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Lion | Characteristics, Habitat, & Facts | Britannica
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id="content" class="md-content"> <div class="md-article-container template-desktop infinite-pagination"> <div class="infinite-scroll-container article last"> <article class="article-content container-lg qa-content px-0 pt-0 pb-40 py-lg-20 content md-expanded" data-topic-id="342664"> <div class="grid gx-0"> <div class="col-auto"> <div class="topic-left-rail md-article-drawer position-relative d-flex border-right-sm border-left-sm open"> <div class="drawer d-flex flex-column open"> <div class="left-rail-section-content"> <div class="topic-left-rail-header text-truncate bg-gray-50 position-relative text-right d-flex align-items-center"> <div class="tlr-title px-20 py-15 text-left"> <em class="material-icons text-gray-400 d-lg-none" data-icon="toc"></em> <a class="font-serif font-weight-bold text-black link-blue" href="https://www.britannica.com/animal/lion">lion</a> </div> <button aria-label="Close" class="js-sections-close-button btn-link btn-sm btn d-lg-none position-absolute top-0 p-10 right-0" > <em class="material-icons font-26" data-icon="close"></em> </button> </div> <div class="section-content pl-10 pr-20 pl-sm-50 pr-sm-60 pl-lg-5 pr-lg-10 pt-10 pt-lg-0 bg-gray-50 clear-catfish-ad"> <div class="toc mb-20"> <div class="font-serif font-14 font-weight-bold mx-15 mb-15 mt-20"> Table of Contents </div> <ul class="list-unstyled my-0" data-level="h1"><li data-target="#ref1"><div class="pl-25"><a class="link-gray-900 w-100" href="/animal/lion">Introduction & Top Questions</a></div><div class="ml-40 toc-drawer sub-toc-drawer"></div></li><li data-target="#ref325410"><div class="d-flex align-items-center"><div class="ml-25"></div><a class="w-100 link-gray-900" href="/animal/lion#ref325410">General characteristics</a></div><div class="ml-40 toc-drawer sub-toc-drawer"></div></li><li data-target="#ref325411"><div class="d-flex align-items-center"><div class="ml-25"></div><a class="w-100 link-gray-900" href="/animal/lion#ref325411">Prides</a></div><div class="ml-40 toc-drawer sub-toc-drawer"></div></li><li data-target="#ref325412"><div class="d-flex align-items-center"><div class="ml-25"></div><a class="w-100 link-gray-900" href="/animal/lion#ref325412">Hunting</a></div><div class="ml-40 toc-drawer sub-toc-drawer"></div></li><li data-target="#ref325413"><div class="d-flex align-items-center"><div class="ml-25"></div><a class="w-100 link-gray-900" href="/animal/lion/Reproduction-and-life-cycle">Reproduction and life cycle</a></div><div class="ml-40 toc-drawer sub-toc-drawer"></div></li><li data-target="#ref325414"><div class="d-flex align-items-center"><div class="ml-25"></div><a class="w-100 link-gray-900" href="/animal/lion/Reproduction-and-life-cycle#ref325414">Classification and distribution</a></div><div class="ml-40 toc-drawer sub-toc-drawer"></div></li><li data-target="#ref378991"><div class="d-flex align-items-center"><div class="ml-25"></div><a class="w-100 link-gray-900" href="/animal/lion/Reproduction-and-life-cycle#ref378991">Conservation status</a></div><div class="ml-40 toc-drawer sub-toc-drawer"></div></li></ul> <a class="toc-extra-link link-gray-900" href="https://www.britannica.com/animal/lion/additional-info">References & Edit History</a> <a class="toc-extra-link link-gray-900" href="/facts/lion">Quick Facts & Related Topics</a> </div> <div class="tlr-media-slider pb-10 mb-30"> <a class="section-header link-gray-900 font-serif font-14 font-weight-bold mb-10 mx-10" href="https://www.britannica.com/animal/lion/images-videos">Images & Videos</a> <div class="slider js-slider position-relative d-inline-flex align-items-center mw-100 "> <div class="slider-container js-slider-container overflow-hidden d-flex overflow-hidden text-nowrap ml-15"> <a href="https://cdn.britannica.com/55/2155-050-604F5A4A/lion.jpg" data-href="/media/1/342664/6392" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/55/2155-004-C6B1F0F4/lion.jpg" alt="A male lion" height="50" /> </a> <a href="https://cdn.britannica.com/29/150929-050-547070A1/lion-Kenya-Masai-Mara-National-Reserve.jpg" data-href="/media/1/342664/159005" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/29/150929-004-4E4D59AF/lion-Kenya-Masai-Mara-National-Reserve.jpg" alt="male lion" height="50" /> </a> <a href="https://cdn.britannica.com/70/92770-050-1648428C/Lioness.jpg" data-href="/media/1/342664/97911" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/70/92770-004-3A0BC4D6/Lioness.jpg" alt="lion" height="50" /> </a> <a href="https://cdn.britannica.com/05/75105-050-AE61BF35/Pride-lions.jpg" data-href="/media/1/342664/159007" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/05/75105-004-DB434D32/Pride-lions.jpg" alt="pride of lions" height="50" /> </a> <a href="https://cdn.britannica.com/97/92697-050-39C05D91/Lions-warthog.jpg" data-href="/media/1/342664/159006" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/97/92697-004-696FD40F/Lions-warthog.jpg" alt="lions chasing a warthog" height="50" /> </a> <a href="https://cdn.britannica.com/56/2156-050-52DEB3CA/Lioness-cubs.jpg" data-href="/media/1/342664/6393" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/56/2156-004-6A4AC3C1/Lioness-cubs.jpg" alt="lioness with cubs" height="50" /> </a> <a href="https://cdn.britannica.com/46/92746-050-83104CD7/Lioness-cubs.jpg" data-href="/media/1/342664/159008" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/46/92746-004-D3C0E5D1/Lioness-cubs.jpg" alt="lioness with cubs" height="50" /> </a> <a href="https://cdn.britannica.com/00/171900-050-7F00EE20/Cave-lion.jpg" data-href="/media/1/342664/188075" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/00/171900-004-43B0A07D/Cave-lion.jpg" alt="cave lion" height="50" /> </a> <a href="https://cdn.britannica.com/30/150930-050-51878468/lion-Namibia.jpg" data-href="/media/1/342664/159009" class="media-overlay-link d-inline-block mr-5"> <img loading="lazy" src="https://cdn.britannica.com/30/150930-120-D3D93F1E/lion-Namibia.jpg" alt="male lion" height="50" /> </a> <a href="/video/efforts-lions-Namibia/196661" data-href="/media/1/342664/196661" class="media-overlay-link