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He was fascinated by the different sounds and gestures among the peoples of Tierra del Fuego, and on his return from the voyage he started recording observations in a set of notebooks, later labelled 'Metaphysics on Morals & Speculations on Expression'. Details about the facial and bodily" /> <meta name="generator" content="Drupal 7 (https://www.drupal.org)" /> <link rel="canonical" href="https://www.darwinproject.ac.uk/letters/darwins-works-letters/expression" /> <link rel="shortlink" href="https://www.darwinproject.ac.uk/node/575" /> <meta property="og:site_name" content="Darwin Correspondence Project" /> <meta property="og:type" content="article" /> <meta property="og:url" content="https://www.darwinproject.ac.uk/letters/darwins-works-letters/expression" /> <meta property="og:title" content="Expression" /> <meta property="og:description" content="Darwin's interest in emotional expression can be traced as far back as the Beagle voyage. He was fascinated by the different sounds and gestures among the peoples of Tierra del Fuego, and on his return from the voyage he started recording observations in a set of notebooks, later labelled 'Metaphysics on Morals & Speculations on Expression'. Details about the facial and bodily movements accompanying various emotions were entered, alongside remarks on the origins of language, aesthetic taste, and sympathy, comparing the behaviour of humans and animals." /> <meta property="og:updated_time" content="2022-02-12T19:01:42+00:00" /> <meta property="og:image" content="https://www.darwinproject.ac.uk/sites/default/files/DAR%2053.1%20C125r.jpg" /> <meta name="twitter:card" content="summary" /> <meta name="twitter:url" content="https://www.darwinproject.ac.uk/letters/darwins-works-letters/expression" /> <meta name="twitter:title" content="Expression" /> <meta property="article:published_time" content="2019-02-05T10:33:14+00:00" /> <meta property="article:modified_time" content="2019-02-05T10:33:14+00:00" /> <title>Expression | Darwin Correspondence Project</title> <meta name="viewport" content="width=device-width, initial-scale=1.0"/> <style type="text/css" media="all"> @import 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easy-breadcrumb_segment-title" itemprop="title">Expression</span></li> </ul> </div> </div> </div> </div> <h1 class="campl-page-title">Darwin Correspondence Project</h1> </div> </div> </div> </div> <div class="campl-row campl-page-header"> <div class="campl-wrap"> <div class="region region-horizontal-navigation"> <div id="block-menu-block-1" class="block block-menu-block"> <div> <div class="menu-block-wrapper menu-block-1 menu-name-main-menu parent-mlid-0 menu-level-1"> <div class="campl-wrap clearfix campl-local-navigation"><div class="campl-local-navigation-container"><ul class="campl-unstyled-list"><li class="first leaf menu-mlid-198"><a href="/">Home</a></li> <li class="expanded menu-mlid-2309"><a href="/about-darwin">About Darwin</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2309"><a href="/about-darwin">About Darwin overview</a></li> <li class="expanded menu-mlid-869"><a href="/people/about-darwin/family-life">Family life</a><ul 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href="/about-darwin/origin-species">On the Origin of Species overview</a></li> <li class="leaf menu-mlid-1060"><a href="/people/about-darwin/origin-species/writing-origin">The writing of "Origin"</a></li> <li class="leaf menu-mlid-1084"><a href="/people/about-darwin/origin-species/abstract-darwin-s-theory">Abstract of Darwin’s theory</a></li> <li class="leaf menu-mlid-1093"><a href="/people/about-darwin/origin-species/alfred-russel-wallace-s-essay-varieties">Alfred Russel Wallace’s essay on varieties</a></li> <li class="leaf menu-mlid-1094"><a href="/charles-darwin-and-his-publisher">Charles Darwin and his publisher</a></li> <li class="last leaf menu-mlid-1147"><a href="/people/about-darwin/origin-species/review-origin-species">Review: The Origin of Species</a></li> </ul></li> <li class="leaf menu-mlid-1126"><a href="/people/about-darwin/darwins-health">Darwin’s health</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first last leaf menu-mlid-2917"><a href="/tags/darwin/darwin-on-his-health">Darwin's notes for his physician, 1865</a></li> </ul></li> <li class="leaf menu-mlid-1128"><a href="/about-darwin/darwin-s-photographic-portraits">Darwin’s photographic portraits</a></li> <li class="leaf menu-mlid-2896"><a href="/have-you-read-one-about">Have you read the one about....</a></li> <li class="expanded menu-mlid-2654"><a href="/people/about-darwin/six-things-darwin-never-said">Six things Darwin never said – and one he did</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2654"><a href="/people/about-darwin/six-things-darwin-never-said">Six things Darwin never said – and one he did overview</a></li> <li class="last leaf menu-mlid-2892"><a href="/people/about-darwin/six-things-darwin-never-said/evolution-misquotation">The evolution of a misquotation</a></li> </ul></li> <li class="expanded menu-mlid-3673"><a href="/about-darwin/portraits-charles-darwin-catalogue">Portraits of Charles Darwin: a catalogue</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-3673"><a href="/about-darwin/portraits-charles-darwin-catalogue">Portraits of Charles Darwin: a catalogue overview</a></li> <li class="leaf menu-mlid-3676"><a href="/about-darwin/portraits-charles-darwin-catalogue/11-ellen-sharples-pastel">1.1 Ellen Sharples pastel</a></li> <li class="leaf menu-mlid-3709"><a href="/about-darwin/portraits-charles-darwin-catalogue/12-george-richmond-marriage-portrait">1.2 George Richmond, marriage portrait</a></li> <li class="leaf menu-mlid-3718"><a href="/about-darwin/portraits-charles-darwin-catalogue/13-thomas-herbert-maguire-lithograph">1.3 Thomas Herbert Maguire, lithograph</a></li> <li class="leaf menu-mlid-3721"><a href="/about-darwin/portraits-charles-darwin-catalogue/14-samuel-laurence-drawing-1">1.4 Samuel Laurence drawing 1</a></li> <li class="leaf menu-mlid-3724"><a href="/about-darwin/portraits-charles-darwin-catalogue/15-samuel-laurence-drawing-2">1.5 Samuel Laurence drawing 2</a></li> <li class="leaf menu-mlid-3727"><a href="/about-darwin/portraits-charles-darwin-catalogue/16-ouless-oil-portrait">1.6 Ouless oil portrait</a></li> <li class="leaf menu-mlid-3730"><a href="/about-darwin/portraits-charles-darwin-catalogue/17-ouless-replica">1.7 Ouless replica</a></li> <li class="leaf menu-mlid-3733"><a href="/about-darwin/portraits-charles-darwin-catalogue/18-anonymous-drawing-after-ouless">1.8 anonymous drawing, after Ouless</a></li> <li class="leaf menu-mlid-3736"><a href="/about-darwin/portraits-charles-darwin-catalogue/19-rajon-etching-after-ouless">1.9 Rajon, etching after Ouless</a></li> <li class="leaf menu-mlid-3679"><a href="/about-darwin/portraits-charles-darwin-catalogue/110-rajon-etching-variant-state">1.10 Rajon etching, variant state</a></li> <li class="leaf menu-mlid-3682"><a href="/about-darwin/portraits-charles-darwin-catalogue/111-laura-russell-oil">1.11 Laura Russell, oil</a></li> <li class="leaf menu-mlid-3685"><a href="/about-darwin/portraits-charles-darwin-catalogue/112-marian-huxley-drawing">1.12 Marian Huxley, drawing</a></li> <li class="leaf menu-mlid-3688"><a href="/about-darwin/portraits-charles-darwin-catalogue/113-louisa-nash-drawing">1.13 Louisa Nash, drawing</a></li> <li class="leaf menu-mlid-3691"><a href="/about-darwin/portraits-charles-darwin-catalogue/114-william-richmond-oil">1.14 William Richmond, oil</a></li> <li class="leaf menu-mlid-3694"><a href="/about-darwin/portraits-charles-darwin-catalogue/115-albert-goodwin-watercolour">1.15 Albert Goodwin, watercolour</a></li> <li class="leaf menu-mlid-3697"><a href="/about-darwin/portraits-charles-darwin-catalogue/116-alphonse-legros-drypoint">1.16 Alphonse Legros, drypoint</a></li> <li class="leaf menu-mlid-3700"><a href="/about-darwin/portraits-charles-darwin-catalogue/117-alphonse-legros-drawing">1.17 Alphonse Legros drawing</a></li> <li class="leaf menu-mlid-3703"><a href="/about-darwin/portraits-charles-darwin-catalogue/118-john-collier-oil-linnean">1.18 John Collier, oil in Linnean</a></li> <li class="leaf menu-mlid-3706"><a href="/about-darwin/portraits-charles-darwin-catalogue/119-john-collier-oil-npg">1.19 John Collier, oil in NPG</a></li> <li class="leaf menu-mlid-3712"><a href="/about-darwin/portraits-charles-darwin-catalogue/120-leopold-flameng-etching-after-collier">1.20 Leopold Flameng etching, after Collier</a></li> <li class="leaf menu-mlid-3715"><a href="/about-darwin/portraits-charles-darwin-catalogue/121-window-christs-college-cambridge">1.21 window at Christ's College Cambridge</a></li> <li class="leaf menu-mlid-3739"><a href="/about-darwin/portraits-charles-darwin-catalogue/21-thomas-woolner-bust">2.1 Thomas Woolner bust</a></li> <li class="leaf menu-mlid-3772"><a href="/about-darwin/portraits-charles-darwin-catalogue/22-thomas-woolner-metal-plaque">2.2 Thomas Woolner metal plaque</a></li> <li class="leaf menu-mlid-3802"><a href="/about-darwin/portraits-charles-darwin-catalogue/23-wedgwood-medallions">2.3 Wedgwood medallions</a></li> <li class="leaf menu-mlid-3805"><a href="/about-darwin/portraits-charles-darwin-catalogue/24-wedgwood-plaque">2.4 Wedgwood plaque</a></li> <li class="leaf menu-mlid-3808"><a href="/about-darwin/portraits-charles-darwin-catalogue/25-wedgwood-medallions-2nd-type">2.5 Wedgwood medallions, 2nd type</a></li> <li class="leaf menu-mlid-3811"><a href="/about-darwin/portraits-charles-darwin-catalogue/26-adolf-von-hildebrand-bust">2.6 Adolf von Hildebrand bust</a></li> <li class="leaf menu-mlid-3814"><a href="/about-darwin/portraits-charles-darwin-catalogue/27-joseph-moore-midland-union-medal">2.7 Joseph Moore, Midland Union medal</a></li> <li class="leaf menu-mlid-3817"><a href="/about-darwin/portraits-charles-darwin-catalogue/28-alphonse-legros-medallion">2.8 Alphonse Legros medallion</a></li> <li class="leaf menu-mlid-3820"><a href="/about-darwin/portraits-charles-darwin-catalogue/29-legros-medallion-plaster-model">2.9 Legros medallion, plaster model</a></li> <li class="leaf menu-mlid-3742"><a href="/about-darwin/portraits-charles-darwin-catalogue/210-moritz-klinkicht-print-legros">2.10 Moritz Klinkicht, print from Legros</a></li> <li class="leaf menu-mlid-3745"><a href="/about-darwin/portraits-charles-darwin-catalogue/211-christian-lehr-plaster-bust">2.11 Christian Lehr, plaster bust</a></li> <li class="leaf menu-mlid-3748"><a href="/about-darwin/portraits-charles-darwin-catalogue/212-allan-wyon-royal-society-medal">2.12 Allan Wyon, Royal Society medal</a></li> <li class="leaf menu-mlid-3751"><a href="/about-darwin/portraits-charles-darwin-catalogue/213-edgar-boehm-statue-nhm">2.13 Edgar Boehm, statue in the NHM</a></li> <li class="leaf menu-mlid-3754"><a href="/about-darwin/portraits-charles-darwin-catalogue/214-boehm-westminster-abbey-roundel">2.14 Boehm, Westminster Abbey roundel</a></li> <li class="leaf menu-mlid-3757"><a href="/about-darwin/portraits-charles-darwin-catalogue/215-boehm-terracotta-bust-npg">2.15 Boehm terracotta bust (NPG)</a></li> <li class="leaf menu-mlid-3760"><a href="/about-darwin/portraits-charles-darwin-catalogue/216-horace-montford-statue-shrewsbury">2.16 Horace Montford statue, Shrewsbury</a></li> <li class="leaf menu-mlid-3763"><a href="/about-darwin/portraits-charles-darwin-catalogue/217-montford-statuette">2.17 Montford, statuette</a></li> <li class="leaf menu-mlid-3766"><a href="/about-darwin/portraits-charles-darwin-catalogue/218-montford-carnegie-bust">2.18 Montford, Carnegie bust</a></li> <li class="leaf menu-mlid-3769"><a href="/about-darwin/portraits-charles-darwin-catalogue/219-montford-bust-royal-society">2.19 Montford, bust at the Royal Society</a></li> <li class="leaf menu-mlid-3775"><a href="/about-darwin/portraits-charles-darwin-catalogue/220-montford-terracotta-bust-npg">2.20 Montford, terracotta bust, NPG</a></li> <li class="leaf menu-mlid-3778"><a href="/about-darwin/portraits-charles-darwin-catalogue/221-montford-relief-christs-college">2.21 Montford, relief at Christ's College</a></li> <li class="leaf menu-mlid-3781"><a href="/about-darwin/portraits-charles-darwin-catalogue/222-l-j-chavalliaud-statue-liverpool">2.22 L.