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Available in 44 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-44" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">44 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AE%D9%86%D8%A7%D9%81%D8%B3_%D9%86%D9%87%D9%85%D8%A9" title="خنافس نهمة – Arabic" lang="ar" hreflang="ar" data-title="خنافس نهمة" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Asturian" lang="ast" hreflang="ast" data-title="Polyphaga" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A0%D0%B0%D0%B7%D0%BD%D0%B0%D0%B5%D0%B4%D0%BD%D1%8B%D1%8F_%D0%B6%D1%83%D0%BA%D1%96" title="Разнаедныя жукі – Belarusian" lang="be" hreflang="be" data-title="Разнаедныя жукі" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Pol%C3%ADfags" title="Polífags – Catalan" lang="ca" hreflang="ca" data-title="Polífags" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/V%C5%A1e%C5%BErav%C3%AD" title="Všežraví – Czech" lang="cs" hreflang="cs" data-title="Všežraví" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Danish" lang="da" hreflang="da" data-title="Polyphaga" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Polyphaga" title="Polyphaga – German" lang="de" hreflang="de" data-title="Polyphaga" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Segatoidumardikalised" title="Segatoidumardikalised – Estonian" lang="et" hreflang="et" data-title="Segatoidumardikalised" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Spanish" lang="es" hreflang="es" data-title="Polyphaga" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Basque" lang="eu" hreflang="eu" data-title="Polyphaga" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%D9%88%D8%B3%DA%A9%E2%80%8C%D9%87%D8%A7%DB%8C_%D9%87%D9%85%D9%87%E2%80%8C%DA%86%DB%8C%D8%B2%D8%AE%D9%88%D8%A7%D8%B1" title="سوسک‌های همه‌چیزخوار – Persian" lang="fa" hreflang="fa" data-title="سوسک‌های همه‌چیزخوار" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Polyphaga" title="Polyphaga – French" lang="fr" hreflang="fr" data-title="Polyphaga" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Galician" lang="gl" hreflang="gl" data-title="Polyphaga" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%92%8D%EB%8E%85%EC%9D%B4%EC%95%84%EB%AA%A9" title="풍뎅이아목 – Korean" lang="ko" hreflang="ko" data-title="풍뎅이아목" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Raznojedi" title="Raznojedi – Croatian" lang="hr" hreflang="hr" data-title="Raznojedi" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Indonesian" lang="id" hreflang="id" data-title="Polyphaga" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Italian" lang="it" hreflang="it" data-title="Polyphaga" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%9C%E1%83%90%E1%83%98%E1%83%A0%E1%83%9B%E1%83%AD%E1%83%90%E1%83%9B%E1%83%94%E1%83%9A%E1%83%98_%E1%83%AE%E1%83%9D%E1%83%AD%E1%83%9D%E1%83%94%E1%83%91%E1%83%98" title="ნაირმჭამელი ხოჭოები – Georgian" lang="ka" hreflang="ka" data-title="ნაირმჭამელი ხოჭოები" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Bungo_(mdudu)" title="Bungo (mdudu) – Swahili" lang="sw" hreflang="sw" data-title="Bungo (mdudu)" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%A2%D1%83%D1%88_%D0%BA%D0%B5%D0%BB%D0%B4%D0%B8_%D0%B6%D0%B5%D1%87%D2%AF_%D0%BA%D0%BE%D2%A3%D1%83%D0%B7%D0%B4%D0%B0%D1%80" title="Туш келди жечү коңуздар – Kyrgyz" lang="ky" hreflang="ky" data-title="Туш келди жечү коңуздар" data-language-autonym="Кыргызча" data-language-local-name="Kyrgyz" class="interlanguage-link-target"><span>Кыргызча</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Latin" lang="la" hreflang="la" data-title="Polyphaga" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Vis%C4%93d%C4%81jas_vaboles" title="Visēdājas vaboles – Latvian" lang="lv" hreflang="lv" data-title="Visēdājas vaboles" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/%C4%AEvairia%C4%97d%C5%BEiai_vabalai" title="Įvairiaėdžiai vabalai – Lithuanian" lang="lt" hreflang="lt" data-title="Įvairiaėdžiai vabalai" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Mindenev%C5%91_bogarak" title="Mindenevő bogarak – Hungarian" lang="hu" hreflang="hu" data-title="Mindenevő bogarak" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AE%D9%86%D8%A7%D9%81%D8%B3_%D9%86%D9%87%D9%85%D9%87" title="خنافس نهمه – Egyptian Arabic" lang="arz" hreflang="arz" data-title="خنافس نهمه" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Malay" lang="ms" hreflang="ms" data-title="Polyphaga" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Polyphaga_(kevers)" title="Polyphaga (kevers) – Dutch" lang="nl" hreflang="nl" data-title="Polyphaga (kevers)" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%A4%9A%E9%A3%9F%E4%BA%9C%E7%9B%AE" title="多食亜目 – Japanese" lang="ja" hreflang="ja" data-title="多食亜目" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Polyphaga" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Norwegian Nynorsk" lang="nn" hreflang="nn" data-title="Polyphaga" data-language-autonym="Norsk nynorsk" data-language-local-name="Norwegian Nynorsk" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Low German" lang="nds" hreflang="nds" data-title="Polyphaga" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Chrz%C4%85szcze_wielo%C5%BCerne" title="Chrząszcze wielożerne – Polish" lang="pl" hreflang="pl" data-title="Chrząszcze wielożerne" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Portuguese" lang="pt" hreflang="pt" data-title="Polyphaga" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Romanian" lang="ro" hreflang="ro" data-title="Polyphaga" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A0%D0%B0%D0%B7%D0%BD%D0%BE%D1%8F%D0%B4%D0%BD%D1%8B%D0%B5_%D0%B6%D1%83%D0%BA%D0%B8" title="Разноядные жуки – Russian" lang="ru" hreflang="ru" data-title="Разноядные жуки" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Simple English" lang="en-simple" hreflang="en-simple" data-title="Polyphaga" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Serbian" lang="sr" hreflang="sr" data-title="Polyphaga" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Erilaisruokaiset" title="Erilaisruokaiset – Finnish" lang="fi" hreflang="fi" data-title="Erilaisruokaiset" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/All%C3%A4tarbaggar" title="Allätarbaggar – Swedish" lang="sv" hreflang="sv" data-title="Allätarbaggar" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Turkish" lang="tr" hreflang="tr" data-title="Polyphaga" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%92%D1%81%D0%B5%D1%97%D0%B4%D0%BD%D1%96_%D0%B6%D1%83%D0%BA%D0%B8" title="Всеїдні жуки – Ukrainian" lang="uk" hreflang="uk" data-title="Всеїдні жуки" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Polyphaga" title="Polyphaga – Vietnamese" lang="vi" hreflang="vi" data-title="Polyphaga" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E5%A4%9A%E9%A3%9F%E4%BA%9E%E7%9B%AE" title="多食亞目 – Cantonese" lang="yue" hreflang="yue" data-title="多食亞目" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%A4%9A%E9%A3%9F%E4%BA%9E%E7%9B%AE" title="多食亞目 – Chinese" lang="zh" hreflang="zh" data-title="多食亞目" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit 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<div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Suborder of beetles</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Not to be confused with <a href="/wiki/Polyphagia" title="Polyphagia">Polyphagia</a>.</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">For the genus of Blattodea (cockroaches), see <a href="/wiki/Polyphaga_(cockroach)" title="Polyphaga (cockroach)">Polyphaga (cockroach)</a>.</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Polyphaga<br /><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">Triassic–Present</span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:132.16584615385px; width:87.834153846154px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:134.74086153846px; width:85.259138461538px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:135.74086153846px; width:83.259138461538px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Cetonia-aurata.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Cetonia-aurata.jpg/220px-Cetonia-aurata.jpg" decoding="async" width="220" height="183" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Cetonia-aurata.jpg/330px-Cetonia-aurata.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Cetonia-aurata.jpg/440px-Cetonia-aurata.jpg 2x" data-file-width="1800" data-file-height="1500" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i><a href="/wiki/Cetonia_aurata" title="Cetonia aurata">Cetonia aurata</a></i> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Polyphaga" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Insect" title="Insect">Insecta</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Beetle" title="Beetle">Coleoptera</a> </td></tr> <tr> <td>Suborder: </td> <td><a class="mw-selflink selflink">Polyphaga</a><br /><small><a href="/wiki/Carlo_Emery" title="Carlo Emery">Emery</a>, 1886</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Infraorders </th></tr> <tr> <td colspan="2" style="text-align: left"> <p><a href="/wiki/Bostrichiformia" title="Bostrichiformia">Bostrichiformia</a><br /> <a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a><br /> <a href="/wiki/Elateriformia" title="Elateriformia">Elateriformia</a><br /> <a href="/wiki/Scarabaeiformia" class="mw-redirect" title="Scarabaeiformia">Scarabaeiformia</a><br /> <a href="/wiki/Staphyliniformia" title="Staphyliniformia">Staphyliniformia</a> </p> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Polyphaga</b> is the largest and most diverse <a href="/wiki/Suborder" class="mw-redirect" title="Suborder">suborder</a> of <a href="/wiki/Beetle" title="Beetle">beetles</a>. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the beetle species discovered thus far. </p><p>Key characteristics of Polyphaga are that the hind <a href="/wiki/Arthropod_coxa" class="mw-redirect" title="Arthropod coxa">coxa</a> (base of the leg) does not divide the first and second abdominal/ventral plates which are known as <a href="/wiki/Sternites" class="mw-redirect" title="Sternites">sternites</a>. Also, the <a href="/wiki/Notopleuron" title="Notopleuron">notopleural</a> suture (found under the pronotal shield) is not present.<sup id="cite_ref-Johnson_1-0" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Etymology">Etymology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Polyphaga&action=edit&section=1" title="Edit section: Etymology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The name of <i>polyphaga</i> is derived from two Greek words: <span title="Ancient Greek (to 1453)-language romanization"><i lang="grc-Latn">poly-</i></span>, meaning 'many', and <span title="Ancient Greek (to 1453)-language romanization"><i lang="grc-Latn">phagein</i></span>, meaning 'to eat', so the suborder is called the “eaters of many things”. </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Polyphaga&action=edit&section=2" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The five main infraorders are: </p> <ul><li><a href="/wiki/Bostrichiformia" title="Bostrichiformia">Bostrichiformia</a> — including <a href="/wiki/Furniture_beetle" class="mw-redirect" title="Furniture beetle">furniture beetles</a> and <a href="/wiki/Skin_beetle" class="mw-redirect" title="Skin beetle">skin beetles</a></li> <li><a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a> — includes <a href="/wiki/Lady_beetle" class="mw-redirect" title="Lady beetle">lady beetles</a>, <a href="/wiki/Longhorn_beetle" title="Longhorn beetle">longhorn beetles</a>, <a href="/wiki/Weevil" title="Weevil">weevils</a>, <a href="/wiki/Checkered_beetle" class="mw-redirect" title="Checkered beetle">checkered beetles</a> and <a href="/wiki/Leaf_beetle" title="Leaf beetle">leaf beetles</a></li> <li><a href="/wiki/Elateriformia" title="Elateriformia">Elateriformia</a> — includes <a href="/wiki/Click_beetle" title="Click beetle">click beetles</a> and <a href="/wiki/Firefly" title="Firefly">fireflies</a></li> <li><a href="/wiki/Scarabaeiformia" class="mw-redirect" title="Scarabaeiformia">Scarabaeiformia</a> — includes <a href="/wiki/Scarabaeidae" title="Scarabaeidae">scarab beetles</a>, <a href="/wiki/Stag_beetle" title="Stag beetle">stag beetles</a>, and <a href="/wiki/Dung_beetles" class="mw-redirect" title="Dung beetles">dung beetles</a></li> <li><a href="/wiki/Staphyliniformia" title="Staphyliniformia">Staphyliniformia</a> — includes <a href="/wiki/Rove_beetle" title="Rove beetle">rove beetles</a> and <a href="/wiki/Hydrophilidae" title="Hydrophilidae">water scavenger beetles</a></li></ul> <p>Phylogenetic studies have also suggested that Scirtoidea (<a href="/wiki/Scirtidae" title="Scirtidae">Scirtidae</a>, <a href="/wiki/Decliniidae" class="mw-redirect" title="Decliniidae">Decliniidae</a>), Clamboidea (<a href="/wiki/Clambidae" title="Clambidae">Clambidae</a>, <a href="/wiki/Derodontidae" title="Derodontidae">Derodontidae</a>, <a href="/wiki/Eucinetidae" title="Eucinetidae">Eucinetidae</a>), <i><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipus</a></i> and <a href="/wiki/Nosodendridae" title="Nosodendridae">Nosodendridae</a> are independent lineages of Polyphaga that lie outside these groups.