d-inline-block mr-5"> <div class="position-relative --aspect-ratio: 16/9"> <img loading="lazy" src="https://cdn.britannica.com/37/179737-138-C574548A/efforts-lions-Namibia.jpg?w=400&h=225&c=crop" alt="Inside the Afri-Leo Foundation: Protecting lions in Namibia" class="col-100" /> <div class="btn btn-sm btn-white btn-circle position-absolute shadow" style="top: 50%; left:50%; transform: translate(-50%, -50%)"> <em class="material-icons font-14" data-icon="play_arrow" > </em> </div> </div> </a> </div> <button disabled class="prev-button js-prev-button position-absolute btn btn-circle shadow btn-blue " aria-label="Previous"> <span class="material-icons md-24" data-icon="keyboard_arrow_left"></span> </button> <button disabled class="next-button js-next-button position-absolute btn btn-circle shadow btn-blue " aria-label="Next"> <span class="material-icons md-24" data-icon="keyboard_arrow_right"></span> </button> </div> </div> <div class="mb-30 tlr-student-links"> <div class="text-gray-900 p-5 font-serif font-14 font-weight-bold mx-10 mb-10"> For Students </div> <div class="imagelink-with-image-on-the-side card card-horizontal tlr-img-with-side-link ml-15 link-gray-900 mb-10" > <div class="position-relative card-media" style="flex: 0;"> <a class="ilf-image position-relative" href="/summary/lion"> <img loading="lazy" src="https://cdn.britannica.com/55/2155-050-604F5A4A/lion.jpg?w=200&h=200&c=crop" alt="A male lion" width="200" height="200" /> </a> </div> <div class="card-body ilf-content"> <a class="font-weight-semi-bold d-block mb-5 font-16 ilf-title" href="/summary/lion" >lion summary</a> </div> </div> </div> <div class="mb-30 tlr-related-quizzes"> <div class="text-gray-900 p-5 font-serif font-14 font-weight-bold mx-10 mb-10"> Quizzes </div> <div class="imagelink-with-image-on-the-side card card-horizontal tlr-img-with-side-link ml-15 link-gray-900 mb-10" > <div class="position-relative card-media" style="flex: 0;"> <a class="ilf-image position-relative" href="/quiz/ultimate-animals-quiz"> <img loading="lazy" src="https://cdn.britannica.com/80/140480-131-28E57753/Dromedary-camels.jpg?w=200&h=200&c=crop" alt="Dromedary camels (Camelus dromedarius). Animals, mammals." width="200" height="200" /> </a> </div> <div class="card-body ilf-content"> <a class="font-weight-semi-bold d-block mb-5 font-16 ilf-title" href="/quiz/ultimate-animals-quiz" >Ultimate Animals Quiz</a> </div> </div> <div class="imagelink-with-image-on-the-side card card-horizontal tlr-img-with-side-link ml-15 link-gray-900 mb-10" > <div class="position-relative card-media" style="flex: 0;"> <a class="ilf-image position-relative" href="/quiz/animal-group-names"> <img loading="lazy" src="https://cdn.britannica.com/65/161265-131-DCC952FE/Sea-otter.jpg?w=200&h=200&c=crop" alt="Sea otter (Enhydra lutris), also called great sea otter, rare, completely marine otter of the northern Pacific, usually found in kelp beds. Floats on back. Looks like sea otter laughing. saltwater otters" width="200" height="200" /> </a> </div> <div class="card-body ilf-content"> <a class="font-weight-semi-bold d-block mb-5 font-16 ilf-title" href="/quiz/animal-group-names" >Animal Group Names</a> </div> </div> <div class="imagelink-with-image-on-the-side card card-horizontal tlr-img-with-side-link ml-15 link-gray-900 mb-10" > <div class="position-relative card-media" style="flex: 0;"> <a class="ilf-image position-relative" href="/quiz/deadliest-animals-quiz"> <img loading="lazy" src="https://cdn.britannica.com/54/92754-131-661B61DB/lion-Panthera-leo.jpg?w=200&h=200&c=crop" alt="Lion (panthera leo)" width="200" height="200" /> </a> </div> <div class="card-body ilf-content"> <a class="font-weight-semi-bold d-block mb-5 font-16 ilf-title" href="/quiz/deadliest-animals-quiz" >Deadliest Animals Quiz</a> </div> </div> <div class="imagelink-with-image-on-the-side card card-horizontal tlr-img-with-side-link ml-15 link-gray-900 mb-10" > <div class="position-relative card-media" style="flex: 0;"> <a class="ilf-image position-relative" href="/quiz/wild-words-from-the-animal-kingdom-vocabulary-quiz"> <img loading="lazy" src="https://cdn.britannica.com/09/237309-131-E04A5A59/chimp-scratching-head.jpg?w=200&h=200&c=crop" alt="Young chimpanzee dressed in a shirt and sweater vest, scratching his head thinking. (primates)" width="200" height="200" /> </a> </div> <div class="card-body ilf-content"> <a class="font-weight-semi-bold d-block mb-5 font-16 ilf-title" href="/quiz/wild-words-from-the-animal-kingdom-vocabulary-quiz" >Wild Words from the Animal Kingdom Vocabulary Quiz</a> </div> </div> <div class="imagelink-with-image-on-the-side card card-horizontal tlr-img-with-side-link ml-15 link-gray-900 mb-10" > <div class="position-relative card-media" style="flex: 0;"> <a class="ilf-image position-relative" href="/quiz/match-the-baby-animal-to-its-mama-quiz"> <img loading="lazy" src="https://cdn.britannica.com/57/243957-131-4BD7E5E6/mute-swan-with-cygnet.jpg?w=200&h=200&c=crop" alt="Mute swan with cygnet. (birds)" width="200" height="200" /> </a> </div> <div class="card-body ilf-content"> <a class="font-weight-semi-bold d-block mb-5 font-16 ilf-title" href="/quiz/match-the-baby-animal-to-its-mama-quiz" >Match the Baby Animal to Its Mama Quiz</a> </div> </div> </div> <div class="mb-30 tlr-related-questions"> <div class="text-gray-900 p-5 pb-0 font-serif font-14 font-weight-bold mx-10 mb-15"> Related Questions </div> <ul> <li class="link-gray-900 mb-15"><a class="" href="/question/What-is-a-pride-of-lions">What is a pride of lions?</a> </li> <li class="link-gray-900 mb-15"><a class="" href="/question/Where-did-Barbary-lions-historically-live"> Where did Barbary lions historically live? </a> </li> <li class="link-gray-900 mb-15"><a class="" href="/question/What-were-Barbary-lions-known-for"> What were Barbary lions known for? </a> </li> <li class="link-gray-900 mb-15"><a class="" href="/question/Where-can-descendants-of-Barbary-lions-be-found-today"> Where can descendants of Barbary lions be found today? </a> </li> </ul> </div> <div class="mb-30 tlr-read-next"> <div class="text-gray-900 p-5 font-serif font-14 font-weight-bold mx-10 mb-10"> Read Next </div> <div class="imagelink-with-image-on-the-side card card-horizontal tlr-img-with-side-link ml-15 link-gray-900 mb-10" > <div class="position-relative card-media" style="flex: 0;"> <a class="ilf-image position-relative" href="/list/abundant-animals-the-most-numerous-organisms-in-the-world"> <img loading="lazy" src="https://cdn.britannica.com/48/173648-131-B3E416AD/animal-Frog-Amphibian-grass-Anura-Ranidae.jpg?w=200&h=200&c=crop" alt="animal. 