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Paraphylie — Wikipédia
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mw-ui-icon-wikimedia-expand"></span> <span>Afficher / masquer la sous-section Définitions</span> </button> <ul id="toc-Définitions-sublist" class="vector-toc-list"> <li id="toc-Sens_biologique" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Sens_biologique"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Sens biologique</span> </div> </a> <ul id="toc-Sens_biologique-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Propriétés_mathématiques" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Propriétés_mathématiques"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2</span> <span>Propriétés mathématiques</span> </div> </a> <ul id="toc-Propriétés_mathématiques-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Exemples" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Exemples"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Exemples</span> </div> </a> <ul id="toc-Exemples-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Statut_de_la_paraphylie" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Statut_de_la_paraphylie"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Statut de la paraphylie</span> </div> </a> <ul id="toc-Statut_de_la_paraphylie-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Voir_aussi" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Voir_aussi"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Voir aussi</span> </div> </a> <ul id="toc-Voir_aussi-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notes_et_références" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notes_et_références"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Notes et références</span> </div> </a> <ul id="toc-Notes_et_références-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Sommaire" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Basculer la table des matières" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Basculer la table des matières</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Paraphylie</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Aller à un article dans une autre langue. Disponible en 50 langues." > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-50" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">50 langues</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Parafileties" title="Parafileties – afrikaans" lang="af" hreflang="af" data-title="Parafileties" data-language-autonym="Afrikaans" data-language-local-name="afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-an mw-list-item"><a href="https://an.wikipedia.org/wiki/Parafilia_(filochenetica)" title="Parafilia (filochenetica) – aragonais" lang="an" hreflang="an" data-title="Parafilia (filochenetica)" data-language-autonym="Aragonés" data-language-local-name="aragonais" class="interlanguage-link-target"><span>Aragonés</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B4%D8%A8%D9%87_%D8%B9%D8%B1%D9%82" title="شبه عرق – arabe" lang="ar" hreflang="ar" data-title="شبه عرق" data-language-autonym="العربية" data-language-local-name="arabe" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Parafil%C3%A9ticu" title="Parafiléticu – asturien" lang="ast" hreflang="ast" data-title="Parafiléticu" data-language-autonym="Asturianu" data-language-local-name="asturien" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AA%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%B0%E0%A6%BE%E0%A6%AB%E0%A6%BE%E0%A6%87%E0%A6%B2%E0%A6%BF" title="প্যারাফাইলি – bengali" lang="bn" hreflang="bn" data-title="প্যারাফাইলি" data-language-autonym="বাংলা" data-language-local-name="bengali" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Parafileti%C4%8Dnost" title="Parafiletičnost – bosniaque" lang="bs" hreflang="bs" data-title="Parafiletičnost" data-language-autonym="Bosanski" data-language-local-name="bosniaque" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Parafiletisme" title="Parafiletisme – catalan" lang="ca" hreflang="ca" data-title="Parafiletisme" data-language-autonym="Català" data-language-local-name="catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Parafyletismus" title="Parafyletismus – tchèque" lang="cs" hreflang="cs" data-title="Parafyletismus" data-language-autonym="Čeština" data-language-local-name="tchèque" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Parafyletisk_gruppe" title="Parafyletisk gruppe – danois" lang="da" hreflang="da" data-title="Parafyletisk gruppe" data-language-autonym="Dansk" data-language-local-name="danois" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Paraphyly" title="Paraphyly – anglais" lang="en" hreflang="en" data-title="Paraphyly" data-language-autonym="English" data-language-local-name="anglais" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Parafiletika" title="Parafiletika – espéranto" lang="eo" hreflang="eo" data-title="Parafiletika" data-language-autonym="Esperanto" data-language-local-name="espéranto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Parafil%C3%A9tico" title="Parafilético – espagnol" lang="es" hreflang="es" data-title="Parafilético" data-language-autonym="Español" data-language-local-name="espagnol" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Paraf%C3%BCleetiline_r%C3%BChm" title="Parafüleetiline rühm – estonien" lang="et" hreflang="et" data-title="Parafüleetiline rühm" data-language-autonym="Eesti" data-language-local-name="estonien" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Parafiletiko" title="Parafiletiko – basque" lang="eu" hreflang="eu" data-title="Parafiletiko" data-language-autonym="Euskara" data-language-local-name="basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%BE%DB%8C%D8%B1%D8%A7%D8%AA%D8%A8%D8%A7%D8%B1" title="پیراتبار – persan" lang="fa" hreflang="fa" data-title="پیراتبار" data-language-autonym="فارسی" data-language-local-name="persan" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Parafyleettinen_ryhm%C3%A4" title="Parafyleettinen ryhmä – finnois" lang="fi" hreflang="fi" data-title="Parafyleettinen ryhmä" data-language-autonym="Suomi" data-language-local-name="finnois" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fy mw-list-item"><a href="https://fy.wikipedia.org/wiki/Parafyly" title="Parafyly – frison occidental" lang="fy" hreflang="fy" data-title="Parafyly" data-language-autonym="Frysk" data-language-local-name="frison occidental" class="interlanguage-link-target"><span>Frysk</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Paraif%C3%ADl%C3%A9iteach" title="Paraifíléiteach – irlandais" lang="ga" hreflang="ga" data-title="Paraifíléiteach" data-language-autonym="Gaeilge" data-language-local-name="irlandais" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Parafil%C3%A9tico" title="Parafilético – galicien" lang="gl" hreflang="gl" data-title="Parafilético" data-language-autonym="Galego" data-language-local-name="galicien" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A4%D7%A8%D7%A4%D7%99%D7%9C%D7%98%D7%99%D7%95%D7%AA" title="פרפילטיות – hébreu" lang="he" hreflang="he" data-title="פרפילטיות" data-language-autonym="עברית" data-language-local-name="hébreu" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%AA%E0%A4%BE%E0%A4%B0%E0%A4%B5%E0%A4%B0%E0%A5%8D%E0%A4%97%E0%A5%80" title="पारवर्गी – hindi" lang="hi" hreflang="hi" data-title="पारवर्गी" data-language-autonym="हिन्दी" data-language-local-name="hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Parafiletikus_csoport" title="Parafiletikus csoport – hongrois" lang="hu" hreflang="hu" data-title="Parafiletikus csoport" data-language-autonym="Magyar" data-language-local-name="hongrois" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ia mw-list-item"><a href="https://ia.wikipedia.org/wiki/Paraphylia" title="Paraphylia – interlingua" lang="ia" hreflang="ia" data-title="Paraphylia" data-language-autonym="Interlingua" data-language-local-name="interlingua" class="interlanguage-link-target"><span>Interlingua</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Parafili" title="Parafili – indonésien" lang="id" hreflang="id" data-title="Parafili" data-language-autonym="Bahasa Indonesia" data-language-local-name="indonésien" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Parafilia_(filogenesi)" title="Parafilia (filogenesi) – italien" lang="it" hreflang="it" data-title="Parafilia (filogenesi)" data-language-autonym="Italiano" data-language-local-name="italien" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%81%B4%E7%B3%BB%E7%B5%B1%E7%BE%A4" title="側系統群 – japonais" lang="ja" hreflang="ja" data-title="側系統群" data-language-autonym="日本語" data-language-local-name="japonais" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Parafiletik" title="Parafiletik – javanais" lang="jv" hreflang="jv" data-title="Parafiletik" data-language-autonym="Jawa" data-language-local-name="javanais" class="interlanguage-link-target"><span>Jawa</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%B8%A1%EA%B3%84%ED%86%B5%EA%B5%B0" title="측계통군 – coréen" lang="ko" hreflang="ko" data-title="측계통군" data-language-autonym="한국어" data-language-local-name="coréen" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-li mw-list-item"><a href="https://li.wikipedia.org/wiki/Parafyletisch_taxon" title="Parafyletisch taxon – limbourgeois" lang="li" hreflang="li" data-title="Parafyletisch taxon" data-language-autonym="Limburgs" data-language-local-name="limbourgeois" class="interlanguage-link-target"><span>Limburgs</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Parafili" title="Parafili – malais" lang="ms" hreflang="ms" data-title="Parafili" data-language-autonym="Bahasa Melayu" data-language-local-name="malais" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Parafyletische_groep" title="Parafyletische groep – néerlandais" lang="nl" hreflang="nl" data-title="Parafyletische groep" data-language-autonym="Nederlands" data-language-local-name="néerlandais" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Parafyletisk_gruppe" title="Parafyletisk gruppe – norvégien nynorsk" lang="nn" hreflang="nn" data-title="Parafyletisk gruppe" data-language-autonym="Norsk nynorsk" data-language-local-name="norvégien nynorsk" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Parafyletisk_gruppe" title="Parafyletisk gruppe – norvégien bokmål" lang="nb" hreflang="nb" data-title="Parafyletisk gruppe" data-language-autonym="Norsk bokmål" data-language-local-name="norvégien bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Takson_parafiletyczny" title="Takson parafiletyczny – polonais" lang="pl" hreflang="pl" data-title="Takson parafiletyczny" data-language-autonym="Polski" data-language-local-name="polonais" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Parafil%C3%A9tico" title="Parafilético – portugais" lang="pt" hreflang="pt" data-title="Parafilético" data-language-autonym="Português" data-language-local-name="portugais" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Grup_parafiletic" title="Grup parafiletic – roumain" lang="ro" hreflang="ro" data-title="Grup parafiletic" data-language-autonym="Română" data-language-local-name="roumain" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D0%B0%D1%80%D0%B0%D1%84%D0%B8%D0%BB%D0%B8%D1%8F_(%D1%82%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%BE%D0%BC%D0%B8%D1%8F)" title="Парафилия (таксономия) – russe" lang="ru" hreflang="ru" data-title="Парафилия (таксономия)" data-language-autonym="Русский" data-language-local-name="russe" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Parafileti%C4%8Dnost" title="Parafiletičnost – serbo-croate" lang="sh" hreflang="sh" data-title="Parafiletičnost" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="serbo-croate" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Paraphyly" title="Paraphyly – Simple English" lang="en-simple" hreflang="en-simple" data-title="Paraphyly" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Parafyletizmus" title="Parafyletizmus – slovaque" lang="sk" hreflang="sk" data-title="Parafyletizmus" data-language-autonym="Slovenčina" data-language-local-name="slovaque" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Parafilija" title="Parafilija – slovène" lang="sl" hreflang="sl" data-title="Parafilija" data-language-autonym="Slovenščina" data-language-local-name="slovène" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Parafileti%C4%8Dnost" title="Parafiletičnost – serbe" lang="sr" hreflang="sr" data-title="Parafiletičnost" data-language-autonym="Српски / srpski" data-language-local-name="serbe" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-stq mw-list-item"><a href="https://stq.wikipedia.org/wiki/Paraphylie" title="Paraphylie – saterlandais" lang="stq" hreflang="stq" data-title="Paraphylie" data-language-autonym="Seeltersk" data-language-local-name="saterlandais" class="interlanguage-link-target"><span>Seeltersk</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Parafyli" title="Parafyli – suédois" lang="sv" hreflang="sv" data-title="Parafyli" data-language-autonym="Svenska" data-language-local-name="suédois" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Parafili_(filogenetik)" title="Parafili (filogenetik) – turc" lang="tr" hreflang="tr" data-title="Parafili (filogenetik)" data-language-autonym="Türkçe" data-language-local-name="turc" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9F%D0%B0%D1%80%D0%B0%D1%84%D1%96%D0%BB%D1%96%D1%8F" title="Парафілія – ukrainien" lang="uk" hreflang="uk" data-title="Парафілія" data-language-autonym="Українська" data-language-local-name="ukrainien" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/C%E1%BA%ADn_ng%C3%A0nh" title="Cận ngành – vietnamien" lang="vi" hreflang="vi" data-title="Cận ngành" data-language-autonym="Tiếng Việt" data-language-local-name="vietnamien" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wa mw-list-item"><a href="https://wa.wikipedia.org/wiki/Parafiletike" title="Parafiletike – wallon" lang="wa" hreflang="wa" data-title="Parafiletike" data-language-autonym="Walon" data-language-local-name="wallon" class="interlanguage-link-target"><span>Walon</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E4%B8%A6%E7%B3%BB%E7%BE%A4" title="並系群 – chinois" lang="zh" hreflang="zh" data-title="並系群" data-language-autonym="中文" data-language-local-name="chinois" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E4%B8%A6%E7%B3%BB%E7%BE%A4" title="並系群 – cantonais" lang="yue" hreflang="yue" data-title="並系群" data-language-autonym="粵語" data-language-local-name="cantonais" class="interlanguage-link-target"><span>粵語</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Apparence</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">déplacer vers la barre latérale</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">masquer</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">Un article de Wikipédia, l'encyclopédie libre.</div> </div> <div id="contentSub"><div id="mw-content-subtitle"><span class="mw-redirectedfrom">(Redirigé depuis <a href="/w/index.php?title=Paraphyl%C3%A9tique&redirect=no" class="mw-redirect" title="Paraphylétique">Paraphylétique</a>)</span></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="fr" dir="ltr"><div class="bandeau-container metadata homonymie hatnote"><div class="bandeau-cell bandeau-icone" style="display:table-cell;padding-right:0.5em"><span class="noviewer" typeof="mw:File"><a href="/wiki/Aide:Homonymie" title="Aide:Homonymie"><img alt="Page d’aide sur l’homonymie" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Logo_disambig-homophone.svg/20px-Logo_disambig-homophone.svg.png" decoding="async" width="20" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Logo_disambig-homophone.svg/30px-Logo_disambig-homophone.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/44/Logo_disambig-homophone.svg/40px-Logo_disambig-homophone.svg.png 2x" data-file-width="512" data-file-height="375" /></a></span></div><div class="bandeau-cell" style="display:table-cell;padding-right:0.5em"> <p>Pour l’article ayant un titre homophone, voir <a href="/wiki/Paraphilie" title="Paraphilie">Paraphilie</a>. </p> </div></div> <div class="bandeau-container metadata bandeau-article bandeau-niveau-modere"><figure class="mw-halign-right noviewer" typeof="mw:File"><a href="/wiki/Mod%C3%A8le:%C3%80_sourcer" title="Si ce bandeau n'est plus pertinent, retirez-le. Cliquez ici pour en savoir plus."><img alt="Si ce bandeau n'est plus pertinent, retirez-le. Cliquez ici pour en savoir plus." src="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Info_Simple.svg/12px-Info_Simple.svg.png" decoding="async" width="12" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Info_Simple.svg/18px-Info_Simple.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/38/Info_Simple.svg/24px-Info_Simple.svg.png 2x" data-file-width="512" data-file-height="512" /></a><figcaption>Si ce bandeau n'est plus pertinent, retirez-le. Cliquez ici pour en savoir plus.