to produce spindle fibers which are used to separate chromosomes during cell division.</li></ul> <p>Eukaryotic cells may also be composed of the following molecular components: </p> <ul><li><a href="/wiki/Chromatin" title="Chromatin">Chromatin</a>: This makes up <a href="/wiki/Chromosome" title="Chromosome">chromosomes</a> and is a mixture of DNA with various proteins.</li> <li><a href="/wiki/Cilium" title="Cilium">Cilia</a>: They help to propel substances and can also be used for sensory purposes.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading3"><h3 id="Cell_metabolism">Cell metabolism</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=6" title="Edit section: Cell metabolism"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Cell metabolism is necessary for the production of energy for the cell and therefore its survival and includes many pathways and also sustaining the main cell organelles such as the nucleus, the mitochondria, the cell membrane etc. For <a href="/wiki/Cellular_respiration" title="Cellular respiration">cellular respiration</a>, once glucose is available, glycolysis occurs within the cytosol of the cell to produce pyruvate. Pyruvate undergoes decarboxylation using the multi-enzyme complex to form acetyl coA which can readily be used in the <a href="/wiki/Citric_acid_cycle" title="Citric acid cycle">TCA cycle</a> to produce NADH and FADH<sub>2</sub>. These products are involved in the <a href="/wiki/Electron_transport_chain" title="Electron transport chain">electron transport chain</a> to ultimately form a proton gradient across the inner mitochondrial membrane. This gradient can then drive the production of ATP and <style data-mw-deduplicate="TemplateStyles:r1123817410">.mw-parser-output .template-chem2-su{display:inline-block;font-size:80%;line-height:1;vertical-align:-0.35em}.mw-parser-output .template-chem2-su>span{display:block;text-align:left}.mw-parser-output sub.template-chem2-sub{font-size:80%;vertical-align:-0.35em}.mw-parser-output sup.template-chem2-sup{font-size:80%;vertical-align:0.65em}</style><span class="chemf nowrap">H<sub class="template-chem2-sub">2</sub>O</span> during <a href="/wiki/Oxidative_phosphorylation" title="Oxidative phosphorylation">oxidative phosphorylation</a>.<sup id="cite_ref-:0_27-0" class="reference"><a href="#cite_note-:0-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> Metabolism in plant cells includes <a href="/wiki/Photosynthesis" title="Photosynthesis">photosynthesis</a> which is simply the exact opposite of respiration as it ultimately produces molecules of glucose. </p> <div class="mw-heading mw-heading3"><h3 id="Cell_signaling">Cell signaling</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=7" title="Edit section: Cell signaling"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Cell_signaling" title="Cell signaling">Cell signaling</a></div> <p><a href="/wiki/Cell_signaling" title="Cell signaling">Cell signaling</a> or cell communication is important for cell regulation and for cells to process information from the environment and respond accordingly. Signaling can occur through direct cell contact or <a href="/wiki/Endocrine_system" title="Endocrine system">endocrine</a>, <a href="/wiki/Paracrine_signaling" title="Paracrine signaling">paracrine</a>, and <a href="/wiki/Autocrine_signaling" title="Autocrine signaling">autocrine signaling</a>. Direct cell-cell contact is when a receptor on a cell binds a molecule that is attached to the membrane of another cell. Endocrine signaling occurs through molecules secreted into the bloodstream. Paracrine signaling uses molecules diffusing between two cells to communicate. Autocrine is a cell sending a signal to itself by secreting a molecule that binds to a receptor on its surface. Forms of communication can be through: </p> <ul><li><a href="/wiki/Ion_channel" title="Ion channel">Ion channels</a>: Can be of different types such as voltage or ligand gated ion channels. They allow for the outflow and inflow of molecules and ions.</li> <li><a href="/wiki/G_protein-coupled_receptor" title="G protein-coupled receptor">G-protein coupled receptor</a> (GPCR): Is widely recognized to contain seven transmembrane domains. The ligand binds on the extracellular domain and once the ligand binds, this signals a guanine exchange factor to convert GDP to GTP and activate the G-α subunit. G-α can target other proteins such as adenyl cyclase or phospholipase C, which ultimately produce secondary messengers such as cAMP, Ip3, DAG, and calcium. These secondary messengers function to amplify signals and can target ion channels or other enzymes. One example for amplification of a signal is cAMP binding to and activating PKA by removing the regulatory subunits and releasing the catalytic subunit. The catalytic subunit has a nuclear localization sequence which prompts it to go into the nucleus and phosphorylate other proteins to either repress or activate gene activity.<sup id="cite_ref-:0_27-1" class="reference"><a href="#cite_note-:0-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Receptor_tyrosine_kinase" title="Receptor tyrosine kinase">Receptor tyrosine kinases</a>: Bind growth factors, further promoting the tyrosine on the intracellular portion of the protein to cross phosphorylate. The phosphorylated tyrosine becomes a landing pad for proteins containing an SH2 domain allowing for the activation of Ras and the involvement of the <a href="/wiki/Mitogen-activated_protein_kinase" title="Mitogen-activated protein kinase">MAP kinase pathway</a>.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Growth_and_development">Growth and development</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=8" title="Edit section: Growth and development"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Eukaryotic_cell_cycle">Eukaryotic cell cycle</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=9" title="Edit section: Eukaryotic cell cycle"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Cell_cycle" title="Cell cycle">Cell cycle</a></div> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Animal_cell_cycle-en.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Animal_cell_cycle-en.svg/386px-Animal_cell_cycle-en.svg.png" decoding="async" width="386" height="375" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Animal_cell_cycle-en.svg/579px-Animal_cell_cycle-en.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Animal_cell_cycle-en.svg/772px-Animal_cell_cycle-en.svg.png 2x" data-file-width="7207" data-file-height="7003" /></a><figcaption>The process of <a href="/wiki/Cell_division" title="Cell division">cell division</a> in the animal <a href="/wiki/Cell_cycle" title="Cell cycle">cell cycle</a></figcaption></figure> <p>Cells are the foundation of all organisms and are the fundamental units of life. The growth and development of cells are essential for the maintenance of the host and survival of the organism. For this process, the cell goes through the steps of the <a href="/wiki/Cell_cycle" title="Cell cycle">cell cycle</a> and development which involves cell growth, <a href="/wiki/DNA_replication" title="DNA replication">DNA replication</a>, <a href="/wiki/Cell_division" title="Cell division">cell division</a>, regeneration, and <a href="/wiki/Cell_death" title="Cell death">cell death</a>. </p><p>The cell cycle is divided into four distinct <a href="/wiki/Cell_cycle#Phases" title="Cell cycle">phases</a>: G1, S, G2, and M. The G phase – which is the cell growth phase – makes up approximately 95% of the cycle. The proliferation of cells is instigated by progenitors. All cells start out in an identical form and can essentially become any type of cells. Cell signaling such as induction can influence nearby cells to determinate the type of cell it will become. Moreover, this allows cells of the same type to aggregate and form tissues, then organs, and ultimately systems. The G1, G2, and S phase (DNA replication, damage and repair) are considered to be the interphase portion of the cycle, while the M phase (<a href="/wiki/Mitosis" title="Mitosis">mitosis</a>) is the <a href="/wiki/Cell_division" title="Cell division">cell division</a> portion of the cycle. Mitosis is composed of many stages which include, prophase, metaphase, anaphase, telophase, and cytokinesis, respectively. The ultimate result of mitosis is the formation of two identical daughter cells. </p><p>The cell cycle is regulated in <a href="/wiki/Cell_cycle_checkpoint" title="Cell cycle checkpoint">cell cycle checkpoints</a>, by a series of signaling factors and complexes such as cyclins, <a href="/wiki/Cyclin-dependent_kinase" title="Cyclin-dependent kinase">cyclin-dependent kinase</a>, and <a href="/wiki/P53" title="P53">p53</a>. When the cell has completed its growth process and if it is found to be damaged or altered, it undergoes cell death, either by <a href="/wiki/Apoptosis" title="Apoptosis">apoptosis</a> or <a href="/wiki/Necrosis" title="Necrosis">necrosis</a>, to eliminate the threat it can cause to the organism's survival.<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Cell_mortality,_cell_lineage_immortality"><span id="Cell_mortality.2C_cell_lineage_immortality"></span>Cell mortality, cell lineage immortality</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=10" title="Edit section: Cell mortality, cell lineage immortality"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The ancestry of each present day cell presumably traces back, in an unbroken lineage for over 3 billion years to the <a href="/wiki/Abiogenesis" title="Abiogenesis">origin of life</a>. It is not actually cells that are <a href="/wiki/Immortality" title="Immortality">immortal</a> but multi-generational cell lineages.<sup id="cite_ref-Bernstein2000_30-0" class="reference"><a href="#cite_note-Bernstein2000-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> The immortality of a cell lineage depends on the maintenance of <a href="/wiki/Cell_division" title="Cell division">cell division</a> potential. This potential may be lost in any particular lineage because of cell damage, <a href="/wiki/Cellular_differentiation" title="Cellular differentiation">terminal differentiation</a> as occurs in nerve cells, or programmed cell death (<a href="/wiki/Apoptosis" title="Apoptosis">apoptosis</a>) during development. Maintenance of cell division potential over successive generations depends on the avoidance and the accurate repair of cellular damage, particularly <a href="/wiki/DNA_damage_(naturally_occurring)" title="DNA damage (naturally occurring)">DNA damage</a>. In sexual organisms, continuity of the <a href="/wiki/Germline" title="Germline">germline</a> depends on the effectiveness of processes for avoiding DNA damage and <a href="/wiki/DNA_repair" title="DNA repair">repairing those DNA damages</a> that do occur. Sexual processes in <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotes</a>, as well as in <a href="/wiki/Prokaryote" title="Prokaryote">prokaryotes</a>, provide an opportunity for effective repair of DNA damages in the germ line by <a href="/wiki/Homologous_recombination" title="Homologous recombination">homologous recombination</a>.