-J. Chavalliaud statue in Liverpool</a></li> <li class="leaf menu-mlid-3784"><a href="/about-darwin/portraits-charles-darwin-catalogue/223-hope-pinker-statue-oxford-museum">2.23 Hope Pinker statue, Oxford Museum</a></li> <li class="leaf menu-mlid-3787"><a href="/about-darwin/portraits-charles-darwin-catalogue/224-herbert-hampton-statue-lancaster">2.24 Herbert Hampton statue, Lancaster</a></li> <li class="leaf menu-mlid-3790"><a href="/about-darwin/portraits-charles-darwin-catalogue/225-henry-pegram-statue-birmingham">2.25 Henry Pegram statue, Birmingham</a></li> <li class="leaf menu-mlid-3793"><a href="/about-darwin/portraits-charles-darwin-catalogue/226-linnean-society-medal">2.26 Linnean Society medal</a></li> <li class="leaf menu-mlid-3796"><a href="/about-darwin/portraits-charles-darwin-catalogue/227-william-couper-bust-new-york">2.27 William Couper bust, New York</a></li> <li class="leaf menu-mlid-3799"><a href="/about-darwin/portraits-charles-darwin-catalogue/228-couper-bust-cambridge">2.28 Couper bust in Cambridge</a></li> <li class="leaf menu-mlid-3823"><a href="/about-darwin/portraits-charles-darwin-catalogue/31-antoine-claudet-daguerreotype">3.1 Antoine Claudet, daguerreotype</a></li> <li class="leaf menu-mlid-3856"><a href="/about-darwin/portraits-charles-darwin-catalogue/32-maull-and-polyblank-photo-1">3.2 Maull and Polyblank photo 1</a></li> <li class="leaf menu-mlid-3865"><a href="/about-darwin/portraits-charles-darwin-catalogue/33-maull-and-polyblank-photo-2">3.3 Maull and Polyblank photo 2</a></li> <li class="leaf menu-mlid-3868"><a href="/about-darwin/portraits-charles-darwin-catalogue/34-william-darwin-photo-1">3.4 William Darwin, photo 1</a></li> <li class="leaf menu-mlid-3871"><a href="/about-darwin/portraits-charles-darwin-catalogue/35-william-darwin-photo-2">3.5 William Darwin, photo 2</a></li> <li class="leaf menu-mlid-3874"><a href="/about-darwin/portraits-charles-darwin-catalogue/36-william-darwin-photo-3">3.6 William Darwin, photo 3</a></li> <li class="leaf menu-mlid-3877"><a href="/about-darwin/portraits-charles-darwin-catalogue/37-leonard-darwin-photo-verandah">3.7 Leonard Darwin, photo on verandah</a></li> <li class="leaf menu-mlid-3880"><a href="/about-darwin/portraits-charles-darwin-catalogue/38-leonard-darwin-interior-photo">3.8 Leonard Darwin, interior photo</a></li> <li class="leaf menu-mlid-3883"><a href="/about-darwin/portraits-charles-darwin-catalogue/39-leonard-darwin-photo-horseback">3.9 Leonard Darwin, photo on horseback</a></li> <li class="leaf menu-mlid-3826"><a href="/about-darwin/portraits-charles-darwin-catalogue/310-ernest-edwards-men-eminence">3.10 Ernest Edwards, 'Men of Eminence'</a></li> <li class="leaf menu-mlid-3829"><a href="/about-darwin/portraits-charles-darwin-catalogue/311-edwards-illustrated-london-news">3.11 Edwards, in Illustrated London News</a></li> <li class="leaf menu-mlid-3832"><a href="/about-darwin/portraits-charles-darwin-catalogue/312-edwards-second-group-photos">3.12 Edwards, second group of photos</a></li> <li class="leaf menu-mlid-3835"><a href="/about-darwin/portraits-charles-darwin-catalogue/313-edwards-representative-men">3.13 Edwards 'Representative Men'</a></li> <li class="leaf menu-mlid-3838"><a href="/about-darwin/portraits-charles-darwin-catalogue/314-julia-margaret-cameron-photos">3.14 Julia Margaret Cameron, photos</a></li> <li class="leaf menu-mlid-3841"><a href="/about-darwin/portraits-charles-darwin-catalogue/315-george-charles-wallich-photo">3.15 George Charles Wallich, photo</a></li> <li class="leaf menu-mlid-3844"><a href="/about-darwin/portraits-charles-darwin-catalogue/316-oscar-rejlander-photos">3.16 Oscar Rejlander, photos</a></li> <li class="leaf menu-mlid-3847"><a href="/about-darwin/portraits-charles-darwin-catalogue/317-lock-and-whitfield-men-mark">3.17 Lock and Whitfield, 'Men of Mark'</a></li> <li class="leaf menu-mlid-3850"><a href="/about-darwin/portraits-charles-darwin-catalogue/318-elliott-and-fry-photos-c1869-1871">3.18 Elliott and Fry photos, c.1869-1871</a></li> <li class="leaf menu-mlid-3853"><a href="/about-darwin/portraits-charles-darwin-catalogue/319-elliott-and-fry-photos-c1880-1">3.19 Elliott and Fry photos c.1880-1</a></li> <li class="leaf menu-mlid-3859"><a href="/about-darwin/portraits-charles-darwin-catalogue/320-elliott-and-fry-c1880-1-verandah">3.20 Elliott and Fry, c.1880-1, verandah</a></li> <li class="leaf menu-mlid-3862"><a href="/about-darwin/portraits-charles-darwin-catalogue/321-herbert-rose-barraud-photos">3.21 Herbert Rose Barraud, photos</a></li> <li class="leaf menu-mlid-3886"><a href="/about-darwin/portraits-charles-darwin-catalogue/41-albert-way-comic-drawings">4.1 Albert Way, comic drawings</a></li> <li class="leaf menu-mlid-3919"><a href="/about-darwin/portraits-charles-darwin-catalogue/42-augustus-earle-caricature-drawing">4.2 Augustus Earle, caricature drawing</a></li> <li class="leaf menu-mlid-3952"><a href="/about-darwin/portraits-charles-darwin-catalogue/43-alfred-crowquill-caricature">4.3 Alfred Crowquill, caricature</a></li> <li class="leaf menu-mlid-3985"><a href="/about-darwin/portraits-charles-darwin-catalogue/44-thomas-huxley-caricature-sketch">4.4 Thomas Huxley, caricature sketch</a></li> <li class="leaf menu-mlid-4018"><a href="/about-darwin/portraits-charles-darwin-catalogue/45-william-beard-comic-painting">4.5 William Beard, comic painting</a></li> <li class="leaf menu-mlid-4051"><a href="/about-darwin/portraits-charles-darwin-catalogue/46-thomas-nast-cartoon">4.6 Thomas Nast, cartoon</a></li> <li class="leaf menu-mlid-4054"><a href="/about-darwin/portraits-charles-darwin-catalogue/47-vanity-fair-caricature">4.7 'Vanity Fair', caricature</a></li> <li class="leaf menu-mlid-4057"><a href="/about-darwin/portraits-charles-darwin-catalogue/48-vanity-fair-preliminary-study">4.8 'Vanity Fair', preliminary study</a></li> <li class="leaf menu-mlid-4060"><a href="/about-darwin/portraits-charles-darwin-catalogue/49-graphic-cartoon">4.9 'Graphic', cartoon</a></li> <li class="leaf menu-mlid-3889"><a href="/about-darwin/portraits-charles-darwin-catalogue/410-hornet-caricature-darwin">4.10 'Hornet' caricature of Darwin</a></li> <li class="leaf menu-mlid-3892"><a href="/about-darwin/portraits-charles-darwin-catalogue/411-fun-cartoon-little-lecture">4.11 'Fun' cartoon, 'A little lecture'</a></li> <li class="leaf menu-mlid-3895"><a href="/about-darwin/portraits-charles-darwin-catalogue/412-fun-wedding-procession">4.12 'Fun', Wedding procession</a></li> <li class="leaf menu-mlid-3898"><a href="/about-darwin/portraits-charles-darwin-catalogue/413-fun-cartoon-griset-emotional">4.13 'Fun' cartoon by Griset, 'Emotional'</a></li> <li class="leaf menu-mlid-3901"><a href="/about-darwin/portraits-charles-darwin-catalogue/414-fun-cartoon-troubles">4.14 'Fun' cartoon, 'That troubles'</a></li> <li class="leaf menu-mlid-3904"><a href="/about-darwin/portraits-charles-darwin-catalogue/415-george-cruikshank-comic-drawing">4.15 George Cruikshank, comic drawing</a></li> <li class="leaf menu-mlid-3907"><a href="/about-darwin/portraits-charles-darwin-catalogue/416-joseph-simms-physiognomy">4.16 Joseph Simms, physiognomy</a></li> <li class="leaf menu-mlid-3910"><a href="/about-darwin/portraits-charles-darwin-catalogue/417-figaro-unidentifiable-1871">4.17 'Figaro', unidentifiable 1871</a></li> <li class="leaf menu-mlid-3913"><a href="/about-darwin/portraits-charles-darwin-catalogue/418-figaro-chromolithograph-1">4.18 'Figaro' chromolithograph 1</a></li> <li class="leaf menu-mlid-3916"><a href="/about-darwin/portraits-charles-darwin-catalogue/419-george-montbard-caricature">4.19 George Montbard, caricature</a></li> <li class="leaf menu-mlid-3922"><a href="/about-darwin/portraits-charles-darwin-catalogue/420-frederick-waddy-caricature">4.20 Frederick Waddy, caricature</a></li> <li class="leaf menu-mlid-3925"><a href="/about-darwin/portraits-charles-darwin-catalogue/421-gegeef-our-national-church-1">4.21 Gegeef, 'Our National Church', 1</a></li> <li class="leaf menu-mlid-3928"><a href="/about-darwin/portraits-charles-darwin-catalogue/422-gegeef-et-al-our-national-church-2">4.22 Gegeef et al., 'Our National Church', 2</a></li> <li class="leaf menu-mlid-3931"><a href="/about-darwin/portraits-charles-darwin-catalogue/423-gegeef-battle-field-science">4.23 Gegeef, 'Battle Field of Science'</a></li> <li class="leaf menu-mlid-3934"><a href="/about-darwin/portraits-charles-darwin-catalogue/424-daily-graphic-nast-satire">4.24 'Daily Graphic', Nast satire</a></li> <li class="leaf menu-mlid-3937"><a href="/about-darwin/portraits-charles-darwin-catalogue/425-punch-1877-re-cambridge-doctorate">4.25 'Punch' 1877 re. Cambridge doctorate</a></li> <li class="leaf menu-mlid-3940"><a href="/about-darwin/portraits-charles-darwin-catalogue/426-christmas-card-caricature-monkeys">4.26 Christmas card caricature, monkeys</a></li> <li class="leaf menu-mlid-3943"><a href="/about-darwin/portraits-charles-darwin-catalogue/427-four-founders-darwinismus">4.27 'Four founders of Darwinismus'</a></li> <li class="leaf menu-mlid-3946"><a href="/about-darwin/portraits-charles-darwin-catalogue/428-english-celebrities-montage">4.28 'English celebrities' montage</a></li> <li class="leaf menu-mlid-3949"><a href="/about-darwin/portraits-charles-darwin-catalogue/429-richard-grant-white-fall-man">4.29 Richard Grant White, 'Fall of man'</a></li> <li class="leaf menu-mlid-3955"><a href="/about-darwin/portraits-charles-darwin-catalogue/430-la-petite-lune-gill-cartoon">4.30 'La Petite Lune', Gill cartoon</a></li> <li class="leaf menu-mlid-3958"><a href="/about-darwin/portraits-charles-darwin-catalogue/431-la-lune-rousse-gill-cartoon">4.31 'La Lune Rousse', Gill cartoon</a></li> <li class="leaf menu-mlid-3961"><a href="/about-darwin/portraits-charles-darwin-catalogue/432-anis-liqueur-label">4.32 Anis liqueur label</a></li> <li class="leaf menu-mlid-3964"><a href="/about-darwin/portraits-charles-darwin-catalogue/433-harpers-weekly-bellew-caricature">4.33 'Harper's Weekly', Bellew caricature</a></li> <li class="leaf menu-mlid-3967"><a href="/about-darwin/portraits-charles-darwin-catalogue/434-punch-sambourne-cartoon-1">4.34 'Punch', Sambourne cartoon 1</a></li> <li class="leaf menu-mlid-3970"><a href="/about-darwin/portraits-charles-darwin-catalogue/435-frederick-sem-caricature">4.35 Frederick Sem, caricature</a></li> <li class="leaf menu-mlid-3973"><a href="/about-darwin/portraits-charles-darwin-catalogue/436-sem-chistmas-card">4.36 Sem, Chistmas card</a></li> <li class="leaf menu-mlid-3976"><a href="/about-darwin/portraits-charles-darwin-catalogue/437-mosquito-satire">4.37 'Mosquito' satire</a></li> <li class="leaf menu-mlid-3979"><a href="/about-darwin/portraits-charles-darwin-catalogue/438-franz-goedecker-caricature">4.38 Franz Goedecker, caricature</a></li> <li class="leaf menu-mlid-3982"><a href="/about-darwin/portraits-charles-darwin-catalogue/439-moonshine-magazine-cartoon">4.39 'Moonshine' magazine cartoon</a></li> <li class="leaf menu-mlid-3988"><a href="/about-darwin/portraits-charles-darwin-catalogue/440-phrenological-magazine">4.40 'Phrenological Magazine'</a></li> <li class="leaf menu-mlid-3991"><a href="/about-darwin/portraits-charles-darwin-catalogue/441-punch-sambourne-cartoon-2">4.41 'Punch', Sambourne cartoon 2</a></li> <li class="leaf menu-mlid-3994"><a href="/about-darwin/portraits-charles-darwin-catalogue/442-punch-sambourne-cartoon-3">4.42 'Punch' Sambourne cartoon 3</a></li> <li class="leaf menu-mlid-3997"><a href="/about-darwin/portraits-charles-darwin-catalogue/443-illustrated-london-news-article">4.43 'Illustrated London News' article</a></li> <li class="leaf menu-mlid-4000"><a href="/about-darwin/portraits-charles-darwin-catalogue/444-puck-cartoon-1">4.44 'Puck' cartoon 1</a></li> <li class="leaf menu-mlid-4003"><a href="/about-darwin/portraits-charles-darwin-catalogue/445-puck-cartoon-2">4.45 'Puck' cartoon 2</a></li> <li class="leaf menu-mlid-4006"><a href="/about-darwin/portraits-charles-darwin-catalogue/446-puck-cartoon-3">4.46 'Puck' cartoon 3</a></li> <li class="leaf menu-mlid-4009"><a href="/about-darwin/portraits-charles-darwin-catalogue/447-puck-cartoon-4">4.47 'Puck' cartoon 4</a></li> <li class="leaf menu-mlid-4012"><a href="/about-darwin/portraits-charles-darwin-catalogue/448-puck-cartoon-5">4.48 'Puck', cartoon 5</a></li> <li class="leaf menu-mlid-4015"><a href="/about-darwin/portraits-charles-darwin-catalogue/449-alfred-bryan-caricature">4.49 Alfred Bryan, caricature</a></li> <li class="leaf menu-mlid-4021"><a href="/about-darwin/portraits-charles-darwin-catalogue/450-cigar-box-lid-design">4.50 Cigar box lid design</a></li> <li class="leaf menu-mlid-4024"><a href="/about-darwin/portraits-charles-darwin-catalogue/451-frederick-holder-life-and-work">4.51 Frederick Holder 'Life and Work'</a></li> <li class="leaf menu-mlid-4027"><a href="/about-darwin/portraits-charles-darwin-catalogue/452-wasp-caricature">4.52 'Wasp' caricature</a></li> <li class="leaf menu-mlid-4030"><a href="/about-darwin/portraits-charles-darwin-catalogue/453-claud-warren-outlines-hands">4.53 Claud Warren, 'Outlines of Hands'</a></li> <li class="leaf menu-mlid-4033"><a href="/about-darwin/portraits-charles-darwin-catalogue/454-jubilees-queen-victoria">4.54 jubilees of Queen Victoria</a></li> <li class="leaf menu-mlid-4036"><a href="/about-darwin/portraits-charles-darwin-catalogue/455-harry-furniss-caricature">4.55 Harry Furniss caricature</a></li> <li class="leaf menu-mlid-4039"><a href="/about-darwin/portraits-charles-darwin-catalogue/456-larks-cartoon">4.56 'Larks' cartoon</a></li> <li class="leaf menu-mlid-4042"><a href="/about-darwin/portraits-charles-darwin-catalogue/457-silhouette-cartoon">4.57 silhouette cartoon</a></li> <li class="leaf menu-mlid-4045"><a href="/about-darwin/portraits-charles-darwin-catalogue/458-simian-savage-drawings">4.58 'Simian, savage' . . . drawings</a></li> <li class="last leaf menu-mlid-4048"><a href="/about-darwin/portraits-charles-darwin-catalogue/459-simplicissimus-cartoon">4.59 'Simplicissimus' cartoon</a></li> </ul></li> <li class="expanded menu-mlid-4066"><a href="/about-darwin/darwin-and-experimental-life">Darwin and the experimental life</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-4066"><a href="/about-darwin/darwin-and-experimental-life">Darwin and the experimental life overview</a></li> <li class="leaf menu-mlid-4069"><a href="/about-darwin/darwin-and-experimental-life/what-experiment">What is an experiment?