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Metatheria - Wikipedia
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id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Metatheria</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 31 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-31" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">31 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A8%D8%B9%D8%AF_%D9%88%D8%AD%D8%B4%D9%8A%D8%A7%D8%AA" title="بعد وحشيات – Arabic" lang="ar" hreflang="ar" data-title="بعد وحشيات" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-an mw-list-item"><a href="https://an.wikipedia.org/wiki/Metatheria" title="Metatheria – Aragonese" lang="an" hreflang="an" data-title="Metatheria" data-language-autonym="Aragonés" data-language-local-name="Aragonese" class="interlanguage-link-target"><span>Aragonés</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Metateris" title="Metateris – Catalan" lang="ca" hreflang="ca" data-title="Metateris" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Va%C4%8Dnat%C3%AD" title="Vačnatí – Czech" lang="cs" hreflang="cs" data-title="Vačnatí" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de badge-Q70894304 mw-list-item" title=""><a href="https://de.wikipedia.org/wiki/Metatheria" title="Metatheria – German" lang="de" hreflang="de" data-title="Metatheria" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%9C%CE%B5%CF%84%CE%B1%CE%B8%CE%AE%CF%81%CE%B9%CE%B1" title="Μεταθήρια – Greek" lang="el" hreflang="el" data-title="Μεταθήρια" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Metatheria" title="Metatheria – Spanish" lang="es" hreflang="es" data-title="Metatheria" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Metatheria" title="Metatheria – Basque" lang="eu" hreflang="eu" data-title="Metatheria" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%BE%D8%B3%E2%80%8C%D8%AF%D8%AF%D8%A7%D9%86" title="پسددان – Persian" lang="fa" hreflang="fa" data-title="پسددان" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Metatheria" title="Metatheria – French" lang="fr" hreflang="fr" data-title="Metatheria" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%9B%84%EC%88%98%ED%95%98%EA%B0%95" title="후수하강 – Korean" lang="ko" hreflang="ko" data-title="후수하강" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Metatheria" title="Metatheria – Indonesian" lang="id" hreflang="id" data-title="Metatheria" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%9E%D7%98%D7%90%D7%AA%D7%A8%D7%99%D7%94" title="מטאתריה – Hebrew" lang="he" hreflang="he" data-title="מטאתריה" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A2%D3%A9%D0%BC%D0%B5%D0%BD%D0%B3%D1%96_%D1%81%D0%B0%D1%82%D1%8B%D0%B4%D0%B0%D2%93%D1%8B_%D0%B0%D2%A3%D0%B4%D0%B0%D1%80" title="Төменгі сатыдағы аңдар – Kazakh" lang="kk" hreflang="kk" data-title="Төменгі сатыдағы аңдар" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%91%D0%B0%D1%88%D1%82%D1%8B%D0%BA%D1%87%D0%B0%D0%BD%D0%B4%D0%B0%D1%80" title="Баштыкчандар – Kyrgyz" lang="ky" hreflang="ky" data-title="Баштыкчандар" data-language-autonym="Кыргызча" data-language-local-name="Kyrgyz" class="interlanguage-link-target"><span>Кыргызча</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Apvienot%C4%81_somai%C5%86u_infraklase" title="Apvienotā somaiņu infraklase – Latvian" lang="lv" hreflang="lv" data-title="Apvienotā somaiņu infraklase" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lij mw-list-item"><a href="https://lij.wikipedia.org/wiki/Metatheria" title="Metatheria – Ligurian" lang="lij" hreflang="lij" data-title="Metatheria" data-language-autonym="Ligure" data-language-local-name="Ligurian" class="interlanguage-link-target"><span>Ligure</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Metatheria" title="Metatheria – Hungarian" lang="hu" hreflang="hu" data-title="Metatheria" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A8%D8%B9%D8%AF_%D9%88%D8%AD%D8%B4%D9%8A%D8%A7%D8%AA" title="بعد وحشيات – Egyptian Arabic" lang="arz" hreflang="arz" data-title="بعد وحشيات" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Metatheria" title="Metatheria – Dutch" lang="nl" hreflang="nl" data-title="Metatheria" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%BE%8C%E7%8D%A3%E4%B8%8B%E7%B6%B1" title="後獣下綱 – Japanese" lang="ja" hreflang="ja" data-title="後獣下綱" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Ssaki_ni%C5%BCsze" title="Ssaki niższe – Polish" lang="pl" hreflang="pl" data-title="Ssaki niższe" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9C%D0%B5%D1%82%D0%B0%D1%82%D0%B5%D1%80%D0%B8%D0%B8" title="Метатерии – Russian" lang="ru" hreflang="ru" data-title="Метатерии" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Metatheria" title="Metatheria – Simple English" lang="en-simple" hreflang="en-simple" data-title="Metatheria" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Metatheria" title="Metatheria – Serbian" lang="sr" hreflang="sr" data-title="Metatheria" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Metatheria" title="Metatheria – Finnish" lang="fi" hreflang="fi" data-title="Metatheria" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B9%80%E0%B8%A1%E0%B8%97%E0%B8%B2%E0%B9%80%E0%B8%98%E0%B8%AD%E0%B9%80%E0%B8%A3%E0%B8%B5%E0%B8%A2" title="เมทาเธอเรีย – Thai" lang="th" hreflang="th" data-title="เมทาเธอเรีย" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tg mw-list-item"><a href="https://tg.wikipedia.org/wiki/%D0%A5%D0%B0%D0%BB%D1%82%D0%B0%D0%B4%D0%BE%D1%80%D2%B3%D0%BE" title="Халтадорҳо – Tajik" lang="tg" hreflang="tg" data-title="Халтадорҳо" data-language-autonym="Тоҷикӣ" data-language-local-name="Tajik" class="interlanguage-link-target"><span>Тоҷикӣ</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9D%D0%B8%D0%B6%D1%87%D1%96_%D0%B7%D0%B2%D1%96%D1%80%D1%96" title="Нижчі звірі – Ukrainian" lang="uk" hreflang="uk" data-title="Нижчі звірі" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E5%BE%8C%E7%8D%B8%E4%BA%9E%E7%B6%B1" title="後獸亞綱 – Cantonese" lang="yue" hreflang="yue" data-title="後獸亞綱" data-language-autonym="粵語" data-language-local-name="Cantonese" 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rgb(235,235,210)">Metatheria<br /><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;"><a href="/wiki/Early_Cretaceous" title="Early Cretaceous">Early Cretaceous</a>–recent<sup id="cite_ref-SCI-20130208_1-0" class="reference"><a href="#cite_note-SCI-20130208-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Bennett_2018_2-0" class="reference"><a href="#cite_note-Bennett_2018-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Bi_2018_3-0" class="reference"><a href="#cite_note-Bi_2018-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:17.583076923077px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.22px; width:14.807692307692px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:70.027692307692px; width:7.9470769230769px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:77.974769230769px; width:20.564923076923px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.539692307692px; width:20.294153846154px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.732307692308px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:171.56615384615px; width:26.095384615385px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.540307692308px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20184615385px; width:6.9249230769231px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:181.68615384615px; width:38.313846153846px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:181.68615384615px; width:38.313846153846px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:182.68615384615px; width:36.313846153846px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Lycopsis_longirostris.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ab/Lycopsis_longirostris.JPG/220px-Lycopsis_longirostris.JPG" decoding="async" width="220" height="154" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ab/Lycopsis_longirostris.JPG/330px-Lycopsis_longirostris.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ab/Lycopsis_longirostris.JPG/440px-Lycopsis_longirostris.JPG 2x" data-file-width="2253" data-file-height="1572" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i><a href="/wiki/Lycopsis_longirostris" class="mw-redirect" title="Lycopsis longirostris">Lycopsis longirostris</a></i>, an extinct <a href="/wiki/Sparassodonta" title="Sparassodonta">sparassodont</a>, a relative of the <a href="/wiki/Marsupial" title="Marsupial">marsupials</a> </td></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Mouse_Possum_-Tambopata_Reserve_-Peru-8.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/12/Mouse_Possum_-Tambopata_Reserve_-Peru-8.jpg/220px-Mouse_Possum_-Tambopata_Reserve_-Peru-8.jpg" decoding="async" width="220" height="151" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/12/Mouse_Possum_-Tambopata_Reserve_-Peru-8.jpg/330px-Mouse_Possum_-Tambopata_Reserve_-Peru-8.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/12/Mouse_Possum_-Tambopata_Reserve_-Peru-8.jpg/440px-Mouse_Possum_-Tambopata_Reserve_-Peru-8.jpg 2x" data-file-width="2808" data-file-height="1926" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">A mouse opossum (<i><a href="/wiki/Marmosa" title="Marmosa">Marmosa</a></i>) </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Metatheria" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Mammal" title="Mammal">Mammalia</a> </td></tr> <tr> <td>Subclass: </td> <td><a href="/wiki/Theria" title="Theria">Theria</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a class="mw-selflink selflink">Metatheria</a><br /><small><a href="/wiki/Thomas_Henry_Huxley" title="Thomas Henry Huxley">Huxley</a>, 1880</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Subgroups </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i>?