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class="firstHeading mw-first-heading"><span class="mw-page-title-main">George Beadle</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 55 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-55" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">55 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/George_Beadle" title="George Beadle – Afrikaans" lang="af" hreflang="af" data-title="George Beadle" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D8%A8%D9%8A%D8%AF%D9%84" title="جورج بيدل – Arabic" lang="ar" hreflang="ar" data-title="جورج بيدل" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Corc_Bidl" title="Corc Bidl – Azerbaijani" lang="az" hreflang="az" data-title="Corc Bidl" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D8%A8%DB%8C%D8%AF%D9%84" title="جورج بیدل – South Azerbaijani" lang="azb" hreflang="azb" data-title="جورج بیدل" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%9C%E0%A6%B0%E0%A7%8D%E0%A6%9C_%E0%A6%93%E0%A6%AF%E0%A6%BC%E0%A7%87%E0%A6%B2%E0%A6%B8_%E0%A6%AC%E0%A6%BF%E0%A6%A1%E0%A7%87%E0%A6%B2" title="জর্জ ওয়েলস বিডেল – Bangla" lang="bn" hreflang="bn" data-title="জর্জ ওয়েলস বিডেল" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6_%D0%A3%D1%8D%D0%BB%D1%81_%D0%91%D1%96%D0%B4%D0%BB" title="Джордж Уэлс Бідл – Belarusian" lang="be" hreflang="be" data-title="Джордж Уэлс Бідл" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6_%D0%91%D0%B8%D0%B4%D1%8A%D0%BB" title="Джордж Бидъл – Bulgarian" lang="bg" hreflang="bg" data-title="Джордж Бидъл" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Catalan" lang="ca" hreflang="ca" data-title="George Wells Beadle" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Czech" lang="cs" hreflang="cs" data-title="George Wells Beadle" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – German" lang="de" hreflang="de" data-title="George Wells Beadle" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Estonian" lang="et" hreflang="et" data-title="George Wells Beadle" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Spanish" lang="es" hreflang="es" data-title="George Wells Beadle" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Basque" lang="eu" hreflang="eu" data-title="George Wells Beadle" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AC%D8%B1%D8%AC_%D8%A8%DB%8C%D8%AF%D9%84" title="جرج بیدل – Persian" lang="fa" hreflang="fa" data-title="جرج بیدل" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – French" lang="fr" hreflang="fr" data-title="George Wells Beadle" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Irish" lang="ga" hreflang="ga" data-title="George Wells Beadle" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Galician" lang="gl" hreflang="gl" data-title="George Wells Beadle" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%A1%B0%EC%A7%80_%EC%9B%B0%EC%8A%A4_%EB%B9%84%EB%93%A4" title="조지 웰스 비들 – Korean" lang="ko" hreflang="ko" data-title="조지 웰스 비들" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Croatian" lang="hr" hreflang="hr" data-title="George Wells Beadle" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Ido" lang="io" hreflang="io" data-title="George Wells Beadle" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/George_Beadle" title="George Beadle – Indonesian" lang="id" hreflang="id" data-title="George Beadle" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Italian" lang="it" hreflang="it" data-title="George Wells Beadle" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%92%27%D7%95%D7%A8%D7%92%27_%D7%91%D7%99%D7%93%D7%9C" title="ג'ורג' בידל – Hebrew" lang="he" hreflang="he" data-title="ג'ורג' בידל" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6_%D0%A3%D1%8D%D0%BB%D1%81_%D0%91%D0%B8%D0%B4%D0%BB" title="Джордж Уэлс Бидл – Kazakh" lang="kk" hreflang="kk" data-title="Джордж Уэлс Бидл" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/George_Beadle" title="George Beadle – Swahili" lang="sw" hreflang="sw" data-title="George Beadle" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Georgius_Wells_Beadle" title="Georgius Wells Beadle – Latin" lang="la" hreflang="la" data-title="Georgius Wells Beadle" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Hungarian" lang="hu" hreflang="hu" data-title="George Wells Beadle" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%AF%E1%83%9D%E1%83%A0%E1%83%AF_%E1%83%91%E1%83%98%E1%83%93%E1%83%9A%E1%83%98" title="ჯორჯ ბიდლი – Mingrelian" lang="xmf" hreflang="xmf" data-title="ჯორჯ ბიდლი" data-language-autonym="მარგალური" data-language-local-name="Mingrelian" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D8%A8%DB%8C%D8%AF%D9%84" title="جورج بیدل – Mazanderani" lang="mzn" hreflang="mzn" data-title="جورج بیدل" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/George_Beadle" title="George Beadle – Malay" lang="ms" hreflang="ms" data-title="George Beadle" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Dutch" lang="nl" hreflang="nl" data-title="George Wells Beadle" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B8%E3%83%A7%E3%83%BC%E3%82%B8%E3%83%BB%E3%82%A6%E3%82%A7%E3%83%AB%E3%82%BA%E3%83%BB%E3%83%93%E3%83%BC%E3%83%89%E3%83%AB" title="ジョージ・ウェルズ・ビードル – Japanese" lang="ja" hreflang="ja" data-title="ジョージ・ウェルズ・ビードル" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="George Wells Beadle" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Occitan" lang="oc" hreflang="oc" data-title="George Wells Beadle" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/George_Beadle" title="George Beadle – Uzbek" lang="uz" hreflang="uz" data-title="George Beadle" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%D8%A7%D8%B1%D8%AC_%D9%88%DB%8C%D9%84%D8%B2_%D8%A8%DB%8C%DA%88%D9%84" title="جارج ویلز بیڈل – Western Punjabi" lang="pnb" hreflang="pnb" data-title="جارج ویلز بیڈل" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Low German" lang="nds" hreflang="nds" data-title="George Wells Beadle" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Polish" lang="pl" hreflang="pl" data-title="George Wells Beadle" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/George_Beadle" title="George Beadle – Portuguese" lang="pt" hreflang="pt" data-title="George Beadle" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Romanian" lang="ro" hreflang="ro" data-title="George Wells Beadle" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%91%D0%B8%D0%B4%D0%BB,_%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6_%D0%A3%D1%8D%D0%BB%D1%81" title="Бидл, Джордж Уэлс – Russian" lang="ru" hreflang="ru" data-title="Бидл, Джордж Уэлс" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/George_Beadle" title="George Beadle – Simple English" lang="en-simple" hreflang="en-simple" data-title="George Beadle" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Slovak" lang="sk" hreflang="sk" data-title="George Wells Beadle" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Slovenian" lang="sl" hreflang="sl" data-title="George Wells Beadle" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D8%AC%DB%86%D8%B1%D8%AC_%D9%88%DB%8E%DA%B5%D8%B3_%D8%A8%DB%8E%D8%AF%DA%B5" title="جۆرج وێڵس بێدڵ – Central Kurdish" lang="ckb" hreflang="ckb" data-title="جۆرج وێڵس بێدڵ" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Finnish" lang="fi" hreflang="fi" data-title="George Wells Beadle" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Swedish" lang="sv" hreflang="sv" data-title="George Wells Beadle" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D2%96%D0%BE%D1%80%D2%97_%D0%92%D0%B5%D0%BB%D0%BB%D1%81_%D0%91%D0%B8%D0%B4%D0%BB" title="Җорҗ Веллс Бидл – Tatar" lang="tt" hreflang="tt" data-title="Җорҗ Веллс Бидл" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-tg mw-list-item"><a href="https://tg.wikipedia.org/wiki/%D0%91%D0%B8%D0%B4%D0%BB,_%D2%B6%D0%BE%D1%80%D2%B7_%D0%A3%D1%8D%D0%BB%D1%81" title="Бидл, Ҷорҷ Уэлс – Tajik" lang="tg" hreflang="tg" data-title="Бидл, Ҷорҷ Уэлс" data-language-autonym="Тоҷикӣ" data-language-local-name="Tajik" class="interlanguage-link-target"><span>Тоҷикӣ</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/George_Wells_Beadle" title="George Wells Beadle – Turkish" lang="tr" hreflang="tr" data-title="George Wells Beadle" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk 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class="mw-redirect" title="George Wells Beadle">George Wells Beadle</a>)</span></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">American geneticist (1903–1989)</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">For New Zealand rugby league international, see <a href="/wiki/George_Beadle_(rugby_league)" title="George Beadle (rugby league)">George Beadle (rugby league)</a>.</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">George Beadle</div><div class="honorific-suffix" style="font-size: 77%; font-weight: normal;"><span class="nobold noexcerpt nowraplinks" style="font-size:;"><span style="font-size: 100%;"><a href="/wiki/Foreign_Member_of_the_Royal_Society" class="mw-redirect" title="Foreign Member of the Royal Society">ForMemRS</a></span></span></div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:George_Wells_Beadle.