</figcaption></figure><div class="bandeau-cell bandeau-icone" style="display:table-cell;padding-right:0.5em"><span class="noviewer" typeof="mw:File"><a href="/wiki/Fichier:2017-fr.wp-orange-source.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a1/2017-fr.wp-orange-source.svg/45px-2017-fr.wp-orange-source.svg.png" decoding="async" width="45" height="45" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a1/2017-fr.wp-orange-source.svg/68px-2017-fr.wp-orange-source.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a1/2017-fr.wp-orange-source.svg/90px-2017-fr.wp-orange-source.svg.png 2x" data-file-width="512" data-file-height="512" /></a></span></div><div class="bandeau-cell" style="display:table-cell;padding-right:0.5em"> <p><strong class="bandeau-titre">Cet article <a href="/wiki/Wikip%C3%A9dia:Citez_vos_sources" title="Wikipédia:Citez vos sources">ne cite pas suffisamment ses sources</a></strong> <small>(<time class="nowrap" datetime="2013-02" data-sort-value="2013-02">février 2013</time>).</small> </p><p>Si vous disposez d'ouvrages ou d'articles de référence ou si vous connaissez des sites web de qualité traitant du thème abordé ici, merci de compléter l'article en donnant les <b>références utiles à sa <a href="/wiki/Wikip%C3%A9dia:V%C3%A9rifiabilit%C3%A9" title="Wikipédia:Vérifiabilité">vérifiabilité</a></b> et en les liant à la section « <a href="/wiki/Aide:Ins%C3%A9rer_une_r%C3%A9f%C3%A9rence" title="Aide:Insérer une référence">Notes et références</a> ». </p><p><b>En pratique :</b> <a href="/wiki/Wikip%C3%A9dia:Citez_vos_sources#Qualité_des_sources" title="Wikipédia:Citez vos sources">Quelles sources sont attendues ?</a> <a href="/wiki/Aide:Ins%C3%A9rer_une_r%C3%A9f%C3%A9rence_(%C3%89diteur_visuel)" title="Aide:Insérer une référence (Éditeur visuel)">Comment ajouter mes sources ?</a> </p> </div></div> <figure typeof="mw:File/Thumb"><a href="/wiki/Fichier:Traditional_Reptilia.jpg" class="mw-file-description"><img alt="Arbre d’évolution. Ce groupe comprend un ancêtre et une partie de ses descendants mais pas tous puisque les oiseaux et les mammifères en sont exclus. C'est un groupe paraphylétique de degré deux." src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Traditional_Reptilia.jpg/350px-Traditional_Reptilia.jpg" decoding="async" width="350" height="282" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Traditional_Reptilia.jpg/525px-Traditional_Reptilia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Traditional_Reptilia.jpg/700px-Traditional_Reptilia.jpg 2x" data-file-width="2439" data-file-height="1967" /></a><figcaption>Le groupe traditionnel paraphylétique des <a href="/wiki/Reptile" class="mw-redirect" title="Reptile">reptiles</a> apparaît en vert. Ce groupe comprend un ancêtre et une partie de ses descendants mais pas tous puisque les <a href="/wiki/Oiseau" title="Oiseau">oiseaux</a> et les <a href="/wiki/Mammif%C3%A8re" title="Mammifère">mammifères</a> en sont exclus. C'est un groupe paraphylétique de degré deux. </figcaption></figure> <p>En <a href="/wiki/Syst%C3%A9matique" title="Systématique">systématique</a>, un groupe est dit <b>paraphylétique</b> quand il ne rassemble pas tous les descendants d'une <a href="/wiki/Souche_(biologie)" title="Souche (biologie)">espèce souche</a> qu'il contient<sup id="cite_ref-aub15_1-0" class="reference"><a href="#cite_note-aub15-1"><span class="cite_crochet">[</span>1<span class="cite_crochet">]</span></a></sup>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Définitions"><span id="D.C3.A9finitions"></span>Définitions</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paraphylie&veaction=edit&section=1" title="Modifier la section : Définitions" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paraphylie&action=edit&section=1" title="Modifier le code source de la section : Définitions"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Sens_biologique">Sens biologique</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paraphylie&veaction=edit&section=2" title="Modifier la section : Sens biologique" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paraphylie&action=edit&section=2" title="Modifier le code source de la section : Sens biologique"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Contrairement à la <a href="/wiki/Polyphylie" title="Polyphylie">polyphylie</a>, le groupe paraphylétique est fondé sur une similitude héritée d'un ancêtre commun (<a href="/wiki/Homologie_(%C3%A9volution)" title="Homologie (évolution)">homologie</a>).