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href="https://cs.wikipedia.org/wiki/Heteroceridae" title="Heteroceridae – Czech" lang="cs" hreflang="cs" data-title="Heteroceridae" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/S%C3%A4gek%C3%A4fer" title="Sägekäfer – German" lang="de" hreflang="de" data-title="Sägekäfer" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/Ch%C3%A9%C5%82chaa%CA%BC_nahashch%CA%BCid%C3%ADg%C3%AD%C3%AD" title="Chéłchaaʼ nahashchʼidígíí – Navajo" lang="nv" hreflang="nv" data-title="Chéłchaaʼ nahashchʼidígíí" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Heteroceridae" title="Heteroceridae – Italian" lang="it" hreflang="it" data-title="Heteroceridae" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Pj%C5%ABklavabaliai" title="Pjūklavabaliai – Lithuanian" lang="lt" hreflang="lt" data-title="Pjūklavabaliai" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Iszapbog%C3%A1rf%C3%A9l%C3%A9k" title="Iszapbogárfélék – Hungarian" lang="hu" hreflang="hu" data-title="Iszapbogárfélék" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Oevergraafkevers" title="Oevergraafkevers – Dutch" lang="nl" hreflang="nl" data-title="Oevergraafkevers" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Strandgravere" title="Strandgravere – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Strandgravere" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D0%B8%D0%BB%D0%BE%D1%83%D1%81%D1%8B" title="Пилоусы – Russian" lang="ru" hreflang="ru" data-title="Пилоусы" data-language-autonym="Русский" 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width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Variegated mud-loving beetles<br /><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">Aptian–Recent</span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:178.91076923077px; width:41.089230769231px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:178.91076923077px; width:41.089230769231px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:179.91076923077px; width:39.089230769231px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:COLE_Heteroceridae_Heterocerus_novaeselandiae.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/03/COLE_Heteroceridae_Heterocerus_novaeselandiae.png/220px-COLE_Heteroceridae_Heterocerus_novaeselandiae.png" decoding="async" width="220" height="262" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/03/COLE_Heteroceridae_Heterocerus_novaeselandiae.png/330px-COLE_Heteroceridae_Heterocerus_novaeselandiae.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/03/COLE_Heteroceridae_Heterocerus_novaeselandiae.png/440px-COLE_Heteroceridae_Heterocerus_novaeselandiae.png 2x" data-file-width="3125" data-file-height="3726" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i><a href="/w/index.php?title=Heterocerus_novaeselandiae&action=edit&redlink=1" class="new" title="Heterocerus novaeselandiae (page does not exist)">Heterocerus novaeselandiae</a></i> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Heteroceridae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Insect" title="Insect">Insecta</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Beetle" title="Beetle">Coleoptera</a> </td></tr> <tr> <td>Suborder: </td> <td><a href="/wiki/Polyphaga" title="Polyphaga">Polyphaga</a> </td></tr> <tr> <td>Infraorder: </td> <td><a href="/wiki/Elateriformia" title="Elateriformia">Elateriformia</a> </td></tr> <tr> <td>Superfamily: </td> <td><a href="/wiki/Byrrhoidea" title="Byrrhoidea">Byrrhoidea</a> </td></tr> <tr> <td>Family: </td> <td><a class="mw-selflink selflink">Heteroceridae</a><br /><small>MacLeay, 1825</small> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Heteroceridae</b>, the variegated mud-loving beetles, are a widespread and relatively common family of <a href="/wiki/Beetle" title="Beetle">beetles</a> found on every continent except for <a href="/wiki/Antarctica" title="Antarctica">Antarctica</a>. </p><p>Around two hundred and fifty species of heterocerids are known to occur worldwide.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> They are most diverse in tropical and sub-tropical regions. Currently, 87 species are known from the New World,<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> including 34 from the United States.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>Variegated mud-loving beetles are brownish, dorsoventrally depressed shoreline inhabitants. Superficially they resemble small <a href="/wiki/Scarabaeidae" title="Scarabaeidae">scarabs</a> with the tibiae armed with rows of robust flattened spines. The beetles live in shallow tunnels that they dig in damp soil around fresh and brackish lakes, rivers and ponds. Heterocerids have been reported to live in intertidal sandflats<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> and on remote oceanic islands.<sup id="cite_ref-:0_5-0" class="reference"><a href="#cite_note-:0-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> The uniform way in which they live seems to have favored the conservation of a "phenotypical uniformity in external morphology".<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> Consequently, it is often quite difficult to identify one of these beetles to species relying on external morphology alone. Therefore, male genitalia are most often relied upon to identify species. Although few studies have been conducted on their ecology, heterocerids have been shown to be an important prey group for <a href="/wiki/Passerine" title="Passerine">passerine</a> birds and frogs (Schmidt et al., 2003; Turner, 1959), and they appear to play a significant role in seed dispersal and burial in sandy soils.