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/en/2/24/George_Wells_Beadle.jpg" decoding="async" width="220" height="311" class="mw-file-element" data-file-width="220" data-file-height="311" /></a></span></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">George Wells Beadle</div><br /><span style="display:none">(<span class="bday">1903-10-22</span>)</span>October 22, 1903<sup id="cite_ref-frs_2-0" class="reference"><a href="#cite_note-frs-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><br /><div style="display:inline" class="birthplace"><a href="/wiki/Wahoo,_Nebraska" title="Wahoo, Nebraska">Wahoo</a>, <a href="/wiki/Nebraska" title="Nebraska">Nebraska</a>, U.S.</div></td></tr><tr><th scope="row" class="infobox-label">Died</th><td class="infobox-data">June 9, 1989<span style="display:none">(1989-06-09)</span> (aged 85)<br /><div style="display:inline" class="deathplace"><a href="/wiki/Pomona,_California" title="Pomona, California">Pomona, California</a>, U.S.</div></td></tr><tr><th scope="row" class="infobox-label">Education</th><td class="infobox-data"><a href="/wiki/University_of_Nebraska" class="mw-redirect" title="University of Nebraska">University of Nebraska</a> (<a href="/wiki/Bachelor_of_Science" title="Bachelor of Science">BS</a>) <br /><a href="/wiki/Cornell_University" title="Cornell University">Cornell University</a> (<a href="/wiki/Master_of_Science" title="Master of Science">MS</a>, <a href="/wiki/Doctor_of_Philosophy" title="Doctor of Philosophy">PhD</a>)</td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style><div class="plainlist"><ul><li><a href="/wiki/One_gene-one_enzyme_hypothesis" class="mw-redirect" title="One gene-one enzyme hypothesis">One gene-one enzyme hypothesis</a></li><li><a href="/wiki/Gene" title="Gene">Gene</a> regulation of biochemical events within cells</li></ul></div></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /><div class="plainlist"> <ul><li><a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a> <span style="font-size:85%;">(1950)</span></li> <li><a href="/wiki/Mendel_Medal_(genetics)" class="mw-redirect" title="Mendel Medal (genetics)">Mendel Medal</a> <span style="font-size:85%;">(1958)</span></li> <li><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> <span style="font-size:85%;">(1958)</span></li> <li><a href="/wiki/Thomas_Hunt_Morgan_Medal" title="Thomas Hunt Morgan Medal">Thomas Hunt Morgan Medal</a> <span style="font-size:85%;">(1984)</span></li></ul> </div></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546" /><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Genetics" title="Genetics">Genetics</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /><div class="plainlist"><ul><li><a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">California Institute of Technology</a></li><li><a href="/wiki/University_of_Chicago" title="University of Chicago">University of Chicago</a></li><li><a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a></li><li><a href="/wiki/Stanford_University" title="Stanford University">Stanford University</a></li></ul></div></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Thesis" title="Thesis">Thesis</a></th><td class="infobox-data"><i><a rel="nofollow" class="external text" href="https://www.proquest.com/docview/301772751/">Genetical and Cytological Studies of Mendelian Asynapsis in Zea mays</a></i> <span style="font-size:97%;">(1930)</span></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Doctoral_advisor" title="Doctoral advisor">Doctoral advisor</a></th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /><div class="plainlist"> <ul><li><a href="/wiki/Rollins_A._Emerson" title="Rollins A. Emerson">Rollins A. Emerson</a></li> <li><a href="/wiki/Lester_W._Sharp" title="Lester W. Sharp">Lester W. Sharp</a><sup id="cite_ref-DevTree_1-0" class="reference"><a href="#cite_note-DevTree-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label">Other academic advisors</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /><div class="plainlist"> <ul><li><a href="/wiki/Franklin_David_Keim" title="Franklin David Keim">Franklin D. Keim</a><sup id="cite_ref-DevTree_1-1" class="reference"><a href="#cite_note-DevTree-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Hunt Morgan</a> <span style="font-size:85%;">(post-doc)</span><sup id="cite_ref-DevTree_1-2" class="reference"><a href="#cite_note-DevTree-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label">Doctoral students</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /><div class="plainlist"> <ul><li><a href="/wiki/Robert_Metzenberg" title="Robert Metzenberg">Robert Metzenberg</a><sup id="cite_ref-DevTree_1-3" class="reference"><a href="#cite_note-DevTree-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label">Other notable students</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /><div class="plainlist"> <ul><li><a href="/wiki/Norman_Horowitz" title="Norman Horowitz">Norman Horowitz</a> <span style="font-size:85%;">(post-doc)</span></li> <li><a href="/wiki/Herschel_K._Mitchell" title="Herschel K. Mitchell">Herschel K. Mitchell</a> <span style="font-size:85%;">(post-doc)</span></li> <li><a href="/wiki/William_D._McElroy" title="William D. McElroy">William D. McElroy</a> <span style="font-size:85%;">(post-doc)</span></li> <li><a href="/wiki/Clement_Markert" title="Clement Markert">Clement Markert</a> <span style="font-size:85%;">(post-doc)</span></li></ul> </div></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>George Wells Beadle</b> (October 22, 1903 – June 9, 1989) was an American geneticist. In 1958 he shared one-half of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> with <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> for their discovery of the role of <a href="/wiki/Gene" title="Gene">genes</a> in regulating biochemical events within cells.<sup id="cite_ref-nas_3-0" class="reference"><a href="#cite_note-nas-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> He also served as the 7th President of the <a href="/wiki/University_of_Chicago" title="University of Chicago">University of Chicago</a>.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>Beadle and Tatum's key experiments involved exposing the bread <a href="/wiki/Mold_(fungus)" class="mw-redirect" title="Mold (fungus)">mold</a> <i><a href="/wiki/Neurospora_crassa" title="Neurospora crassa">Neurospora crassa</a></i> to <a href="/wiki/X-ray" title="X-ray">x-rays</a>, causing <a href="/wiki/Mutation" title="Mutation">mutations</a>. In a series of experiments, they showed that these mutations caused changes in specific <a href="/wiki/Enzyme" title="Enzyme">enzymes</a> involved in <a href="/wiki/Metabolic_pathway" title="Metabolic pathway">metabolic pathways</a>. These experiments led them to propose a direct link between genes and enzymatic reactions, known as the <a href="/wiki/One_gene-one_enzyme_hypothesis" class="mw-redirect" title="One gene-one enzyme hypothesis">One gene-one enzyme hypothesis</a>.<sup id="cite_ref-onegeneonenz_6-0" class="reference"><a href="#cite_note-onegeneonenz-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Education_and_early_life">Education and early life</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Beadle&action=edit&section=1" title="Edit section: Education and early life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>George Wells Beadle was born in <a href="/wiki/Wahoo,_Nebraska" title="Wahoo, Nebraska">Wahoo, Nebraska</a>. He was the son of Chauncey Elmer Beadle and Hattie Albro, who owned and operated a 40-acre (160,000 m<sup>2</sup>) farm nearby.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> George was educated at the Wahoo High School and might himself have become a farmer if one of his teachers at school had not directed his mind towards science and persuaded him to go to the College of Agriculture in Lincoln, Nebraska. In 1926 he earned his <a href="/wiki/Bachelor_of_Science" title="Bachelor of Science">Bachelor of Science</a> degree at the <a href="/wiki/University_of_Nebraska" class="mw-redirect" title="University of Nebraska">University of Nebraska</a> and subsequently worked for a year with Professor <a href="/wiki/Franklin_David_Keim" title="Franklin David Keim">F.D. Keim</a>, who was studying hybrid wheat. In 1927 he earned his <a href="/wiki/Master_of_Science" title="Master of Science">Master of Science</a> degree, and Professor Keim secured for him a post as Teaching Assistant at <a href="/wiki/Cornell_University" title="Cornell University">Cornell University</a>, where he worked, until 1931, with Professors <a href="/wiki/Rollins_A._Emerson" title="Rollins A. Emerson">R.A. Emerson</a> and <a href="/wiki/Lester_W._Sharp" title="Lester W. Sharp">L.W. Sharp</a> on Mendelian asynapsis in <i><a href="/wiki/Zea_mays" class="mw-redirect" title="Zea mays">Zea mays</a></i>.