<sup class="need_ref_tag" style="padding-left:2px;"><a href="/wiki/Aide:R%C3%A9f%C3%A9rence_n%C3%A9cessaire" title="Aide:Référence nécessaire">[<abbr class="abbr" title="référence">réf.</abbr> souhaitée]</a></sup> L'exclusion de certaines lignées nécessite de considérer les <a href="/wiki/Caract%C3%A8re_ancestral_et_caract%C3%A8re_d%C3%A9riv%C3%A9" title="Caractère ancestral et caractère dérivé">états ancestraux</a> de certains caractères (encore appelé <a href="/wiki/Sympl%C3%A9siomorphie" title="Symplésiomorphie">symplésiomorphie</a>) par opposition à leurs <a href="/wiki/Caract%C3%A8re_ancestral_et_caract%C3%A8re_d%C3%A9riv%C3%A9" title="Caractère ancestral et caractère dérivé">états dérivés</a> (ou <a href="/wiki/Synapomorphie" title="Synapomorphie">synapomorphie</a>) dans les lignées exclues. </p><p>Le groupe des <a href="/wiki/Reptiles" title="Reptiles">reptiles</a> (<a href="/wiki/Amniota" title="Amniota">amniotes</a> desquels ont été exclus les <a href="/wiki/Oiseau" title="Oiseau">oiseaux</a> et les <a href="/wiki/Mammif%C3%A8re" title="Mammifère">mammifères</a>) en est un exemple : les <a href="/wiki/Crocodilia" title="Crocodilia">crocodiliens</a>, inclus dans les reptiles, sont généalogiquement plus proches des <a href="/wiki/Oiseau" title="Oiseau">oiseaux</a>, exclus des reptiles, que des autres <a href="/wiki/Sauropsida" title="Sauropsida">sauropsides</a> (<a href="/wiki/Tortue" title="Tortue">chéloniens</a>, <a href="/wiki/Squamata" title="Squamata">squamates</a> et <a href="/wiki/Rhynchocephalia" title="Rhynchocephalia">sphénodontiens</a>) et les reptiles mammaliens, inclus, sont plus proches des mammifères, exclus, que des autres <a href="/wiki/Amniota" title="Amniota">amniotes</a>. L'ancêtre commun le plus proche de tous les reptiles est également l'ancêtre commun le plus proche des oiseaux et des mammifères. Le groupe des reptiles ainsi défini est donc paraphylétique. </p> <pre> <a href="/wiki/Amniota" title="Amniota">amniotes</a> ├─<a href="/wiki/Synapsida" title="Synapsida">Synapsides</a> │ ├─<a href="/wiki/Mammif%C3%A8re" title="Mammifère">Mammifères</a> │ └─<a href="/wiki/Synapsida#Les_«_reptiles_mammaliens_»" title="Synapsida">Reptiles mammaliens</a>----------| | └─<a href="/wiki/Sauropsida" title="Sauropsida">Sauropsides</a> ├─<b><a href="/wiki/Tortue" title="Tortue">Chéloniens</a></b>-------------------| │ | « <b><a href="/wiki/Reptiles" title="Reptiles">Reptiles</a></b> » └─<a href="/wiki/Diapsida" title="Diapsida">Diapsides</a>--------------------| ├─<b><a href="/wiki/Lepidosauria" title="Lepidosauria">Lépidosauriens</a></b>-----------| │ ├─<b><a href="/wiki/Squamata" title="Squamata">Squamates</a></b>-----------| │ └─<b><a href="/wiki/Rhynchocephalia" title="Rhynchocephalia">Sphénodontiens</a></b>------| │ | └─<a href="/wiki/Archosauria" title="Archosauria">Archosauriens</a> | ├─<b><a href="/wiki/Crocodilia" title="Crocodilia">Crocodiliens</a></b>--------| └─<a href="/wiki/Oiseau" title="Oiseau">Oiseaux</a> </pre> <p><span style="text-decoration: underline;">Note</span> : Les <a href="/wiki/Crustac%C3%A9s" title="Crustacés">crustacés</a>, les <a href="/wiki/Amphibia" title="Amphibia">amphibiens</a> ou les <a href="/wiki/Poisson" title="Poisson">poissons</a> sont d'autres exemples de groupes paraphylétiques. </p> <div class="mw-heading mw-heading3"><h3 id="Propriétés_mathématiques"><span id="Propri.C3.A9t.C3.A9s_math.C3.A9matiques"></span>Propriétés mathématiques</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paraphylie&veaction=edit&section=3" title="Modifier la section : Propriétés mathématiques" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paraphylie&action=edit&section=3" title="Modifier le code source de la section : Propriétés mathématiques"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>En <a href="/wiki/Th%C3%A9orie_des_graphes" title="Théorie des graphes">théorie des graphes</a>, on peut définir le « degré de paraphylie » d'un groupe comme étant le nombre minimum de branches qu'il faudrait retirer à un groupe <a href="/wiki/Monophylie" title="Monophylie">holophylétique</a><sup