</a></li> <li class="leaf menu-mlid-4081"><a href="/about-darwin/darwin-and-experimental-life/morphology-movement-observation-and-experiment">From morphology to movement: observation and experiment</a></li> <li class="leaf menu-mlid-4078"><a href="/about-darwin/darwin-and-experimental-life/fools-experiments">Fool's experiments</a></li> <li class="leaf menu-mlid-4075"><a href="/about-darwin/darwin-and-experimental-life/experimenting-emotions">Experimenting with emotions</a></li> <li class="last leaf menu-mlid-4072"><a href="/about-darwin/darwin-and-experimental-life/animals-ethics-and-progress-science">Animals, ethics, and the progress of science</a></li> </ul></li> <li class="leaf menu-mlid-3366"><a href="/fake-darwin">Fake Darwin: myths and misconceptions</a></li> <li class="leaf menu-mlid-3402"><a href="/darwins-bad-days">Darwin's bad days</a></li> <li class="last leaf menu-mlid-3644"><a href="/people/about-darwin/darwin-s-first-love">Darwin’s first love</a></li> </ul></li> <li class="expanded active-trail menu-mlid-800"><a href="/letters" title="" class="active-trail campl-selected">The letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed active-trail menu-mlid-800"><a href="/letters" title="" class="active-trail">The letters overview</a></li> <li class="expanded menu-mlid-2080"><a href="/letters/darwins-life-letters">Darwin's life in letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2080"><a href="/letters/darwins-life-letters">Darwin's life in letters overview</a></li> <li class="leaf menu-mlid-1097"><a href="/letters/darwins-life-letters/darwin-letters-1821-1836-childhood-beagle-voyage">1821-1836: Childhood to the Beagle voyage</a></li> <li class="leaf menu-mlid-1106"><a href="/letters/darwins-life-letters/darwin-letters-1837-1843-london-years-natural-selection" title="Charles Darwin's life seen through his letters, 1837-43">1837-43: The London years to 'natural selection'</a></li> <li class="leaf menu-mlid-1107"><a href="/letters/darwins-life-letters/darwin-letters-1844-1846-building-scientific-network">1844-1846: Building a scientific network</a></li> <li class="leaf menu-mlid-1108"><a href="/letters/darwins-life-letters/darwin-letters-1847-1850-microscopes-and-barnacles">1847-1850: Microscopes and barnacles</a></li> <li class="leaf menu-mlid-1109"><a href="/letters/darwins-life-letters/darwin-letters-1851-1855-death-daughter">1851-1855: Death of a daughter</a></li> <li class="leaf menu-mlid-1110"><a href="/letters/darwins-life-letters/darwin-letters-1856-1857-big-book">1856-1857: The 'Big Book'</a></li> <li class="leaf menu-mlid-936"><a href="/letters/darwins-life-letters/darwin-letters-1858-1859-origin">1858-1859: Origin</a></li> <li class="leaf menu-mlid-937"><a href="/letters/darwins-life-letters/darwin-letters-1860-answering-critics">1860: Answering critics</a></li> <li class="leaf menu-mlid-1111"><a href="/letters/darwins-life-letters/darwin-letters-1861-gaining-allies">1861: Gaining allies</a></li> <li class="leaf menu-mlid-1098"><a href="/letters/darwins-life-letters/darwin-letters-1862-multiplicity-experiments">1862: A multiplicity of experiments</a></li> <li class="leaf menu-mlid-1099"><a href="/letters/darwins-life-letters/darwin-letters-1863-quarrels-home-honours-abroad">1863: Quarrels at home, honours abroad</a></li> <li class="leaf menu-mlid-1100"><a href="/letters/darwins-life-letters/darwin-letters-1864-failing-health">1864: Failing health</a></li> <li class="leaf menu-mlid-935"><a href="/letters/darwins-life-letters/darwin-letters-1865-delays-and-disappointments">1865: Delays and disappointments</a></li> <li class="leaf menu-mlid-1101"><a href="/letters/darwins-life-letters/darwin-letters1866-survival-fittest">1866: Survival of the fittest</a></li> <li class="leaf menu-mlid-1102"><a href="/letters/darwins-life-letters/darwin-letters-1867-civilised-dispute">1867: A civilised dispute</a></li> <li class="leaf menu-mlid-1103"><a href="/letters/darwins-life-letters/darwin-letters-1868-studying-sex">1868: Studying sex</a></li> <li class="leaf menu-mlid-1104"><a href="/letters/darwins-life-letters/darwin-letters-1869-forward-all-fronts">1869: Forward on all fronts</a></li> <li class="leaf menu-mlid-1105"><a href="/letters/darwins-life-letters/darwin-letters1870-human-evolution">1870: Human evolution</a></li> <li class="leaf menu-mlid-1151"><a href="/letters/darwins-life-letters/darwin-letters-1871-emptying-nest">1871: An emptying nest</a></li> <li class="leaf menu-mlid-1152"><a href="/letters/darwins-life-letters/darwin-letters-1872-job-done">1872: Job done?</a></li> <li class="leaf menu-mlid-1049"><a href="/letters/darwins-life-letters/darwins-letters-1873-animal-or-vegetable">1873: Animal or vegetable?</a></li> <li class="leaf menu-mlid-1050"><a href="/letters/darwins-life-letters/darwin-letters-1874-turbulent-year">1874: A turbulent year</a></li> <li class="leaf menu-mlid-2275"><a href="/letters/darwins-life-letters/darwin-letters-1875-pulling-strings">1875: Pulling strings</a></li> <li class="leaf menu-mlid-2894"><a href="/letters/darwins-life-letters/darwin-letters-1876-midst-life">1876: In the midst of life</a></li> <li class="leaf menu-mlid-3363"><a href="/letters/darwins-life-letters/darwin-letters-1877-flowers-and-honours">1877: Flowers and honours</a></li> <li class="leaf menu-mlid-3383"><a href="/letters/darwins-life-letters/darwin-letters-1878-movement-and-sleep">1878: Movement and sleep</a></li> <li class="leaf menu-mlid-3394"><a href="/letters/darwins-life-letters/darwin-letters-1879-tracing-roots">1879: Tracing roots</a></li> <li class="leaf menu-mlid-3661"><a href="/letters/darwins-life-letters/darwin-letters-1880-sensitivity-and-worms">1880: Sensitivity and worms</a></li> <li class="leaf menu-mlid-4063"><a href="/letters/darwins-life-letters/darwin-letters-1881-old-friends-and-new-admirers">1881: Old friends and new admirers</a></li> <li class="last leaf menu-mlid-4102"><a href="/letters/darwins-life-letters/darwin-letters-1882-nothing-too-great-or-too-small">1882: Nothing too great or too small</a></li> </ul></li> <li class="expanded active-trail menu-mlid-3384"><a href="/letters/darwins-works-letters" class="active-trail">Darwin's works in letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed active-trail menu-mlid-3384"><a href="/letters/darwins-works-letters" class="active-trail">Darwin's works in letters overview</a></li> <li class="leaf menu-mlid-3385"><a href="/letters/darwins-works-letters/journal-researches">Journal of researches</a></li> <li class="leaf menu-mlid-3386"><a href="/letters/darwins-works-letters/living-and-fossil-cirripedia">Living and fossil cirripedia</a></li> <li class="expanded menu-mlid-3400"><a href="/letters/darwins-works-letters/origin-big-book">Before Origin: the ‘big book’</a><ul class="menu"><li class="first collapsed menu-mlid-3400"><a href="/letters/darwins-works-letters/origin-big-book">Before Origin: the ‘big book’ overview</a></li> <li class="last leaf menu-mlid-3401"><a href="/letters/darwins-works-letters/origin-big-book/dates-composition-darwins-manuscript-species">Dates of composition of Darwin's manuscript on species</a></li> </ul></li> <li class="leaf menu-mlid-3406"><a href="/letters/darwins-works-letters/origin">Origin</a></li> <li class="expanded menu-mlid-3407"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions">Rewriting Origin - the later editions</a><ul class="menu"><li class="first collapsed menu-mlid-3407"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions">Rewriting Origin - the later editions overview</a></li> <li class="leaf menu-mlid-3412"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions/how-old-earth">How old is the earth?</a></li> <li class="leaf menu-mlid-3411"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions/whale-bear">The whale-bear</a></li> <li class="last leaf menu-mlid-3413"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions/origin-lost-changes-second-german">Origin: the lost changes for the second German edition</a></li> </ul></li> <li class="leaf menu-mlid-3404"><a href="/letters/darwins-works-letters/orchids">Orchids</a></li> <li class="leaf menu-mlid-3662"><a href="/letters/darwins-works-letters/climbing-plants">Climbing plants</a></li> <li class="leaf menu-mlid-3387"><a href="/letters/darwins-works-letters/descent">Descent</a></li> <li class="leaf active-trail active menu-mlid-3388 campl-current-page"><a href="/letters/darwins-works-letters/expression" class="active-trail active">Expression</a></li> <li class="leaf menu-mlid-3389"><a href="/letters/darwins-works-letters/insectivorous-plants">Insectivorous plants</a></li> <li class="leaf menu-mlid-3663"><a href="/letters/darwins-works-letters/forms-flowers">Forms of flowers</a></li> <li class="leaf menu-mlid-4084"><a href="/letters/darwins-works-letters/cross-and-self-fertilisation">Cross and self fertilisation</a></li> <li class="leaf menu-mlid-3405"><a href="/letters/darwins-works-letters/life-erasmus-darwin">Life of Erasmus Darwin</a></li> <li class="last leaf menu-mlid-3665"><a href="/letters/darwins-works-letters/movement-plants">Movement in Plants</a></li> </ul></li> <li class="leaf menu-mlid-2264"><a href="/letters/about-letters">About the letters</a></li> <li class="expanded menu-mlid-2408"><a href="/letters/lifecycle-letter-film">Lifecycle of a letter film</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2408"><a href="/letters/lifecycle-letter-film">Lifecycle of a letter film overview</a></li> <li class="leaf menu-mlid-2411"><a href="/letters/lifecycle-letter-film/editing-letter">Editing a Letter</a></li> <li class="last leaf menu-mlid-2409"><a href="/letters/lifecycle-letter-film/working-darwin-archive">Working in the Darwin archive</a></li> </ul></li> <li class="leaf menu-mlid-2296"><a href="/letters/capturing-darwin-s-voice-audio-selected-letters">Capturing Darwin’s voice: audio of selected letters</a></li> <li class="leaf menu-mlid-2266"><a href="/letters/correspondence-women">Correspondence with women</a></li> <li class="leaf menu-mlid-878"><a href="/hunt-new-letters">The hunt for new letters</a></li> <li class="expanded menu-mlid-1136"><a href="/letters/editorial-policy-and-practice">Editorial policy and practice</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1136"><a href="/letters/editorial-policy-and-practice">Editorial policy and practice overview</a></li> <li class="last leaf menu-mlid-3646"><a href="/letters/editorial-policy-and-practice/full-notes-editorial-policy">Full notes on editorial policy</a></li> </ul></li> <li class="leaf menu-mlid-2263"><a href="/letters/symbols-and-abbreviations">Symbols and abbreviations</a></li> <li class="leaf menu-mlid-2905"><a href="/letters/darwins-letters-timeline">Darwin's letters: a timeline</a></li> <li class="leaf menu-mlid-2909"><a href="/world-map">Darwin's letters: World Map</a></li> <li class="leaf menu-mlid-3364"><a href="https://www.darwinproject.ac.uk/have-you-read-one-about" title="">Have you read the one about...