<sup id="cite_ref-Bernstein2000_30-1" class="reference"><a href="#cite_note-Bernstein2000-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Cell_cycle_phases">Cell cycle phases</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=11" title="Edit section: Cell cycle phases"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The cell cycle is a four-stage process that a cell goes through as it develops and divides. It includes Gap 1 (G1), synthesis (S), Gap 2 (G2), and mitosis (M). The cell either restarts the cycle from G1 or leaves the cycle through G0 after completing the cycle. The cell can progress from G0 through terminal differentiation. Finally, the interphase refers to the phases of the cell cycle that occur between one mitosis and the next, and includes G1, S, and G2. Thus, the phases are: </p> <ul><li><b>G1 phase</b>: the cell grows in size and its contents are replicated.</li> <li><b>S phase</b>: the cell replicates each of the 46 chromosomes.</li> <li><b>G2 phase</b>: in preparation for cell division, new organelles and proteins form.</li> <li><b>M phase</b>: cytokinesis occurs, resulting in two identical daughter cells.</li> <li><b>G0 phase</b>: the two cells enter a resting stage where they do their job without actively preparing to divide.<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Pathology">Pathology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=12" title="Edit section: Pathology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Cytopathology" title="Cytopathology">Cytopathology</a></div> <p>The scientific branch that studies and diagnoses diseases on the cellular level is called <a href="/wiki/Cytopathology" title="Cytopathology">cytopathology</a>. Cytopathology is generally used on samples of free cells or tissue fragments, in contrast to the <a href="/wiki/Pathology" title="Pathology">pathology</a> branch of <a href="/wiki/Histopathology" title="Histopathology">histopathology</a>, which studies whole tissues. Cytopathology is commonly used to investigate diseases involving a wide range of body sites, often to aid in the diagnosis of cancer but also in the diagnosis of some infectious diseases and other inflammatory conditions. For example, a common application of cytopathology is the <a href="/wiki/Pap_smear" class="mw-redirect" title="Pap smear">Pap smear</a>, a <a href="/wiki/Screening_(medicine)" title="Screening (medicine)">screening test</a> used to detect <a href="/wiki/Cervical_cancer" title="Cervical cancer">cervical cancer</a>, and <a href="/wiki/Cervical_intraepithelial_neoplasia" title="Cervical intraepithelial neoplasia">precancerous cervical lesions</a> that may lead to cervical cancer.<sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Cell_cycle_checkpoints_and_DNA_damage_repair_system">Cell cycle checkpoints and DNA damage repair system</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=13" title="Edit section: Cell cycle checkpoints and DNA damage repair system"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The cell cycle is composed of a number of well-ordered, consecutive stages that result in cellular division. The fact that cells do not begin the next stage until the last one is finished, is a significant element of cell cycle regulation. Cell cycle checkpoints are characteristics that constitute an excellent monitoring strategy for accurate cell cycle and divisions. Cdks, associated cyclin counterparts, protein kinases, and phosphatases regulate cell growth and division from one stage to another.<sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> The cell cycle is controlled by the temporal activation of Cdks, which is governed by cyclin partner interaction, phosphorylation by particular protein kinases, and de-phosphorylation by Cdc25 family phosphatases. In response to DNA damage, a cell's DNA repair reaction is a cascade of signaling pathways that leads to checkpoint engagement, regulates, the repairing mechanism in DNA, cell cycle alterations, and apoptosis. Numerous biochemical structures, as well as processes that detect damage in DNA, are ATM and ATR, which induce the DNA repair checkpoints<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup> </p><p>The cell cycle is a sequence of activities in which cell organelles are duplicated and subsequently separated into daughter cells with precision. There are major events that happen during a cell cycle. The processes that happen in the cell cycle include cell development, replication and segregation of chromosomes.  The cell cycle checkpoints are surveillance systems that keep track of the cell cycle's integrity, accuracy, and chronology. Each checkpoint serves as an alternative cell cycle endpoint, wherein the cell's parameters are examined and only when desirable characteristics are fulfilled does the cell cycle advance through the distinct steps. The cell cycle's goal is to precisely copy each organism's DNA and afterwards equally split the cell and its components between the two new cells. Four main stages occur in the eukaryotes. In G1, the cell is usually active and continues to grow rapidly, while in G2, the cell growth continues while protein molecules become ready for separation. These are not dormant times; they are when cells gain mass, integrate growth factor receptors, establish a replicated genome, and prepare for chromosome segregation. DNA replication is restricted to a separate Synthesis in eukaryotes, which is also known as the S-phase. During mitosis, which is also known as the M-phase, the segregation of the chromosomes occur.<sup id="cite_ref-Giacomello_204–224_36-0" class="reference"><a href="#cite_note-Giacomello_204–224-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> DNA, like every other molecule, is capable of undergoing a wide range of chemical reactions. Modifications in DNA's sequence, on the other hand, have a considerably bigger impact than modifications in other cellular constituents like RNAs or proteins because DNA acts as a permanent copy of the cell genome. When erroneous nucleotides are incorporated during DNA replication, mutations can occur. The majority of DNA damage is fixed by removing the defective bases and then re-synthesizing the excised area. On the other hand, some DNA lesions can be mended by reversing the damage, which may be a more effective method of coping with common types of DNA damage. Only a few forms of DNA damage are mended in this fashion, including pyrimidine dimers caused by ultraviolet (UV) light changed by the insertion of methyl or ethyl groups at the purine ring's O6 position.<sup id="cite_ref-37" class="reference"><a href="#cite_note-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Mitochondrial_membrane_dynamics">Mitochondrial membrane dynamics</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=14" title="Edit section: Mitochondrial membrane dynamics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Mitochondria are commonly referred to as the cell's "powerhouses" because of their capacity to effectively produce ATP which is essential to maintain cellular homeostasis and metabolism. Moreover, researchers have gained a better knowledge of mitochondria's significance in cell biology because of the discovery of cell signaling pathways by mitochondria which are crucial platforms for cell function regulation such as apoptosis. Its physiological adaptability is strongly linked to the cell mitochondrial channel's ongoing reconfiguration through a range of mechanisms known as mitochondrial membrane dynamics, including endomembrane fusion and fragmentation (separation) and ultrastructural membrane remodeling. As a result, mitochondrial dynamics regulate and frequently choreograph not only metabolic but also complicated cell signaling processes such as cell pluripotent stem cells, proliferation, maturation, aging, and mortality. Mutually, post-translational alterations of mitochondrial apparatus and the development of transmembrane contact sites among mitochondria and other structures, which both have the potential to link signals from diverse routes that affect mitochondrial membrane dynamics substantially,<sup id="cite_ref-Giacomello_204–224_36-1" class="reference"><a href="#cite_note-Giacomello_204–224-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> Mitochondria are wrapped by two membranes: an inner mitochondrial membrane (IMM) and an outer mitochondrial membrane (OMM), each with a distinctive function and structure, which parallels their dual role as cellular powerhouses and signaling organelles. The inner mitochondrial membrane divides the mitochondrial lumen into two parts: the inner border membrane, which runs parallel to the OMM, and the cristae, which are deeply twisted, multinucleated invaginations that give room for surface area enlargement and house the mitochondrial respiration apparatus. The outer mitochondrial membrane, on the other hand, is soft and permeable. It, therefore, acts as a foundation for cell signaling pathways to congregate, be deciphered, and be transported into mitochondria. Furthermore, the OMM connects to other cellular organelles, such as the endoplasmic reticulum (ER), lysosomes, endosomes, and the plasma membrane. Mitochondria play a wide range of roles in cell biology, which is reflected in their morphological diversity. Ever since the beginning of the mitochondrial study, it has been well documented that mitochondria can have a variety of forms, with both their general and ultra-structural morphology varying greatly among cells, during the cell cycle, and in response to metabolic or cellular cues. Mitochondria can exist as independent organelles or as part of larger systems; they can also be unequally distributed in the cytosol through regulated mitochondrial transport and placement to meet the cell's localized energy requirements. Mitochondrial dynamics refers to the adaptive and variable aspect of mitochondria, including their shape and subcellular distribution.