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Ophioglossidae - Wikipedia
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class="mw-redirectedfrom">(Redirected from <a href="/w/index.php?title=Psilotopsida&redirect=no" class="mw-redirect" title="Psilotopsida">Psilotopsida</a>)</span></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Subclass of ferns</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)">Ophioglossidae </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Botrychium-4.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Botrychium-4.jpg/220px-Botrychium-4.jpg" decoding="async" width="220" height="307" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Botrychium-4.jpg/330px-Botrychium-4.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Botrychium-4.jpg/440px-Botrychium-4.jpg 2x" data-file-width="576" data-file-height="803" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i><a href="/wiki/Botrychium_lunaria" title="Botrychium lunaria">Botrychium lunaria</a></i> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(180,250,180)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Ophioglossidae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Plant" title="Plant">Plantae</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Vascular_plant" title="Vascular plant">Tracheophytes</a> </td></tr> <tr> <td>Division: </td> <td><a href="/wiki/Fern" title="Fern">Polypodiophyta</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Fern" title="Fern">Polypodiopsida</a> </td></tr> <tr> <td>Subclass: </td> <td><a class="mw-selflink selflink">Ophioglossidae</a><br /><small>Klinge 1882<sup id="cite_ref-FOOTNOTEPteridophyte_Phylogeny_Group2016_1-0" class="reference"><a href="#cite_note-FOOTNOTEPteridophyte_Phylogeny_Group2016-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)"><a href="/wiki/Type_genus" title="Type genus">Type genus</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><i><a href="/wiki/Ophioglossum" title="Ophioglossum">Ophioglossum</a></i><br /><div style="font-size: 85%;"><a href="/wiki/Carl_Linnaeus" title="Carl Linnaeus">L.</a></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)">Orders </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><a href="/wiki/Ophioglossales" class="mw-redirect" title="Ophioglossales">Ophioglossales</a></li> <li><a href="/wiki/Psilotales" class="mw-redirect" title="Psilotales">Psilotales</a></li></ul> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li>Psilotopsida</li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Ophioglossidae</b> is one of the four <a href="/wiki/Class_(biology)" title="Class (biology)">subclasses</a> of <a href="/wiki/Fern" title="Fern">Polypodiopsida</a> (ferns). This subclass consists of the ferns commonly known as whisk ferns, grape ferns, adder's-tongues and moonworts. It is equivalent to the class <b>Psilotopsida</b> in previous treatments, including Smith <i>et al.</i> (2006).<sup id="cite_ref-FOOTNOTESmith_et_al._2006_2-0" class="reference"><a href="#cite_note-FOOTNOTESmith_et_al._2006-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The subclass contains two orders, <a href="/wiki/Psilotales" class="mw-redirect" title="Psilotales">Psilotales</a> and <a href="/wiki/Ophioglossales" class="mw-redirect" title="Ophioglossales">Ophioglossales</a>, whose relationship was only confirmed by <a href="/wiki/Molecular_phylogenetics" title="Molecular phylogenetics">molecular phylogenetic</a> studies. </p> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Ophioglossidae&action=edit&section=1" title="Edit section: Taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; font-style: italic;"><a href="https://commons.wikimedia.org/wiki/Category:Psilotopsida" class="extiw" title="commons:Category:Psilotopsida">Psilotopsida</a></span>.</div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1237033735"><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/34px-Wikispecies-logo.svg.png" decoding="async" width="34" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/51px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/68px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></span></span></div> <div class="side-box-text plainlist"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a> has information related to <i><b><a href="https://species.wikimedia.org/wiki/Psilotopsida" class="extiw" title="wikispecies:Psilotopsida">Psilotopsida</a></b></i>.