<a href="/wiki/Sinodelphys" title="Sinodelphys">Sinodelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Holoclemensia" title="Holoclemensia">Holoclemensia</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Adinodon&action=edit&redlink=1" class="new" title="Adinodon (page does not exist)">Adinodon</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Deltatheroida" title="Deltatheroida">Deltatheroida</a></li> <li><b>Marsupialiformes</b> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Adelodelphys&action=edit&redlink=1" class="new" title="Adelodelphys (page does not exist)">Adelodelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Alphadontidae" title="Alphadontidae">Alphadontidae</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Asiadelphia" class="mw-redirect" title="Asiadelphia">Asiadelphia</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Sinbadelphys&action=edit&redlink=1" class="new" title="Sinbadelphys (page does not exist)">Sinbadelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Camptomus&action=edit&redlink=1" class="new" title="Camptomus (page does not exist)">Camptomus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><b>Archimetatheria</b> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Stagodontidae" title="Stagodontidae">Stagodontidae</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Iqualadelphis&action=edit&redlink=1" class="new" title="Iqualadelphis (page does not exist)">Iqualadelphis</a></i></li></ul></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Sparassodonta" title="Sparassodonta">Sparassodonta</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/w/index.php?title=Pucadelphyidae&action=edit&redlink=1" class="new" title="Pucadelphyidae (page does not exist)">Pucadelphyidae</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Herpetotheriidae" title="Herpetotheriidae">Herpetotheriidae</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Anatoliadelphyidae" title="Anatoliadelphyidae">Anatoliadelphyidae</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Polydolopimorphia" title="Polydolopimorphia">Polydolopimorphia</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Peradectidae" title="Peradectidae">Peradectidae</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/w/index.php?title=Pediomyidae&action=edit&redlink=1" class="new" title="Pediomyidae (page does not exist)">Pediomyidae</a></li> <li><a href="/wiki/Marsupialia" class="mw-redirect" title="Marsupialia">Marsupialia</a></li></ul></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Metatheria</b> is a mammalian <a href="/wiki/Clade" title="Clade">clade</a> that includes all <a href="/wiki/Mammal" title="Mammal">mammals</a> more closely related to <a href="/wiki/Marsupial" title="Marsupial">marsupials</a> than to <a href="/wiki/Placental" class="mw-redirect" title="Placental">placentals</a>. First proposed by <a href="/wiki/Thomas_Henry_Huxley" title="Thomas Henry Huxley">Thomas Henry Huxley</a> in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well as many extinct non-marsupial relatives. It is one of two groups placed in the clade <a href="/wiki/Theria" title="Theria">Theria</a> alongside <a href="/wiki/Eutheria" title="Eutheria">Eutheria</a>, which contains the placentals. Remains of metatherians have been found on all of Earth’s continents. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Metatheria&action=edit&section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Distinctive characteristics (<a href="/wiki/Synapomorphies" class="mw-redirect" title="Synapomorphies">synapomorphies</a>) of Metatheria include a <a href="/wiki/Prehensile_tail" title="Prehensile tail">prehensile tail</a>, the development of a capitular tail on the humerus, the loss of tooth replacement on the second and fifth <a href="/wiki/Premolars" class="mw-redirect" title="Premolars">premolars</a>, lower <a href="/wiki/Canine_tooth" title="Canine tooth">canines</a> that outwardly diverge from each other, an <a href="/wiki/Angular_process" class="mw-redirect" title="Angular process">angular process</a> on the dentary bone—which additionally bears a posterior shelf in its masseteric fossa in Metatheria<sup id="cite_ref-:2_4-0" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup>—that is equal to or greater than the length of the <a href="/wiki/Ramus_mandibulae" class="mw-redirect" title="Ramus mandibulae">ramus</a>,<sup id="cite_ref-:2_4-1" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup class="reference nowrap"><span title="Page / location: Appendix 2">: Appendix 2 </span></sup> and the lower fifth premolar with a "very trenchant"<sup id="cite_ref-:2_4-2" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> cristid obliqua/ectolophid. The permanent, retained deciduous fifth premolars are <a href="/wiki/Molar_(tooth)" title="Molar (tooth)">molar</a> like and were historically identified as first molars, with the third premolar found in basal <a href="/wiki/Theria" title="Theria">therians</a> being lost, leaving four premolars in the halves of each jaw.<sup id="cite_ref-:2_4-3" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> <i><a href="/wiki/Sinodelphys" title="Sinodelphys">Sinodelphys</a></i> differs in its dentition and may represent a basal metatherian.<sup id="cite_ref-:2_4-4" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Evolutionary_history">Evolutionary history</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Metatheria&action=edit&section=2" title="Edit section: Evolutionary history"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The relationships between the three extant divisions of mammals (<a href="/wiki/Monotreme" title="Monotreme">monotremes</a>, marsupials, and <a href="/wiki/Placentalia" title="Placentalia">placentals</a>) was long a matter of debate among <a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">taxonomists</a>.<sup id="cite_ref-Moyal2004_6-0" class="reference"><a href="#cite_note-Moyal2004-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> Most <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphological</a> evidence comparing traits, such as the <a href="/wiki/Dentition" title="Dentition">number and arrangement of teeth</a> and the structure of the <a href="/wiki/Genitourinary_system" title="Genitourinary system">reproductive and waste elimination systems</a>, favors a closer evolutionary relationship between marsupials and placental mammals than either has with the monotremes, as does most <a href="/wiki/Molecular_genetics" title="Molecular genetics">genetic and molecular</a> evidence.<sup id="cite_ref-Rheede2005_7-0" class="reference"><a href="#cite_note-Rheede2005-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>The earliest possible known metatherian is <i><a href="/wiki/Sinodelphys" title="Sinodelphys">Sinodelphys szalayi</a></i>, which lived in <a href="/wiki/China" title="China">China</a> during the <a href="/wiki/Early_Cretaceous" title="Early Cretaceous">Early Cretaceous</a> around 125 million years ago (mya).<sup id="cite_ref-:2_4-5" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> This makes it a contemporary to some early eutherian species that have been found in the same area.<sup id="cite_ref-acristatherium_9-0" class="reference"><a href="#cite_note-acristatherium-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> However, Bi <i>et al.</i> (2018) reinterpreted <i>Sinodelphys</i> as an early member of Eutheria. The oldest uncontested metatherians are now 110 million year old fossils from western North America.<sup id="cite_ref-Bi_2018_3-1" class="reference"><a href="#cite_note-Bi_2018-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Metatherians were widespread in Asia and North America during the Late Cretaceous, including both <a href="/wiki/Deltatheroida" title="Deltatheroida">Deltatheroida</a> and Marsupialiformes,<sup id="cite_ref-:0_10-0" class="reference"><a href="#cite_note-:0-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> with fossils also known from Europe during this time. During the Late Cretaceous, metatherians were more diverse than eutherians in North America.<sup id="cite_ref-:2_4-6" class="reference"><a href="#cite_note-:2-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Metatherians underwent a severe decline during the <a href="/wiki/Cretaceous%E2%80%93Paleogene_extinction_event" title="Cretaceous–Paleogene extinction event">Cretaceous–Paleogene extinction event</a>, more severe than that suffered by contemporary eutherians and <a href="/wiki/Multituberculates" class="mw-redirect" title="Multituberculates">multituberculates</a>, and were slower to recover diversity.<sup id="cite_ref-:0_10-1" class="reference"><a href="#cite_note-:0-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>Morphological and species diversity of metatherians in <a href="/wiki/Laurasia" title="Laurasia">Laurasia</a> remained low in comparison to eutherians throughout the Cenozoic.<sup id="cite_ref-:1_11-0" class="reference"><a href="#cite_note-:1-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> The two major groups of Cenozoic Laurasian metatherians, the opossum-like <a href="/wiki/Herpetotheriidae" title="Herpetotheriidae">herpetotheriids</a> and <a href="/wiki/Peradectidae" title="Peradectidae">peradectids</a> persisted into the <a href="/wiki/Miocene" title="Miocene">Miocene</a> before becoming extinct, with the North American herpetotheriid <i><a href="/wiki/Herpetotherium" title="Herpetotherium">Herpetotherium</a></i>, the European herpetotheriid <i><a href="/wiki/Amphiperatherium" title="Amphiperatherium">Amphiperatherium</a></i> and the peradectids <i><a href="/wiki/Siamoperadectes" title="Siamoperadectes">Siamoperadectes</a></i> and <i><a href="/w/index.php?title=Sinoperadectes&action=edit&redlink=1" class="new" title="Sinoperadectes (page does not exist)">Sinoperadectes</a></i> from Asia being the youngest Laurasian non-marsupial metatherians (with marsupials invading North America during the <a href="/wiki/Pliocene" title="Pliocene">Pliocene</a>-<a href="/wiki/Pleistocene" title="Pleistocene">Pleistocene</a> as part of the <a href="/wiki/Great_American_Interchange" title="Great American Interchange">Great American interchange</a>).<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:0_10-2" class="reference"><a href="#cite_note-:0-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> Metatherians first arrived in Afro-Arabia during the <a href="/wiki/Paleogene" title="Paleogene">Paleogene</a>, probably from Europe, including the possible peradectoid <i><a href="/w/index.php?title=Kasserinotherium&action=edit&redlink=1" class="new" title="Kasserinotherium (page does not exist)">Kasserinotherium</a></i> from the Early Eocene of Tunisia and the herpetotheriid <i><a href="/wiki/Peratherium_africanum" class="mw-redirect" title="Peratherium africanum">Peratherium africanum</a></i> from the Early <a href="/wiki/Oligocene" title="Oligocene">Oligocene</a> of Egypt and Oman. The youngest African metatherian is the possible herpetotheriid <i><a href="/w/index.php?title=Morotodon&action=edit&redlink=1" class="new" title="Morotodon (page does not exist)">Morotodon</a></i> from the late <a href="/wiki/Early_Miocene" title="Early Miocene">Early Miocene</a> of <a href="/wiki/Uganda" title="Uganda">Uganda</a>.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p><p>Metatherians arrived in South America from North America during the latest Cretaceous or <a href="/wiki/Paleocene" title="Paleocene">Paleocene</a> and underwent a major diversificiation, with South American metatherians including both the ancestors of extant marsupials as well as the extinct <a href="/wiki/Sparassodonta" title="Sparassodonta">Sparassodonta</a>, which were major predators in South American ecosystems during most of the <a href="/wiki/Cenozoic" title="Cenozoic">Cenozoic</a>, up until their extinction in the <a href="/wiki/Pliocene" title="Pliocene">Pliocene</a>, as well as the <a href="/wiki/Polydolopimorphia" title="Polydolopimorphia">Polydolopimorphia</a>, which likely had a wide range of diets.<sup id="cite_ref-:1_11-1" class="reference"><a href="#cite_note-:1-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Metatherians then declined in diversity in South America during the Late Eocene as well as the later Oligocene epoch. The oldest known Australian marsupials are from the early Eocene, and are thought to have arrived in the region after having dispersed via Antarctica from South America.<sup id="cite_ref-:1_11-2" class="reference"><a href="#cite_note-:1-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> During the Oligocene epoch, Australian metatherians radiated rapidly, which contributed most to the global peak in metatherian diversity during the Early Miocene. The only known Antarctic metatherians are from the Early Eocene <a href="/wiki/La_Meseta_Formation" title="La Meseta Formation">La Meseta Formation</a> of the <a href="/wiki/Antarctic_Peninsula" title="Antarctic Peninsula">Antarctic Peninsula</a>, where they are the most diverse group of mammals, and include marsupials as well as polydolopimorphians.