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> They are thought to be <a href="/wiki/Detritivore" title="Detritivore">detritivores</a>, consuming the substrate to sift for organic matter, microorganisms, and algae.<sup id="cite_ref-:0_5-1" class="reference"><a href="#cite_note-:0-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>In the most recent revision of the family, <a href="/wiki/Francisco_Pacheco" title="Francisco Pacheco">Francisco Pacheco</a> (1964) split the heterocerids into 20 different genera, erecting 17 new generic names based almost entirely on the features of male <a href="/wiki/Genitalia" class="mw-redirect" title="Genitalia">genitalia</a>. Most <a href="/wiki/Systematics" title="Systematics">systematists</a> have found Pacheco's system complex and impractical because the ratio of genera to species is high and females cannot be identified. Consequently, his generic circumscriptions have not been widely accepted.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Heteroceridae&action=edit&section=1" title="Edit section: Taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Thomas_Say" title="Thomas Say">Thomas Say</a> (1823) described <i>Heterocerus pallidus</i> and <i>Heterocerus pusillus</i>, the first New World species in the family. By the time <a href="/wiki/George_H._Horn" class="mw-redirect" title="George H. Horn">George H. Horn</a> (1890) produced his key to the genus <i>Heterocerus</i>, there were eleven described North American species in two genera. Horn's key relied heavily on general coloration and elytral pattern for species diagnosis. No other major taxonomic work concerning Heteroceridae was published until 1964, when Pacheco published his dissertation on the systematics, phylogeny and distribution of the family in the New World. Pacheco described 25 additional New World species. He produced worldwide (excluding Africa) identification keys based largely on characters of the male genitalia, leaving it impossible to identify females to species. Pacheco also split the heterocerids into 20 different genera, erecting 17 new generic names, once again relying heavily on characters of the male genitalia. Distribution maps are included in Pacheco's work; however, in his introduction, Pacheco admits that his work did not include enough material to create particularly informative distributions for many New World heterocerids. The only other monograph on the family Heteroceridae was published by <a href="/w/index.php?title=Reinhold_Charpentier&action=edit&redlink=1" class="new" title="Reinhold Charpentier (page does not exist)">Reinhold Charpentier</a> (1965), and this encompassed the species of the Ethiopian region. Charpentier's work included two species level keys, one based on male genitalia, the other on external characteristics. Charpentier saw no justification in dividing the family into any new taxa and he retained all 35 Ethiopian species in the genus <i>Heterocerus</i>. He did note, however, the great difference between the male genitalia of various groups of species. </p><p>Little subsequent taxonomic work was conducted with the family until the late 1980s, when W. V. Miller began describing species from around the world (Miller 1988, 1992). Miller's work included the descriptions of seven new species from North America. These additions brought the number of North American species to 34. A key to 21 northeastern species, produced by Miller, appeared in Downie and Arnett's <i>Beetles of Northeastern North America</i> (1995). This key relies mostly on elytral color patterns for species identification. In his work, Miller chose to apply a taxonomic scheme in which the heterocerids are divided into only five genera worldwide. European authors, namely S. Skalický (Czech Republic) and A. Mascagni (Italy), have described numerous species in the last ten years. </p><p>The oldest fossils of the genus are from the Early Cretaceous (<a href="/wiki/Aptian" title="Aptian">Aptian</a>) of China and Mongolia, belonging to the genus <i><a href="/w/index.php?title=Heterocerites&action=edit&redlink=1" class="new" title="Heterocerites (page does not exist)">Heterocerites</a>.</i><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Genera">Genera</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Heteroceridae&action=edit&section=2" title="Edit section: Genera"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>These 15 genera belong to the family Heteroceridae: </p> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 22em;"> <ul><li><i><a href="/wiki/Augyles" title="Augyles">Augyles</a></i> <small>Schiødte, 1866</small><span style="color:gray"><sup> i c g b</sup></span></li> <li><i><a href="/w/index.php?title=Centuriatus&action=edit&redlink=1" class="new" title="Centuriatus (page does not exist)">Centuriatus</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Efflagitatus&action=edit&redlink=1" class="new" title="Efflagitatus (page does not exist)">Efflagitatus</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Explorator&action=edit&redlink=1" class="new" title="Explorator (page does not exist)">Explorator</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Heterocerites&action=edit&redlink=1" class="new" title="Heterocerites (page does not exist)">Heterocerites</a></i> <small>Ponomarenko, 1986</small><span style="color:gray"><sup> g</sup></span></li> <li><i><a href="/wiki/Heterocerus" title="Heterocerus">Heterocerus</a></i> <small>Fabricius, 1792</small><span style="color:gray"><sup> i c g b</sup></span></li> <li><i><a href="/w/index.php?title=Lanternarius&action=edit&redlink=1" class="new" title="Lanternarius (page does not