</div></div> </div> <p>Smith et al. (2006) carried out the first higher-level <a href="/wiki/Pteridophyte" title="Pteridophyte">pteridophyte</a> classification published in the <a href="/wiki/Molecular_phylogenetic" class="mw-redirect" title="Molecular phylogenetic">molecular phylogenetic</a> era, and considered the ferns (monilophytes), with four <a href="/wiki/Class_(biology)" title="Class (biology)">classes</a>. They placed the whisk ferns and related taxa in the class Psilotopsida, with two <a href="/wiki/Order_(biology)" title="Order (biology)">orders</a>.<sup id="cite_ref-FOOTNOTESmith_et_al._2006_2-1" class="reference"><a href="#cite_note-FOOTNOTESmith_et_al._2006-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> <a href="/wiki/Mark_W._Chase" class="mw-redirect" title="Mark W. Chase">Mark W. Chase</a> and <a href="/wiki/James_L._Reveal" title="James L. Reveal">James L. Reveal</a> (2009) classified them as two separate subclasses, Psilotidae and Ophioglossidae, corresponding to those orders within a much broader grouping, the class <a href="/wiki/Equisetopsida_sensu_lato" class="mw-redirect" title="Equisetopsida sensu lato">Equisetopsida</a> <i>sensu lato</i>.<sup id="cite_ref-FOOTNOTEChaseReveal2009_3-0" class="reference"><a href="#cite_note-FOOTNOTEChaseReveal2009-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Christenhusz <i>et al.</i>, 2011, included both the <a href="/wiki/Ophioglossales" class="mw-redirect" title="Ophioglossales">Ophioglossales</a> and <a href="/wiki/Psilotales" class="mw-redirect" title="Psilotales">Psilotales</a> orders in the Ophioglossidae subclass.<sup id="cite_ref-FOOTNOTEChristenhusz_et_al.2011_4-0" class="reference"><a href="#cite_note-FOOTNOTEChristenhusz_et_al.2011-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> This was continued by both <a href="/wiki/Christenhusz" class="mw-redirect" title="Christenhusz">Christenhusz</a> and <a href="/wiki/Mark_W._Chase" class="mw-redirect" title="Mark W. Chase">Chase</a> (2014)<sup id="cite_ref-FOOTNOTEChristenhuszChase2014_5-0" class="reference"><a href="#cite_note-FOOTNOTEChristenhuszChase2014-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> and by the Pteridophyte Phylogeny Group (2016). Under the latter the subclass is one of four subclasses of <a href="/wiki/Polypodiopsida" class="mw-redirect" title="Polypodiopsida">Polypodiopsida</a> (ferns) and contains two orders, two families, 12 genera, and an estimated 129 species.<sup id="cite_ref-FOOTNOTEPteridophyte_Phylogeny_Group2016_1-1" class="reference"><a href="#cite_note-FOOTNOTEPteridophyte_Phylogeny_Group2016-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> The relationships between the two orders, Psilotales and Ophioglossales, has long been unclear and was only confirmed by molecular systematic studies. Psilotales have <a href="/wiki/Rhizome" title="Rhizome">rhizomes</a> instead of real roots, and the roots of Ophioglossales lack both branching and <a href="/wiki/Root_hair" title="Root hair">root hairs</a>. The <a href="/wiki/Gametophyte" title="Gametophyte">gametophytes</a> of both orders are <a href="/wiki/Heterotrophic" class="mw-redirect" title="Heterotrophic">heterotrophic</a> and often subterranean, obtaining nutrients from <a href="/wiki/Mycorrhiza" title="Mycorrhiza">mycorrhiza</a> instead of light. <a href="/wiki/Photosynthesis" title="Photosynthesis">Photosynthesis</a> happens exclusively in the <a href="/wiki/Sporophyte" title="Sporophyte">sporophyte</a>.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>The following cladogram shows a likely phylogenic relationship between subclass Ophioglossidae and the other Polypodiopsida subclasses. The first three small subclasses are sometimes informally referred to as <a href="/wiki/Eusporangiate_fern" title="Eusporangiate fern">eusporangiate ferns</a>, in contrast to the largest subclass, Polypodiidae or <a href="/wiki/Leptosporangiate_fern" title="Leptosporangiate fern">leptosporangiate ferns</a>.