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Let us know if you have suggestions to improve this article (requires login). </div> <div class="type-menu"> <label for="feedback-type" class="label mb-10">Feedback Type</label> <select id="feedback-type" class="form-select mb-30" name="feedbackTypeId" required> <option value="" selected="selected">Select a type (Required)</option> <option value="1">Factual Correction</option> <option value="2">Spelling/Grammar Correction</option> <option value="3">Link Correction</option> <option value="4">Additional Information</option> <option value="5">Other</option> </select> </div> <label for="feedback" class="label mb-10">Your Feedback</label> <textarea id="feedback" class="form-control mb-30" name="feedback" maxlength="3000" rows="7" required></textarea> <button class="btn btn-blue" type="submit">Submit Feedback</button> </form> <div class="success-messaging d-none mt-30"> <div class="title">Thank you for your feedback</div> <p>Our editors will review what you’ve submitted and determine whether to revise the article.</p> </div> </div> </div> <div class="md-websites-modal size-lg d-none"> <div class="md-modal-body"> <div class="h2 font-serif pb-15 border-bottom font-weight-bold"> External Websites </div> <div class="pb-20"> <ul class="list-unstyled mt-20 lh-lg"> <li><a class="external" href="https://animalcorner.org/animals/lion/" target="_blank" rel="noopener ">Animal Corner - Lion</a></li> <li><a class="external" href="https://a-z-animals.com/animals/lion/" target="_blank" rel="noopener ">A-Z Animals - Lion</a></li> <li><a class="external" href="https://animaldiversity.org/accounts/Panthera_leo/" target="_blank" rel="noopener ">Animal Diversity Web - Lion</a></li> <li><a class="external" href="https://www.livescience.com/27404-lion-facts.html" target="_blank" rel="noopener ">Live Science - Lions: Facts, behavior and news</a></li> <li><a class="external" href="https://animals.sandiegozoo.org/animals/lion" target="_blank" rel="noopener ">San Diego Zoo - Animals & Plants - Lion</a></li> </ul> </div> <div class="md-websites-ebk-title">Britannica Websites</div> <div class="md-websites-ebk-subtitle">Articles from Britannica Encyclopedias for elementary and high school students.</div> <ul class="list-unstyled bps-topic-web-sites lh-lg"> <li><a class="external" href="https://kids.britannica.com/kids/article/lion/353389" target="_blank" rel="noopener">lion - Children's Encyclopedia (Ages 8-11)</a></li> <li><a class="external" href="https://kids.britannica.com/students/article/lion/275490" target="_blank" rel="noopener">lion - Student Encyclopedia (Ages 11 and up)</a></li> </ul> </div> </div> </div> </div> <div class="toc-header-marker"></div> <button class="ai-ask-button btn border-2 js-header-ai-ask-button d-none btn-sm btn-outline-red-400 border-red-400 mr-0 mr-lg-10 ml-5 ml-sm-10 ml-lg-0 p-10"> Ask the Chatbot a Question </button> <div class="caption alternate-titles">Also known as: Panthera leo</div> <div class="md-byline module-spacing "> <div class="font-serif font-12"> <span class="written-by text-gray-700"> Written by </span> <div class="editor-popover popover p-0"> <a class="d-block p-20 gtm-byline font-12 byline-contributor" href="/contributor/Roland-W-Kays/5443" > <div class="editor-title font-16 font-weight-bold">Roland W. Kays</div> <div class="editor-description font-12 font-serif mt-5 clamp-description text-black">Curator of Mammals, New York State Museum, Albany.</div> </a> <div data-popper-arrow></div> </div> <span class="btn btn-link editor-link p-0 qa-byline-link gtm-byline font-12 byline-contributor text-decoration-underline"> Roland W. Kays</span></div> <div class="font-serif font-12 text-gray-700"> <span class="qa-fact-checked-by">Fact-checked by</span> <div class="editor-popover popover p-0"> <a class="d-block p-20 font-12" href="/editor/The-Editors-of-Encyclopaedia-Britannica/4419" > <div class="editor-title font-16 font-weight-bold">The Editors of Encyclopaedia Britannica</div> <div class="editor-description font-12 font-serif mt-5 text-black">Encyclopaedia Britannica's editors oversee subject areas in which they have extensive knowledge, whether from years of experience gained by working on that content or via study for an advanced degree. They write new content and verify and edit content received from contributors.</div> </a> <div data-popper-arrow></div> </div> <span class="btn btn-link editor-link p-0 qa-byline-link font-12 "> The Editors of Encyclopaedia Britannica</span></div> <div class="last-updated font-12 font-serif"> <span class="text-gray-700"> Last Updated: <time datetime="2024-11-23T00:00:00CST" >Nov 23, 2024</time> •</span> <a class="byline-edit-history" href="https://www.britannica.com/animal/lion/additional-info#history" rel="nofollow">Article History</a> </div></div> </div> <button class="d-flex d-lg-none btn btn-outline-blue border rounded-sm shadow-sm mobile-toc-button gtm-mobile-toc-inline-button d-none d-sm-block js-sections-inline-button module-spacing btn d-lg-none"> <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="toc"></em> Table of Contents </button> <div class="d-flex d-sm-none flex-row"> <button class="d-flex d-lg-none btn btn-outline-blue border rounded-sm shadow-sm mobile-toc-button gtm-mobile-toc-inline-button js-sections-inline-button module-spacing"> <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="toc"></em> Table of Contents </button> <button class="ai-ask-button btn border-2 ai-ask-button btn border-2 module-spacing btn-sm js-inline-ai-ask-button btn-outline-red-400 border-red-400 p-10 ml-5"> Ask the Chatbot a Question </button> </div> <div class="js-qf-module qf-module px-40 px-sm-20 py-15 mx-auto module-spacing font-14 bg-gray-50 