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</div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contenidos" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Cambiar a la tabla de contenidos" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Cambiar a la tabla de contenidos</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Yoshinori Ohsumi</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Ir a un artículo en otro idioma. Disponible en 66 idiomas" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-66" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">66 idiomas</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%8A%D9%88%D8%B4%D9%8A%D9%86%D9%88%D8%B1%D9%8A_%D8%A3%D9%88%D8%B3%D9%88%D9%85%D9%8A" title="يوشينوري أوسومي (árabe)" lang="ar" hreflang="ar" data-title="يوشينوري أوسومي" data-language-autonym="العربية" data-language-local-name="árabe" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-as mw-list-item"><a href="https://as.wikipedia.org/wiki/%E0%A6%AF%E0%A6%BC%E0%A6%B6%E0%A7%8D%E0%A6%AC%E0%A6%BF%E0%A6%A8%E0%A7%B0%E0%A7%80_%E0%A6%85%E0%A6%B9%E0%A6%9A%E0%A7%81%E0%A6%AE%E0%A6%BF" title="য়শ্বিনৰী অহচুমি (asamés)" lang="as" hreflang="as" data-title="য়শ্বিনৰী অহচুমি" data-language-autonym="অসমীয়া" data-language-local-name="asamés" class="interlanguage-link-target"><span>অসমীয়া</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Yo%C5%9Finori_Osumi" title="Yoşinori Osumi (azerbaiyano)" lang="az" hreflang="az" data-title="Yoşinori Osumi" data-language-autonym="Azərbaycanca" data-language-local-name="azerbaiyano" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اوسومی (South Azerbaijani)" lang="azb" hreflang="azb" data-title="یوشینوری اوسومی" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%81%D1%81%D1%96%D0%BD%D0%BE%D1%80%D1%8B_%D0%90%D1%81%D1%83%D0%BC%D1%96" title="Ёсіноры Асумі (bielorruso)" lang="be" hreflang="be" data-title="Ёсіноры Асумі" data-language-autonym="Беларуская" data-language-local-name="bielorruso" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми (búlgaro)" lang="bg" hreflang="bg" data-title="Йошинори Осуми" data-language-autonym="Български" data-language-local-name="búlgaro" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%87%E0%A6%AF%E0%A6%BC%E0%A7%8B%E0%A6%B6%E0%A6%BF%E0%A6%A8%E0%A7%8B%E0%A6%B0%E0%A6%BF_%E0%A6%93%E0%A6%B8%E0%A7%81%E0%A6%AE%E0%A6%BF" title="ইয়োশিনোরি ওসুমি (bengalí)" lang="bn" hreflang="bn" data-title="ইয়োশিনোরি ওসুমি" data-language-autonym="বাংলা" data-language-local-name="bengalí" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (bosnio)" lang="bs" hreflang="bs" data-title="Yoshinori Ohsumi" data-language-autonym="Bosanski" data-language-local-name="bosnio" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (catalán)" lang="ca" hreflang="ca" data-title="Yoshinori Ohsumi" data-language-autonym="Català" data-language-local-name="catalán" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Jo%C5%A1inori_%C3%93sumi" title="Jošinori Ósumi (checo)" lang="cs" hreflang="cs" data-title="Jošinori Ósumi" data-language-autonym="Čeština" data-language-local-name="checo" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (danés)" lang="da" hreflang="da" data-title="Yoshinori Ohsumi" data-language-autonym="Dansk" data-language-local-name="danés" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (alemán)" lang="de" hreflang="de" data-title="Yoshinori Ōsumi" data-language-autonym="Deutsch" data-language-local-name="alemán" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%93%CE%B9%CE%BF%CF%83%CE%B9%CE%BD%CF%8C%CF%81%CE%B9_%CE%9F%CF%87%CF%83%CE%BF%CF%8D%CE%BC%CE%B9" title="Γιοσινόρι Οχσούμι (griego)" lang="el" hreflang="el" data-title="Γιοσινόρι Οχσούμι" data-language-autonym="Ελληνικά" data-language-local-name="griego" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (inglés)" lang="en" hreflang="en" data-title="Yoshinori Ohsumi" data-language-autonym="English" data-language-local-name="inglés" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (estonio)" lang="et" hreflang="et" data-title="Yoshinori Ōsumi" data-language-autonym="Eesti" data-language-local-name="estonio" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (euskera)" lang="eu" hreflang="eu" data-title="Yoshinori Ohsumi" data-language-autonym="Euskara" data-language-local-name="euskera" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اسومی (persa)" lang="fa" hreflang="fa" data-title="یوشینوری اسومی" data-language-autonym="فارسی" data-language-local-name="persa" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (finés)" lang="fi" hreflang="fi" data-title="Yoshinori Ōsumi" data-language-autonym="Suomi" data-language-local-name="finés" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (francés)" lang="fr" hreflang="fr" data-title="Yoshinori Ohsumi" data-language-autonym="Français" data-language-local-name="francés" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (gallego)" lang="gl" hreflang="gl" data-title="Yoshinori Ōsumi" data-language-autonym="Galego" data-language-local-name="gallego" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%99%D7%95%D7%A9%D7%99%D7%A0%D7%95%D7%A8%D7%99_%D7%90%D7%95%D7%A1%D7%95%D7%9E%D7%99" title="יושינורי אוסומי (hebreo)" lang="he" hreflang="he" data-title="יושינורי אוסומי" data-language-autonym="עברית" data-language-local-name="hebreo" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/%C3%93szumi_Josinori" title="Ószumi Josinori (húngaro)" lang="hu" hreflang="hu" data-title="Ószumi Josinori" data-language-autonym="Magyar" data-language-local-name="húngaro" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%85%D5%B8%D5%BD%D5%AB%D5%B6%D5%B8%D6%80%D5%AB_%D5%95%D5%BD%D5%B8%D6%82%D5%B4%D5%AB" title="Յոսինորի Օսումի (armenio)" lang="hy" hreflang="hy" data-title="Յոսինորի Օսումի" data-language-autonym="Հայերեն" data-language-local-name="armenio" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (indonesio)" lang="id" hreflang="id" data-title="Yoshinori Ohsumi" data-language-autonym="Bahasa Indonesia" data-language-local-name="indonesio" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (ido)" lang="io" hreflang="io" data-title="Yoshinori Ohsumi" data-language-autonym="Ido" data-language-local-name="ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it badge-Q17437798 badge-goodarticle mw-list-item" title="artículo bueno"><a href="https://it.