</a></li> <li class="leaf menu-mlid-4183"><a href="/letters/charles-darwin-life-letters">Charles Darwin: A Life in Letters</a></li> <li class="leaf menu-mlid-4158"><a href="/darwin-conversation-exhibition">Darwin in Conversation exhibition</a></li> <li class="leaf menu-mlid-4099"><a href="/letters/diagrams-and-drawings-letters">Diagrams and drawings in letters</a></li> <li class="expanded menu-mlid-4105"><a href="/letters/favourite-letters">Favourite Letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-4105"><a href="/letters/favourite-letters">Favourite Letters overview</a></li> <li class="leaf menu-mlid-4162"><a href="/letters/favourite-letters/be-envious-ripe-oranges-w-d-fox-may-1832">Be envious of ripe oranges: To W. D. Fox, May 1832</a></li> <li class="leaf menu-mlid-4111"><a href="/letters/favourite-letters/monstrous-stain-j-m-herbert-2-june-1833">That monstrous stain: To J. M. Herbert, 2 June 1833</a></li> <li class="leaf menu-mlid-4166"><a href="/letters/favourite-letters/my-most-solemn-request-emma-darwin-5-july-1844">My most solemn request: To Emma Darwin, 5 July 1844</a></li> <li class="leaf menu-mlid-4178"><a href="/letters/favourite-letters/our-poor-dear-dear-child-emma-darwin-23-april-1851">Our poor dear dear child: To Emma Darwin, [23 April 1851]</a></li> <li class="leaf menu-mlid-4174"><a href="/letters/favourite-letters/i-beg-million-pardons-john-lubbock-3-september-1862">I beg a million pardons: To John Lubbock, [3 September 1862]</a></li> <li class="leaf menu-mlid-4129"><a href="/letters/favourite-letters/prize-possessions-henry-denny-17-january-1865">Prize possessions: To Henry Denny, 17 January [1865]</a></li> <li class="leaf menu-mlid-4114"><a href="/letters/favourite-letters/how-manage-it-j-d-hooker-17-june-1865">How to manage it: To J. D. Hooker, [17 June 1865]</a></li> <li class="leaf menu-mlid-4117"><a href="/letters/favourite-letters/fly-flower-hermann-m-ller-23-october-1867">A fly on the flower: From Hermann Müller, 23 October 1867</a></li> <li class="leaf menu-mlid-4138"><a href="/letters/favourite-letters/reading-my-roommate-s-illustrious-ancestor-t-h-huxley-10-june-1868">Reading my roommate’s illustrious ancestor: To T. H. Huxley, 10 June 1868</a></li> <li class="leaf menu-mlid-4169"><a href="/letters/favourite-letters/beginning-something-j-d-hooker-22-january-1869">A beginning, & that is something: To J. D. Hooker, [22 January 1869]</a></li> <li class="leaf menu-mlid-4135"><a href="/letters/favourite-letters/perfect-copper-plate-hand-adolf-reuter-30-may-1869">Perfect copper-plate hand: From Adolf Reuter, 30 May 1869</a></li> <li class="leaf menu-mlid-4132"><a href="/letters/favourite-letters/darwin-s-favourite-photographer-o-g-rejlander-30-april-1871">Darwin’s favourite photographer: From O. G. Rejlander, 30 April 1871</a></li> <li class="leaf menu-mlid-4172"><a href="/letters/favourite-letters/your-letter-eternalized-us-n-d-doedes-27-march-1873">Your letter eternalized before us: From N. D. Doedes, 27 March 1873</a></li> <li class="leaf menu-mlid-4120"><a href="/letters/favourite-letters/lost-translation-auguste-forel-12-november-1874">Lost in translation: From Auguste Forel, 12 November 1874</a></li> <li class="leaf menu-mlid-4182"><a href="/letters/favourite-letters/i-never-trusted-drosera-e-f-lubbock-after-2-july-1875">I never trusted Drosera: From E. F. Lubbock, [after 2 July] 1875</a></li> <li class="leaf menu-mlid-4126"><a href="/letters/favourite-letters/argus-pheasant-mivart-r-wallace-17-june-1876">From Argus pheasant to Mivart: To A. R. Wallace, 17 June 1876</a></li> <li class="leaf menu-mlid-4141"><a href="/letters/favourite-letters/wearing-his-knowledge-lightly-fritz-m-ller-5-april-1878">Wearing his knowledge lightly: From Fritz Müller, 5 April 1878</a></li> <li class="leaf menu-mlid-4123"><a href="/letters/favourite-letters/terms-engagement-julius-wiesner-25-october-1881">Terms of engagement: To Julius Wiesner, 25 October 1881</a></li> <li class="last leaf menu-mlid-4108"><a href="/letters/favourite-letters/intellectual-capacities-caroline-kennard-26-december-1881">Intellectual capacities: From Caroline Kennard, 26 December 1881</a></li> </ul></li> <li class="expanded menu-mlid-4144"><a href="/letters/darwin-plays">Darwin plays</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-4144"><a href="/letters/darwin-plays">Darwin plays overview</a></li> <li class="leaf menu-mlid-4150"><a href="/letters/darwin-plays/emma-audio-play">'Emma' audio play</a></li> <li class="leaf menu-mlid-4147"><a href="/letters/darwin-plays/frank-audio-play">'Frank' audio play</a></li> <li class="leaf menu-mlid-4153"><a href="/letters/darwin-plays/confessing-murder-audio-play">'Like confessing a murder' audio play</a></li> <li class="last expanded menu-mlid-2294"><a href="/letters/darwin-plays/re-design-dramatisation">'Re: Design' dramatisation</a><ul class="menu"><li class="first collapsed menu-mlid-2294"><a href="/letters/darwin-plays/re-design-dramatisation">'Re: Design' dramatisation overview</a></li> <li class="last leaf menu-mlid-1134"><a href="/commentary/religion/re-design-dramatisation/dramatisation-script">Dramatisation script</a></li> </ul></li> </ul></li> <li class="leaf menu-mlid-3377"><a href="https://www.darwinproject.ac.uk/search?sort=date&keyword=darwin&f1-document-type=letter" title="">Browse all Darwin letters in date order</a></li> <li class="last leaf menu-mlid-3391"><a href="/letters/list-correspondents">List of correspondents</a></li> </ul></li> <li class="expanded menu-mlid-824"><a href="/commentary">Commentary</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-824"><a href="/commentary">Commentary overview</a></li> <li class="expanded menu-mlid-2269"><a href="/commentary/evolution" title="">Evolution</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2269"><a href="/commentary/evolution" title="">Evolution overview</a></li> <li class="leaf menu-mlid-2262"><a href="/commentary/evolution/natural-selection">Natural selection</a></li> <li class="leaf menu-mlid-2293"><a href="/commentary/evolution/sexual-selection">Sexual selection</a></li> <li class="leaf menu-mlid-2299"><a href="/commentary/evolution/inheritance">Inheritance</a></li> <li class="leaf menu-mlid-2298"><a href="/commentary/evolution/divergence">Divergence</a></li> <li class="leaf menu-mlid-3410"><a href="/commentary/evolution/correlation-growth-deaf-blue-eyed-cats-pigs-and-poison">Correlation of growth: deaf blue-eyed cats, pigs, and poison</a></li> <li class="leaf menu-mlid-3414"><a href="/commentary/evolution/natural-selection-trouble-terminology-part-i">Natural Selection: the trouble with terminology Part I</a></li> <li class="leaf menu-mlid-3415"><a href="/commentary/evolution/survival-fittest-trouble-terminology-part-ii">Survival of the fittest: the trouble with terminology Part II</a></li> <li class="last leaf menu-mlid-3671"><a href="/commentary/evolution/darwin-s-species-notebooks-i-think">Darwin’s species notebooks: ‘I think . . .’</a></li> </ul></li> <li class="expanded menu-mlid-871"><a href="/commentary/geology" title="">Geology</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-871"><a href="/commentary/geology" title="">Geology overview</a></li> <li class="leaf menu-mlid-2259"><a href="/commentary/geology/darwin-geology">Darwin and geology</a></li> <li class="leaf menu-mlid-1127"><a href="/topics/geology/darwin-s-introduction-geology">Darwin’s introduction to geology</a></li> <li class="leaf menu-mlid-1270"><a href="/commentary/geology/geology-beagle-voyage">The geology of the Beagle voyage</a></li> <li class="leaf menu-mlid-1122"><a href="/commentary/geology/darwin-coral-reefs">Darwin and coral reefs</a></li> <li class="leaf menu-mlid-2258"><a href="/commentary/geology/darwin-s-earthquakes">Darwin’s earthquakes</a></li> <li class="leaf menu-mlid-2257"><a href="/topics/geology/darwin-geological-society" title="">Darwin and the Geological Society</a></li> <li class="leaf menu-mlid-1123"><a href="/commentary/geology/darwin-glen-roy">Darwin and Glen Roy</a></li> <li class="last leaf menu-mlid-1087"><a href="/topics/geology/bibliography-darwin-s-geological-publications">Bibliography of Darwin’s geological publications</a></li> </ul></li> <li class="expanded menu-mlid-2247"><a href="/commentary/life-sciences" title="">Life sciences</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2247"><a href="/commentary/life-sciences" title="">Life sciences overview</a></li> <li class="expanded menu-mlid-1117"><a href="/commentary/life-sciences/darwin-and-down">Darwin and Down</a><ul class="menu"><li class="first collapsed menu-mlid-1117"><a href="/commentary/life-sciences/darwin-and-down">Darwin and Down overview</a></li> <li class="last leaf menu-mlid-1149"><a href="/commentary/life-sciences/darwin-and-down/darwin-s-hothouse-and-lists-hothouse-plants">Darwin’s hothouse and lists of hothouse plants</a></li> </ul></li> <li class="leaf menu-mlid-4087"><a href="/commentary/life-sciences/species-and-varieties">Species and varieties</a></li> <li class="leaf menu-mlid-1058"><a href="/commentary/life-sciences/evolution-honeycomb">The evolution of honeycomb</a></li> <li class="leaf menu-mlid-1083"><a href="/commentary/life-sciences/tale-two-bees">A tale of two bees</a></li> <li class="expanded menu-mlid-1088"><a href="/commentary/life-sciences/beauty-and-seed">Beauty and the seed</a><ul class="menu"><li class="first collapsed menu-mlid-1088"><a href="/commentary/life-sciences/beauty-and-seed">Beauty and the seed overview</a></li> <li class="last leaf menu-mlid-1055"><a href="/commentary/life-sciences/beauty-and-seed/mauro-galetti-profile-ecologist">Mauro Galetti: profile of an ecologist</a></li> </ul></li> <li class="leaf menu-mlid-2261"><a href="/commentary/life-sciences/casting-about-darwin-worms">Casting about: Darwin on worms</a></li> <li class="leaf menu-mlid-1081"><a href="/commentary/life-sciences/was-darwin-ecologist">Was Darwin an ecologist?</a></li> <li class="leaf menu-mlid-3641"><a href="/commentary/life-sciences/dipsacus-and-drosera-frank-s-favourite-carnivores">Dipsacus and Drosera</a></li> <li class="expanded menu-mlid-2318"><a href="/commentary/life-sciences/darwin-and-barnacles">Darwin and barnacles</a><ul class="menu"><li class="first collapsed menu-mlid-2318"><a href="/commentary/life-sciences/darwin-and-barnacles">Darwin and barnacles overview</a></li> <li class="last leaf menu-mlid-2317"><a href="/commentary/life-sciences/darwin-and-barnacles/darwin-s-study-cirripedia">Darwin’s study of the Cirripedia</a></li> </ul></li> <li class="expanded menu-mlid-2280"><a href="/commentary/life-sciences/darwin-and-vivisection">Darwin and vivisection</a><ul class="menu"><li class="first collapsed menu-mlid-2280"><a href="/commentary/life-sciences/darwin-and-vivisection">Darwin and vivisection overview</a></li> <li class="leaf menu-mlid-2284"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-draft-petition">Vivisection: draft petition</a></li> <li class="leaf menu-mlid-2285"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-baas-committee-report">Vivisection: BAAS committee report</a></li> <li class="leaf menu-mlid-2286"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-first-sketch-bill">Vivisection: first sketch of the bill</a></li> <li class="leaf menu-mlid-2287"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-darwins-testimony-royal-commission">Vivisection: Darwin's testimony</a></li> <li class="last leaf menu-mlid-2288"><a href="/topics/life-sciences/darwin-and-vivisection/appeal-against-animal-cruelty">'An Appeal' against animal cruelty</a></li> </ul></li> <li class="last leaf menu-mlid-2916"><a href="/commentary/life-sciences/biodiversity-and-its-histories">Biodiversity and its histories</a></li> </ul></li> <li class="expanded menu-mlid-873"><a href="/commentary/human-nature" title="">Human nature</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-873"><a href="/commentary/human-nature" title="">Human nature overview</a></li> <li class="leaf menu-mlid-2604"><a href="/commentary/human-nature/darwin-human-evolution">Darwin on human evolution</a></li> <li class="expanded menu-mlid-1269"><a href="/commentary/human-nature/expression-emotions">The expression of emotions</a><ul class="menu"><li