<sup id="cite_ref-:1_11-3" class="reference"><a href="#cite_note-:1-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Metatheria&action=edit&section=3" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Below is a metatherian <a href="/wiki/Cladogram" title="Cladogram">cladogram</a> from Wilson et al. (2016):<sup id="cite_ref-wilson2016_15-0" class="reference"><a href="#cite_note-wilson2016-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1261294616">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto;display:table}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leaf p{padding-right:5px;padding-left:2px}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leafR p{padding-left:5px;padding-right:2px}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse p{padding-left:5px;padding-right:2px}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:85%; line-height:85%"> <tbody><tr> <td class="clade-label first">Metatheria </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Holoclemensia" title="Holoclemensia">Holoclemensia</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Deltatheroida" title="Deltatheroida">Deltatheroida</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Pappotheriidae" class="mw-redirect" title="Pappotheriidae">Pappotheriidae</a> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Pappotherium" title="Pappotherium">Pappotherium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Deltatheridiidae" title="Deltatheridiidae">Deltatheridiidae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Sulestes" title="Sulestes">Sulestes</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Oklatheridium" title="Oklatheridium">Oklatheridium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Tsagandelta" title="Tsagandelta">Tsagandelta</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Lotheridium" title="Lotheridium">Lotheridium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Deltatheroides" title="Deltatheroides">Deltatheroides</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Deltatheridium" title="Deltatheridium">Deltatheridium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Nanocuris" title="Nanocuris">Nanocuris</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Atokatheridium" title="Atokatheridium">Atokatheridium</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label">Marsupialiformes </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Gurlin_Tsav_skull" title="Gurlin Tsav skull">Gurlin Tsav skull</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Borhyaenidae" title="Borhyaenidae">Borhyaenidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Mayulestes" title="Mayulestes">Mayulestes</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Jaskhadelphys&action=edit&redlink=1" class="new" title="Jaskhadelphys (page does not exist)">Jaskhadelphys</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Andinodelphys" title="Andinodelphys">Andinodelphys</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Pucadelphys" title="Pucadelphys">Pucadelphys</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Asiatherium" title="Asiatherium">Asiatherium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Iugomortiferum&action=edit&redlink=1" class="new" title="Iugomortiferum (page does not exist)">Iugomortiferum</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Kokopellia&action=edit&redlink=1" class="new" title="Kokopellia (page does not exist)">Kokopellia</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Aenigmadelphys&action=edit&redlink=1" class="new" title="Aenigmadelphys (page does not exist)">Aenigmadelphys</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Anchistodelphys&action=edit&redlink=1" class="new" title="Anchistodelphys (page does not exist)">Anchistodelphys</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Glasbiidae&action=edit&redlink=1" class="new" title="Glasbiidae (page does not exist)">Glasbiidae</a> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Glasbius&action=edit&redlink=1" class="new" title="Glasbius (page does not exist)">Glasbius</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Pediomyidae&action=edit&redlink=1" class="new" title="Pediomyidae (page does not exist)">Pediomyidae</a> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Pediomys" title="Pediomys">Pediomys</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Stagodontidae" title="Stagodontidae">Stagodontidae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Pariadens&action=edit&redlink=1" class="new" title="Pariadens (page does not exist)">Pariadens</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Eodelphis" title="Eodelphis">Eodelphis</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Didelphodon" title="Didelphodon">Didelphodon</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf" style="border-right:2px solid green;border-bottom:0;border-top:0;"> <p><i><a href="/wiki/Turgidodon" title="Turgidodon">Turgidodon</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf" style="border-right:2px solid green;border-bottom:0;border-top:0;"> <p><i><a href="/wiki/Alphadon" title="Alphadon">Alphadon</a></i> </p> </td> <td rowspan="2" class="clade-bar" style=""><a href="/wiki/Alphadontidae" title="Alphadontidae">Alphadontidae</a> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf" style="border-right:2px solid green;border-bottom:0;border-top:0;"> <p><i><a href="/wiki/Albertatherium" title="Albertatherium">Albertatherium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Marsupial" title="Marsupial">Marsupialia</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div><p>Cladogram after<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup>:</p><div class="clade"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first">Metatheria </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Deltatheroida" title="Deltatheroida">Deltatheriidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/w/index.php?title=Kokopellia&action=edit&redlink=1" class="new" title="Kokopellia (page does not exist)">Kokopellia</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Asiatherium" title="Asiatherium">Asiatherium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Peradectidae" title="Peradectidae">Peradectidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Stagodontidae" title="Stagodontidae">Stagodontidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/w/index.php?title=Pucadelphyidae&action=edit&redlink=1" class="new" title="Pucadelphyidae (page does not exist)">Pucadelphyidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Sparassodonta" title="Sparassodonta">Sparassodonta</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf" style="border-right:2px solid green;border-bottom:0;border-top:0;"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Amphiperatherium" title="Amphiperatherium">Amphiperatherium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Peratherium" title="Peratherium">Peratherium</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td> <td rowspan="2" class="clade-bar" style="">"<a href="/wiki/Herpetotheriidae" title="Herpetotheriidae">Herpetotheriidae</a>" </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf" style="border-right:2px solid green;border-bottom:0;border-top:0;"> <p><i><a href="/wiki/Herpetotherium" title="Herpetotherium">Herpetotherium</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Marsupialia" class="mw-redirect" title="Marsupialia">Marsupialia</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <p><br /> Below is a listing of metatherians that do not fall readily into well-defined groups. </p><p>Basal Metatheria </p> <ul><li>†<i><a href="/w/index.php?title=Archaeonothos&action=edit&redlink=1" class="new" title="Archaeonothos (page does not exist)">Archaeonothos henkgodthelpi</a></i> <small>Beck 2015</small></li> <li>†<i><a href="/w/index.php?title=Esteslestes&action=edit&redlink=1" class="new" title="Esteslestes (page does not exist)">Esteslestes ensis</a></i> <small>Novacek et al. 1991</small></li> <li>†<i><a href="/w/index.php?title=Ghamidtherium&action=edit&redlink=1" class="new" title="Ghamidtherium (page does not exist)">Ghamidtherium dimaiensis</a></i> <small>Sánches-Villagra et al. 2007</small></li> <li>†<i><a href="/w/index.php?title=Kasserinotherium&action=edit&redlink=1" class="new" title="Kasserinotherium (page does not exist)">Kasserinotherium tunisiense</a></i> <small>Crochet 1989</small></li> <li>†<i><a href="/w/index.php?title=Palangania&action=edit&redlink=1" class="new" title="Palangania (page does not exist)">Palangania brandmayri</a></i> <small>Goin et al. 1998</small></li> <li>†<i><a href="/w/index.php?title=Perrodelphys&action=edit&redlink=1" class="new" title="Perrodelphys (page does not exist)">Perrodelphys coquinense</a></i> <small>Goin et al. 1999</small></li></ul> <p>Ameridelphia incertae sedis: </p> <ul><li>†<i><a href="/w/index.php?title=Apistodon&action=edit&redlink=1" class="new" title="Apistodon (page does not exist)">Apistodon exiguus</a></i> <small>(Fox 1971) Davis 2007</small></li> <li>†<i><a href="/wiki/Cocatherium" title="Cocatherium">Cocatherium lefipanum</a></i> <small>Goin et al. 2006</small></li> <li>†<i><a href="/w/index.php?title=Dakotadens&action=edit&redlink=1" class="new" title="Dakotadens (page does not exist)">Dakotadens morrowi</a></i> <small>Eaton 1993</small></li> <li>†<i><a href="/w/index.php?title=Iugomortiferum&action=edit&redlink=1" class="new" title="Iugomortiferum (page does not exist)">Iugomortiferum thoringtoni</a></i> <small>Cifelli 1990b</small></li> <li>†<i><a href="/w/index.php?title=Marambiotherium&action=edit&redlink=1" class="new" title="Marambiotherium (page does not exist)">Marambiotherium glacialis</a></i> <small>Goin et al. 1999</small></li> <li>†<i><a href="/w/index.php?title=Marmosopsis&action=edit&redlink=1" class="new" title="Marmosopsis (page does not exist)">Marmosopsis juradoi</a></i> <small>Paula Couto 1962</small> [Marmosopsini <small>Kirsch & Palma 1995</small>]</li> <li>†<i><a href="/w/index.php?title=Pascualdelphys&action=edit&redlink=1" class="new" title="Pascualdelphys (page does not exist)">Pascualdelphys fierroensis</a></i></li> <li>†<i><a href="/w/index.php?title=Progarzonia&action=edit&redlink=1" class="new" title="Progarzonia (page does not exist)">Progarzonia notostylopense</a></i> <small>Ameghino 1904</small></li> <li>†<i><a href="/wiki/Protalphadon" title="Protalphadon">Protalphadon</a></i> <small>Cifelli 1990</small> <ul><li>†<i><a href="/wiki/Protalphadon" title="Protalphadon">P. lulli</a></i> <small>(Clemens 1966) Cifelli 1990a</small></li> <li>†<i><a href="/wiki/Protalphadon" title="Protalphadon">P. foxi</a></i> <small>Johnson 1996</small></li></ul></li></ul> <p>Marsupialia incertae sedis: </p> <ul><li>†<i><a href="/w/index.php?title=Itaboraidelphys&action=edit&redlink=1" class="new" title="Itaboraidelphys (page does not exist)">Itaboraidelphys camposi</a></i> <small>Marshall & de Muizon 1984</small></li> <li>†<i><a href="/w/index.php?title=Mizquedelphys&action=edit&redlink=1" class="new" title="Mizquedelphys (page does not exist)">Mizquedelphys pilpinensis</a></i> <small>Marshall & de Muizon 1988</small></li> <li>†<i><a href="/w/index.php?title=Numbigilga&action=edit&redlink=1" class="new" title="Numbigilga (page does not exist)">Numbigilga ernielundeliusi</a></i> <small>Beck et al. 2008</small> {Numbigilgidae <small>Beck et al. 2008</small>}</li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Metatheria&action=edit&section=4" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 25em;"> <ol class="references"> <li id="cite_note-SCI-20130208-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-SCI-20130208_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFO'LearyBlochFlynnGaudin2013" class="citation journal cs1">O'Leary, Maureen A.; Bloch, Jonathan I.; Flynn, John J.; Gaudin, Timothy J.; Giallombardo, Andres; Giannini, Norberto P.; Goldberg, Suzann L.; Kraatz, Brian P.; Luo, Zhe-Xi; Meng, Jin; Ni, Michael J.; Novacek, Fernando A.; Perini, Zachary S.; Randall, Guillermo; Rougier, Eric J.; Sargis, Mary T.; Silcox, Nancy b.; Simmons, Micelle; Spaulding, Paul M.; Velazco, Marcelo; Weksler, John R.; Wible, Andrea L.; Cirranello, A. L. (8 February 2013). "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". <i><a href="/wiki/Science_(journal)" title="Science (journal)">Science</a></i>. <b>339</b> (6120): <span class="nowrap">662–</span>667. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2013Sci...339..662O">2013Sci...339..662O</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1229237">10.1126/science.1229237</a>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/11336%2F7302">11336/7302</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/23393258">23393258</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:206544776">206544776</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=The+Placental+Mammal+Ancestor+and+the+Post%E2%80%93K-Pg+Radiation+of+Placentals&rft.volume=339&rft.issue=6120&rft.pages=%3Cspan+class%3D%22nowrap%22%3E662-%3C%2Fspan%3E667&rft.date=2013-02-08&rft_id=info%3Ahdl%2F11336%2F7302&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A206544776%23id-name%3DS2CID&rft_id=info%3Abibcode%2F2013Sci...339..662O&rft_id=info%3Apmid%2F23393258&rft_id=info%3Adoi%2F10.1126%2Fscience.1229237&rft.aulast=O%27Leary&rft.aufirst=Maureen+A.&rft.au=Bloch%2C+Jonathan+I.&rft.au=Flynn%2C+John+J.&rft.au=Gaudin%2C+Timothy+J.&rft.au=Giallombardo%2C+Andres&rft.au=Giannini%2C+Norberto+P.&rft.au=Goldberg%2C+Suzann+L.&rft.au=Kraatz%2C+Brian+P.&rft.au=Luo%2C+Zhe-Xi&rft.au=Meng%2C+Jin&rft.au=Ni%2C+Michael+J.&rft.au=Novacek%2C+Fernando+A.&rft.au=Perini%2C+Zachary+S.&rft.au=Randall%2C+Guillermo&rft.au=Rougier%2C+Eric+J.&rft.au=Sargis%2C+Mary+T.&rft.au=Silcox%2C+Nancy+b.&rft.au=Simmons%2C+Micelle&rft.au=Spaulding%2C+Paul+M.&rft.au=Velazco%2C+Marcelo&rft.au=Weksler%2C+John+R.&rft.au=Wible%2C+Andrea+L.&rft.au=Cirranello%2C+A.+L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-Bennett_2018-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-Bennett_2018_2-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFC.V._BennettP._FranciscoF._J._GoinA._Goswami2018" class="citation journal cs1">C.V. Bennett; P. Francisco; F. J. Goin; A. Goswami (2018). <a rel="nofollow" class="external text" href="https://doi.org/10.1017%2Fpab.2017.34">"Deep time diversity of metatherian mammals: implications for evolutionary history and fossil-record quality"</a>. <i>Paleobiology</i>. <b>44</b> (2): <span class="nowrap">171–</span>198. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2018Pbio...44..171B">2018Pbio...44..171B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1017%2Fpab.2017.34">10.1017/pab.2017.34</a></span>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/11336%2F94590">11336/94590</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Paleobiology&rft.atitle=Deep+time+diversity+of+metatherian+mammals%3A+implications+for+evolutionary+history+and+fossil-record+quality&rft.volume=44&rft.issue=2&rft.pages=%3Cspan+class%3D%22nowrap%22%3E171-%3C%2Fspan%3E198&rft.date=2018&rft_id=info%3Ahdl%2F11336%2F94590&rft_id=info%3Adoi%2F10.1017%2Fpab.2017.34&rft_id=info%3Abibcode%2F2018Pbio...44..171B&rft.au=C.V.+Bennett&rft.au=P.+Francisco&rft.au=F.+J.+Goin&rft.au=A.+Goswami&rft_id=https%3A%2F%2Fdoi.org%2F10.1017%252Fpab.2017.34&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-Bi_2018-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-Bi_2018_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Bi_2018_3-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFS._BiX._ZhengX._WangN.E._Cignetti2018" class="citation journal cs1">S. Bi; X. Zheng; X. Wang; N.E. Cignetti; S. Yang; J.R. Wible (2018). "An Early Cretaceous eutherian and the placental–marsupial dichotomy". <i>Nature</i>. <b>558</b> (7710): <span class="nowrap">390–</span>395. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2018Natur.558..390B">2018Natur.558..390B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fs41586-018-0210-3">10.1038/s41586-018-0210-3</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29899454">29899454</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:49183466">49183466</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=An+Early+Cretaceous+eutherian+and+the+placental%E2%80%93marsupial+dichotomy&rft.volume=558&rft.issue=7710&rft.pages=%3Cspan+class%3D%22nowrap%22%3E390-%3C%2Fspan%3E395&rft.date=2018&rft_id=info%3Adoi%2F10.1038%2Fs41586-018-0210-3&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A49183466%23id-name%3DS2CID&rft_id=info%3Apmid%2F29899454&rft_id=info%3Abibcode%2F2018Natur.558..390B&rft.au=S.+Bi&rft.au=X.+Zheng&rft.au=X.+Wang&rft.au=N.E.+Cignetti&rft.au=S.+Yang&rft.au=J.R.+Wible&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-:2-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-:2_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:2_4-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:2_4-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-:2_4-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-:2_4-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-:2_4-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-:2_4-6"><sup><i><b>g</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilliamsonBrusatteWilson2014" class="citation journal cs1">Williamson, Thomas E.; Brusatte, Stephen L.; Wilson, Gregory P. (17 December 2014). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4284630">"The origin and early evolution of metatherian mammals: the Cretaceous record"</a>. <i>ZooKeys</i> (465): <span class="nowrap">1–</span>76. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2014ZooK..465....1W">2014ZooK..465....1W</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3897%2Fzookeys.465.8178">10.3897/zookeys.465.8178</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1313-2970">1313-2970</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4284630">4284630</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/25589872">25589872</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=ZooKeys&rft.atitle=The+origin+and+early+evolution+of+metatherian+mammals%3A+the+Cretaceous+record&rft.issue=465&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1-%3C%2Fspan%3E76&rft.date=2014-12-17&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC4284630%23id-name%3DPMC&rft_id=info%3Abibcode%2F2014ZooK..465....1W&rft_id=info%3Apmid%2F25589872&rft_id=info%3Adoi%2F10.3897%2Fzookeys.465.8178&rft.issn=1313-2970&rft.aulast=Williamson&rft.aufirst=Thomas+E.&rft.au=Brusatte%2C+Stephen+L.&rft.au=Wilson%2C+Gregory+P.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC4284630&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWibleRougierNovacekAsher2009" class="citation journal cs1">Wible, John R.; Rougier, Guillermo R.; Novacek, Michael J.; Asher, Robert J. (2009). <a rel="nofollow" class="external text" href="https://doi.org/10.1206%2F623.1">"The Eutherian Mammal <i>Maelestes gobiensis</i> from the Late Cretaceous of Mongolia and the phylogeny of cretaceous eutheria"</a>. <i>Bulletin of the American Museum of Natural History</i>. <b>2009</b> (327): <span class="nowrap">1–</span>123. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1206%2F623.1">10.1206/623.1</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Bulletin+of+the+American+Museum+of+Natural+History&rft.atitle=The+Eutherian+Mammal+Maelestes+gobiensis+from+the+Late+Cretaceous+of+Mongolia+and+the+phylogeny+of+cretaceous+eutheria&rft.volume=2009&rft.issue=327&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1-%3C%2Fspan%3E123&rft.date=2009&rft_id=info%3Adoi%2F10.1206%2F623.1&rft.aulast=Wible&rft.aufirst=John+R.&rft.au=Rougier%2C+Guillermo+R.&rft.au=Novacek%2C+Michael+J.&rft.au=Asher%2C+Robert+J.&rft_id=https%3A%2F%2Fdoi.org%2F10.1206%252F623.1&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-Moyal2004-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-Moyal2004_6-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMoyal,_Ann_Mozley2004" class="citation book cs1">Moyal, Ann Mozley (2004). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=5DkezNMhSTYC"><i>Platypus: The Extraordinary Story of How a Curious Creature Baffled the World</i></a>. Baltimore: The Johns Hopkins University Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-8018-8052-0" title="Special:BookSources/978-0-8018-8052-0"><bdi>978-0-8018-8052-0</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Platypus%3A+The+Extraordinary+Story+of+How+a+Curious+Creature+Baffled+the+World&rft.place=Baltimore&rft.pub=The+Johns+Hopkins+University+Press&rft.date=2004&rft.isbn=978-0-8018-8052-0&rft.au=Moyal%2C+Ann+Mozley&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3D5DkezNMhSTYC&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-Rheede2005-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-Rheede2005_7-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFvan_RheedeBastiaansBooneHedges2006" class="citation journal cs1">van Rheede, T.; Bastiaans, T.; Boone, D.; Hedges, S.; De Jong, W.; Madsen, O. (2006). <a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fmolbev%2Fmsj064">"The platypus is in its place: nuclear genes and indels confirm the sister group relation of monotremes and therians"</a>. <i>Molecular Biology and Evolution</i>. <b>23</b> (3): <span class="nowrap">587–</span>597. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fmolbev%2Fmsj064">10.1093/molbev/msj064</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16291999">16291999</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Molecular+Biology+and+Evolution&rft.atitle=The+platypus+is+in+its+place%3A+nuclear+genes+and+indels+confirm+the+sister+group+relation+of+monotremes+and+therians&rft.volume=23&rft.issue=3&rft.pages=%3Cspan+class%3D%22nowrap%22%3E587-%3C%2Fspan%3E597&rft.date=2006&rft_id=info%3Adoi%2F10.1093%2Fmolbev%2Fmsj064&rft_id=info%3Apmid%2F16291999&rft.aulast=van+Rheede&rft.aufirst=T.&rft.au=Bastiaans%2C+T.&rft.au=Boone%2C+D.&rft.au=Hedges%2C+S.&rft.au=De+Jong%2C+W.&rft.au=Madsen%2C+O.&rft_id=https%3A%2F%2Fdoi.org%2F10.1093%252Fmolbev%252Fmsj064&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRincon2003" class="citation web cs1">Rincon, Paul (12 December 2003). <a rel="nofollow" class="external text" href="http://news.bbc.co.uk/2/hi/science/nature/3311911.stm">"Oldest Marsupial Ancestor Found, BBC, Dec 2003"</a>. BBC News<span class="reference-accessdate">. Retrieved <span class="nowrap">16 March</span> 2010</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Oldest+Marsupial+Ancestor+Found%2C+BBC%2C+Dec+2003&rft.pub=BBC+News&rft.date=2003-12-12&rft.aulast=Rincon&rft.aufirst=Paul&rft_id=http%3A%2F%2Fnews.bbc.co.uk%2F2%2Fhi%2Fscience%2Fnature%2F3311911.stm&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-acristatherium-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-acristatherium_9-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHuMengLiWang2010" class="citation journal cs1">Hu, Yaoming; Meng, Jin; Li, Chuankui; Wang, Yuanqing (2010). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2842663">"New basal eutherian mammal from the Early Cretaceous Jehol biota, Liaoning, China"</a>. <i><a href="/wiki/Proceedings_of_the_Royal_Society_B" class="mw-redirect" title="Proceedings of the Royal Society B">Proceedings of the Royal Society B</a></i>. <b>277</b> (1679): <span class="nowrap">229–</span>236. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frspb.2009.0203">10.1098/rspb.2009.0203</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2842663">2842663</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/19419990">19419990</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+Royal+Society+B&rft.atitle=New+basal+eutherian+mammal+from+the+Early+Cretaceous+Jehol+biota%2C+Liaoning%2C+China&rft.volume=277&rft.issue=1679&rft.pages=%3Cspan+class%3D%22nowrap%22%3E229-%3C%2Fspan%3E236&rft.date=2010&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2842663%23id-name%3DPMC&rft_id=info%3Apmid%2F19419990&rft_id=info%3Adoi%2F10.1098%2Frspb.2009.0203&rft.aulast=Hu&rft.aufirst=Yaoming&rft.au=Meng%2C+Jin&rft.au=Li%2C+Chuankui&rft.au=Wang%2C+Yuanqing&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2842663&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-:0-10"><span class="mw-cite-backlink">^ <a href="#cite_ref-:0_10-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:0_10-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:0_10-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBennettUpchurchGoinGoswami2018" class="citation journal cs1">Bennett, C. Verity; Upchurch, Paul; Goin, Francisco J.; Goswami, Anjali (6 February 2018). <a rel="nofollow" class="external text" href="https://www.cambridge.org/core/journals/paleobiology/article/deep-time-diversity-of-metatherian-mammals-implications-for-evolutionary-history-and-fossilrecord-quality/79E914107366FFD593A831E0486E356C">"Deep time diversity of metatherian mammals: implications for evolutionary history and fossil-record quality"</a>. <i><a href="/wiki/Paleobiology_(journal)" title="Paleobiology (journal)">Paleobiology</a></i>. <b>44</b> (2): <span class="nowrap">171–</span>198. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2018Pbio...44..171B">2018Pbio...44..171B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1017%2Fpab.2017.34">10.1017/pab.2017.34</a></span>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/11336%2F94590">11336/94590</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0094-8373">0094-8373</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:46796692">46796692</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Paleobiology&rft.atitle=Deep+time+diversity+of+metatherian+mammals%3A+implications+for+evolutionary+history+and+fossil-record+quality&rft.volume=44&rft.issue=2&rft.pages=%3Cspan+class%3D%22nowrap%22%3E171-%3C%2Fspan%3E198&rft.date=2018-02-06&rft_id=info%3Ahdl%2F11336%2F94590&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A46796692%23id-name%3DS2CID&rft_id=info%3Abibcode%2F2018Pbio...44..171B&rft.issn=0094-8373&rft_id=info%3Adoi%2F10.1017%2Fpab.2017.34&rft.aulast=Bennett&rft.aufirst=C.+Verity&rft.au=Upchurch%2C+Paul&rft.au=Goin%2C+Francisco+J.&rft.au=Goswami%2C+Anjali&rft_id=https%3A%2F%2Fwww.cambridge.org%2Fcore%2Fjournals%2Fpaleobiology%2Farticle%2Fdeep-time-diversity-of-metatherian-mammals-implications-for-evolutionary-history-and-fossilrecord-quality%2F79E914107366FFD593A831E0486E356C&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-:1-11"><span class="mw-cite-backlink">^ <a href="#cite_ref-:1_11-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:1_11-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:1_11-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-:1_11-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFEldridgeBeckCroftTravouillon2019" class="citation journal cs1">Eldridge, Mark D B; Beck, Robin M D; Croft, Darin A; Travouillon, Kenny J; Fox, Barry J (23 May 2019). <a rel="nofollow" class="external text" href="https://academic.oup.com/jmammal/article/100/3/802/5498022">"An emerging consensus in the evolution, phylogeny, and systematics of marsupials and their fossil relatives (Metatheria)"</a>. <i><a href="/wiki/Journal_of_Mammalogy" title="Journal of Mammalogy">Journal of Mammalogy</a></i>. <b>100</b> (3): <span class="nowrap">802–</span>837. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fjmammal%2Fgyz018">10.1093/jmammal/gyz018</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0022-2372">0022-2372</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Mammalogy&rft.atitle=An+emerging+consensus+in+the+evolution%2C+phylogeny%2C+and+systematics+of+marsupials+and+their+fossil+relatives+%28Metatheria%29&rft.volume=100&rft.issue=3&rft.pages=%3Cspan+class%3D%22nowrap%22%3E802-%3C%2Fspan%3E837&rft.date=2019-05-23&rft_id=info%3Adoi%2F10.1093%2Fjmammal%2Fgyz018&rft.issn=0022-2372&rft.aulast=Eldridge&rft.aufirst=Mark+D+B&rft.au=Beck%2C+Robin+M+D&rft.au=Croft%2C+Darin+A&rft.au=Travouillon%2C+Kenny+J&rft.au=Fox%2C+Barry+J&rft_id=https%3A%2F%2Facademic.oup.com%2Fjmammal%2Farticle%2F100%2F3%2F802%2F5498022&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFurióRuiz-SánchezCrespoFreudenthal2012" class="citation journal cs1">Furió, Marc; Ruiz-Sánchez, Francisco J.; Crespo, Vicente D.; Freudenthal, Matthijs; Montoya, Plinio (July 2012). <a rel="nofollow" class="external text" href="https://www.sciencedirect.com/science/article/pii/S1631068312000504?via%3Dihub">"The southernmost Miocene occurrence of the last European herpetotheriid Amphiperatherium frequens (Metatheria, Mammalia)"</a>. <i><a href="/wiki/Comptes_Rendus_Palevol" class="mw-redirect" title="Comptes Rendus Palevol">Comptes Rendus Palevol</a></i>. <b>11</b> (5): <span class="nowrap">371–</span>377. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012CRPal..11..371F">2012CRPal..11..371F</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.crpv.2012.01.004">10.1016/j.crpv.2012.01.004</a><span class="reference-accessdate">. Retrieved <span class="nowrap">26 October</span> 2024</span> – via Elsevier Science Direct.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Comptes+Rendus+Palevol&rft.atitle=The+southernmost+Miocene+occurrence+of+the+last+European+herpetotheriid+Amphiperatherium+frequens+%28Metatheria%2C+Mammalia%29&rft.volume=11&rft.issue=5&rft.pages=%3Cspan+class%3D%22nowrap%22%3E371-%3C%2Fspan%3E377&rft.date=2012-07&rft_id=info%3Adoi%2F10.1016%2Fj.crpv.2012.01.004&rft_id=info%3Abibcode%2F2012CRPal..11..371F&rft.aulast=Furi%C3%B3&rft.aufirst=Marc&rft.au=Ruiz-S%C3%A1nchez%2C+Francisco+J.&rft.au=Crespo%2C+Vicente+D.&rft.au=Freudenthal%2C+Matthijs&rft.au=Montoya%2C+Plinio&rft_id=https%3A%2F%2Fwww.sciencedirect.com%2Fscience%2Farticle%2Fpii%2FS1631068312000504%3Fvia%253Dihub&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCrespoGoin2021" class="citation journal cs1">Crespo, Vicente D.; Goin, Francisco J. (21 June 2021). <a rel="nofollow" class="external text" href="https://ojs.uv.es/index.php/sjpalaeontology/article/view/20974">"Taxonomy and Affinities of African Cenozoic Metatherians"</a>. <i>Spanish Journal of Palaeontology</i>. <b>36</b> (2). <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.7203%2Fsjp.36.2.20974">10.7203/sjp.36.2.20974</a></span>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/11336%2F165007">11336/165007</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/2255-0550">2255-0550</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:237387495">237387495</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Spanish+Journal+of+Palaeontology&rft.atitle=Taxonomy+and+Affinities+of+African+Cenozoic+Metatherians&rft.volume=36&rft.issue=2&rft.date=2021-06-21&rft_id=info%3Ahdl%2F11336%2F165007&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A237387495%23id-name%3DS2CID&rft.issn=2255-0550&rft_id=info%3Adoi%2F10.7203%2Fsjp.36.2.20974&rft.aulast=Crespo&rft.aufirst=Vicente+D.&rft.au=Goin%2C+Francisco+J.&rft_id=https%3A%2F%2Fojs.uv.es%2Findex.php%2Fsjpalaeontology%2Farticle%2Fview%2F20974&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCrespoGoinPickford2022" class="citation journal cs1">Crespo, Vicente D.; Goin, Francisco J.; Pickford, Martin (3 June 2022). <a rel="nofollow" class="external text" href="https://fr.pensoft.net/article/80706/">"The last African metatherian"</a>. <i><a href="/wiki/Fossil_Record" title="Fossil Record">Fossil Record</a></i>. <b>25</b> (1): <span class="nowrap">173–</span>186. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3897%2Ffr.25.80706">10.3897/fr.25.80706</a></span>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/10362%2F151025">10362/151025</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/2193-0074">2193-0074</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:249349445">249349445</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Fossil+Record&rft.atitle=The+last+African+metatherian&rft.volume=25&rft.issue=1&rft.pages=%3Cspan+class%3D%22nowrap%22%3E173-%3C%2Fspan%3E186&rft.date=2022-06-03&rft_id=info%3Ahdl%2F10362%2F151025&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A249349445%23id-name%3DS2CID&rft.issn=2193-0074&rft_id=info%3Adoi%2F10.3897%2Ffr.25.80706&rft.aulast=Crespo&rft.aufirst=Vicente+D.&rft.au=Goin%2C+Francisco+J.&rft.au=Pickford%2C+Martin&rft_id=https%3A%2F%2Ffr.pensoft.net%2Farticle%2F80706%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-wilson2016-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-wilson2016_15-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilsonEkdaleHogansonCalede2016" class="citation journal cs1">Wilson, G.P.; Ekdale, E.G.; Hoganson, J.W.; Calede, J.J.; Linden, A.V. (December 2016). <a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/311523654_A_large_carnivorous_mammal_from_the_Late_Cretaceous_and_the_North_American_origin_of_marsupials">"A large carnivorous mammal from the Late Cretaceous and the North American origin of marsupials"</a>. <i><a href="/wiki/Nature_Communications" title="Nature Communications">Nature Communications</a></i>. <b>7</b>: 13734. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2016NatCo...713734W">2016NatCo...713734W</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fncomms13734">10.1038/ncomms13734</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5155139">5155139</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/27929063">27929063</a><span class="reference-accessdate">. Retrieved <span class="nowrap">26 October</span> 2024</span> – via ResearchGate.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature+Communications&rft.atitle=A+large+carnivorous+mammal+from+the+Late+Cretaceous+and+the+North+American+origin+of+marsupials&rft.volume=7&rft.pages=13734&rft.date=2016-12&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5155139%23id-name%3DPMC&rft_id=info%3Apmid%2F27929063&rft_id=info%3Adoi%2F10.1038%2Fncomms13734&rft_id=info%3Abibcode%2F2016NatCo...713734W&rft.aulast=Wilson&rft.aufirst=G.P.&rft.au=Ekdale%2C+E.G.&rft.au=Hoganson%2C+J.W.&rft.au=Calede%2C+J.J.&rft.au=Linden%2C+A.V.