id="cite_ref-aub15_1-1" class="reference"><a href="#cite_note-aub15-1"><span class="cite_crochet">[</span>1<span class="cite_crochet">]</span></a></sup>. Un groupe holophylétique a donc un degré de paraphylie de zéro. </p><p>La définition traditionnelle des reptiles (<a href="/wiki/Sauropsida" title="Sauropsida">sauropsides</a> + <a href="/wiki/Synapsida" title="Synapsida">synapsides</a>) en fait un groupe ayant un degré de paraphylie de 2, puisqu'il faut retirer au groupe des <a href="/wiki/Amniota" title="Amniota">amniotes</a> la branche des <a href="/wiki/Mammif%C3%A8re" title="Mammifère">mammifères</a> et celle des <a href="/wiki/Oiseau" title="Oiseau">oiseaux</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Exemples">Exemples</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paraphylie&veaction=edit&section=4" title="Modifier la section : Exemples" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paraphylie&action=edit&section=4" title="Modifier le code source de la section : Exemples"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Les <a href="/wiki/Prokaryota" title="Prokaryota">procaryotes</a> (excluent les <a href="/wiki/Eukaryota" title="Eukaryota">eucaryotes</a>)</li> <li>Les <a href="/wiki/Micro-organisme" title="Micro-organisme">microorganismes</a></li> <li>Les <a href="/wiki/Pr%C3%A9spermaphytes" title="Préspermaphytes">préspermaphytes</a></li> <li>Les <a href="/wiki/Herbe" title="Herbe">herbes</a></li> <li>Les <a href="/wiki/Invert%C3%A9br%C3%A9" title="Invertébré">invertébrés</a></li> <li>Les <a href="/wiki/Crustac%C3%A9s" title="Crustacés">crustacés</a> (excluent les <a href="/wiki/Insecte" title="Insecte">insectes</a>)</li> <li>Les <a href="/wiki/Poisson" title="Poisson">poissons</a> (excluent les <a href="/wiki/Tetrapoda" title="Tetrapoda">tétrapodes</a>)</li> <li>Les <a href="/wiki/Amphibia" title="Amphibia">amphibiens</a> (excluent les <a href="/wiki/Amniota" title="Amniota">amniotes</a>)</li> <li>Les <a href="/wiki/Reptile" class="mw-redirect" title="Reptile">reptiles</a> (excluent les <a href="/wiki/Oiseau" title="Oiseau">oiseaux</a> et les mammifères)</li> <li>Les <a href="/wiki/Antilopes" title="Antilopes">antilopes</a> (excluent les <a href="/wiki/Bovini" title="Bovini">bovins</a> et <a href="/wiki/Caprinae" title="Caprinae">caprins</a>)</li> <li>Les reptiles mammaliens (incluent tous les <a href="/wiki/Synapsida" title="Synapsida">synapsides</a> sauf les mammifères, qui en sont exclus)</li> <li>Les <a href="/wiki/Micromammif%C3%A8re" title="Micromammifère">micromammifères</a></li> <li>Les <a href="/wiki/Renard" title="Renard">renards</a></li> <li>Les <a href="/wiki/Singe" title="Singe">singes</a> (Débat concernant l'inclusion, ou non, de l'humain dans ce groupe ; il peut, donc, être aussi vu comme <a href="/wiki/Monophylie" title="Monophylie">monophylétique</a>, ce qui va donner au groupe des <a href="/wiki/Simien" class="mw-disambig" title="Simien">simiens</a>).</li></ul> <div class="mw-heading mw-heading2"><h2 id="Statut_de_la_paraphylie">Statut de la paraphylie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paraphylie&veaction=edit&section=5" title="Modifier la section : Statut de la paraphylie" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paraphylie&action=edit&section=5" title="Modifier le code source de la section : Statut de la paraphylie"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Pour les <a href="/wiki/Classification_phylog%C3%A9n%C3%A9tique" title="Classification phylogénétique">cladistes</a>, les <a href="/wiki/Taxon" title="Taxon">taxons</a> paraphylétiques sont inacceptables dans la classification. Au contraire pour les <a href="/wiki/Syst%C3%A9matique_%C3%A9volutionniste" title="Systématique évolutionniste">évolutionnistes</a> il n'y a aucun problème dans le fait d'accepter de tels <a href="/wiki/Taxon" title="Taxon">taxons</a>. Selon la définition qui en est donnée (voir l'article <a href="/wiki/Monophylie" title="Monophylie">Monophylie</a>), la paraphylie est parfois considérée comme une forme de <a href="/wiki/Monophylie" title="Monophylie">monophylie</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Voir_aussi">Voir aussi</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paraphylie&veaction=edit&section=6" title="Modifier la section : Voir aussi" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paraphylie&action=edit&section=6" title="Modifier le code source de la section : Voir aussi"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Termes associés : </p> <ul><li><a href="/wiki/H%C3%A9t%C3%A9rophylie" title="Hétérophylie">hétérophylétique</a> ;</li> <li><a href="/wiki/Monophylie" title="Monophylie">holophylétique et monophylétique</a> ;</li> <li><a href="/wiki/Polyphylie" title="Polyphylie">polyphylétique</a>.