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for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Har Gobind Khorana</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 71 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-71" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">71 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1_%D8%BA%D9%88%D8%A8%D9%86%D8%AF_%D8%AE%D9%88%D8%B1%D8%A7%D9%86%D8%A7" title="هار غوبند خورانا – Arabic" lang="ar" hreflang="ar" data-title="هار غوبند خورانا" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-awa mw-list-item"><a href="https://awa.wikipedia.org/wiki/%E0%A4%B9%E0%A4%B0%E0%A4%97%E0%A5%8B%E0%A4%B5%E0%A4%BF%E0%A4%A8%E0%A5%8D%E0%A4%A6_%E0%A4%96%E0%A5%81%E0%A4%B0%E0%A4%BE%E0%A4%A8%E0%A4%BE" title="हरगोविन्द खुराना – Awadhi" lang="awa" hreflang="awa" data-title="हरगोविन्द खुराना" data-language-autonym="अवधी" data-language-local-name="Awadhi" class="interlanguage-link-target"><span>अवधी</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1_%DA%AF%D9%88%D8%A8%DB%8C%D9%86%D8%AF_%DA%A9%D9%88%D8%B1%D8%A7%D9%86%D8%A7" title="هار گوبیند کورانا – South Azerbaijani" lang="azb" hreflang="azb" data-title="هار گوبیند کورانا" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B9%E0%A6%B0_%E0%A6%97%E0%A7%8B%E0%A6%AC%E0%A6%BF%E0%A6%A8%E0%A7%8D%E0%A6%A6_%E0%A6%96%E0%A7%8B%E0%A6%B0%E0%A6%BE%E0%A6%A8%E0%A6%BE" title="হর গোবিন্দ খোরানা – Bangla" lang="bn" hreflang="bn" data-title="হর গোবিন্দ খোরানা" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A5%D0%B0%D1%80_%D0%93%D0%BE%D0%B1%D1%96%D0%BD%D0%B4_%D0%9A%D0%B0%D1%80%D0%B0%D0%BD%D0%B0" title="Хар Гобінд Карана – Belarusian" lang="be" hreflang="be" data-title="Хар Гобінд Карана" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Catalan" lang="ca" hreflang="ca" data-title="Har Gobind Khorana" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Czech" lang="cs" hreflang="cs" data-title="Har Gobind Khorana" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Danish" lang="da" hreflang="da" data-title="Har Gobind Khorana" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – German" lang="de" hreflang="de" data-title="Har Gobind Khorana" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Estonian" lang="et" hreflang="et" data-title="Har Gobind Khorana" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A7%CE%B1%CF%81_%CE%93%CE%BA%CE%BF%CE%BC%CF%80%CE%AF%CE%BD%CF%84_%CE%A7%CE%BF%CF%81%CE%AC%CE%BD%CE%B1" title="Χαρ Γκομπίντ Χοράνα – Greek" lang="el" hreflang="el" data-title="Χαρ Γκομπίντ Χοράνα" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – English" lang="en" hreflang="en" data-title="Har Gobind Khorana" data-language-autonym="English" data-language-local-name="English" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Spanish" lang="es" hreflang="es" data-title="Har Gobind Khorana" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Esperanto" lang="eo" hreflang="eo" data-title="Har Gobind Khorana" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Basque" lang="eu" hreflang="eu" data-title="Har Gobind Khorana" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1_%DA%AF%D9%88%D8%A8%DB%8C%D9%86%D8%AF_%DA%A9%D9%88%D8%B1%D8%A7%D9%86%D8%A7" title="هار گوبیند کورانا – Persian" lang="fa" hreflang="fa" data-title="هار گوبیند کورانا" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – French" lang="fr" hreflang="fr" data-title="Har Gobind Khorana" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Irish" lang="ga" hreflang="ga" data-title="Har Gobind Khorana" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Galician" lang="gl" hreflang="gl" data-title="Har Gobind Khorana" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-gu mw-list-item"><a href="https://gu.wikipedia.org/wiki/%E0%AA%A1%E0%AB%8B._%E0%AA%B9%E0%AA%B0%E0%AA%97%E0%AB%8B%E0%AA%B5%E0%AA%BF%E0%AA%82%E0%AA%A6_%E0%AA%96%E0%AB%81%E0%AA%B0%E0%AA%BE%E0%AA%A8%E0%AA%BE" title="ડો. હરગોવિંદ ખુરાના – Gujarati" lang="gu" hreflang="gu" data-title="ડો. હરગોવિંદ ખુરાના" data-language-autonym="ગુજરાતી" data-language-local-name="Gujarati" class="interlanguage-link-target"><span>ગુજરાતી</span></a></li><li class="interlanguage-link interwiki-gom mw-list-item"><a href="https://gom.