rounded"> <div class="facts-list mt-10"> <div class=""> <div class="js-fact mb-10 line-clamp clamp-3"> <dl> <dt>Related Topics: </dt> <dd><a href="/animal/Barbary-lion" topicid="1181310">Barbary lion</a></dd> <dd><a href="/animal/Asiatic-lion" topicid="38673">Asiatic lion</a></dd> <dd><a href="/animal/American-lion-extinct-mammal" topicid="2123175">American lion</a></dd> <dd><a href="/animal/cave-lion" topicid="2009903">cave lion</a></dd> <dd><a href="/animal/leopon" topicid="2123169">leopon</a></dd> </dl> <button class="js-more-btn d-none btn btn-unstyled font-12 bg-gray-50" aria-label="Toggle more/less fact data"> <em class="js-content link-blue">(Show more)</em> </button> </div> <div class="text-center"> <a class="btn btn-sm btn-link p-0" href="/facts/lion"> See all related content </a> </div> </div> </div> </div><script type="application/ld+json"> { "@context": "https://schema.org", "@type": "FAQPage", "mainEntity": [ { "@type" : "Question", "name" : "<div>How long do lions live?<\/div><em><\/em>", "acceptedAnswer" : { "@type" : "Answer", "text" : "In the wild, lions usually live no more than 8 to 10 years because of attacks by humans or other lions, or the effects of goring or kicks from intended prey. In captivity, they may live over 25 years." } } , { "@type" : "Question", "name" : "<div>Where do lions live?<\/div><em><\/em>", "acceptedAnswer" : { "@type" : "Answer", "text" : "Lions live in a variety of habitats but prefer grassland, savanna, dense scrub, and open woodland. Historically, they lived across much of Europe, Asia, and Africa, but now they are mainly found in parts of Africa south of the <a href=\"https:\/\/www.britannica.com\/place\/Sahara-desert-Africa\">Sahara<\/a>. " } } , { "@type" : "Question", "name" : "<div>What is a pride of lions?<\/div><em><\/em>", "acceptedAnswer" : { "@type" : "Answer", "text" : "A pride is a group of lions that live together. The members of a pride spend days in several scattered groups that meet to hunt or share a meal. Each pride has its own territory that it defends, ranging from 20 square km (8 square miles) if food is abundant to 400 square km (around 150 square miles) if food is sparse. " } } , { "@type" : "Question", "name" : "<div>What is the purpose of a lion\u2019s mane?<\/div><em><\/em>", "acceptedAnswer" : { "@type" : "Answer", "text" : "Manes make male lions look larger than they really are, which may function to intimidate rivals and impress prospective mates. Manes vary from one lion to another and may be entirely absent. " } } , { "@type" : "Question", "name" : "<div>What do lions eat?<\/div><em><\/em>", "acceptedAnswer" : { "@type" : "Answer", "text" : "Lions usually hunt and eat medium-sized to large hoofed animals like wildebeests, zebras, and antelopes. They occasionally also <a href=\"https:\/\/www.britannica.com\/science\/predation\">prey on<\/a> larger animals, especially sick or injured ones, and eat found meat such as carrion." } } ] } </script> <div class="top-questions qa-accordion d-flex flex-column module-spacing"><div class="font-weight-bold font-14 mb-5"> Top Questions </div><div id="intent-accordion" class="md-intent-accordion"><div class="top-question bg-gray-50 rounded" data-value="1"> <h3 class="accordion--question font-14 font-weight-normal cursor-pointer rounded"><div class="pe-none d-flex justify-content-between align-items-center"><div>How long do lions live?</div><em class="material-icons" data-icon="expand_more"></em></div></h3> <div class="accordion--answer hidden p-15 pt-5 font-16 mb-5"><div class="accordion--answer-copy"><p>In the wild, lions usually live no more than 8 to 10 years because of attacks by humans or other lions, or the effects of goring or kicks from intended prey. In captivity, they may live over 25 years.</p></div></div> </div><div class="top-question bg-gray-50 rounded" data-value="2"> <h3 class="accordion--question font-14 font-weight-normal cursor-pointer rounded"><div class="pe-none d-flex justify-content-between align-items-center"><div>Where do lions live?</div><em class="material-icons" data-icon="expand_more"></em></div></h3> <div class="accordion--answer hidden p-15 pt-5 font-16 mb-5"><div class="accordion--answer-copy"><p>Lions live in a variety of habitats but prefer grassland, savanna, dense scrub, and open woodland. Historically, they lived across much of Europe, Asia, and Africa, but now they are mainly found in parts of Africa south of the <a href="https://www.britannica.com/place/Sahara-desert-Africa" class="md-crosslink " data-show-preview="true">Sahara</a>. </p></div></div> </div><div class="top-question bg-gray-50 rounded" data-value="3"> <h3 class="accordion--question font-14 font-weight-normal cursor-pointer rounded"><div class="pe-none d-flex justify-content-between align-items-center"><div>What is a pride of lions?</div><em class="material-icons" data-icon="expand_more"></em></div></h3> <div class="accordion--answer hidden p-15 pt-5 font-16 mb-5"><div class="accordion--answer-copy"><p>A pride is a group of lions that live together. The members of a pride spend days in several scattered groups that meet to hunt or share a meal. Each pride has its own territory that it defends, ranging from 20 square km (8 square miles) if food is abundant to 400 square km (around 150 square miles) if food is sparse. </p></div></div> </div><div class="top-question bg-gray-50 rounded" data-value="4"> <h3 class="accordion--question font-14 font-weight-normal cursor-pointer rounded"><div class="pe-none d-flex justify-content-between align-items-center"><div>What is the purpose of a lion’s mane?</div><em class="material-icons" data-icon="expand_more"></em></div></h3> <div class="accordion--answer hidden p-15 pt-5 font-16 mb-5"><div class="accordion--answer-copy"><p>Manes make male lions look larger than they really are, which may function to intimidate rivals and impress prospective mates. Manes vary from one lion to another and may be entirely absent. </p></div></div> </div><div class="top-question bg-gray-50 rounded" data-value="5"> <h3 class="accordion--question font-14 font-weight-normal cursor-pointer rounded"><div class="pe-none d-flex justify-content-between align-items-center"><div>What do lions eat?