<sup id="cite_ref-FOOTNOTEPteridophyte_Phylogeny_Group2016_1-2" class="reference"><a href="#cite_note-FOOTNOTEPteridophyte_Phylogeny_Group2016-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Polypodiopsida" class="mw-redirect" title="Polypodiopsida">Polypodiopsida</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Equisetidae" title="Equisetidae">Equisetidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><b>Ophioglossidae</b> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Marattiidae" class="mw-redirect" title="Marattiidae">Marattiidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Polypodiidae_(plant)" class="mw-redirect" title="Polypodiidae (plant)">Polypodiidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <p>The two orders, <a href="/wiki/Ophioglossales" class="mw-redirect" title="Ophioglossales">Ophioglossales</a> and <a href="/wiki/Psilotales" class="mw-redirect" title="Psilotales">Psilotales</a> are <a href="/wiki/Sister_groups" class="mw-redirect" title="Sister groups">sister groups</a> to each other. </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Ophioglossidae&action=edit&section=2" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 20em;"> <ol class="references"> <li id="cite_note-FOOTNOTEPteridophyte_Phylogeny_Group2016-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-FOOTNOTEPteridophyte_Phylogeny_Group2016_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-FOOTNOTEPteridophyte_Phylogeny_Group2016_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-FOOTNOTEPteridophyte_Phylogeny_Group2016_1-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><a href="#CITEREFPteridophyte_Phylogeny_Group2016">Pteridophyte Phylogeny Group 2016</a>.</span> </li> <li id="cite_note-FOOTNOTESmith_et_al._2006-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-FOOTNOTESmith_et_al._2006_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-FOOTNOTESmith_et_al._2006_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><a href="#CITEREFSmith_et_al._2006">Smith et al. 2006</a>.</span> </li> <li id="cite_note-FOOTNOTEChaseReveal2009-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEChaseReveal2009_3-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFChaseReveal2009">Chase & Reveal 2009</a>.</span> </li> <li id="cite_note-FOOTNOTEChristenhusz_et_al.2011-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEChristenhusz_et_al.2011_4-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFChristenhusz_et_al.2011">Christenhusz et al. 2011</a>.</span> </li> <li id="cite_note-FOOTNOTEChristenhuszChase2014-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEChristenhuszChase2014_5-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFChristenhuszChase2014">Christenhusz & Chase 2014</a>.</span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://books.google.com/books?id=dj8KRImgyf4C&pg=PA94">Plant Systematics</a></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Bibliography">Bibliography</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Ophioglossidae&action=edit&section=3" title="Edit section: Bibliography"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239549316">.mw-parser-output .refbegin{margin-bottom:0.5em}.mw-parser-output .refbegin-hanging-indents>ul{margin-left:0}.mw-parser-output .refbegin-hanging-indents>ul>li{margin-left:0;padding-left:3.2em;text-indent:-3.2em}.mw-parser-output .refbegin-hanging-indents ul,.mw-parser-output .refbegin-hanging-indents ul li{list-style:none}@media(max-width:720px){.mw-parser-output .refbegin-hanging-indents>ul>li{padding-left:1.6em;text-indent:-1.6em}}.mw-parser-output .refbegin-columns{margin-top:0.3em}.mw-parser-output .refbegin-columns ul{margin-top:0}.mw-parser-output .refbegin-columns li{page-break-inside:avoid;break-inside:avoid-column}@media screen{.mw-parser-output .refbegin{font-size:90%}}</style><div class="refbegin refbegin-columns references-column-width" style="column-width: 30em"> <ul><li><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFChaseReveal2009" class="citation journal cs1"><a href="/wiki/Mark_W._Chase" class="mw-redirect" title="Mark W. Chase">Chase, Mark W.</a>; <a href="/wiki/James_L._Reveal" title="James L. 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.navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Plant_classification" title="Template:Plant classification"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Plant_classification" title="Template talk:Plant classification"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Plant_classification" title="Special:EditPage/Template:Plant classification"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Classification_of_Archaeplastida_or_Plantae_s.l." style="font-size:114%;margin:0 4em">Classification of <a href="/wiki/Archaeplastida" title="Archaeplastida">Archaeplastida</a> or <a href="/wiki/Plant" title="Plant">Plantae <span class="rt-commentedText tooltip tooltip-dotted" title="sensu lato; in the broad sense"><i>s.l.