class="first collapsed menu-mlid-1269"><a href="/commentary/human-nature/expression-emotions">The expression of emotions overview</a></li> <li class="expanded menu-mlid-1262"><a href="/commentary/human-nature/expression-emotions/emotion-experiment">Emotion experiment</a><ul class="menu"><li class="first collapsed menu-mlid-1262"><a href="/commentary/human-nature/expression-emotions/emotion-experiment">Emotion experiment overview</a></li> <li class="last leaf menu-mlid-1263"><a href="/commentary/human-nature/expression-emotions/emotion-experiment/results-darwin-online-emotions">Results of the Darwin Online Emotions Experiment</a></li> </ul></li> <li class="leaf menu-mlid-2256"><a href="/commentary/human-nature/expression-emotions/face-emotion">Face of emotion</a></li> <li class="last leaf menu-mlid-2292"><a href="/commentary/human-nature/expression-emotions/darwin-s-queries-expression">Darwin’s queries on expression</a></li> </ul></li> <li class="expanded menu-mlid-1052"><a href="/commentary/human-nature/origin-language">The origin of language</a><ul class="menu"><li class="first collapsed menu-mlid-1052"><a href="/commentary/human-nature/origin-language">The origin of language overview</a></li> <li class="leaf menu-mlid-1053"><a href="/commentary/human-nature/origin-language/language-key-letters">Language: key letters</a></li> <li class="last leaf menu-mlid-1153"><a href="/commentary/human-nature/origin-language/language-interview-gregory-radick">Language: Interview with Gregory Radick</a></li> </ul></li> <li class="last leaf menu-mlid-1140"><a href="/commentary/human-nature/darwin-and-human-nature-film-series">Film series podcasts</a></li> </ul></li> <li class="expanded menu-mlid-874"><a href="/commentary/religion" title="">Religion</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-874"><a href="/commentary/religion" title="">Religion overview</a></li> <li class="leaf menu-mlid-1033"><a href="/commentary/religion/darwin-and-design">Darwin and design</a></li> <li class="leaf menu-mlid-1288"><a href="/commentary/religion/what-did-darwin-believe">What did Darwin believe?</a></li> <li class="leaf menu-mlid-1034"><a href="/commentary/religion/darwin-and-church">Darwin and the Church</a></li> <li class="leaf menu-mlid-1089"><a href="/commentary/religion/british-association-meeting-1860">British Association meeting 1860</a></li> <li class="leaf menu-mlid-1121"><a href="/commentary/religion/darwin-and-religion-america">Darwin and religion in America</a></li> <li class="expanded menu-mlid-1045"><a href="/commentary/religion/essays-reviews-asa-gray">Essays and reviews by Asa Gray</a><ul class="menu"><li class="first collapsed menu-mlid-1045"><a href="/commentary/religion/essays-reviews-asa-gray">Essays and reviews by Asa Gray overview</a></li> <li class="leaf menu-mlid-2271"><a href="/commentary/religion/essays-reviews-asa-gray/darwiniana-preface">Darwiniana – Preface</a></li> <li class="leaf menu-mlid-2272"><a href="/commentary/religion/essays-reviews-asa-gray/essay-design-versus-necessity">Essay: Design versus necessity</a></li> <li class="leaf menu-mlid-1046"><a href="/commentary/religion/essays-reviews-asa-gray/essay-natural-selection-natural-theology">Essay: Natural selection & natural theology</a></li> <li class="leaf menu-mlid-1146"><a href="/commentary/religion/essays-reviews-asa-gray/essay-evolution-theology">Essay: Evolution and theology</a></li> <li class="leaf menu-mlid-2273"><a href="/commentary/religion/essays-reviews-asa-gray/essay-what-darwinism">Essay: What is Darwinism?</a></li> <li class="last leaf menu-mlid-2274"><a href="/commentary/religion/essays-reviews-asa-gray/essay-evolutionary-teleology">Essay: Evolutionary teleology</a></li> </ul></li> <li class="last expanded menu-mlid-2249"><a href="/commentary/religion/science-and-religion-interviews">Science and religion Interviews</a><ul class="menu"><li class="first collapsed menu-mlid-2249"><a href="/commentary/religion/science-and-religion-interviews">Science and religion Interviews overview</a></li> <li class="leaf menu-mlid-2250"><a href="/commentary/religion/science-and-religion-interviews/interview-emily-ballou">Interview with Emily Ballou</a></li> <li class="leaf menu-mlid-2251"><a href="/commentary/religion/science-and-religion-interviews/interview-simon-conway-morris">Interview with Simon Conway Morris</a></li> <li class="leaf menu-mlid-2252"><a href="/commentary/religion/science-and-religion-interviews/interview-john-hedley-brooke">Interview with John Hedley Brooke</a></li> <li class="leaf menu-mlid-2253"><a href="/commentary/religion/science-and-religion-interviews/interview-randal-keynes">Interview with Randal Keynes</a></li> <li class="leaf menu-mlid-2254"><a href="/commentary/religion/science-and-religion-interviews/interview-tim-lewens">Interview with Tim Lewens</a></li> <li class="last leaf menu-mlid-2255"><a href="/commentary/religion/science-and-religion-interviews/interview-pietro-corsi">Interview with Pietro Corsi</a></li> </ul></li> </ul></li> <li class="last expanded menu-mlid-3395"><a href="/commentary/curious" title="">For the curious...</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-3395"><a href="/commentary/curious" title="">For the curious... overview</a></li> <li class="leaf menu-mlid-4155"><a href="/commentary/curious/cordillera-beagle-expedition">Cordillera Beagle expedition</a></li> <li class="leaf menu-mlid-4096"><a href="/commentary/curious/darwin-family">The Darwin family</a></li> <li class="leaf menu-mlid-4093"><a href="/commentary/curious/darwin-s-plant-experiments">Darwin’s plant experiments</a></li> <li class="leaf menu-mlid-4090"><a href="/commentary/curious/behind-scenes">Behind the scenes</a></li> <li class="leaf menu-mlid-3664"><a href="/commentary/curious/darwin-s-networks">Darwin’s Networks</a></li> <li class="leaf menu-mlid-3643"><a href="/commentary/curious/darwin-and-beagle-voyage">Darwin and the Beagle voyage</a></li> <li class="leaf menu-mlid-3403"><a href="/commentary/curious/darwin-and-working-home">Darwin and working from home</a></li> <li class="leaf menu-mlid-3409"><a href="/commentary/curious/darwin-cats-and-cat-shows">Darwin, cats and cat shows</a></li> <li class="leaf menu-mlid-3396"><a href="/commentary/curious/darwin-and-dogs">Darwin and dogs</a></li> <li class="leaf menu-mlid-3397"><a href="/commentary/curious/darwins-illness">Darwin's illness</a></li> <li class="leaf menu-mlid-3398"><a href="/commentary/curious/plant-or-animal-or-don-t-try-home">Plant or animal? (Or: Don’t try this at home!)</a></li> <li class="last leaf menu-mlid-3399"><a href="/commentary/curious/strange-things-sent-darwin-post">Strange things sent to Darwin in the post</a></li> </ul></li> </ul></li> <li class="expanded menu-mlid-823"><a href="/people">People</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-823"><a href="/people">People overview</a></li> <li class="leaf menu-mlid-2150"><a href="/commentary/key-correspondents" title="">Key correspondents</a></li> <li class="leaf menu-mlid-2153"><a href="/commentary/beagle-voyage-networks" title="">Beagle voyage networks</a></li> <li class="leaf menu-mlid-2152"><a href="/commentary/family-and-friends" title="">Family and friends</a></li> <li class="leaf menu-mlid-2159"><a href="/commentary/darwins-scientific-network" title="">Darwin's scientific network</a></li> <li class="leaf menu-mlid-2155"><a href="/tags/readers-and-critics" title="">Readers and critics</a></li> <li class="leaf menu-mlid-2162"><a href="/tags/publishers-artists-and-illustrators" title="">Publishers, artists and illustrators</a></li> <li class="leaf menu-mlid-3381"><a href="/correspondents-alphabetical" title="">People pages in alphabetical order</a></li> <li class="expanded menu-mlid-3367"><a href="/people/german-and-dutch-photograph-albums">German and Dutch photograph albums</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-3367"><a href="/people/german-and-dutch-photograph-albums">German and Dutch photograph albums overview</a></li> <li class="leaf menu-mlid-3372"><a href="/people/german-and-dutch-photograph-albums/photograph-album-german-and-austrian-scientists">Photograph album of German and Austrian scientists</a></li> <li class="leaf menu-mlid-3371"><a href="/people/german-and-dutch-photograph-albums/photograph-album-dutch-admirers">Photograph album of Dutch admirers</a></li> <li class="last leaf menu-mlid-3370"><a href="/german-poems-presented-darwin">German poems presented to Darwin</a></li> </ul></li> <li class="last leaf menu-mlid-3380"><a href="https://www.darwinproject.ac.uk/search?text=%22%22;f1-document-type=people;sort=name" title="">List of all people mentioned in letters</a></li> </ul></li> <li class="expanded menu-mlid-825"><a href="/learning-resources" title="">Learning</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-825"><a href="/learning-resources" title="">Learning overview</a></li> <li class="expanded menu-mlid-1739"><a href="/learning/7-11">Ages 7-11</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1739"><a href="/learning/7-11">Ages 7-11 overview</a></li> <li class="leaf menu-mlid-2601"><a href="/learning/7-11/darwin-the-collector">Darwin The Collector</a></li> <li class="leaf menu-mlid-2602"><a href="/learning/7-11/detecting-darwin">Detecting Darwin</a></li> <li class="leaf menu-mlid-2603"><a href="/learning/7-11/darwin-and-evolution">Darwin And Evolution</a></li> <li class="leaf menu-mlid-2600"><a href="/learning/7-11/darwins-fantastical-voyage">Darwin's Fantastical Voyage</a></li> <li class="last leaf menu-mlid-3408"><a href="/learning/7-11/home-learning-7-11-years">Home learning: 7-11 years</a></li> </ul></li> <li class="expanded menu-mlid-1744"><a href="/learning/11-14">Ages 11-14</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1744"><a href="/learning/11-14">Ages 11-14 overview</a></li> <li class="leaf menu-mlid-2899"><a href="/learning/11-14/darwin-and-religion">Darwin and Religion</a></li> <li class="leaf menu-mlid-2898"><a href="/learning/11-14/doing-darwins-experiments">Doing Darwin’s Experiments</a></li> <li class="leaf menu-mlid-2900"><a href="/learning/11-14/how-dangerous-was-darwin">How dangerous was Darwin?</a></li> <li class="leaf menu-mlid-2901"><a href="/learning/11-14/offer-of-a-lifetime">Offer of a lifetime</a></li> <li class="leaf menu-mlid-2903"><a href="/learning/11-14/darwin-and-slavery">Darwin and slavery</a></li> <li class="leaf menu-mlid-2902"><a href="/learning/11-14/beagle-voyage">Beagle Voyage</a></li> <li class="leaf menu-mlid-2904"><a href="/learning/11-14/darwins-scientific-women">Darwin’s scientific women</a></li> <li class="last leaf menu-mlid-1154"><a href="/case-studies-using-darwin%E2%80%99s-letters-classroom">Schools Gallery: Using Darwin’s letters in the classroom</a></li> </ul></li> <li class="expanded menu-mlid-833"><a href="/learning/universities">Universities</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-833"><a href="/learning/universities">Universities overview</a></li> <li class="expanded menu-mlid-1063"><a href="/learning/universities/letters-primary-source" title="">Letters as a primary source</a><ul class="menu"><li class="first collapsed menu-mlid-1063"><a href="/learning/universities/letters-primary-source" title="">Letters as a primary source overview</a></li> <li class="leaf menu-mlid-2276"><a href="/learning/universities/letters-primary-source/scientific-networks">Scientific networks</a></li> <li class="leaf menu-mlid-2277"><a href="/learning/universities/letters-primary-source/scientific-practice">Scientific practice</a></li> <li class="leaf menu-mlid-2278"><a href="/learning/universities/letters-primary-source/controversy">Controversy</a></li> <li class="leaf menu-mlid-2279"><a href="/learning/universities/letters-primary-source/religion">Religion</a></li> <li class="leaf menu-mlid-1066"><a href="/learning/universities/letters-primary-source/discussion-questions-and-essay-questions">Discussion questions and essay questions</a></li> <li class="last leaf menu-mlid-1286"><a href="/learning/universities/letters-primary-source/suggested-reading">Suggested reading</a></li> </ul></li> <li class="expanded menu-mlid-1550"><a href="/learning/universities/getting-know-darwins-science">Getting to know Darwin's science</a><ul class="menu"><li class="first collapsed menu-mlid-1550"><a href="/learning/universities/getting-know-darwins-science">Getting to know Darwin's science overview</a></li> <li class="leaf menu-mlid-2144"><a