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<div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Kanpo estekak</span> </div> </a> <ul id="toc-Kanpo_estekak-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Edukiak" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Eduki taularen ikusgarritasuna aldatu" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Eduki taularen ikusgarritasuna aldatu</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Nikolaas Tinbergen</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Joan beste hizkuntza batean idatzitako artikulu batera. 58 hizkuntzatan eskuragarri." > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-58" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">58 hizkuntza</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%86%D9%8A%D9%83%D9%88%D9%84%D8%A7%D8%B3_%D8%AA%D9%8A%D9%86%D8%A8%D8%B1%D8%AE%D9%86" title="نيكولاس تينبرخن – arabiera" lang="ar" hreflang="ar" data-title="نيكولاس تينبرخن" data-language-autonym="العربية" data-language-local-name="arabiera" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%86%D9%8A%D9%83%D9%88%D9%84%D8%A7%D8%B3_%D8%AA%D9%8A%D9%86%D8%A8%D8%B1%D8%AC%D9%86" title="نيكولاس تينبرجن – Egyptian Arabic" lang="arz" hreflang="arz" data-title="نيكولاس تينبرجن" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%86%DB%8C%DA%A9%D9%88%D9%84%D8%A7%D8%B3_%D8%AA%DB%8C%D9%86%D8%A8%D8%B1%D9%82%D9%86" title="نیکولاس تینبرقن – South Azerbaijani" lang="azb" hreflang="azb" data-title="نیکولاس تینبرقن" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%9D%D1%96%D0%BA%D0%B0%D0%BB%D0%B0%D1%81_%D0%A2%D1%8B%D0%BD%D0%B1%D0%B5%D1%80%D0%B3%D0%B5%D0%BD" title="Нікалас Тынберген – bielorrusiera" lang="be" hreflang="be" data-title="Нікалас Тынберген" data-language-autonym="Беларуская" data-language-local-name="bielorrusiera" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9D%D0%B8%D0%BA%D0%BE%D0%BB%D0%B0%D1%81_%D0%A2%D0%B8%D0%BD%D0%B1%D0%B5%D1%80%D0%B3%D0%B5%D0%BD" title="Николас Тинберген – bulgariera" lang="bg" hreflang="bg" data-title="Николас Тинберген" data-language-autonym="Български" data-language-local-name="bulgariera" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – katalana" lang="ca" hreflang="ca" data-title="Nikolaas Tinbergen" data-language-autonym="Català" data-language-local-name="katalana" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D9%86%DB%8C%DA%A9%DB%86%D9%84%D8%A7%D8%B3_%D8%AA%DB%8E%D9%86%D8%A8%DB%95%D8%B1%DA%AF%DB%8C%D9%86" title="نیکۆلاس تێنبەرگین – erdialdeko kurduera" lang="ckb" hreflang="ckb" data-title="نیکۆلاس تێنبەرگین" data-language-autonym="کوردی" data-language-local-name="erdialdeko kurduera" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – txekiera" lang="cs" hreflang="cs" data-title="Nikolaas Tinbergen" data-language-autonym="Čeština" data-language-local-name="txekiera" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/Niko_Tinbergen" title="Niko Tinbergen – galesa" lang="cy" hreflang="cy" data-title="Niko Tinbergen" data-language-autonym="Cymraeg" data-language-local-name="galesa" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – alemana" lang="de" hreflang="de" data-title="Nikolaas Tinbergen" data-language-autonym="Deutsch" data-language-local-name="alemana" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – ingelesa" lang="en" hreflang="en" data-title="Nikolaas Tinbergen" data-language-autonym="English" data-language-local-name="ingelesa" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – esperantoa" lang="eo" hreflang="eo" data-title="Nikolaas Tinbergen" data-language-autonym="Esperanto" data-language-local-name="esperantoa" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – gaztelania" lang="es" hreflang="es" data-title="Nikolaas Tinbergen" data-language-autonym="Español" data-language-local-name="gaztelania" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Niko_Tinbergen" title="Niko Tinbergen – estoniera" lang="et" hreflang="et" data-title="Niko Tinbergen" data-language-autonym="Eesti" data-language-local-name="estoniera" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%86%DB%8C%DA%A9%D9%88%D9%84%D8%A7%D8%B3_%D8%AA%DB%8C%D9%86%D8%A8%D8%B1%DA%AF%D9%86" title="نیکولاس تینبرگن – persiera" lang="fa" hreflang="fa" data-title="نیکولاس تینبرگن" data-language-autonym="فارسی" data-language-local-name="persiera" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – finlandiera" lang="fi" hreflang="fi" data-title="Nikolaas Tinbergen" data-language-autonym="Suomi" data-language-local-name="finlandiera" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – frantsesa" lang="fr" hreflang="fr" data-title="Nikolaas Tinbergen" data-language-autonym="Français" data-language-local-name="frantsesa" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – irlandera" lang="ga" hreflang="ga" data-title="Nikolaas Tinbergen" data-language-autonym="Gaeilge" data-language-local-name="irlandera" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Niko_Tinbergen" title="Niko Tinbergen – galiziera" lang="gl" hreflang="gl" data-title="Niko Tinbergen" data-language-autonym="Galego" data-language-local-name="galiziera" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A0%D7%99%D7%A7%D7%95%D7%9C%D7%90%D7%A1_%D7%98%D7%99%D7%A0%D7%91%D7%A8%D7%92%D7%9F" title="ניקולאס טינברגן – hebreera" lang="he" hreflang="he" data-title="ניקולאס טינברגן" data-language-autonym="עברית" data-language-local-name="hebreera" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – kroaziera" lang="hr" hreflang="hr" data-title="Nikolaas Tinbergen" data-language-autonym="Hrvatski" data-language-local-name="kroaziera" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – hungariera" lang="hu" hreflang="hu" data-title="Nikolaas Tinbergen" data-language-autonym="Magyar" data-language-local-name="hungariera" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – indonesiera" lang="id" hreflang="id" data-title="Nikolaas Tinbergen" data-language-autonym="Bahasa