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Available in 44 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-44" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">44 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D8%B3%D9%86%D9%8A%D9%84" title="جورج سنيل – Arabic" lang="ar" hreflang="ar" data-title="جورج سنيل" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D8%AF%DB%8C%D9%88%DB%8C%D8%B3_%D8%B3%D9%86%D9%84" title="جورج دیویس سنل – South Azerbaijani" lang="azb" hreflang="azb" data-title="جورج دیویس سنل" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6_%D0%94%D1%8D%D0%B9%D0%B2%D1%96%D1%81_%D0%A1%D0%BD%D0%B5%D0%BB" title="Джордж Дэйвіс Снел – Belarusian" lang="be" hreflang="be" data-title="Джордж Дэйвіс Снел" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Catalan" lang="ca" hreflang="ca" data-title="George Davis Snell" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Czech" lang="cs" hreflang="cs" data-title="George Davis Snell" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – German" lang="de" hreflang="de" data-title="George Davis Snell" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/George_Snell" title="George Snell – Estonian" lang="et" hreflang="et" data-title="George Snell" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/George_D._Snell" title="George D. Snell – Spanish" lang="es" hreflang="es" data-title="George D. Snell" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Basque" lang="eu" hreflang="eu" data-title="George Davis Snell" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AC%D8%B1%D8%AC_%D8%AF%DB%8C%D9%88%DB%8C%D8%B3_%D8%B3%D9%86%D9%84" title="جرج دیویس سنل – Persian" lang="fa" hreflang="fa" data-title="جرج دیویس سنل" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/George_Snell" title="George Snell – French" lang="fr" hreflang="fr" data-title="George Snell" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Irish" lang="ga" hreflang="ga" data-title="George Davis Snell" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%A1%B0%EC%A7%80_%EB%8D%B0%EC%9D%B4%EB%B9%84%EC%8A%A4_%EC%8A%A4%EB%84%AC" title="조지 데이비스 스넬 – Korean" lang="ko" hreflang="ko" data-title="조지 데이비스 스넬" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/George_D._Snell" title="George D. Snell – Croatian" lang="hr" hreflang="hr" data-title="George D. Snell" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Ido" lang="io" hreflang="io" data-title="George Davis Snell" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/George_Snell" title="George Snell – Indonesian" lang="id" hreflang="id" data-title="George Snell" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/George_Snell" title="George Snell – Italian" lang="it" hreflang="it" data-title="George Snell" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%92%27%D7%95%D7%A8%D7%92%27_%D7%93._%D7%A1%D7%A0%D7%9C" title="ג'ורג' ד. סנל – Hebrew" lang="he" hreflang="he" data-title="ג'ורג' ד. סנל" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6_%D0%A1%D0%BD%D0%B5%D0%BB%D0%BB" title="Джордж Снелл – Kazakh" lang="kk" hreflang="kk" data-title="Джордж Снелл" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/George_Snell" title="George Snell – Swahili" lang="sw" hreflang="sw" data-title="George Snell" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Georgius_Snell" title="Georgius Snell – Latin" lang="la" hreflang="la" data-title="Georgius Snell" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/George_D._Snell" title="George D. Snell – Hungarian" lang="hu" hreflang="hu" data-title="George D. Snell" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D8%B3%D9%86%D9%8A%D9%84" title="جورج سنيل – Egyptian Arabic" lang="arz" hreflang="arz" data-title="جورج سنيل" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%D8%B1%D8%AC_%D8%AF%DB%8C%D9%88%DB%8C%D8%B3_%D8%B3%D9%86%D9%84" title="جرج دیویس سنل – Mazanderani" lang="mzn" hreflang="mzn" data-title="جرج دیویس سنل" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Malay" lang="ms" hreflang="ms" data-title="George Davis Snell" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Dutch" lang="nl" hreflang="nl" data-title="George Davis Snell" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B8%E3%83%A7%E3%83%BC%E3%82%B8%E3%83%BB%E3%82%B9%E3%83%8D%E3%83%AB" title="ジョージ・スネル – Japanese" lang="ja" hreflang="ja" data-title="ジョージ・スネル" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="George Davis Snell" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Occitan" lang="oc" hreflang="oc" data-title="George Davis Snell" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Uzbek" lang="uz" hreflang="uz" data-title="George Davis Snell" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%D8%A7%D8%B1%D8%AC_%DA%88%DB%8C%D9%88%D8%B3_%D8%B3%D9%86%D9%84" title="جارج ڈیوس سنل – Western Punjabi" lang="pnb" hreflang="pnb" data-title="جارج ڈیوس سنل" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/George_Snell" title="George Snell – Polish" lang="pl" hreflang="pl" data-title="George Snell" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Portuguese" lang="pt" hreflang="pt" data-title="George