&rft_id=https%3A%2F%2Fwww.researchgate.net%2Fpublication%2F311523654_A_large_carnivorous_mammal_from_the_Late_Cretaceous_and_the_North_American_origin_of_marsupials&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> <li id="cite_note-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-16">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLadevèzeSelvade_Muizon2020" class="citation journal cs1">Ladevèze, Sandrine; Selva, Charlène; de Muizon, Christian (1 September 2020). <a rel="nofollow" class="external text" href="https://www.tandfonline.com/doi/full/10.1080/14772019.2020.1772387">"What are "opossum-like" fossils? The phylogeny of herpetotheriid and peradectid metatherians, based on new features from the petrosal anatomy"</a>. <i><a href="/wiki/Journal_of_Systematic_Palaeontology" title="Journal of Systematic Palaeontology">Journal of Systematic Palaeontology</a></i>. <b>18</b> (17): <span class="nowrap">1463–</span>1479. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1080%2F14772019.2020.1772387">10.1080/14772019.2020.1772387</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1477-2019">1477-2019</a><span class="reference-accessdate">. Retrieved <span class="nowrap">26 October</span> 2024</span> – via Taylor and Francis Online.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Systematic+Palaeontology&rft.atitle=What+are+%E2%80%9Copossum-like%E2%80%9D+fossils%3F+The+phylogeny+of+herpetotheriid+and+peradectid+metatherians%2C+based+on+new+features+from+the+petrosal+anatomy&rft.volume=18&rft.issue=17&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1463-%3C%2Fspan%3E1479&rft.date=2020-09-01&rft_id=info%3Adoi%2F10.1080%2F14772019.2020.1772387&rft.issn=1477-2019&rft.aulast=Ladev%C3%A8ze&rft.aufirst=Sandrine&rft.au=Selva%2C+Charl%C3%A8ne&rft.au=de+Muizon%2C+Christian&rft_id=https%3A%2F%2Fwww.tandfonline.com%2Fdoi%2Ffull%2F10.1080%2F14772019.2020.1772387&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMetatheria" class="Z3988"></span></span> </li> </ol></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 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"}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Metatheria" title="Template:Metatheria"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Metatheria" title="Template talk:Metatheria"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Metatheria" title="Special:EditPage/Template:Metatheria"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Metatheria263" style="font-size:114%;margin:0 4em"><a class="mw-selflink selflink">Metatheria</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*_Kingdom:_Animalia_*_Phylum:_Chordata_*_Class:_Mammalia_*_Subclass:_Theria263" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size:85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Mammal" title="Mammal">Mammalia</a></li> <li>Subclass: <a href="/wiki/Theria" title="Theria">Theria</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Synapsida99" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg);;"><a href="/wiki/Synapsida" title="Synapsida">Synapsida</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Synapsida" title="Template:Synapsida">Synapsida</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cynodont" class="mw-redirect" title="Cynodont">Cynodontia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Cynodontia" title="Template:Cynodontia">Cynodontia</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Mammal" title="Mammal">Mammalia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Mammalia" title="Template:Mammalia">Mammalia</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a class="mw-selflink selflink">Metatheria</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Metatheria263" style="font-size:114%;margin:0 4em"><a class="mw-selflink selflink">Metatheria</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Metatheria100" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg);;"><a class="mw-selflink selflink">Metatheria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Adinodon&action=edit&redlink=1" class="new" title="Adinodon (page does not exist)">Adinodon</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Archaeonothos&action=edit&redlink=1" class="new" title="Archaeonothos (page does not exist)">Archaeonothos</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Esteslestes&action=edit&redlink=1" class="new" title="Esteslestes (page does not exist)">Esteslestes</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Ghamidtherium&action=edit&redlink=1" class="new" title="Ghamidtherium (page does not exist)">Ghamidtherium</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Holoclemensia" title="Holoclemensia">Holoclemensia</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Kasserinotherium&action=edit&redlink=1" class="new" title="Kasserinotherium (page does not exist)">Kasserinotherium</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Protalphadon" title="Protalphadon">Protalphadon</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Sinodelphys" title="Sinodelphys">Sinodelphys</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Deltatheroida" title="Deltatheroida">Deltatheroida</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Pappotherium" title="Pappotherium">Pappotherium</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Deltatheridiidae" title="Deltatheridiidae">Deltatheridiidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Atokatheridium" title="Atokatheridium">Atokatheridium</a></i>?</li> <li><i><a href="/wiki/Deltatheridium" title="Deltatheridium">Deltatheridium</a></i></li> <li><i><a href="/wiki/Deltatheroides" title="Deltatheroides">Deltatheroides</a></i></li> <li><i><a href="/wiki/Gurbanodelta" title="Gurbanodelta">Gurbanodelta</a></i></li> <li><i><a href="/wiki/Lotheridium" title="Lotheridium">Lotheridium</a></i></li> <li><i><a href="/wiki/Nanocuris" title="Nanocuris">Nanocuris</a></i></li> <li><i><a href="/wiki/Oklatheridium" title="Oklatheridium">Oklatheridium</a></i>?</li> <li><i><a href="/w/index.php?title=Prodeltitheridium&action=edit&redlink=1" class="new" title="Prodeltitheridium (page does not exist)">Prodeltitheridium</a></i></li> <li><i><a href="/wiki/Sulestes" title="Sulestes">Sulestes</a></i></li> <li><i><a href="/wiki/Tsagandelta" title="Tsagandelta">Tsagandelta</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg);;"><a href="/wiki/Marsupialiformes" class="mw-redirect" title="Marsupialiformes">Marsupialiformes</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Adelodelphys&action=edit&redlink=1" class="new" title="Adelodelphys (page does not exist)">Adelodelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Aenigmadelphys&action=edit&redlink=1" class="new" title="Aenigmadelphys (page does not exist)">Aenigmadelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Anchistodelphys&action=edit&redlink=1" class="new" title="Anchistodelphys (page does not exist)">Anchistodelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Andinodelphys" title="Andinodelphys">Andinodelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Arcantiodelphys" title="Arcantiodelphys">Arcantiodelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Camptomus&action=edit&redlink=1" class="new" title="Camptomus (page does not exist)">Camptomus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Glasbius&action=edit&redlink=1" class="new" title="Glasbius (page does not exist)">Glasbius</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Gurlin_Tsav_skull" title="Gurlin Tsav skull">Gurlin Tsav skull</a></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Iugomortiferum&action=edit&redlink=1" class="new" title="Iugomortiferum (page does not exist)">Iugomortiferum</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Jaskhadelphys&action=edit&redlink=1" class="new" title="Jaskhadelphys (page does not exist)">Jaskhadelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Mayulestes" title="Mayulestes">Mayulestes</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Pucadelphys" title="Pucadelphys">Pucadelphys</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Sinbadelphys&action=edit&redlink=1" class="new" title="Sinbadelphys (page does not exist)">Sinbadelphys</a></i></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Sparassodonta" title="Sparassodonta">Sparassodonta</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Asiatheriidae" class="mw-redirect" title="Asiatheriidae">Asiatheriidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Asiatherium" title="Asiatherium">Asiatherium</a></i></li> <li><i><a href="/w/index.php?title=Kokopellia&action=edit&redlink=1" class="new" title="Kokopellia (page does not exist)">Kokopellia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/w/index.php?title=Archimetatheria&action=edit&redlink=1" class="new" title="Archimetatheria (page does not exist)">Archimetatheria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Iqualadelphis&action=edit&redlink=1" class="new" title="Iqualadelphis (page does not exist)">Iqualadelphis</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Stagodontidae" title="Stagodontidae">Stagodontidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Didelphodon" title="Didelphodon">Didelphodon</a></i></li> <li><i><a href="/w/index.php?title=Eobrasilia&action=edit&redlink=1" class="new" title="Eobrasilia (page does not exist)">Eobrasilia</a></i>?</li> <li><i><a href="/wiki/Eodelphis" title="Eodelphis">Eodelphis</a></i></li> <li><i><a href="/w/index.php?title=Fumodelphodon&action=edit&redlink=1" class="new" title="Fumodelphodon (page does not exist)">Fumodelphodon</a></i></li> <li><i><a href="/w/index.php?title=Hoodootherium&action=edit&redlink=1" class="new" title="Hoodootherium (page does not exist)">Hoodootherium</a></i></li> <li><i><a href="/w/index.php?title=Pariadens&action=edit&redlink=1" class="new" title="Pariadens (page does not exist)">Pariadens</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Pediomyoidea&action=edit&redlink=1" class="new" title="Pediomyoidea (page does not exist)">Pediomyoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Aquiladelphis&action=edit&redlink=1" class="new" title="Aquiladelphis (page does not exist)">Aquiladelphis</a></i></li> <li><i><a href="/w/index.php?title=Austropediomys&action=edit&redlink=1" class="new" title="Austropediomys (page does not exist)">Austropediomys</a></i></li> <li><i><a href="/w/index.php?title=Dakotadens&action=edit&redlink=1" class="new" title="Dakotadens (page does not exist)">Dakotadens</a></i></li> <li><i><a href="/w/index.php?title=Monodelphopsis&action=edit&redlink=1" class="new" title="Monodelphopsis (page does not exist)">Monodelphopsis</a></i></li> <li><i><a href="/wiki/Pediomys" title="Pediomys">Pediomys</a></i></li> <li><i><a href="/w/index.php?title=Protolambda&action=edit&redlink=1" class="new" title="Protolambda (page does not exist)">Protolambda</a></i></li> <li><i><a href="/wiki/Scalaridelphys" title="Scalaridelphys">Scalaridelphys</a></i></li> <li><i><a href="/wiki/Unnuakomys" title="Unnuakomys">Unnuakomys</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Peradectidae" title="Peradectidae">Peradectidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Armintodelphys&action=edit&redlink=1" class="new" title="Armintodelphys (page does not exist)">Armintodelphys</a></i></li> <li><i><a href="/w/index.php?title=Didelphidectes&action=edit&redlink=1" class="new" title="Didelphidectes (page does not exist)">Didelphidectes</a></i></li> <li><i><a