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"https://www.academia.edu/login?post_login_redirect_url=https%3A%2F%2Fwww.academia.edu%2F110076275%2FTheropod_Tooth_from_the_Upper_Cretaceous_Himenoura_Group_in_the_Koshikijima_Islands_Southwestern_Japan%3Fauto%3Ddownload"; window.loswp.translateUrl = "https://www.academia.edu/login?post_login_redirect_url=https%3A%2F%2Fwww.academia.edu%2F110076275%2FTheropod_Tooth_from_the_Upper_Cretaceous_Himenoura_Group_in_the_Koshikijima_Islands_Southwestern_Japan%3Fshow_translation%3Dtrue"; window.loswp.previewableAttachments = [{"id":108006187,"identifier":"Attachment_108006187","shouldShowBulkDownload":false}]; window.loswp.shouldDetectTimezone = true; window.loswp.shouldShowBulkDownload = true; window.loswp.showSignupCaptcha = false window.loswp.willEdgeCache = false; window.loswp.work = {"work":{"id":110076275,"created_at":"2023-11-28T18:54:32.528-08:00","from_world_paper_id":244263499,"updated_at":"2024-11-25T00:06:16.069-08:00","_data":{"publisher":"BioOne (Paleontological Society of Japan)","grobid_abstract":"A theropod tooth found in the Upper Cretaceous Himenoura Group in the Koshikijima Islands, Kagoshima Prefecture, is described. Recent paleontological work in the region suggests a middle Campanian age for the specimen. Lacking a feature diagnostic of any specific theropod clade, this specimen is considered merely as Theropoda incertae sedis. The present finding confirms the potential of the Himenoura Group as a new source of information on the diversity of Theropoda during the Campanian.","publication_date":"2013,4,1","publication_name":"Paleontological Research","grobid_abstract_attachment_id":"108006187"},"document_type":"paper","pre_hit_view_count_baseline":null,"quality":"high","language":"en","title":"Theropod Tooth from the Upper Cretaceous Himenoura Group in the Koshikijima Islands, Southwestern Japan","broadcastable":false,"draft":null,"has_indexable_attachment":true,"indexable":true}}["work"]; window.loswp.workCoauthors = [39916965]; window.loswp.locale = "en"; window.loswp.countryCode = "SG"; window.loswp.cwvAbTestBucket = ""; window.loswp.designVariant = "ds_vanilla"; window.loswp.fullPageMobileSutdModalVariant = "control"; window.loswp.useOptimizedScribd4genScript = false; window.loswp.appleClientId = 'edu.academia.applesignon';</script><script defer="" src="https://accounts.google.com/gsi/client"></script><div 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data-landing_url="https://www.academia.edu/110076275/Theropod_Tooth_from_the_Upper_Cretaceous_Himenoura_Group_in_the_Koshikijima_Islands_Southwestern_Japan" data-login_uri="https://www.academia.edu/registrations/google_one_tap" data-moment_callback="onGoogleOneTapEvent" id="g_id_onload"></div><div class="ds-top-related-works--grid-container"><div class="ds-related-content--container ds-top-related-works--container"><h2 class="ds-related-content--heading">Related papers</h2><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="0" data-entity-id="101318477" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/101318477/A_diverse_theropod_footprint_assemblage_from_the_Lower_Cretaceous_Kitadani_Formation_Tetori_Group_central_Japan">A diverse theropod footprint assemblage from the Lower Cretaceous Kitadani Formation, Tetori Group, central Japan</a><div 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Nearly all these discoveries are referable to the previously known coelurosaurian subgroups. Here, we report a new theropod, Fukuivenator paradoxus, gen. et sp. nov., based on a nearly complete specimen from the Lower Cretaceous Kitadani Formation of the Tetori Group, Fukui, Japan. While Fukuivenator possesses a large number of morphological features unknown in any other theropod, it has a combination of primitive and derived features seen in different theropod subgroups, notably dromaeosaurid dinosaurs. Computed-tomography data indicate that Fukuivenator possesses inner ears whose morphology is intermediate between those of birds and non-avian dinosaurs. Our phylogenetic analysis recovers Fukuivenator as a basally branching maniraptoran theropod, yet is unable to refer it to any known coelurosaurian subgroups. The disc...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A bizarre theropod from the Early Cretaceous of Japan highlighting mosaic evolution among coelurosaurians","attachmentId":81755501,"attachmentType":"pdf","work_url":"https://www.academia.edu/73101309/A_bizarre_theropod_from_the_Early_Cretaceous_of_Japan_highlighting_mosaic_evolution_among_coelurosaurians","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/73101309/A_bizarre_theropod_from_the_Early_Cretaceous_of_Japan_highlighting_mosaic_evolution_among_coelurosaurians"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="5" data-entity-id="10010828" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/10010828/A_theropod_dinosaur_Saurischia_Maniraptora_from_the_Upper_Cretaceous_Yezo_Group_of_Hokkaido_Northeastern_Japan">A theropod dinosaur (Saurischia: Maniraptora) from the Upper Cretaceous Yezo Group　of Hokkaido, Northeastern Japan</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="53125158" href="https://independent.academia.edu/%E7%96%8B%E5%90%89%E8%AD%98">疋. 