Indonesia" data-language-local-name="indonesiera" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – idoa" lang="io" hreflang="io" data-title="Nikolaas Tinbergen" data-language-autonym="Ido" data-language-local-name="idoa" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – italiera" lang="it" hreflang="it" data-title="Nikolaas Tinbergen" data-language-autonym="Italiano" data-language-local-name="italiera" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%8B%E3%82%B3%E3%83%BB%E3%83%86%E3%82%A3%E3%83%B3%E3%83%90%E3%83%BC%E3%82%B2%E3%83%B3" title="ニコ・ティンバーゲン – japoniera" lang="ja" hreflang="ja" data-title="ニコ・ティンバーゲン" data-language-autonym="日本語" data-language-local-name="japoniera" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%9D%D0%B8%D0%BA%D0%BE%D0%BB%D0%B0%D0%B0%D1%81_%D0%A2%D0%B8%D0%BD%D0%B1%D0%B5%D1%80%D0%B3%D0%B5%D0%BD" title="Николаас Тинберген – kazakhera" lang="kk" hreflang="kk" data-title="Николаас Тинберген" data-language-autonym="Қазақша" data-language-local-name="kazakhera" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%8B%88%EC%BD%9C%EB%9D%BC%EC%8A%A4_%ED%8B%B4%EB%B2%A0%EB%A5%B4%ED%97%8C" title="니콜라스 틴베르헌 – koreera" lang="ko" hreflang="ko" data-title="니콜라스 틴베르헌" data-language-autonym="한국어" data-language-local-name="koreera" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Nicolaus_Tinbergen" title="Nicolaus Tinbergen – latina" lang="la" hreflang="la" data-title="Nicolaus Tinbergen" data-language-autonym="Latina" data-language-local-name="latina" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Nikolass_Tinbergens" title="Nikolass Tinbergens – letoniera" lang="lv" hreflang="lv" data-title="Nikolass Tinbergens" data-language-autonym="Latviešu" data-language-local-name="letoniera" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%9D%D0%B8%D0%BA%D0%BE%D0%BB%D0%B0%D1%81_%D0%A2%D0%B8%D0%BD%D0%B1%D0%B5%D1%80%D0%B3%D0%B5%D0%BD" title="Николас Тинберген – mazedoniera" lang="mk" hreflang="mk" data-title="Николас Тинберген" data-language-autonym="Македонски" data-language-local-name="mazedoniera" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%A8%E0%B4%BF%E0%B4%95%E0%B5%8D%E0%B4%95%E0%B5%8B%E0%B4%B3%E0%B4%BE%E0%B4%B8%E0%B5%8D_%E0%B4%9F%E0%B4%BF%E0%B5%BB%E0%B4%AC%E0%B5%BC%E0%B4%9C%E0%B5%86%E0%B5%BB" title="നിക്കോളാസ് ടിൻബർജെൻ – malabarera" lang="ml" hreflang="ml" data-title="നിക്കോളാസ് ടിൻബർജെൻ" data-language-autonym="മലയാളം" data-language-local-name="malabarera" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – malaysiera" lang="ms" hreflang="ms" data-title="Nikolaas Tinbergen" data-language-autonym="Bahasa Melayu" data-language-local-name="malaysiera" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%86%DB%8C%DA%A9%D9%84%D8%A7%D8%B3_%D8%AA%DB%8C%D9%86%D8%A8%D8%B1%DA%AF%D9%86" title="نیکلاس تینبرگن – mazandarandera" lang="mzn" hreflang="mzn" data-title="نیکلاس تینبرگن" data-language-autonym="مازِرونی" data-language-local-name="mazandarandera" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Niko_Tinbergen" title="Niko Tinbergen – nederlandera" lang="nl" hreflang="nl" data-title="Niko Tinbergen" data-language-autonym="Nederlands" data-language-local-name="nederlandera" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – nynorsk (norvegiera)" lang="nn" hreflang="nn" data-title="Nikolaas Tinbergen" data-language-autonym="Norsk nynorsk" data-language-local-name="nynorsk (norvegiera)" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – bokmål (norvegiera)" lang="nb" hreflang="nb" data-title="Nikolaas Tinbergen" data-language-autonym="Norsk bokmål" data-language-local-name="bokmål (norvegiera)" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – okzitaniera" lang="oc" hreflang="oc" data-title="Nikolaas Tinbergen" data-language-autonym="Occitan" data-language-local-name="okzitaniera" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – poloniera" lang="pl" hreflang="pl" data-title="Nikolaas Tinbergen" data-language-autonym="Polski" data-language-local-name="poloniera" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%86%DA%A9%D9%88%D9%84%D8%A7%D8%B3_%D9%B9%D9%86%D8%A8%D8%B1%DA%AF%D9%86" title="نکولاس ٹنبرگن – Western Punjabi" lang="pnb" hreflang="pnb" data-title="نکولاس ٹنبرگن" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – portugesa" lang="pt" hreflang="pt" data-title="Nikolaas Tinbergen" data-language-autonym="Português" data-language-local-name="portugesa" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – errumaniera" lang="ro" hreflang="ro" data-title="Nikolaas Tinbergen" data-language-autonym="Română" data-language-local-name="errumaniera" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A2%D0%B8%D0%BD%D0%B1%D0%B5%D1%80%D0%B3%D0%B5%D0%BD,_%D0%9D%D0%B8%D0%BA%D0%BE%D0%BB%D0%B0%D1%81" title="Тинберген, Николас – errusiera" lang="ru" hreflang="ru" data-title="Тинберген, Николас" data-language-autonym="Русский" data-language-local-name="errusiera" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Niko_Tinbergen" title="Niko Tinbergen – Simple English" lang="en-simple" hreflang="en-simple" data-title="Niko Tinbergen" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – eslovakiera" lang="sk" hreflang="sk" data-title="Nikolaas Tinbergen" data-language-autonym="Slovenčina" data-language-local-name="eslovakiera" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – esloveniera" lang="sl" hreflang="sl" data-title="Nikolaas Tinbergen" data-language-autonym="Slovenščina" data-language-local-name="esloveniera" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%9D%D0%B8%D0%BA%D0%BE%D0%BB%D0%B0%D1%81_%D0%A2%D0%B8%D0%BD%D0%B1%D0%B5%D1%80%D0%B3%D0%B5%D0%BD" title="Николас Тинберген – serbiera" lang="sr" hreflang="sr" data-title="Николас Тинберген" data-language-autonym="Српски / srpski" data-language-local-name="serbiera" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – suediera" lang="sv" hreflang="sv" data-title="Nikolaas Tinbergen" data-language-autonym="Svenska" data-language-local-name="suediera" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – swahilia" lang="sw" hreflang="sw" data-title="Nikolaas