<sup id="cite_ref-Giacomello_204–224_36-2" class="reference"><a href="#cite_note-Giacomello_204–224-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Autophagy">Autophagy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=15" title="Edit section: Autophagy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Autophagy" title="Autophagy">Autophagy</a></div> <p>Autophagy is a self-degradative mechanism that regulates energy sources during growth and reaction to dietary stress. Autophagy also cleans up after itself, clearing aggregated proteins, cleaning damaged structures including mitochondria and endoplasmic reticulum and eradicating intracellular infections. Additionally, autophagy has antiviral and antibacterial roles within the cell, and it is involved at the beginning of distinctive and adaptive immune responses to viral and bacterial contamination. Some viruses include virulence proteins that prevent autophagy, while others utilize autophagy elements for intracellular development or cellular splitting.<sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> Macro autophagy, micro autophagy, and chaperon-mediated autophagy are the three basic types of autophagy. When macro autophagy is triggered, an exclusion membrane incorporates a section of the cytoplasm, generating the autophagosome, a distinctive double-membraned organelle. The autophagosome then joins the lysosome to create an autolysosome, with lysosomal enzymes degrading the components. In micro autophagy, the lysosome or vacuole engulfs a piece of the cytoplasm by invaginating or protruding the lysosomal membrane to enclose the cytosol or organelles. The <a href="/wiki/Chaperone-mediated_autophagy" title="Chaperone-mediated autophagy">chaperone-mediated autophagy</a> (CMA) protein quality assurance by digesting oxidized and altered proteins under stressful circumstances and supplying amino acids through protein denaturation.<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">[</span>39<span class="cite-bracket">]</span></a></sup> Autophagy is the primary intrinsic degradative system for peptides, fats, carbohydrates, and other cellular structures. In both physiologic and stressful situations, this cellular progression is vital for upholding the correct cellular balance. Autophagy instability leads to a variety of illness symptoms, including inflammation, biochemical disturbances, aging, and neurodegenerative, due to its involvement in controlling cell integrity. The modification of the autophagy-lysosomal networks is a typical hallmark of many neurological and muscular illnesses. As a result, autophagy has been identified as a potential strategy for the prevention and treatment of various disorders. Many of these disorders are prevented or improved by consuming polyphenol in the meal. As a result, natural compounds with the ability to modify the autophagy mechanism are seen as a potential therapeutic option.<sup id="cite_ref-40" class="reference"><a href="#cite_note-40"><span class="cite-bracket">[</span>40<span class="cite-bracket">]</span></a></sup> The creation of the double membrane (phagophore), which would be known as nucleation, is the first step in macro-autophagy. The phagophore approach indicates dysregulated polypeptides or defective organelles that come from the cell membrane, Golgi apparatus, endoplasmic reticulum, and mitochondria. With the conclusion of the autophagocyte, the phagophore's enlargement comes to an end. The auto-phagosome combines with the lysosomal vesicles to formulate an auto-lysosome that degrades the encapsulated substances, referred to as phagocytosis.<sup id="cite_ref-41" class="reference"><a href="#cite_note-41"><span class="cite-bracket">[</span>41<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Notable_cell_biologists">Notable cell biologists</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=16" title="Edit section: Notable cell biologists"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 22em;"> <ul><li><a href="/wiki/Jean_Baptiste_Carnoy" title="Jean Baptiste Carnoy">Jean Baptiste Carnoy</a></li> <li><a href="/wiki/Peter_Agre" title="Peter Agre">Peter Agre</a></li> <li><a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></li> <li><a href="/wiki/Robert_Brown_(Scottish_botanist_from_Montrose)" class="mw-redirect" title="Robert Brown (Scottish botanist from Montrose)">Robert Brown</a></li> <li><a href="/wiki/Geoffrey_M._Cooper" title="Geoffrey M. Cooper">Geoffrey M. Cooper</a></li> <li><a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a></li> <li><a href="/wiki/Henri_Dutrochet" title="Henri Dutrochet">Henri Dutrochet</a></li> <li><a href="/wiki/Robert_Hooke" title="Robert Hooke">Robert Hooke</a></li> <li><a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a></li> <li><a href="/wiki/Marc_Kirschner" title="Marc Kirschner">Marc Kirschner</a></li> <li><a href="/wiki/Anton_van_Leeuwenhoek" class="mw-redirect" title="Anton van Leeuwenhoek">Anton van Leeuwenhoek</a></li> <li><a href="/wiki/Ira_Mellman" title="Ira Mellman">Ira Mellman</a></li> <li><a href="/wiki/Marta_Mi%C4%85czy%C5%84ska" title="Marta Miączyńska">Marta Miączyńska</a><sup id="cite_ref-42" class="reference"><a href="#cite_note-42"><span class="cite-bracket">[</span>42<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Peter_D._Mitchell" title="Peter D. Mitchell">Peter D. Mitchell</a></li> <li><a href="/wiki/Rudolf_Virchow" title="Rudolf Virchow">Rudolf Virchow</a></li> <li><a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></li> <li><a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Emil Palade</a></li> <li><a href="/wiki/Keith_R._Porter" title="Keith R. Porter">Keith R. Porter</a></li> <li><a href="/wiki/Ray_Rappaport" title="Ray Rappaport">Ray Rappaport</a></li> <li><a href="/wiki/Michael_Swann" title="Michael Swann">Michael Swann</a></li> <li><a href="/wiki/Roger_Tsien" class="mw-redirect" title="Roger Tsien">Roger Tsien</a></li> <li><a href="/wiki/Edmund_Beecher_Wilson" title="Edmund Beecher Wilson">Edmund Beecher Wilson</a></li> <li><a href="/wiki/Kenneth_R._Miller" title="Kenneth R. Miller">Kenneth R. Miller</a></li> <li><a href="/wiki/Matthias_Jakob_Schleiden" title="Matthias Jakob Schleiden">Matthias Jakob Schleiden</a></li> <li><a href="/wiki/Theodor_Schwann" title="Theodor Schwann">Theodor Schwann</a></li> <li><a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></li> <li><a href="/wiki/Jan_Evangelista_Purkyn%C4%9B" title="Jan Evangelista Purkyně">Jan Evangelista Purkyně</a></li></ul> </div> <style data-mw-deduplicate="TemplateStyles:r1248256098">@media all and (max-width:720px){.mw-parser-output .mod-gallery{width:100%!important}}.mw-parser-output .mod-gallery{display:table}.mw-parser-output .mod-gallery-default{background:transparent;margin-top:4px}.mw-parser-output .mod-gallery-center{margin-left:auto;margin-right:auto}.mw-parser-output .mod-gallery-left{float:left}.mw-parser-output .mod-gallery-right{float:right}.mw-parser-output .mod-gallery-none{float:none}.mw-parser-output .mod-gallery-collapsible{width:100%}.mw-parser-output .mod-gallery .title,.mw-parser-output .mod-gallery .main,.mw-parser-output .mod-gallery .footer{display:table-row}.mw-parser-output .mod-gallery .title>div{display:table-cell;padding:0 4px 4px;text-align:center;font-weight:bold}.mw-parser-output .mod-gallery .main>div{display:table-cell}.mw-parser-output .mod-gallery .gallery{line-height:1.35em}.mw-parser-output .mod-gallery .footer>div{display:table-cell;padding:4px;text-align:right;font-size:85%;line-height:1em}.mw-parser-output .mod-gallery .title>div *,.mw-parser-output .mod-gallery .footer>div *{overflow:visible}.mw-parser-output .mod-gallery .gallerybox img{background:none!important}.mw-parser-output .mod-gallery .bordered-images .thumb img{border:solid var(--background-color-neutral,#eaecf0)1px}.mw-parser-output .mod-gallery .whitebg .thumb{background:var(--background-color-base,#fff)!important}</style><div class="mod-gallery mod-gallery-default"><div class="main"><div><ul class="gallery mw-gallery-traditional nochecker bordered-images whitebg"> <li class="gallerybox" style="width: 215px"> <div class="thumb" style="width: 210px; height: 210px;"><span typeof="mw:File"><a href="/wiki/File:Jan_Evangelista_Purkyne_2_(cropped).jpg" class="mw-file-description" title="Czech anatomist Jan Evangelista Purkyně is best known for his 1837 discovery of Purkinje cells."><img alt="Czech anatomist Jan Evangelista Purkyně is best known for his 1837 discovery of Purkinje cells." src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Jan_Evangelista_Purkyne_2_%28cropped%29.jpg/136px-Jan_Evangelista_Purkyne_2_%28cropped%29.jpg" decoding="async" width="136" height="180" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Jan_Evangelista_Purkyne_2_%28cropped%29.jpg/204px-Jan_Evangelista_Purkyne_2_%28cropped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Jan_Evangelista_Purkyne_2_%28cropped%29.jpg/273px-Jan_Evangelista_Purkyne_2_%28cropped%29.jpg 2x" data-file-width="1144" data-file-height="1510" /></a></span></div> <div class="gallerytext">Czech anatomist <a href="/wiki/Jan_Evangelista_Purkyn%C4%9B" title="Jan Evangelista Purkyně">Jan Evangelista Purkyně</a> is best known for his 1837 discovery of <a href="/wiki/Purkinje_cell" title="Purkinje cell">Purkinje cells</a>.</div> </li> <li class="gallerybox" style="width: 215px"> <div class="thumb" style="width: 210px; height: 210px;"><span typeof="mw:File"><a href="/wiki/File:Theodor_Schwann_Litho.jpg" class="mw-file-description" title="Theodor Schwann, discoverer of the Schwann cell"><img alt="Theodor Schwann, discoverer of the Schwann cell" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Theodor_Schwann_Litho.jpg/134px-Theodor_Schwann_Litho.jpg" decoding="async" width="134" height="180" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Theodor_Schwann_Litho.jpg/201px-Theodor_Schwann_Litho.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Theodor_Schwann_Litho.jpg/267px-Theodor_Schwann_Litho.jpg 2x" data-file-width="1260" data-file-height="1695" /></a></span></div> <div class="gallerytext"><a href="/wiki/Theodor_Schwann" title="Theodor Schwann">Theodor Schwann</a>, discoverer of the <a href="/wiki/Schwann_cell" title="Schwann cell">Schwann cell</a></div> </li> <li class="gallerybox" style="width: 215px"> <div class="thumb" style="width: 210px; height: 210px;"><span typeof="mw:File"><a href="/wiki/File:Nobel_Laureates_1042_(30647248184).jpg" class="mw-file-description" title="Yoshinori Ohsumi, Nobel Prize winner for work on autophagy"><img alt="Yoshinori Ohsumi, Nobel Prize winner for work on autophagy" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/67/Nobel_Laureates_1042_%2830647248184%29.jpg/135px-Nobel_Laureates_1042_%2830647248184%29.jpg" decoding="async" width="135" height="180" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/67/Nobel_Laureates_1042_%2830647248184%29.jpg/202px-Nobel_Laureates_1042_%2830647248184%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/67/Nobel_Laureates_1042_%2830647248184%29.jpg/270px-Nobel_Laureates_1042_%2830647248184%29.jpg 2x" data-file-width="3456" data-file-height="4608" /></a></span></div> <div class="gallerytext"><a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a>, Nobel Prize winner for work on <a href="/wiki/Autophagy" title="Autophagy">autophagy</a></div> </li> </ul></div></div></div> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=17" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239009302">.mw-parser-output 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.citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFAlbertsJohnsonMorganRaff2015" class="citation book cs1">Alberts, Bruce; Johnson, Alexander D.; Morgan, David; Raff, Martin; Roberts, Keith; Walter, Peter (2015). 