Davis Snell" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A1%D0%BD%D0%B5%D0%BB%D0%BB,_%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6" title="Снелл, Джордж – Russian" lang="ru" hreflang="ru" data-title="Снелл, Джордж" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Simple English" lang="en-simple" hreflang="en-simple" data-title="George Davis Snell" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Slovak" lang="sk" hreflang="sk" data-title="George Davis Snell" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Finnish" lang="fi" hreflang="fi" data-title="George Davis Snell" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/George_D._Snell" title="George D. Snell – Swedish" lang="sv" hreflang="sv" data-title="George D. Snell" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Turkish" lang="tr" hreflang="tr" data-title="George Davis Snell" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%94%D0%B6%D0%BE%D1%80%D0%B4%D0%B6_%D0%A1%D0%BD%D0%B5%D0%BB%D0%BB" title="Джордж Снелл – Ukrainian" lang="uk" hreflang="uk" data-title="Джордж Снелл" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%AC%D8%A7%D8%B1%D8%AC_%DA%88%DB%8C%D9%88%DA%88_%D8%B3%D9%86%D9%84" title="جارج ڈیوڈ سنل – Urdu" lang="ur" hreflang="ur" data-title="جارج ڈیوڈ سنل" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Vietnamese" lang="vi" hreflang="vi" data-title="George Davis Snell" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/George_Davis_Snell" title="George Davis Snell – Yoruba" lang="yo" hreflang="yo" data-title="George Davis Snell" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E4%B9%94%E6%B2%BB%C2%B7%E6%96%AF%E5%86%85%E5%B0%94" title="乔治·斯内尔 – Chinese" lang="zh" hreflang="zh" data-title="乔治·斯内尔" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q295666#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected 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<div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">American geneticist</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">For other people named George Snell, see <a href="/wiki/George_Snell_(disambiguation)" class="mw-redirect mw-disambig" title="George Snell (disambiguation)">George Snell 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.infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">George Davis Snell</div></th></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data">December 19, 1903<br /><div style="display:inline" class="birthplace"><a href="/wiki/Bradford,_Massachusetts" title="Bradford, Massachusetts">Bradford, Massachusetts</a>, U.S</div></td></tr><tr><th scope="row" class="infobox-label">Died</th><td class="infobox-data">June 6, 1996<span style="display:none">(1996-06-06)</span> (aged 92)<br /><div style="display:inline" class="deathplace"><a href="/wiki/Bar_Harbor,_Maine" title="Bar Harbor, Maine">Bar Harbor, Maine</a>, U.S</div></td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/Dartmouth_College" title="Dartmouth College">Dartmouth College</a> <span style="font-size:85%;">(BS, 1926)</span><br /><a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a> <span style="font-size:85%;">(D.Sc, 1930)</span></td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data">Discovery of <a href="/wiki/Major_histocompatibility_complex" title="Major histocompatibility complex">H2</a></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data">1980 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Genetics" title="Genetics">genetics</a><br /> <a href="/wiki/Immunologist" class="mw-redirect" title="Immunologist">immunologist</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/Jackson_Laboratory" title="Jackson Laboratory">Jackson Laboratory</a><br /><a href="/wiki/Brown_University" title="Brown University">Brown University</a><br /> <a href="/wiki/Washington_University_School_of_Medicine" title="Washington University School of Medicine">Washington University School of Medicine</a></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Doctoral_advisor" title="Doctoral advisor">Doctoral advisor</a></th><td class="infobox-data"><a href="/wiki/William_E._Castle" title="William E. Castle">William E. Castle</a></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>George Davis Snell</b> <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">NAS</a> (December 19, 1903 – June 6, 1996) was an American mouse <a href="/wiki/Geneticist" title="Geneticist">geneticist</a> and basic <a href="/wiki/Organ_transplant" class="mw-redirect" title="Organ transplant">transplant</a> <a href="/wiki/Immunologist" class="mw-redirect" title="Immunologist">immunologist</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Work">Work</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Davis_Snell&action=edit&section=1" title="Edit section: Work"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>George Snell shared the 1980 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> with <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> and <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> for their discoveries concerning "genetically determined structures on the cell surface that regulate immunological reactions". Snell specifically "discovered the genetic factors that determine the possibilities of transplanting tissue from one individual to another. It was Snell who introduced the concept of H <a href="/wiki/Antigen" title="Antigen">antigens</a>."