Tinbergen" data-language-autonym="Kiswahili" data-language-local-name="swahilia" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%A8%E0%AE%BF%E0%AE%95%E0%AF%8D%E0%AE%95%E0%AF%8B_%E0%AE%9F%E0%AE%BF%E0%AE%A9%E0%AF%8D%E0%AE%AA%E0%AF%86%E0%AE%B0%E0%AF%8D%E0%AE%9C%E0%AF%86%E0%AE%A9%E0%AF%8D" title="நிக்கோ டின்பெர்ஜென் – tamilera" lang="ta" hreflang="ta" data-title="நிக்கோ டின்பெர்ஜென்" data-language-autonym="தமிழ்" data-language-local-name="tamilera" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – turkiera" lang="tr" hreflang="tr" data-title="Nikolaas Tinbergen" data-language-autonym="Türkçe" data-language-local-name="turkiera" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9D%D1%96%D0%BA%D0%BE%D0%BB%D0%B0%D1%81_%D0%A2%D1%96%D0%BD%D0%B1%D0%B5%D1%80%D0%B3%D0%B5%D0%BD" title="Ніколас Тінберген – ukrainera" lang="uk" hreflang="uk" data-title="Ніколас Тінберген" data-language-autonym="Українська" data-language-local-name="ukrainera" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D9%86%DA%A9%D9%88%D9%84%D8%A7%D8%B3_%D9%B9%D9%86%D8%A8%D8%B1%D8%AC%D9%86" title="نکولاس ٹنبرجن – urdua" lang="ur" hreflang="ur" data-title="نکولاس ٹنبرجن" data-language-autonym="اردو" data-language-local-name="urdua" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – uzbekera" lang="uz" hreflang="uz" data-title="Nikolaas Tinbergen" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="uzbekera" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – jorubera" lang="yo" hreflang="yo" data-title="Nikolaas Tinbergen" data-language-autonym="Yorùbá" data-language-local-name="jorubera" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%B0%BC%E5%8F%A4%E6%8B%89%E6%96%AF%C2%B7%E5%BB%B7%E8%B4%9D%E4%BA%A8" title="尼古拉斯·廷贝亨 – txinera" lang="zh" hreflang="zh" data-title="尼古拉斯·廷贝亨" data-language-autonym="中文" data-language-local-name="txinera" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen – Minnan" lang="nan" hreflang="nan" data-title="Nikolaas Tinbergen" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E5%B0%BC%E5%8F%A4%E6%8B%89%E6%96%AF%C2%B7%E5%BB%B7%E8%B2%9D%E6%A0%B9" title="尼古拉斯·廷貝根 – kantonera" lang="yue" hreflang="yue" data-title="尼古拉斯·廷貝根" data-language-autonym="粵語" data-language-local-name="kantonera" class="interlanguage-link-target"><span>粵語</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q153890#sitelinks-wikipedia" title="Aldatu hizkuntzen arteko loturak" class="wbc-editpage">Aldatu loturak</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Izen-tarteak"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul 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href="/wiki/Berezi:ZerkLotzenDuHona/Nikolaas_Tinbergen" title="Orri honetaranzko esteka duten wiki orri guztien zerrenda [j]" accesskey="j"><span>Honanzko esteka duten orriak</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Berezi:RecentChangesLinked/Nikolaas_Tinbergen" rel="nofollow" title="Orri honetatik esteka duten orrietako azken aldaketak [k]" accesskey="k"><span>Lotutako orrietako aldaketak</span></a></li><li id="t-upload" class="mw-list-item"><a href="//commons.wikimedia.org/wiki/Special:UploadWizard?uselang=eu" title="Irudiak edo media fitxategiak igo [u]" accesskey="u"><span>Fitxategia igo</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Berezi:OrrialdeBereziak" title="Orri berezi guztien zerrenda [q]" accesskey="q"><span>Orri bereziak</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Nikolaas_Tinbergen&oldid=9793281" title="Orriaren bertsio honetaranzko esteka 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id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="eu" dir="ltr"><table class="infobox" style="width:22em"><tbody><tr><th colspan="2" class="hd" style="text-align:center;font-size:125%;font-weight:bold;background-color: lightsteelblue">Nikolaas Tinbergen</th></tr><tr><td colspan="2" class="infobox-full-data infobox-data" style="text-align:center"><span typeof="mw:File"><a href="/wiki/Fitxategi:Nikolaas_Tinbergen_1978.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/84/Nikolaas_Tinbergen_1978.jpg/260px-Nikolaas_Tinbergen_1978.jpg" decoding="async" width="260" height="309" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/84/Nikolaas_Tinbergen_1978.jpg/390px-Nikolaas_Tinbergen_1978.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/8/84/Nikolaas_Tinbergen_1978.jpg 2x" data-file-width="470" data-file-height="559" /></a></span><br /><div><span typeof="mw:File"><a href="https://commons.wikimedia.org/wiki/Category:Nikolaas_Tinbergen" title="c:Category:Nikolaas Tinbergen"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/10px-Commons-logo.svg.png" decoding="async" width="10" height="13" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/15px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/20px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span> <a href="https://commons.wikimedia.org/wiki/Category:Nikolaas_Tinbergen" class="extiw" title="c:Category:Nikolaas Tinbergen">Irudi gehiago</a></div></td></tr><tr><th colspan="2" class="infobox-header" style="text-align:center;background-color: lightsteelblue">Bizitza</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Jaiotza</th><td class="infobox-data"><a href="/wiki/Haga" title="Haga">Haga</a>, <a href="/wiki/1907" title="1907">1907ko</a> <a href="/wiki/Apirilaren_15" title="Apirilaren 15">apirilaren 15a</a></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Herrialdea</th><td class="infobox-data"><span class="flagicon"><span class="mw-image-border" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/20/Flag_of_the_Netherlands.svg/22px-Flag_of_the_Netherlands.svg.png" decoding="async" width="22" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/20/Flag_of_the_Netherlands.svg/33px-Flag_of_the_Netherlands.