吉識</a><span>, </span><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="23984245" href="https://independent.academia.edu/%E5%BB%89%E5%B9%B3%E5%B1%B1">Ren Hirayama</a></div><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"A theropod dinosaur (Saurischia: Maniraptora) from the Upper Cretaceous Yezo Group　of Hokkaido, Northeastern Japan","attachmentId":36147331,"attachmentType":"pdf","work_url":"https://www.academia.edu/10010828/A_theropod_dinosaur_Saurischia_Maniraptora_from_the_Upper_Cretaceous_Yezo_Group_of_Hokkaido_Northeastern_Japan","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/10010828/A_theropod_dinosaur_Saurischia_Maniraptora_from_the_Upper_Cretaceous_Yezo_Group_of_Hokkaido_Northeastern_Japan"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="6" data-entity-id="52474826" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/52474826/New_specimens_including_a_growth_series_of_Fukuiraptor_Dinosauria_Theropoda_from_the_Lower_Cretaceous_Kitadani_Quarry_of_Japan">New specimens, including a growth series, of Fukuiraptor (Dinosauria, Theropoda) from the Lower Cretaceous Kitadani Quarry of Japan</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="683178" href="https://ualberta.academia.edu/PhilipJCurrie">Philip J. Currie</a></div><p class="ds-related-work--metadata ds2-5-body-xs">JOURNAL-PALEONTOLOGICAL SOCIETY …, 2006</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"New specimens, including a growth series, of Fukuiraptor (Dinosauria, Theropoda) from the Lower Cretaceous Kitadani Quarry of Japan","attachmentId":69724054,"attachmentType":"pdf","work_url":"https://www.academia.edu/52474826/New_specimens_including_a_growth_series_of_Fukuiraptor_Dinosauria_Theropoda_from_the_Lower_Cretaceous_Kitadani_Quarry_of_Japan","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/52474826/New_specimens_including_a_growth_series_of_Fukuiraptor_Dinosauria_Theropoda_from_the_Lower_Cretaceous_Kitadani_Quarry_of_Japan"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="7" data-entity-id="69978540" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/69978540/An_unusual_bird_Theropoda_Avialae_from_the_Early_Cretaceous_of_Japan_suggests_complex_evolutionary_history_of_basal_birds">An unusual bird (Theropoda, Avialae) from the Early Cretaceous of Japan suggests complex evolutionary history of basal birds</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="924143" href="https://independent.academia.edu/SoichiroKawabe">Soichiro Kawabe</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Communications Biology</p><p class="ds-related-work--abstract ds2-5-body-sm">The Early Cretaceous basal birds were known largely from just two-dimensionally preserved specimens from north-eastern China (Jehol Biota), which has hindered our understanding of the early evolution of birds. Here, we present a three-dimensionally-preserved skeleton (FPDM-V-9769) of a basal bird from the Early Cretaceous of Fukui, central Japan. Unique features in the pygostyle and humerus allow the assignment of FPDM-V-9769 to a new taxon, Fukuipteryx prima. FPDM-V-9769 exhibits a set of features comparable to that of other basalmost birds including Archaeopteryx. Osteohistological analyses indicate that FPDM-V-9769 is subadult. Phylogenetic analyses resolve F. prima as a non-ornithothoracine avialan basal to Jeholornis and outgroup of the Pygostylia. This phylogenetic result may imply a complex evolutionary history of basal birds. To our knowledge, FPDM-V-9769 represents the first record of the Early Cretaceous non-ornithothoracine avialan outside of the Jehol Biota and increases...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"An unusual bird (Theropoda, Avialae) from the Early Cretaceous of Japan suggests complex evolutionary history of basal birds","attachmentId":79874646,"attachmentType":"pdf","work_url":"https://www.academia.edu/69978540/An_unusual_bird_Theropoda_Avialae_from_the_Early_Cretaceous_of_Japan_suggests_complex_evolutionary_history_of_basal_birds","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/69978540/An_unusual_bird_Theropoda_Avialae_from_the_Early_Cretaceous_of_Japan_suggests_complex_evolutionary_history_of_basal_birds"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="8" data-entity-id="86193156" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/86193156/An_Early_Cretaceous_Mammal_from_the_Kuwajima_Formation_Tetori_Group_Japan_and_a_Reassessment_of_Triconodont_Phylogeny">An Early Cretaceous Mammal from the Kuwajima Formation (Tetori Group), Japan, and a Reassessment of Triconodont Phylogeny</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="34164059" href="https://independent.academia.edu/GuillermoRougier">Guillermo Rougier</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Annals of Carnegie Museum, 2007</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"An Early Cretaceous Mammal from the Kuwajima Formation (Tetori Group), Japan, and a Reassessment of Triconodont