<sup id="cite_ref-phd_10-0" class="reference"><a href="#cite_note-phd-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> For this work he obtained, in 1931, his <a href="/wiki/Doctor_of_Philosophy" title="Doctor of Philosophy">Doctor of Philosophy</a> degree.<sup id="cite_ref-phd_10-1" class="reference"><a href="#cite_note-phd-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Career_and_research">Career and research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Beadle&action=edit&section=2" title="Edit section: Career and research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In 1931 Fellowship at the <a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">California Institute of Technology</a> at Pasadena, where he remained from 1931 until 1936. During this period he continued his work on Indian corn and began, in collaboration with Professors <a href="/wiki/Theodosius_Dobzhansky" title="Theodosius Dobzhansky">Theodosius Dobzhansky</a>, S. Emerson, and <a href="/wiki/Alfred_Sturtevant" title="Alfred Sturtevant">Alfred Sturtevant</a>, work on crossing-over in the fruit fly, <i><a href="/wiki/Drosophila_melanogaster" title="Drosophila melanogaster">Drosophila melanogaster</a></i>.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (January 2014)">citation needed</span></a></i>]</sup> </p><p>In 1935 Beadle visited <a href="/wiki/Paris" title="Paris">Paris</a> for six months to work with Professor <a href="/wiki/Boris_Ephrussi" title="Boris Ephrussi">Boris Ephrussi</a> at the <a href="/wiki/Institut_de_biologie_physico-chimique" title="Institut de biologie physico-chimique">Institut de Biologie physico-chimique</a>. Together they began the study of the development of eye pigment in Drosophila which later led to the work on the biochemistry of the genetics of the fungus Neurospora for which Beadle and Edward Lawrie Tatum were together awarded the 1958 Nobel Prize for Physiology or Medicine. </p><p>In 1936 Beadle left the California Institute of Technology to become Assistant Professor of Genetics at <a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a>. A year later he was appointed Professor of Biology (Genetics) at <a href="/wiki/Stanford_University" title="Stanford University">Stanford University</a> and there he remained for nine years, working for most of this period in collaboration with Tatum. This work of Beadle and Tatum led to an important generalization. This was that most mutants unable to grow on minimal medium, but able to grow on “complete” medium, each require addition of only one particular supplement for growth on minimal medium. If the synthesis of a particular nutrient (such as an <a href="/wiki/Amino_acid" title="Amino acid">amino acid</a> or <a href="/wiki/Vitamin" title="Vitamin">vitamin</a>) was disrupted by <a href="/wiki/Mutation" title="Mutation">mutation</a>, that mutant strain could be grown by adding the necessary nutrient to the minimal medium.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> This finding suggested that most mutations affected only a single <a href="/wiki/Metabolic_pathway" title="Metabolic pathway">metabolic pathway</a>. Further evidence obtained soon after the initial findings tended to show that generally only a single step in the pathway is blocked. Following their first report of three such <a href="/wiki/Auxotrophy" title="Auxotrophy">auxotroph</a> mutants in 1941, Beadle and Tatum used this method to create series of related mutants and determined the order in which amino acids and some other <a href="/wiki/Metabolite" title="Metabolite">metabolites</a> were synthesized in several metabolic pathways. The obvious inference from these experiments was that each gene mutation affects the activity of a single enzyme. This led directly to the <a href="/wiki/One_gene-one_enzyme_hypothesis" class="mw-redirect" title="One gene-one enzyme hypothesis">one gene-one enzyme hypothesis</a>, which, with certain qualifications and refinements, has remained essentially valid to the present day. As recalled by Horowitz,<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> the work of Beadle and Tatum also demonstrated that <a href="/wiki/Gene" title="Gene">genes</a> have an essential role in <a href="/wiki/Biosynthesis" title="Biosynthesis">biosynthesis</a>. At the time of the experiments (1941), non-geneticists still generally believed that genes governed only trivial biological traits, such as eye color, and bristle arrangement in fruit flies, while basic biochemistry was determined in the cytoplasm by unknown processes. Also, many respected geneticists thought that gene action was far too complicated to be resolved by any simple experiment. Thus Beadle and Tatum brought about a fundamental revolution in our understanding of genetics. </p><p>In 1946 Beadle returned to the California Institute of Technology as Professor of Biology and Chairman of the Division of Biology. Here he remained until January 1961 when he was elected Chancellor of the University of Chicago and, in the autumn of the same year, President of this university. </p><p>After retiring, Beadle undertook a remarkable experiment in <a href="/wiki/Maize" title="Maize">maize</a> genetics. In several laboratories he grew a series of <a href="/wiki/Teosinte" class="mw-redirect" title="Teosinte">Teosinte</a>/Maize crosses. Then he crossed these progeny with each other. He looked for the rate of appearance of parent phenotypes among this second generation. The vast majority of these plants were intermediate between maize and Teosinte in their features, but about 1 in 500 of the plants were identical to either the parent maize or the parent teosinte. Using the mathematics of Mendelian genetics, he calculated that this showed a difference between maize and teosinte of about 5 or 6 genetic loci. This demonstration was so compelling that most scientists now agree that Teosinte is the wild progenitor of maize.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p><p>During his career, Beadle has received many honors. These include the Honorary <a href="/wiki/Doctor_of_Science" title="Doctor of Science">Doctor of Science</a> of the following Universities: Yale (1947), Nebraska (1949), Northwestern University (1952), Rutgers University (1954), Kenyon College (1955), Wesleyan University (1956), the <a href="/wiki/University_of_Birmingham" title="University of Birmingham">University of Birmingham</a> and the <a href="/wiki/University_of_Oxford" title="University of Oxford">University of Oxford</a>, England (1959), Pomona College (1961), and Lake Forest College (1962). In 1962 he was also given the honorary degree of LL.D. by the University of California, Los Angeles. He was elected a Fellow of the <a href="/wiki/American_Academy_of_Arts_and_Sciences" title="American Academy of Arts and Sciences">American Academy of Arts and Sciences</a> in 1946.<sup id="cite_ref-AAAS_15-0" class="reference"><a href="#cite_note-AAAS-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> He also received the Lasker Award of the American Public Health Association (1950), the Dyer Award (1951), the Emil Christian Hansen Prize of Denmark (1953), the Albert Einstein Commemorative Award in Science (1958), the Nobel Prize in Physiology or Medicine 1958 with Edward Tatum and <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a>, the National Award of the American Cancer Society (1959), and the Kimber Genetics Award of the National Academy of Sciences (1960). </p> <div class="mw-heading mw-heading3"><h3 id="Awards_and_honors">Awards and honors</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Beadle&action=edit&section=3" title="Edit section: Awards and honors"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In addition to the Nobel Prize, Beadle received numerous other awards. Beadle was a member of several learned societies, he was a <a href="/wiki/Member_of_the_National_Academy_of_Sciences" title="Member of the National Academy of Sciences">Member of the National Academy of Sciences</a><sup id="cite_ref-nasmember_16-0" class="reference"><a href="#cite_note-nasmember-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> (and Chairman of Committee on Genetic Effects of Atomic Radiation), the <a href="/wiki/Genetics_Society_of_America" title="Genetics Society of America">Genetics Society of America</a> (President in 1946), the <a href="/wiki/American_Association_for_the_Advancement_of_Science" title="American Association for the Advancement of Science">American Association for the Advancement of Science</a> (President in 1955), the <a href="/wiki/American_Cancer_Society" title="American Cancer Society">American Cancer Society</a> (Chairman of Scientific Advisory Council), a <a href="/wiki/Foreign_Member_of_the_Royal_Society" class="mw-redirect" title="Foreign Member of the Royal Society">Foreign Member of the Royal Society</a> (ForMemRS) of London,<sup id="cite_ref-frs_2-1" class="reference"><a href="#cite_note-frs-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> the Danish Royal Academy of Science and the <a href="/wiki/American_Philosophical_Society" title="American Philosophical Society">American Philosophical Society</a>.