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/20/Flag_of_the_Netherlands.svg/44px-Flag_of_the_Netherlands.svg.png 2x" data-file-width="900" data-file-height="600" /></span></span> </span><a href="/wiki/Herbehereetako_Erresuma" title="Herbehereetako Erresuma">Herbehereetako Erresuma</a> <br /><span class="flagicon"><span class="mw-image-border" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/83/Flag_of_the_United_Kingdom_%283-5%29.svg/22px-Flag_of_the_United_Kingdom_%283-5%29.svg.png" decoding="async" width="22" height="13" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/83/Flag_of_the_United_Kingdom_%283-5%29.svg/33px-Flag_of_the_United_Kingdom_%283-5%29.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/83/Flag_of_the_United_Kingdom_%283-5%29.svg/44px-Flag_of_the_United_Kingdom_%283-5%29.svg.png 2x" data-file-width="1000" data-file-height="600" /></span></span> </span><a href="/wiki/Erresuma_Batua" title="Erresuma Batua">Erresuma Batua</a> <small> (1955 -</small></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Heriotza</th><td class="infobox-data"><a href="/wiki/Oxford" title="Oxford">Oxford</a>, <a href="/wiki/1988" title="1988">1988ko</a> <a href="/wiki/Abenduaren_21" title="Abenduaren 21">abenduaren 21a</a> (81 urte)</td></tr><tr><th colspan="2" class="infobox-header" style="text-align:center;background-color: lightsteelblue">Familia</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Aita</th><td class="infobox-data"><a href="/w/index.php?title=D._C._Tinbergen&action=edit&redlink=1" class="new" title="D. C. Tinbergen (sortu gabe)">D. C. Tinbergen</a></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Haurrideak</th><td class="infobox-data"><div class="NavFrame collapsed" style="border: none; padding: 0;"> <div class="NavHead" style="font-size: 105%; background: transparent; text-align: left;">ikusi</div> <ul class="NavContent" style="list-style: none none; margin-left: 0; text-align: left; font-size: 105%; margin-top: 0; margin-bottom: 0; line-height: inherit;"><li style="line-height: inherit; margin: 0"><a href="/wiki/Jan_Tinbergen" title="Jan Tinbergen">Jan Tinbergen</a><br /><a href="/w/index.php?title=Luuk_Tinbergen&action=edit&redlink=1" class="new" title="Luuk Tinbergen (sortu gabe)">Luuk Tinbergen</a> <sup>(mul)</sup> <span class="mw-valign-baseline skin-invert" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q331505?uselang=eu"><img alt="Itzuli" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/10px-Noun_Project_label_icon_1116097_cc_mirror.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/15px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/20px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 2x" data-file-width="158" data-file-height="161" /></a></span></li></ul> </div></td></tr><tr><th colspan="2" class="infobox-header" style="text-align:center;background-color: lightsteelblue">Hezkuntza</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Heziketa</th><td class="infobox-data"><a href="/wiki/Leidengo_Unibertsitatea" title="Leidengo Unibertsitatea">Leidengo Unibertsitatea</a> <br /><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q57046996" class="extiw" title="d:Special:EntityPage/Q57046996">Stedelijk Gymnasium Haarlem</a> <sup>(en)</sup> <span class="mw-valign-baseline skin-invert" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q57046996?uselang=eu"><img alt="Itzuli" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/10px-Noun_Project_label_icon_1116097_cc_mirror.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/15px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/20px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 2x" data-file-width="158" data-file-height="161" /></a></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Doktorego ikaslea(k)</th><td class="infobox-data"><a href="/wiki/Richard_Dawkins" title="Richard Dawkins">Richard Dawkins</a><br /><a href="/w/index.php?title=Colin_Gordon_Beer&action=edit&redlink=1" class="new" title="Colin Gordon Beer (sortu gabe)">Colin Gordon Beer</a> <sup>(en)</sup> <span class="mw-valign-baseline skin-invert" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q113949916?uselang=eu"><img alt="Itzuli" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/10px-Noun_Project_label_icon_1116097_cc_mirror.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/15px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/20px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 2x" data-file-width="158" data-file-height="161" /></a></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Hizkuntzak</th><td class="infobox-data"><a href="/wiki/Ingeles" title="Ingeles">ingelesa</a><br /><a href="/wiki/Nederlandera" title="Nederlandera">nederlandera</a></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Ikaslea(k)</th><td class="infobox-data"><div class="NavFrame collapsed" style="border: none; padding: 0;"> <div class="NavHead" style="font-size: 105%; background: transparent; text-align: left;">ikusi</div> <ul class="NavContent" style="list-style: none none; margin-left: 0; text-align: left; font-size: 105%; margin-top: 0; margin-bottom: 0; line-height: inherit;"><li style="line-height: inherit; margin: 0"><i><a href="/wiki/Richard_Dawkins" title="Richard Dawkins">Richard Dawkins</a></i></li></ul> </div></td></tr><tr><th colspan="2" class="infobox-header" style="text-align:center;background-color: lightsteelblue">Jarduerak</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Jarduerak</th><td class="infobox-data"><a href="/wiki/Biologo" class="mw-redirect" title="Biologo">biologoa</a>, <a href="/wiki/Zoologo" class="mw-redirect" title="Zoologo">zoologoa</a>, <a href="/wiki/Ornitologo" class="mw-redirect" title="Ornitologo">ornitologoa</a>, <a href="/w/index.php?title=Etologo&action=edit&redlink=1" class="new" title="Etologo (sortu gabe)">etologoa</a>, <a href="/wiki/Unibertsitateko_irakasle" class="mw-redirect" title="Unibertsitateko irakasle">unibertsitateko irakaslea</a> eta <a href="/wiki/Argazkilari" class="mw-redirect" title="Argazkilari">argazkilaria</a></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Enplegatzailea(k)</th><td