<a rel="nofollow" class="external autonumber" href="http://nobelprize.org/nobel_prizes/medicine/laureates/1980/press.html">[1]</a> Snell's work in mice led to the discovery of <a href="/wiki/Human_leukocyte_antigen" title="Human leukocyte antigen">HLA</a>, the major histocompatibility complex, in humans (and all vertebrates) that is analogous to the H-2 complex in mice. Recognition of these key genes was prerequisite to successful tissue and <a href="/wiki/Organ_transplant" class="mw-redirect" title="Organ transplant">organ transplantation</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Life">Life</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Davis_Snell&action=edit&section=2" title="Edit section: Life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>George Snell was born in <a href="/wiki/Bradford,_Massachusetts" title="Bradford, Massachusetts">Bradford, Massachusetts</a>, the youngest of three children. His father (who was born in <a href="/wiki/Minnesota" title="Minnesota">Minnesota</a>) worked as a secretary for the local <a href="/wiki/YMCA" title="YMCA">YMCA</a>; he invented a device for winding induction coils for motorboat engines. Snell was educated in the <a href="/wiki/Brookline,_Massachusetts" title="Brookline, Massachusetts">Brookline, Massachusetts</a> schools and then enrolled at <a href="/wiki/Dartmouth_College" title="Dartmouth College">Dartmouth College</a> in <a href="/wiki/Hanover,_New_Hampshire" title="Hanover, New Hampshire">Hanover, New Hampshire</a> where he continued his passion for mathematics and science, focusing on genetics. He received his <a href="/wiki/Bachelor%27s_degree" title="Bachelor's degree">bachelor's degree</a> from Dartmouth in 1926. </p><p>On the recommendation of John Gerould, his genetics professor at Dartmouth, Snell did graduate work at <a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a> with <a href="/wiki/William_E._Castle" title="William E. Castle">William E. Castle</a>, the first American biologist to look for <a href="/wiki/Mendelian_inheritance" title="Mendelian inheritance">Mendelian inheritance</a> in mammals. Snell earned his <a href="/wiki/PhD" class="mw-redirect" title="PhD">PhD</a> from Harvard in 1930. His doctoral thesis was on <a href="/wiki/Genetic_linkage" title="Genetic linkage">genetic linkage</a> in mice. </p><p>Upon receiving the PhD from <a href="/wiki/Harvard" class="mw-redirect" title="Harvard">Harvard</a>, George Snell was employed as a teacher at <a href="/wiki/Brown_University" title="Brown University">Brown University</a>, from 1930 to 1931. </p><p>Snell then spent two years as a postdoctoral fellow at the <a href="/wiki/University_of_Texas" class="mw-redirect" title="University of Texas">University of Texas</a> with <a href="/wiki/H.J._Muller" class="mw-redirect" title="H.J. Muller">H.J. Muller</a>, who pioneered radiation genetics (and was also to win a <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a>). Snell studied the genetic effects of x-rays on mice with Muller. </p><p>This experience "served to convince me that research was my real love," Snell wrote in his autobiography.<a rel="nofollow" class="external autonumber" href="http://nobelprize.org/nobel_prizes/medicine/laureates/1980/snell-autobio.html">[2]</a>"If it were to be research, mouse genetics was the clear choice and the <a href="/wiki/Jackson_Laboratory" title="Jackson Laboratory">Jackson Laboratory</a>, founded in 1929 by <a href="/wiki/C._C._Little" title="C. C. Little">Dr. Clarence Cook Little</a>, one of Castle's earlier students, almost the inevitable selection as a place to work." The Jackson Laboratory was (and still is) the world's mecca for mouse genetics. </p><p>From 1933 to 1934, Snell was a teacher at <a href="/wiki/Washington_University_in_St._Louis" title="Washington University in St. Louis">Washington University in St. Louis</a>. </p><p>After brief stints as teachers, in 1935 Snell joined the staff of The Jackson Laboratory in <a href="/wiki/Bar_Harbor,_Maine" title="Bar Harbor, Maine">Bar Harbor</a> on <a href="/wiki/Mount_Desert_Island" title="Mount Desert Island">Mount Desert Island</a> on the coast of <a href="/wiki/Maine" title="Maine">Maine</a> and he remained there for the entire balance of his long career. In Bar Harbor, he met and married Rhoda Carson. Together they had three sons, Thomas, Roy, and Peter. In his leisure time, Snell enjoyed skiing, a passion he developed during his years at Dartmouth, as well as tennis. </p><p>Snell received the Cancer Research Institute <a href="/wiki/William_B._Coley_Award" title="William B. Coley Award">William B. Coley Award</a> in 