</i></span></a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: #C3EEC3;"><div><div class="hlist"> <dl><dt><span class="nobold"><a href="/wiki/Domain_(biology)" title="Domain (biology)">Domain</a></span></dt> <dd><a href="/wiki/Template:Archaea_classification" title="Template:Archaea classification">Archaea</a></dd> <dd><a href="/wiki/Template:Bacteria_classification" title="Template:Bacteria classification">Bacteria</a></dd> <dd><a href="/wiki/Template:Eukaryota" title="Template:Eukaryota">Eukaryota</a></dd> <dt><span class="nobold">(major groups</span></dt> <dd><a href="/wiki/Template:Excavata" title="Template:Excavata">Excavata</a></dd> <dd><a href="/wiki/Diaphoretickes" title="Diaphoretickes">Diaphoretickes</a> <dl><dd><a href="/wiki/Template:Hacrobia" title="Template:Hacrobia">Hacrobia</a> <dl><dd><a href="/wiki/Template:Cryptista" title="Template:Cryptista">Cryptista</a></dd></dl></dd> <dd><a href="/wiki/Template:Rhizaria" title="Template:Rhizaria">Rhizaria</a></dd> <dd><a href="/wiki/Template:Alveolata" title="Template:Alveolata">Alveolata</a></dd> <dd><a href="/wiki/Template:Stramenopile" title="Template:Stramenopile">Stramenopiles</a></dd> <dd><a href="/wiki/Template:Plant_classification" title="Template:Plant classification">Plants</a></dd></dl></dd> <dd><a href="/wiki/Amorphea" title="Amorphea">Amorphea</a> <dl><dd><a href="/wiki/Template:Amoebozoa" title="Template:Amoebozoa">Amoebozoa</a></dd> <dd><a href="/wiki/Template:Opisthokont_protists" title="Template:Opisthokont protists">Opisthokonta</a> <dl><dd><a href="/wiki/Template:Animalia" title="Template:Animalia">Animals</a></dd> <dd><a href="/wiki/Template:Fungi_classification" title="Template:Fungi classification">Fungi</a></dd> <dd><a href="/wiki/Template:Mesomycetozoea" title="Template:Mesomycetozoea">Mesomycetozoea</a>)</dd></dl></dd></dl></dd></dl> </div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Archaeplastida" scope="row" class="navbox-group" style="width:1%;line-height:1.0em; background: #D4EED4;"><span style="margin:-40px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg);transform: rotate(270deg);"><a href="/wiki/Archaeplastida" title="Archaeplastida">Archaeplastida</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Picozoa" title="Picozoa">Picozoa</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Picomonas" class="mw-redirect" title="Picomonas">Picomonas</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Rhodelphis" title="Rhodelphis">Rhodelphidia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Rhodelphis" title="Rhodelphis">Rhodelphis</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Red_algae" title="Red algae">Rhodophyta</a><br /><span class="nobold">(red algae)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cyanidiophyceae" title="Cyanidiophyceae">Cyanidiophyceae</a></li> <li><a href="/wiki/Porphyridiophyceae" title="Porphyridiophyceae">Porphyridiophyceae</a></li> <li><a href="/wiki/Stylonematophyceae" title="Stylonematophyceae">Stylonematophyceae</a></li> <li><a href="/wiki/Compsopogonophyceae" title="Compsopogonophyceae">Compsopogonophyceae</a></li> <li><a href="/wiki/Rhodellophyceae" title="Rhodellophyceae">Rhodellophyceae</a></li> <li><a href="/wiki/Bangiophyceae" title="Bangiophyceae">Bangiophyceae</a></li> <li><a href="/wiki/Florideophyceae" title="Florideophyceae">Florideophyceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Glaucophyte" title="Glaucophyte">Glaucophyta</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glaucophyte" title="Glaucophyte">Glaucocystophyceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><i>incertae sedis</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Algospongia" title="Algospongia">Algospongia</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Viridiplantae" title="Viridiplantae">Viridiplantae</a> <br /> <span class="nobold">or <a href="/wiki/Plant" title="Plant">Plantae <span class="rt-commentedText tooltip tooltip-dotted" title="sensu stricto; in the strict sense"><i>s.s.