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>The <a href="/wiki/George_W._Beadle_Award" title="George W. Beadle Award">George W. Beadle Award</a> of the <a href="/wiki/Genetics_Society_of_America" title="Genetics Society of America">Genetics Society of America</a> is named in his honor. George Beadle Middle School in <a href="/wiki/Millard,_Nebraska" class="mw-redirect" title="Millard, Nebraska">Millard, Nebraska</a> (Part of the <a href="/wiki/Millard_Public_Schools" title="Millard Public Schools">Millard Public Schools</a> district) was named after him. It opened in 2001. The Beadle Center, which houses the Department of Biochemistry at the <a href="/wiki/University_of_Nebraska%E2%80%93Lincoln" title="University of Nebraska–Lincoln">University of Nebraska-Lincoln</a>, is also named after George Beadle. </p> <div class="mw-heading mw-heading2"><h2 id="Personal_life">Personal life</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Beadle&action=edit&section=4" title="Edit section: Personal life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Beadle was married twice. By his first wife he had a son, David, who now lives at The Hague, the Netherlands.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (March 2020)">citation needed</span></a></i>]</sup> His second wife, Muriel McClure (1915–1994), a well-known writer, was born in California. Beadle's chief hobbies were rockclimbing, skiing, and gardening. He is credited with the first ascent of <a href="/wiki/Mount_Doonerak" title="Mount Doonerak">Mount Doonerak</a> in Alaska.<sup id="cite_ref-Wood_18-0" class="reference"><a href="#cite_note-Wood-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> He was a member of <a href="/wiki/FarmHouse" title="FarmHouse">FarmHouse</a> <a href="/wiki/Fraternity" title="Fraternity">fraternity</a> while at the University of Nebraska.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (April 2019)">citation needed</span></a></i>]</sup> </p><p>Beadle died on June 9, 1989 at a retirement community in <a href="/wiki/Pomona,_California" title="Pomona, California">Pomona, California</a> from complications of <a href="/wiki/Alzheimer%27s_disease" title="Alzheimer's disease">Alzheimer's disease</a>, aged 85.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> He was an atheist.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Beadle&action=edit&section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-DevTree-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-DevTree_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-DevTree_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-DevTree_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-DevTree_1-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://academictree.org/dev/peopleinfo.php?pid=13579">"DevTree - George Wells Beadle"</a>. <i>academictree.org</i>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=academictree.org&rft.atitle=DevTree+-+George+Wells+Beadle&rft_id=https%3A%2F%2Facademictree.org%2Fdev%2Fpeopleinfo.php%3Fpid%3D13579&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-frs-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-frs_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-frs_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFHorowitz1995" class="citation journal cs1"><a href="/wiki/Norman_Horowitz" title="Norman Horowitz">Horowitz, Norman H.</a> (1995). <a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frsbm.1995.0003">"George Wells Beadle. 23 October 1903-9 June 1989"</a>. <i><a href="/wiki/Biographical_Memoirs_of_Fellows_of_the_Royal_Society" title="Biographical Memoirs of Fellows of the Royal Society">Biographical Memoirs of Fellows of the Royal Society</a></i>. <b>41</b>. <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a>: <span class="nowrap">44–</span>54. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frsbm.1995.0003">10.1098/rsbm.1995.0003</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11615361">11615361</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biographical+Memoirs+of+Fellows+of+the+Royal+Society&rft.atitle=George+Wells+Beadle.+23+October+1903-9+June+1989&rft.volume=41&rft.pages=%3Cspan+class%3D%22nowrap%22%3E44-%3C%2Fspan%3E54&rft.date=1995&rft_id=info%3Adoi%2F10.1098%2Frsbm.1995.0003&rft_id=info%3Apmid%2F11615361&rft.aulast=Horowitz&rft.aufirst=Norman+H.&rft_id=https%3A%2F%2Fdoi.org%2F10.1098%252Frsbm.1995.0003&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-nas-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-nas_3-0">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.nasonline.org/publications/biographical-memoirs/memoir-pdfs/beadle-g-w.pdf">George W. Beadle</a> — <i><a href="/wiki/Biographical_Memoirs_of_the_National_Academy_of_Sciences" title="Biographical Memoirs of the National Academy of Sciences">Biographical Memoirs</a></i> of the <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFStern1954" class="citation journal cs1">Stern, C. (1954). <a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.119.3086.229">"George W. Beadle"</a>. <i>Science</i>. <b>119</b> (3086): <span class="nowrap">229–</span>230. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1954Sci...119..229S">1954Sci...119..229S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.119.3086.229">10.1126/science.119.3086.229</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/13135519">13135519</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=George+W.+Beadle&rft.volume=119&rft.issue=3086&rft.pages=%3Cspan+class%3D%22nowrap%22%3E229-%3C%2Fspan%3E230&rft.date=1954&rft_id=info%3Apmid%2F13135519&rft_id=info%3Adoi%2F10.1126%2Fscience.119.3086.229&rft_id=info%3Abibcode%2F1954Sci...119..229S&rft.aulast=Stern&rft.aufirst=C.&rft_id=https%3A%2F%2Fdoi.org%2F10.1126%252Fscience.119.3086.229&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://president.uchicago.edu/directory/george-w-beadle">"George W. Beadle | Office of the President | The University of Chicago"</a>. <i>president.uchicago.edu</i>. 12 June 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">2019-06-22</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=president.uchicago.edu&rft.atitle=George+W.+Beadle+%7C+Office+of+the+President+%7C+The+University+of+Chicago&rft.date=2012-06-12&rft_id=https%3A%2F%2Fpresident.uchicago.edu%2Fdirectory%2Fgeorge-w-beadle&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-onegeneonenz-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-onegeneonenz_6-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFBeadleTatum1941" class="citation journal cs1">Beadle, G. W.; Tatum, E. L. (1941). <a rel="nofollow" class="external text" href="http://authors.library.caltech.edu/13093/1/BEApnas41.pdf">"Genetic Control of Biochemical Reactions in Neurospora"</a> <span class="cs1-format">(PDF)</span>. <i>Proceedings of the National Academy of Sciences</i>. <b>27</b> (11): <span class="nowrap">499–</span>506. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1941PNAS...27..499B">1941PNAS...27..499B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.27.11.499">10.1073/pnas.27.11.499</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1078370">1078370</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16588492">16588492</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences&rft.atitle=Genetic+Control+of+Biochemical+Reactions+in+Neurospora&rft.volume=27&rft.issue=11&rft.pages=%3Cspan+class%3D%22nowrap%22%3E499-%3C%2Fspan%3E506&rft.date=1941&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1078370%23id-name%3DPMC&rft_id=info%3Apmid%2F16588492&rft_id=info%3Adoi%2F10.1073%2Fpnas.27.11.499&rft_id=info%3Abibcode%2F1941PNAS...27..499B&rft.aulast=Beadle&rft.aufirst=G.+W.&rft.au=Tatum%2C+E.+L.&rft_id=http%3A%2F%2Fauthors.library.caltech.edu%2F13093%2F1%2FBEApnas41.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text">Paul Berg and Maxine Singer. <i>George Beadle: An Uncommon Farmer. The Emergence of Genetics in the 20th Century</i>. Cold Springs Harbor Laboratory Press, 2003. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-87969-688-5" title="Special:BookSources/0-87969-688-5">0-87969-688-5</a></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://osulibrary.oregonstate.edu/specialcollections/coll/pauling/blood/people/beadle.html">Key Participants: George Beadle</a> - <i>It's in the Blood! A Documentary History of Linus Pauling, Hemoglobin, and Sickle Cell Anemia</i></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFBeadle1974" class="citation journal cs1">Beadle, G. W. (1974). <a rel="nofollow" class="external text" href="https://doi.org/10.1146%2Fannurev.bi.43.070174.000245">"Recollections"</a>. <i>Annual Review of Biochemistry</i>. <b>43</b>: <span class="nowrap">1–</span>13. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1146%2Fannurev.bi.43.070174.000245">10.1146/annurev.bi.43.070174.000245</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/4605017">4605017</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Annual+Review+of+Biochemistry&rft.atitle=Recollections&rft.volume=43&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1-%3C%2Fspan%3E13&rft.date=1974&rft_id=info%3Adoi%2F10.1146%2Fannurev.bi.43.070174.000245&rft_id=info%3Apmid%2F4605017&rft.aulast=Beadle&rft.aufirst=G.+W.&rft_id=https%3A%2F%2Fdoi.org%2F10.1146%252Fannurev.bi.43.070174.000245&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-phd-10"><span class="mw-cite-backlink">^ <a href="#cite_ref-phd_10-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-phd_10-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFBeadle1930" class="citation thesis cs1"><a href="/wiki/George_Wells_Beadle" class="mw-redirect" title="George Wells Beadle">Beadle, George Wells</a> (1930). <a rel="nofollow" class="external text" href="https://www.proquest.com/docview/301772751/"><i>Genetical and Cytological Studies of Mendelian Asynapsis in Zea mays</i></a> (PhD thesis). Cornell University.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adissertation&rft.title=Genetical+and+Cytological+Studies+of+Mendelian+Asynapsis+in+Zea+mays&rft.degree=PhD&rft.inst=Cornell+University&rft.date=1930&rft.aulast=Beadle&rft.aufirst=George+Wells&rft_id=https%3A%2F%2Fwww.proquest.com%2Fdocview%2F301772751%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text">Beadle, G. W. (1966) "Biochemical genetics: some recollections", pp. 23-32 in Phage and the Origins of Molecular Biology, edited by J. Cairns, G. S. Stent and J. D. Watson. Cold Spring Harbor Symposia, Cold Spring Harbor Laboratory of Quantitative Biology, NY. ASIN: B005F08IQ8</span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text">Horowitz NH (May, 1996). "The sixtieth anniversary of biochemical genetics". Genetics. 143 (1): 1-4. doi:10.1093/genetics/143.1.1.PMC 1207243. PMID 8722756</span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFBeadle1980" class="citation journal cs1">Beadle, G. W. (1980). "The ancestry of corn". <i>Scientific American</i>. <b>242</b> (1): <span class="nowrap">112–</span>119. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1980SciAm.242a.112B">1980SciAm.242a.112B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fscientificamerican0180-112">10.1038/scientificamerican0180-112</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Scientific+American&rft.atitle=The+ancestry+of+corn&rft.volume=242&rft.issue=1&rft.pages=%3Cspan+class%3D%22nowrap%22%3E112-%3C%2Fspan%3E119&rft.date=1980&rft_id=info%3Adoi%2F10.1038%2Fscientificamerican0180-112&rft_id=info%3Abibcode%2F1980SciAm.242a.112B&rft.aulast=Beadle&rft.aufirst=G.+W.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFBeadle" class="citation web cs1">Beadle, George. <a rel="nofollow" class="external text" href="http://www.clas.ufl.edu/users/dcgrove/mexarchreadings/corn.pdf">"The Ancestry of Corn"</a> <span class="cs1-format">(PDF)</span><span class="reference-accessdate">. Retrieved <span class="nowrap">2 July</span> 2014</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=The+Ancestry+of+Corn&rft.aulast=Beadle&rft.aufirst=George&rft_id=http%3A%2F%2Fwww.clas.ufl.edu%2Fusers%2Fdcgrove%2Fmexarchreadings%2Fcorn.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-AAAS-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-AAAS_15-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.amacad.org/publications/BookofMembers/ChapterB.pdf">"Book of Members, 1780-2010: Chapter B"</a> <span class="cs1-format">(PDF)</span>. American Academy of Arts and Sciences<span class="reference-accessdate">. Retrieved <span class="nowrap">May 28,</span> 2011</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Book+of+Members%2C+1780-2010%3A+Chapter+B&rft.pub=American+Academy+of+Arts+and+Sciences&rft_id=http%3A%2F%2Fwww.amacad.org%2Fpublications%2FBookofMembers%2FChapterB.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-nasmember-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-nasmember_16-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.nasonline.org/member-directory/deceased-members/57907.html">"G. Beadle"</a>. <i>Nasonline.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">29 April</span> 2019</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Nasonline.org&rft.atitle=G.+Beadle&rft_id=http%3A%2F%2Fwww.nasonline.org%2Fmember-directory%2Fdeceased-members%2F57907.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://search.amphilsoc.org/memhist/search?smode=advanced;subject=2.%20Biological%20Sciences;startDoc=21">"APS Member History"</a>. <i>Search.amphilsoc.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2019-04-07</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Search.amphilsoc.org&rft.atitle=APS+Member+History&rft_id=https%3A%2F%2Fsearch.amphilsoc.org%2Fmemhist%2Fsearch%3Fsmode%3Dadvanced%3Bsubject%3D2.%2520Biological%2520Sciences%3BstartDoc%3D21&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-Wood-18"><span class="mw-cite-backlink"><b><a href="#cite_ref-Wood_18-0">^</a></b></span> <span class="reference-text">Michael Wood and Colby Coombs, <i>Alaska: A Climbing Guide</i>, The Mountaineers Books, 2002, page 40.</span> </li> <li id="cite_note-19"><span class="mw-cite-backlink"><b><a href="#cite_ref-19">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.nytimes.com/1989/06/12/obituaries/george-w-beadle-85-geneticist-and-nobel-prize-winner-is-dead.html">"George W. Beadle, 85, Geneticist And Nobel Prize Winner, Is Dead"</a>. <a href="/wiki/The_New_York_Times" title="The New York Times">The New York Times</a>. June 12, 1989<span class="reference-accessdate">. Retrieved <span class="nowrap">January 8,</span> 2025</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=George+W.+Beadle%2C+85%2C+Geneticist+And+Nobel+Prize+Winner%2C+Is+Dead&rft.pub=The+New+York+Times&rft.date=1989-06-12&rft_id=https%3A%2F%2Fwww.nytimes.com%2F1989%2F06%2F12%2Fobituaries%2Fgeorge-w-beadle-85-geneticist-and-nobel-prize-winner-is-dead.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Beadle" class="Z3988"></span></span> </li> <li id="cite_note-20"><span class="mw-cite-backlink"><b><a href="#cite_ref-20">^</a></b></span> <span class="reference-text">George Beadle, An Uncommon Farmer: The Emergence of Genetics in the 20th Century. CSHL Press. 2003. p. 273. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780879696887" title="Special:BookSources/9780879696887">9780879696887</a>. Beadle's views on this occasion were somewhat more tempered than David's characterization of him as a "vehement atheist," and from his earliest days "intolerant of religion and other forms of superstition."</span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Beadle&action=edit&section=6" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Wikiquote-logo.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Wikiquote-logo.svg/34px-Wikiquote-logo.svg.png" decoding="async" width="34" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Wikiquote-logo.svg/51px-Wikiquote-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Wikiquote-logo.svg/68px-Wikiquote-logo.svg.png 2x" data-file-width="300" data-file-height="355" /></a></span></div> <div class="side-box-text plainlist">Wikiquote has quotations related to <i><b><a href="https://en.wikiquote.org/wiki/Special:Search/George_Beadle" class="extiw" title="q:Special:Search/George Beadle">George Beadle</a></b></i>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="https://www.lib.uchicago.edu/e/scrc/findingaids/view.php?eadid=ICU.SPCL.BEADLEG">Guide to the George Wells Beadle Papers 1908-1981</a> at the <a rel="nofollow" class="external text" href="https://www.lib.uchicago.edu/scrc/">University of Chicago Special Collections Research Center</a></li> <li><a rel="nofollow" class="external text" href="https://www.lib.uchicago.edu/e/scrc/findingaids/view.php?eadid=ICU.SPCL.OFCPRESBEADLE">Guide to the University of Chicago Office of the President, Beadle Administration Records 1916-1968</a> at the <a rel="nofollow" class="external text" href="https://www.lib.uchicago.edu/scrc/">University of Chicago Special Collections Research Center</a></li> <li><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/364">George Beadle</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q238739#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></li></ul> <table class="wikitable succession-box noprint" style="margin:0.5em auto; font-size:small;clear:both;"> <tbody><tr> <th colspan="3" style="border-top: 5px solid #DAA520;">Academic offices </th></tr> <tr style="text-align:center;"> <td