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"Chapter 2. Technologies for Detecting Metals in Single Cells. Section 4. Intrinsic X-Ray Fluorescence". In Bani, Lucia (ed.). <i>Metallomics and the Cell</i>. Metal Ions in Life Sciences. Vol. 12. Springer. pp. 15–40. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2F978-94-007-5561-1_2">10.1007/978-94-007-5561-1_2</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-94-007-5560-4" title="Special:BookSources/978-94-007-5560-4"><bdi>978-94-007-5560-4</bdi></a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/23595669">23595669</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Chapter+2.+Technologies+for+Detecting+Metals+in+Single+Cells.+Section+4.+Intrinsic+X-Ray+Fluorescence&rft.btitle=Metallomics+and+the+Cell&rft.series=Metal+Ions+in+Life+Sciences&rft.pages=15-40&rft.pub=Springer&rft.date=2013&rft_id=info%3Apmid%2F23595669&rft_id=info%3Adoi%2F10.1007%2F978-94-007-5561-1_2&rft.isbn=978-94-007-5560-4&rft.aulast=Penner-Hahn&rft.aufirst=James+E.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACell+biology" class="Z3988"></span>electronic-book <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-94-007-5561-1" title="Special:BookSources/978-94-007-5561-1">978-94-007-5561-1</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://www.worldcat.org/search?fq=x0:jrnl&q=n2:1559-0836">1559-0836</a>electronic-<link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://www.worldcat.org/search?fq=x0:jrnl&q=n2:1868-0402">1868-0402</a></li> <li>Cell and Molecular Biology by Karp 5th Ed., <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-471-46580-1" title="Special:BookSources/0-471-46580-1">0-471-46580-1</a></li> <li><span class="noviewer" typeof="mw:File"><span><img alt="Public Domain" src="//upload.wikimedia.org/wikipedia/en/thumb/6/62/PD-icon.svg/12px-PD-icon.svg.png" decoding="async" width="12" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/6/62/PD-icon.svg/18px-PD-icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/6/62/PD-icon.svg/24px-PD-icon.svg.png 2x" data-file-width="196" data-file-height="196" /></span></span> This article incorporates <a href="/wiki/Copyright_status_of_works_by_the_federal_government_of_the_United_States" title="Copyright status of works by the federal government of the United States">public domain material</a> from <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20091208004309/https://www.ncbi.nlm.nih.gov/About/primer/index.html"><i>Science Primer</i></a>. <a href="/wiki/NCBI" class="mw-redirect" title="NCBI">NCBI</a>. Archived from <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/About/primer/index.html">the original</a> on 8 December 2009.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Science+Primer&rft.pub=NCBI&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2FAbout%2Fprimer%2Findex.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACell+biology" class="Z3988"></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cell_biology&action=edit&section=20" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikibooks-logo-en-noslogan.svg/40px-Wikibooks-logo-en-noslogan.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikibooks-logo-en-noslogan.svg/60px-Wikibooks-logo-en-noslogan.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikibooks-logo-en-noslogan.svg/80px-Wikibooks-logo-en-noslogan.svg.png 2x" data-file-width="400" data-file-height="400" /></span></span></div> <div class="side-box-text plainlist">Wikibooks has more on the topic of: <i><b><a href="https://en.wikibooks.org/wiki/Special:Search/Cell_biology" class="extiw" title="wikibooks:Special:Search/Cell biology">Cell biology</a></b></i></div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985"><div class="side-box metadata side-box-right"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-abovebelow"> <a href="/wiki/Wikipedia:The_Wikipedia_Library" title="Wikipedia:The Wikipedia Library">Library resources</a> about <br /> <b>Cell biology</b> <hr /></div> <div class="side-box-flex"> <div class="side-box-text plainlist"><ul><li><a class="external text" href="https://ftl.toolforge.org/cgi-bin/ftl?st=wp&su=Cell+biology">Resources in your library</a></li> </ul></div></div> </div> <ul><li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Commons-logo.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span> Media related to <a href="https://commons.wikimedia.org/wiki/Category:Cell_biology" class="extiw" title="commons:Category:Cell biology">Cell biology</a> at Wikimedia Commons</li> <li><i><a rel="nofollow" class="external text" href="https://onlinelibrary.wiley.com/journal/14749726">Aging Cell</a></i></li> <li><a rel="nofollow" class="external text" href="https://embryo.asu.edu/pages/francis-harry-compton-crick-1916-2004">"Francis Harry Compton Crick (1916–2004)" by A. Andrei at the Embryo Project Encyclopedia</a></li> <li><a rel="nofollow" class="external text" href="https://docs.google.com/presentation/d/1oLBCXt6wVUb9-wh1WvAhpMJuqY4z4UxrMBpwGRPolHo/edit?usp=sharing">"Biology Resource By Professor Lin."</a></li></ul> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output 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srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span><b><a href="/wiki/Category:Cell_biology" title="Category:Cell biology">Cell biology</a></b></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Structures_of_the_cell_/_organelles" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" 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href="/wiki/Endomembrane_system" title="Endomembrane system">Endomembrane <br />system</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cell_membrane" title="Cell membrane">Cell membrane</a></li> <li><a href="/wiki/Cell_nucleus" title="Cell nucleus">Nucleus</a></li> <li><a href="/wiki/Endoplasmic_reticulum" title="Endoplasmic reticulum">Endoplasmic reticulum</a></li> <li><a href="/wiki/Golgi_apparatus" title="Golgi apparatus">Golgi apparatus</a></li> <li><a href="/wiki/Parenthesome" title="Parenthesome">Parenthesome</a></li> <li><a href="/wiki/Autophagosome" title="Autophagosome">Autophagosome</a></li> <li><a href="/wiki/Vesicle_(biology_and_chemistry)" title="Vesicle (biology and chemistry)">Vesicle</a> <ul><li><a href="/wiki/Exosome_(vesicle)" title="Exosome (vesicle)">Exosome</a></li> <li><a href="/wiki/Lysosome" title="Lysosome">Lysosome</a></li> <li><a href="/wiki/Endosome" title="Endosome">Endosome</a></li> <li><a href="/wiki/Phagosome" title="Phagosome">Phagosome</a></li> <li><a href="/wiki/Vacuole" title="Vacuole">Vacuole</a></li> <li><a href="/wiki/Acrosome" title="Acrosome">Acrosome</a></li></ul></li> <li><a href="/wiki/Granule_(cell_biology)" title="Granule (cell biology)">Cytoplasmic granule</a> <ul><li><a href="/wiki/Melanosome" title="Melanosome">Melanosome</a></li> <li><a href="/wiki/Microbody" title="Microbody">Microbody</a></li> <li><a href="/wiki/Glyoxysome" title="Glyoxysome">Glyoxysome</a></li> <li><a href="/wiki/Peroxisome" title="Peroxisome">Peroxisome</a></li> <li><a href="/wiki/Weibel%E2%80%93Palade_body" title="Weibel–Palade body">Weibel–Palade body</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cytoskeleton" title="Cytoskeleton">Cytoskeleton</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Microfilament" title="Microfilament">Microfilament</a></li> <li><a href="/wiki/Intermediate_filament" title="Intermediate filament">Intermediate filament</a></li> <li><a href="/wiki/Microtubule" title="Microtubule">Microtubule</a></li> <li><a href="/wiki/Prokaryotic_cytoskeleton" title="Prokaryotic cytoskeleton">Prokaryotic cytoskeleton</a></li> <li><a href="/wiki/Microtubule_organizing_center" title="Microtubule organizing center">Microtubule organizing center</a> <ul><li><a href="/wiki/Centrosome" title="Centrosome">Centrosome</a></li> <li><a href="/wiki/Centriole" title="Centriole">Centriole</a></li> <li><a href="/wiki/Basal_body" title="Basal body">Basal body</a></li> <li><a href="/wiki/Spindle_pole_body" title="Spindle pole body">Spindle pole body</a></li></ul></li> <li><a href="/wiki/Myofibril" title="Myofibril">Myofibril</a></li> <li><a href="/wiki/Undulipodium" title="Undulipodium">Undulipodium</a> <ul><li><a href="/wiki/Cilium" title="Cilium">Cilium</a></li> <li><a href="/wiki/Flagellum#Eukaryotic" title="Flagellum">Flagellum</a></li> <li><a href="/wiki/Axoneme" title="Axoneme">Axoneme</a></li> <li><a href="/wiki/Radial_spoke" title="Radial spoke">Radial spoke</a></li></ul></li> <li><a href="/wiki/Pseudopodium" class="mw-redirect" title="Pseudopodium">Pseudopodium</a> <ul><li><a href="/wiki/Lamellipodium" title="Lamellipodium">Lamellipodium</a></li> <li><a href="/wiki/Filopodium" class="mw-redirect" title="Filopodium">Filopodium</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Endosymbiont" title="Endosymbiont">Endosymbionts</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mitochondrion" title="Mitochondrion">Mitochondrion</a></li> <li><a href="/wiki/Plastid" title="Plastid">Plastid</a> <ul><li><a href="/wiki/Chloroplast" title="Chloroplast">Chloroplast</a></li> <li><a href="/wiki/Chromoplast" title="Chromoplast">Chromoplast</a></li> <li><a href="/wiki/Gerontoplast" title="Gerontoplast">Gerontoplast</a></li> <li><a href="/wiki/Leucoplast" title="Leucoplast">Leucoplast</a></li> <li><a href="/wiki/Amyloplast" title="Amyloplast">Amyloplast</a></li> <li><a href="/wiki/Elaioplast" title="Elaioplast">Elaioplast</a></li> <li><a href="/wiki/Proteinoplast" title="Proteinoplast">Proteinoplast</a></li> <li><a href="/wiki/Tannosome" title="Tannosome">Tannosome</a></li> <li><a href="/wiki/Apicoplast" title="Apicoplast">Apicoplast</a></li></ul></li> <li><a href="/wiki/Nitroplast" title="Nitroplast">Nitroplast</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other internal</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nucleolus" title="Nucleolus">Nucleolus</a></li> <li><a href="/wiki/RNA" title="RNA">RNA</a> <ul><li><a href="/wiki/Ribosome" title="Ribosome">Ribosome</a></li> <li><a href="/wiki/Spliceosome" title="Spliceosome">Spliceosome</a></li> <li><a href="/wiki/Vault_(organelle)" title="Vault (organelle)">Vault</a></li></ul></li> <li><a href="/wiki/Cytoplasm" title="Cytoplasm">Cytoplasm</a> <ul><li><a href="/wiki/Cytosol" title="Cytosol">Cytosol</a></li> <li><a