1978 for distinguished research in immunology. In 1988, he authored a substantial book, <i>Search for a Rational Ethic</i>, on the nature of ethics and the rules by which we live. It includes an <a href="/wiki/Evolution" title="Evolution">evolution</a>-based ethic founded on biological realities that he believed to be applicable to all human beings. </p><p>Snell died in Bar Harbor on June 6, 1996. His wife died in 1994. </p> <div class="mw-heading mw-heading2"><h2 id="Awards_and_honors">Awards and honors</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Davis_Snell&action=edit&section=3" title="Edit section: Awards and honors"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>1935-68 The Jackson Laboratory, staff scientist</li> <li>1952 Elected to American Academy of Arts and Sciences</li> <li>1955 Hekteon Medal of American Medical Association</li> <li>1962 Griffen Animal Care Panel Award</li> <li>1962 Bertner Foundation Award</li> <li>1967 Gregor Mendel Medal, Czechoslovak Academy of Sciences</li> <li>1968-96 The Jackson Laboratory, senior staff scientist emeritus</li> <li>1970 Elected to National Academy of Sciences</li> <li>1976 <a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a></li> <li>1978 National Cancer Institute Award</li> <li>1978 Elected to British Transplantation Society, honorary</li> <li>1978 <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a></li> <li>1979 Elected to French Academy of Sciences, foreign associate</li> <li>1980 Nobel Prize in Physiology or Medicine</li> <li>1981 Founding member of the <a href="/wiki/World_Cultural_Council" title="World Cultural Council">World Cultural Council</a><sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li> <li>1982 Elected to American Philosophical Society</li> <li>1982 Golden Plate Award of the <a href="/wiki/Academy_of_Achievement" title="Academy of Achievement">American Academy of Achievement</a><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li>1983 Elected to British Society of Immunology, honorary</li></ul> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Davis_Snell&action=edit&section=4" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/C.C._Little" class="mw-redirect" title="C.C. Little">C.C. Little</a></li> <li><a href="/wiki/Peter_Alfred_Gorer" title="Peter Alfred Gorer">Peter Alfred Gorer</a></li> <li><a href="/wiki/Leroy_Stevens" title="Leroy Stevens">Leroy Stevens</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Davis_Snell&action=edit&section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.consejoculturalmundial.org/about-us/">"About Us"</a>. <a href="/wiki/World_Cultural_Council" title="World Cultural Council">World Cultural Council</a><span class="reference-accessdate">. Retrieved <span class="nowrap">November 8,</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=About+Us&rft.pub=World+Cultural+Council&rft_id=http%3A%2F%2Fwww.consejoculturalmundial.org%2Fabout-us%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Davis+Snell" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://achievement.org/our-history/golden-plate-awards/#science-exploration">"Golden Plate Awardees of the American Academy of Achievement"</a>. <i>www.achievement.org</i>. <a href="/wiki/American_Academy_of_Achievement" class="mw-redirect" title="American Academy of Achievement">American Academy of Achievement</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.achievement.org&rft.atitle=Golden+Plate+Awardees+of+the+American+Academy+of+Achievement&rft_id=https%3A%2F%2Fachievement.org%2Four-history%2Fgolden-plate-awards%2F%23science-exploration&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGeorge+Davis+Snell" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=George_Davis_Snell&action=edit&section=6" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/421">George D. Snell</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q295666#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> including the Nobel Lecture <i>Studies in Histocompatibility</i></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist 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abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Wolf_Prize_in_Medicine" title="Template:Wolf Prize in Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Wolf_Prize_in_Medicine" title="Template talk:Wolf Prize in Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Wolf_Prize_in_Medicine" title="Special:EditPage/Template:Wolf Prize in Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Wolf_Prize_in_Medicine" style="font-size:114%;margin:0 4em">Laureates of the <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1970s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">George Snell</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/Jon_van_Rood" title="Jon van Rood">Jon J. van Rood</a> (1978)</li> <li><a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Oleh_Hornykiewicz" title="Oleh Hornykiewicz">Oleh