style="width:30%;" rowspan="1">Preceded by<div style="font-weight: bold"><a href="/wiki/Lawrence_A._Kimpton" title="Lawrence A. Kimpton">Lawrence A. Kimpton</a></div> </td> <td style="width: 40%; text-align: center;" rowspan="1"><b> <a href="/wiki/List_of_University_of_Chicago_faculty#University_Presidents" title="List of University of Chicago faculty">President of the University of Chicago</a> </b><br />1961–1968 </td> <td style="width: 30%; text-align: center;" rowspan="1">Succeeded by<div style="font-weight: bold"><a href="/wiki/Edward_H._Levi" title="Edward H. Levi">Edward H. Levi</a></div> </td></tr> </tbody></table> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist 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template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Special:EditPage/Template:Nobel Prize in Physiology or Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine170" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Physiology_or_Medicine" title="List of Nobel laureates in Physiology or Medicine">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_Macleod_(physiologist)" title="John Macleod (physiologist)">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1925</span></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Krebs_(biochemist)" title="Hans Krebs (biochemist)">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a class="mw-selflink selflink">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_MacLeod_Cormack" title="Allan MacLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards_(physiologist)" title="Robert Edwards (physiologist)">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>posthumously</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe_(neuroscientist)" title="John O'Keefe (neuroscientist)">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap"><a href="/wiki/2020_Nobel_Prize_in_Physiology_or_Medicine" title="2020 Nobel Prize in Physiology or Medicine">2020</a>: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton_(virologist)" class="mw-redirect" title="Michael Houghton (virologist)">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap"><a href="/wiki/2021_Nobel_Prize_in_Physiology_or_Medicine" title="2021 Nobel Prize in Physiology or Medicine">2021</a>: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap"><a href="/wiki/2022_Nobel_Prize_in_Physiology_or_Medicine" title="2022 Nobel Prize in Physiology or Medicine">2022</a>: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap"><a href="/wiki/2023_Nobel_Prize_in_Physiology_or_Medicine" title="2023 Nobel Prize in Physiology or Medicine">2023</a>: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="1958_Nobel_Prize_laureates913" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231" /><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:1958_Nobel_Prize_winners" title="Template:1958 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:1958_Nobel_Prize_winners" title="Template talk:1958 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:1958_Nobel_Prize_winners" title="Special:EditPage/Template:1958 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="1958_Nobel_Prize_laureates913" style="font-size:114%;margin:0 4em">1958 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Frederick_Sanger" title="Frederick Sanger">Frederick Sanger</a> (Great Britain)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/1958_Nobel_Prize_in_Literature" title="1958 Nobel Prize in Literature">1958</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Boris_Pasternak" title="Boris Pasternak">Boris Pasternak</a> (Soviet Union)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dominique_Pire" title="Dominique Pire">Dominique Pire</a> (Belgium)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pavel_Cherenkov" title="Pavel Cherenkov">Pavel Cherenkov</a> (Soviet Union)</li> <li><a href="/wiki/Ilya_Frank" title="Ilya Frank">Ilya Frank</a> (Soviet Union)</li> <li><a href="/wiki/Igor_Tamm" title="Igor Tamm">Igor Tamm</a> (Soviet Union)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">George Wells Beadle</a> (United States)</li> <li><a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Lawrie Tatum</a> (United States)</li> <li><a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a> (United States)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/Template:1953_Nobel_Prize_winners" title="Template:1953 Nobel Prize winners">1953</a></dd> <dd><a href="/wiki/Template:1954_Nobel_Prize_winners" title="Template:1954 Nobel Prize winners">1954</a></dd> <dd><a href="/wiki/Template:1955_Nobel_Prize_winners" title="Template:1955 Nobel Prize winners">1955</a></dd> <dd><a href="/wiki/Template:1956_Nobel_Prize_winners" title="Template:1956 Nobel Prize winners">1956</a></dd> <dd><a href="/wiki/Template:1957_Nobel_Prize_winners" title="Template:1957 Nobel Prize winners">1957</a></dd> <dd><b>1958 </b></dd> <dd><a href="/wiki/Template:1959_Nobel_Prize_winners" title="Template:1959 Nobel Prize winners">1959</a></dd> <dd><a href="/wiki/Template:1960_Nobel_Prize_winners" title="Template:1960 Nobel Prize winners">1960</a></dd> <dd><a href="/wiki/Template:1961_Nobel_Prize_winners" title="Template:1961 Nobel Prize winners">1961</a></dd> <dd><a href="/wiki/Template:1962_Nobel_Prize_winners" title="Template:1962 Nobel Prize winners">1962</a></dd> <dd><a href="/wiki/Template:1963_Nobel_Prize_winners" title="Template:1963 Nobel Prize winners">1963</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox" aria-labelledby="Presidents_of_the_University_of_Chicago45" style="padding:3px"><table class="nowraplinks mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background-color:#800000;color:white;"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231" /><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:University_of_Chicago_presidents" title="Template:University of Chicago presidents"><abbr title="View this template" style="color:white">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:University_of_Chicago_presidents" title="Template talk:University of Chicago presidents"><abbr title="Discuss this template" style="color:white">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:University_of_Chicago_presidents" title="Special:EditPage/Template:University of Chicago presidents"><abbr title="Edit this template" style="color:white">e</abbr></a></li></ul></div><div id="Presidents_of_the_University_of_Chicago45" style="font-size:114%;margin:0 4em"><b>Presidents of the University of Chicago</b></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/William_Rainey_Harper" title="William Rainey Harper">Harper</a></li> <li><a href="/wiki/Harry_Pratt_Judson" title="Harry Pratt Judson">Judson</a></li> <li><a href="/wiki/Ernest_DeWitt_Burton" title="Ernest DeWitt Burton">Burton</a></li> <li><a href="/wiki/Max_Mason" title="Max Mason">Mason</a></li> <li><a href="/wiki/Robert_Maynard_Hutchins" title="Robert Maynard Hutchins">Hutchins</a></li> <li><a href="/wiki/Lawrence_A._Kimpton" title="Lawrence A. Kimpton">Kimpton</a></li> <li><a class="mw-selflink selflink">Beadle</a></li> <li><a href="/wiki/Edward_H._Levi" title="Edward H. Levi">Levi</a></li> <li><a href="/wiki/John_T._Wilson" title="John T. Wilson">Wilson</a></li> <li><a href="/wiki/Hanna_Holborn_Gray" title="Hanna Holborn Gray">Gray</a></li> <li><a href="/wiki/Hugo_F._Sonnenschein" title="Hugo F. Sonnenschein">Sonnenschein</a></li> <li><a href="/wiki/Don_Michael_Randel" title="Don Michael Randel">Randel</a></li> <li><a href="/wiki/Robert_Zimmer" title="Robert Zimmer">Zimmer</a></li> <li><a href="/wiki/Paul_Alivisatos" title="Paul Alivisatos">Alivisatos</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox" aria-labelledby="Time_Persons_of_the_Year103" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231" /><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Time_Persons_of_the_Year" title="Template:Time Persons of the Year"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Time_Persons_of_the_Year" title="Template talk:Time Persons of the Year"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Time_Persons_of_the_Year" title="Special:EditPage/Template:Time Persons of the Year"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Time_Persons_of_the_Year103" style="font-size:114%;margin:0 4em"><a href="/wiki/Time_Person_of_the_Year" title="Time Person of the Year"><i>Time</i> Persons of the Year</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1927–1950</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap"><a href="/wiki/Charles_Lindbergh" title="Charles Lindbergh">Charles Lindbergh</a> (1927)</span></li> <li><span class="nowrap"><a href="/wiki/Walter_Chrysler" title="Walter Chrysler">Walter Chrysler</a> (1928)</span></li> <li><span class="nowrap"><a href="/wiki/Owen_D._Young" title="Owen D. Young">Owen D. Young</a> (1929)</span></li> <li><span class="nowrap"><a href="/wiki/Mahatma_Gandhi" title="Mahatma Gandhi">Mohandas