exist)">Lanternarius</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Lapsus_(beetle)&action=edit&redlink=1" class="new" title="Lapsus (beetle) (page does not exist)">Lapsus</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Littorimus&action=edit&redlink=1" class="new" title="Littorimus (page does not exist)">Littorimus</a></i><span style="color:gray"><sup> g</sup></span></li> <li><i><a href="/w/index.php?title=Micilus&action=edit&redlink=1" class="new" title="Micilus (page does not exist)">Micilus</a></i> <small>Mulsant & Rey, 1872</small><span style="color:gray"><sup> g</sup></span></li> <li><i><a href="/w/index.php?title=Microaugyles&action=edit&redlink=1" class="new" title="Microaugyles (page does not exist)">Microaugyles</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Neoheterocerus&action=edit&redlink=1" class="new" title="Neoheterocerus (page does not exist)">Neoheterocerus</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Peditatus&action=edit&redlink=1" class="new" title="Peditatus (page does not exist)">Peditatus</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g</sup></span></li> <li><i><a href="/w/index.php?title=Protoallopygmephorus&action=edit&redlink=1" class="new" title="Protoallopygmephorus (page does not exist)">Protoallopygmephorus</a></i><span style="color:gray"><sup> g</sup></span></li> <li><i><a href="/wiki/Tropicus" title="Tropicus">Tropicus</a></i> <small>Pacheco, 1964</small><span style="color:gray"><sup> i c g b</sup></span></li></ul> </div> <p><small>Data sources: i = ITIS,<sup id="cite_ref-itis_10-0" class="reference"><a href="#cite_note-itis-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> c = Catalogue of Life,<sup id="cite_ref-catlife_11-0" class="reference"><a href="#cite_note-catlife-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> g = GBIF,<sup id="cite_ref-gbif_12-0" class="reference"><a href="#cite_note-gbif-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> b = Bugguide.net<sup id="cite_ref-bugref_13-0" class="reference"><a href="#cite_note-bugref-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup></small> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Heteroceridae&action=edit&section=3" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text">Bamuel, 1995; Charpentier, 1965; Ivie, 1984; Mascogni, all works listed; Miller, all works listed; Pacheco, 1964, 1969; Skalický, all works listed</span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text">Bamuel, 1995; Ivie, 1984; Mascogni, 1988, 1989, 1994; Miller, all works listed; Pacheco, 1964, 1969</span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text">Marske and Ivie, 2003</span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text">Good, 1999</span> </li> <li id="cite_note-:0-5"><span class="mw-cite-backlink">^ <a href="#cite_ref-:0_5-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:0_5-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFSazhnev2020" class="citation journal cs1">Sazhnev, Alexey S. (2020). <a rel="nofollow" class="external text" href="http://en.ecosysttrans.com/publikatsii/ecosystem-transformation-volume-3-no-2-2020/beetles-of-the-family-heteroceridae-insecta-coleoptera-in-extreme-environments/">"Beetles of the family Heteroceridae (Insecta: Coleoptera) in extreme environments"</a>. <i>Ecosystem Transformation</i>. <b>3</b> (2). <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.23859%2Festr-200323a">10.23859/estr-200323a</a></span>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:226725701">226725701</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Ecosystem+Transformation&rft.atitle=Beetles+of+the+family+Heteroceridae+%28Insecta%3A+Coleoptera%29+in+extreme+environments&rft.volume=3&rft.issue=2&rft.date=2020&rft_id=info%3Adoi%2F10.23859%2Festr-200323a&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A226725701%23id-name%3DS2CID&rft.aulast=Sazhnev&rft.aufirst=Alexey+S.&rft_id=http%3A%2F%2Fen.ecosysttrans.com%2Fpublikatsii%2Fecosystem-transformation-volume-3-no-2-2020%2Fbeetles-of-the-family-heteroceridae-insecta-coleoptera-in-extreme-environments%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AHeteroceridae" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text">Charpentier, 1965</span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text">Bernhardt, 1995</span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text">see Arnett 2002</span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFProkinRen2011" class="citation journal cs1">Prokin, A. A.; Ren, D. (May 2011). <a rel="nofollow" class="external text" href="http://link.springer.com/10.1134/S003103011102016X">"New species of variegated mud-loving beetles (Coleoptera: Heteroceridae) from mesozoic deposits of China"</a>. <i>Paleontological Journal</i>. <b>45</b> (3): 284–286. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2011PalJ...45..284P">2011PalJ...45..284P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1134%2FS003103011102016X">10.1134/S003103011102016X</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0031-0301">0031-0301</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:129824564">129824564</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Paleontological+Journal&rft.atitle=New+species+of+variegated+mud-loving+beetles+%28Coleoptera%3A+Heteroceridae%29+from+mesozoic+deposits+of+China&rft.volume=45&rft.issue=3&rft.pages=284-286&rft.date=2011-05&rft_id=info%3Adoi%2F10.1134%2FS003103011102016X&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A129824564%23id-name%3DS2CID&rft.issn=0031-0301&rft_id=info%3Abibcode%2F2011PalJ...45..284P&rft.aulast=Prokin&rft.aufirst=A.+A.&rft.au=Ren%2C+D.