</div><em class="material-icons" data-icon="expand_more"></em></div></h3> <div class="accordion--answer hidden p-15 pt-5 font-16 mb-5"><div class="accordion--answer-copy"><p>Lions usually hunt and eat medium-sized to large hoofed animals like wildebeests, zebras, and antelopes. They occasionally also <a href="https://www.britannica.com/science/predation" class="md-crosslink " data-show-preview="true">prey on</a> larger animals, especially sick or injured ones, and eat found meat such as carrion.</p></div></div> </div><button class="show-more-button js-toggle-top-questions btn btn-unstyled font-14 d-flex pr-10 rounded-sm"><em class="material-icons" data-icon="expand_more"></em></button></div></div><div class="bg-gray-50 p-15 rounded module-spacing recent-news d-flex flex-column float-false"> <div> <h2 class="font-weight-bold font-14 m-0 d-inline"> News <span class="text-gray-600">•</span> </h2> <div class="recent-news-item first-recent-news-item d-inline"> <a class="font-14 gtm-ap-news-link" href="https://www.nytimes.com/2024/11/21/world/asia/putin-russia-north-korea-animals.html" rel="nofollow">Russia Gifts Bears and a Lion to North Korea</a> <span class="font-14 text-gray-600"> <span>•</span> Nov. 22, 2024, 1:44 AM ET (New York Times) <button class="btn btn-link d-inline p-0 font-12 js-toggle-recent-news"> <span class="text-gray-500">...</span><span>(Show more)</span> </button> </span> </div> </div> <div class="rest-of-recent-news-items"> <div class="recent-news-item mt-5"> <a class="font-14 gtm-ap-news-link" href="/news/342664/e371d8463f7baab8730896c69e60094a" rel="nofollow">A lion cub evacuated from Lebanon to a South African sanctuary escapes airstrikes and abuse</a> <span class="font-14 text-gray-600"> <span>•</span> Nov. 15, 2024, 8:25 AM ET (AP) </span> </div> <button class="js-toggle-recent-news d-flex btn btn-unstyled font-14 pr-10 rounded-sm mt-10" aria-label="Toggle additional news items"> Show less <em class="material-icons" data-icon="expand_less"></em> </button> </div> </div><!--[BEFORE-ARTICLE]--><span class="marker before-article"></span><section data-level="1" id="ref1"><!--[PREMOD1]--><span class="marker PREMOD1 mod-inline"></span><p class="topic-paragraph"><strong><span id="ref75417"></span>lion</strong>, (<em>Panthera leo</em>), large, powerfully built <a href="https://www.britannica.com/animal/cat" class="md-crosslink autoxref " data-show-preview="true">cat</a> (family Felidae) that is second in size only to the <a href="https://www.britannica.com/animal/tiger" class="md-crosslink " data-show-preview="true">tiger</a>; it is a famous <span id="ref1321423"></span><a href="https://www.britannica.com/science/apex-predator" class="md-crosslink " data-show-preview="true">apex predator</a> (meaning without a natural predator or enemy). The proverbial “king of beasts,” the lion has been one of the best-known wild animals since earliest times. Lions are most active at night and live in a variety of habitats but prefer <a href="https://www.britannica.com/science/grassland" class="md-crosslink " data-show-preview="true">grassland</a>, <a href="https://www.britannica.com/science/savanna" class="md-crosslink " data-show-preview="true">savanna</a>, dense <a href="https://www.britannica.com/science/scrubland" class="md-crosslink " data-show-preview="true">scrub</a>, and open woodland. Historically, they ranged across much of <a href="https://www.britannica.com/place/Europe" class="md-crosslink " data-show-preview="true">Europe</a>, <a href="https://www.britannica.com/place/Asia" class="md-crosslink " data-show-preview="true">Asia</a>, and <a href="https://www.britannica.com/place/Africa" class="md-crosslink " data-show-preview="true">Africa</a>, but now they are found mainly in parts of Africa south of the <a href="https://www.britannica.com/place/Sahara-desert-Africa" class="md-crosslink " data-show-preview="true">Sahara</a>. An isolated population of about between 500 and 700 wild Asiatic lions <a class="md-dictionary-link md-dictionary-tt-off mw" data-term="constitute" href="https://www.merriam-webster.com/dictionary/constitute" data-type="MW">constitute</a> a slightly smaller population that lives under strict protection in <a href="https://www.britannica.com/place/India" class="md-crosslink " data-show-preview="true">India</a>’s <span id="ref1265179"></span><a href="https://www.britannica.com/place/Gir-National-Park" class="md-crosslink " data-show-preview="true">Gir National Park</a> and Wildlife Sanctuary.</p><!--[MOD1]--><span class="marker MOD1 mod-inline"></span></section> <!--[H2]--><span class="marker h2"></span><section data-level="1" id="ref325410"> <h2 class="h1">General characteristics</h2> <!--[PREMOD2]--><span class="marker PREMOD2 mod-inline"></span><div class="assemblies multiple medialist slider js-slider position-relative d-inline-flex align-items-center mw-100" data-type="other"><div class="slider-container js-slider-container overflow-hidden d-flex"><div class="rw-track d-flex align-items-center"><div class="position-relative rw-slide col-100 px-20 "><figure class="md-assembly m-0 mb-md-0 card card-borderless print-false" data-assembly-id="159005" data-asm-type="image"><div class="md-assembly-wrapper card-media " data-type="image"><a href="https://cdn.britannica.com/29/150929-050-547070A1/lion-Kenya-Masai-Mara-National-Reserve.jpg" class="gtm-assembly-link position-relative d-flex align-items-center justify-content-center media-overlay-link card-media" data-href="/media/1/342664/159005"><picture><source media="(min-width: 680px)" srcset="https://cdn.britannica.com/29/150929-050-547070A1/lion-Kenya-Masai-Mara-National-Reserve.jpg?w=300"><img src="https://cdn.britannica.com/29/150929-050-547070A1/lion-Kenya-Masai-Mara-National-Reserve.jpg?w=300" alt="male lion" data-width="1600" data-height="1085" loading="eager"></picture><div class="position-absolute top-10 left-10 assembly-slide-tag rounded-lg">1 of 2</div><button class="magnifying-glass btn btn-circle position-absolute shadow btn-white top-10 right-10" aria-label="Zoom in"><em class="material-icons link-blue" data-icon="zoom_in"></em></button></a></div><figcaption class="card-body"><div class="md-assembly-caption text-muted font-14 font-serif line-clamp"><span><a class="gtm-assembly-link md-assembly-title font-weight-bold d-inline font-sans-serif mr-5 media-overlay-link" href="https://cdn.britannica.com/29/150929-050-547070A1/lion-Kenya-Masai-Mara-National-Reserve.jpg" data-href="/media/1/342664/159005">male lion</a><span>Male lion (<em>Panthera leo</em>) in the Masai Mara National Reserve, Kenya.