href="/wiki/Cytoplasmic_inclusion" class="mw-redirect" title="Cytoplasmic inclusion">Inclusions</a></li></ul></li> <li><a href="/wiki/Proteasome" title="Proteasome">Proteasome</a></li> <li><a href="/wiki/Magnetosome" title="Magnetosome">Magnetosome</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">External</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cell_wall" title="Cell wall">Cell wall</a></li> <li><a href="/wiki/Extracellular_matrix" title="Extracellular matrix">Extracellular matrix</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Branches_of_biology" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Branches_of_biology" title="Template:Branches of biology"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Branches_of_biology" title="Template talk:Branches of biology"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Branches_of_biology" title="Special:EditPage/Template:Branches of biology"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Branches_of_biology" style="font-size:114%;margin:0 4em"><a href="/wiki/Outline_of_biology#Branches" title="Outline of biology">Branches of biology</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></li> <li><a href="/wiki/Aerobiology" title="Aerobiology">Aerobiology</a></li> <li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Agrostology" title="Agrostology">Agrostology</a></li> <li><a href="/wiki/Anatomy" title="Anatomy">Anatomy</a></li> <li><a href="/wiki/Astrobiology" title="Astrobiology">Astrobiology</a></li> <li><a href="/wiki/Bacteriology" title="Bacteriology">Bacteriology</a></li> <li><a href="/wiki/Biochemistry" title="Biochemistry">Biochemistry</a></li> <li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Biogeology" title="Biogeology">Biogeology</a></li> <li><a href="/wiki/Bioinformatics" title="Bioinformatics">Bioinformatics</a></li> <li><a href="/wiki/Biological_engineering" title="Biological engineering">Biological engineering</a></li> <li><a href="/wiki/Biomechanics" title="Biomechanics">Biomechanics</a></li> <li><a href="/wiki/Biophysics" title="Biophysics">Biophysics</a></li> <li><a href="/wiki/Biosemiotics" title="Biosemiotics">Biosemiotics</a></li> <li><a href="/wiki/Biostatistics" title="Biostatistics">Biostatistics</a></li> <li><a href="/wiki/Biotechnology" title="Biotechnology">Biotechnology</a></li> <li><a href="/wiki/Botany" title="Botany">Botany</a></li> <li><a class="mw-selflink selflink">Cell biology</a></li> <li><a href="/wiki/Cellular_microbiology" title="Cellular microbiology">Cellular microbiology</a></li> <li><a href="/wiki/Chemical_biology" title="Chemical biology">Chemical biology</a></li> <li><a href="/wiki/Chronobiology" title="Chronobiology">Chronobiology</a></li> <li><a href="/wiki/Cognitive_biology" title="Cognitive biology">Cognitive biology</a></li> <li><a href="/wiki/Computational_biology" title="Computational biology">Computational biology</a></li> <li><a href="/wiki/Conservation_biology" title="Conservation biology">Conservation biology</a></li> <li><a href="/wiki/Cryobiology" title="Cryobiology">Cryobiology</a></li> <li><a href="/wiki/Cytogenetics" title="Cytogenetics">Cytogenetics</a></li> <li><a href="/wiki/Dendrology" title="Dendrology">Dendrology</a></li> <li><a href="/wiki/Developmental_biology" title="Developmental biology">Developmental biology</a></li> <li><a href="/wiki/Ecological_genetics" title="Ecological genetics">Ecological genetics</a></li> <li><a href="/wiki/Ecology" title="Ecology">Ecology</a></li> <li><a href="/wiki/Embryology" title="Embryology">Embryology</a></li> <li><a href="/wiki/Epidemiology" title="Epidemiology">Epidemiology</a></li> <li><a href="/wiki/Epigenetics" title="Epigenetics">Epigenetics</a></li> <li><a href="/wiki/Evolutionary_biology" title="Evolutionary biology">Evolutionary biology</a></li> <li><a href="/wiki/Freshwater_biology" title="Freshwater biology">Freshwater biology</a></li> <li><a href="/wiki/Generative_biology" class="mw-redirect" title="Generative biology">Generative biology</a></li> <li><a href="/wiki/Genetics" title="Genetics">Genetics</a></li> <li><a href="/wiki/Genomics" title="Genomics">Genomics</a></li> <li><a href="/wiki/Geobiology" title="Geobiology">Geobiology</a></li> <li><a href="/wiki/Gerontology" title="Gerontology">Gerontology</a></li> <li><a href="/wiki/Herpetology" title="Herpetology">Herpetology</a></li> <li><a href="/wiki/Histology" title="Histology">Histology</a></li> <li><a href="/wiki/Human_biology" title="Human biology">Human biology</a></li> <li><a href="/wiki/Ichthyology" title="Ichthyology">Ichthyology</a></li> <li><a href="/wiki/Immunology" title="Immunology">Immunology</a></li> <li><a href="/wiki/Lipidology" title="Lipidology">Lipidology</a></li> <li><a href="/wiki/Mammalogy" title="Mammalogy">Mammalogy</a></li> <li><a href="/wiki/Marine_biology" title="Marine biology">Marine biology</a></li> <li><a href="/wiki/Mathematical_and_theoretical_biology" title="Mathematical and theoretical biology">Mathematical biology</a></li> <li><a href="/wiki/Microbiology" title="Microbiology">Microbiology</a></li> <li><a href="/wiki/Molecular_biology" title="Molecular biology">Molecular biology</a></li> <li><a href="/wiki/Mycology" title="Mycology">Mycology</a></li> <li><a href="/wiki/Neontology" title="Neontology">Neontology</a></li> <li><a href="/wiki/Neuroscience" title="Neuroscience">Neuroscience</a></li> <li><a href="/wiki/Nutritional_science" title="Nutritional science">Nutrition</a></li> <li><a href="/wiki/Ornithology" title="Ornithology">Ornithology</a></li> <li><a href="/wiki/Osteology" title="Osteology">Osteology</a></li> <li><a href="/wiki/Paleontology" title="Paleontology">Paleontology</a></li> <li><a href="/wiki/Parasitology" title="Parasitology">Parasitology</a></li> <li><a href="/wiki/Pathology" title="Pathology">Pathology</a></li> <li><a href="/wiki/Pharmacology" title="Pharmacology">Pharmacology</a></li> <li><a href="/wiki/Photobiology" title="Photobiology">Photobiology</a></li> <li><a href="/wiki/Phycology" title="Phycology">Phycology</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a></li> <li><a href="/wiki/Physiology" title="Physiology">Physiology</a></li> <li><a href="/wiki/Pomology" title="Pomology">Pomology</a></li> <li><a href="/wiki/Primatology" title="Primatology">Primatology</a></li> <li><a href="/wiki/Proteomics" title="Proteomics">Proteomics</a></li> <li><a href="/wiki/Protistology" title="Protistology">Protistology</a></li> <li><a href="/wiki/Quantum_biology" title="Quantum biology">Quantum biology</a></li> <li><a href="/wiki/Relational_biology" class="mw-redirect" title="Relational biology">Relational biology</a></li> <li><a href="/wiki/Reproductive_biology" title="Reproductive biology">Reproductive biology</a></li> <li><a href="/wiki/Sociobiology" title="Sociobiology">Sociobiology</a></li> <li><a href="/wiki/Structural_biology" title="Structural biology">Structural biology</a></li> <li><a href="/wiki/Synthetic_biology" title="Synthetic biology">Synthetic biology</a></li> <li><a href="/wiki/Systematics" title="Systematics">Systematics</a></li> <li><a href="/wiki/Systems_biology" title="Systems biology">Systems biology</a></li> <li><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Taxonomy</a></li> <li><a href="/wiki/Teratology" title="Teratology">Teratology</a></li> <li><a href="/wiki/Toxicology" title="Toxicology">Toxicology</a></li> <li><a href="/wiki/Virology" title="Virology">Virology</a></li> <li><a href="/wiki/Virophysics" title="Virophysics">Virophysics</a></li> <li><a href="/wiki/Xenobiology" title="Xenobiology">Xenobiology</a></li> <li><a href="/wiki/Zoology" title="Zoology">Zoology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">See also</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/History_of_biology" title="History of biology">History of biology</a></li> <li><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></li> <li><a href="/wiki/Timeline_of_biology_and_organic_chemistry" title="Timeline of biology and organic chemistry">Timeline of biology and organic chemistry</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1066933788"></div><div role="navigation" class="navbox" aria-labelledby="Biology" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Biology_nav" title="Template:Biology nav"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Biology_nav" title="Template talk:Biology nav"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Biology_nav" title="Special:EditPage/Template:Biology nav"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Biology" style="font-size:114%;margin:0 4em"><a href="/wiki/Biology" title="Biology">Biology</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li>Introduction (<a href="/wiki/Introduction_to_genetics" title="Introduction to genetics">Genetics</a>, <a href="/wiki/Introduction_to_evolution" title="Introduction to evolution">Evolution</a>)</li> <li><a href="/wiki/Outline_of_biology" title="Outline of biology">Outline</a></li> <li><a href="/wiki/History_of_biology" title="History of biology">History</a> <ul><li><a href="/wiki/Timeline_of_biology_and_organic_chemistry" title="Timeline of biology and organic chemistry">Timeline</a></li></ul></li> <li><a href="/wiki/Index_of_biology_articles" title="Index of biology articles">Index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Biology" title="Biology">Biology</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Overview</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Science" title="Science">Science</a></li> <li><a href="/wiki/Life" title="Life">Life</a></li> <li>Properties (<a href="/wiki/Adaptation" title="Adaptation">Adaptation</a>, <a href="/wiki/Metabolism" title="Metabolism"> Energy processing</a>, <a href="/wiki/Developmental_biology" title="Developmental biology">Growth</a>, <a href="/wiki/Structure#Biological" title="Structure">Order</a>, <a href="/wiki/Homeostasis" title="Homeostasis">Regulation</a>, <a href="/wiki/Reproduction" title="Reproduction">Reproduction</a> (<a href="/wiki/Self-replication" title="Self-replication">Self-replication</a>), <a href="/wiki/Stimulus_(physiology)" title="Stimulus (physiology)">Response to environment</a>)</li> <li><a href="/wiki/Biological_organisation" title="Biological organisation">Hierarchy of life</a> (<a href="/wiki/Atom" title="Atom">Atom</a> > <a href="/wiki/Molecule" title="Molecule">Molecule</a> > <a href="/wiki/Organelle" title="Organelle">Organelle</a> > <a href="/wiki/Cell_(biology)" title="Cell (biology)">Cell</a> > <a href="/wiki/Tissue_(biology)" title="Tissue (biology)">Tissue</a> > <a href="/wiki/Organ_(biology)" title="Organ (biology)">Organ</a> > <a href="/wiki/Biological_system" title="Biological system">Organ system</a> > <a href="/wiki/Organism" title="Organism">Organism</a> > <a href="/wiki/Population" title="Population">Population</a> > <a