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (italiano)" lang="it" hreflang="it" data-title="Yoshinori Ōsumi" data-language-autonym="Italiano" data-language-local-name="italiano" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 (japonés)" lang="ja" hreflang="ja" data-title="大隅良典" data-language-autonym="日本語" data-language-local-name="japonés" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми (kazajo)" lang="kk" hreflang="kk" data-title="Йошинори Осуми" data-language-autonym="Қазақша" data-language-local-name="kazajo" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%AF%E0%B3%8A%E0%B2%B6%E0%B2%BF%E0%B2%A8%E0%B3%8B%E0%B2%B0%E0%B2%BF_%E0%B2%93%E0%B2%B6%E0%B3%81%E0%B2%AE%E0%B2%BF" title="ಯೊಶಿನೋರಿ ಓಶುಮಿ (canarés)" lang="kn" hreflang="kn" data-title="ಯೊಶಿನೋರಿ ಓಶುಮಿ" data-language-autonym="ಕನ್ನಡ" data-language-local-name="canarés" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%98%A4%EC%8A%A4%EB%AF%B8_%EC%9A%94%EC%8B%9C%EB%85%B8%EB%A6%AC" title="오스미 요시노리 (coreano)" lang="ko" hreflang="ko" data-title="오스미 요시노리" data-language-autonym="한국어" data-language-local-name="coreano" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (latín)" lang="la" hreflang="la" data-title="Yoshinori Ohsumi" data-language-autonym="Latina" data-language-local-name="latín" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%88%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Јошинори Осуми (macedonio)" lang="mk" hreflang="mk" data-title="Јошинори Осуми" data-language-autonym="Македонски" data-language-local-name="macedonio" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%AF%E0%B5%8B%E0%B4%B7%E0%B4%BF%E0%B4%A8%E0%B5%8B%E0%B4%B1%E0%B4%BF_%E0%B4%89%E0%B4%B7%E0%B5%81%E0%B4%AE%E0%B4%BF" title="യോഷിനോറി ഉഷുമി (malayálam)" lang="ml" hreflang="ml" data-title="യോഷിനോറി ഉഷുമി" data-language-autonym="മലയാളം" data-language-local-name="malayálam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (malayo)" lang="ms" hreflang="ms" data-title="Yoshinori Ohsumi" data-language-autonym="Bahasa Melayu" data-language-local-name="malayo" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D8%B3%D9%88%D9%85%DB%8C" title="یوشینوری اسومی (mazandaraní)" lang="mzn" hreflang="mzn" data-title="یوشینوری اسومی" data-language-autonym="مازِرونی" data-language-local-name="mazandaraní" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (bajo alemán)" lang="nds" hreflang="nds" data-title="Yoshinori Ōsumi" data-language-autonym="Plattdüütsch" data-language-local-name="bajo alemán" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-ne mw-list-item"><a href="https://ne.wikipedia.org/wiki/%E0%A4%AF%E0%A5%8B%E0%A4%B8%E0%A4%BF%E0%A4%A8%E0%A5%8B%E0%A4%B0%E0%A5%80_%E0%A4%93%E0%A4%B8%E0%A5%81%E0%A4%AE%E0%A5%80" title="योसिनोरी ओसुमी (nepalí)" lang="ne" hreflang="ne" data-title="योसिनोरी ओसुमी" data-language-autonym="नेपाली" data-language-local-name="nepalí" class="interlanguage-link-target"><span>नेपाली</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Yoshinori_Osumi" title="Yoshinori Osumi (neerlandés)" lang="nl" hreflang="nl" data-title="Yoshinori Osumi" data-language-autonym="Nederlands" data-language-local-name="neerlandés" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (noruego bokmal)" lang="nb" hreflang="nb" data-title="Yoshinori Ohsumi" data-language-autonym="Norsk bokmål" data-language-local-name="noruego bokmal" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (occitano)" lang="oc" hreflang="oc" data-title="Yoshinori Ōsumi" data-language-autonym="Occitan" data-language-local-name="occitano" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pa mw-list-item"><a href="https://pa.wikipedia.org/wiki/%E0%A8%AF%E0%A9%8B%E0%A8%B8%E0%A8%BC%E0%A9%80%E0%A8%A8%E0%A9%8B%E0%A8%B0%E0%A9%80_%E0%A8%93%E0%A8%B8%E0%A9%81%E0%A8%AE%E0%A9%80" title="ਯੋਸ਼ੀਨੋਰੀ ਓਸੁਮੀ (punyabí)" lang="pa" hreflang="pa" data-title="ਯੋਸ਼ੀਨੋਰੀ ਓਸੁਮੀ" data-language-autonym="ਪੰਜਾਬੀ" data-language-local-name="punyabí" class="interlanguage-link-target"><span>ਪੰਜਾਬੀ</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (polaco)" lang="pl" hreflang="pl" data-title="Yoshinori Ōsumi" data-language-autonym="Polski" data-language-local-name="polaco" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%DB%81%D8%B3%D9%85%DB%8C" title="یوشینوری اوہسمی (Western Punjabi)" lang="pnb" hreflang="pnb" data-title="یوشینوری اوہسمی" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (portugués)" lang="pt" hreflang="pt" data-title="Yoshinori Ohsumi" data-language-autonym="Português" data-language-local-name="portugués" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (rumano)" lang="ro" hreflang="ro" data-title="Yoshinori Ohsumi" data-language-autonym="Română" data-language-local-name="rumano" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9E%D1%81%D1%83%D0%BC%D0%B8,_%D0%81%D1%81%D0%B8%D0%BD%D0%BE%D1%80%D0%B8" title="Осуми, Ёсинори (ruso)" lang="ru" hreflang="ru" data-title="Осуми, Ёсинори" data-language-autonym="Русский" data-language-local-name="ruso" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (escocés)" lang="sco" hreflang="sco" data-title="Yoshinori