href="/learning/universities/getting-know-darwins-science/early-days">Early days</a></li> <li class="leaf menu-mlid-1086"><a href="/learning/universities/getting-know-darwins-science/barnacles">Barnacles</a></li> <li class="leaf menu-mlid-932"><a href="/learning/universities/getting-know-darwins-science/biogeography">Biogeography</a></li> <li class="leaf menu-mlid-1287"><a href="/learning/universities/getting-know-darwins-science/variation-under-domestication">Variation under domestication</a></li> <li class="leaf menu-mlid-1259"><a href="/learning/universities/getting-know-darwins-science/orchids">Orchids</a></li> <li class="leaf menu-mlid-1150"><a href="/learning/universities/getting-know-darwins-science/instinct-and-evolution-mind">Instinct and the evolution of mind</a></li> <li class="leaf menu-mlid-1048"><a href="/learning/universities/getting-know-darwins-science/insectivorous-plants">Insectivorous plants</a></li> <li class="leaf menu-mlid-1095"><a href="/learning/universities/getting-know-darwins-science/climbing-plants">Climbing plants</a></li> <li class="leaf menu-mlid-1141"><a href="/learning/universities/getting-know-darwins-science/floral-dimorphism">Floral dimorphism</a></li> <li class="leaf menu-mlid-1260"><a href="/learning/universities/getting-know-darwins-science/power-movement-plants">Power of movement in plants</a></li> <li class="leaf menu-mlid-1135"><a href="/learning/universities/getting-know-darwins-science/earthworms">Earthworms</a></li> <li class="last leaf menu-mlid-1132"><a href="/learning/universities/getting-know-darwins-science/dining-down-house">Dining at Down House</a></li> </ul></li> <li class="expanded menu-mlid-2314"><a href="/learning/universities/darwin-and-human-nature">Darwin and human nature</a><ul class="menu"><li class="first collapsed menu-mlid-2314"><a href="/learning/universities/darwin-and-human-nature">Darwin and human nature overview</a></li> <li class="leaf menu-mlid-1155"><a href="/learning/universities/darwin-and-human-nature/moral-nature">Moral nature</a></li> <li class="last leaf menu-mlid-1261"><a href="/learning/universities/darwin-and-human-nature/race-civilization-and-progress">Race, civilization, and progress</a></li> </ul></li> <li class="last expanded menu-mlid-1289"><a href="/learning/universities/women-and-science">Women and science</a><ul class="menu"><li class="first collapsed menu-mlid-1289"><a href="/learning/universities/women-and-science">Women and science overview</a></li> <li class="leaf menu-mlid-2281"><a href="/learning/universities/women-and-science/women-s-scientific-participation">Women’s scientific participation</a></li> <li class="leaf menu-mlid-1290"><a href="/learning/universities/women-and-science/women-scientific-audience">Women as a scientific audience</a></li> <li class="leaf menu-mlid-1041"><a href="/learning/universities/women-and-science/referencing-women-s-work">Referencing women’s work</a></li> <li class="leaf menu-mlid-2312"><a href="/learning/universities/women-and-science/darwin-public-and-private">Darwin in public and private</a></li> <li class="leaf menu-mlid-2283"><a href="/learning/universities/women-and-science/darwin-mentor">Darwin as mentor</a></li> <li class="leaf menu-mlid-2313"><a href="/learning/universities/women-and-science/discussion-questions">Discussion questions</a></li> <li class="last leaf menu-mlid-2311"><a href="/learning/universities/women-and-science/suggested-reading">Suggested reading</a></li> </ul></li> </ul></li> <li class="leaf menu-mlid-2897"><a href="/learning-resources/timeline" title="">Darwin timeline</a></li> <li class="leaf menu-mlid-3668"><a href="/learning-resources/exploring-evolution">Teacher training</a></li> <li class="last expanded menu-mlid-4186"><a href="/commentary/curious" title="">For the curious...</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-4186"><a href="/commentary/curious" title="">For the curious... overview</a></li> <li class="leaf menu-mlid-4189"><a href="/commentary/curious/cordillera-beagle-expedition" title="">Cordillera Beagle expedition</a></li> <li class="leaf menu-mlid-4192"><a href="/commentary/curious/darwin-family" title="">The Darwin family</a></li> <li class="leaf menu-mlid-4195"><a href="/commentary/curious/darwin-s-plant-experiments" title="">Darwin’s plant experiments</a></li> <li class="leaf menu-mlid-4198"><a href="/commentary/curious/behind-scenes" title="">Behind the scenes</a></li> <li class="leaf menu-mlid-4201"><a href="/commentary/curious/darwin-s-networks" title="">Darwin’s Networks</a></li> <li class="leaf menu-mlid-4204"><a 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href="https://www.facebook.com/sharer/sharer.php?u=https%3A%2F%2Fwww.darwinproject.ac.uk%2Fletters%2Fdarwins-works-letters%2Fexpression" title="Share on Facebook" target="_blank"><i class="fab fa-facebook-f"></i></a> <a class="icon-sm darwin-twitter" href="https://twitter.com/intent/tweet?text=Expression&url=https%3A%2F%2Fwww.darwinproject.ac.uk%2Fletters%2Fdarwins-works-letters%2Fexpression" title="Share on Twitter" target="_blank"><i class="fab fa-twitter"></i></a> <a class="icon-sm darwin-email" href="mailto:?&subject=Expression&body=https%3A%2F%2Fwww.darwinproject.ac.uk%2Fletters%2Fdarwins-works-letters%2Fexpression" title="Share by email"><i class="fas fa-envelope"></i></a></div>  </div> </div> <div id="block-system-main" class="block block-system"> <div> <div id="node-575" class="node node-page clearfix" about="/letters/darwins-works-letters/expression" typeof="foaf:Document"> <span property="dc:title" content="Expression" class="rdf-meta element-hidden"></span><span property="sioc:num_replies" content="0" datatype="xsd:integer" class="rdf-meta element-hidden"></span> <div class="content campl-content-container"> <div class="field field-name-field-image field-type-file field-label-hidden"><div class="field-items"><div class="field-item even"><div id="file-918" class="file file-image file-image-jpeg"> <h2 class="element-invisible"><a href="/file/dar-531-c125rjpg">DAR 53.1 C125r.jpg</a></h2> <div class="content"> <a href="https://www.darwinproject.ac.uk/sites/default/files/DAR%2053.1%20C125r.jpg"><img typeof="foaf:Image" class="campl-scale-with-grid" src="https://www.darwinproject.ac.uk/sites/default/files/DAR%2053.1%20C125r.jpg" width="424" height="687" alt="Sulky child from Expression" title="Sulky child from Expression" /></a><div class="field field-name-field-file-caption field-type-text field-label-hidden"><div class="field-items"><div class="field-item even">Sulky child from Expression, pl. 2; DAR 53.1 C125r</div></div></div><div class="field field-name-field-credit field-type-text field-label-hidden"><div class="field-items"><div class="field-item even">By permission of the Syndics of Cambridge University Library</div></div></div> </div> </div> </div></div></div><div class="field field-name-body field-type-text-with-summary field-label-hidden"><div class="field-items"><div class="field-item even" property="content:encoded"><p>Darwin's interest in emotional expression can be traced as far back as the <em>Beagle</em> voyage. He was fascinated by the different sounds and gestures among the peoples of Tierra del Fuego, and on his return from the voyage he started recording observations in a set of notebooks, later labelled 'Metaphysics on Morals & Speculations on Expression'. Details about the facial and bodily movements accompanying various emotions were entered, alongside remarks on the origins of language, aesthetic taste, and sympathy, comparing the behaviour of humans and animals. Darwin's work on expression was thus closely tied to his larger research programme on the modification of species and the animal ancestry of all our physical and mental attributes. Like the other notebooks on transmutation, these were kept private. A vast amount of research accumulated over time, and when the book finally appeared in 1872, it had been over thirty years in gestation.</p> <p>Darwin sometimes referred to expression as '<a href="/letter/?docId=letters/DCP-LETT-5330.xml">an old hobby horse</a>' and even made light of the work. '<a href="/letter/?docId=letters/DCP-LETT-3875.xml">I have got some funny notions on the subject</a>', he wrote to Hooker in 1862. After <em>Origin</em> was published, he suffered an extended period of illness and despaired that he would never bring the project to a conclusion. At a particularly low point in 1864, he even offered all his material to Alfred Russel Wallace: '<a href="/letter/?docId=letters/DCP-LETT-4510.xml">I have collected a few notes on man but I do not suppose I shall ever use them ... There is much more that I shd like to write but I have not strength</a>'. Darwin persevered, however. He planned at first to present all his conclusions on human evolution in <a href="/letter/?docId=letters/DCP-LETT-5348.xml">a single chapter in <em>Variation</em></a> (treating man as a domesticated animal). But that book swelled to two volumes, and then so did <em>Descent</em>. By 1870, Darwin had amassed so much material that it required separate publication (<em>Expression</em> ran to 367 pages).</p> <p>One of Darwin's main objects of study was infants. As soon as his first child (William) was born he began recording the boy's development in a separate notebook. But even before his marriage, he shared his interest in expression with Emma (then his fiancée), and asked her to observe the '<a href="/letter/?docId=letters/DCP-LETT-490.xml">nods & winks</a>' of her nephew, Ernest Wedgwood. The couple would continue recording observations of their children through the early 1850s. Other family relations and friends were asked to contribute. Perhaps unsurprisingly, many of the observations of babies and young children were made by mothers, and their communications <a href="/letter/?docId=letters/DCP-LETT-5332.xml">were often addressed either to Emma or her eldest daughter</a>. 'I am afraid you will think I have taken a long time in answering your questions,' wrote Margaret Vaughan Williams to Henrietta in 1869, '<a href="/letter/?docId=letters/DCP-LETT-6535.xml">but we do find it so hard to observe Baby’s expression accurately, & besides I seldom can attend to his eyebrows, when he is crying and I am wanting to comfort him</a>'. Unlike the many men who contributed to Darwin's research, and whose authority as observers was often highlighted in his text, these women observers were rarely acknowledged by name.</p> <p>While he was observing his children at home, Darwin was also studying expression in the family pets. His beloved dogs were favourite subjects and appeared frequently both in <em>Expression</em> and <em>Descent</em>. The family also kept cats, and Henrietta made a series of observations on their behaviour: one '<a href="/letter/?docId=letters/DCP-LETT-4260F.xml">about the age of 10 months when put upon a soft cloak or gown used to take a piece in her mouth & then pound with her claws out & purr</a>'. Observations extended to caged creatures. Darwin requested his niece Lucy Wedgwood to '<a href="/letter/?docId=letters/DCP-LETT-7223.xml">think of any fact about expression of any emotion in any of your birds.</a>' J. J. Weir described canaries and goldfinches ruffling their feathers when frightened or angry: '<a href="/letter/?docId=letters/DCP-LETT-7247.xml">I have a Goldfinch male, often in my dining room, of the most irascible disposition</a>'. The keeper of the Zoological Gardens at Regents Park was asked whether squirrel monkeys screamed like a baby, with wrinkled up skin around the eyes: '<a href="/letter/?docId=letters/DCP-LETT-7072.xml">Could you make it scream without hurting it much?</a>' When in London, Darwin sometimes went to the zoo himself. One day <a href="/letter/?docId=letters/DCP-LETT-6067.xml">he saw elephants shedding tears when they trumpeted</a>, and he asked a correspondent in Ceylon, where elephants were captured and bound, to confirm: '<a href="/letter/?docId=letters/DCP-LETT-2670.xml">I am going to beg a favor of you, which will appear one of the oddest ever asked</a>'.</p> <p>Another main object of Darwin's investigation was indigenous peoples, especially 'savages' or 'less civilised natives' who had had little contact with Europeans. He began writing to naturalists, travellers and settlers in European colonies in the mid-1850s. His questions about expression were often conjoined with others on beauty and mate selection: '<a href="/letter/?docId=letters/DCP-LETT-6520.xml">would the handsomest woman in a tribe in the eyes of a New Zealander, be the handsomest woman in our eyes?