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>The internal classification of Polyphaga involves several superfamilies or series, whose constituents are relatively stable, although some smaller families (whose rank even is disputed) are allocated to different clades by different authors. Large superfamilies include <a href="/wiki/Hydrophiloidea" title="Hydrophiloidea">Hydrophiloidea</a>, <a href="/wiki/Staphylinoidea" title="Staphylinoidea">Staphylinoidea</a>, <a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeoidea</a>, <a href="/wiki/Buprestoidea" title="Buprestoidea">Buprestoidea</a>, <a href="/wiki/Byrrhoidea" title="Byrrhoidea">Byrrhoidea</a>, <a href="/wiki/Elateroidea" title="Elateroidea">Elateroidea</a>, and <a href="/wiki/Bostrichoidea" title="Bostrichoidea">Bostrichoidea</a>. </p><p>The infraorder <a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a> includes the vast majority of <a href="/wiki/Phytophagous" class="mw-redirect" title="Phytophagous">phytophagous</a> (plant-eating) beetles, united by cryptonephric <a href="/wiki/Malpighian_tubule" class="mw-redirect" title="Malpighian tubule">Malpighian tubules</a> of the normal type, a cone <a href="/wiki/Ommatidium" title="Ommatidium">ommatidium</a> with open <a href="/wiki/Rhabdom" class="mw-redirect" title="Rhabdom">rhabdom</a>, and lack of functional <a href="/wiki/Spiracle_(arthropods)" title="Spiracle (arthropods)">spiracles</a> on the eighth abdominal segment. Constituent superfamilies of Cucujiformia are <a href="/wiki/Cleroidea" title="Cleroidea">Cleroidea</a>, <a href="/wiki/Cucujoidea" title="Cucujoidea">Cucujoidea</a>, <a href="/wiki/Tenebrionoidea" title="Tenebrionoidea">Tenebrionoidea</a>, <a href="/wiki/Chrysomeloidea" title="Chrysomeloidea">Chrysomeloidea</a>, and <a href="/wiki/Curculionoidea" class="mw-redirect" title="Curculionoidea">Curculionoidea</a>. Evidently adoption of a phytophagous lifestyle correlates with taxon diversity in beetles, with Cucujiformia, especially <a href="/wiki/Weevil" title="Weevil">weevils</a> (<a href="/wiki/Curculionoidea" class="mw-redirect" title="Curculionoidea">Curculionoidea</a>), forming a major radiation. </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Polyphaga&action=edit&section=3" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/List_of_subgroups_of_the_order_Coleoptera" title="List of subgroups of the order Coleoptera">List of subgroups of the order Coleoptera</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Polyphaga&action=edit&section=4" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-Johnson-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-Johnson_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFJohnsonTriplehorn2004" class="citation book cs1">Johnson, Norman F.; Triplehorn, Charles A. (2004). <i>Borror and DeLong's Introduction to the Study of Insects</i> (7th ed.). Belmont: Brooks/Cole. pp. 365–400, 428–429. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-03-096835-6" title="Special:BookSources/0-03-096835-6"><bdi>0-03-096835-6</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Borror+and+DeLong%27s+Introduction+to+the+Study+of+Insects&rft.place=Belmont&rft.pages=365-400%2C+428-429&rft.edition=7th&rft.pub=Brooks%2FCole&rft.date=2004&rft.isbn=0-03-096835-6&rft.aulast=Johnson&rft.aufirst=Norman+F.&rft.au=Triplehorn%2C+Charles+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3APolyphaga" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCaiTihelkaGiacomelliLawrence2022" class="citation journal cs1">Cai, Chenyang; Tihelka, Erik; Giacomelli, Mattia; Lawrence, John F.; Ślipiński, Adam; Kundrata, Robin; Yamamoto, Shûhei; Thayer, Margaret K.; Newton, Alfred F.; Leschen, Richard A. B.; Gimmel, Matthew L.; Lü, Liang; Engel, Michael S.; Bouchard, Patrice; Huang, Diying (March 2022). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8941382">"Integrated phylogenomics and fossil data illuminate the evolution of beetles"</a>. <i>Royal Society Open Science</i>. <b>9</b> (3): 211771. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2022RSOS....911771C">2022RSOS....911771C</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frsos.211771">10.1098/rsos.211771</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/2054-5703">2054-5703</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8941382">8941382</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/35345430">35345430</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Royal+Society+Open+Science&rft.atitle=Integrated+phylogenomics+and+fossil+data+illuminate+the+evolution+of+beetles&rft.volume=9&rft.issue=3&rft.pages=211771&rft.date=2022-03&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8941382%23id-name%3DPMC&rft_id=info%3Abibcode%2F2022RSOS....911771C&rft_id=info%3Apmid%2F35345430&rft_id=info%3Adoi%2F10.1098%2Frsos.211771&rft.issn=2054-5703&rft.aulast=Cai&rft.aufirst=Chenyang&rft.au=Tihelka%2C+Erik&rft.au=Giacomelli%2C+Mattia&rft.au=Lawrence%2C+John+F.