Phylogeny","attachmentId":90702535,"attachmentType":"pdf","work_url":"https://www.academia.edu/86193156/An_Early_Cretaceous_Mammal_from_the_Kuwajima_Formation_Tetori_Group_Japan_and_a_Reassessment_of_Triconodont_Phylogeny","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/86193156/An_Early_Cretaceous_Mammal_from_the_Kuwajima_Formation_Tetori_Group_Japan_and_a_Reassessment_of_Triconodont_Phylogeny"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="9" data-entity-id="112887851" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/112887851/Theropod_teeth_from_the_Lower_Cretaceous_Ilek_Formation_of_Western_Siberia_Russia">Theropod teeth from the Lower Cretaceous Ilek Formation of Western Siberia, Russia</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="94834754" href="https://independent.academia.edu/PavelSkutchas">Pavel Skutchas</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Trudy Zoologičeskogo instituta, 2019</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Theropod teeth from the Lower Cretaceous Ilek Formation of Western Siberia, Russia","attachmentId":109986079,"attachmentType":"pdf","work_url":"https://www.academia.edu/112887851/Theropod_teeth_from_the_Lower_Cretaceous_Ilek_Formation_of_Western_Siberia_Russia","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/112887851/Theropod_teeth_from_the_Lower_Cretaceous_Ilek_Formation_of_Western_Siberia_Russia"><span class="ds2-5-text-link__content">View PDF</span><span 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You can download the paper by clicking the button above.</p></div></div></div></div><div class="ds-sidebar--container js-work-sidebar"><div class="ds-related-content--container"><h2 class="ds-related-content--heading">Related papers</h2><div class="ds-related-work--container js-related-work-sidebar-card" data-collection-position="0" data-entity-id="78416119" data-sort-order="default"><a class="ds-related-work--title js-related-work-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/78416119/A_new_carnosaur_Dinosauria_Theropoda_from_the_Lower_Cretaceous_of_Japan">A new carnosaur (Dinosauria: Theropoda) from the Lower Cretaceous of Japan</a><div class="ds-related-work--metadata"><a class="js-related-work-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="42314779" href="https://independent.academia.edu/YoichiAzuma">Yoichi Azuma</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Canadian Journal of Earth Sciences, 2000</p><div 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href="https://www.academia.edu/121850745/An_Early_Cretaceous_mammal_from_the_Duwajima_Formation_Tetori_Group_Japan_and_a_reassessment_of_triconodont_phylogeny">An Early Cretaceous mammal from the Duwajima Formation (Tetori Group), Japan, and a reassessment of triconodont phylogeny</a><div class="ds-related-work--metadata"><a class="js-related-work-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="34164059" href="https://independent.academia.edu/GuillermoRougier">Guillermo Rougier</a></div><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"An Early Cretaceous mammal from the Duwajima Formation (Tetori Group), Japan, and a reassessment of triconodont phylogeny","attachmentId":116634811,"attachmentType":"pdf","work_url":"https://www.academia.edu/121850745/An_Early_Cretaceous_mammal_from_the_Duwajima_Formation_Tetori_Group_Japan_and_a_reassessment_of_triconodont_phylogeny","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-related-work-grid-card-view-pdf" href="https://www.academia.edu/121850745/An_Early_Cretaceous_mammal_from_the_Duwajima_Formation_Tetori_Group_Japan_and_a_reassessment_of_triconodont_phylogeny"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-related-work-sidebar-card" data-collection-position="3" data-entity-id="62100063" data-sort-order="default"><a class="ds-related-work--title js-related-work-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/62100063/Cretaceous_theropods_from_India_A_review_of_specimens_described_by_Huene_and_Matley_1933_">Cretaceous theropods from India: A review of specimens described by Huene and Matley (1933)</a><div class="ds-related-work--metadata"><a class="js-related-work-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="7495089" href="https://independent.academia.edu/SaswatiBandyopadhyay">Saswati Bandyopadhyay</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Revista del Museo Argentino de Ciencias Naturales, 2004</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Cretaceous theropods from India: A review of specimens described by Huene and Matley (1933)","attachmentId":74954163,"attachmentType":"pdf","work_url":"https://www.academia.edu/62100063/Cretaceous_theropods_from_India_A_review_of_specimens_described_by_Huene_and_Matley_1933_","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-related-work-grid-card-view-pdf" href="https://www.academia.edu/62100063/Cretaceous_theropods_from_India_A_review_of_specimens_described_by_Huene_and_Matley_1933_"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-related-work-sidebar-card" data-collection-position="4" data-entity-id="24898189" data-sort-order="default"><a class="ds-related-work--title js-related-work-grid-card-title 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