</a>' By 1860, his inquiries had become more systematic, and he sent a long list to Thomas Bridges, a missionary stationed in the Falkland's, including: '<a href="/letter/?docId=letters/DCP-LETT-2640.xml">Do the Fuegians or Patagonians, or both, nod their heads vertically to express assent, and shake their heads horizontally to express dissent?</a>'. Darwin continued to refine his questions, and for ease of distribution he had the list printed around 1867, and began circulating it more widely through his correspondence network. Some of his contacts were able to distribute additional copies locally, so that <a href="/letter/?docId=letters/DCP-LETT-6314.xml">Darwin's questionnaire reached some remote parts of the world</a>. Responses were received from China, India, Malaysia, Australia, North and South America, and South Africa.</p> <p>Despite Darwin's precisely worded questions, which could have been answered with a simple 'yes' or 'no', many of the replies contained lengthy descriptions of the natives peoples and their interactions with Europeans, some in highly charged settings such as criminal trials, or altercations with colonial officials in the streets. Observations of expression were often mingled with judgments about the emotional nature or moral character of the observed. '<a href="/letter/?docId=letters/DCP-LETT-6054.xml">Guilty sly and jealous expressions are distinctly seen in these two tribes notwithstanding that they are otherwise great adepts in concealing their feelings.</a>' '<a href="/letter/?docId=letters/DCP-LETT-6160.xml">Defiance is indeed grotesquely expressed by the lower class of Bengalees. inter se. Again and again I have watched them in their brawls and squabbles, and really they strike me more as the snarling contentions of cowardly dogs</a>'. Darwin received one response from a native in South Africa, the brother of a local chief who had been educated by missionaries and was <a href="/letter/?docId=letters/DCP-LETT-5581.xml">employed by the colonial government as a constable</a>. Christian Gaika's replies, written out in English, so impressed Darwin that he remarked to his colonial contact John Weale: '<a href="/letter/?docId=letters/DCP-LETT-5617.xml">That I shd receive answers written by the brother of a Kaffir chief is a truly wonderful fact in the progress of civilization</a>'. Darwin's outlook was immediately condemned, however, by Weale: '<a href="/letter/?docId=letters/DCP-LETT-5722.xml">You speak sanguinely about the civilization of the natives, & the fact that Christian Gaika can write. This appears to me an error into which most people in England fall ...  the longer a Kafir has been on a Mission Station the worse servant he is.</a>' Most of the overt racism of Darwin's correspondents was passed over when he came to summarize his findings for publication. Many of the cultural differences between peoples were also erased. Such exclusions allowed Darwin to conclude more readily that emotional response was universal across the globe, a key support to his more general argument about the common descent of the human races from a single ancestral species.</p> <p>Another class of subjects whom Darwin thought were particularly important were the insane, for he believed that they shared with children the inability to control or conceal strong emotions. In 1869, he contacted the well-known alienist Henry Maudsley who forwarded Darwin's queries to his friend, <a href="/letter/?docId=letters/DCP-LETT-6752.xml">James Crichton Browne, the superintendent of an asylum in West Riding</a>. Browne was trying to make the institution a centre of research on madness and diseases of the brain, and he composed detailed descriptions of patients who suffered from emotional disorders such as extreme fear, rage, and melancholy. He also used photography to document and classify his patients, and he sent Darwin <a href="/letter/?docId=letters/DCP-LETT-7658.xml">a large number of medical portraits, with his own hand-written diagnoses at the bottom</a>. Darwin eventually used one of these, a patient named <a href="/letter/?docId=letters/DCP-LETT-8253.xml">Ruth Lockwood whose hair bristled in horror similar to that of some animals when under threat</a>. For Browne, Darwin's letters and the opportunity to assist him offered a 'genuine solace' in his 'house of bondage': '<a href="/letter/?docId=letters/DCP-LETT-7484.xml">The exigencies of the public service have already ruined my health, & curtailed my capacities. They now threaten to shorten my life</a>'.</p> <p>On learning of Browne's interest in photography, Darwin lent him another set of photographs that had been produced in a medical setting. This was the folio-sized album of <em>Mécanisme de la physionomie humaine</em> by the French physiologist Emile Duchenne. It contained over a hundred full-page images showing a wide range of expressions that had been produced using a galvanic technique. By applying electrical probes directly to the skin, Duchenne claimed he could reproduce expressions more authentically than the often exaggerated depictions of artists. When Darwin requested permission to use some of the photographs from his album, Duchenne refused to accept payment: '<a href="/letter/?docId=letters/DCP-LETT-7623.xml">So you will understand, dear Sir, that between men of science the question of money cannot exist</a>.'  Darwin was extremely interested in Duchenne's work in part because it drew on the physiology of reflexes, rather than subjective emotional states, to explain expression. This was consistent with Darwin's theory that emotional response was fundamentally instinctive, a set of behaviours that had once given competitive advantage to animals, and that had persisted in humans, forming an automatic system of nervous discharge and muscular movement, such as the gaping mouth and eyes, the bristling hair and racing heart that accompanied fear and that (had once) prepared a creature to flee or fight.</p> <p>Darwin drew heavily on the experimental approach to expression pioneered by Charles Bell in the early decades of the nineteenth century, even though he took issue with Bell's view that humans had a separately designed set of facial muscles to communicate higher moral and spiritual feelings. Darwin corresponded with a number of medical specialists to confirm some of Bell's findings on the physiology of weeping. He asked the ophthalmic surgeon William Bowman to observe if '<a href="/letter/?docId=letters/DCP-LETT-5589.xml">when an infant screams violently, it closes the orbicular muscles so as to compress the eyes & prevent them becoming gorged with blood</a>'. The Dutch eye specialist Franz Donders performed detailed experiments on Darwin's behalf to determine the specific nerve fibres responsible for tear secretion: 'this is difficult and laborious, but not impossible... '<a href="/letter/?docId=letters/DCP-LETT-7207.xml">where there is a will, there is a way</a>''. Some of Darwin's questions stretched the limits of physiological knowledge and challenged experts to investigate new phenomena. One particularly puzzling expression was blushing, which seemed to be uniquely human, and to involve the operation of mental attention upon the blood (hence its association with shame and embarrassment). Darwin found evidence of the phenomenon in dark-skinned races, even though it was scarcely visible, and <a href="/letter/?docId=letters/DCP-LETT-6069.xml">among people </a>who were <a href="/letter/?docId=letters/DCP-LETT-7857.xml">born blind</a>. But how could an act of self-attention (a pang of guilt), cause the face to redden? <a href="/letter/?docId=letters/DCP-LETT-7800.xml">And how could such a stimulus-response mechanism have arisen?</a></p> <p><img alt="Rejlander posing with baby Jinx, CUL DAR 53.1 C96r" src="https://www.darwinproject.ac.uk/sites/default/files/DAR%2053.1%20C96r.jpg" style="float: right; margin: 4px;" />One of the striking features of <em>Expression</em> was its combination of highly technical discussions (on the physiology of tears, for example), and deeply sentimental material (portraits of weeping babies). Darwin had started collecting photographs from the mid-1860s, when studio methods and technologies began to reduce exposure times sufficiently to allow emotional expressions to be captured with apparent authenticity. Darwin was particularly keen on portraits of children or infants smiling, frowning, or crying. He made contact with several studio photographers and succeeded in getting one of the best known, Oscar Rejlander, to work on commission. Rejlander took such an interest in Darwin's work that he <a href="/letter/?docId=letters/DCP-LETT-7418.xml">sent detailed notes, posed for a number of pictures himself, and also took photographs of his wife, mimicking natural expressions</a>. '<a href="/letter/?docId=letters/DCP-LETT-7728.xml">It is very difficult to get, at will—those expressions you wish ... In time—I might catch some —So I have tried in propria persona— —even cut my moustache shorter to try to please you in the last batch</a>'.</p> <p>At the time, the use of photographs was quite novel in a scientific publication, and it posed considerable difficulties. <a href="/letter/?docId=letters/DCP-LETT-7705.xml">Darwin warned his publisher about the potential costs</a>. He also tried to maximize the <a href="/letter/?docId=letters/DCP-LETT-7744.xml">use of photos by mounting a number of images on individual plates</a>. This had the effect of de-personalizing the images, making readers attend more closely to the details of outward expression (the shape of the mouth in crying, the furrowed brow in grief), rather than the inner feelings of the subject.</p> <p>Though costly and time-consuming to produce, the photographs and other illustrations were extremely well-received by readers. Reviews also commented on the book's 'popular nature', the charming animal stories and entertaining anecdotes of children. Darwin's manner of involving his readers in the process of investigation may have encouraged them to share their own observations of expression directly with the author. For years after publication, letters arrived about dogs who grin, cats who beg for food, students wiggling their ears in excitement during lectures, children stomping the ground when angry, <a href="/letter/?docId=letters/DCP-LETT-8708.xml">Maori men who rub noses and weep in the street</a>. Darwin collected the material and made notes for revision, but a second edition was not published until after his death, edited by his son Francis. This may have been partly because the demand for reprinting slowed considerably after the first year, but Darwin was also keen to leave the subject of human evolution behind. Darwin's observations of children and his global survey of expression were among the very few areas of his research that focused directly on humans. After <em>Expression</em>, he returned at once to his favourite creatures on earth, plants, which would occupy him for the rest of his life.