class="infobox-data"><a href="/wiki/Oxfordeko_Unibertsitatea" title="Oxfordeko Unibertsitatea">Oxfordeko Unibertsitatea</a> <br /><a href="/wiki/Leidengo_Unibertsitatea" title="Leidengo Unibertsitatea">Leidengo Unibertsitatea</a> <small> (<a href="/wiki/1939" title="1939">1939ko</a> <a href="/wiki/Apirilaren_18" title="Apirilaren 18">apirilaren 18a</a> -</small> <br /><a href="/wiki/Leidengo_Unibertsitatea" title="Leidengo Unibertsitatea">Leidengo Unibertsitatea</a> <small> (<a href="/wiki/1947" title="1947">1947ko</a> <a href="/wiki/Urtarrilaren_23" title="Urtarrilaren 23">urtarrilaren 23a</a> -</small> <small> <a href="/wiki/1949" title="1949">1949ko</a> <a href="/wiki/Ekainaren_16" title="Ekainaren 16">ekainaren 16a</a></small>)</td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Jasotako sariak</th><td class="infobox-data"><div class="NavFrame collapsed" style="border: none; padding: 0;"> <div class="NavHead" style="font-size: 105%; background: transparent; text-align: left;">ikusi</div> <ul class="NavContent" style="list-style: none none; margin-left: 0; text-align: left; font-size: 105%; margin-top: 0; margin-bottom: 0; line-height: inherit;"><li style="line-height: inherit; margin: 0"><span typeof="mw:File"><a href="/wiki/Fitxategi:Nobel_prize_medal.svg" class="mw-file-description" title="Medikuntzako Nobel Saria"><img alt="Medikuntzako Nobel Saria" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/20px-Nobel_prize_medal.svg.png" decoding="async" width="20" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/30px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/40px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span> <a href="/wiki/Medikuntzako_Nobel_Saria" class="mw-redirect" title="Medikuntzako Nobel Saria">Medikuntzako Nobel Saria</a> <small> (1973)</small> <br /> <a href="/wiki/Royal_Societyko_kidea" class="mw-redirect" title="Royal Societyko kidea">Royal Societyko kidea</a> <br /> <a href="/w/index.php?title=APA_Award_for_Distinguished_Scientific_Contributions_to_Psychology&action=edit&redlink=1" class="new" title="APA Award for Distinguished Scientific Contributions to Psychology (sortu gabe)">APA Award for Distinguished Scientific Contributions to Psychology</a> <small> (1987)</small> <br /> <a href="/w/index.php?title=Godman-Salvin_Medal&action=edit&redlink=1" class="new" title="Godman-Salvin Medal (sortu gabe)">Godman-Salvin Medal</a> <small> (1969)</small> <br /> <a href="/w/index.php?title=Croonian_Medal_and_Lecture&action=edit&redlink=1" class="new" title="Croonian Medal and Lecture (sortu gabe)">Croonian Medal and Lecture</a> <small> (1972)</small> <br /> <a href="/w/index.php?title=Honorary_doctor_of_the_University_of_Leicester&action=edit&redlink=1" class="new" title="Honorary doctor of the University of Leicester (sortu gabe)">honorary doctor of the University of Leicester</a> <br /> <a href="/w/index.php?title=Elliott_Coues_Award&action=edit&redlink=1" class="new" title="Elliott Coues Award (sortu gabe)">Elliott Coues Award</a> <small> (1972)</small> </li></ul> </div></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Kidetza</th><td class="infobox-data"><a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a><br /><a href="/wiki/Alemaniako_Natur_Zientzien_Leopoldina_Akademia" title="Alemaniako Natur Zientzien Leopoldina Akademia">Alemaniako Natur Zientzien Leopoldina Akademia</a><br /><a href="/w/index.php?title=Arteen_eta_Zientzien_Herbehereetako_Errege_Akademia&action=edit&redlink=1" class="new" title="Arteen eta Zientzien Herbehereetako Errege Akademia (sortu gabe)">Arteen eta Zientzien Herbehereetako Errege Akademia</a><br /><a href="/wiki/Arteen_eta_Zientzien_Ameriketako_Estatu_Batuetako_Akademia" title="Arteen eta Zientzien Ameriketako Estatu Batuetako Akademia">Arteen eta Zientzien Ameriketako Estatu Batuetako Akademia</a><br /><a href="/wiki/Ameriketako_Estatu_Batuetako_Zientzien_Akademia_Nazionala" title="Ameriketako Estatu Batuetako Zientzien Akademia Nazionala">Ameriketako Estatu Batuetako Zientzien Akademia Nazionala</a></td></tr><tr><td colspan="2" class="infobox-full-data infobox-data" style="text-align:center"><hr /></td></tr><tr><td colspan="2" class="infobox-full-data infobox-data" style="text-align:center"><span typeof="mw:File"><a href="https://www.findagrave.com/memorial/86505019" title="Find a Grave: 86505019" rel="nofollow"><img alt="Find a Grave: 86505019" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0e/Creative-Tail-Halloween-grave.svg/22px-Creative-Tail-Halloween-grave.svg.png" decoding="async" width="22" height="22" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0e/Creative-Tail-Halloween-grave.svg/33px-Creative-Tail-Halloween-grave.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0e/Creative-Tail-Halloween-grave.svg/44px-Creative-Tail-Halloween-grave.svg.png 2x" data-file-width="140" data-file-height="140" /></a></span> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q153890?uselang=eu#identifiers" title="Edit the value on Wikidata"><img alt="Edit the value on Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr></tbody></table> <p><b>Nikolaas "Niko" Tinbergen</b> (<a href="/wiki/Haga" title="Haga">Haga</a>, <a href="/wiki/1907" title="1907">1907ko</a> <a href="/wiki/Apirilaren_15" title="Apirilaren 15">apirilaren 15a</a> - <a href="/wiki/Oxford" title="Oxford">Oxford</a>, <a href="/wiki/1988" title="1988">1988ko</a> <a href="/wiki/Abenduaren_21" title="Abenduaren 21">abenduaren 21a</a>) herbeheretar <a href="/wiki/Etologia" title="Etologia">etologoa</a> izan zen. Zoologiako ikasketak egin zituen, <a href="/wiki/Leidengo_Unibertsitatea" title="Leidengo Unibertsitatea">Leidengo Unibertsitatean</a>. Gero, Zoologia Esperimentaleko irakasle izan zen unibertsitate hartan bertan. 