Gandhi</a> (1930)</span></li> <li><span class="nowrap"><a href="/wiki/Pierre_Laval" title="Pierre Laval">Pierre Laval</a> (1931)</span></li> <li><span class="nowrap"><a href="/wiki/Franklin_D._Roosevelt" title="Franklin D. Roosevelt">Franklin D. Roosevelt</a> (1932)</span></li> <li><span class="nowrap"><a href="/wiki/Hugh_S._Johnson" title="Hugh S. Johnson">Hugh S. Johnson</a> (1933)</span></li> <li><span class="nowrap"><a href="/wiki/Franklin_D._Roosevelt" title="Franklin D. Roosevelt">Franklin D. Roosevelt</a> (1934)</span></li> <li><span class="nowrap"><a href="/wiki/Haile_Selassie" title="Haile Selassie">Haile Selassie</a> (1935)</span></li> <li><span class="nowrap"><a href="/wiki/Wallis_Simpson" title="Wallis Simpson">Wallis Simpson</a> (1936)</span></li> <li><span class="nowrap"><a href="/wiki/Chiang_Kai-shek" title="Chiang Kai-shek">Chiang Kai-shek</a> / <a href="/wiki/Soong_Mei-ling" title="Soong Mei-ling">Soong Mei-ling</a> (1937)</span></li> <li><span class="nowrap"><a href="/wiki/Adolf_Hitler" title="Adolf Hitler">Adolf Hitler</a> (1938)</span></li> <li><span class="nowrap"><a href="/wiki/Joseph_Stalin" title="Joseph Stalin">Joseph Stalin</a> (1939)</span></li> <li><span class="nowrap"><a href="/wiki/Winston_Churchill" title="Winston Churchill">Winston Churchill</a> (1940)</span></li> <li><span class="nowrap"><a href="/wiki/Franklin_D._Roosevelt" title="Franklin D. Roosevelt">Franklin D. Roosevelt</a> (1941)</span></li> <li><span class="nowrap"><a href="/wiki/Joseph_Stalin" title="Joseph Stalin">Joseph Stalin</a> (1942)</span></li> <li><span class="nowrap"><a href="/wiki/George_C._Marshall" title="George C. Marshall">George Marshall</a> (1943)</span></li> <li><span class="nowrap"><a href="/wiki/Dwight_D._Eisenhower" title="Dwight D. Eisenhower">Dwight D. Eisenhower</a> (1944)</span></li> <li><span class="nowrap"><a href="/wiki/Harry_S._Truman" title="Harry S. Truman">Harry S. Truman</a> (1945)</span></li> <li><span class="nowrap"><a href="/wiki/James_F._Byrnes" title="James F. Byrnes">James F. Byrnes</a> (1946)</span></li> <li><span class="nowrap"><a href="/wiki/George_C._Marshall" title="George C. Marshall">George Marshall</a> (1947)</span></li> <li><span class="nowrap"><a href="/wiki/Harry_S._Truman" title="Harry S. Truman">Harry S. Truman</a> (1948)</span></li> <li><span class="nowrap"><a href="/wiki/Winston_Churchill" title="Winston Churchill">Winston Churchill</a> (1949)</span></li> <li><span class="nowrap"><a href="/wiki/Korean_War" title="Korean War">The American Fighting-Man</a> (1950)</span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap"><a href="/wiki/Mohammad_Mosaddegh" title="Mohammad Mosaddegh">Mohammed Mosaddeq</a> (1951)</span></li> <li><span class="nowrap"><a href="/wiki/Elizabeth_II" title="Elizabeth II">Elizabeth II </a> (1952)</span></li> <li><span class="nowrap"><a href="/wiki/Konrad_Adenauer" title="Konrad Adenauer">Konrad Adenauer</a> (1953)</span></li> <li><span class="nowrap"><a href="/wiki/John_Foster_Dulles" title="John Foster Dulles">John Foster Dulles</a> (1954)</span></li> <li><span class="nowrap"><a href="/wiki/Harlow_Curtice" title="Harlow Curtice">Harlow Curtice</a> (1955)</span></li> <li><span class="nowrap"><a href="/wiki/Hungarian_Revolution_of_1956" title="Hungarian Revolution of 1956">Hungarian Freedom Fighters</a> (1956)</span></li> <li><span class="nowrap"><a href="/wiki/Nikita_Khrushchev" title="Nikita Khrushchev">Nikita Khrushchev</a> (1957)</span></li> <li><span class="nowrap"><a href="/wiki/Charles_de_Gaulle" title="Charles de Gaulle">Charles de Gaulle</a> (1958)</span></li> <li><span class="nowrap"><a href="/wiki/Dwight_D._Eisenhower" title="Dwight D. Eisenhower">Dwight D. Eisenhower</a> (1959)</span></li> <li><span class="nowrap"><span class="wrap"> U.S. Scientists: <a class="mw-selflink selflink">George Beadle</a> / <a href="/wiki/Charles_Stark_Draper" title="Charles Stark Draper">Charles Draper</a> / <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Donald_A._Glaser" title="Donald A. Glaser">Donald A. Glaser</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a> / <a href="/wiki/Willard_Libby" title="Willard Libby">Willard Libby</a> / <a href="/wiki/Linus_Pauling" title="Linus Pauling">Linus Pauling</a> / <a href="/wiki/Edward_Mills_Purcell" title="Edward Mills Purcell">Edward Purcell</a> / <a href="/wiki/Isidor_Isaac_Rabi" title="Isidor Isaac Rabi">Isidor Rabi</a> / <a href="/wiki/Emilio_Segr%C3%A8" title="Emilio Segrè">Emilio Segrè</a> / <a href="/wiki/William_Shockley" title="William Shockley">William Shockley</a> / <a href="/wiki/Edward_Teller" title="Edward Teller">Edward Teller</a> / <a href="/wiki/Charles_H._Townes" title="Charles H. Townes">Charles Townes</a> / <a href="/wiki/James_Van_Allen" title="James Van Allen">James Van Allen</a> / <a href="/wiki/Robert_Burns_Woodward" title="Robert Burns Woodward">Robert Woodward</a> </span> (1960)</span></li> <li><span class="nowrap"><a href="/wiki/John_F._Kennedy" title="John F. Kennedy">John F. Kennedy</a> (1961)</span></li> <li><span class="nowrap"><a href="/wiki/Pope_John_XXIII" title="Pope John XXIII">Pope John XXIII</a> (1962)</span></li> <li><span class="nowrap"><a href="/wiki/Martin_Luther_King_Jr." title="Martin Luther King Jr.">Martin Luther King Jr.</a> (1963)</span></li> <li><span class="nowrap"><a href="/wiki/Lyndon_B._Johnson" title="Lyndon B. Johnson">Lyndon B. Johnson</a> (1964)</span></li> <li><span class="nowrap"><a href="/wiki/William_Westmoreland" title="William Westmoreland">William Westmoreland</a> (1965)</span></li> <li><span class="nowrap"><a href="/wiki/Baby_boomers" title="Baby boomers">The Generation Twenty-Five and Under</a> (1966)</span></li> <li><span class="nowrap"><a href="/wiki/Lyndon_B._Johnson" title="Lyndon B. Johnson">Lyndon B. Johnson</a> (1967)</span></li> <li><span class="nowrap"><span class="wrap"> <a href="/wiki/Apollo_8" title="Apollo 8">The Apollo 8 Astronauts</a>: <a href="/wiki/William_Anders" title="William Anders">William Anders</a> / <a href="/wiki/Frank_Borman" title="Frank Borman">Frank Borman</a> / <a href="/wiki/Jim_Lovell" title="Jim Lovell">Jim Lovell</a> </span> (1968)</span></li> <li><span class="nowrap"><a href="/wiki/Middle_America_(United_States)" title="Middle America (United States)">The Middle Americans</a> (1969)</span></li> <li><span class="nowrap"><a href="/wiki/Willy_Brandt" title="Willy Brandt">Willy Brandt</a> (1970)</span></li> <li><span class="nowrap"><a href="/wiki/Richard_Nixon" title="Richard Nixon">Richard Nixon</a> (1971)</span></li> <li><span class="nowrap"><a href="/wiki/Henry_Kissinger" title="Henry Kissinger">Henry Kissinger</a> / <a href="/wiki/Richard_Nixon" title="Richard Nixon">Richard Nixon</a> (1972)</span></li> <li><span class="nowrap"><a href="/wiki/John_Sirica" title="John Sirica">John Sirica</a> (1973)</span></li> <li><span class="nowrap"><a href="/wiki/Faisal_of_Saudi_Arabia" title="Faisal of Saudi Arabia">King Faisal</a> (1974)</span></li> <li><span class="nowrap"><span class="wrap"> American Women: <a href="/wiki/Susan_Brownmiller" title="Susan Brownmiller">Susan Brownmiller</a> / <a href="/wiki/Kathleen_Byerly" title="Kathleen Byerly">Kathleen Byerly</a> / <a href="/wiki/Alison_Cheek" title="Alison Cheek">Alison Cheek</a> / <a href="/wiki/Jill_Ker_Conway" title="Jill Ker Conway">Jill Conway</a> / <a href="/wiki/Betty_Ford" title="Betty Ford">Betty Ford</a> / <a href="/wiki/Ella_Grasso" title="Ella Grasso">Ella Grasso</a> / <a href="/wiki/Carla_Anderson_Hills" title="Carla Anderson Hills">Carla Hills</a> / <a href="/wiki/Barbara_Jordan" title="Barbara Jordan">Barbara Jordan</a> / <a href="/wiki/Billie_Jean_King" title="Billie Jean King">Billie Jean King</a> / <a href="/wiki/Susie_Sharp" title="Susie Sharp">Susie Sharp</a> / <a href="/wiki/Carol_Sutton_(journalist)" title="Carol Sutton (journalist)">Carol Sutton</a> / <a href="/wiki/Addie_L._Wyatt" title="Addie L. Wyatt">Addie Wyatt</a> </span> (1975)</span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap"><a href="/wiki/Jimmy_Carter" title="Jimmy Carter">Jimmy Carter</a> (1976)</span></li> <li><span class="nowrap"><a href="/wiki/Anwar_Sadat" title="Anwar Sadat">Anwar Sadat</a> (1977)</span></li> <li><span class="nowrap"><a href="/wiki/Deng_Xiaoping" title="Deng Xiaoping">Deng Xiaoping</a> (1978)</span></li> <li><span class="nowrap"><a href="/wiki/Ruhollah_Khomeini" title="Ruhollah Khomeini">Ayatollah Khomeini</a> (1979)</span></li> <li><span class="nowrap"><a href="/wiki/Ronald_Reagan" title="Ronald Reagan">Ronald Reagan</a> (1980)</span></li> <li><span class="nowrap"><a href="/wiki/Lech_Wa%C5%82%C4%99sa" title="Lech Wałęsa">Lech Wałęsa</a> (1981)</span></li> <li><span class="nowrap"><a href="/wiki/Personal_computer" title="Personal computer">The Computer</a> (1982)</span></li> <li><span class="nowrap"><a href="/wiki/Ronald_Reagan" title="Ronald Reagan">Ronald Reagan</a> / <a href="/wiki/Yuri_Andropov" title="Yuri Andropov">Yuri Andropov</a> (1983)</span></li> <li><span