Ohsumi" data-language-autonym="Scots" data-language-local-name="escocés" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Yoshinori_%C5%8Csumi" title="Yoshinori Ōsumi (serbocroata)" lang="sh" hreflang="sh" data-title="Yoshinori Ōsumi" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="serbocroata" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-si mw-list-item"><a href="https://si.wikipedia.org/wiki/%E0%B6%BA%E0%B7%9D%E0%B7%82%E0%B7%92%E0%B6%B1%E0%B7%9C%E0%B6%BB%E0%B7%92_%E0%B6%94%E0%B7%83%E0%B7%94%E0%B6%B8%E0%B7%92" title="යෝෂිනොරි ඔසුමි (cingalés)" lang="si" hreflang="si" data-title="යෝෂිනොරි ඔසුමි" data-language-autonym="සිංහල" data-language-local-name="cingalés" class="interlanguage-link-target"><span>සිංහල</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (Simple English)" lang="en-simple" hreflang="en-simple" data-title="Yoshinori Ohsumi" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Jo%C5%A1inori_Ohsumi" title="Jošinori Ohsumi (esloveno)" lang="sl" hreflang="sl" data-title="Jošinori Ohsumi" data-language-autonym="Slovenščina" data-language-local-name="esloveno" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (albanés)" lang="sq" hreflang="sq" data-title="Yoshinori Ohsumi" data-language-autonym="Shqip" data-language-local-name="albanés" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%88%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Јошинори Осуми (serbio)" lang="sr" hreflang="sr" data-title="Јошинори Осуми" data-language-autonym="Српски / srpski" data-language-local-name="serbio" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (sueco)" lang="sv" hreflang="sv" data-title="Yoshinori Ohsumi" data-language-autonym="Svenska" data-language-local-name="sueco" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (suajili)" lang="sw" hreflang="sw" data-title="Yoshinori Ohsumi" data-language-autonym="Kiswahili" data-language-local-name="suajili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%87%E0%AE%AF%E0%AF%8B%E0%AE%9A%E0%AE%BF%E0%AE%A9%E0%AF%8B%E0%AE%B0%E0%AE%BF_%E0%AE%93%E0%AE%9A%E0%AF%82%E0%AE%AE%E0%AE%BF" title="இயோசினோரி ஓசூமி (tamil)" lang="ta" hreflang="ta" data-title="இயோசினோரி ஓசூமி" data-language-autonym="தமிழ்" data-language-local-name="tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a href="https://te.wikipedia.org/wiki/%E0%B0%AF%E0%B1%8B%E0%B0%B7%E0%B0%BF%E0%B0%A8%E0%B1%8B%E0%B0%B0%E0%B0%BF_%E0%B0%93%E0%B0%B9%E0%B1%8D%E0%B0%B8%E0%B1%81%E0%B0%AE%E0%B1%80" title="యోషినోరి ఓహ్సుమీ (telugu)" lang="te" hreflang="te" data-title="యోషినోరి ఓహ్సుమీ" data-language-autonym="తెలుగు" data-language-local-name="telugu" class="interlanguage-link-target"><span>తెలుగు</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B9%82%E0%B8%A2%E0%B8%8A%E0%B8%B4%E0%B9%82%E0%B8%99%E0%B8%A3%E0%B8%B4_%E0%B9%82%E0%B8%AD%E0%B8%8B%E0%B8%B9%E0%B8%A1%E0%B8%B4" title="โยชิโนริ โอซูมิ (tailandés)" lang="th" hreflang="th" data-title="โยชิโนริ โอซูมิ" data-language-autonym="ไทย" data-language-local-name="tailandés" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi (turco)" lang="tr" hreflang="tr" data-title="Yoshinori Ohsumi" data-language-autonym="Türkçe" data-language-local-name="turco" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%99%D0%BE%D1%88%D0%B8%D0%BD%D0%BE%D1%80%D0%B8_%D0%9E%D1%81%D1%83%D0%BC%D0%B8" title="Йошинори Осуми (tártaro)" lang="tt" hreflang="tt" data-title="Йошинори Осуми" data-language-autonym="Татарча / tatarça" data-language-local-name="tártaro" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9E%D1%81%D1%83%D0%BC%D1%96_%D0%99%D0%BE%D1%81%D1%96%D0%BD%D0%BE%D1%80%D1%96" title="Осумі Йосінорі (ucraniano)" lang="uk" hreflang="uk" data-title="Осумі Йосінорі" data-language-autonym="Українська" data-language-local-name="ucraniano" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%DB%8C%D9%88%D8%B4%DB%8C%D9%86%D9%88%D8%B1%DB%8C_%D8%A7%D9%88%D8%B4%D9%88%D9%85%DB%8C" title="یوشینوری اوشومی (urdu)" lang="ur" hreflang="ur" data-title="یوشینوری اوشومی" data-language-autonym="اردو" data-language-local-name="urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/%C5%8Csumi_Yoshinori" title="Ōsumi Yoshinori (vietnamita)" lang="vi" hreflang="vi" data-title="Ōsumi Yoshinori" data-language-autonym="Tiếng Việt" data-language-local-name="vietnamita" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 (chino wu)" lang="wuu" hreflang="wuu" data-title="大隅良典" data-language-autonym="吴语" data-language-local-name="chino wu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%A4%A7%E9%9A%85%E8%89%AF%E5%85%B8" title="大隅良典 (chino)" lang="zh" hreflang="zh" data-title="大隅良典" data-language-autonym="中文" data-language-local-name="chino" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/%C3%94sumi_Yosinori" title="Ôsumi Yosinori (chino min nan)" lang="nan" hreflang="nan" data-title="Ôsumi Yosinori" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="chino min nan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q8056214#sitelinks-wikipedia" title="Editar enlaces interlingüísticos" class="wbc-editpage">Editar enlaces</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Espacios de nombres"> <div 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vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Apariencia</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">mover a la barra lateral</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">ocultar</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">De Wikipedia, la enciclopedia libre</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="es" dir="ltr"><style