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Goan Konkani" lang="gom" hreflang="gom" data-title="Har Gobind Khorana" data-language-autonym="गोंयची कोंकणी / Gõychi Konknni" data-language-local-name="Goan Konkani" class="interlanguage-link-target"><span>गोंयची कोंकणी / Gõychi Konknni</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%95%98%EB%A5%B4_%EA%B3%A0%EB%B9%88%EB%93%9C_%EC%BD%94%EB%9D%BC%EB%82%98" title="하르 고빈드 코라나 – Korean" lang="ko" hreflang="ko" data-title="하르 고빈드 코라나" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B9%E0%A4%B0%E0%A4%97%E0%A5%8B%E0%A4%B5%E0%A4%BF%E0%A4%A8%E0%A5%8D%E0%A4%A6_%E0%A4%96%E0%A5%81%E0%A4%B0%E0%A4%BE%E0%A4%A8%E0%A4%BE" title="हरगोविन्द खुराना – Hindi" lang="hi" hreflang="hi" data-title="हरगोविन्द खुराना" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Croatian" lang="hr" hreflang="hr" data-title="Har Gobind Khorana" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Ido" lang="io" hreflang="io" data-title="Har Gobind Khorana" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Indonesian" lang="id" hreflang="id" data-title="Har Gobind Khorana" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Italian" lang="it" hreflang="it" data-title="Har Gobind Khorana" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%94%D7%A8_%D7%92%D7%95%D7%91%D7%99%D7%A0%D7%93_%D7%A7%D7%95%D7%A8%D7%90%D7%A0%D7%94" title="הר גובינד קוראנה – Hebrew" lang="he" hreflang="he" data-title="הר גובינד קוראנה" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%B9%E0%B2%B0%E0%B2%97%E0%B3%8B%E0%B2%AC%E0%B2%BF%E0%B2%82%E0%B2%A6_%E0%B2%96%E0%B3%81%E0%B2%B0%E0%B2%BE%E0%B2%A8" title="ಹರಗೋಬಿಂದ ಖುರಾನ – Kannada" lang="kn" hreflang="kn" data-title="ಹರಗೋಬಿಂದ ಖುರಾನ" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%B0%E1%83%90%E1%83%A0_%E1%83%92%E1%83%9D%E1%83%91%E1%83%98%E1%83%9C%E1%83%93_%E1%83%AE%E1%83%9D%E1%83%A0%E1%83%90%E1%83%9C%E1%83%90" title="ჰარ გობინდ ხორანა – Georgian" lang="ka" hreflang="ka" data-title="ჰარ გობინდ ხორანა" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A5%D0%B0%D1%80_%D0%93%D0%BE%D0%B1%D0%B8%D0%BD%D0%B4_%D0%9A%D0%BE%D1%80%D0%B0%D0%BD%D0%B0" title="Хар Гобинд Корана – Kazakh" lang="kk" hreflang="kk" data-title="Хар Гобинд Корана" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Har_Khorana" title="Har Khorana – Swahili" lang="sw" hreflang="sw" data-title="Har Khorana" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Hars_Gobinds_Korana" title="Hars Gobinds Korana – Latvian" lang="lv" hreflang="lv" data-title="Hars Gobinds Korana" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Hungarian" lang="hu" hreflang="hu" data-title="Har Gobind Khorana" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mai mw-list-item"><a href="https://mai.wikipedia.org/wiki/%E0%A4%B9%E0%A4%B0%E0%A4%97%E0%A5%8B%E0%A4%B5%E0%A4%BF%E0%A4%A8%E0%A5%8D%E0%A4%A6_%E0%A4%96%E0%A5%81%E0%A4%B0%E0%A4%BE%E0%A4%A8%E0%A4%BE" title="हरगोविन्द खुराना – Maithili" lang="mai" hreflang="mai" data-title="हरगोविन्द खुराना" data-language-autonym="मैथिली" data-language-local-name="Maithili" class="interlanguage-link-target"><span>मैथिली</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%B9%E0%B5%BC_%E0%B4%97%E0%B5%8B%E0%B4%B5%E0%B4%BF%E0%B4%A8%E0%B5%8D%E0%B4%A6%E0%B5%8D%E2%80%8C_%E0%B4%96%E0%B5%81%E0%B4%B0%E0%B4%BE%E0%B4%A8" title="ഹർ ഗോവിന്ദ്‌ ഖുരാന – Malayalam" lang="ml" hreflang="ml" data-title="ഹർ ഗോവിന്ദ്‌ ഖുരാന" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-mr mw-list-item"><a href="https://mr.wikipedia.org/wiki/%E0%A4%B9%E0%A4%B0%E0%A4%97%E0%A5%8B%E0%A4%B5%E0%A4%BF%E0%A4%82%E0%A4%A6_%E0%A4%96%E0%A5%81%E0%A4%B0%E0%A4%BE%E0%A4%A3%E0%A4%BE" title="हरगोविंद खुराणा – Marathi" lang="mr" hreflang="mr" data-title="हरगोविंद खुराणा" data-language-autonym="मराठी" data-language-local-name="Marathi" class="interlanguage-link-target"><span>मराठी</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%B0%E1%83%90%E1%83%A0_%E1%83%92%E1%83%9D%E1%83%91%E1%83%98%E1%83%9C%E1%83%93_%E1%83%99%E1%83%9D%E1%83%A0%E1%83%90%E1%83%9C%E1%83%90" title="ჰარ გობინდ კორანა – Mingrelian" lang="xmf" hreflang="xmf" data-title="ჰარ გობინდ კორანა" data-language-autonym="მარგალური" data-language-local-name="Mingrelian" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1_%D8%AC%D9%88%D8%A8%D9%86%D8%AF_%D8%AE%D9%88%D8%B1%D8%A7%D9%86%D8%A7" title="هار جوبند خورانا – Egyptian Arabic" lang="arz" hreflang="arz" data-title="هار جوبند خورانا" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1_%DA%AF%D9%88%D8%A8%DB%8C%D9%86%D8%AF_%DA%A9%D9%88%D8%B1%D8%A7%D9%86%D8%A7" title="هار گوبیند کورانا – Mazanderani" lang="mzn" hreflang="mzn" data-title="هار گوبیند کورانا" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Malay" lang="ms" hreflang="ms" data-title="Har Gobind Khorana" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Dutch" lang="nl" hreflang="nl" data-title="Har Gobind Khorana" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%8F%E3%83%BC%E3%83%BB%E3%82%B4%E3%83%93%E3%83%B3%E3%83%89%E3%83%BB%E3%82%B3%E3%83%A9%E3%83%8A" title="ハー・ゴビンド・コラナ – Japanese" lang="ja" hreflang="ja" data-title="ハー・ゴビンド・コラナ" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Har Gobind Khorana" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Norwegian Nynorsk" lang="nn" hreflang="nn" data-title="Har Gobind Khorana" data-language-autonym="Norsk nynorsk" data-language-local-name="Norwegian