</i></span></a><br />(<a href="/wiki/Green_algae" title="Green algae">green algae</a> &<br /> land plants)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Prasinodermophyta" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Prasinodermophyta" title="Prasinodermophyta">Prasinodermophyta</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Prasinoderma" title="Prasinoderma">Prasinodermophyceae</a></li> <li><a href="/wiki/Palmophyllophyceae" title="Palmophyllophyceae">Palmophyllophyceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"> <a href="/wiki/Chlorophyta" title="Chlorophyta">Chlorophyta</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Prasinophytina</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mamiellophyceae" title="Mamiellophyceae">Mamiellophyceae</a></li> <li><a href="/wiki/Pyramimonadophyceae" title="Pyramimonadophyceae">Pyramimonadophyceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Chlorophytina</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nephroselmidaceae" title="Nephroselmidaceae">Nephroselmidophyceae</a></li> <li><a href="/wiki/Picocystis" title="Picocystis">Picocystophyceae</a></li> <li><a href="/wiki/Chloropicophyceae" class="mw-redirect" title="Chloropicophyceae">Chloropicophyceae</a></li> <li><a href="/wiki/Pedinophyceae" title="Pedinophyceae">Pedinophyceae</a></li> <li><a href="/wiki/Chlorodendrales" title="Chlorodendrales">Chlorodendrophyceae</a></li> <li><a href="/wiki/UTC_clade" title="UTC clade">UTC clade</a> <ul><li><a href="/wiki/Ulvophyceae" title="Ulvophyceae">Ulvophyceae</a></li> <li><a href="/wiki/Trebouxiophyceae" title="Trebouxiophyceae">Trebouxiophyceae</a></li> <li><a href="/wiki/Chlorophyceae" title="Chlorophyceae">Chlorophyceae</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin:-29px;display: inline-block;-webkit-transform: rotate(270deg);-moz-transform: rotate(270deg);-ms-transform: rotate(270deg);-o-transform: rotate(270deg);transform: rotate(270deg);"><a href="/wiki/Streptophyta" title="Streptophyta">Streptophyta</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mesostigmatophyceae" title="Mesostigmatophyceae">Mesostigmatophyceae</a></li> <li><a href="/wiki/Chlorokybophyceae" class="mw-redirect" title="Chlorokybophyceae">Chlorokybophyceae</a></li> <li><i><a href="/wiki/Spirotaenia" title="Spirotaenia">Spirotaenia</a></i></li> <li><a href="/wiki/Klebsormidiophyceae" class="mw-redirect" title="Klebsormidiophyceae">Klebsormidiophyceae</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Phragmoplastophyta" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Phragmoplastophyta" title="Phragmoplastophyta">Phragmoplastophyta</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Charophyceae" title="Charophyceae">Charophyceae</a></li> <li><a href="/wiki/Coleochaetophyceae" title="Coleochaetophyceae">Coleochaetophyceae</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Anydrophyta</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Zygnematophyceae" title="Zygnematophyceae">Zygnematophyceae</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Embryophyta" class="mw-redirect" title="Embryophyta">Embryophyta</a><br /><span class="nobold">(land plants)</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>(see below↓)</li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background: #C3EEC3;"><div id="Embryophytes_or_Plantae_sensu_strictissimo" style="font-size:114%;margin:0 4em"><a href="/wiki/Embryophyte" title="Embryophyte">Embryophytes</a> or <a href="/wiki/Plant" title="Plant">Plantae <span class="rt-commentedText tooltip tooltip-dotted" title="in the strictest sense"><i>sensu strictissimo</i></span></a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Bryophytes" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -25px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Bryophyte" title="Bryophyte">Bryophytes</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Marchantiophyta" title="Marchantiophyta">Marchantiophyta</a><br /><span class="nobold">(liverworts)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Haplomitriopsida" title="Haplomitriopsida">Haplomitriopsida</a></li> <li><a href="/wiki/Marchantiopsida" title="Marchantiopsida">Marchantiopsida</a></li> <li><a href="/wiki/Jungermanniopsida" title="Jungermanniopsida">Jungermanniopsida</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Hornwort" title="Hornwort">Anthocerotophyta</a><br /><span class="nobold">(hornworts)</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Leiosporoceros" title="Leiosporoceros">Leiosporocerotopsida</a></li> <li><a