data-mw-deduplicate="TemplateStyles:r157776694">.mw-parser-output .infobox .imagen{max-width:100%;margin:0 auto}.mw-parser-output .infobox .imagen img{max-width:100%;height:auto}.mw-parser-output .infobox .mw-kartographer-container .thumbinner,.mw-parser-output .infobox .mw-kartographer-map{box-sizing:border-box;width:100%!important}body.skin-timeless .mw-parser-output .infobox .imagen a.image>img{max-width:100%!important;height:auto!important}</style><table class="infobox biography vcard" style="width:22.7em; line-height: 1.4em; text-align:left; padding:.23em;"><tbody><tr><th colspan="3" class="cabecera persona" style="text-align:center;background-color:transparent;color:inherit;background-color:#8DB1C3;color:#000;">Yoshinori Ōsumi</th></tr><tr><td colspan="3" class="imagen" style="text-align:center;line-height:1.3em; vertical-align:middle;;"> <span typeof="mw:File"><a href="/wiki/Archivo:Nobel_Laureates_7428_(30679389523)_(cropped).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/71/Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg/220px-Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg" decoding="async" width="220" height="301" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/71/Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg/330px-Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/71/Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg/440px-Nobel_Laureates_7428_%2830679389523%29_%28cropped%29.jpg 2x" data-file-width="3418" data-file-height="4684" /></a></span><br /><div style="display:inline;">Yoshinori Ōsumi en 2016</div></td></tr><tr><th colspan="3" style="text-align:center;background-color:#8DB1C3;color:#000;">Información personal</th></tr><tr><th scope="row" style="text-align:left;width:33%;">Nombre en japonés</th><td colspan="2" style="line-height:1.3em; vertical-align:middle;;"> 大隅良典<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P1559" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Nacimiento</th><td colspan="2" style="line-height:1.3em; vertical-align:middle;;"> 9 de febrero de 1945<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P569" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span> (79 años)<br /><a href="/wiki/Fukuoka" title="Fukuoka">Fukuoka</a> (<a href="/wiki/Prefectura_de_Fukuoka" title="Prefectura de Fukuoka">Prefectura de Fukuoka</a>, <a href="/wiki/Jap%C3%B3n" title="Japón">Japón</a>)<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P19" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Nacionalidad</th><td colspan="2" class="category" style="line-height:1.3em; vertical-align:middle;;"> Japonesa</td></tr><tr><th colspan="3" style="text-align:center;background-color:#8DB1C3;color:#000;">Familia</th></tr><tr><th scope="row" style="text-align:left;width:33%;">Padre</th><td colspan="2" class="plainlist" style="line-height:1.3em; vertical-align:middle;;"> Yoshio Ōsumi<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P22" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Cónyuge</th><td colspan="2" class="plainlist" style="line-height:1.3em; vertical-align:middle;;"> Mariko Osumi<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P26" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th colspan="3" style="text-align:center;background-color:#8DB1C3;color:#000;">Educación</th></tr><tr><th scope="row" style="text-align:left;width:33%;">Educación</th><td colspan="2" style="line-height:1.3em; vertical-align:middle;;"> doctorado en ciencias<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P512" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Educado en</th><td colspan="2" class="plainlist" style="line-height:1.3em; vertical-align:middle;;"> <ul><li>Fukuoka Prefectural Fukuoka High School</li><li><a href="/wiki/Universidad_de_Tokio" title="Universidad de Tokio">Universidad de Tokio</a> <small>(1963-1972)</small></li><li><a href="/wiki/Universidad_Rockefeller" title="Universidad Rockefeller">Universidad Rockefeller</a> <small>(1974-1977)</small><span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P69" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></li></ul></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Supervisor doctoral</th><td colspan="2" style="line-height:1.3em; vertical-align:middle;;"> Kazutomo Imahori<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P184" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th colspan="3" style="text-align:center;background-color:#8DB1C3;color:#000;">Información profesional</th></tr><tr><th scope="row" style="text-align:left;width:33%;">Ocupación</th><td colspan="2" class="role" style="line-height:1.3em; vertical-align:middle;;"> <a href="/wiki/Bi%C3%B3logo" title="Biólogo">Biólogo</a><span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P106" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Área</th><td colspan="2" style="line-height:1.3em; vertical-align:middle;;"> <a href="/wiki/Biolog%C3%ADa_molecular" title="Biología molecular">Biología molecular</a><span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P101" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Empleador</th><td colspan="2" class="plainlist" style="line-height:1.3em; vertical-align:middle;;"> <ul><li>Sokendai</li><li><a href="/wiki/Instituto_Tecnol%C3%B3gico_de_Tokio" title="Instituto Tecnológico de Tokio">Instituto Tecnológico de Tokio</a></li><li>National Institute for Basic Biology, Japan<span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P108" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></li></ul></td></tr><tr><th scope="row" style="text-align:left;width:33%;">Sitio web</th><td colspan="2" class="biography