&rft.au=%C5%9Alipi%C5%84ski%2C+Adam&rft.au=Kundrata%2C+Robin&rft.au=Yamamoto%2C+Sh%C3%BBhei&rft.au=Thayer%2C+Margaret+K.&rft.au=Newton%2C+Alfred+F.&rft.au=Leschen%2C+Richard+A.+B.&rft.au=Gimmel%2C+Matthew+L.&rft.au=L%C3%BC%2C+Liang&rft.au=Engel%2C+Michael+S.&rft.au=Bouchard%2C+Patrice&rft.au=Huang%2C+Diying&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8941382&rfr_id=info%3Asid%2Fen.wikipedia.org%3APolyphaga" class="Z3988"></span></span> </li> </ol></div></div> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCranstonGullan2009" class="citation book cs1">Cranston, Peter S.; Gullan, Penny J. (2009). "Phylogeny of Insects". In Resh, Vincent H.; Cardé, Ring T. (eds.). <i>Encyclopedia of Insects</i> (Second ed.). Academic Press. pp. 780–793. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-12-374144-8" title="Special:BookSources/978-0-12-374144-8"><bdi>978-0-12-374144-8</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Phylogeny+of+Insects&rft.btitle=Encyclopedia+of+Insects&rft.pages=780-793&rft.edition=Second&rft.pub=Academic+Press&rft.date=2009&rft.isbn=978-0-12-374144-8&rft.aulast=Cranston&rft.aufirst=Peter+S.&rft.au=Gullan%2C+Penny+J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3APolyphaga" class="Z3988"></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Polyphaga&action=edit&section=5" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output 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srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/51px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/68px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></span></span></div> <div class="side-box-text plainlist"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a> has information related to <i><b><a href="https://species.wikimedia.org/wiki/Polyphaga_(Coleoptera)" class="extiw" title="wikispecies:Polyphaga (Coleoptera)">Polyphaga (Coleoptera)</a></b></i>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20160819092349/https://www.zin.ru/animalia/coleoptera/pdf/Gullan_Insects_2014.pdf">Phylogeny of Insects</a></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20051018185209/http://www.kbinirsnb.be/collections/entomo/staphy/">Site devoted to Staphyliniformia</a></li> <li><a rel="nofollow" class="external text" href="http://www.fieldmuseum.org/peet_staph/project_sum.html">Project devoted to studying rove beetles</a></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20050307184527/http://www.insects.org/entophiles/coleoptera/cole_en005.html">Image of a click beetle</a></li> <li><a rel="nofollow" class="external text" href="http://maria.fremlin.de/stagbeetles/">site devoted to stag beetles</a></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20060610061035/http://www.aces.uiuc.edu/longhorned_beetle/">news article highlighting the damage caused by Longhorn beetles</a></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://bie.ala.org.au/species/urn:lsid:biodiversity.org.au:afd.taxon:17babfba-1818-4371-bb03-f09595452fa6">"<i>Polyphaga</i>"</a>. <i>Atlas of Living Australia</i>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Atlas+of+Living+Australia&rft.atitle=Polyphaga&rft_id=https%3A%2F%2Fbie.ala.org.au%2Fspecies%2Furn%3Alsid%3Abiodiversity.org.au%3Aafd.taxon%3A17babfba-1818-4371-bb03-f09595452fa6&rfr_id=info%3Asid%2Fen.wikipedia.org%3APolyphaga" class="Z3988"></span></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output 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href="/wiki/Special:EditPage/Template:Coleoptera" title="Special:EditPage/Template:Coleoptera"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Extant_Coleoptera_families" style="font-size:114%;margin:0 4em">Extant <a href="/wiki/Beetle" title="Beetle">Coleoptera</a> families</div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a></li> <li>Class: <a href="/wiki/Insect" title="Insect">Insecta</a></li> <li>Subclass: <a href="/wiki/Pterygota" title="Pterygota">Pterygota</a></li> <li>Infraclass: <a href="/wiki/Neoptera" title="Neoptera">Neoptera</a></li> <li>Superorder: <a href="/wiki/Holometabola" title="Holometabola">Holometabola</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Archostemata" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Archostemata" title="Archostemata">Archostemata</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Crowsoniella" title="Crowsoniella">Crowsoniellidae <small>(<i>Crowsoniella relicta</i>)</small></a></li> <li><a