href="/wiki/Hornwort" title="Hornwort">Anthocerotopsida</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Moss" title="Moss">Bryophyta</a><br /><span class="nobold">(mosses)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Takakia" title="Takakia">Takakiopsida</a></li> <li><a href="/wiki/Sphagnopsida" title="Sphagnopsida">Sphagnopsida</a></li> <li><a href="/wiki/Andreaeobryum" title="Andreaeobryum">Andreaeobryopsida</a></li> <li><a href="/wiki/Andreaeaceae" title="Andreaeaceae">Andreaeopsida</a></li> <li><a href="/wiki/Oedipodium" title="Oedipodium">Oedipodiopsida</a></li> <li><a href="/wiki/Tetraphidaceae" title="Tetraphidaceae">Tetraphidopsida</a></li> <li><a href="/wiki/Polytrichaceae" title="Polytrichaceae">Polytrichopsida</a></li> <li><a href="/wiki/Bryopsida" title="Bryopsida">Bryopsida</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -55px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"> <a href="/wiki/Polysporangiophyte" title="Polysporangiophyte">Polysporangiophytes</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Protracheophytes*" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span>Protracheophytes*</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Horneophytopsida" title="Horneophytopsida">Horneophytopsida</a></li> <li>†<i><a href="/wiki/Aglaophyton" title="Aglaophyton">Aglaophyton</a></i></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -25px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Vascular_plant" title="Vascular plant">Tracheophytes</a><br /><span class="nobold">(vascular plants)</span></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Paratracheophytes*" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span>Paratracheophytes*</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/w/index.php?title=Cooksoniopsida&action=edit&redlink=1" class="new" title="Cooksoniopsida (page does not exist)">Cooksoniopsida</a></li> <li>†<a href="/wiki/Rhyniophyte" title="Rhyniophyte">Rhyniopsida</a></li> <li>†<a href="/w/index.php?title=Renaliales&action=edit&redlink=1" class="new" title="Renaliales (page does not exist)">Renaliales</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -40px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);">Eutracheophytes</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Lycophytes" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Lycophyte" title="Lycophyte">Lycophytes</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Barinophyte" title="Barinophyte">Barinophytopsida</a></li> <li>†<a href="/wiki/Zosterophyll" title="Zosterophyll">Zosterophyllopsida</a></li> <li><a href="/wiki/Lycopodiopsida" title="Lycopodiopsida">Lycopodiopsida</a> (<a href="/wiki/Lycopodiaceae" title="Lycopodiaceae">clubmosses</a>, <a href="/wiki/Selaginella" title="Selaginella">spikemosses</a> & <a href="/wiki/Isoetes" title="Isoetes">quillworts</a>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -35px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Euphyllophyte" title="Euphyllophyte">Euphyllophytes</a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Eophyllophyton" title="Eophyllophyton">Eophyllophytopsida</a></li> <li>†<a href="/wiki/Trimerophytopsida" title="Trimerophytopsida">Trimerophytopsida</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Moniliformopses" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;">Moniliformopses</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/w/index.php?title=Ibykales&action=edit&redlink=1" class="new" title="Ibykales (page does not exist)">Ibykales</a></li> <li>†<a href="/wiki/Cladoxylopsida" title="Cladoxylopsida">Cladoxylopsida</a></li> <li><a href="/wiki/Fern" title="Fern">Polypodiopsida</a> (ferns and <a href="/wiki/Equisetum" title="Equisetum">horsetails</a>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -30px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);">Lignophytes</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Progymnosperms*" scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span><a href="/wiki/Progymnosperm" title="Progymnosperm">Progymnosperms</a>*</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/w/index.php?title=Noeggerathiopsida&action=edit&redlink=1" class="new" title="Noeggerathiopsida (page does not exist)">Noeggerathiopsida</a></li> <li>†<a