vcard" style="line-height:1.3em; vertical-align:middle;;"> <a rel="nofollow" class="external text" href="http://www.ohsumilab.aro.iri.titech.ac.jp/english.html">www.ohsumilab.aro.iri.titech.ac.jp/english.html</a><span class="wikidata-link lapiz noprint"> <span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q8056214?uselang=es#P856" title="Ver y modificar los datos en Wikidata"><img alt="Ver y modificar los datos en Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><td class="noprint" colspan="3" style="text-align:left;"><div class="plainlinks wikidata-link" style="font-size: 0.85em">[<a href="https://www.wikidata.org/wiki/Q8056214" class="extiw" title="d:Q8056214">editar datos en Wikidata</a>]</div></td></tr></tbody></table> <p><b>Yoshinori Ōsumi</b> (<a href="/wiki/Fukuoka" title="Fukuoka">Fukuoka</a>, <a href="/wiki/Jap%C3%B3n" title="Japón">Japón</a>, 9 de febrero de 1945) es un <a href="/wiki/Biolog%C3%ADa_celular" title="Biología celular">biólogo celular</a> <a href="/wiki/Jap%C3%B3n" title="Japón">japonés</a>, quien descubrió la existencia de la <a href="/wiki/Autofagia" title="Autofagia">autofagia</a> en <a href="/wiki/Levaduras" class="mw-redirect" title="Levaduras">levaduras</a>, utilizando estas últimas para identificar los <a href="/wiki/Genes" class="mw-redirect" title="Genes">genes</a> implicados en el proceso en sí. Gracias a este cribado genético cada vez más preciso, ha identificado algunas de las funciones importantes de la autofagia en los procesos fisiológicos humanos. Aún se están investigando otras funciones. Desde 2014 es <a href="/wiki/Catedr%C3%A1tico_de_universidad" title="Catedrático de universidad">catedrático</a> (<a href="/wiki/Em%C3%A9rito" title="Emérito">emérito</a>) del <a href="/wiki/Instituto_Tecnol%C3%B3gico_de_Tokio" title="Instituto Tecnológico de Tokio">Instituto Tecnológico de Tokio</a>.<sup id="cite_ref-1" class="reference separada"><a href="#cite_note-1"><span class="corchete-llamada">[</span>1<span class="corchete-llamada">]</span></a></sup> Recibió el <a href="/wiki/Premio_Kyoto_de_Ciencias_B%C3%A1sicas" title="Premio Kyoto de Ciencias Básicas">Premio Kioto</a> en 2012 y,<sup id="cite_ref-2" class="reference separada"><a href="#cite_note-2"><span class="corchete-llamada">[</span>2<span class="corchete-llamada">]</span></a></sup> en 2016, el <a href="/wiki/Premio_Nobel_de_Medicina" class="mw-redirect" title="Premio Nobel de Medicina">Premio Nobel de Medicina</a> «por sus descubrimientos sobre los mecanismos de la <a href="/wiki/Autofagia" title="Autofagia">autofagia</a>».<sup id="cite_ref-3" class="reference separada"><a href="#cite_note-3"><span class="corchete-llamada">[</span>3<span class="corchete-llamada">]</span></a></sup><sup id="cite_ref-4" class="reference separada"><a href="#cite_note-4"><span class="corchete-llamada">[</span>4<span class="corchete-llamada">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Obra">Obra</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_Ohsumi&action=edit&section=1" title="Editar sección: Obra"><span>editar</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Publicaciones">Publicaciones</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_Ohsumi&action=edit&section=2" title="Editar sección: Publicaciones"><span>editar</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><span id="CITAREFTsukadaŌsumi25_de_octubre_de_1993" class="citation publicación">Tsukada, M; Ōsumi, Y (25 de octubre de 1993). «Isolation and characterization of autophagy-defective mutants of Saccharomyces cerevisiae». <i>FEBS Letters</i> <b>333</b> (1-2): 169-74. <small><a href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/8224160">8224160</a></small>. <small><a href="/wiki/Digital_object_identifier" class="mw-redirect" title="Digital object identifier">doi</a>:<a rel="nofollow" class="external text" href="https://dx.doi.org/10.1016%2F0014-5793%2893%2980398-E">10.1016/0014-5793(93)80398-E</a></small>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=Isolation+and+characterization+of+autophagy-defective+mutants+of+Saccharomyces+cerevisiae&rft.au=%C5%8Csumi%2C+Y&rft.au=Tsukada%2C+M&rft.aufirst=M&rft.aulast=Tsukada&rft.date=25+de+octubre+de+1993&rft.genre=article&rft.issue=1-2&rft.jtitle=FEBS+Letters&rft.pages=169-74&rft.volume=333&rft_id=info%3Adoi%2F10.1016%2F0014-5793%2893%2980398-E&rft_id=info%3Apmid%2F8224160&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span></li> <li><span id="CITAREFMizushimaNodaYoshimoriTanaka24_de_septiembre_de_1998" class="citation publicación">Mizushima, N; Noda, T; Yoshimori, T; Tanaka, Y; Ishii, T; George, MD; Klionsky, DJ; Ohsumi, M <i>et al.</i> (24 de septiembre de 1998). «A protein conjugation system essential for autophagy». <i>Nature</i> <b>395</b> (6700): 395-8. <small><a href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/9759731">9759731</a></small>. <small><a href="/wiki/Digital_object_identifier" class="mw-redirect" title="Digital object identifier">doi</a>:<a rel="nofollow" class="external text" href="https://dx.doi.org/10.1038%2F26506">10.1038/26506</a></small>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=A+protein+conjugation+system+essential+for+autophagy&rft.au=%C5%8Csumi%2C+Y&rft.au=George%2C+MD&rft.au=Ishii%2C+T&rft.au=Klionsky%2C+DJ&rft.au=Mizushima%2C+N&rft.au=Noda%2C+T&rft.au=Ohsumi%2C+M&rft.au=Tanaka%2C+Y&rft.au=Yoshimori%2C+T&rft.aufirst=N&rft.aulast=Mizushima&rft.date=24+de+septiembre+de+1998&rft.genre=article&rft.issue=6700&rft.jtitle=Nature&rft.pages=395-8&rft.volume=395&rft_id=info%3Adoi%2F10.1038%2F26506&rft_id=info%3Apmid%2F9759731&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span> <span style="display:none;font-size:100%" class="error citation-comment">Se sugiere usar <code>|número-autores=</code> (<a href="/wiki/Ayuda:Errores_en_las_referencias#displayauthors" title="Ayuda:Errores en las referencias">ayuda</a>)</span></li> <li><span id="CITAREFKabeyaMizushimaUenoYamamoto2000" class="citation publicación">Kabeya, Y.; Mizushima, N.; Ueno, T.; Yamamoto, A.; Kirisako, T.; Noda, T.; Kominami, E.; Ōsumi, Y. <i>et al.