Nynorsk" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Occitan" lang="oc" hreflang="oc" data-title="Har Gobind Khorana" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-or mw-list-item"><a href="https://or.wikipedia.org/wiki/%E0%AC%B9%E0%AC%B0%E0%AC%97%E0%AD%8B%E0%AC%AC%E0%AC%BF%E0%AC%A8%E0%AD%8D%E0%AC%A6_%E0%AC%96%E0%AD%81%E0%AC%B0%E0%AC%BE%E0%AC%A8%E0%AC%BE" title="ହରଗୋବିନ୍ଦ ଖୁରାନା – Odia" lang="or" hreflang="or" data-title="ହରଗୋବିନ୍ଦ ଖୁରାନା" data-language-autonym="ଓଡ଼ିଆ" data-language-local-name="Odia" class="interlanguage-link-target"><span>ଓଡ଼ିଆ</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Uzbek" lang="uz" hreflang="uz" data-title="Har Gobind Khorana" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pa mw-list-item"><a href="https://pa.wikipedia.org/wiki/%E0%A8%B9%E0%A8%B0%E0%A8%97%E0%A9%8B%E0%A8%AC%E0%A8%BF%E0%A9%B0%E0%A8%A6_%E0%A8%96%E0%A9%81%E0%A8%B0%E0%A8%BE%E0%A8%A3%E0%A8%BE" title="ਹਰਗੋਬਿੰਦ ਖੁਰਾਣਾ – Punjabi" lang="pa" hreflang="pa" data-title="ਹਰਗੋਬਿੰਦ ਖੁਰਾਣਾ" data-language-autonym="ਪੰਜਾਬੀ" data-language-local-name="Punjabi" class="interlanguage-link-target"><span>ਪੰਜਾਬੀ</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DB%81%D8%B1_%DA%AF%D9%88%D8%A8%D9%86%D8%AF_%DA%A9%DA%BE%D9%8F%D8%B1%D8%A7%D9%86%D8%A7" title="ہر گوبند کھُرانا – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ہر گوبند کھُرانا" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Low German" lang="nds" hreflang="nds" data-title="Har Gobind Khorana" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Polish" lang="pl" hreflang="pl" data-title="Har Gobind Khorana" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Har_Khorana" title="Har Khorana – Portuguese" lang="pt" hreflang="pt" data-title="Har Khorana" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9A%D0%BE%D1%80%D0%B0%D0%BD%D0%B0,_%D0%A5%D0%B0%D1%80_%D0%93%D0%BE%D0%B1%D0%B8%D0%BD%D0%B4" title="Корана, Хар Гобинд – Russian" lang="ru" hreflang="ru" data-title="Корана, Хар Гобинд" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sa mw-list-item"><a href="https://sa.wikipedia.org/wiki/%E0%A4%B9%E0%A4%B0%E0%A4%97%E0%A5%8B%E0%A4%B5%E0%A4%BF%E0%A4%A8%E0%A5%8D%E0%A4%A6_%E0%A4%96%E0%A5%81%E0%A4%B0%E0%A4%BE%E0%A4%A8%E0%A4%BE" title="हरगोविन्द खुराना – Sanskrit" lang="sa" hreflang="sa" data-title="हरगोविन्द खुराना" data-language-autonym="संस्कृतम्" data-language-local-name="Sanskrit" class="interlanguage-link-target"><span>संस्कृतम्</span></a></li><li class="interlanguage-link interwiki-skr mw-list-item"><a href="https://skr.wikipedia.org/wiki/%DB%81%D8%B1%DA%AF%D9%88%D8%A8%D9%86%D8%AF_%DA%A9%DA%BE%D8%B1%D8%A7%DD%A8%DB%81" title="ہرگوبند کھراݨہ – Saraiki" lang="skr" hreflang="skr" data-title="ہرگوبند کھراݨہ" data-language-autonym="سرائیکی" data-language-local-name="Saraiki" class="interlanguage-link-target"><span>سرائیکی</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Scots" lang="sco" hreflang="sco" data-title="Har Gobind Khorana" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-sd mw-list-item"><a href="https://sd.wikipedia.org/wiki/%D9%87%D8%B1%D8%A8_%DA%AF%D9%88%D8%A8%D9%86%D8%AF_%DA%B3%D9%88%D8%B1%D8%A7%D9%86%D8%A7" title="هرب گوبند ڳورانا – Sindhi" lang="sd" hreflang="sd" data-title="هرب گوبند ڳورانا" data-language-autonym="سنڌي" data-language-local-name="Sindhi" class="interlanguage-link-target"><span>سنڌي</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%DA%BE%D8%A7%D8%B1_%DA%AF%DB%86%D8%A8%DB%8C%D8%AF_%D8%AE%D9%88%D8%B1%D8%A7%D9%86%D8%A7" title="ھار گۆبید خورانا – Central Kurdish" lang="ckb" hreflang="ckb" data-title="ھار گۆبید خورانا" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Har_Gobind_Korana" title="Har Gobind Korana – Serbian" lang="sr" hreflang="sr" data-title="Har Gobind Korana" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Har_Gobind_Korana" title="Har Gobind Korana – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Har Gobind Korana" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Finnish" lang="fi" hreflang="fi" data-title="Har Gobind Khorana" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Swedish" lang="sv" hreflang="sv" data-title="Har Gobind Khorana" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%B9%E0%AE%B0%E0%AF%8D_%E0%AE%95%E0%AF%8B%E0%AE%B5%E0%AE%BF%E0%AE%A8%E0%AF%8D%E0%AE%A4%E0%AF%8D_%E0%AE%95%E0%AF%8A%E0%AE%B0%E0%AE%BE%E0%AE%A9%E0%AE%BE" title="ஹர் கோவிந்த் கொரானா – Tamil" lang="ta" hreflang="ta" data-title="ஹர் கோவிந்த் கொரானா" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a href="https://te.wikipedia.org/wiki/%E0%B0%B9%E0%B0%B0%E0%B0%97%E0%B1%8B%E0%B0%B5%E0%B0%BF%E0%B0%82%E0%B0%A6%E0%B1%8D_%E0%B0%96%E0%B1%8A%E0%B0%B0%E0%B0%BE%E0%B0%A8%E0%B0%BE" title="హరగోవింద్ ఖొరానా – Telugu" lang="te" hreflang="te" data-title="హరగోవింద్ ఖొరానా" data-language-autonym="తెలుగు" data-language-local-name="Telugu" class="interlanguage-link-target"><span>తెలుగు</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Turkish" lang="tr" hreflang="tr" data-title="Har Gobind Khorana" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%93%D0%B0%D1%80_%D0%93%D0%BE%D0%B1%D1%96%D0%BD%D0%B4_%D0%9A%D0%BE%D1%80%D0%B0%D0%BD%D0%B0" title="Гар Гобінд Корана – Ukrainian" lang="uk" hreflang="uk" data-title="Гар Гобінд Корана" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%DB%81%D8%B1_%DA%AF%D9%88%D8%A8%D9%86%D8%AF_%DA%A9%DA%BE%D8%B1%D8%A7%D9%86%D8%A7" title="ہر گوبند کھرانا – Urdu" lang="ur" hreflang="ur" data-title="ہر گوبند کھرانا" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E5%93%88%E5%B0%94%C2%B7%E8%91%9B%E5%AE%BE%C2%B7%E7%A7%91%E6%8B%89%E7%BA%B3" title="哈尔·葛宾·科拉纳 – Wu" lang="wuu" hreflang="wuu" data-title="哈尔·葛宾·科拉纳" data-language-autonym="吴语" data-language-local-name="Wu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana – Yoruba" lang="yo" hreflang="yo" data-title="Har Gobind Khorana" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%93%88%E7%88%BE%C2%B7%E8%91%9B%E8%B3%93%C2%B7%E7%A7%91%E6%8B%89%E7%B4%8D" title="哈爾·葛賓·科拉納 – Chinese" lang="zh" hreflang="zh" data-title="哈爾·葛賓·科拉納" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q107462#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Change links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div 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.mw-parser-output .infobox-header,body.skin-minerva .mw-parser-output .infobox-subheader,body.skin-minerva .mw-parser-output .infobox-above,body.skin-minerva .mw-parser-output .infobox-title,body.skin-minerva .mw-parser-output .infobox-image,body.skin-minerva .mw-parser-output .infobox-full-data,body.skin-minerva .mw-parser-output .infobox-below{text-align:center}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data div{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data div{background:#1f1f23!important;color:#f8f9fa}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn" style="display:inline-block">Har Gobind Khorana <br /> هار گوبند خرانه</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Har_Gobind_Khorana.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e8/Har_Gobind_Khorana.jpg/220px-Har_Gobind_Khorana.jpg" decoding="async" width="220" height="310" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e8/Har_Gobind_Khorana.jpg/330px-Har_Gobind_Khorana.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/e/e8/Har_Gobind_Khorana.jpg 2x" data-file-width="400" data-file-height="563" /></a></span></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data">9 January, 1922<br /><div style="display:inline" class="birthplace"><a href="/w/index.php?title=Raipur,_Pakistan&action=edit&redlink=1" class="new" title="Raipur, Pakistan (not yet started)">Raipur</a>, <a href="/wiki/Multan" title="Multan">Multan</a>, <a href="/wiki/Punjab_Province_(British_India)" title="Punjab Province (British India)">Punjab Province</a>, <a href="/wiki/British_India" class="mw-redirect" title="British India">British India</a> (present day <a href="/wiki/Punjab,_Pakistan" class="mw-redirect" title="Punjab, Pakistan">Punjab</a>, <a href="/wiki/Pakistan" title="Pakistan">Pakistan</a>)</div></td></tr><tr><th scope="row" class="infobox-label">Died</th><td class="infobox-data">9 November, 2011<br /><div style="display:inline" class="deathplace"><a href="/wiki/Boston" title="Boston">Boston</a>, <a href="/wiki/Massachusetts" title="Massachusetts">Massachusetts</a></div></td></tr><tr><th scope="row" class="infobox-label">Nationality</th><td class="infobox-data category"><a href="/wiki/British_Raj" title="British Raj">British Indian</a></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" class="mw-redirect" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></td></tr></tbody></table> <p><b>Har Gobind Khorana</b> (<a href="/wiki/Sindhi_language" title="Sindhi language">Sindhi</a>: هار گوبند خرانه) (also known as <b>Hargobind Khorana</b>; 9 January 1922 – 9 November 2011)<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> was a <a href="/wiki/Biochemist" title="Biochemist">biochemist</a>. He shared the 1968 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" class="mw-redirect" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> with <a href="/wiki/Marshall_W._Nirenberg" class="mw-redirect" title="Marshall W. Nirenberg">Marshall W. Nirenberg</a> and <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a>. </p><p>He worked in <a href="/wiki/Madison,_Wisconsin" title="Madison, Wisconsin">Madison</a>, <a href="/wiki/Columbia,_South_Carolina" title="Columbia, South Carolina">Columbia</a>, and <a href="/wiki/Boston" title="Boston">Boston</a>. </p><p>Their work was on the order of <a