href="/w/index.php?title=Aneurophytopsida&action=edit&redlink=1" class="new" title="Aneurophytopsida (page does not exist)">Aneurophytopsida</a></li> <li>†<a href="/w/index.php?title=Archaeopteridopsida&action=edit&redlink=1" class="new" title="Archaeopteridopsida (page does not exist)">Archaeopteridopsida</a></li> <li>†<a href="/w/index.php?title=Protopityales&action=edit&redlink=1" class="new" title="Protopityales (page does not exist)">Protopityales</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span style="margin: -25px; display: inline-block; -webkit-transform: rotate(270deg); -moz-transform: rotate(270deg); -ms-transform: rotate(270deg); -o-transform: rotate(270deg); transform: rotate(270deg);"><a href="/wiki/Spermatophyte" class="mw-redirect" title="Spermatophyte">Spermatophytes</a><br /><span class="nobold">(seed plants)</span></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><span class="nobold">†</span><a href="/wiki/Pteridospermatophyta" title="Pteridospermatophyta">Pteridosperms</a><span class="nobold">*</span><br /><span class="nobold">(seed ferns)<br />and other extinct<br /> seed plant groups</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>†<a href="/wiki/Bennettitales" title="Bennettitales">Bennettitales</a></li> <li>†<a href="/wiki/Calamopityales" class="mw-redirect" title="Calamopityales">Calamopityales</a></li> <li>†<a href="/wiki/Callistophytales" title="Callistophytales">Callistophytales</a></li> <li>†<a href="/wiki/Caytoniales" title="Caytoniales">Caytoniales</a></li> <li>†<a href="/wiki/Cordaitales" title="Cordaitales">Cordaitales</a></li> <li>†<a href="/wiki/Corystospermaceae" title="Corystospermaceae">Corystospermales</a></li> <li>†<a href="/wiki/Czekanowskiales" title="Czekanowskiales">Czekanowskiales</a></li> <li>†<a href="/wiki/Gigantopteridales" class="mw-redirect" title="Gigantopteridales">Gigantopteridales</a></li> <li>†<a href="/wiki/Glossopteridales" title="Glossopteridales">Glossopteridales</a></li> <li>†<a href="/wiki/Lyginopteridales" title="Lyginopteridales">Lyginopteridopsida</a></li> <li>†<a href="/wiki/Medullosales" title="Medullosales">Medullosales</a></li> <li>†<a href="/wiki/Peltaspermales" title="Peltaspermales">Peltaspermopsida</a></li> <li>†<a href="/wiki/Pentoxylales" title="Pentoxylales">Pentoxylopsida</a></li> <li>†<a href="/w/index.php?title=Petriellales&action=edit&redlink=1" class="new" title="Petriellales (page does not exist)">Petriellales</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Gymnosperm" title="Gymnosperm">Acrogymnospermae</a><br /><span class="nobold">(living gymnosperms)</span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cycad" title="Cycad">Cycadopsida (cycads)</a></li> <li><a href="/wiki/Ginkgoales" title="Ginkgoales">Ginkgoales (<i>Ginkgo</i>)</a></li> <li><a href="/wiki/Conifer" title="Conifer">Pinopsida (conifers)</a></li> <li><a href="/wiki/Gnetophyta" title="Gnetophyta">Gnetopsida</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: #D4EED4;"><a href="/wiki/Flowering_plant" title="Flowering plant">Angiospermae</a><br /><span class="nobold">(flowering plants)</span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Basal_angiosperms" title="Basal angiosperms">Basal angiosperms</a></li> <li><a href="/wiki/Mesangiospermae" title="Mesangiospermae">Core angiosperms</a> <ul><li><a href="/wiki/Magnoliids" title="Magnoliids">Magnoliids</a></li> <li><a href="/wiki/Monocotyledon" title="Monocotyledon">Monocots</a></li> <li><a href="/wiki/Eudicots" title="Eudicots">Eudicots</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: #C3EEC3;"><div> <ul><li>*<a href="/wiki/Paraphyletic" class="mw-redirect" title="Paraphyletic">paraphyletic</a> groups</li> <li>† = <a href="/wiki/Extinction" title="Extinction">extinct</a></li> <li><span class="noviewer" typeof="mw:File"><span title="List-Class article"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/16px-Symbol_list_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/23px-Symbol_list_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/31px-Symbol_list_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/List_of_plant_orders" title="List of plant orders">List of plant orders</a></li> <li><a href="/wiki/Plant#Current_definitions_of_Plantae" title="Plant">Current definitions of