</span><button class="js-more-btn d-none btn btn-unstyled font-12 bg-white js-content" aria-label="Toggle more/less fact data"><span class="link-blue">(more)</span></button></span></div></figcaption></figure></div><div class="position-relative rw-slide col-100 px-20 "><figure class="md-assembly m-0 mb-md-0 card card-borderless print-false" data-assembly-id="97911" data-asm-type="image"><div class="md-assembly-wrapper card-media " data-type="image"><a href="https://cdn.britannica.com/70/92770-050-1648428C/Lioness.jpg" class="gtm-assembly-link position-relative d-flex align-items-center justify-content-center media-overlay-link card-media" data-href="/media/1/342664/97911"><picture><source media="(min-width: 680px)" srcset="https://cdn.britannica.com/70/92770-050-1648428C/Lioness.jpg?w=300"><img src="https://cdn.britannica.com/70/92770-050-1648428C/Lioness.jpg?w=300" alt="lion" data-width="1040" data-height="1600" loading="eager"></picture><div class="position-absolute top-10 left-10 assembly-slide-tag rounded-lg">2 of 2</div><button class="magnifying-glass btn btn-circle position-absolute shadow btn-white top-10 right-10" aria-label="Zoom in"><em class="material-icons link-blue" data-icon="zoom_in"></em></button></a></div><figcaption class="card-body"><div class="md-assembly-caption text-muted font-14 font-serif line-clamp"><span><a class="gtm-assembly-link md-assembly-title font-weight-bold d-inline font-sans-serif mr-5 media-overlay-link" href="https://cdn.britannica.com/70/92770-050-1648428C/Lioness.jpg" data-href="/media/1/342664/97911">lion</a><span>Lioness (<em>Panthera leo</em>).</span><button class="js-more-btn d-none btn btn-unstyled font-12 bg-white js-content" aria-label="Toggle more/less fact data"><span class="link-blue">(more)</span></button></span></div></figcaption></figure></div></div></div><button disabled="true" class="prev-button js-prev-button position-absolute btn btn-circle shadow btn-lg btn-blue-dark m-20"><span class="material-icons" data-icon="keyboard_arrow_left"></span></button><button disabled="true" class="next-button js-next-button position-absolute btn btn-circle shadow btn-lg btn-blue-dark m-20"><span class="material-icons" data-icon="keyboard_arrow_right"></span></button></div><p class="topic-paragraph">The lion is a well-muscled cat with a long body, large head, and short legs. Size and appearance vary considerably between the sexes. The male’s outstanding characteristic is his <span id="ref75418"></span>mane, which varies between different individuals and populations. It may be entirely lacking; it may fringe the face; or it may be full and shaggy, covering the back of the head, neck, and shoulders and continuing onto the throat and chest to join a fringe along the belly. In some lions the mane and fringe are very dark, almost black, giving the cat a <a class="md-dictionary-link md-dictionary-tt-off eb" data-term="majestic" href="https://www.britannica.com/dictionary/majestic" data-type="EB">majestic</a> appearance. Manes make males look larger and may serve to intimidate rivals or impress prospective mates. A full-grown male is about 1.8–2.1 metres (6–7 feet) long, excluding the 1-metre tail; he stands about 1.2 metres high at the shoulder and weighs 170–230 kg (370–500 pounds). The female, or lioness, is smaller, with a body length of 1.5 metres, a shoulder height of 0.9–1.1 metres, and a weight of 120–180 kg. The lion’s coat is short and varies in colour from buff yellow, orange-brown, or silvery gray to dark brown, with a tuft on the tail tip that is usually darker than the rest of the coat.</p><!--[MOD2]--><span class="marker MOD2 mod-inline"></span> </section> <!--[H3]--><span class="marker h3"></span><section data-level="1" id="ref325411"> <h2 class="h1"><span id="ref75419"></span><a href="https://www.britannica.com/science/pride-animal-behavior" class="md-crosslink ">Prides</a></h2> <!--[PREMOD3]--><span class="marker PREMOD3 mod-inline"></span><div class="assemblies"><div class="w-100"><figure class="md-assembly m-0 mb-md-0 card card-borderless print-false" data-assembly-id="159007" data-asm-type="image"><div class="md-assembly-wrapper card-media " data-type="image"><a href="https://cdn.britannica.com/05/75105-050-AE61BF35/Pride-lions.jpg" class="gtm-assembly-link position-relative d-flex align-items-center justify-content-center media-overlay-link card-media" data-href="/media/1/342664/159007"><picture><source media="(min-width: 680px)" srcset="https://cdn.britannica.com/05/75105-050-AE61BF35/Pride-lions.jpg?w=300"><img src="https://cdn.britannica.com/05/75105-050-AE61BF35/Pride-lions.jpg?w=300" alt="pride of lions" data-width="1600" data-height="926" loading="eager"></picture><button class="magnifying-glass btn btn-circle position-absolute shadow btn-white top-10 right-10" aria-label="Zoom in"><em class="material-icons link-blue" data-icon="zoom_in"></em></button></a></div><figcaption class="card-body"><div class="md-assembly-caption text-muted font-14 font-serif line-clamp"><span><a class="gtm-assembly-link md-assembly-title font-weight-bold d-inline font-sans-serif mr-5 media-overlay-link" href="https://cdn.britannica.com/05/75105-050-AE61BF35/Pride-lions.jpg" data-href="/media/1/342664/159007">pride of lions</a><span>Pride of lions (<em>Panthera leo</em>).