Hornykiewicz</a> (1979)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1980s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> / <a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a> / <a href="/wiki/James_Learmonth_Gowans" title="James Learmonth Gowans">James L. Gowans</a> (1980)</li> <li><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a> / <a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a> (1981)</li> <li><a href="/wiki/Jean-Pierre_Changeux" title="Jean-Pierre Changeux">Jean-Pierre Changeux</a> / <a href="/wiki/Solomon_H._Snyder" title="Solomon H. Snyder">Solomon H. Snyder</a> / <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> (1982)</li> <li><a href="/wiki/Donald_F._Steiner" title="Donald F. Steiner">Donald F. Steiner</a> (1984/5)</li> <li><a href="/wiki/Osamu_Hayaishi" title="Osamu Hayaishi">Osamu Hayaishi</a> (1986)</li> <li><a href="/wiki/Pedro_Cuatrecasas" title="Pedro Cuatrecasas">Pedro Cuatrecasas</a> / <a href="/wiki/Meir_Wilchek" title="Meir Wilchek">Meir Wilchek</a> (1987)</li> <li><a href="/wiki/Henri_G._Hers" title="Henri G. Hers">Henri G. Hers</a> / <a href="/wiki/Elizabeth_F._Neufeld" title="Elizabeth F. Neufeld">Elizabeth F. Neufeld</a> (1988)</li> <li><a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> (1989)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1990s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Maclyn_McCarty" title="Maclyn McCarty">Maclyn McCarty</a> (1990)</li> <li><a href="/wiki/Seymour_Benzer" title="Seymour Benzer">Seymour Benzer</a> (1991)</li> <li><a href="/wiki/Judah_Folkman" title="Judah Folkman">Judah Folkman</a> (1992)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> / <a href="/wiki/Yasutomi_Nishizuka" title="Yasutomi Nishizuka">Yasutomi Nishizuka</a> (1994/5)</li> <li><a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a> (1995/6)</li> <li><a href="/wiki/Mary_F._Lyon" title="Mary F. Lyon">Mary F. Lyon</a> (1996/7)</li> <li><a href="/wiki/Michael_Sela" title="Michael Sela">Michael Sela</a> / <a href="/wiki/Ruth_Arnon" title="Ruth Arnon">Ruth Arnon</a> (1998)</li> <li><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a> (1999)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2000s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Avram_Hershko" title="Avram Hershko">Avram Hershko</a> / <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a> (2001)</li> <li><a href="/wiki/Ralph_L._Brinster" title="Ralph L. Brinster">Ralph L. Brinster</a> / <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> (2002/3)</li> <li><a href="/wiki/Robert_Weinberg_(biologist)" title="Robert Weinberg (biologist)">Robert Weinberg</a> / <a href="/wiki/Roger_Y._Tsien" title="Roger Y. Tsien">Roger Y. Tsien</a> (2004)</li> <li><a href="/wiki/Alexander_Levitzki" title="Alexander Levitzki">Alexander Levitzki</a> / <a href="/wiki/Anthony_R._Hunter" title="Anthony R. Hunter">Anthony R. Hunter</a> / <a href="/wiki/Tony_Pawson_(biochemist)" title="Tony Pawson (biochemist)">Tony Pawson</a> (2005)</li> <li><a href="/wiki/Howard_Cedar" title="Howard Cedar">Howard Cedar</a> / <a href="/wiki/Aharon_Razin" title="Aharon Razin">Aharon Razin</a> (2008)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2010s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Axel_Ullrich" title="Axel Ullrich">Axel Ullrich</a> (2010)</li> <li><a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> / <a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a> (2011)</li> <li><a href="/wiki/Ronald_M._Evans" title="Ronald M. Evans">Ronald M. Evans</a> (2012)</li> <li><a href="/wiki/Nahum_Sonenberg" title="Nahum Sonenberg">Nahum Sonenberg</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a> / <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> (2014)</li> <li><a href="/wiki/John_Kappler" title="John Kappler">John Kappler</a> / <a href="/wiki/Philippa_Marrack" title="Philippa Marrack">Philippa Marrack</a> / <a href="/wiki/Jeffrey_V._Ravetch" title="Jeffrey V. Ravetch">Jeffrey V. Ravetch</a> (2015)</li> <li><a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a> / <a href="/wiki/C._Ronald_Kahn" title="C. Ronald Kahn">C. Ronald Kahn</a> (2016)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> (2017)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a> (2019)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2020s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> / <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> (2020)</li> <li><a href="/wiki/Joan_Steitz" class="mw-redirect" title="Joan Steitz">Joan Steitz</a> / <a href="/wiki/Lynne_Maquat" class="mw-redirect" title="Lynne Maquat">Lynne Elizabeth Maquat</a> / <a href="/wiki/Adrian_R._Krainer" class="mw-redirect" title="Adrian R. Krainer">Adrian Krainer</a> (2021)</li> <li><a href="/wiki/Daniel_J._Drucker" title="Daniel J. Drucker">Daniel J. Drucker</a> (2023)</li> <li><a href="/wiki/Botond_Roska" title="Botond Roska">Botond Roska</a> / <a href="/wiki/Jos%C3%A9-Alain_Sahel" title="José-Alain Sahel">José-Alain Sahel</a> (2024)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1239334494">@media screen{html.skin-theme-clientpref-night .