1947an Oxforden izan zen, hango Unibertsitatean lan egin zuen eta hango irakasle izan zen; Animalien Portaeraren Ikerketa saileko arduradun izan zen. Animalien jokabideari buruzko lanak egin zituen han, aske zeuden eta laborategian zeuden animalien portaera desberdintasunak aztertzeko. Artikoko azeriak, itsas hegaztiak, itsas otsoak, barraskiloak eta liztorrak aztertu zituen, besteak beste. Animalien portaeraren ikaskuntza eta gizakien sena, autismoa eta oldarkortasuna aztertu zituen, besteak beste. </p><p>1973an <a href="/wiki/Medikuntzako_Nobel_Saria" class="mw-redirect" title="Medikuntzako Nobel Saria">Fisiologia eta Medikuntzako Nobel saria</a> jaso zuen, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> eta <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frischekin</a> batera. Lanik aipagarrienak: <i>The Study of Instinct</i> (1951, <i>Senaren azterketa</i>) eta <i>The Animal in Its World. Field Studies</i> (1932-1972). </p> <div class="mw-heading mw-heading2"><h2 id="Erreferentziak">Erreferentziak</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nikolaas_Tinbergen&veaction=edit&section=1" title="Aldatu atal hau: «Erreferentziak»" class="mw-editsection-visualeditor"><span>aldatu</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Nikolaas_Tinbergen&action=edit&section=1" title="Edit section's source code: Erreferentziak"><span>aldatu iturburu kodea</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="boilerplate" style="text-align:left; font-size: 90%"> <ul><li>Artikulu honen edukiaren zati bat <i>Lur hiztegi entziklopediko</i>tik edo <i>Lur entziklopedia tematiko</i>tik txertatu zen 2011/12/30 egunean. Egile-eskubideen jabeak, <a href="/wiki/Eusko_Jaurlaritza" title="Eusko Jaurlaritza">Eusko Jaurlaritzak</a>, hiztegi horiek <a rel="nofollow" class="external text" href="http://creativecommons.org/licenses/by/3.0/es/deed.eu">CC-BY 3.0 lizentziarekin</a> argitaratu ditu, <a rel="nofollow" class="external text" href="http://opendata.euskadi.eus/katalogoa/-/lur-entziklopedia-tematikoa/">Open Data Euskadi webgunean</a>.</li></ul></div> <div class="reflist" style="list-style-type: decimal;"> </div> <div class="mw-heading mw-heading2"><h2 id="Kanpo_estekak">Kanpo estekak</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nikolaas_Tinbergen&veaction=edit&section=2" title="Aldatu atal hau: «Kanpo estekak»" class="mw-editsection-visualeditor"><span>aldatu</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Nikolaas_Tinbergen&action=edit&section=2" title="Edit section's source code: Kanpo estekak"><span>aldatu iturburu kodea</span></a><span class="mw-editsection-bracket">]</span></span></div> <div style="clear:both;"></div><style data-mw-deduplicate="TemplateStyles:r7786466">.mw-parser-output .mw-authority-control{margin-top:1.5em}.mw-parser-output .mw-authority-control .navbox hr:last-child{display:none}.mw-parser-output .mw-authority-control .navbox+.mw-mf-linked-projects{display:none}.mw-parser-output .mw-authority-control,.mw-parser-output .mw-mf-linked-projects{border:1px solid #a2a9b1;font-size:88%}.mw-parser-output .mw-authority-control .mw-mf-linked-projects ul li{margin-bottom:0}</style><div class="mw-authority-control"><div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r9236167">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output 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.navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}</style></div><div role="navigation" class="navbox" aria-labelledby="Autoritate_kontrola" style="width: inherit;padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th id="Autoritate_kontrola" scope="row" class="navbox-group" style="width:1%;width: 12%; text-align:center;"><a href="/wiki/Laguntza:Autoritate_kontrola" title="Laguntza:Autoritate kontrola">Autoritate kontrola</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Wikimedia proiektuak</b></li> <li><span style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikidata" title="Wikidata"><img alt="Wd" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/20px-Wikidata-logo.svg.png" decoding="async" width="20" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/30px-Wikidata-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/40px-Wikidata-logo.svg.png 2x" data-file-width="1050" data-file-height="590" /></a></span> Datuak:</span> <span class="uid"><a href="https://www.wikidata.org/wiki/Q153890" class="extiw" title="wikidata:Q153890">Q153890</a></span></li> <li><span style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikimedia_Commons" title="Commonscat"><img alt="Commonscat" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/15px-Commons-logo.svg.png" decoding="async" width="15" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/23px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span> Multimedia:</span> <span class="uid"><span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Category:Nikolaas_Tinbergen">Nikolaas Tinbergen</a></span> / <span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Special:MediaSearch?type=image&search=%22Q153890%22">Q153890</a></span></span></li></ul> <hr /> <ul><li><b>Identifikadoreak</b></li> <li><a rel="nofollow" class="external text" href="//www.worldcat.org/identities/viaf-36979006">WorldCat</a></li> <li><span style="white-space:nowrap;"><a href="/wiki/Virtual_International_Authority_File" class="mw-redirect" title="Virtual International Authority File">VIAF</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/36979006/">36979006</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/International_Standard_Name_Identifier" title="International Standard Name Identifier">ISNI</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/0000000110249174">0000000110249174</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Biblioteca_Nacional_de_Espa%C3%B1a" class="mw-redirect" title="Biblioteca Nacional de España">BNE</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://datos.bne.es/resource/XX1066689">XX1066689</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Biblioth%C3%A8que_nationale_de_France" class="mw-redirect" title="Bibliothèque nationale de France">BNF</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb12280443t">12280443t</a> <a