href="/wiki/Biocoenosis" title="Biocoenosis">Community</a> > <a href="/wiki/Ecosystem" title="Ecosystem">Ecosystem</a> > <a href="/wiki/Biosphere" title="Biosphere">Biosphere</a>)</li> <li><a href="/wiki/Reductionism#In_science" title="Reductionism">Reductionistic</a></li> <li><a href="/wiki/Emergence" title="Emergence"> Emergent property</a></li> <li><a href="/wiki/Mechanical_philosophy" class="mw-redirect" title="Mechanical philosophy">Mechanistic</a></li> <li><a href="/wiki/Scientific_method" title="Scientific method">Scientific method</a></li> <li><a href="/wiki/Taxonomic_rank" title="Taxonomic rank">Taxonomic rank</a></li> <li><a href="/wiki/Scientific_theory" title="Scientific theory"> Theory</a></li> <li><a href="/wiki/Scientific_law" title="Scientific law"> Law</a></li> <li><a href="/wiki/Peer_review" title="Peer review">Peer review</a></li> <li><a href="/wiki/List_of_biology_journals" title="List of biology journals">Biology journals</a></li> <li><a href="/wiki/Common_name" title="Common name">Common name</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Chemical basis</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Atom" title="Atom">Atoms</a></li> <li><a href="/wiki/Amino_acid" title="Amino acid">Amino acids</a></li> <li><a href="/wiki/Carbohydrate" title="Carbohydrate">Carbohydrates</a></li> <li><a href="/wiki/Chemical_bond" title="Chemical bond">Chemical bond</a></li> <li><a href="/wiki/Chemical_element" title="Chemical element">Chemical element</a></li> <li><a href="/wiki/Lipid" title="Lipid">Lipids</a></li> <li><a href="/wiki/Matter" title="Matter">Matter</a> <ul><li><a href="/wiki/Quantum_biology" title="Quantum biology">Quantum</a></li></ul></li> <li><a href="/wiki/Molecule" title="Molecule">Molecules</a></li> <li><a href="/wiki/Monomer" title="Monomer">Monomer</a></li> <li><a href="/wiki/Nucleic_acid" title="Nucleic acid">Nucleic acids</a></li> <li><a href="/wiki/Organic_compound" title="Organic compound">Organic compounds</a></li> <li><a href="/wiki/PH" title="PH">pH</a></li> <li><a href="/wiki/Polymer" title="Polymer">Polymer</a></li> <li><a href="/wiki/Protein" title="Protein">Proteins</a></li> <li><a href="/wiki/Water" title="Water">Water</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cell_(biology)" title="Cell (biology)">Cells</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Adenosine_triphosphate" title="Adenosine triphosphate">ATP</a></li> <li><a href="/wiki/Cell_cycle" title="Cell cycle">Cell cycle</a></li> <li><a href="/wiki/Cell_theory" title="Cell theory">Cell theory</a></li> <li><a href="/wiki/Cell_signaling" title="Cell signaling">Cell signaling</a></li> <li><a href="/wiki/Cellular_respiration" title="Cellular respiration">Cellular respiration</a></li> <li><a href="/wiki/Energy" title="Energy">Energy transformation</a></li> <li><a href="/wiki/Enzyme" title="Enzyme">Enzyme</a></li> <li><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryote</a></li> <li><a href="/wiki/Fermentation" title="Fermentation">Fermentation</a></li> <li><a href="/wiki/Metabolism" title="Metabolism">Metabolism</a></li> <li><a href="/wiki/Meiosis" title="Meiosis">Meiosis</a></li> <li><a href="/wiki/Mitosis" title="Mitosis">Mitosis</a></li> <li><a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a></li> <li><a href="/wiki/Prokaryote" title="Prokaryote">Prokaryote</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Genetics" title="Genetics">Genetics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/DNA" title="DNA">DNA</a></li> <li><a href="/wiki/Epigenetics" title="Epigenetics">Epigenetics</a></li> <li><a href="/wiki/Evolutionary_developmental_biology" title="Evolutionary developmental biology">Evolutionary developmental biology</a></li> <li><a href="/wiki/Gene_expression" title="Gene expression">Gene expression</a></li> <li><a href="/wiki/Regulation_of_gene_expression" title="Regulation of gene expression">Gene regulation</a></li> <li><a href="/wiki/Genome" title="Genome">Genomes</a></li> <li><a href="/wiki/Mendelian_inheritance" title="Mendelian inheritance">Mendelian inheritance</a></li> <li><a href="/wiki/Post-transcriptional_modification" title="Post-transcriptional modification">Post-transcriptional modification</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Evolution" title="Evolution">Evolution</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Adaptation" title="Adaptation">Adaptation</a></li> <li><a href="/wiki/Earliest_known_life_forms" title="Earliest known life forms">Earliest known life forms</a></li> <li><a href="/wiki/Function_(biology)" title="Function (biology)">Function</a></li> <li><a href="/wiki/Genetic_drift" title="Genetic drift">Genetic drift</a></li> <li><a href="/wiki/Gene_flow" title="Gene flow">Gene flow</a></li> <li><a href="/wiki/History_of_life" title="History of life">History of life</a></li> <li><a href="/wiki/Macroevolution" title="Macroevolution">Macroevolution</a></li> <li><a href="/wiki/Microevolution" title="Microevolution">Microevolution</a></li> <li><a href="/wiki/Mutation" title="Mutation">Mutation</a></li> <li><a href="/wiki/Natural_selection" title="Natural selection">Natural selection</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a></li> <li><a href="/wiki/Speciation" title="Speciation">Speciation</a></li> <li><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Taxonomy</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Biodiversity" title="Biodiversity">Diversity</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Bacteria" title="Bacteria">Bacteria</a></li> <li><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryote</a> <ul><li><a href="/wiki/Alga" class="mw-redirect" title="Alga">Alga</a></li> <li><a href="/wiki/Animal" title="Animal">Animal</a></li> <li><a href="/wiki/Fungus" title="Fungus">Fungus</a></li> <li><a href="/wiki/Plant" title="Plant">Plant</a></li> <li><a href="/wiki/Protist" title="Protist">Protist</a></li></ul></li> <li><a href="/wiki/Virus" title="Virus">Virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Plant form <br /> and function</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Epidermis_(botany)" title="Epidermis (botany)">Epidermis (botany)</a></li> <li><a href="/wiki/Flower" title="Flower">Flower</a></li> <li><a href="/wiki/Ground_tissue" title="Ground tissue">Ground tissue</a></li> <li><a href="/wiki/Leaf" title="Leaf">Leaf</a></li> <li><a href="/wiki/Phloem" title="Phloem">Phloem</a></li> <li><a href="/wiki/Plant_stem" title="Plant stem">Plant stem</a></li> <li><a href="/wiki/Root" title="Root">Root</a></li> <li><a href="/wiki/Shoot_(botany)" title="Shoot (botany)">Shoot</a></li> <li><a href="/wiki/Vascular_plant" title="Vascular plant">Vascular plant</a></li> <li><a href="/wiki/Vascular_tissue" title="Vascular tissue">Vascular tissue</a></li> <li><a href="/wiki/Xylem" title="Xylem">Xylem</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Animal form <br />and function</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Breathing" title="Breathing">Breathing</a></li> <li><a href="/wiki/Circulatory_system" title="Circulatory system">Circulatory system</a></li> <li><a href="/wiki/Endocrine_system" title="Endocrine system">Endocrine system</a></li> <li><a href="/wiki/Human_digestive_system" title="Human digestive system">Digestive system</a></li> <li><a href="/wiki/Homeostasis" title="Homeostasis">Homeostasis</a></li> <li><a href="/wiki/Immune_system" title="Immune system">Immune system</a></li> <li><a href="/wiki/Internal_environment" title="Internal environment">Internal environment</a></li> <li><a href="/wiki/Muscular_system" title="Muscular system">Muscular system</a></li> <li><a href="/wiki/Nervous_system" title="Nervous system">Nervous system</a></li> <li><a href="/wiki/Reproductive_system" title="Reproductive system">Reproductive system</a></li> <li><a href="/wiki/Respiratory_system" title="Respiratory system">Respiratory system</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ecology" title="Ecology">Ecology</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biogeochemical_cycle" title="Biogeochemical cycle">Biogeochemical cycle</a></li> <li><a href="/wiki/Biological_interaction" title="Biological interaction">Biological interaction</a></li> <li><a href="/wiki/Biomass" title="Biomass">Biomass</a></li> <li><a href="/wiki/Biome" title="Biome">Biomes</a></li> <li><a href="/wiki/Biosphere" title="Biosphere">Biosphere</a></li> <li><a href="/wiki/Climate" title="Climate">Climate</a></li> <li><a href="/wiki/Climate_change" title="Climate change">Climate change</a></li> <li><a href="/wiki/Community_(ecology)" title="Community (ecology)">Community</a></li> <li><a href="/wiki/Conservation_biology" title="Conservation biology">Conservation</a></li> <li><a href="/wiki/Ecosystem_ecology" title="Ecosystem ecology">Ecosystem</a></li> <li><a href="/wiki/Habitat" title="Habitat">Habitat</a> <ul><li><a href="/wiki/Ecological_niche" title="Ecological niche">niche</a></li></ul></li> <li><a href="/wiki/Microbiome" title="Microbiome">Microbiome</a></li> <li><a href="/wiki/Population_ecology" title="Population ecology">Population dynamics</a></li> <li><a href="/wiki/Resource_(biology)" title="Resource (biology)">Resources</a></li></ul> </div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="4" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Tree_of_life_by_Haeckel.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/de/Tree_of_life_by_Haeckel.jpg/140px-Tree_of_life_by_Haeckel.jpg" decoding="async" width="140" height="222" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/de/Tree_of_life_by_Haeckel.jpg/210px-Tree_of_life_by_Haeckel.