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"></div><div role="navigation" class="navbox" aria-labelledby="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Template:Nobel Prize in Physiology or Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Nobel_Prize_in_Physiology_or_Medicine" title="Template talk:Nobel Prize in Physiology or Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Special:EditPage/Template:Nobel Prize in Physiology or Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Physiology_or_Medicine" title="List of Nobel laureates in Physiology or Medicine">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_Macleod_(physiologist)" title="John Macleod (physiologist)">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1925</span></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Krebs_(biochemist)" title="Hans Krebs (biochemist)">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Beadle" title="George Beadle">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_MacLeod_Cormack" title="Allan MacLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a class="mw-selflink selflink">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards_(physiologist)" title="Robert Edwards (physiologist)">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>posthumously</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe_(neuroscientist)" title="John O'Keefe (neuroscientist)">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap"><a href="/wiki/2020_Nobel_Prize_in_Physiology_or_Medicine" title="2020 Nobel Prize in Physiology or Medicine">2020</a>: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton_(virologist)" class="mw-redirect" title="Michael Houghton (virologist)">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap"><a href="/wiki/2021_Nobel_Prize_in_Physiology_or_Medicine" title="2021 Nobel Prize in Physiology or Medicine">2021</a>: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap"><a href="/wiki/2022_Nobel_Prize_in_Physiology_or_Medicine" title="2022 Nobel Prize in Physiology or Medicine">2022</a>: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap"><a href="/wiki/2023_Nobel_Prize_in_Physiology_or_Medicine" title="2023 Nobel Prize in Physiology or Medicine">2023</a>: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="1980_Nobel_Prize_laureates" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:1980_Nobel_Prize_winners" title="Template:1980 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:1980_Nobel_Prize_winners" title="Template talk:1980 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:1980_Nobel_Prize_winners" title="Special:EditPage/Template:1980 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="1980_Nobel_Prize_laureates" style="font-size:114%;margin:0 4em">1980 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Paul_Berg" title="Paul Berg">Paul Berg</a> (United States)</li> <li><a href="/wiki/Walter_Gilbert" title="Walter Gilbert">Walter Gilbert</a> (United States)</li> <li><a href="/wiki/Frederick_Sanger" title="Frederick Sanger">Frederick Sanger</a> (Great Britain)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/1980_Nobel_Prize_in_Literature" title="1980 Nobel Prize in Literature">1980</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Czes%C5%82aw_Mi%C5%82osz" title="Czesław Miłosz">Czesław Miłosz</a> (Poland/United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Adolfo_P%C3%A9rez_Esquivel" title="Adolfo Pérez Esquivel">Adolfo Pérez Esquivel</a> (Argentina)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/James_Cronin" title="James Cronin">James Cronin</a> (United States)</li> <li><a href="/wiki/Val_Logsdon_Fitch" title="Val Logsdon Fitch">Val Logsdon Fitch</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> (United States)</li> <li><a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> (France)</li> <li><a class="mw-selflink selflink">George Davis Snell</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economic Sciences</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lawrence_Klein" title="Lawrence Klein">Lawrence Klein</a> (United States)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/Template:1975_Nobel_Prize_winners" title="Template:1975 Nobel Prize winners">1975</a></dd> <dd><a href="/wiki/Template:1976_Nobel_Prize_winners" title="Template:1976 Nobel Prize winners">1976</a></dd> <dd><a href="/wiki/Template:1977_Nobel_Prize_winners" title="Template:1977 Nobel Prize winners">1977</a></dd> <dd><a href="/wiki/Template:1978_Nobel_Prize_winners" title="Template:1978 Nobel Prize winners">1978</a></dd> <dd><a href="/wiki/Template:1979_Nobel_Prize_winners" title="Template:1979 Nobel Prize winners">1979</a></dd> <dd><b>1980 </b></dd> <dd><a href="/wiki/Template:1981_Nobel_Prize_winners" title="Template:1981 Nobel Prize winners">1981</a></dd> <dd><a href="/wiki/Template:1982_Nobel_Prize_winners" title="Template:1982 Nobel Prize winners">1982</a></dd> <dd><a href="/wiki/Template:1983_Nobel_Prize_winners" title="Template:1983 Nobel Prize winners">1983</a></dd> <dd><a