</li></ul> <p>Concepts associés : </p> <ul><li><a href="/wiki/Taxonomie" title="Taxonomie">taxinomie</a> ;</li> <li><a href="/wiki/Taxon_poubelle" title="Taxon poubelle">taxon poubelle</a> ;</li> <li><a href="/wiki/Arbre_phylog%C3%A9n%C3%A9tique" title="Arbre phylogénétique">arbre phylogénétique</a> ;</li> <li><a href="/wiki/Syst%C3%A9matique_%C3%A9volutionniste" title="Systématique évolutionniste">évolutionnisme (biologie)</a> ;</li> <li><a href="/wiki/Classification_phylog%C3%A9n%C3%A9tique" title="Classification phylogénétique">cladisme</a>.</li></ul> <div class="mw-heading mw-heading2"><h2 id="Notes_et_références"><span id="Notes_et_r.C3.A9f.C3.A9rences"></span>Notes et références</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paraphylie&veaction=edit&section=7" title="Modifier la section : Notes et références" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paraphylie&action=edit&section=7" title="Modifier le code source de la section : Notes et références"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="references-small decimal" style=""><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-aub15-1"><span class="mw-cite-backlink noprint">↑ <sup><a href="#cite_ref-aub15_1-0">a</a> et <a href="#cite_ref-aub15_1-1">b</a></sup> </span><span class="reference-text">Aubert, D. 2015. A formal analysis of phylogenetic terminology: Towards a reconsideration of the current paradigm in systematics. <i>Phytoneuron</i> <b>2015-66</b>:1–54. <a rel="nofollow" class="external free" href="https://hal.archives-ouvertes.fr/hal-01240878v1">https://hal.archives-ouvertes.fr/hal-01240878v1</a></span> </li> </ol></div> </div> <ul id="bandeau-portail" class="bandeau-portail"><li><span class="bandeau-portail-element"><span class="bandeau-portail-icone"><span class="noviewer" typeof="mw:File"><a href="/wiki/Portail:Origine_et_%C3%A9volution_du_vivant" title="Portail origine et évolution du vivant"><img alt="icône décorative" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e2/Icone_OEV2.jpg/48px-Icone_OEV2.jpg" decoding="async" width="48" height="23" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e2/Icone_OEV2.jpg/72px-Icone_OEV2.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e2/Icone_OEV2.jpg/96px-Icone_OEV2.jpg 2x" data-file-width="170" data-file-height="83" /></a></span></span> <span class="bandeau-portail-texte"><a href="/wiki/Portail:Origine_et_%C3%A9volution_du_vivant" title="Portail:Origine et évolution du vivant">Portail origine et évolution du vivant</a></span> </span></li> </ul> <!-- NewPP limit report Parsed by mw‐web.eqiad.main‐6f4956b788‐7gsw9 Cached time: 20241128184305 Cache expiry: 2592000 Reduced expiry: false Complications: [show‐toc] CPU time usage: 0.138 seconds Real time usage: 0.437 seconds Preprocessor visited node count: 582/1000000 Post‐expand include size: 10327/2097152 bytes Template argument size: 690/2097152 bytes Highest expansion depth: 12/100 Expensive parser function count: 1/500 Unstrip recursion depth: 0/20 Unstrip post‐expand size: 1041/5000000 bytes Lua time usage: 0.046/10.000 seconds Lua memory usage: 1785087/52428800 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 152.849 1 -total 42.57% 65.063 1 Modèle:Portail 26.17% 40.004 1 Modèle:Homophonie 23.22% 35.488 1 Modèle:Méta_bandeau_de_note 21.93% 33.521 1 Modèle:Méta_bandeau 21.55% 32.932 1 Modèle:Catégorisation_badges 18.63% 28.480 1 Modèle:À_sourcer 14.47% 22.123 1 Modèle:Suivi_des_biographies 9.80% 14.976 1 Modèle:Méta_bandeau_d'avertissement 7.01% 10.713 1 Modèle:Contrôle_date_bandeau --> <!-- Saved in parser cache with key frwiki:pcache:173695:|#|:idhash:canonical and timestamp 20241128184305 and revision id 220066725. 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