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/de/Tree_of_life_by_Haeckel.jpg/280px-Tree_of_life_by_Haeckel.jpg 2x" data-file-width="1804" data-file-height="2856" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/List_of_research_methods_in_biology" title="List of research methods in biology">Research <br /> methods</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Laboratory <br /> techniques</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Genetic_engineering" title="Genetic engineering">Genetic engineering</a></li> <li><a href="/wiki/Transformation_(genetics)" class="mw-redirect" title="Transformation (genetics)">Transformation</a></li> <li><a href="/wiki/Gel_electrophoresis" title="Gel electrophoresis">Gel electrophoresis</a></li> <li><a href="/wiki/Chromatography" title="Chromatography">Chromatography</a></li> <li><a href="/wiki/Centrifugation" title="Centrifugation">Centrifugation</a></li> <li><a href="/wiki/Cell_culture" title="Cell culture">Cell culture</a></li> <li><a href="/wiki/DNA_sequencing" title="DNA sequencing">DNA sequencing</a></li> <li><a href="/wiki/DNA_microarray" title="DNA microarray">DNA microarray</a></li> <li><a href="/wiki/Green_fluorescent_protein" title="Green fluorescent protein">Green fluorescent protein</a></li> <li><a href="/wiki/Vector_(molecular_biology)" title="Vector (molecular biology)">vector</a></li> <li><a href="/wiki/Enzyme_assay" title="Enzyme assay">Enzyme assay</a></li> <li><a href="/wiki/Protein_purification" title="Protein purification">Protein purification</a></li> <li><a href="/wiki/Western_blot" title="Western blot">Western blot</a></li> <li><a href="/wiki/Northern_blot" title="Northern blot">Northern blot</a></li> <li><a href="/wiki/Southern_blot" title="Southern blot">Southern blot</a></li> <li><a href="/wiki/Restriction_enzyme" title="Restriction enzyme">Restriction enzyme</a></li> <li><a href="/wiki/Polymerase_chain_reaction" title="Polymerase chain reaction">Polymerase chain reaction</a></li> <li><a href="/wiki/Two-hybrid_screening" title="Two-hybrid screening">Two-hybrid screening</a></li> <li><i><a href="/wiki/In_vivo" title="In vivo">in vivo</a></i></li> <li><i><a href="/wiki/In_vitro" title="In vitro">in vitro</a></i></li> <li><i><a href="/wiki/In_silico" title="In silico">in silico</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Field techniques</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Belt_transect" title="Belt transect">Belt transect</a></li> <li><a href="/wiki/Mark_and_recapture" title="Mark and recapture">mark and recapture</a></li> <li><a href="/wiki/Species_discovery_curve" title="Species discovery curve">species discovery curve</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Branches</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><div class="excerpt-block"><div class="excerpt"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></li> <li><a href="/wiki/Aerobiology" title="Aerobiology">Aerobiology</a></li> <li><a href="/wiki/Agronomy" title="Agronomy">Agronomy</a></li> <li><a href="/wiki/Agrostology" title="Agrostology">Agrostology</a></li> <li><a href="/wiki/Anatomy" title="Anatomy">Anatomy</a></li> <li><a href="/wiki/Astrobiology" title="Astrobiology">Astrobiology</a></li> <li><a href="/wiki/Bacteriology" title="Bacteriology">Bacteriology</a></li> <li><a href="/wiki/Biochemistry" title="Biochemistry">Biochemistry</a></li> <li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Biogeology" title="Biogeology">Biogeology</a></li> <li><a href="/wiki/Bioinformatics" title="Bioinformatics">Bioinformatics</a></li> <li><a href="/wiki/Biological_engineering" title="Biological engineering">Biological engineering</a></li> <li><a href="/wiki/Biomechanics" title="Biomechanics">Biomechanics</a></li> <li><a href="/wiki/Biophysics" title="Biophysics">Biophysics</a></li> <li><a href="/wiki/Biosemiotics" title="Biosemiotics">Biosemiotics</a></li> <li><a href="/wiki/Biostatistics" title="Biostatistics">Biostatistics</a></li> <li><a href="/wiki/Biotechnology" title="Biotechnology">Biotechnology</a></li> <li><a href="/wiki/Botany" title="Botany">Botany</a></li> <li>Cell biology</li> <li><a href="/wiki/Cellular_microbiology" title="Cellular microbiology">Cellular microbiology</a></li> <li><a href="/wiki/Chemical_biology" title="Chemical biology">Chemical biology</a></li> <li><a href="/wiki/Chronobiology" title="Chronobiology">Chronobiology</a></li> <li><a href="/wiki/Cognitive_biology" title="Cognitive biology">Cognitive biology</a></li> <li><a href="/wiki/Computational_biology" title="Computational biology">Computational biology</a></li> <li><a href="/wiki/Conservation_biology" title="Conservation biology">Conservation biology</a></li> <li><a href="/wiki/Cryobiology" title="Cryobiology">Cryobiology</a></li> <li><a href="/wiki/Cytogenetics" title="Cytogenetics">Cytogenetics</a></li> <li><a href="/wiki/Dendrology" title="Dendrology">Dendrology</a></li> <li><a href="/wiki/Developmental_biology" title="Developmental biology">Developmental biology</a></li> <li><a href="/wiki/Ecological_genetics" title="Ecological genetics">Ecological genetics</a></li> <li><a href="/wiki/Ecology" title="Ecology">Ecology</a></li> <li><a href="/wiki/Embryology" title="Embryology">Embryology</a></li> <li><a href="/wiki/Epidemiology" title="Epidemiology">Epidemiology</a></li> <li><a href="/wiki/Epigenetics" title="Epigenetics">Epigenetics</a></li> <li><a href="/wiki/Evolutionary_biology" title="Evolutionary biology">Evolutionary biology</a></li> <li><a href="/wiki/Freshwater_biology" title="Freshwater biology">Freshwater biology</a></li> <li><a href="/wiki/Generative_biology" class="mw-redirect" title="Generative biology">Generative biology</a></li> <li><a href="/wiki/Genetics" title="Genetics">Genetics</a></li> <li><a href="/wiki/Genomics" title="Genomics">Genomics</a></li> <li><a href="/wiki/Geobiology" title="Geobiology">Geobiology</a></li> <li><a href="/wiki/Gerontology" title="Gerontology">Gerontology</a></li> <li><a href="/wiki/Herpetology" title="Herpetology">Herpetology</a></li> <li><a href="/wiki/Histology" title="Histology">Histology</a></li> <li><a href="/wiki/Human_biology" title="Human biology">Human biology</a></li> <li><a href="/wiki/Ichthyology" title="Ichthyology">Ichthyology</a></li> <li><a href="/wiki/Immunology" title="Immunology">Immunology</a></li> <li><a href="/wiki/Lipidology" title="Lipidology">Lipidology</a></li> <li><a href="/wiki/Mammalogy" title="Mammalogy">Mammalogy</a></li> <li><a href="/wiki/Marine_biology" title="Marine biology">Marine biology</a></li> <li><a href="/wiki/Mathematical_and_theoretical_biology" title="Mathematical and theoretical biology">Mathematical biology</a></li> <li><a href="/wiki/Microbiology" title="Microbiology">Microbiology</a></li> <li><a href="/wiki/Molecular_biology" title="Molecular biology">Molecular biology</a></li> <li><a href="/wiki/Mycology" title="Mycology">Mycology</a></li> <li><a href="/wiki/Neontology" title="Neontology">Neontology</a></li> <li><a href="/wiki/Neuroscience" title="Neuroscience">Neuroscience</a></li> <li><a href="/wiki/Nutritional_science" title="Nutritional science">Nutrition</a></li> <li><a href="/wiki/Ornithology" title="Ornithology">Ornithology</a></li> <li><a href="/wiki/Osteology" title="Osteology">Osteology</a></li> <li><a href="/wiki/Paleontology" title="Paleontology">Paleontology</a></li> <li><a href="/wiki/Parasitology" title="Parasitology">Parasitology</a></li> <li><a href="/wiki/Pathology" title="Pathology">Pathology</a></li> <li><a href="/wiki/Pharmacology" title="Pharmacology">Pharmacology</a></li> <li><a href="/wiki/Photobiology" title="Photobiology">Photobiology</a></li> <li><a href="/wiki/Phycology" title="Phycology">Phycology</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a></li> <li><a href="/wiki/Physiology" title="Physiology">Physiology</a></li> <li><a href="/wiki/Pomology" title="Pomology">Pomology</a></li> <li><a href="/wiki/Primatology" title="Primatology">Primatology</a></li> <li><a href="/wiki/Proteomics" title="Proteomics">Proteomics</a></li> <li><a href="/wiki/Protistology" title="Protistology">Protistology</a></li> <li><a href="/wiki/Quantum_biology" title="Quantum biology">Quantum biology</a></li> <li><a href="/wiki/Relational_biology" class="mw-redirect" title="Relational biology">Relational biology</a></li> <li><a href="/wiki/Reproductive_biology" title="Reproductive biology">Reproductive biology</a></li> <li><a href="/wiki/Sociobiology" title="Sociobiology">Sociobiology</a></li> <li><a href="/wiki/Structural_biology" title="Structural biology">Structural biology</a></li> <li><a href="/wiki/Synthetic_biology" title="Synthetic biology">Synthetic biology</a></li> <li><a href="/wiki/Systematics" title="Systematics">Systematics</a></li> <li><a href="/wiki/Systems_biology" title="Systems biology">Systems biology</a></li> <li><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Taxonomy</a></li> <li><a href="/wiki/Teratology" title="Teratology">Teratology</a></li> <li><a href="/wiki/Toxicology" title="Toxicology">Toxicology</a></li> <li><a href="/wiki/Virology" title="Virology">Virology</a></li> <li><a href="/wiki/Virophysics" title="Virophysics">Virophysics</a></li> <li><a href="/wiki/Xenobiology" title="Xenobiology">Xenobiology</a></li> <li><a href="/wiki/Zoology" title="Zoology">Zoology</a></li></ul></div></div> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Glossaries</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glossary_of_biology" title="Glossary of biology">Biology</a></li> <li><a href="/wiki/Glossary_of_botanical_terms" title="Glossary of botanical terms">Botanical terms</a></li> <li><a href="/wiki/Glossary_of_ecology" title="Glossary of ecology">Ecological terms</a></li> <li><a href="/wiki/Glossary_of_plant_morphology" title="Glossary of plant morphology">Plant morphology terms</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li><span class="nowrap"><span class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Issoria_lathonia.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/16px-Issoria_lathonia.jpg" decoding="async" width="16" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/24px-Issoria_lathonia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/32px-Issoria_lathonia.jpg 2x" data-file-width="629" data-file-height="445" /></a></span> </span><a href="/wiki/Portal:Biology" title="Portal:Biology">Biology portal</a></span></li> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" 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style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Branches_of_chemistry" title="Template:Branches of chemistry"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Branches_of_chemistry" title="Template talk:Branches of chemistry"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Branches_of_chemistry" title="Special:EditPage/Template:Branches of chemistry"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Branches_of_chemistry" style="font-size:114%;margin:0 4em">Branches of <a href="/wiki/Chemistry" title="Chemistry">chemistry</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><a href="/wiki/Glossary_of_chemical_formulae" title="Glossary of chemical formulae">Glossary of chemical formulae</a></li> <li><a href="/wiki/List_of_biomolecules" title="List of biomolecules">List of biomolecules</a></li> <li><a href="/wiki/List_of_inorganic_compounds" title="List of inorganic compounds">List of inorganic compounds</a></li> <li><a href="/wiki/Periodic_table" title="Periodic table">Periodic table</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Analytical_chemistry" title="Analytical chemistry">Analytical</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Instrumental_chemistry" title="Instrumental chemistry">Instrumental chemistry</a></li> 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class="navbox-group" style="width:1%"><a href="/wiki/Theoretical_chemistry" title="Theoretical