</span><button class="js-more-btn d-none btn btn-unstyled font-12 bg-white js-content" aria-label="Toggle more/less fact data"><span class="link-blue">(more)</span></button></span></div></figcaption></figure></div></div><p class="topic-paragraph">Lions are <a class="md-dictionary-link md-dictionary-tt-off eb" data-term="unique" href="https://www.britannica.com/dictionary/unique" data-type="EB">unique</a> among cats in that they live in a group, or pride. The members of a pride typically spend the day in several scattered groups that may unite to hunt or share a meal. A pride consists of several generations of lionesses, some of which are related, a smaller number of breeding males, and their cubs. The group may consist of as few as 4 or as many as 37 members, but about 15 is the average size. Each pride has a well-defined <span id="ref75420"></span><a href="https://www.britannica.com/science/territorial-behaviour" class="md-crosslink " data-show-preview="true">territory</a> consisting of a core area that is strictly defended against intruding lions and a fringe area where some overlap is tolerated. Where prey is abundant, a territory area may be as small as 20 square km (8 square miles), but if game is <a class="md-dictionary-link md-dictionary-tt-off eb" data-term="sparse" href="https://www.britannica.com/dictionary/sparse" data-type="EB">sparse</a>, it may cover up to 400 square km. Some prides have been known to use the same territory for decades, passing the area on between females. Lions proclaim their territory by roaring and by <span id="ref75421"></span><a href="https://www.britannica.com/science/scent-mark" class="md-crosslink ">scent marking</a>. Their distinctive roar is generally delivered in the evening before a night’s <a href="https://www.britannica.com/sports/hunting-sport" class="md-crosslink " data-show-preview="true">hunting</a> and again before getting up at dawn. Males also proclaim their presence by urinating on bushes, trees, or simply on the ground, leaving a pungent scent behind. <a href="https://www.britannica.com/science/defecation-physiology" class="md-crosslink " data-show-preview="true">Defecation</a> and rubbing against bushes leave different scent markings.</p><!--[MOD3]--><span class="marker MOD3 mod-inline"></span> <!--[PREMOD4]--><span class="marker PREMOD4 mod-inline"></span><p class="topic-paragraph">There are a number of competing evolutionary explanations for why lions form groups. Large body size and high <a class="md-dictionary-link md-dictionary-tt-off eb" data-term="density" href="https://www.britannica.com/dictionary/density" data-type="EB">density</a> of their main prey probably make group life more efficient for females in terms of energy expenditure. Groups of females, for example, hunt more effectively and are better able to defend cubs against infanticidal males and their hunting territory against other females. The relative importance of these factors is debated, and it is not clear which was responsible for the establishment of group life and which are secondary benefits.</p><!--[MOD4]--><span class="marker MOD4 mod-inline"></span> </section> <!--[H4]--><span class="marker h4"></span><section data-level="1" id="ref325412"> <h2 class="h1">Hunting</h2> <!--[PREMOD5]--><span class="marker PREMOD5 mod-inline"></span><div class="assemblies"><div class="w-100"><figure class="md-assembly m-0 mb-md-0 card card-borderless print-false" data-assembly-id="159006" data-asm-type="image"><div class="md-assembly-wrapper card-media " data-type="image"><a href="https://cdn.britannica.com/97/92697-050-39C05D91/Lions-warthog.jpg" class="gtm-assembly-link position-relative d-flex align-items-center justify-content-center media-overlay-link card-media" data-href="/media/1/342664/159006"><picture><source media="(min-width: 680px)" srcset="https://cdn.britannica.com/97/92697-050-39C05D91/Lions-warthog.jpg?w=300"><img src="https://cdn.britannica.com/97/92697-050-39C05D91/Lions-warthog.jpg?w=300" alt="lions chasing a warthog" data-width="1600" data-height="1064" loading="eager"></picture><button class="magnifying-glass btn btn-circle position-absolute shadow btn-white top-10 right-10" aria-label="Zoom in"><em class="material-icons link-blue" data-icon="zoom_in"></em></button></a></div><figcaption class="card-body"><div class="md-assembly-caption text-muted font-14 font-serif line-clamp"><span><a class="gtm-assembly-link md-assembly-title font-weight-bold d-inline font-sans-serif mr-5 media-overlay-link" href="https://cdn.britannica.com/97/92697-050-39C05D91/Lions-warthog.jpg" data-href="/media/1/342664/159006">lions chasing a warthog</a><button class="js-more-btn d-none btn btn-unstyled font-12 bg-white js-content" aria-label="Toggle more/less fact data"><span class="link-blue">(more)</span></button></span></div></figcaption></figure></div></div><p class="topic-paragraph">Lions <span id="ref75422"></span><a href="https://www.britannica.com/science/predation" class="md-crosslink " data-show-preview="true">prey</a> on a large variety of animals ranging in size from <a href="https://www.britannica.com/animal/rodent" class="md-crosslink " data-show-preview="true">rodents</a> and <a href="https://www.britannica.com/animal/baboon" class="md-crosslink " data-show-preview="true">baboon</a>s to <a href="https://www.britannica.com/animal/Cape-buffalo" class="md-crosslink " data-show-preview="true">Cape (or African) buffalo</a> and <a href="https://www.britannica.com/animal/hippopotamus-mammal-species" class="md-crosslink " data-show-preview="true">hippopotamuses</a>, but they predominantly hunt medium- to large-sized hoofed animals such as wildebeests, <a href="https://www.britannica.com/animal/zebra" class="md-crosslink " data-show-preview="true">zebras</a>, and <a href="https://www.britannica.com/animal/antelope-mammal" class="md-crosslink " data-show-preview="true">antelopes</a>. Prey preferences vary geographically as well as between neighbouring prides. Lions are known to take <a href="https://www.britannica.com/animal/elephant-mammal" class="md-crosslink " data-show-preview="true">elephants</a> and <a href="https://www.britannica.com/animal/giraffe" class="md-crosslink " data-show-preview="true">giraffes</a>, but only if the individual is young or especially sick. They readily eat any meat they can find, including carrion and fresh kills that they scavenge or forcefully steal from <a href="https://www.britannica.com/animal/hyena" class="md-crosslink " data-show-preview="true">hyenas</a>, <a href="https://www.britannica.com/animal/cheetah-mammal" class="md-crosslink " data-show-preview="true">cheetahs</a>, or wild dogs. Lionesses living in open savanna do most of the hunting, whereas males typically appropriate their meals from the female’s kills. However, male lions are also adept hunters, and in some areas they hunt frequently. Pride males in scrub or wooded habitat spend less time with the females and hunt most of their own meals. Nomadic males must always secure their own food.