href="/wiki/Cupedidae" title="Cupedidae">Cupedidae <small>(reticulated beetles)</small></a></li> <li><a href="/wiki/Jurodidae" title="Jurodidae">Jurodidae <small>(<i>Sikhotealinia zhiltzovae</i>)</small></a></li> <li><a href="/wiki/Micromalthidae" title="Micromalthidae">Micromalthidae</a></li> <li><a href="/wiki/Ommatidae" title="Ommatidae">Ommatidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Adephaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Adephaga" title="Adephaga">Adephaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Extant_families" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Extant_taxon" class="mw-redirect" title="Extant taxon">Extant</a> families</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amphizoa" title="Amphizoa">Amphizoidae <small>(trout-stream beetles)</small></a></li> <li><a href="/wiki/Aspidytidae" title="Aspidytidae">Aspidytidae</a></li> <li><a href="/wiki/Ground_beetle" title="Ground beetle">Carabidae <small>(ground beetles)</small></a></li> <li><a href="/wiki/Tiger_beetle" title="Tiger beetle">Cicindelidae <small>(tiger beetles)</small></a></li> <li><a href="/wiki/Dytiscidae" title="Dytiscidae">Dytiscidae <small>(predaceous diving beetles)</small></a></li> <li><a href="/wiki/Whirligig_beetle" title="Whirligig beetle">Gyrinidae <small>(whirligig beetles)</small></a></li> <li><a href="/wiki/Haliplidae" title="Haliplidae">Haliplidae <small>(crawling water beetles)</small></a></li> <li><a href="/wiki/Hygrobia" title="Hygrobia">Hygrobiidae</a></li> <li><a href="/wiki/Meru_phyllisae" title="Meru phyllisae">Meruidae <small>(<i>Meru phyllisae</i>)</small></a></li> <li><a href="/wiki/Noteridae" title="Noteridae">Noteridae <small>(burrowing water beetles)</small></a></li> <li><a href="/wiki/Trachypachidae" title="Trachypachidae">Trachypachidae <small>(false ground beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Myxophaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Myxophaga" title="Myxophaga">Myxophaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hydroscaphidae" title="Hydroscaphidae">Hydroscaphidae <small>(skiff beetles)</small></a></li> <li><a href="/wiki/Lepicerus" title="Lepicerus">Lepiceridae</a></li> <li><a href="/wiki/Sphaerius" title="Sphaerius">Sphaeriusidae</a></li> <li><a href="/wiki/Torridincolidae" title="Torridincolidae">Torridincolidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Polyphaga" style="font-size:114%;margin:0 4em">Suborder <a class="mw-selflink selflink">Polyphaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bostrichiformia" title="Bostrichiformia">Bostrichiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Bostrichoidea" title="Bostrichoidea">Bostrichoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bostrichidae" title="Bostrichidae">Bostrichidae <small>(auger beetles)</small></a></li> <li><a href="/wiki/Dermestidae" title="Dermestidae">Dermestidae <small>(skin beetles)</small></a></li> <li><a href="/wiki/Endecatomus" title="Endecatomus">Endecatomidae</a></li> <li><a href="/wiki/Jacobsoniidae" title="Jacobsoniidae">Jacobsoniidae <small>(Jacobson's beetles)</small></a></li> <li><a href="/wiki/Nosodendridae" title="Nosodendridae">Nosodendridae <small>(wounded-tree beetles)</small></a></li> <li><a href="/wiki/Ptinidae" title="Ptinidae">Ptiniidae <small>(furniture beetles, death watch beetles, spider beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Derodontidae" title="Derodontidae">Derodontoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Derodontidae" title="Derodontidae">Derodontidae <small>(tooth-necked fungus beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Chrysomeloidea" title="Chrysomeloidea">Chrysomeloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Longhorn_beetle" title="Longhorn beetle">Cerambycidae <small>(longhorn beetles)</small></a></li> <li><a href="/wiki/Leaf_beetle" title="Leaf beetle">Chrysomelidae <small>(leaf beetles)</small></a></li> <li><a href="/wiki/Disteniidae" title="Disteniidae">Disteniidae</a></li> <li><a href="/wiki/Megalopodidae" title="Megalopodidae">Megalopodidae</a></li> <li><a href="/wiki/Orsodacnidae" title="Orsodacnidae">Orsodacnidae</a></li> <li><a href="/wiki/Oxypeltidae" title="Oxypeltidae">Oxypeltidae</a></li> <li><a href="/wiki/Vesperidae" title="Vesperidae">Vesperidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cleroidea" title="Cleroidea">Cleroidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acanthocnemus" title="Acanthocnemus">Acanthocnemidae <small>(<i>Acanthocnemus nigricans</i>)</small></a></li> <li><a