</i> (2000). <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pmc/articles/PMC305793">«LC3, a mammalian homologue of yeast Apg8p, is localized in autophagosome membranes after processing»</a>. <i>The EMBO Journal</i> <b>19</b> (21): 5720-5728. <small><a href="/wiki/PubMed_Central" title="PubMed Central">PMC</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pmc/articles/PMC305793">305793</a></small>. <small><a href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/11060023">11060023</a></small>. <small><a href="/wiki/Digital_object_identifier" class="mw-redirect" title="Digital object identifier">doi</a>:<a rel="nofollow" class="external text" href="https://dx.doi.org/10.1093%2Femboj%2F19.21.5720">10.1093/emboj/19.21.5720</a></small>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=LC3%2C+a+mammalian+homologue+of+yeast+Apg8p%2C+is+localized+in+autophagosome+membranes+after+processing&rft.au=%C5%8Csumi%2C+Y.&rft.au=Kabeya%2C+Y.&rft.au=Kirisako%2C+T.&rft.au=Kominami%2C+E.&rft.au=Mizushima%2C+N.&rft.au=Noda%2C+T.&rft.au=Ueno%2C+T.&rft.au=Yamamoto%2C+A.&rft.au=Yoshimori%2C+T.&rft.aufirst=Y.&rft.aulast=Kabeya&rft.date=2000&rft.genre=article&rft.issue=21&rft.jtitle=The+EMBO+Journal&rft.pages=5720-5728&rft.volume=19&rft_id=%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC305793&rft_id=info%3Adoi%2F10.1093%2Femboj%2F19.21.5720&rft_id=info%3Apmc%2F305793&rft_id=info%3Apmid%2F11060023&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span> <span style="display:none;font-size:100%" class="error citation-comment">Se sugiere usar <code>|número-autores=</code> (<a href="/wiki/Ayuda:Errores_en_las_referencias#displayauthors" title="Ayuda:Errores en las referencias">ayuda</a>)</span></li> <li><span id="CITAREFIchimuraKirisakoTakaoSatomi23_de_noviembre_de_2000" class="citation publicación">Ichimura, Y; Kirisako, T; Takao, T; Satomi, Y; Shimonishi, Y; Ishihara, N; Mizushima, N; Tanida, I; Kominami, E; Ōsumi, M; Noda, T; Ōsumi, Y (23 de noviembre de 2000). «A ubiquitin-like system mediates protein lipidation». <i>Nature</i> <b>408</b> (6811): 488-92. <small><a href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/11100732">11100732</a></small>. <small><a href="/wiki/Digital_object_identifier" class="mw-redirect" title="Digital object identifier">doi</a>:<a rel="nofollow" class="external text" href="https://dx.doi.org/10.1038%2F35044114">10.1038/35044114</a></small>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=A+ubiquitin-like+system+mediates+protein+lipidation&rft.au=%C5%8Csumi%2C+M&rft.au=%C5%8Csumi%2C+Y&rft.au=Ichimura%2C+Y&rft.au=Ishihara%2C+N&rft.au=Kirisako%2C+T&rft.au=Kominami%2C+E&rft.au=Mizushima%2C+N&rft.au=Noda%2C+T&rft.au=Satomi%2C+Y&rft.au=Shimonishi%2C+Y&rft.au=Takao%2C+T&rft.au=Tanida%2C+I&rft.aufirst=Y&rft.aulast=Ichimura&rft.date=23+de+noviembre+de+2000&rft.genre=article&rft.issue=6811&rft.jtitle=Nature&rft.pages=488-92&rft.volume=408&rft_id=info%3Adoi%2F10.1038%2F35044114&rft_id=info%3Apmid%2F11100732&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span></li> <li><span id="CITAREFŌsumimarzo_de_2001" class="citation publicación">Ōsumi, Y (marzo de 2001). «Molecular dissection of autophagy: two ubiquitin-like systems». <i>Nature</i> <b>2</b> (3): 211-6. <small><a href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/11265251">11265251</a></small>. <small><a href="/wiki/Digital_object_identifier" class="mw-redirect" title="Digital object identifier">doi</a>:<a rel="nofollow" class="external text" href="https://dx.doi.org/10.1038%2F35044114">10.1038/35044114</a></small>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=Molecular+dissection+of+autophagy%3A+two+ubiquitin-like+systems&rft.au=%C5%8Csumi%2C+Y&rft.aufirst=Y&rft.aulast=%C5%8Csumi&rft.date=marzo+de+2001&rft.genre=article&rft.issue=3&rft.jtitle=Nature&rft.pages=211-6&rft.volume=2&rft_id=info%3Adoi%2F10.1038%2F35044114&rft_id=info%3Apmid%2F11265251&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span></li> <li><span id="CITAREFKumaHatanoMatsuiYamamoto23_de_diciembre_de_2004" class="citation publicación">Kuma, A; Hatano, M; Matsui, M; Yamamoto, A; Nakaya, H; Yoshimori, T; Ōsumi, Y; Tokuhisa, T <i>et al.</i> (23 de diciembre de 2004). «The role of autophagy during the early neonatal starvation period». <i>Nature</i> <b>432</b> (7020): 1032-6. <small><a href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/15525940">15525940</a></small>. <small><a href="/wiki/Digital_object_identifier" class="mw-redirect" title="Digital object identifier">doi</a>:<a rel="nofollow" class="external text" href="https://dx.doi.org/10.1038%2Fnature03029">10.1038/nature03029</a></small>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=The+role+of+autophagy+during+the+early+neonatal+starvation+period&rft.au=%C5%8Csumi%2C+Y&rft.au=Hatano%2C+M&rft.au=Kuma%2C+A&rft.au=Matsui%2C+M&rft.au=Mizushima%2C+N&rft.au=Nakaya%2C+H&rft.au=Tokuhisa%2C+T&rft.au=Yamamoto%2C+A&rft.au=Yoshimori%2C+T&rft.aufirst=A&rft.aulast=Kuma&rft.date=23+de+diciembre+de+2004&rft.genre=article&rft.issue=7020&rft.jtitle=Nature&rft.pages=1032-6&rft.volume=432&rft_id=info%3Adoi%2F10.1038%2Fnature03029&rft_id=info%3Apmid%2F15525940&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span> <span style="display:none;font-size:100%" class="error