rel="nofollow" class="external text" href="http://data.bnf.fr/ark:/12148/cb12280443t">(data)</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Kataluniako_Liburutegia" title="Kataluniako Liburutegia">CANTIC</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://cantic.bnc.cat/registres/CUCId/a10337076">a10337076</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Authority_File" class="mw-redirect" title="Integrated Authority File">GND</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/11880233X">11880233X</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Library_of_Congress_Control_Number" title="Library of Congress Control Number">LCCN</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n79125115">n79125115</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Diet_Library" class="mw-redirect" title="National Diet Library">NDL</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00458808">00458808</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Library_of_the_Czech_Republic" class="mw-redirect" title="National Library of the Czech Republic">NKC</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=nlk20000083425">nlk20000083425</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Library_of_Australia" class="mw-redirect" title="National Library of Australia">NLA</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://librariesaustralia.nla.gov.au/search/display?dbid=auth&id=36181334">36181334</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/CiNii" title="CiNii">CiNii</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://ci.nii.ac.jp/author/DA0076758X">DA0076758X</a></span></li> <li><span style="white-space:nowrap;"><a href="https://www.wikidata.org/wiki/Q29861311" class="extiw" title="d:Q29861311">SNAC</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://snaccooperative.org/ark:/99166/w61r7hch">w61r7hch</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Syst%C3%A8me_universitaire_de_documentation" title="Système universitaire de documentation">SUDOC</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/031625371">031625371</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/BIBSYS" title="BIBSYS">BIBSYS</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://authority.bibsys.no/authority/rest/authorities/html/90097590">90097590</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Bartzelonako_Unibertsitatea" title="Bartzelonako Unibertsitatea">UB</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://web.archive.org/web/*/https://crai.ub.edu/sites/default/files/autoritats/permanent/a1246936">a1246936</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Istituto_Centrale_per_il_Catalogo_Unico" title="Istituto Centrale per il Catalogo Unico">ICCU</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://opac.sbn.it/risultati-autori/-/opac-autori/detail/CFIV032166?core=autoriall">CFIV032166</a></span></li> <li><span style="white-space:nowrap;"><a href="/w/index.php?title=Herbehereetako_Arte_eta_Zientzien_Errege_Akademia&action=edit&redlink=1" class="new" title="Herbehereetako Arte eta Zientzien Errege Akademia (sortu gabe)">KNAW</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="http://www.dwc.knaw.nl/biografie/pmknaw/?pagetype=authorDetail&aId=PE00003350">PE00003350</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Deutsche_Akademie_der_Naturforscher_Leopoldina" class="mw-redirect" title="Deutsche Akademie der Naturforscher Leopoldina">Leopoldina</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://web.archive.org/web/https://www.leopoldina.org/mitglieder/mitgliederverzeichnis/mitglieder/member/Member/show/6931/">6931</a></span></li> <li><span style="white-space:nowrap;"><a href="/w/index.php?title=Biografisch_Portaal&action=edit&redlink=1" class="new" title="Biografisch Portaal (sortu gabe)">BPN</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="http://www.biografischportaal.nl/persoon/34530648">34530648</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Library" title="Open Library">Open Library</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://openlibrary.org/works/OL4869829A?mode=all">OL4869829A</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Rijksbureau_voor_Kunsthistorische_Documentatie" title="Rijksbureau voor Kunsthistorische Documentatie">RKD</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://rkd.nl/artists/389123">389123</a></span></li> <li><b>Hiztegiak eta entziklopediak</b></li> <li><span style="white-space:nowrap;"><a href="/wiki/Lur_Hiztegi_Entziklopedikoa" title="Lur Hiztegi Entziklopedikoa">Lur</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://www.euskadi.eus/web01-a2lurhiz/eu/contenidos/termino/_c05631/eu_t_1017/t1017.html">05631/eu_t_1017/t1017</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclop%C3%A6dia_Britannica" title="Encyclopædia Britannica">Britannica</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://www.britannica.com/biography/Nikolaas-Tinbergen">url</a></span></li></ul> </div></td></tr></tbody></table></div><div class="mw-mf-linked-projects hlist"> <ul><li><span style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikidata" title="Wikidata"><img alt="Wd" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/20px-Wikidata-logo.svg.png" decoding="async" width="20" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/30px-Wikidata-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/40px-Wikidata-logo.svg.png 2x" data-file-width="1050" data-file-height="590" /></a></span> Datuak:</span> <span class="uid"><a href="https://www.wikidata.org/wiki/Q153890" class="extiw" title="wikidata:Q153890">Q153890</a></span></li> <li><span style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikimedia_Commons" title="Commonscat"><img alt="Commonscat" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/15px-Commons-logo.svg.png" decoding="async" width="15" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/23px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span> Multimedia:</span> <span