&rft_id=http%3A%2F%2Flink.springer.com%2F10.1134%2FS003103011102016X&rfr_id=info%3Asid%2Fen.wikipedia.org%3AHeteroceridae" class="Z3988"></span></span> </li> <li id="cite_note-itis-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-itis_10-0">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=114262">"Heteroceridae Report"</a>. <i>Integrated Taxonomic Information System</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2018-05-06</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Integrated+Taxonomic+Information+System&rft.atitle=Heteroceridae+Report&rft_id=https%3A%2F%2Fwww.itis.gov%2Fservlet%2FSingleRpt%2FSingleRpt%3Fsearch_topic%3DTSN%26search_value%3D114262&rfr_id=info%3Asid%2Fen.wikipedia.org%3AHeteroceridae" class="Z3988"></span></span> </li> <li id="cite_note-catlife-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-catlife_11-0">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.catalogueoflife.org/col/browse/tree/id/0fa366749483b11a00cd4414413bc5b4">"Browse Heteroceridae"</a>. <i>Catalogue of Life</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2018-05-06</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Catalogue+of+Life&rft.atitle=Browse+Heteroceridae&rft_id=http%3A%2F%2Fwww.catalogueoflife.org%2Fcol%2Fbrowse%2Ftree%2Fid%2F0fa366749483b11a00cd4414413bc5b4&rfr_id=info%3Asid%2Fen.wikipedia.org%3AHeteroceridae" class="Z3988"></span></span> </li> <li id="cite_note-gbif-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-gbif_12-0">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/6986">"Heteroceridae"</a>. <i>GBIF</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2018-05-06</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=GBIF&rft.atitle=Heteroceridae&rft_id=https%3A%2F%2Fwww.gbif.org%2Fspecies%2F6986&rfr_id=info%3Asid%2Fen.wikipedia.org%3AHeteroceridae" class="Z3988"></span></span> </li> <li id="cite_note-bugref-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-bugref_13-0">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://bugguide.net/node/view/30091">"Heteroceridae Family Information"</a>. <i>BugGuide.net</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2018-05-06</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=BugGuide.net&rft.atitle=Heteroceridae+Family+Information&rft_id=https%3A%2F%2Fbugguide.net%2Fnode%2Fview%2F30091&rfr_id=info%3Asid%2Fen.wikipedia.org%3AHeteroceridae" class="Z3988"></span></span> </li> </ol></div></div> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output 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href="/wiki/Special:EditPage/Template:Coleoptera" title="Special:EditPage/Template:Coleoptera"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Extant_Coleoptera_families" style="font-size:114%;margin:0 4em">Extant <a href="/wiki/Beetle" title="Beetle">Coleoptera</a> families</div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a></li> <li>Class: <a href="/wiki/Insect" title="Insect">Insecta</a></li> <li>Subclass: <a href="/wiki/Pterygota" title="Pterygota">Pterygota</a></li> <li>Infraclass: <a href="/wiki/Neoptera" title="Neoptera">Neoptera</a></li> <li>Superorder: <a href="/wiki/Holometabola" title="Holometabola">Holometabola</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Archostemata" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Archostemata" title="Archostemata">Archostemata</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Crowsoniella" title="Crowsoniella">Crowsoniellidae <small>(<i>Crowsoniella relicta</i>)</small></a></li> <li><a href="/wiki/Cupedidae" title="Cupedidae">Cupedidae <small>(reticulated beetles)</small></a></li> <li><a href="/wiki/Jurodidae" title="Jurodidae">Jurodidae <small>(<i>Sikhotealinia zhiltzovae</i>)</small></a></li> <li><a href="/wiki/Micromalthidae" title="Micromalthidae">Micromalthidae</a></li> <li><a href="/wiki/Ommatidae" title="Ommatidae">Ommatidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Adephaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Adephaga" title="Adephaga">Adephaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Extant_families" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Extant_taxon" class="mw-redirect" title="Extant taxon">Extant</a> families</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amphizoa" title="Amphizoa">Amphizoidae <small>(trout-stream beetles)</small></a></li> <li><a href="/wiki/Aspidytidae" title="Aspidytidae">Aspidytidae</a></li> <li><a href="/wiki/Ground_beetle" title="Ground beetle">Carabidae <small>(ground beetles)</small></a></li> <li><a href="/wiki/Tiger_beetle" title="Tiger beetle">Cicindelidae <small>(tiger beetles)</small></a></li> <li><a href="/wiki/Dytiscidae" title="Dytiscidae">Dytiscidae <small>(predaceous diving beetles)</small></a></li> <li><a href="/wiki/Whirligig_beetle" title="Whirligig beetle">Gyrinidae <small>(whirligig beetles)</small></a></li> <li><a href="/wiki/Haliplidae" title="Haliplidae">Haliplidae <small>(crawling water beetles)</small></a></li> <li><a href="/wiki/Hygrobia" title="Hygrobia">Hygrobiidae</a></li> <li><a href="/wiki/Meru_phyllisae" title="Meru phyllisae">Meruidae <small>(<i>Meru phyllisae</i>)</small></a></li> <li><a href="/wiki/Noteridae" title="Noteridae">Noteridae <small>(burrowing water beetles)</small></a></li> <li><a href="/wiki/Trachypachidae" title="Trachypachidae">Trachypachidae <small>(false ground beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Myxophaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Myxophaga" title="Myxophaga">Myxophaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hydroscaphidae" title="Hydroscaphidae">Hydroscaphidae <small>(skiff beetles)</small></a></li> <li><a href="/wiki/Lepicerus" title="Lepicerus">Lepiceridae</a></li> <li><a href="/wiki/Sphaerius" title="Sphaerius">Sphaeriusidae</a></li> <li><a