citation-comment">Se sugiere usar <code>|número-autores=</code> (<a href="/wiki/Ayuda:Errores_en_las_referencias#displayauthors" title="Ayuda:Errores en las referencias">ayuda</a>)</span></li> <li><span id="CITAREFHanadaNodaSatomiIchimura28_de_diciembre_de_2007" class="citation publicación">Hanada, T; Noda, NN; Satomi, Y; Ichimura, Y; Fujioka, Y; Takao, T; Inagaki, F; Ohsumi, Y (28 de diciembre de 2007). «The Atg12-Atg5 conjugate has a novel E3-like activity for protein lipidation in autophagy». <i>Journal of Biological Chemistry</i> <b>282</b> (52): 37298-302. <small><a href="/wiki/PubMed_Identifier" class="mw-redirect" title="PubMed Identifier">PMID</a> <a rel="nofollow" class="external text" href="//www.ncbi.nlm.nih.gov/pubmed/17986448">17986448</a></small>. <small><a href="/wiki/Digital_object_identifier" class="mw-redirect" title="Digital object identifier">doi</a>:<a rel="nofollow" class="external text" href="https://dx.doi.org/10.1074%2Fjbc.C700195200">10.1074/jbc.C700195200</a></small>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=The+Atg12-Atg5+conjugate+has+a+novel+E3-like+activity+for+protein+lipidation+in+autophagy&rft.au=Fujioka%2C+Y&rft.au=Hanada%2C+T&rft.au=Ichimura%2C+Y&rft.au=Inagaki%2C+F&rft.au=Noda%2C+NN&rft.au=Ohsumi%2C+Y&rft.au=Satomi%2C+Y&rft.au=Takao%2C+T&rft.aufirst=T&rft.aulast=Hanada&rft.date=28+de+diciembre+de+2007&rft.genre=article&rft.issue=52&rft.jtitle=Journal+of+Biological+Chemistry&rft.pages=37298-302&rft.volume=282&rft_id=info%3Adoi%2F10.1074%2Fjbc.C700195200&rft_id=info%3Apmid%2F17986448&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="Referencias">Referencias</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_Ohsumi&action=edit&section=3" title="Editar sección: Referencias"><span>editar</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="listaref" style="list-style-type: decimal;"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><span class="citation web"><a rel="nofollow" class="external text" href="http://www.titech.ac.jp/english/news/2016/036279.html">«Honorary Professor Yoshinori Ohsumi wins Nobel Prize in Physiology or Medicine for 2016»</a>. <i>titech.ac.jp</i> <span style="color:var(--color-subtle, #555 );">(en inglés)</span>. 3 de octubre de 2016<span class="reference-accessdate">. Consultado el 4 de octubre de 2016</span>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=Honorary+Professor+Yoshinori+Ohsumi+wins+Nobel+Prize+in+Physiology+or+Medicine+for+2016&rft.date=3+de+octubre+de+2016&rft.genre=article&rft.jtitle=titech.ac.jp&rft_id=http%3A%2F%2Fwww.titech.ac.jp%2Fenglish%2Fnews%2F2016%2F036279.html&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><span id="CITAREFBiemiller10_de_noviembre_de_2012" class="citation noticia">Biemiller, Lawrence (10 de noviembre de 2012). <a rel="nofollow" class="external text" href="http://www.chronicle.com/blogs/ticker/kyoto-prize-is-awarded-to-3-scholars/51696?cid=gn">«Kyoto Prize Is Awarded to 3 Scholars»</a>. <i>Chronicle</i><span class="reference-accessdate">. Consultado el 4 de octubre de 2016</span>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=Kyoto+Prize+Is+Awarded+to+3+Scholars&rft.au=Biemiller%2C+Lawrence&rft.aufirst=Lawrence&rft.aulast=Biemiller&rft.date=10+de+noviembre+de+2012&rft.genre=article&rft.jtitle=Chronicle&rft_id=http%3A%2F%2Fwww.chronicle.com%2Fblogs%2Fticker%2Fkyoto-prize-is-awarded-to-3-scholars%2F51696%3Fcid%3Dgn&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.elmundo.es/salud/2016/10/03/57f2193e268e3edd358b457d.html">«Premio Nobel de Medicina para Yoshinori Ōsumi, por sus hallazgos sobre el reciclaje celular.» 3 de octubre de 2016.</a> <i>El Mundo</i>.</span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><span class="citation web"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/2016/press.html">«The Nobel Prize in Physiology or Medicine 2016»</a>. <i>nobelprize.org</i> <span style="color:var(--color-subtle, #555 );">(en inglés)</span>. 3 de octubre de 2016<span class="reference-accessdate">. Consultado el 4 de octubre de 2016</span>.</span><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fes.wikipedia.org%3AYoshinori+Ohsumi&rft.atitle=The+Nobel+Prize+in+Physiology+or+Medicine+2016&rft.date=3+de+octubre+de+2016&rft.genre=article&rft.jtitle=nobelprize.org&rft_id=https%3A%2F%2Fwww.nobelprize.org%2Fnobel_prizes%2Fmedicine%2Flaureates%2F2016%2Fpress.html&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal" class="Z3988"><span style="display:none;"> </span></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Enlaces_externos">Enlaces externos</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yoshinori_Ohsumi&action=edit&section=4" title="Editar sección: Enlaces externos"><span>editar</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/15px-Commons-logo.svg.png" decoding="async" width="15" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/23px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span> <a href="/wiki/Wikimedia_Commons" title="Wikimedia Commons">Wikimedia Commons</a> alberga una categoría multimedia sobre <b><a href="https://commons.wikimedia.org/wiki/Category:Yoshinori_%C5%8Csumi" class="extiw" title="commons:Category:Yoshinori Ōsumi">Yoshinori Ohsumi</a></b>.</li> <li><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/24/Wikinews-logo.svg/20px-Wikinews-logo.svg.png" decoding="async" width="20" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/24/Wikinews-logo.svg/30px-Wikinews-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/24/Wikinews-logo.svg/40px-Wikinews-logo.svg.png 2x" data-file-width="759" data-file-height="415" /></span></span> <a href="/wiki/Wikinoticias" title="Wikinoticias">Wikinoticias</a> tiene noticias relacionadas con <b><a href="https://es.wikinews.org/wiki/Special:Search/Otorgan_el_Nobel_de_Medicina_a_Yoshinori_Ohsumi" class="extiw" title="n:Special:Search/Otorgan el Nobel de Medicina a Yoshinori Ohsumi">Yoshinori Ohsumi</a></b>.