</p> <p> </p> <p><strong>Find out more about <a href="https://www.darwinproject.ac.uk/commentary/human-nature/expression-emotions">Darwin's <em>Experssion of emotions</em></a></strong></p> </div></div></div><div class="field field-name-field-terms field-type-taxonomy-term-reference field-label-above"><div class="field-label">Terms: </div><div class="field-items"><div class="field-item even"><a href="/commentary/human-nature" typeof="skos:Concept" property="rdfs:label skos:prefLabel" datatype="">Human nature</a></div></div></div><div class="field field-name-field-term-related-letters field-type-text field-label-above"><div class="field-label">Related letters: </div><div class="field-items"><div class="field-item even">5330</div><div class="field-item odd">6535</div><div class="field-item even">2670</div><div class="field-item odd">4260F</div><div class="field-item even">7418</div><div class="field-item odd">5589</div><div class="field-item even">5581</div><div class="field-item odd">7658</div><div class="field-item even">7207</div><div class="field-item odd">7728</div><div class="field-item even">7744</div><div class="field-item odd">7247</div></div></div> </div> </div> </div> </div> </div> </div> </div> <div class="campl-column3 campl-secondary-content "> <div class="region region-sidebar"> <div id="block-menu-block-3" class="block block-menu-block"> <div class="campl-content-container campl-no-bottom-padding"> <div class="campl-heading-container"> <h2>In this section:</h2> </div> </div> <div class="campl-content-container"> <div class="menu-block-wrapper menu-block-3 menu-name-main-menu parent-mlid-0 menu-level-3"> <ul class="menu"><li class="first leaf menu-mlid-3385 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Barrier%20reef.jpg?itok=2F1QJpfK)"><a href="/letters/darwins-works-letters/journal-researches">Journal of researches</a></li> <li class="leaf menu-mlid-3386 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Nomenclature%20of%20the%20valves.jpg?itok=IPSwrW5J)"><a href="/letters/darwins-works-letters/living-and-fossil-cirripedia">Living and fossil cirripedia</a></li> <li class="collapsed menu-mlid-3400 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Natural%20Selection%20page.jpg?itok=3t8mZVx1)"><a href="/letters/darwins-works-letters/origin-big-book">Before Origin: the ‘big book’</a></li> <li class="leaf menu-mlid-3406 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Origin%20cover_1859.jpg?itok=eUmxW2gz)"><a href="/letters/darwins-works-letters/origin">Origin</a></li> <li class="collapsed menu-mlid-3407 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Origin-Spines.jpg?itok=jHvAD6fT)"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions">Rewriting Origin - the later editions</a></li> <li class="leaf menu-mlid-3404 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/orchids_cover3.jpg?itok=kMlJZfRP)"><a href="/letters/darwins-works-letters/orchids">Orchids</a></li> <li class="leaf menu-mlid-3662 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Solanum%20jasminoides.jpg?itok=FqrfKrMP)"><a href="/letters/darwins-works-letters/climbing-plants">Climbing plants</a></li> <li class="leaf menu-mlid-3387 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/DAR_225_178.jpg?itok=Zc3lfCgr)"><a href="/letters/darwins-works-letters/descent">Descent</a></li> <li class="leaf active-trail active menu-mlid-3388 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/DAR%2053.1%20C125r.jpg?itok=D-Wc1iAl)"><a href="/letters/darwins-works-letters/expression" class="active-trail active">Expression</a></li> <li class="leaf menu-mlid-3389 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Drosera-rotundiflora.jpg?itok=ybw7Urdy)"><a href="/letters/darwins-works-letters/insectivorous-plants">Insectivorous plants</a></li> <li class="leaf menu-mlid-3663 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Long%20and%20short%20styled%20form.jpg?itok=4k9jJCMj)"><a href="/letters/darwins-works-letters/forms-flowers">Forms of flowers</a></li> <li class="leaf menu-mlid-4084 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Cross%20and%20Self%20Diagram.jpg?itok=WOMOPSji)"><a href="/letters/darwins-works-letters/cross-and-self-fertilisation">Cross and self fertilisation</a></li> <li class="leaf menu-mlid-3405 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Erasmus%20Darwin.jpg?itok=4asdQNd3)"><a href="/letters/darwins-works-letters/life-erasmus-darwin">Life of Erasmus Darwin</a></li> <li class="last leaf menu-mlid-3665 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/MS-DAR-00209-00015-000-00089c.jpg?itok=7_bZSSKt)"><a href="/letters/darwins-works-letters/movement-plants">Movement in Plants</a></li> </ul></div> </div> </div> <div id="block-cudl-related-letters" class="block block-cudl"> <div class="campl-content-container campl-no-bottom-padding"> <div class="campl-heading-container"> <h2>Related letters</h2> </div> </div> <div class="campl-content-container"> <div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-4260F.xml">From Henrietta Emma Darwin [1 August 1863]</a></h3> <div class="summary"><p>A memorandum describing the expressive behaviour of a cat with added notes by CD.</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-5330.xml">To B. J. Sulivan 31 December [1866]</a></h3> <div class="summary"><p>Thanks BJS for his account of S. America and the Fuegians.</p><p>Can BJS ask W. H. Stirling to make observations on expression?</p><p>Has asked Hooker about the fossil leaves, and he suggests they be sent to Oswald Heer.</p><p>Has just sent MS on domestic animals [<i>Variation</i>] to the printer.</p></div> </div><div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-5581.xml">From J. P. M. Weale 7 July 1867</a></h3> <div class="summary"><p>Has distributed CD’s questions on expression. Observations on the natives.</p><p>Floral structure encouraging cross-pollination in <i>Polygala</i>.</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-5589.xml">To William Bowman 30 July [1867]</a></h3> <div class="summary"><p>Plans to write a book on expression. Questions WB on orbicular muscle in screaming infant and function of muscle contractions in looking at a distant object.</p></div> </div><div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-2670.xml">To George Henry Kendrick Thwaites 31 January [1868]</a></h3> <div class="summary"><p>Asks GHKT about eyes of screaming elephants.</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-6535.xml">From Margaret Susan Vaughan Williams to Henrietta Emma Darwin [after 14 October 1869]</a></h3> <div class="summary"><p>Describes expression of her baby when crying.</p></div> </div><div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-7207.xml">From F. C. Donders 28 May 1870</a></h3> <div class="summary"><p>A detailed description of the physiological and anatomical processes related to the prolonged involuntary contraction of the orbicular muscles and the secretion of tears (as in retching, violent coughing, or laughing). [See <i>Expression</i>, p. 160.].</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-7247.xml">From J. J. Weir 27 June 1870</a></h3> <div class="summary"><p>On behaviour of birds when frightened and when threatening.</p><p>Purple <i>Cytisus</i> grafted onto yellow stock produces some yellow flowers.</p><p>Mutations in rabbits.</p><p>Cites case of variegated leaf form of one plant apparently spreading to a neighbour.</p></div> </div><div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-7418.xml">From O. G. Rejlander [1871]</a></h3> <div class="summary"><p>Observations on expression.</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-7658.xml">From James Crichton-Browne 3 April 1871</a></h3> <div class="summary"><p>Sends photographs of general paralytics. Expressions of exaltation of [these?] patients do not come out well in the photographs.</p><p>Is experimenting with idiots under his care. Has been unable to produce a blush in any one of them.</p></div> </div><div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-7728.xml">From O. G. Rejlander 30 April 1871</a></h3> <div class="summary"><p>Since it is difficult to catch the expressions CD wants, OGR is posing himself.</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-7744.xml">To R. F. Cooke 8 May 1871</a></h3> <div class="summary"><p>Sends photographs he intends to use in <i>Expression</i> and asks for estimates of cost per thousand of each process [Woodburytype and Heliotype] suggested by RC and for different numbers of photos on each plate.</p></div> </div> </div> </div> <div id="block-cudl-related-people" class="block block-cudl"> <div class="campl-content-container campl-no-bottom-padding"> <div class="campl-heading-container"> <h2>Related people</h2> </div> </div> <div class="campl-content-container"> <div class="people even"> <h3><a href="/letter/?docId=nameregs/nameregs_1354.xml">Donders, F. C.</a></h3> <div class="date">1818–89</div> <div class="summary">Dutch physiologist and ophthalmologist. MD, Leiden, 1840. Professor of anatomy at Utrecht military school; professor of physiology at the University of Utrecht, 1862. President of the science section of the Royal Amsterdam Academy of Sciences, 1865–83. Founder and director of a physiology laboratory at the University of Utrecht, 1866–88. Founder and director of the Netherlands Hospital for Eye Patients, 1858–83. FRS 1866.</div> </div> <div class="people odd"> <h3><a href="/letter/?docId=nameregs/nameregs_1404.xml">Duchenne, G. B. A.</a></h3> <div class="date">1806–75</div> <div class="summary">French physician. Studied medicine at Paris, then practised in his native town of Boulogne-sur-Mer. Returned to Paris in 1855 and carried out experiments on the therapeutic use of electricity. One of the founders of neurology.</div> </div> <div class="people even"> <h3><a href="/letter/?docId=nameregs/nameregs_3973.xml">Rejlander, O. G.</a></h3> <div class="date">1813–75</div> <div class="summary">Swedish?-born photographer. Studied art in Rome. Opened a painter's studio in Wolverhampton in 1845. Took up photography in 1853. Moved to London in 1862. Famous for genre photographs. CD used some of his photographs in Expression.</div> </div> <div class="people odd"> <h3><a href="/letter/?docId=nameregs/nameregs_5719.xml">Gaika, Christian</a></h3> <div class="date">1831–1905</div> <div class="summary">Xhosa interpreter and constable. Educated by missionaries. Interpreter and constable in the division of Bedford, Cape Colony, from 1867 or before until 1887 or later. Son of Ngqika (Gaika), a Xhosa chief. Brother of Chief Sandili.</div> </div> <div class="people even"> <h3><a href="/letter/?docId=nameregs/nameregs_621.xml">Bridges, Thomas (b)</a></h3> <div class="date">b. c. 1842 d. 1898</div> <div class="summary">Dictionary compiler, missionary, and sheep farmer. Missionary on Keppel Island, West Falklands, 1856–68; established a mission at Ushuaia in the Beagle Channel, 1867. Settled permanently in Ushuaia with his wife, Mary Ann Varder, and daughter in 1871. Abandoned missionary work in 1884 and became a sheep farmer near Ushuaia from 1887. Author of a Yahgan–English dictionary and Yahgan grammar.</div> </div> <div class="people odd"> <h3><a href="/letter/?docId=nameregs/nameregs_677.xml">Crichton-Browne, James</a></h3> <div class="date">1840–1938</div> <div class="summary">Scottish physician, psychiatrist, and author. Graduated from the College of Surgeons, Edinburgh, in 1862. Medical director, West Riding Asylum, Wakefield, 1866–75. Lord chancellor’s visitor in lunacy, 1875–1922. Pioneering researcher into the causes and management of mental illness. Editor, West Riding Lunatic Asylum Medical Reports, 1871–5. One of the founders of Brain, 1878; co-editor, 1878–85. Author of a number of popular works on mental health and social policy. Knighted, 1886. FRS 1883.</div> </div> </div> </div> <div id="block-views-my-sidebar-block" class="block block-views"> <div> <div class="view view-my-sidebar view-id-my_sidebar view-display-id-block view-dom-id-507a53a7582c10348f5a49b107b660d1"> <div class="view-content"> <div class="views-row views-row-1 views-row-odd views-row-first views-row-last"> <div class="views-field views-field-field-side-text"> <div class="field-content"><h3>Resources</h3> <ul> <li><strong>Take part</strong> in our <a href="/emotion-experiment">online recreation of Darwin’s expression experiment</a></li> <li><strong>Read</strong> <a href="/tags/about-darwin/family-life/darwin-s-observations-his-children">Darwin’s notebook of observations on his children</a></li> <li><strong>See</strong> the responses from around the world to <a href="/expression-emotions/darwin-s-queries-expression">Darwin’s expression questionnaire</a></li> <li><strong>Listen</strong> to the talks on how Darwin’s work has influenced contemporary scientific research, from our event <a href="/expression-emotions/face-emotion">Face of Emotion</a></li> </ul> </div> </div> </div> </div> </div> </div> </div> </div> </div> </div> </div> <div class="campl-row campl-local-footer"> <div class="campl-wrap clearfix"> <div class="campl-column3 campl-footer-navigation"> <div class="region region-footer-1"> <div id="block-block-1" class="block block-block campl-content-container campl-navigation-list"> <div class="content"> <p>Darwin Correspondence Project<br /> <a href="mailto:darwin@lib.cam.ac.uk">darwin@lib.cam.ac.uk</a></p> <p>© <a href="http://www.cam.ac.uk/">University of Cambridge</a> 2022</p> <p><a href="/copyright-declaration">Copyright declaration</a></p> <p><a href="https://www.darwinproject.ac.uk/about/privacy-policy">Privacy policy</a></p> <p>Website by <a href="http://www.surfaceimpression.digital/">Surface Impression</a></p> </div> </div> </div> </div> <div class="campl-column3 campl-footer-navigation"> <div class="region 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