href="/wiki/Template:1984_Nobel_Prize_winners" title="Template:1984 Nobel Prize winners">1984</a></dd> <dd><a href="/wiki/Template:1985_Nobel_Prize_winners" title="Template:1985 Nobel Prize winners">1985</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Founding_members_of_the_World_Cultural_Council" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Founding_members_of_the_World_Cultural_Council" title="Template:Founding members of the World Cultural Council"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Founding_members_of_the_World_Cultural_Council" title="Template talk:Founding members of the World Cultural Council"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Founding_members_of_the_World_Cultural_Council" title="Special:EditPage/Template:Founding members of the World Cultural Council"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Founding_members_of_the_World_Cultural_Council" style="font-size:114%;margin:0 4em">Founding members of the <a href="/wiki/World_Cultural_Council" title="World Cultural Council">World Cultural Council</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Christian_B._Anfinsen" title="Christian B. Anfinsen">Christian B. Anfinsen</a></li> <li><a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a></li> <li><a href="/wiki/James_Baddiley" title="James Baddiley">James Baddiley</a></li> <li>M. Balasegaram</li> <li><a href="/wiki/Frank_Barnaby" title="Frank Barnaby">Frank Barnaby</a></li> <li><a href="/wiki/Christiaan_Barnard" title="Christiaan Barnard">Christiaan Barnard</a></li> <li><a href="/wiki/Colin_Blakemore" title="Colin Blakemore">Colin Blakemore</a></li> <li><a href="/wiki/Aage_Bohr" title="Aage Bohr">Aage N. Bohr</a></li> <li><a href="/wiki/Norman_Borlaug" title="Norman Borlaug">Norman Borlaug</a></li> <li><a href="/wiki/H._G._Callan" title="H. G. Callan">Harold G. Callan</a></li> <li><a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Frédéric Cournand</a></li> <li>William J. Darby</li> <li><a href="/wiki/Eduardo_de_Robertis" title="Eduardo de Robertis">Eduardo de Robertis</a></li> <li><a href="/wiki/Cornelis_de_Jager" class="mw-redirect" title="Cornelis de Jager">Cornelis de Jager</a></li> <li>Guy Blaudin de Thé</li> <li>Jean-François Denisse</li> <li>Venancio Deulofeu</li> <li><a href="/wiki/Frank_J._Dixon" title="Frank J. Dixon">Frank J. Dixon</a></li> <li><a href="/wiki/Richard_Doll" title="Richard Doll">Richard S. Doll</a></li> <li><a href="/wiki/Audouin_Dollfus" title="Audouin Dollfus">Audouin Dollfus</a></li> <li><a href="/wiki/Jacques-%C3%89mile_Dubois" title="Jacques-Émile Dubois">Jacques-Émile Dubois</a></li> <li><a href="/wiki/Gerald_Durrell" title="Gerald Durrell">Gerald Durrell</a></li> <li>Francisco J. Dy</li> <li><a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John C. Eccles</a></li> <li><a href="/wiki/S._Paul_Ehrlich_Jr." title="S. Paul Ehrlich Jr.">Paul Ehrlich</a></li> <li><a href="/wiki/Manfred_Eigen" title="Manfred Eigen">Manfred Eigen</a></li> <li>Mohammed El Fasi</li> <li><a href="/wiki/Ernest_L._Eliel" title="Ernest L. Eliel">Ernest L. Eliel</a></li> <li>Kenneth O. Emergy</li> <li>José Rafael Estrada</li> <li><a href="/wiki/Hans_J._Eysenck" class="mw-redirect" title="Hans J. Eysenck">Hans J. Eysenck</a></li> <li><a href="/wiki/Don_W._Fawcett" title="Don W. Fawcett">Don W. Fawcett</a></li> <li><a href="/wiki/D._J._Finney" title="D. J. Finney">David J. Finney</a></li> <li><a href="/wiki/Val_Logsdon_Fitch" title="Val Logsdon Fitch">Val L. Fitch</a></li> <li>Carl G. Gahmberg</li> <li><a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred D. Hershey</a></li> <li><a href="/wiki/Gerhard_Herzberg" title="Gerhard Herzberg">Gerhard Herzberg</a></li> <li><a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a></li> <li>Osmo H. Järvi</li> <li><a href="/wiki/Reginald_Victor_Jones" title="Reginald Victor Jones">Reginald V. Jones</a></li> <li><a href="/wiki/Adrian_Kantrowitz" title="Adrian Kantrowitz">Adrian Kantrowitz</a></li> <li><a href="/wiki/Nathan_O._Kaplan" title="Nathan O. Kaplan">Nathan O. Kaplan</a></li> <li>Leo A. Kaprio</li> <li><a href="/wiki/Vassos_Karageorghis" title="Vassos Karageorghis">Vassos Karageorghis</a></li> <li>Peter E. Kent</li> <li><a href="/wiki/Donald_William_Kerst" title="Donald William Kerst">Donald W. Kerst</a></li> <li><a href="/wiki/Seymour_S._Kety" title="Seymour S. Kety">Seymour S. Kety</a></li> <li>Prem N. Kirpal</li> <li>Georges B. Koelle</li> <li>Walther Manshard</li> <li><a href="/wiki/Georges_Math%C3%A9" title="Georges Mathé">Georges Mathé</a></li> <li><a href="/wiki/William_D._McElroy" title="William D. McElroy">William D. McElroy</a></li> <li>Henry McIlwain</li> <li>John McMichael</li> <li><a href="/wiki/Jerrold_Meinwald" title="Jerrold Meinwald">Jerrold Meinwald</a></li> <li><a href="/wiki/Harry_Melville_(chemist)" title="Harry Melville (chemist)">Harry Melville</a></li> <li><a href="/wiki/Desmond_Morris" title="Desmond Morris">Desmond J. Morris</a></li> <li><a href="/wiki/Giuseppe_Moruzzi" title="Giuseppe Moruzzi">Giuseppe Moruzzi</a></li> <li><a