Plantae</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Ophioglossidae</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q7097935" class="extiw" title="wikidata:Q7097935">Q7097935</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Plant_Name_Index" title="Australian Plant Name Index">APNI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://id.biodiversity.org.au/name/apni/224850">224850</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/8796847">8796847</a></span></span></li> <li><span style="white-space:nowrap;">Euro+Med PlantBase: <span class="uid"><a rel="nofollow" class="external text" href="https://www.europlusmed.org/cdm_dataportal/taxon/f0322bb2-959b-4716-b9ce-cc847d879fd9">f0322bb2-959b-4716-b9ce-cc847d879fd9</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=11907106">11907106</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=846533">846533</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=1521257">1521257</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/6a9c960c-21d9-4290-aeb1-badba1e7bcf4">6a9c960c-21d9-4290-aeb1-badba1e7bcf4</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=687390">687390</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=55182">55182</a></span></span></li> <li><span style="white-space:nowrap;">VASCAN: <span class="uid"><a rel="nofollow" class="external text" href="https://data.canadensys.net/vascan/taxon/48">48</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=450775">450775</a></span></span></li></ul> </div></td></tr></tbody></table></div> <p><br /> </p> <style data-mw-deduplicate="TemplateStyles:r1012311289">.mw-parser-output .asbox{position:relative;overflow:hidden}.mw-parser-output .asbox table{background:transparent}.mw-parser-output .asbox p{margin:0}.mw-parser-output .asbox p+p{margin-top:0.25em}.mw-parser-output .asbox-body{font-style:italic}.mw-parser-output .asbox-note{font-size:smaller}.mw-parser-output .asbox .navbar{position:absolute;top:-0.75em;right:1em;display:none}</style><div role="note" class="metadata plainlinks asbox stub"><table role="presentation"><tbody><tr class="noresize"><td><span typeof="mw:File"><a href="/wiki/File:Silver-fern.jpg" class="mw-file-description"><img alt="Stub icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/Silver-fern.jpg/40px-Silver-fern.jpg" decoding="async" width="40" height="30" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/Silver-fern.jpg/60px-Silver-fern.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/97/Silver-fern.jpg/80px-Silver-fern.jpg 2x" data-file-width="1184" data-file-height="890" /></a></span></td><td><p class="asbox-body">This <a href="/wiki/Fern" 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You can help Wikipedia by <a class="external text" href="https://en.wikipedia.org/w/index.php?title=Ophioglossidae&action=edit">expanding it</a>.</p></td></tr></tbody></table><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Fern-stub" title="Template:Fern-stub"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Fern-stub" title="Template talk:Fern-stub"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Fern-stub" title="Special:EditPage/Template:Fern-stub"><abbr title="Edit this template">e</abbr></a></li></ul></div></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐f69cdc8f6‐nhjg5 Cached time: 20241123134127 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1] CPU time usage: 0.843 seconds Real time usage: 1.173 seconds Preprocessor visited node count: 5712/1000000 Post‐expand include size: 303246/2097152 bytes Template argument size: 6601/2097152 bytes Highest expansion depth: 24/100 Expensive parser function count: 14/500 Unstrip recursion depth: 0/20 Unstrip post‐expand size: 53731/5000000 bytes Lua time usage: 0.572/10.000 seconds Lua memory usage: 9493631/52428800 bytes Number of Wikibase entities loaded: 12/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 992.099 1 -total 25.95% 257.481 27 Template:Navbox 21.97% 217.972 1 Template:Automatic_taxobox 20.05% 198.962 1 Template:Plant_classification 19.81% 196.576 1 Template:Navbox_with_collapsible_groups 13.37% 132.688 7 Template:Cite_journal 12.67% 125.680 1 Template:Taxonbar 12.22% 121.242 4 Template:Clade 9.05% 89.778 1 Template:Short_description 6.96% 69.058 1 Template:Commons_category --> <!-- Saved in parser cache with key enwiki:pcache:34945384:|#|:idhash:canonical and timestamp 20241123134127 and revision id 1213904569. 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