href="/wiki/Torridincolidae" title="Torridincolidae">Torridincolidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Polyphaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Polyphaga" title="Polyphaga">Polyphaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bostrichiformia" title="Bostrichiformia">Bostrichiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Bostrichoidea" title="Bostrichoidea">Bostrichoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bostrichidae" title="Bostrichidae">Bostrichidae <small>(auger beetles)</small></a></li> <li><a href="/wiki/Dermestidae" title="Dermestidae">Dermestidae <small>(skin beetles)</small></a></li> <li><a href="/wiki/Endecatomus" title="Endecatomus">Endecatomidae</a></li> <li><a href="/wiki/Jacobsoniidae" title="Jacobsoniidae">Jacobsoniidae <small>(Jacobson's beetles)</small></a></li> <li><a href="/wiki/Nosodendridae" title="Nosodendridae">Nosodendridae <small>(wounded-tree beetles)</small></a></li> <li><a href="/wiki/Ptinidae" title="Ptinidae">Ptiniidae <small>(furniture beetles, death watch beetles, spider beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Derodontidae" title="Derodontidae">Derodontoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Derodontidae" title="Derodontidae">Derodontidae <small>(tooth-necked fungus beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Chrysomeloidea" title="Chrysomeloidea">Chrysomeloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Longhorn_beetle" title="Longhorn beetle">Cerambycidae <small>(longhorn beetles)</small></a></li> <li><a href="/wiki/Leaf_beetle" title="Leaf beetle">Chrysomelidae <small>(leaf beetles)</small></a></li> <li><a href="/wiki/Disteniidae" title="Disteniidae">Disteniidae</a></li> <li><a href="/wiki/Megalopodidae" title="Megalopodidae">Megalopodidae</a></li> <li><a href="/wiki/Orsodacnidae" title="Orsodacnidae">Orsodacnidae</a></li> <li><a href="/wiki/Oxypeltidae" title="Oxypeltidae">Oxypeltidae</a></li> <li><a href="/wiki/Vesperidae" title="Vesperidae">Vesperidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cleroidea" title="Cleroidea">Cleroidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acanthocnemus" title="Acanthocnemus">Acanthocnemidae <small>(<i>Acanthocnemus nigricans</i>)</small></a></li> <li><a href="/wiki/Biphyllidae" title="Biphyllidae">Biphyllidae <small>(false skin beetles)</small></a></li> <li><a href="/wiki/Byturidae" title="Byturidae">Byturidae <small>(fruitworm beetles)</small></a></li> <li><a href="/wiki/Chaetosomatidae" title="Chaetosomatidae">Chaetosomatidae</a></li> <li><a href="/wiki/Cleridae" title="Cleridae">Cleridae <small>(checkered beetles)</small></a></li> <li><a href="/wiki/Lophocateridae" title="Lophocateridae">Lophocateridae</a></li> <li><a href="/wiki/Mauroniscidae" title="Mauroniscidae">Mauroniscidae</a></li> <li><a href="/wiki/Melyridae" title="Melyridae">Melyridae <small>(soft-wing flower beetles)</small></a></li> <li><a href="/wiki/Metaxina" title="Metaxina">Metaxinidae <small>(<i>Metaxina ornata</i>)</small></a></li> <li><a href="/wiki/Peltis" title="Peltis">Peltidae</a></li> <li><a href="/wiki/Phloiophilus" title="Phloiophilus">Phloiophilidae <small>(<i>Phloiophilus edwardsi</i>)</small></a></li> <li><a href="/wiki/Phycosecidae" title="Phycosecidae">Phycosecidae</a></li> <li><a href="/wiki/Prionoceridae" title="Prionoceridae">Prionoceridae</a></li> <li><a href="/wiki/Protopeltis" title="Protopeltis">Protopeltidae</a></li> <li><a href="/wiki/Rentoniidae" title="Rentoniidae">Rentoniidae</a></li> <li><a href="/wiki/Rhadalidae" title="Rhadalidae">Rhadalidae</a></li> <li><a href="/wiki/Thanerocleridae" title="Thanerocleridae">Thanerocleridae</a></li> <li><a href="/wiki/Thymalidae" title="Thymalidae">Thymalidae</a></li> <li><a href="/wiki/Trogossitidae" title="Trogossitidae">Trogossitidae <small>(bark-gnawing beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Coccinelloidea" title="Coccinelloidea">Coccinelloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Akalyptoischion" title="Akalyptoischion">Akalyptoischiidae</a></li> <li><a href="/wiki/Alexiidae" title="Alexiidae">Alexiidae</a></li> <li><a href="/wiki/Anamorphidae" title="Anamorphidae">Anamorphidae</a></li> <li><a href="/wiki/Bothrideridae" title="Bothrideridae">Bothrideridae <small>(dry bark beetles)</small></a></li> <li><a href="/wiki/Cerylonidae" title="Cerylonidae">Cerylonidae <small>(minute bark beetles)</small></a></li> <li><a href="/wiki/Coccinellidae" title="Coccinellidae">Coccinellidae <small>(lady beetles, or God's cows)</small></a></li> <li><a href="/wiki/Corylophidae" title="Corylophidae">Corylophidae <small>(minute fungus beetles)</small></a></li> <li><a href="/wiki/Discolomatidae" title="Discolomatidae">Discolomatidae</a></li> <li><a href="/wiki/Endomychidae" title="Endomychidae">Endomychidae <small>(handsome fungus beetles)</small></a></li> <li><a href="/wiki/Eupsilobiidae" title="Eupsilobiidae">Eupsilobiidae</a></li> <li><a href="/wiki/Euxestidae" title="Euxestidae">Euxestidae <small>(well polished beetles)</small></a></li> <li><a href="/wiki/Latridiidae" title="Latridiidae">Latridiidae <small>(minute brown scavenger beetles)</small></a></li> <li><a href="/wiki/Murmidiidae" title="Murmidiidae">Murmidiidae</a></li> <li><a href="/wiki/Mycetaeidae" title="Mycetaeidae">Mycetaeidae</a></li> <li><a href="/wiki/Teredidae" title="Teredidae">Teredidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cucujoidea" title="Cucujoidea">Cucujoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agapytho" title="Agapytho">Agapythidae <small>(<i>Agapytho foveicollis</i>)</small></a></li> <li><a href="/wiki/Boganiidae" title="Boganiidae">Boganiidae</a></li> <li><a