href="/wiki/Nevill_Francis_Mott" title="Nevill Francis Mott">Nevill F. Mott</a></li> <li><a href="/wiki/Vernon_Benjamin_Mountcastle" title="Vernon Benjamin Mountcastle">Vernon B. Mountcastle</a></li> <li><a href="/wiki/Robert_S._Mulliken" title="Robert S. Mulliken">Robert S. Mulliken</a></li> <li><a href="/wiki/Walter_H._Munk" class="mw-redirect" title="Walter H. Munk">Walter H. Munk</a></li> <li><a href="/wiki/Ilie_G._Murgulescu" title="Ilie G. Murgulescu">Ilie G. Murgulescu</a></li> <li><a href="/wiki/Jayant_Narlikar" title="Jayant Narlikar">Jayant V. Narlikar</a></li> <li><a href="/wiki/Louis_N%C3%A9el" title="Louis Néel">Louis E. F. Néel</a></li> <li><a href="/wiki/Yuval_Ne%27eman" title="Yuval Ne'eman">Yuval Ne'eman</a></li> <li><a href="/wiki/Bernhard_Neumann" title="Bernhard Neumann">Bernhard H. Neumann</a></li> <li><a href="/wiki/William_Nierenberg" title="William Nierenberg">William A. Nierenberg</a></li> <li><a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall W. Nirenberg</a></li> <li><a href="/wiki/George_Emil_Palade" title="George Emil Palade">George E. Palade</a></li> <li><a href="/wiki/Arthur_Pardee" title="Arthur Pardee">Arthur B. Pardee</a></li> <li><a href="/wiki/David_Phillips_(chemist)" title="David Phillips (chemist)">David Phillips</a></li> <li><a href="/wiki/Jacques_Piccard" title="Jacques Piccard">Jacques Piccard</a></li> <li>Jens J. Pindborg</li> <li><a href="/wiki/Comlan_A._A._Quenum" title="Comlan A. A. Quenum">Comlan A. A. Quenum</a></li> <li><a href="/wiki/Hermann_Rahn" title="Hermann Rahn">Hermann Rahn</a></li> <li><a href="/wiki/G._N._Ramachandran" title="G. N. Ramachandran">G. N. Ramachandran</a></li> <li><a href="/wiki/Gunnar_Randers" title="Gunnar Randers">Gunnar Randers</a></li> <li><a href="/wiki/C._N._R._Rao" title="C. N. R. Rao">Chintamani N. R. Rao</a></li> <li><a href="/wiki/Rex_Richards_(chemist)" title="Rex Richards (chemist)">Rex Richards</a></li> <li>Jean Rösch</li> <li>Abraham J. A. Roux</li> <li><a href="/wiki/Keith_Runcorn" title="Keith Runcorn">Stanley K. Runcorn</a></li> <li><a href="/wiki/Donald_Sadler" title="Donald Sadler">Donald H. Sadler</a></li> <li><a href="/wiki/Hakim_Said" class="mw-redirect" title="Hakim Said">Hakim Muhammad Saeed</a></li> <li>Nobufusa Saito</li> <li><a href="/wiki/Abdus_Salam" title="Abdus Salam">Abdus Salam</a></li> <li>Stuart J. Saunders</li> <li><a href="/wiki/Menahem_Max_Schiffer" title="Menahem Max Schiffer">Menahem Max Schiffer</a></li> <li><a href="/wiki/William_G._Schneider" title="William G. Schneider">William G. Schneider</a></li> <li><a href="/wiki/Glenn_T._Seaborg" title="Glenn T. Seaborg">Glenn T. Seaborg</a></li> <li><a href="/wiki/Ernest_Robert_Sears" title="Ernest Robert Sears">Ernest R. Sears</a></li> <li><a href="/wiki/Frederick_Seitz" title="Frederick Seitz">Frederick Seitz</a></li> <li>Leonard T. Skeggs</li> <li><a href="/wiki/Stefan_%C5%9Alopek" title="Stefan Ślopek">Stefan Ślopek</a></li> <li>George J. Smets</li> <li><a class="mw-selflink selflink">George D. Snell</a></li> <li>Leonard Sosnowky</li> <li><a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger W. Sperry</a></li> <li><a href="/wiki/Lyman_Spitzer" title="Lyman Spitzer">Lyman Spitzer</a></li> <li><a href="/wiki/Frederick_Stewart_(geologist)" title="Frederick Stewart (geologist)">Frederick Stewart</a></li> <li>Heikki Suomalainen</li> <li><a href="/wiki/Pol_Swings" title="Pol Swings">Pol Swings</a></li> <li><a href="/wiki/Charles_Tanford" title="Charles Tanford">Charles Tanford</a></li> <li><a href="/wiki/Henry_Taube" title="Henry Taube">Henry Taube</a></li> <li><a href="/wiki/John_Tedder,_2nd_Baron_Tedder" title="John Tedder, 2nd Baron Tedder">John M. Tedder</a></li> <li><a href="/wiki/Edward_Teller" title="Edward Teller">Edward Teller</a></li> <li><a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></li> <li><a href="/wiki/Harold_Warris_Thompson" class="mw-redirect" title="Harold Warris Thompson">Harold Thompson</a></li> <li>Peter C. Thonemann</li> <li><a href="/wiki/Phillip_V._Tobias" class="mw-redirect" title="Phillip V. Tobias">Phillip V. Tobias</a></li> <li><a href="/wiki/Alexander_R._Todd" class="mw-redirect" title="Alexander R. Todd">Alexander R. Todd</a></li> <li><a href="/wiki/Jan_Peter_Toennies" title="Jan Peter Toennies">Jan Peter Toennies</a></li> <li><a href="/wiki/Andrzej_Trautman" title="Andrzej Trautman">Andrzej Trautman</a></li> <li><a href="/wiki/Jean_Tricart" title="Jean Tricart">Jean L. F. Tricart</a></li> <li>Ioan Ursu</li> <li>Constantin Vago</li> <li><a href="/wiki/Eugene_van_Tamelen" title="Eugene van Tamelen">Eugene van Tamelen</a></li> <li><a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf S. von Euler</a></li> <li><a href="/wiki/Alan_Walsh_(physicist)" title="Alan Walsh (physicist)">Alan Walsh</a></li> <li><a href="/wiki/William_Joseph_Whelan" title="William Joseph Whelan">William J. Whelan</a></li> <li><a href="/wiki/Karel_Wiesner" title="Karel Wiesner">Karel F. Wiesner</a></li> <li><a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn S. 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