href="/wiki/Nucleotides" class="mw-redirect" title="Nucleotides">nucleotides</a> in <a href="/wiki/Nucleic_acid" title="Nucleic acid">nucleic acids</a>. Each triplet of nucleotides codes for a particular <a href="/wiki/Amino_acid" title="Amino acid">amino acid</a>. Therefore the nucleic acid sequence controls how <a href="/wiki/Proteins" class="mw-redirect" title="Proteins">proteins</a> are built. Khorana was the first to build a synthetic <a href="/wiki/Gene" title="Gene">gene</a>, in 1970. This helped a lot, with decoding the <a href="/wiki/Genetic_code" title="Genetic code">genetic code</a>, which is based on amino acids. </p> <div class="mw-heading mw-heading2"><h2 id="Related_pages">Related pages</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Har_Gobind_Khorana&veaction=edit&section=1" title="Change section: Related pages" class="mw-editsection-visualeditor"><span>change</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Har_Gobind_Khorana&action=edit&section=1" title="Edit section's source code: Related pages"><span>change source</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/List_of_Indian_scientists" title="List of Indian scientists">List of Indian scientists</a></li> <li><a href="/wiki/List_of_Nobel_Prize_winners_from_India" class="mw-redirect" title="List of Nobel Prize winners from India">List of Nobel Prize winners from India</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Har_Gobind_Khorana&veaction=edit&section=2" title="Change section: References" class="mw-editsection-visualeditor"><span>change</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Har_Gobind_Khorana&action=edit&section=2" title="Edit section's source code: References"><span>change source</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r9724317">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r9724332">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFCaruthers,_M.;_Wells,_R.2011" class="citation journal cs1">Caruthers, M.; Wells, R. (2011). "Har Gobind Khorana (1922-2011)". <i>Science</i>. <b>334</b> (6062): 1511. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2011Sci...334.1511C">2011Sci...334.1511C</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1217138">10.1126/science.1217138</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/22174242">22174242</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:206539004">206539004</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Har+Gobind+Khorana+%281922-2011%29&rft.volume=334&rft.issue=6062&rft.pages=1511&rft.date=2011&rft_id=info%3Adoi%2F10.1126%2Fscience.1217138&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A206539004%23id-name%3DS2CID&rft_id=info%3Apmid%2F22174242&rft_id=info%3Abibcode%2F2011Sci...334.1511C&rft.au=Caruthers%2C+M.%3B+Wells%2C+R.&rfr_id=info%3Asid%2Fsimple.wikipedia.org%3AHar+Gobind+Khorana" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_journal" title="Template:Cite journal">cite journal</a>}}</code>: CS1 maint: multiple names: authors list (<a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">link</a>)</span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r9724332"><cite id="CITEREFRajbhandary,_U._L.2011" class="citation journal cs1">Rajbhandary, U. L. (2011). "Har Gobind Khorana (1922–2011)". <i>Nature</i>. <b>480</b> (7377): 322. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2011Natur.480..322R">2011Natur.480..322R</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F480322a">10.1038/480322a</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/22170673">22170673</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:205068970">205068970</a>.</cite><span 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title="Template:Nobel Prize in Physiology or Medicine"><abbr title="View this template" style=";background:#d3d3d3;;background:none transparent;border:none;box-shadow:none;padding:0;">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Nobel_Prize_in_Physiology_or_Medicine" title="Template talk:Nobel Prize in Physiology or Medicine"><abbr title="Discuss this template" style=";background:#d3d3d3;;background:none transparent;border:none;box-shadow:none;padding:0;">t</abbr></a></li><li class="nv-edit"><a class="external text" href="https://simple.wikipedia.org/w/index.php?title=Template:Nobel_Prize_in_Physiology_or_Medicine&action=edit"><abbr title="Edit this template" style=";background:#d3d3d3;;background:none transparent;border:none;box-shadow:none;padding:0;">e</abbr></a></li></ul></div><div id="Winners_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="font-size:114%;margin:0 4em">Winners of the <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> in <a href="/wiki/List_of_Nobel_Prize_winners_in_Physiology_or_Medicine" title="List of Nobel Prize winners in Physiology