</li> <li><a rel="nofollow" class="external text" href="http://search.star.titech.ac.jp/titech-ss/pursuer.act?event=outside&key_rid=5000032417&lang=en">Perfil académico</a> (en inglés).</li> <li><a rel="nofollow" class="external text" href="http://www.ohsumilab.aro.iri.titech.ac.jp/ohsumieng.html">Currículo</a> en su página web (en inglés).</li></ul> <style data-mw-deduplicate="TemplateStyles:r161257576">.mw-parser-output .mw-authority-control{margin-top:1.5em}.mw-parser-output .mw-authority-control .navbox 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style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikidata" title="Wikidata"><img alt="Wd" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/20px-Wikidata-logo.svg.png" decoding="async" width="20" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/30px-Wikidata-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/40px-Wikidata-logo.svg.png 2x" data-file-width="1050" data-file-height="590" /></a></span> Datos:</span> <span class="uid"><a href="https://www.wikidata.org/wiki/Q8056214" class="extiw" title="wikidata:Q8056214">Q8056214</a></span></li> <li><span style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikimedia_Commons" title="Commonscat"><img alt="Commonscat" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/15px-Commons-logo.svg.png" decoding="async" width="15" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/23px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span> Multimedia:</span> <span class="uid"><span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Category:Yoshinori_Ōsumi">Yoshinori Ōsumi</a></span> / <span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Special:MediaSearch?type=image&search=%22Q8056214%22">Q8056214</a></span></span></li></ul> <hr /> <ul><li><b>Identificadores</b></li> <li><a rel="nofollow" class="external text" href="//www.worldcat.org/identities/viaf-252902927">WorldCat</a></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fichero_de_Autoridades_Virtual_Internacional" title="Fichero de Autoridades Virtual Internacional">VIAF</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/252902927/">252902927</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/International_Standard_Name_Identifier" title="International Standard Name Identifier">ISNI</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/000000037593330X">000000037593330X</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Gemeinsame_Normdatei" title="Gemeinsame Normdatei">GND</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/1089644000">1089644000</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Library_of_Congress_Control_Number" title="Library of Congress Control Number">LCCN</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n2008083589">n2008083589</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Biblioteca_Nacional_de_la_Dieta" title="Biblioteca Nacional de la Dieta">NDL</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00204204">00204204</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/CiNii" title="CiNii">CiNii</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://ci.nii.ac.jp/author/DA04729130">DA04729130</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/ORCID" title="ORCID">ORCID</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://orcid.org/0000-0003-2384-2166">0000-0003-2384-2166</a></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/ResearcherID" title="ResearcherID">ResearcherID</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://www.webofscience.com/wos/author/record/C-6449-2009">C-6449-2009</a></span></li> <li><b>Diccionarios y enciclopedias</b></li> <li><span style="white-space:nowrap;"><a href="/wiki/Enciclopedia_Brit%C3%A1nica" title="Enciclopedia Británica">Britannica</a>:</span> <span class="uid"><a rel="nofollow" class="external text" href="https://www.britannica.com/biography/Yoshinori-Ohsumi">url</a></span></li></ul> </div></td></tr></tbody></table></div><div class="mw-mf-linked-projects hlist"> <ul><li><span style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikidata" title="Wikidata"><img alt="Wd" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/20px-Wikidata-logo.svg.png" decoding="async" width="20" height="11" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/30px-Wikidata-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Wikidata-logo.svg/40px-Wikidata-logo.svg.png 2x" data-file-width="1050" data-file-height="590" /></a></span> Datos:</span> <span class="uid"><a href="https://www.wikidata.org/wiki/Q8056214" class="extiw" title="wikidata:Q8056214">Q8056214</a></span></li> <li><span style="white-space:nowrap;"><span typeof="mw:File"><a href="/wiki/Wikimedia_Commons" title="Commonscat"><img alt="Commonscat" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/15px-Commons-logo.svg.png" decoding="async" width="15" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/23px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span> Multimedia:</span> <span class="uid"><span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Category:Yoshinori_Ōsumi">Yoshinori Ōsumi</a></span> / <span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Special:MediaSearch?type=image&search=%22Q8056214%22">Q8056214</a></span></span></li></ul> </div></div>   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