href="/wiki/Cavognathidae" title="Cavognathidae">Cavognathidae</a></li> <li><a href="/wiki/Cryptophagidae" title="Cryptophagidae">Cryptophagidae <small>(silken fungus beetles)</small></a></li> <li><a href="/wiki/Cucujidae" title="Cucujidae">Cucujidae <small>(flat bark beetles)</small></a></li> <li><a href="/wiki/Cybocephalidae" title="Cybocephalidae">Cybocephalidae</a></li> <li><a href="/wiki/Cyclaxyridae" title="Cyclaxyridae">Cyclaxyridae</a></li> <li><a href="/wiki/Erotylidae" title="Erotylidae">Erotylidae <small>(pleasing fungus beetles)</small></a></li> <li><a href="/wiki/Helotidae" title="Helotidae">Helotidae</a></li> <li><a href="/wiki/Hobartiidae" title="Hobartiidae">Hobartiidae</a></li> <li><a href="/wiki/Kateretidae" title="Kateretidae">Kateretidae <small>(short-winged flower beetles)</small></a></li> <li><a href="/wiki/Laemophloeidae" title="Laemophloeidae">Laemophloeidae <small>(lined flat bark beetles)</small></a></li> <li><a href="/wiki/Lamingtonium" title="Lamingtonium">Lamingtoniidae <small>(<i>Lamingtonium binnaberrense</i>)</small></a></li> <li><a href="/wiki/Monotomidae" title="Monotomidae">Monotomidae <small>(root-eating beetles)</small></a></li> <li><a href="/wiki/Myraboliidae" class="mw-redirect" title="Myraboliidae">Myraboliidae</a></li> <li><a href="/wiki/Sap_beetle" title="Sap beetle">Nitidulidae <small>(sap beetles)</small></a></li> <li><a href="/wiki/Passandridae" title="Passandridae">Passandridae <small>(parasitic flat bark beetles)</small></a></li> <li><a href="/wiki/Phalacridae" title="Phalacridae">Phalacridae <small>(shining flower beetles)</small></a></li> <li><a href="/wiki/Phloeostichidae" title="Phloeostichidae">Phloeostichidae</a></li> <li><a href="/wiki/Priasilphidae" title="Priasilphidae">Priasilphidae</a></li> <li><a href="/wiki/Propalticidae" title="Propalticidae">Propalticidae</a></li> <li><a href="/wiki/Protocucujidae" title="Protocucujidae">Protocucujidae</a></li> <li><a href="/wiki/Silvanidae" title="Silvanidae">Silvanidae <small>(silvanid flat bark beetles)</small></a></li> <li><a href="/wiki/Smicripidae" title="Smicripidae">Smicripidae <small>(palmetto beetles)</small></a></li> <li><a href="/wiki/Sphindidae" title="Sphindidae">Sphindidae <small>(dry-fungus beetles)</small></a></li> <li><a href="/wiki/Tasmosalpingus" title="Tasmosalpingus">Tasmosalpingidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Weevil" title="Weevil">Curculionoidea</a><br /><small><span style="color:#696969"><span class="nobold">(weevils)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anthribidae" title="Anthribidae">Anthribidae <small>(fungus weevils)</small></a></li> <li><a href="/wiki/Attelabidae" title="Attelabidae">Attelabidae <small>(leaf-rolling weevils)</small></a></li> <li><a href="/wiki/Belidae" title="Belidae">Belidae <small>(primitive weevils)</small></a></li> <li><a href="/wiki/Brentidae" title="Brentidae">Brentidae <small>(straight snout weevils, New York weevil)</small></a></li> <li><a href="/wiki/Caridae" title="Caridae">Caridae</a></li> <li><a href="/wiki/Curculionidae" title="Curculionidae">Curculionidae <small>(true weevils, bark beetles, ambrosia beetles)</small></a></li> <li><a href="/wiki/Nemonychidae" title="Nemonychidae">Nemonychidae <small>(pine flower weevils)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexyloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexylidae <small>(ship-timber beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Tenebrionoidea" title="Tenebrionoidea">Tenebrionoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aderidae" title="Aderidae">Aderidae <small>(ant-like leaf beetles)</small></a></li> <li><a href="/wiki/Anthicidae" title="Anthicidae">Anthicidae <small>(ant-like flower beetles)</small></a></li> <li><a href="/wiki/Archeocrypticidae" title="Archeocrypticidae">Archeocrypticidae <small>(cryptic fungus beetles)</small></a></li> <li><a href="/wiki/Boridae" title="Boridae">Boridae <small>(conifer bark beetles)</small></a></li> <li><a href="/wiki/Chalcodryidae" title="Chalcodryidae">Chalcodryidae</a></li> <li><a href="/wiki/Ciidae" title="Ciidae">Ciidae <small>(minute tree-fungus beetles)</small></a></li> <li><a href="/wiki/Melandryidae" title="Melandryidae">Melandryidae <small>(false darkling beetles)</small></a></li> <li><a href="/wiki/Blister_beetle" title="Blister beetle">Meloidae <small>(blister beetles)</small></a></li> <li><a href="/wiki/Mordellidae" title="Mordellidae">Mordellidae <small>(tumbling flower beetles)</small></a></li> <li><a href="/wiki/Mycetophagidae" title="Mycetophagidae">Mycetophagidae <small>(hairy fungus beetles)</small></a></li> <li><a href="/wiki/Mycteridae" title="Mycteridae">Mycteridae <small>(palm and flower beetles)</small></a></li> <li><a href="/wiki/Oedemeridae" title="Oedemeridae">Oedemeridae <small>(false blister beetle)</small></a></li> <li><a href="/wiki/Perimylopidae" class="mw-redirect" title="Perimylopidae">Perimylopidae, or Promecheilidae</a></li> <li><a href="/wiki/Prostomidae" title="Prostomidae">Prostomidae <small>(jugular-horned beetles)</small></a></li> <li><a href="/wiki/Pterogeniidae" title="Pterogeniidae">Pterogeniidae</a></li> <li><a href="/wiki/Fire-coloured_beetle" title="Fire-coloured beetle">Pyrochroidae <small>(fire-coloured beetles)</small></a></li> <li><a href="/wiki/Pythidae" title="Pythidae">Pythidae <small>(dead log bark beetles)</small></a></li> <li><a href="/wiki/Ripiphoridae" title="Ripiphoridae">Ripiphoridae <small>(wedge-shaped beetles)</small></a></li> <li><a href="/wiki/Salpingidae" title="Salpingidae">Salpingidae <small>(narrow-waisted bark beetles)</small></a></li> <li><a href="/wiki/Scraptiidae" title="Scraptiidae">Scraptiidae <small>(false flower