href="/wiki/Mimoperadectes" title="Mimoperadectes">Mimoperadectes</a></i></li> <li><i><a href="/w/index.php?title=Nanodelphys&action=edit&redlink=1" class="new" title="Nanodelphys (page does not exist)">Nanodelphys</a></i></li> <li><i><a href="/wiki/Peradectes" title="Peradectes">Peradectes</a></i></li> <li><i><a href="/wiki/Siamoperadectes" title="Siamoperadectes">Siamoperadectes</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Alphadontidae" title="Alphadontidae">Alphadontidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Albertatherium" title="Albertatherium">Albertatherium</a></i></li> <li><i><a href="/wiki/Alphadon" title="Alphadon">Alphadon</a></i></li> <li><i><a href="/w/index.php?title=Eoalphadon&action=edit&redlink=1" class="new" title="Eoalphadon (page does not exist)">Eoalphadon</a></i></li> <li><i><a href="/wiki/Turgidodon" title="Turgidodon">Turgidodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Herpetotheriidae" title="Herpetotheriidae">Herpetotheriidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amphiperatherium" title="Amphiperatherium">Amphiperatherium</a></i></li> <li><i><a href="/w/index.php?title=Asiadidelphis&action=edit&redlink=1" class="new" title="Asiadidelphis (page does not exist)">Asiadidelphis</a></i></li> <li><i><a href="/w/index.php?title=Copedelphys&action=edit&redlink=1" class="new" title="Copedelphys (page does not exist)">Copedelphys</a></i></li> <li><i><a href="/w/index.php?title=Ectocentrocristus&action=edit&redlink=1" class="new" title="Ectocentrocristus (page does not exist)">Ectocentrocristus</a></i></li> <li><i><a href="/w/index.php?title=Entomacodon&action=edit&redlink=1" class="new" title="Entomacodon (page does not exist)">Entomacodon</a></i></li> <li><i><a href="/wiki/Galatiadelphys" title="Galatiadelphys">Galatiadelphys</a></i></li> <li><i><a href="/w/index.php?title=Golerdelphys&action=edit&redlink=1" class="new" title="Golerdelphys (page does not exist)">Golerdelphys</a></i></li> <li><i><a href="/wiki/Herpetotherium" title="Herpetotherium">Herpetotherium</a></i></li> <li><i><a href="/wiki/Indodelphis" title="Indodelphis">Indodelphis</a></i></li> <li><i><a href="/wiki/Maastrichtidelphys" title="Maastrichtidelphys">Maastrichtidelphys</a></i></li> <li><i><a href="/w/index.php?title=Morotodon&action=edit&redlink=1" class="new" title="Morotodon (page does not exist)">Morotodon</a></i>?</li> <li><i><a href="/w/index.php?title=Nortedelphys&action=edit&redlink=1" class="new" title="Nortedelphys (page does not exist)">Nortedelphys</a></i></li> <li><i><a href="/wiki/Peratherium" title="Peratherium">Peratherium</a></i></li> <li><i><a href="/w/index.php?title=Rumiodon&action=edit&redlink=1" class="new" title="Rumiodon (page does not exist)">Rumiodon</a></i>?</li> <li><i><a href="/w/index.php?title=Swaindelphys&action=edit&redlink=1" class="new" title="Swaindelphys (page does not exist)">Swaindelphys</a></i></li> <li><i><a href="/wiki/Thylacodon" class="mw-redirect" title="Thylacodon">Thylacodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Anatoliadelphyidae" title="Anatoliadelphyidae">Anatoliadelphyidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Anatoliadelphys" title="Anatoliadelphys">Anatoliadelphys</a></i></li> <li><i><a href="/wiki/Orhaniyeia" title="Orhaniyeia">Orhaniyeia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Polydolopimorphia" title="Polydolopimorphia">Polydolopimorphia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marsupial" title="Marsupial">Marsupialia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b><a href="/wiki/Ameridelphia" title="Ameridelphia">Ameridelphia</a></b> <ul><li><b><a href="/wiki/Opossum" title="Opossum">Didelphimorphia</a></b></li> <li><b><a href="/wiki/Paucituberculata" title="Paucituberculata">Paucituberculata</a></b></li></ul></li> <li><b><a href="/wiki/Australidelphia" title="Australidelphia">Australidelphia</a></b></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Didelphodon_NT_small.jpg" class="mw-file-description" title="Didelphodon vorax"><img alt="Didelphodon vorax" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/Didelphodon_NT_small.jpg/140px-Didelphodon_NT_small.jpg" decoding="async" width="140" height="67" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/Didelphodon_NT_small.jpg/210px-Didelphodon_NT_small.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/97/Didelphodon_NT_small.jpg/280px-Didelphodon_NT_small.jpg 2x" data-file-width="902" data-file-height="430" /></a></span> <p><span typeof="mw:File"><a href="/wiki/File:Alphadon_sp._-_MUSE.jpg" class="mw-file-description" title="Alphadon sp."><img alt="Alphadon sp." src="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/Alphadon_sp._-_MUSE.jpg/140px-Alphadon_sp._-_MUSE.jpg" decoding="async" width="140" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/37/Alphadon_sp._-_MUSE.jpg/210px-Alphadon_sp._-_MUSE.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/37/Alphadon_sp._-_MUSE.jpg/280px-Alphadon_sp._-_MUSE.jpg 2x" data-file-width="3200" data-file-height="1800" /></a></span> </p> <span typeof="mw:File"><a href="/wiki/File:Herpetotherium_NT.jpg" class="mw-file-description" title="Herpetotherium fugax"><img alt="Herpetotherium fugax" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8c/Herpetotherium_NT.jpg/140px-Herpetotherium_NT.jpg" decoding="async" width="140" height="69" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8c/Herpetotherium_NT.jpg/210px-Herpetotherium_NT.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8c/Herpetotherium_NT.jpg/280px-Herpetotherium_NT.jpg 2x" data-file-width="500" data-file-height="245" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Sparassodonta263" style="font-size:114%;margin:0 4em"><a href="/wiki/Sparassodonta" title="Sparassodonta">Sparassodonta</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Sparassodonta103" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg);;"><a href="/wiki/Sparassodonta" title="Sparassodonta">Sparassodonta</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Allqokirus&action=edit&redlink=1" class="new" title="Allqokirus (page does not exist)">Allqokirus</a></i></li> <li><i><a href="/w/index.php?title=Argyrolestes&action=edit&redlink=1" class="new" title="Argyrolestes (page does not exist)">Argyrolestes</a></i></li> <li><i><a href="/wiki/Hondadelphys" title="Hondadelphys">Hondadelphys</a></i></li> <li><i><a href="/w/index.php?title=Nemolestes&action=edit&redlink=1" class="new" title="Nemolestes (page does not exist)">Nemolestes</a></i></li> <li><i><a href="/w/index.php?title=Patene&action=edit&redlink=1" class="new" title="Patene (page does not exist)">Patene</a></i></li> <li><i><a href="/w/index.php?title=Procladosictis&action=edit&redlink=1" class="new" title="Procladosictis (page does not exist)">Procladosictis</a></i></li> <li><i><a href="/w/index.php?title=Stylocynus&action=edit&redlink=1" class="new" title="Stylocynus (page does not exist)">Stylocynus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Hathliacynidae&action=edit&redlink=1" class="new" title="Hathliacynidae (page does not exist)">Hathliacynidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Acyon&action=edit&redlink=1" class="new" title="Acyon (page does not exist)">Acyon</a></i></li> <li><i><a href="/w/index.php?title=Australogale&action=edit&redlink=1" class="new" title="Australogale (page does not exist)">Australogale</a></i></li> <li><i><a href="/w/index.php?title=Borhyaenidium&action=edit&redlink=1" class="new" title="Borhyaenidium (page does not exist)">Borhyaenidium</a></i></li> <li><i><a href="/w/index.php?title=Chasicostylus&action=edit&redlink=1" class="new" title="Chasicostylus (page does not exist)">Chasicostylus</a></i></li> <li><i><a href="/wiki/Cladosictis" title="Cladosictis">Cladosictis</a></i></li> <li><i><a href="/w/index.php?title=Notictis&action=edit&redlink=1" class="new" title="Notictis (page does not exist)">Notictis</a></i></li> <li><i><a href="/w/index.php?title=Notocynus&action=edit&redlink=1" class="new" title="Notocynus (page does not exist)">Notocynus</a></i></li> <li><i><a href="/w/index.php?title=Notogale&action=edit&redlink=1" class="new" title="Notogale (page does not exist)">Notogale</a></i></li> <li><i><a href="/w/index.php?title=Perathereutes&action=edit&redlink=1" class="new" title="Perathereutes (page does not exist)">Perathereutes</a></i></li> <li><i><a href="/w/index.php?title=Pseudonotictis&action=edit&redlink=1" class="new" title="Pseudonotictis (page does not exist)">Pseudonotictis</a></i></li> <li><i><a href="/w/index.php?title=Sallacyon&action=edit&redlink=1" class="new" title="Sallacyon (page does not exist)">Sallacyon</a></i></li> <li><i><a href="/w/index.php?title=Sipalocyon&action=edit&redlink=1" class="new" title="Sipalocyon (page does not exist)">Sipalocyon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Borhyaenoidea" class="mw-redirect" title="Borhyaenoidea">Borhyaenoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Angelocabrerus&action=edit&redlink=1" class="new" title="Angelocabrerus (page does not exist)">Angelocabrerus</a></i></li> <li><i><a href="/w/index.php?title=Chlorocyon&action=edit&redlink=1" class="new" title="Chlorocyon (page does not exist)">Chlorocyon</a></i></li> <li><i><a href="/wiki/Dukecynus" title="Dukecynus">Dukecynus</a></i></li> <li><i><a href="/wiki/Eomakhaira" title="Eomakhaira">Eomakhaira</a></i></li> <li><i><a href="/wiki/Fredszalaya" title="Fredszalaya">Fredszalaya</a></i></li> <li><i><a href="/wiki/Lycopsis" title="Lycopsis">Lycopsis</a></i></li> <li><i><a href="/wiki/Pharsophorus" title="Pharsophorus">Pharsophorus</a></i></li> <li><i><a href="/w/index.php?title=Plesiofelis&action=edit&redlink=1" class="new" title="Plesiofelis (page does not exist)">Plesiofelis</a></i></li> <li><i><a href="/wiki/Prothylacinus" title="Prothylacinus">Prothylacinus</a></i></li> <li><i><a href="/w/index.php?title=Pseudolycopsis&action=edit&redlink=1" class="new" title="Pseudolycopsis (page does not exist)">Pseudolycopsis</a></i></li> <li><i><a href="/w/index.php?title=Pseudothylacynus&action=edit&redlink=1" class="new" title="Pseudothylacynus (page does not exist)">Pseudothylacynus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Borhyaenidae" title="Borhyaenidae">Borhyaenidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Acrocyon&action=edit&redlink=1" class="new" title="Acrocyon (page does not exist)">Acrocyon</a></i></li> <li><i><a href="/w/index.php?title=Arctodictis&action=edit&redlink=1" class="new" title="Arctodictis (page does not exist)">Arctodictis</a></i></li> <li><i><a href="/wiki/Australohyaena" title="Australohyaena">Australohyaena</a></i></li> <li><i><a href="/wiki/Borhyaena" title="Borhyaena">Borhyaena</a></i></li> <li><i><a href="/w/index.php?title=Eutemnodus&action=edit&redlink=1" class="new" title="Eutemnodus (page does not exist)">Eutemnodus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Proborhyaenidae" title="Proborhyaenidae">Proborhyaenidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Arminiheringia" title="Arminiheringia">Arminiheringia</a></i></li> <li><i><a href="/wiki/Callistoe" title="Callistoe">Callistoe</a></i></li> <li><i><a href="/wiki/Paraborhyaena" title="Paraborhyaena">Paraborhyaena</a></i></li> <li><i><a href="/wiki/Proborhyaena" title="Proborhyaena">Proborhyaena</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Thylacosmiliformes" title="Thylacosmiliformes">Thylacosmiliformes</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dimartinia" title="Dimartinia">Dimartinia</a></i></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Thylacosmilidae19" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Thylacosmilidae" title="Thylacosmilidae">Thylacosmilidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Anachlysictis" title="Anachlysictis">Anachlysictis</a></i></li> <li><i><a href="/wiki/Patagosmilus" title="Patagosmilus">Patagosmilus</a></i></li> <li><i><a href="/wiki/Thylacosmilus" title="Thylacosmilus">Thylacosmilus</a></i></li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Thylacosmilus_NT_small.jpg" class="mw-file-description" title="Thylacosmilus atrox"><img alt="Thylacosmilus atrox" src="//upload.wikimedia.org/wikipedia/commons/thumb/b/b4/Thylacosmilus_NT_small.jpg/140px-Thylacosmilus_NT_small.jpg" decoding="async" width="140" height="78" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/b4/Thylacosmilus_NT_small.jpg/210px-Thylacosmilus_NT_small.