chemistry">Theoretical</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Quantum_chemistry" title="Quantum chemistry">Quantum chemistry</a></li> <li><a href="/wiki/Computational_chemistry" title="Computational chemistry">Computational chemistry</a> <ul><li><a href="/wiki/Mathematical_chemistry" title="Mathematical chemistry">Mathematical chemistry</a></li></ul></li> <li><a href="/wiki/Molecular_modelling" title="Molecular modelling">Molecular modelling</a></li> <li><a href="/wiki/Molecular_mechanics" title="Molecular mechanics">Molecular mechanics</a></li> <li><a href="/wiki/Molecular_dynamics" title="Molecular dynamics">Molecular dynamics</a></li> <li><a href="/wiki/Molecular_geometry" title="Molecular geometry">Molecular geometry</a> <ul><li><a href="/wiki/VSEPR_theory" title="VSEPR theory">VSEPR theory</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Physical_chemistry" title="Physical chemistry">Physical</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Electrochemistry" title="Electrochemistry">Electrochemistry</a> <ul><li><a href="/wiki/Spectroelectrochemistry" title="Spectroelectrochemistry">Spectroelectrochemistry</a></li> <li><a href="/wiki/Photoelectrochemistry" title="Photoelectrochemistry">Photoelectrochemistry</a></li></ul></li> <li><a href="/wiki/Thermochemistry" title="Thermochemistry">Thermochemistry</a></li> <li><a href="/wiki/Chemical_thermodynamics" title="Chemical thermodynamics">Chemical thermodynamics</a></li> <li><a href="/wiki/Surface_science" title="Surface science">Surface science</a></li> <li><a href="/wiki/Interface_and_colloid_science" title="Interface and colloid 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<li><a href="/wiki/Atom_cluster" class="mw-redirect" title="Atom cluster">Cluster chemistry</a></li> <li><a href="/wiki/Solid-state_chemistry" title="Solid-state chemistry">Solid-state chemistry</a></li> <li><a href="/wiki/Ceramic_chemistry" class="mw-redirect" title="Ceramic chemistry">Ceramic chemistry</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Organic_chemistry" title="Organic chemistry">Organic</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Stereochemistry" title="Stereochemistry">Stereochemistry</a> <ul><li><a href="/wiki/Alkane_stereochemistry" class="mw-redirect" title="Alkane stereochemistry">Alkane stereochemistry</a></li></ul></li> <li><a href="/wiki/Physical_organic_chemistry" title="Physical organic chemistry">Physical organic chemistry</a></li> <li><a href="/wiki/Organic_reactions" class="mw-redirect" title="Organic reactions">Organic reactions</a></li> <li><a href="/wiki/Organic_synthesis" title="Organic synthesis">Organic synthesis</a></li> <li><a href="/wiki/Retrosynthetic_analysis" title="Retrosynthetic analysis">Retrosynthetic analysis</a></li> <li><a href="/wiki/Enantioselective_synthesis" title="Enantioselective synthesis">Enantioselective synthesis</a></li> <li><a href="/wiki/Total_synthesis" title="Total synthesis">Total synthesis</a> / <a href="/wiki/Semisynthesis" title="Semisynthesis">Semisynthesis</a></li> <li><a href="/wiki/Fullerene_chemistry" title="Fullerene chemistry">Fullerene chemistry</a></li> <li><a href="/wiki/Polymer_chemistry" title="Polymer chemistry">Polymer chemistry</a></li> <li><a href="/wiki/Petrochemistry" class="mw-redirect" title="Petrochemistry">Petrochemistry</a></li> <li><a href="/wiki/Dynamic_covalent_chemistry" title="Dynamic covalent chemistry">Dynamic covalent chemistry</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Biochemistry" title="Biochemistry">Biological</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biochemistry" title="Biochemistry">Biochemistry</a> <ul><li><a href="/wiki/Molecular_biology" title="Molecular biology">Molecular biology</a></li> <li><a class="mw-selflink selflink">Cell biology</a></li></ul></li> <li><a href="/wiki/Chemical_biology" title="Chemical biology">Chemical biology</a> <ul><li><a href="/wiki/Bioorthogonal_chemistry" title="Bioorthogonal chemistry">Bioorthogonal chemistry</a></li></ul></li> <li><a href="/wiki/Medicinal_chemistry" title="Medicinal chemistry">Medicinal chemistry</a> <ul><li><a href="/wiki/Pharmacology" title="Pharmacology">Pharmacology</a></li></ul></li> <li><a href="/wiki/Clinical_chemistry" title="Clinical chemistry">Clinical chemistry</a></li> <li><a href="/wiki/Neurochemistry" title="Neurochemistry">Neurochemistry</a></li> <li><a href="/wiki/Bioorganic_chemistry" title="Bioorganic chemistry">Bioorganic chemistry</a></li> <li><a href="/wiki/Bioorganometallic_chemistry" title="Bioorganometallic chemistry">Bioorganometallic chemistry</a></li> <li><a href="/wiki/Bioinorganic_chemistry" title="Bioinorganic chemistry">Bioinorganic chemistry</a></li> <li><a href="/wiki/Biophysical_chemistry" title="Biophysical chemistry">Biophysical chemistry</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Interdisciplinarity" title="Interdisciplinarity">Interdisciplinarity</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nuclear_chemistry" title="Nuclear chemistry">Nuclear chemistry</a> <ul><li><a href="/wiki/Radiochemistry" title="Radiochemistry">Radiochemistry</a></li> <li><a href="/wiki/Radiation_chemistry" title="Radiation chemistry">Radiation chemistry</a></li> <li><a href="/wiki/Actinide_chemistry" title="Actinide chemistry">Actinide chemistry</a></li></ul></li> <li><a href="/wiki/Cosmochemistry" title="Cosmochemistry">Cosmochemistry</a> / <a href="/wiki/Astrochemistry" title="Astrochemistry">Astrochemistry</a> / <a href="/wiki/Stellar_chemistry" title="Stellar chemistry">Stellar chemistry</a></li> <li><a href="/wiki/Geochemistry" title="Geochemistry">Geochemistry</a> <ul><li><a href="/wiki/Biogeochemistry" title="Biogeochemistry">Biogeochemistry</a></li> <li><a href="/wiki/Photogeochemistry" title="Photogeochemistry">Photogeochemistry</a></li></ul></li></ul> <ul><li><a href="/wiki/Environmental_chemistry" title="Environmental chemistry">Environmental chemistry</a> <ul><li><a href="/wiki/Atmospheric_chemistry" title="Atmospheric chemistry">Atmospheric chemistry</a></li> <li><a href="/wiki/Ocean_chemistry" class="mw-redirect" title="Ocean chemistry">Ocean chemistry</a></li></ul></li> <li><a href="/wiki/Clay_chemistry" title="Clay chemistry">Clay chemistry</a></li> <li><a href="/wiki/Carbochemistry" title="Carbochemistry">Carbochemistry</a></li> <li><a href="/wiki/Food_chemistry" title="Food chemistry">Food chemistry</a> <ul><li><a href="/wiki/Carbohydrate_chemistry" class="mw-redirect" title="Carbohydrate chemistry">Carbohydrate chemistry</a></li> <li><a href="/wiki/Food_physical_chemistry" title="Food physical chemistry">Food physical chemistry</a></li></ul></li> <li><a href="/wiki/Agricultural_chemistry" title="Agricultural chemistry">Agricultural chemistry</a> <ul><li><a href="/wiki/Soil_chemistry" title="Soil chemistry">Soil chemistry</a></li></ul></li></ul> <ul><li><a href="/wiki/Chemistry_education" title="Chemistry education">Chemistry education</a> <ul><li><a href="/wiki/Amateur_chemistry" title="Amateur chemistry">Amateur chemistry</a></li> <li><a href="/wiki/General_chemistry" title="General chemistry">General chemistry</a></li></ul></li> <li><a href="/wiki/Clandestine_chemistry" title="Clandestine chemistry">Clandestine chemistry</a></li> <li><a href="/wiki/Forensic_chemistry" title="Forensic chemistry">Forensic chemistry</a> <ul><li><a href="/wiki/Forensic_toxicology" title="Forensic toxicology">Forensic toxicology</a></li> <li><a href="/wiki/Post-mortem_chemistry" title="Post-mortem chemistry">Post-mortem chemistry</a></li></ul></li></ul> <ul><li><a href="/wiki/Nanochemistry" title="Nanochemistry">Nanochemistry</a> <ul><li><a href="/wiki/Supramolecular_chemistry" title="Supramolecular chemistry">Supramolecular chemistry</a></li></ul></li> <li><a href="/wiki/Chemical_synthesis" title="Chemical synthesis">Chemical synthesis</a> <ul><li><a href="/wiki/Green_chemistry" title="Green chemistry">Green chemistry</a></li> <li><a href="/wiki/Click_chemistry" title="Click chemistry">Click chemistry</a></li> <li><a href="/wiki/Combinatorial_chemistry" title="Combinatorial chemistry">Combinatorial chemistry</a></li> <li><a href="/wiki/Biosynthesis" title="Biosynthesis">Biosynthesis</a></li></ul></li> <li><a href="/wiki/Chemical_engineering" title="Chemical engineering">Chemical engineering</a> <ul><li><a href="/wiki/Stoichiometry" title="Stoichiometry">Stoichiometry</a></li></ul></li> <li><a href="/wiki/Materials_science" title="Materials science">Materials science</a> <ul><li><a href="/wiki/Metallurgy" title="Metallurgy">Metallurgy</a></li> <li><a href="/wiki/Ceramic_engineering" title="Ceramic engineering">Ceramic engineering</a></li> <li><a href="/wiki/Polymer_science" title="Polymer science">Polymer science</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">See also</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/History_of_chemistry" title="History of chemistry">History of chemistry</a></li> <li><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Nobel Prize in Chemistry</a></li> <li><a href="/wiki/Timeline_of_chemistry" title="Timeline of chemistry">Timeline of chemistry</a> <ul><li><a href="/wiki/Discovery_of_chemical_elements" title="Discovery of chemical elements">of element discoveries</a></li></ul></li> <li>"<a href="/wiki/The_central_science" title="The central science">The central science</a>"</li> <li><a href="/wiki/Chemical_reaction" title="Chemical reaction">Chemical reaction</a> <ul><li><a href="/wiki/Catalysis" title="Catalysis">Catalysis</a></li></ul></li> <li><a href="/wiki/Chemical_element" title="Chemical element">Chemical element</a></li> <li><a href="/wiki/Chemical_compound" title="Chemical compound">Chemical compound</a></li> <li><a href="/wiki/Atom" title="Atom">Atom</a></li> <li><a href="/wiki/Molecule" title="Molecule">Molecule</a></li> <li><a href="/wiki/Ion" title="Ion">Ion</a></li> <li><a href="/wiki/Chemical_substance" title="Chemical substance">Chemical substance</a></li> <li><a href="/wiki/Chemical_bond" title="Chemical bond">Chemical bond</a></li> <li><a href="/wiki/Alchemy" title="Alchemy">Alchemy</a></li> <li><a href="/wiki/Quantum_mechanics" title="Quantum mechanics">Quantum mechanics</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <b><a href="/wiki/Category:Chemistry" title="Category:Chemistry">Category</a></b></li> <li><span class="noviewer" typeof="mw:File"><span title="Commons page"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span> <b><a href="https://commons.wikimedia.org/wiki/Category:Chemistry" class="extiw" title="commons:Category:Chemistry">Commons</a></b></li> <li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" class="mw-file-description" title="Portal"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/16px-Symbol_portal_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" 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