</p><a class="link-module shadow-sm d-block qa-quiz-module" href="/quiz/deadliest-animals-quiz" data-link-module-iframe-link=""> <img loading="lazy" src="https://cdn.britannica.com/54/92754-131-661B61DB/lion-Panthera-leo.jpg" alt="Lion (panthera leo)" class="rounded-sm mr-15" width="70" /> <div class="line-clamp clamp-5"> <div class="module-title bg-green">Britannica Quiz</div> <div class="font-weight-semi-bold mt-5">Deadliest Animals Quiz</div> </div> </a><div class="one-good-fact-module"> </div><!--[MOD5]--><span class="marker MOD5 mod-inline"></span> <div class="feature-block callout"><div class="font-weight-bold">Taxonomy</div><ul class="list-none "><li><div>Kingdom: <a href="https://www.britannica.com/animal/animal" class="md-crosslink " data-show-preview="true">Animalia</a></div></li><li><div>Class: <a href="https://www.britannica.com/animal/mammal" class="md-crosslink " data-show-preview="true">Mammalia</a></div></li><li><div>Order: <a href="https://www.britannica.com/animal/carnivore-mammal" class="md-crosslink " data-show-preview="true">Carnivora</a></div></li><li><div>Family: <a href="https://www.britannica.com/animal/feline" class="md-crosslink " data-show-preview="true">Felidae</a></div></li><li><div>Genus: <em>Panthera</em></div></li><li><div>Species: <em>Panthera leo</em></div></li></ul><p class="topic-paragraph"><strong>Vulnerable</strong></p></div> <!--[PREMOD6]--><span class="marker PREMOD6 mod-inline"></span><p class="topic-paragraph">Though a group of hunting lions is potentially nature’s most <a class="md-dictionary-link md-dictionary-tt-off mw" data-term="formidable" href="https://www.merriam-webster.com/dictionary/formidable" data-type="MW">formidable</a> predatory force on land, a high proportion of their hunts fail. The cats pay no attention to the wind’s direction (which can carry their scent to their prey), and they tire after running short distances. Typically, they stalk prey from nearby cover and then burst forth to run it down in a short, rapid rush. After leaping on the prey, the lion lunges at its neck and bites until the animal has been strangled. Other members of the pride quickly crowd around to feed on the kill, usually fighting for access. Hunts are sometimes conducted in groups, with members of a pride encircling a herd or approaching it from opposite directions, then closing in for a kill in the resulting panic. The cats typically <a class="md-dictionary-link md-dictionary-tt-off eb" data-term="gorge" href="https://www.britannica.com/dictionary/gorge" data-type="EB">gorge</a> themselves and then rest for several days in its vicinity. An adult male can consume more than 34 kg (75 pounds) of meat at a single meal and rest for a week before resuming the hunt. If prey is abundant, both sexes typically spend 21 to 22 hours a day resting, sleeping, or sitting and hunt for only 2 or 3 hours a day.</p><!--[MOD6]--><span class="marker MOD6 mod-inline"></span> </section> <!--[END-OF-CONTENT]--><span class="marker end-of-content"></span><!--[AFTER-ARTICLE]--><span class="marker after-article"></span></div> <div id="chatbot-root"></div> </div> </div> </div> <div class="ai-dialog-placeholder"></div> </div> </div> <aside class="col-md-da-320"></aside> </div> </div> </div> </div> </article></div> </div></div> </div> </main> <div id="md-footer"></div> <noscript><iframe src="//www.googletagmanager.com/ns.html?id=GTM-5W6NC8" height="0" width="0" style="display:none;visibility:hidden"></iframe></noscript> <script type="text/javascript" id="_informizely_script_tag"> var IzWidget = IzWidget || {}; (function (d) { var scriptElement = d.createElement('script'); scriptElement.type = 'text/javascript'; scriptElement.async = true; scriptElement.src = "https://insitez.blob.core.windows.net/site/f780f33e-a610-4ac2-af81-3eb184037547.js"; var node = d.getElementById('_informizely_script_tag'); node.parentNode.insertBefore(scriptElement, node); } )(document); </script> <!-- Ortto ebmwprod capture code --> <script> window.ap3c = window.ap3c || {}; var ap3c = window.ap3c; ap3c.cmd = ap3c.cmd || []; ap3c.cmd.push(function() { ap3c.init('ZO4siT4cLwnykPnzZWJtd3Byb2Q', 'https://engage.email.britannica.com/'); ap3c.track({v: 0}); }); ap3c.activity = function(act) { ap3c.act = (ap3c.act || []); ap3c.act.push(act); }; var s, t; s = document.createElement('script'); s.type = 'text/javascript'; s.src = "https://engage.email.britannica.com/app.js"; t = document.getElementsByTagName('script')[0]; t.parentNode.insertBefore(s, t); </script> <script class="marketing-page-info" type="application/json"> {"pageType":"Topic","templateName":"DESKTOP","pageNumber":1,"pagesTotal":2,"pageId":342664,"pageLength":1030,"initialLoad":true,"lastPageOfScroll":false} </script> <script class="marketing-content-info" type="application/json"> [] </script> <script src="https://cdn.britannica.com/mendel-resources/3-130/js/libs/jquery-3.5.0.min.js?v=3.130.14"></script> <script type="text/javascript" data-type="Init Mendel Code Splitting"> (function() { $.ajax({ dataType: 'script', cache: true, url: 'https://cdn.britannica.com/mendel-resources/3-130/dist/topic-page.js?v=3.130.14' }); })(); </script> <script class="analytics-metadata" type="application/json"> {"leg":"C","adLeg":"C","userType":"ANONYMOUS","pageType":"Topic","pageSubtype":null,"articleTemplateType":"PAGINATED","gisted":false,"pageNumber":1,"hasSummarizeButton":false,"hasAskButton":false} </script> <script type="text/javascript"> EBStat={accountId:-1,hostnameOverride:'webstats.eb.com',domain:'www.britannica.com', json:''}; </script> <script type="text/javascript"> ( function() { $.ajax( { dataType: 'script', cache: true, url: '//www.britannica.com/webstats/mendelstats.js?v=1' } ) .done( function() { try {writeStat(null,EBStat);} catch(err){} } ); })(); </script> <div id="bc-fixed-dialogue"></div> </body> </html>