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/b4/Thylacosmilus_NT_small.jpg/280px-Thylacosmilus_NT_small.jpg 2x" data-file-width="949" data-file-height="528" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Polydolopimorphia263" style="font-size:114%;margin:0 4em"><a href="/wiki/Polydolopimorphia" title="Polydolopimorphia">Polydolopimorphia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Polydolopimorphia107" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg);;"><a href="/wiki/Polydolopimorphia" title="Polydolopimorphia">Polydolopimorphia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Apeirodon" title="Apeirodon">Apeirodon</a></i>?</li> <li><i><a href="/w/index.php?title=Bobbschaefferia&action=edit&redlink=1" class="new" title="Bobbschaefferia (page does not exist)">Bobbschaefferia</a></i></li> <li><i><a href="/w/index.php?title=Caroloameghinia&action=edit&redlink=1" class="new" title="Caroloameghinia (page does not exist)">Caroloameghinia</a></i></li> <li><i><a href="/w/index.php?title=Carolocoutoia&action=edit&redlink=1" class="new" title="Carolocoutoia (page does not exist)">Carolocoutoia</a></i></li> <li><i><a href="/w/index.php?title=Guggenheimia&action=edit&redlink=1" class="new" title="Guggenheimia (page does not exist)">Guggenheimia</a></i></li> <li><i><a href="/w/index.php?title=Gashternia&action=edit&redlink=1" class="new" title="Gashternia (page does not exist)">Gashternia</a></i></li> <li><i><a href="/w/index.php?title=Hatcheritherium&action=edit&redlink=1" class="new" title="Hatcheritherium (page does not exist)">Hatcheritherium</a></i></li> <li><i><a href="/w/index.php?title=Palangania&action=edit&redlink=1" class="new" title="Palangania (page does not exist)">Palangania</a></i></li> <li><i><a href="/w/index.php?title=Procaroloameghinia&action=edit&redlink=1" class="new" title="Procaroloameghinia (page does not exist)">Procaroloameghinia</a></i></li> <li><i><a href="/w/index.php?title=Protodidelphis&action=edit&redlink=1" class="new" title="Protodidelphis (page does not exist)">Protodidelphis</a></i></li> <li><i><a href="/w/index.php?title=Pujatodon&action=edit&redlink=1" class="new" title="Pujatodon (page does not exist)">Pujatodon</a></i>?</li> <li><i><a href="/w/index.php?title=Reigia&action=edit&redlink=1" class="new" title="Reigia (page does not exist)">Reigia</a></i></li> <li><i><a href="/w/index.php?title=Robertbutleria&action=edit&redlink=1" class="new" title="Robertbutleria (page does not exist)">Robertbutleria</a></i></li> <li><i><a href="/w/index.php?title=Wamradolops&action=edit&redlink=1" class="new" title="Wamradolops (page does not exist)">Wamradolops</a></i></li> <li><i><a href="/w/index.php?title=Zeusdelphys&action=edit&redlink=1" class="new" title="Zeusdelphys (page does not exist)">Zeusdelphys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Bonapartheriiformes&action=edit&redlink=1" class="new" title="Bonapartheriiformes (page does not exist)">Bonapartheriiformes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Bonapartheriidae&action=edit&redlink=1" class="new" title="Bonapartheriidae (page does not exist)">Bonapartheriidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bonapartherium" title="Bonapartherium">Bonapartherium</a></i></li> <li><i><a href="/wiki/Epidolops" title="Epidolops">Epidolops</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Argyrolagidae&action=edit&redlink=1" class="new" title="Argyrolagidae (page does not exist)">Argyrolagidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Anargyrolagus&action=edit&redlink=1" class="new" title="Anargyrolagus (page does not exist)">Anargyrolagus</a></i></li> <li><i><a href="/wiki/Argyrolagus" title="Argyrolagus">Argyrolagus</a></i></li> <li><i><a href="/wiki/Hondalagus" title="Hondalagus">Hondalagus</a></i></li> <li><i><a href="/w/index.php?title=Microtragulus&action=edit&redlink=1" class="new" title="Microtragulus (page does not exist)">Microtragulus</a></i></li> <li><i><a href="/w/index.php?title=Proargyrolagus&action=edit&redlink=1" class="new" title="Proargyrolagus (page does not exist)">Proargyrolagus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Chulpasiinae&action=edit&redlink=1" class="new" title="Chulpasiinae (page does not exist)">Chulpasiinae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chulpasia" title="Chulpasia">Chulpasia</a></i></li> <li><i><a href="/w/index.php?title=Thylacotinga&action=edit&redlink=1" class="new" title="Thylacotinga (page does not exist)">Thylacotinga</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Prepidolopidae&action=edit&redlink=1" class="new" title="Prepidolopidae (page does not exist)">Prepidolopidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Incadolops&action=edit&redlink=1" class="new" title="Incadolops (page does not exist)">Incadolops</a></i></li> <li><i><a href="/w/index.php?title=Perrodelphys&action=edit&redlink=1" class="new" title="Perrodelphys (page does not exist)">Perrodelphys</a></i></li> <li><i><a href="/w/index.php?title=Prepidolops&action=edit&redlink=1" class="new" title="Prepidolops (page does not exist)">Prepidolops</a></i></li> <li><i><a href="/w/index.php?title=Punadolops&action=edit&redlink=1" class="new" title="Punadolops (page does not exist)">Punadolops</a></i></li> <li><i><a href="/w/index.php?title=Seumandia&action=edit&redlink=1" class="new" title="Seumandia (page does not exist)">Seumandia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Rosendolopidae&action=edit&redlink=1" class="new" title="Rosendolopidae (page does not exist)">Rosendolopidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Hondonadia" title="Hondonadia">Hondonadia</a></i></li> <li><i><a href="/w/index.php?title=Rosendolops&action=edit&redlink=1" class="new" title="Rosendolops (page does not exist)">Rosendolops</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Polydolopiformes&action=edit&redlink=1" class="new" title="Polydolopiformes (page does not exist)">Polydolopiformes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Polydolopidae" class="mw-redirect" title="Polydolopidae">Polydolopidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Amphidolops&action=edit&redlink=1" class="new" title="Amphidolops (page does not exist)">Amphidolops</a></i></li> <li><i><a href="/wiki/Antarctodolops" title="Antarctodolops">Antarctodolops</a></i></li> <li><i><a href="/w/index.php?title=Eudolops&action=edit&redlink=1" class="new" title="Eudolops (page does not exist)">Eudolops</a></i></li> <li><i><a href="/w/index.php?title=Eurydolops&action=edit&redlink=1" class="new" title="Eurydolops (page does not exist)">Eurydolops</a></i></li> <li><i><a href="/w/index.php?title=Hypodolops&action=edit&redlink=1" class="new" title="Hypodolops (page does not exist)">Hypodolops</a></i></li> <li><i><a href="/w/index.php?title=Kramadolops&action=edit&redlink=1" class="new" title="Kramadolops (page does not exist)">Kramadolops</a></i></li> <li><i><a href="/w/index.php?title=Parabderites&action=edit&redlink=1" class="new" title="Parabderites (page does not exist)">Parabderites</a></i></li> <li><i><a href="/w/index.php?title=Polydolops&action=edit&redlink=1" class="new" title="Polydolops (page does not exist)">Polydolops</a></i></li> <li><i><a href="/w/index.php?title=Pseudolops&action=edit&redlink=1" class="new" title="Pseudolops (page does not exist)">Pseudolops</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Sillustaniidae&action=edit&redlink=1" class="new" title="Sillustaniidae (page does not exist)">Sillustaniidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Roberthoffstetteria&action=edit&redlink=1" class="new" title="Roberthoffstetteria (page does not exist)">Roberthoffstetteria</a></i></li> <li><i><a href="/w/index.php?title=Sillustania&action=edit&redlink=1" class="new" title="Sillustania (page does not exist)">Sillustania</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Argyrolagus_NT.jpg" class="mw-file-description" title="Argyrolagus palmeri"><img alt="Argyrolagus palmeri" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f9/Argyrolagus_NT.jpg/140px-Argyrolagus_NT.jpg" decoding="async" width="140" height="105" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f9/Argyrolagus_NT.jpg/210px-Argyrolagus_NT.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f9/Argyrolagus_NT.jpg/280px-Argyrolagus_NT.jpg 2x" data-file-width="800" data-file-height="600" /></a></span></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers1821" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers1821" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Metatheria</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q1096960" class="extiw" title="wikidata:Q1096960">Q1096960</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Metatheria" class="extiw" title="wikispecies:Metatheria">Metatheria</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Animal_Diversity_Web" title="Animal Diversity Web">ADW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://animaldiversity.org/accounts/Metatheria/">Metatheria</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/2844108">2844108</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:faunaeur.org:taxname:12614">12614</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea (new)</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://fauna-eu.org/cdm_dataportal/taxon/26226554-59ee-472b-84eb-87709ad0d830">26226554-59ee-472b-84eb-87709ad0d830</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=179917">179917</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=9263">9263</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=16033">16033</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=39936">39936</a></span></span></li></ul> </div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐5558bc8b6‐zqrbx Cached time: 20250210165440 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 0.898 seconds Real time usage: 1.099 seconds Preprocessor visited node count: 18598/1000000 Post‐expand include size: 452870/2097152 bytes Template argument size: 30314/2097152 bytes Highest expansion depth: 24/100 Expensive parser function count: 11/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 164583/5000000 bytes Lua time usage: 0.578/10.000 seconds Lua memory usage: 7312225/52428800 bytes Number of Wikibase entities loaded: 10/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 953.920 1 -total 60.24% 574.603 32 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