or Medicine">Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1901 – 1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Emil_Adolf_von_Behring" title="Emil Adolf von Behring">Behring</a> (1901) <b>·</b>  <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> (1902) <b>·</b>  <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> (1903) <b>·</b>  <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Pavlov</a> (1904) <b>·</b>  <a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> (1905) <b>·</b>  <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Ramón y Cajal</a> (1906) <b>·</b>  <a href="/wiki/Alphonse_Laveran" title="Alphonse Laveran">Laveran</a> (1907) <b>·</b>  <a href="/wiki/Ilya_Mechnikov" title="Ilya Mechnikov">Mechnikov</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> (1908) <b>·</b>  <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> (1909) <b>·</b>  <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> (1910) <b>·</b>  <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> (1911) <b>·</b>  <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> (1912) <b>·</b>  <a href="/wiki/Charles_Robert_Richet" title="Charles Robert Richet">Richet</a> (1913) <b>·</b>  <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> (1914) <b>·</b>  <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> (1919) <b>·</b>  <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> (1920) <b>·</b>  <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> (1922) <b>·</b>  <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a> / <a href="/wiki/John_Macleod" title="John Macleod">Macleod</a> (1923) <b>·</b>  <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> (1924) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1926 – 1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Fibiger</a> (1926) <b>·</b>  <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> (1927) <b>·</b>  <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> (1928) <b>·</b>  <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a> / <a href="/wiki/Frederick_Hopkins" title="Frederick Hopkins">Hopkins</a> (1929) <b>·</b>  <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> (1930) <b>·</b>  <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> (1931) <b>·</b>  <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a> / <a href="/wiki/Edgar_Adrian,_1st_Baron_Adrian" title="Edgar Adrian, 1st Baron Adrian">Adrian</a> (1932) <b>·</b>  <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> (1933) <b>·</b>  <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">Murphy</a> (1934) <b>·</b>  <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> (1935) <b>·</b>  <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> (1936) <b>·</b>  <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> (1937) <b>·</b>  <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Heymans</a> (1938) <b>·</b>  <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> (1939) <b>·</b>  <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> (1943) <b>·</b>  <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a> / <a href="/wiki/Herbert_Gasser" title="Herbert Gasser">Gasser</a> (1944) <b>·</b>  <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> (1945) <b>·</b>  <a href="/wiki/H._J._Muller" title="H. J. Muller">Muller</a> (1946) <b>·</b>  <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C.Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">G.Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> (1947) <b>·</b>  <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> (1948) <b>·</b>  <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> (1949) <b>·</b>  <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a> / <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> (1950) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1951 – 1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> (1951) <b>·</b>  <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> (1952) <b>·</b>  <a href="/wiki/Hans_Krebs" title="Hans Krebs">Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" class="mw-redirect" title="Fritz Albert Lipmann">Lipmann</a> (1953) <b>·</b>  <a href="/wiki/John_Franklin_Enders" class="mw-redirect" title="John Franklin Enders">Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" class="mw-redirect" title="Thomas Huckle Weller">Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" class="mw-redirect" title="Frederick Chapman Robbins">Robbins</a> (1954) <b>·</b>  <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> (1955) <b>·</b>  <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" class="mw-redirect" title="André Frédéric Cournand">Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> (1956) <b>·</b>  <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> (1957) <b>·</b>  <a href="/wiki/George_Beadle" title="George Beadle">Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> (1958) <b>·</b>  <a href="/wiki/Severo_Ochoa" class="mw-redirect" title="Severo Ochoa">Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> (1959) <b>·</b>  <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> (1960) <b>·</b>  <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> (1961) <b>·</b>  <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a> / <a href="/wiki/James_D._Watson" title="James D. Watson">Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> (1962) <b>·</b>  <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> (1963) <b>·</b>  <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a> / <a href="/wiki/Feodor_Felix_Konrad_Lynen" class="mw-redirect" title="Feodor Felix Konrad Lynen">Lynen</a> (1964) <b>·</b>  <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" class="mw-redirect" title="André Michel Lwoff">Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> (1965) <b>·</b>  <a href="/wiki/Francis_Peyton_Rous" class="mw-redirect" title="Francis Peyton Rous">Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> (1966) <b>·</b>  <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">Wald</a> (1967) <b>·</b>  <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a> / <a class="mw-selflink selflink">Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> (1968) <b>·</b>  <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> (1969) <b>·</b>  <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Euler</a> / <a href="/wiki/Julius_Axelrod" class="mw-redirect" title="Julius Axelrod">Axelrod</a> (1970) <b>·</b>  <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Sutherland</a> (1971) <b>·</b>  <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Porter</a> (1972) <b>·</b>  <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a> / <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> (1973) <b>·</b>  <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> (1974) <b>·</b>  <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a> / <a href="/wiki/Howard_Temin" title="Howard Temin">Temin</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a> (1975) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1976 – 2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> (1976) <b>·</b>  <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a> / <a href="/wiki/Rosalyn_Yalow" title="Rosalyn Yalow">Yalow</a> (1977) <b>·</b>  <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a> / <a href="/wiki/Hamilton_Smith" title="Hamilton Smith">Smith</a> (1978) <b>·</b>  <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" class="mw-redirect" title="Godfrey Hounsfield">Hounsfield</a> (1979) <b>·</b>  <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> (1980) <b>·</b>  <a href="/wiki/Roger_Sperry" class="mw-redirect" title="Roger Sperry">Sperry</a> / <a href="/wiki/David_Hubel" title="David Hubel">Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> (1981) <b>·</b>  <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Samuelsson</a> / <a href="/wiki/John_Robert_Vane" class="mw-redirect" title="John Robert Vane">Vane</a> (1982) <b>·</b>  <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> (1983) <b>·</b>  <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" class="mw-redirect" title="César Milstein">Milstein</a> (1984) <b>·</b>  <a href="/wiki/Michael_Stuart_Brown" class="mw-redirect" title="Michael Stuart Brown">Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Goldstein</a> (1985) <b>·</b>  <a href="/wiki/Stanley_Cohen" title="Stanley Cohen">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a> (1986) <b>·</b>  <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> (1987) <b>·</b>  <a href="/wiki/James_Black" title="James Black">Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> (1988) <b>·</b>  <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">Bishop</a> / <a href="/wiki/Harold_Varmus" title="Harold Varmus">Varmus</a> (1989) <b>·</b>  <a href="/wiki/Joseph_Murray" title="Joseph Murray">Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">Thomas</a> (1990) <b>·</b>  <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> (1991) <b>·</b>  <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Krebs</a> (1992) <b>·</b>  <a href="/wiki/Richard_Roberts" title="Richard Roberts">Roberts</a> / <a href="/wiki/Phillip_Sharp" title="Phillip Sharp">Sharp</a> (1993) <b>·</b>  <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> (1994) <b>·</b>  <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> (1995) <b>·</b>  <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> (1996) <b>·</b>  <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> (1997) <b>·</b>  <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> (1998) <b>·</b>  <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> (1999) <b>·</b>  <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> (2000) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2001 – present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Lee_Hartwell" title="Lee Hartwell">Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> (2001) <b>·</b>  <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">Sulston</a> (2002) <b>·</b>  <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> (2003) <b>·</b>  <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> (2004) <b>·</b>  <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> (2005) <b>·</b>  <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> (2006) <b>·</b>  <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a> (2007) <b>·</b>  <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré-Sinoussi</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> / <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">zur Hausen</a> (2008) <b>·</b>  <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a> / <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a> / <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> (2009) <b>·</b>  <a href="/wiki/Robert_Edwards" title="Robert Edwards">Edwards</a> (2010) <b>·</b>  <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a> / <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>; <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> (2011) <b>·</b>  <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> (2012) <b>·</b>  <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Südhof</a> (2013) <b>·</b>  <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">M-B. Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">E. Moser</a> (2014) <b>·</b>  <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Youyou</a> (2015) <b>·</b>  <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Ohsumi</a> (2016) <b>·</b>  <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> (2017) <b>·</b>  <a href="/wiki/James_P._Allison" title="James P. Allison">Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> (2018) <b>·</b>  <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">Kaelin Jr.</a> (2019) <b>·</b>  <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Atler</a> / <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> (2020) <b>·</b>  <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> / <a href="/wiki/David_Julius" title="David Julius">Julius</a> (2021) <b>·</b>  <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> (2022) <b>·</b>  <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> (2023) <b>·</b>  <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> (2024) </p> </div></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://simple.wikipedia.org/w/index.php?title=Har_Gobind_Khorana&oldid=9241762">https://simple.wikipedia.org/w/index.php?title=Har_Gobind_Khorana&oldid=9241762</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Special:Categories" title="Special:Categories">Categories</a>: <ul><li><a href="/wiki/Category:Biochemists" title="Category:Biochemists">Biochemists</a></li><li><a href="/wiki/Category:1922_births" title="Category:1922 births">1922 births</a></li><li><a href="/wiki/Category:2011_deaths" title="Category:2011 deaths">2011 deaths</a></li><li><a href="/wiki/Category:Naturalized_citizens_of_the_United_States" title="Category:Naturalized citizens of the United States">Naturalized citizens of the United States</a></li><li><a href="/wiki/Category:Nobel_Prize_winners" title="Category:Nobel Prize winners">Nobel Prize winners</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Hidden categories: <ul><li><a href="/wiki/Category:Pages_using_the_JsonConfig_extension" title="Category:Pages using the JsonConfig extension">Pages using the JsonConfig extension</a></li><li><a href="/wiki/Category:CS1_maint:_multiple_names:_authors_list" title="Category:CS1 maint: multiple names: authors list">CS1 maint: multiple names: authors list</a></li><li><a href="/wiki/Category:Articles_with_hCards" title="Category:Articles with hCards">Articles with hCards</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> This page was last changed on 10 December 2023, at 16:38.</li> <li id="footer-info-copyright">Text is available under the <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/">Creative Commons Attribution-ShareAlike License</a> and the <a href="/wiki/Wikipedia:Text_of_the_GNU_Free_Documentation_License" title="Wikipedia:Text of the GNU Free Documentation License">GFDL</a>; additional terms may apply. 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