class="uid"><span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Category:Nikolaas_Tinbergen">Nikolaas Tinbergen</a></span> / <span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Special:MediaSearch?type=image&search=%22Q153890%22">Q153890</a></span></span></li></ul> </div></div> <p><br /> </p> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r9236167"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r9236165"></div><div role="navigation" class="navbox" aria-labelledby="Zerrenda:Medikuntzako_Nobel_Saria" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r9236167"><style data-mw-deduplicate="TemplateStyles:r9235565">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-ikusi"><a href="/w/index.php?title=Txantiloi:Nobel_Prize_in_Physiology_or_Medicine&action=edit&redlink=1" class="new" title="Txantiloi:Nobel Prize in Physiology or Medicine (sortu gabe)"><abbr title="Txantiloi hau ikusi" style=";;background:none transparent;border:none;box-shadow:none;padding:0;">i</abbr></a></li><li class="nv-eztabaida"><a href="/w/index.php?title=Txantiloi_eztabaida:Nobel_Prize_in_Physiology_or_Medicine&action=edit&redlink=1" class="new" title="Txantiloi eztabaida:Nobel Prize in Physiology or Medicine (sortu gabe)"><abbr title="Txantiloi hau eztabaidatu" style=";;background:none transparent;border:none;box-shadow:none;padding:0;">e</abbr></a></li><li class="nv-aldatu"><a class="external text" href="https://eu.wikipedia.org/w/index.php?title=Txantiloi:Nobel_Prize_in_Physiology_or_Medicine&action=edit"><abbr title="Txantiloi hau aldatu" style=";;background:none transparent;border:none;box-shadow:none;padding:0;">a</abbr></a></li></ul></div><div id="Zerrenda:Medikuntzako_Nobel_Saria" style="font-size:114%;margin:0 4em"><a href="/wiki/Zerrenda:Medikuntzako_Nobel_Saria" title="Zerrenda:Medikuntzako Nobel Saria">Zerrenda:Medikuntzako Nobel Saria</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Alphonse_Laveran" title="Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Grant_Banting" title="Frederick Grant Banting">Frederick Banting</a> / <a href="/wiki/John_James_Rickard_Macleod" title="John James Rickard Macleod">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1925</span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Doisy" title="Edward Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Tadeus Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Abraham_Waksman" title="Selman Abraham Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Adolf_Krebs" title="Hans Adolf Krebs">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_For%C3%9Fmann" title="Werner Forßmann">Werner Forßmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">George Beadle</a> / <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Brian_Medawar" title="Peter Brian Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Dewey_Watson" title="James Dewey Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Carew_Eccles" title="John Carew Eccles">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Fielding_Huxley" title="Andrew Fielding Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Lwoff" title="André Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_William_Holley" title="Robert William Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbruck" title="Max Delbruck">Max Delbruck</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a class="mw-selflink selflink">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Blumberg" title="Baruch Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Yalow" title="Rosalyn Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_Smith" title="Hamilton Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Newbold_Hounsfield" title="Godfrey Newbold Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_Samuelsson" title="Bengt Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_Leonard_Goldstein" title="Joseph Leonard Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen" title="Stanley Cohen">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_W._Black" title="James W. Black">James W. Black</a> / <a href="/wiki/Gertrude_Belle_Elion" title="Gertrude Belle Elion">Gertrude B. Elion</a> / <a href="/wiki/George_Herbert_Hitchings" title="George Herbert Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_Henri_Fischer" title="Edmond Henri Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_Krebs" title="Edwin Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Sharp" title="Phillip Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_Wieschaus" title="Eric Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_Zinkernagel" title="Rolf Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_Prusiner" title="Stanley Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_Hartwell" title="Leland Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Timothy_Hunt" title="Timothy Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_E._Sulston" title="John E. Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9_Sinoussi" title="Françoise Barré Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards" title="Robert Edwards">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>hil ostean</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell" title="William C. Campbell">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuko_Honjo" title="Tasuko Honjo">Tasuko Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap">2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton" title="Michael Houghton">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap">2021: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap">2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap">2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div>    </div><script>(RLQ=window.RLQ||[]).push(function(){mw.log.warn("Gadget \"ErrefAurrebista\" was not loaded. Please migrate it to use ResourceLoader. 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