beetles)</small></a></li> <li><a href="/wiki/Stenotrachelidae" title="Stenotrachelidae">Stenotrachelidae <small>(false longhorn beetles)</small></a></li> <li><a href="/wiki/Synchroidae" title="Synchroidae">Synchroidae <small>(synchroa bark beetles)</small></a></li> <li><a href="/wiki/Darkling_beetle" title="Darkling beetle">Tenebrionidae <small>(darkling beetles)</small></a></li> <li><a href="/wiki/Tetratomidae" title="Tetratomidae">Tetratomidae <small>(polypore fungus beetles)</small></a></li> <li><a href="/wiki/Trictenotomidae" title="Trictenotomidae">Trictenotomidae</a></li> <li><a href="/wiki/Ulodidae" title="Ulodidae">Ulodidae</a></li> <li><a href="/wiki/Zopheridae" title="Zopheridae">Zopheridae <small>(ironclad beetles, cylindrical bark beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Elateriformia" title="Elateriformia">Elateriformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Buprestoidea" title="Buprestoidea">Buprestoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Buprestidae" title="Buprestidae">Buprestidae <small>(jewel beetles, or metallic wood-boring beetles)</small></a></li> <li><a href="/wiki/Schizopodidae" title="Schizopodidae">Schizopodidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Byrrhoidea" title="Byrrhoidea">Byrrhoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Byrrhidae" title="Byrrhidae">Byrrhidae <small>(pill beetles)</small></a></li> <li><a href="/wiki/Callirhipidae" title="Callirhipidae">Callirhipidae <small>(cedar beetles)</small></a></li> <li><a href="/wiki/Chelonariidae" title="Chelonariidae">Chelonariidae <small>(turtle beetles)</small></a></li> <li><a href="/wiki/Cneoglossidae" title="Cneoglossidae">Cneoglossidae</a></li> <li><a href="/wiki/Dryopidae" title="Dryopidae">Dryopidae <small>(long-toed water beetles)</small></a></li> <li><a href="/wiki/Elmidae" title="Elmidae">Elmidae <small>(riffle beetles)</small></a></li> <li><a href="/wiki/Eulichadidae" title="Eulichadidae">Eulichadidae <small>(forest stream beetles)</small></a></li> <li><a class="mw-selflink selflink">Heteroceridae <small>(variegated mud-loving beetles)</small></a></li> <li><a href="/wiki/Limnichidae" title="Limnichidae">Limnichidae <small>(minute mud beetles)</small></a></li> <li><a href="/wiki/Lutrochidae" class="mw-redirect" title="Lutrochidae">Lutrochidae <small>(travertine beetles)</small></a></li> <li><a href="/wiki/Water-penny_beetles" class="mw-redirect" title="Water-penny beetles">Psephenidae <small>(water-penny beetles)</small></a></li> <li><a href="/wiki/Ptilodactylidae" title="Ptilodactylidae">Ptilodactylidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Dascilloidea" title="Dascilloidea">Dascilloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dascillidae" title="Dascillidae">Dascillidae <small>(soft bodied plant beetles)</small></a></li> <li><a href="/wiki/Rhipiceridae" title="Rhipiceridae">Rhipiceridae <small>(cicada beetle, cicada parasite beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Elateroidea" title="Elateroidea">Elateroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Artematopodidae" title="Artematopodidae">Artematopodidae <small>(soft-bodied plant beetles)</small></a></li> <li><a href="/wiki/Brachypsectridae" title="Brachypsectridae">Brachypsectridae <small>(Texas beetles)</small></a></li> <li><a href="/wiki/Soldier_beetle" title="Soldier beetle">Cantharidae <small>(soldier beetles)</small></a></li> <li><a href="/wiki/Cerophytidae" title="Cerophytidae">Cerophytidae <small>(rare click beetles)</small></a></li> <li><a href="/wiki/Click_beetle" title="Click beetle">Elateridae <small>(click beetles)</small></a></li> <li><a href="/wiki/Eucnemidae" title="Eucnemidae">Eucnemidae <small>(false click beetles)</small></a></li> <li><a href="/wiki/Jurasaidae" title="Jurasaidae">Jurasaidae</a></li> <li><a href="/wiki/Firefly" title="Firefly">Lampyridae <small>(fireflies)</small></a></li> <li><a href="/wiki/Lycidae" title="Lycidae">Lycidae <small>(net-winged beetles)</small></a></li> <li><a href="/wiki/Omethidae" title="Omethidae">Omethidae <small>(false fireflies, long-lipped beetles)</small></a></li> <li><a href="/wiki/Phengodidae" title="Phengodidae">Phengodidae <small>(glowworm beetles)</small></a></li> <li><a href="/wiki/Rhagophthalmidae" title="Rhagophthalmidae">Rhagophthalmidae</a></li> <li><a href="/wiki/Sinopyrophoridae" class="mw-redirect" title="Sinopyrophoridae">Sinopyrophoridae</a></li> <li><a href="/wiki/Throscidae" title="Throscidae">Throscidae <small>(false metallic wood-boring beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipidae <small>(<i>Rhinorhipus tamborinensis</i>)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scirtoidea" title="Scirtoidea">Scirtoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Clambidae" title="Clambidae">Clambidae</a></li> <li><a href="/wiki/Declinia" title="Declinia">Decliniidae <small>(<i>Declinia relicta</i>)</small></a></li> <li><a href="/wiki/Eucinetidae" title="Eucinetidae">Eucinetidae <small>(plate-thigh beetles)</small></a></li> <li><a href="/wiki/Scirtidae" title="Scirtidae">Scirtidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Belohina" title="Belohina">Belohinidae <small>(<i>Belohina inexpectata</i>)</small></a></li> <li><a href="/wiki/Bolboceratidae" title="Bolboceratidae">Bolboceratidae</a></li> <li><a href="/wiki/False_stag_beetle" title="False stag beetle">Diphyllostomatidae <small>(false stag beetles)</small></a></li> <li><a href="/wiki/Geotrupidae" title="Geotrupidae">Geotrupidae <small>(dor beetles)</small></a></li> <li><a href="/wiki/Glaphyridae" title="Glaphyridae">Glaphyridae <small>(bumble bee scarab beetles)</small></a></li> <li><a href="/wiki/Glaresis" title="Glaresis">Glaresidae <small>(enigmatic scarab beetles)</small></a></li> <li><a href="/wiki/Hybosoridae" title="Hybosoridae">Hybosoridae <small>(scavenger scarab beetles)</small></a></li> <li><a href="/wiki/Stag_beetle" title="Stag beetle">Lucanidae <small>(stag beetles)</small></a></li> <li><a href="/wiki/Ochodaeidae" title="Ochodaeidae">Ochodaeidae <small>(sand-loving scarab beetles)</small></a></li> <li><a href="/wiki/Passalidae" title="Passalidae">Passalidae <small>(betsy beetles)</small></a></li> <li><a href="/wiki/Rain_beetle" title="Rain beetle">Pleocomidae <small>(rain beetles)</small></a></li> <li><a href="/wiki/Scarabaeidae" title="Scarabaeidae">Scarabaeidae <small>(scarabs)</small></a></li> <li><a href="/wiki/Trogidae" title="Trogidae">Trogidae <small>(hide beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Staphyliniformia" title="Staphyliniformia">Staphyliniformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Histeroidea" title="Histeroidea">Histeroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Histeridae" title="Histeridae">Histeridae <small>(clown beetles)</small></a></li> <li><a href="/wiki/Sphaerites" title="Sphaerites">Sphaeritidae <small>(false clown beetles)</small></a></li> <li><a href="/wiki/Syntelia" title="Syntelia">Synteliidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Hydrophiloidea" title="Hydrophiloidea">Hydrophiloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Epimetopidae" title="Epimetopidae">Epimetopidae</a></li> <li><a href="/wiki/Georissus" title="Georissus">Georissidae <small>(minute mud-loving beetles)</small></a></li> <li><a href="/wiki/Helophorus" title="Helophorus">Helophoridae</a></li> <li><a href="/wiki/Hydrochidae" class="mw-redirect" title="Hydrochidae">Hydrochidae</a></li> <li><a href="/wiki/Hydrophilidae" title="Hydrophilidae">Hydrophilidae <small>(water scavenger beetles)</small></a></li> <li><a href="/wiki/Spercheidae" class="mw-redirect" title="Spercheidae">Spercheidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Staphylinoidea" title="Staphylinoidea">Staphylinoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agyrtidae" title="Agyrtidae">Agyrtidae <small>(primitive carrion beetles)</small></a></li> <li><a href="/wiki/Hydraenidae" title="Hydraenidae">Hydraenidae</a></li> <li><a href="/wiki/Leiodidae" title="Leiodidae">Leiodidae <small>(round fungus beetles)</small></a></li> <li><a href="/wiki/Ptiliidae" title="Ptiliidae">Ptiliidae <small>(feather-winged beetles)</small></a></li> <li><a href="/wiki/Silphidae" title="Silphidae">Silphidae <small>(carrion beetles)</small></a></li> <li><a href="/wiki/Rove_beetle" title="Rove beetle">Staphylinidae <small>(rove beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/List_of_subgroups_of_the_order_Coleoptera" title="List of subgroups of the order Coleoptera">List of subgroups of the order Coleoptera</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Heteroceridae</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q617854" class="extiw" title="wikidata:Q617854">Q617854</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Heteroceridae" class="extiw" title="wikispecies:Heteroceridae">Heteroceridae</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Faunal_Directory" title="Australian Faunal Directory">AFD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://biodiversity.org.au/afd/taxa/Heteroceridae">Heteroceridae</a></span></span></li> <li><span style="white-space:nowrap;">BioLib: <span class="uid"><a rel="nofollow" class="external text" href="https://www.biolib.cz/en/taxon/id8290">8290</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=1079">1079</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/BugGuide" title="BugGuide">BugGuide</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://bugguide.net/node/view/30091">30091</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/6255M">6255M</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/362">362</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/EPPO_Code" title="EPPO Code">EPPO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://gd.eppo.int/taxon/1HTRKF">1HTRKF</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:faunaeur.org:taxname:11154">11154</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea (new)</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://fauna-eu.org/cdm_dataportal/taxon/cc281af7-9464-4fd1-9818-1b1b9a06c866">cc281af7-9464-4fd1-9818-1b1b9a06c866</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/6986">6986</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/174627">174627</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=100983">100983</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=114262">114262</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Biodiversity_Network" title="National Biodiversity Network">NBN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://data.nbn.org.uk/Taxa/NHMSYS0001718442">NHMSYS0001718442</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=186979">186979</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/413befdf-542d-4986-820b-19a8e51c7ec6">413befdf-542d-4986-820b-19a8e51c7ec6</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=567884">567884</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=69343">69343</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Plazi" title="Plazi">Plazi</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://treatment.plazi.org/id/921787F1-F100-994A-B3F2-FD32FE19BACA">921787F1-F100-994A-B3F2-FD32FE19BACA</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=150720">150720</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" 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