class="nowrap"><a href="/wiki/Peter_Ueberroth" title="Peter Ueberroth">Peter Ueberroth</a> (1984)</span></li> <li><span class="nowrap"><a href="/wiki/Deng_Xiaoping" title="Deng Xiaoping">Deng Xiaoping</a> (1985)</span></li> <li><span class="nowrap"><a href="/wiki/Corazon_Aquino" title="Corazon Aquino">Corazon Aquino</a> (1986)</span></li> <li><span class="nowrap"><a href="/wiki/Mikhail_Gorbachev" title="Mikhail Gorbachev">Mikhail Gorbachev</a> (1987)</span></li> <li><span class="nowrap"><a href="/wiki/Environmentalism" title="Environmentalism">The Endangered Earth</a> (1988)</span></li> <li><span class="nowrap"><a href="/wiki/Mikhail_Gorbachev" title="Mikhail Gorbachev">Mikhail Gorbachev</a> (1989)</span></li> <li><span class="nowrap"><a href="/wiki/George_H._W._Bush" title="George H. W. Bush">George H. W. Bush</a> (1990)</span></li> <li><span class="nowrap"><a href="/wiki/Ted_Turner" title="Ted Turner">Ted Turner</a> (1991)</span></li> <li><span class="nowrap"><a href="/wiki/Bill_Clinton" title="Bill Clinton">Bill Clinton</a> (1992)</span></li> <li><span class="nowrap"><span class="wrap"> The Peacemakers: <a href="/wiki/Yasser_Arafat" title="Yasser Arafat">Yasser Arafat</a> / <a href="/wiki/F._W._de_Klerk" title="F. W. de Klerk">F. W. de Klerk</a> / <a href="/wiki/Nelson_Mandela" title="Nelson Mandela">Nelson Mandela</a> / <a href="/wiki/Yitzhak_Rabin" title="Yitzhak Rabin">Yitzhak Rabin</a> </span> (1993)</span></li> <li><span class="nowrap"><a href="/wiki/Pope_John_Paul_II" title="Pope John Paul II">Pope John Paul II</a> (1994)</span></li> <li><span class="nowrap"><a href="/wiki/Newt_Gingrich" title="Newt Gingrich">Newt Gingrich</a> (1995)</span></li> <li><span class="nowrap"><a href="/wiki/David_Ho" title="David Ho">David Ho</a> (1996)</span></li> <li><span class="nowrap"><a href="/wiki/Andrew_Grove" title="Andrew Grove">Andrew Grove</a> (1997)</span></li> <li><span class="nowrap"><a href="/wiki/Bill_Clinton" title="Bill Clinton">Bill Clinton</a> / <a href="/wiki/Ken_Starr" title="Ken Starr">Ken Starr</a> (1998)</span></li> <li><span class="nowrap"><a href="/wiki/Jeff_Bezos" title="Jeff Bezos">Jeff Bezos</a> (1999)</span></li> <li><span class="nowrap"><a href="/wiki/George_W._Bush" title="George W. Bush">George W. Bush</a> (2000)</span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap"><a href="/wiki/Rudy_Giuliani" title="Rudy Giuliani">Rudolph Giuliani</a> (2001)</span></li> <li><span class="nowrap"><span class="wrap"> The Whistleblowers: <a href="/wiki/Cynthia_Cooper_(accountant)" title="Cynthia Cooper (accountant)">Cynthia Cooper</a> / <a href="/wiki/Coleen_Rowley" title="Coleen Rowley">Coleen Rowley</a> / <a href="/wiki/Sherron_Watkins" title="Sherron Watkins">Sherron Watkins</a> </span> (2002)</span></li> <li><span class="nowrap"><a href="/wiki/Iraq_War" title="Iraq War">The American Soldier</a> (2003)</span></li> <li><span class="nowrap"><a href="/wiki/George_W._Bush" title="George W. Bush">George W. Bush</a> (2004)</span></li> <li><span class="nowrap"><span class="wrap"> The Good Samaritans: <a href="/wiki/Bono" title="Bono">Bono</a> / <a href="/wiki/Bill_Gates" title="Bill Gates">Bill Gates</a> / <a href="/wiki/Melinda_French_Gates" title="Melinda French Gates">Melinda Gates</a> </span> (2005)</span></li> <li><span class="nowrap"><a href="/wiki/You_(Time_Person_of_the_Year)" title="You (Time Person of the Year)">You</a> (2006)</span></li> <li><span class="nowrap"><a href="/wiki/Vladimir_Putin" title="Vladimir Putin">Vladimir Putin</a> (2007)</span></li> <li><span class="nowrap"><a href="/wiki/Barack_Obama" title="Barack Obama">Barack Obama</a> (2008)</span></li> <li><span class="nowrap"><a href="/wiki/Ben_Bernanke" title="Ben Bernanke">Ben Bernanke</a> (2009)</span></li> <li><span class="nowrap"><a href="/wiki/Mark_Zuckerberg" title="Mark Zuckerberg">Mark Zuckerberg</a> (2010)</span></li> <li><span class="nowrap"><a href="/wiki/Protest" title="Protest">The Protester</a> (2011)</span></li> <li><span class="nowrap"><a href="/wiki/Barack_Obama" title="Barack Obama">Barack Obama</a> (2012)</span></li> <li><span class="nowrap"><a href="/wiki/Pope_Francis" title="Pope Francis">Pope Francis</a> (2013)</span></li> <li><span class="nowrap"><span class="wrap"> <a href="/wiki/Responses_to_the_West_African_Ebola_virus_epidemic" title="Responses to the West African Ebola virus epidemic">Ebola Fighters</a>: Dr. Jerry Brown / Dr. <a href="/wiki/Kent_Brantly" title="Kent Brantly">Kent Brantly</a> / Ella Watson-Stryker / Foday Gollah / <a href="/wiki/Salome_Karwah" title="Salome Karwah">Salome Karwah</a> </span> (2014)</span></li> <li><span class="nowrap"><a href="/wiki/Angela_Merkel" title="Angela Merkel">Angela Merkel</a> (2015)</span></li> <li><span class="nowrap"><a href="/wiki/Donald_Trump" title="Donald Trump">Donald Trump</a> (2016)</span></li> <li><span class="nowrap"><a href="/wiki/MeToo_movement" title="MeToo movement">The Silence Breakers</a> (2017)</span></li> <li><span class="nowrap"><span class="wrap"> The Guardians: <a href="/wiki/Jamal_Khashoggi" title="Jamal Khashoggi">Jamal Khashoggi</a> / <a href="/wiki/Maria_Ressa" title="Maria Ressa">Maria Ressa</a> / <a href="/wiki/Wa_Lone" title="Wa Lone">Wa Lone</a> / <a href="/wiki/Kyaw_Soe_Oo" title="Kyaw Soe Oo">Kyaw Soe Oo</a> / Staff of <i><a href="/wiki/Capital_Gazette_shooting" title="Capital Gazette shooting">The Capital</a></i> </span> (2018)</span></li> <li><span class="nowrap"><a href="/wiki/Greta_Thunberg" title="Greta Thunberg">Greta Thunberg</a> (2019)</span></li> <li><span class="nowrap"><a href="/wiki/Joe_Biden" title="Joe Biden">Joe Biden</a> / <a href="/wiki/Kamala_Harris" title="Kamala Harris">Kamala Harris</a> (2020)</span></li> <li><span class="nowrap"><a href="/wiki/Elon_Musk" title="Elon Musk">Elon Musk</a> (2021)</span></li> <li><span class="nowrap"><a href="/wiki/Volodymyr_Zelenskyy" title="Volodymyr Zelenskyy">Volodymyr Zelenskyy</a> / <a href="/wiki/Ukrainians" title="Ukrainians">Spirit of Ukraine</a> (2022)</span></li> <li><span class="nowrap"><a href="/wiki/Taylor_Swift" title="Taylor Swift">Taylor Swift</a> (2023)</span></li> <li><span class="nowrap"><a href="/wiki/Donald_Trump" title="Donald Trump">Donald Trump</a> (2024)</span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q238739#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata2322" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" 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srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">International</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/0000000114958322">ISNI</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/97657524">VIAF</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://id.worldcat.org/fast/1429/">FAST</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.oclc.org/worldcat/entity/E39PBJmxYWmtqVgtYTwXF34KBP">WorldCat</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/128581301">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n50005708">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb12383464p">France</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb12383464p">BnF data</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00432614">Japan</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=nlk20000080375&CON_LNG=ENG">Czech Republic</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p091017777">Netherlands</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://authority.bibsys.no/authority/rest/authorities/html/3116790">Norway</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://dbn.bn.org.pl/descriptor-details/9810536047305606">Poland</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.nli.org.il/en/authorities/987007258330505171">Israel</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://opac.kbr.be/LIBRARY/doc/AUTHORITY/14077519">Belgium</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Academics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://ci.nii.ac.jp/author/DA02487074?l=en">CiNii</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.uri?authorId=16672941600">Scopus</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">People</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.deutsche-biographie.de/128581301.html?language=en">Deutsche Biographie</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/032893906">IdRef</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalog.archives.gov/id/10581416">NARA</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://snaccooperative.org/ark:/99166/w6708fh4">SNAC</a></span></li></ul></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?useformat=desktop&type=1x1&usesul3=0" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=George_Beadle&oldid=1275055569">https://en.wikipedia.org/w/index.php?title=George_Beadle&oldid=1275055569</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div 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