href="/wiki/Biphyllidae" title="Biphyllidae">Biphyllidae <small>(false skin beetles)</small></a></li> <li><a href="/wiki/Byturidae" title="Byturidae">Byturidae <small>(fruitworm beetles)</small></a></li> <li><a href="/wiki/Chaetosomatidae" title="Chaetosomatidae">Chaetosomatidae</a></li> <li><a href="/wiki/Cleridae" title="Cleridae">Cleridae <small>(checkered beetles)</small></a></li> <li><a href="/wiki/Lophocateridae" title="Lophocateridae">Lophocateridae</a></li> <li><a href="/wiki/Mauroniscidae" title="Mauroniscidae">Mauroniscidae</a></li> <li><a href="/wiki/Melyridae" title="Melyridae">Melyridae <small>(soft-wing flower beetles)</small></a></li> <li><a href="/wiki/Metaxina" title="Metaxina">Metaxinidae <small>(<i>Metaxina ornata</i>)</small></a></li> <li><a href="/wiki/Peltis" title="Peltis">Peltidae</a></li> <li><a href="/wiki/Phloiophilus" title="Phloiophilus">Phloiophilidae <small>(<i>Phloiophilus edwardsi</i>)</small></a></li> <li><a href="/wiki/Phycosecidae" title="Phycosecidae">Phycosecidae</a></li> <li><a href="/wiki/Prionoceridae" title="Prionoceridae">Prionoceridae</a></li> <li><a href="/wiki/Protopeltis" title="Protopeltis">Protopeltidae</a></li> <li><a href="/wiki/Rentoniidae" title="Rentoniidae">Rentoniidae</a></li> <li><a href="/wiki/Rhadalidae" title="Rhadalidae">Rhadalidae</a></li> <li><a href="/wiki/Thanerocleridae" title="Thanerocleridae">Thanerocleridae</a></li> <li><a href="/wiki/Thymalidae" title="Thymalidae">Thymalidae</a></li> <li><a href="/wiki/Trogossitidae" title="Trogossitidae">Trogossitidae <small>(bark-gnawing beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Coccinelloidea" title="Coccinelloidea">Coccinelloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Akalyptoischion" title="Akalyptoischion">Akalyptoischiidae</a></li> <li><a href="/wiki/Alexiidae" title="Alexiidae">Alexiidae</a></li> <li><a href="/wiki/Anamorphidae" title="Anamorphidae">Anamorphidae</a></li> <li><a href="/wiki/Bothrideridae" title="Bothrideridae">Bothrideridae <small>(dry bark beetles)</small></a></li> <li><a href="/wiki/Cerylonidae" title="Cerylonidae">Cerylonidae <small>(minute bark beetles)</small></a></li> <li><a href="/wiki/Coccinellidae" title="Coccinellidae">Coccinellidae <small>(lady beetles, or God's cows)</small></a></li> <li><a href="/wiki/Corylophidae" title="Corylophidae">Corylophidae <small>(minute fungus beetles)</small></a></li> <li><a href="/wiki/Discolomatidae" title="Discolomatidae">Discolomatidae</a></li> <li><a href="/wiki/Endomychidae" title="Endomychidae">Endomychidae <small>(handsome fungus beetles)</small></a></li> <li><a href="/wiki/Eupsilobiidae" title="Eupsilobiidae">Eupsilobiidae</a></li> <li><a href="/wiki/Euxestidae" title="Euxestidae">Euxestidae <small>(well polished beetles)</small></a></li> <li><a href="/wiki/Latridiidae" title="Latridiidae">Latridiidae <small>(minute brown scavenger beetles)</small></a></li> <li><a href="/wiki/Murmidiidae" title="Murmidiidae">Murmidiidae</a></li> <li><a href="/wiki/Mycetaeidae" title="Mycetaeidae">Mycetaeidae</a></li> <li><a href="/wiki/Teredidae" title="Teredidae">Teredidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cucujoidea" title="Cucujoidea">Cucujoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agapytho" title="Agapytho">Agapythidae <small>(<i>Agapytho foveicollis</i>)</small></a></li> <li><a href="/wiki/Boganiidae" title="Boganiidae">Boganiidae</a></li> <li><a href="/wiki/Cavognathidae" title="Cavognathidae">Cavognathidae</a></li> <li><a href="/wiki/Cryptophagidae" title="Cryptophagidae">Cryptophagidae <small>(silken fungus beetles)</small></a></li> <li><a href="/wiki/Cucujidae" title="Cucujidae">Cucujidae <small>(flat bark beetles)</small></a></li> <li><a href="/wiki/Cybocephalidae" title="Cybocephalidae">Cybocephalidae</a></li> <li><a href="/wiki/Cyclaxyridae" title="Cyclaxyridae">Cyclaxyridae</a></li> <li><a href="/wiki/Erotylidae" title="Erotylidae">Erotylidae <small>(pleasing fungus beetles)</small></a></li> <li><a href="/wiki/Helotidae" title="Helotidae">Helotidae</a></li> <li><a href="/wiki/Hobartiidae" title="Hobartiidae">Hobartiidae</a></li> <li><a href="/wiki/Kateretidae" title="Kateretidae">Kateretidae <small>(short-winged flower beetles)</small></a></li> <li><a href="/wiki/Laemophloeidae" title="Laemophloeidae">Laemophloeidae <small>(lined flat bark beetles)</small></a></li> <li><a href="/wiki/Lamingtonium" title="Lamingtonium">Lamingtoniidae <small>(<i>Lamingtonium binnaberrense</i>)</small></a></li> <li><a href="/wiki/Monotomidae" title="Monotomidae">Monotomidae <small>(root-eating beetles)</small></a></li> <li><a href="/wiki/Myraboliidae" class="mw-redirect" title="Myraboliidae">Myraboliidae</a></li> <li><a href="/wiki/Sap_beetle" title="Sap beetle">Nitidulidae <small>(sap beetles)</small></a></li> <li><a href="/wiki/Passandridae" title="Passandridae">Passandridae <small>(parasitic flat bark beetles)</small></a></li> <li><a href="/wiki/Phalacridae" title="Phalacridae">Phalacridae <small>(shining flower beetles)</small></a></li> <li><a href="/wiki/Phloeostichidae" title="Phloeostichidae">Phloeostichidae</a></li> <li><a href="/wiki/Priasilphidae" title="Priasilphidae">Priasilphidae</a></li> <li><a href="/wiki/Propalticidae" title="Propalticidae">Propalticidae</a></li> <li><a href="/wiki/Protocucujidae" title="Protocucujidae">Protocucujidae</a></li> <li><a href="/wiki/Silvanidae" title="Silvanidae">Silvanidae <small>(silvanid flat bark beetles)</small></a></li> <li><a href="/wiki/Smicripidae" title="Smicripidae">Smicripidae <small>(palmetto beetles)</small></a></li> <li><a href="/wiki/Sphindidae" title="Sphindidae">Sphindidae <small>(dry-fungus beetles)</small></a></li> <li><a href="/wiki/Tasmosalpingus" title="Tasmosalpingus">Tasmosalpingidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Weevil" title="Weevil">Curculionoidea</a><br /><small><span style="color:#696969"><span class="nobold">(weevils)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anthribidae" title="Anthribidae">Anthribidae <small>(fungus weevils)</small></a></li> <li><a href="/wiki/Attelabidae" title="Attelabidae">Attelabidae <small>(leaf-rolling weevils)</small></a></li> <li><a href="/wiki/Belidae" title="Belidae">Belidae <small>(primitive weevils)</small></a></li> <li><a href="/wiki/Brentidae" title="Brentidae">Brentidae <small>(straight snout weevils, New York weevil)</small></a></li> <li><a href="/wiki/Caridae" title="Caridae">Caridae</a></li> <li><a href="/wiki/Curculionidae" title="Curculionidae">Curculionidae <small>(true weevils, bark beetles, ambrosia beetles)</small></a></li> <li><a href="/wiki/Nemonychidae" title="Nemonychidae">Nemonychidae <small>(pine flower weevils)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexyloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexylidae <small>(ship-timber beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Tenebrionoidea" title="Tenebrionoidea">Tenebrionoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aderidae" title="Aderidae">Aderidae <small>(ant-like leaf beetles)</small></a></li> <li><a href="/wiki/Anthicidae" title="Anthicidae">Anthicidae <small>(ant-like flower beetles)</small></a></li> <li><a href="/wiki/Archeocrypticidae" title="Archeocrypticidae">Archeocrypticidae <small>(cryptic fungus beetles)</small></a></li> <li><a href="/wiki/Boridae" title="Boridae">Boridae <small>(conifer bark beetles)</small></a></li> <li><a href="/wiki/Chalcodryidae" title="Chalcodryidae">Chalcodryidae</a></li> <li><a href="/wiki/Ciidae" title="Ciidae">Ciidae <small>(minute tree-fungus beetles)</small></a></li> <li><a href="/wiki/Melandryidae" title="Melandryidae">Melandryidae <small>(false darkling beetles)</small></a></li> <li><a href="/wiki/Blister_beetle" title="Blister beetle">Meloidae <small>(blister beetles)</small></a></li> <li><a href="/wiki/Mordellidae" title="Mordellidae">Mordellidae <small>(tumbling flower beetles)</small></a></li> <li><a href="/wiki/Mycetophagidae" title="Mycetophagidae">Mycetophagidae <small>(hairy fungus beetles)</small></a></li> <li><a href="/wiki/Mycteridae" title="Mycteridae">Mycteridae <small>(palm and flower beetles)</small></a></li> <li><a href="/wiki/Oedemeridae" title="Oedemeridae">Oedemeridae <small>(false blister beetle)</small></a></li> <li><a href="/wiki/Perimylopidae" class="mw-redirect" title="Perimylopidae">Perimylopidae, or Promecheilidae</a></li> <li><a href="/wiki/Prostomidae" title="Prostomidae">Prostomidae <small>(jugular-horned beetles)</small></a></li> <li><a href="/wiki/Pterogeniidae" title="Pterogeniidae">Pterogeniidae</a></li> <li><a href="/wiki/Fire-coloured_beetle" title="Fire-coloured beetle">Pyrochroidae <small>(fire-coloured beetles)</small></a></li> <li><a href="/wiki/Pythidae" title="Pythidae">Pythidae <small>(dead log bark beetles)</small></a></li> <li><a href="/wiki/Ripiphoridae" title="Ripiphoridae">Ripiphoridae <small>(wedge-shaped beetles)</small></a></li> <li><a href="/wiki/Salpingidae" title="Salpingidae">Salpingidae <small>(narrow-waisted bark beetles)</small></a></li> <li><a href="/wiki/Scraptiidae" title="Scraptiidae">Scraptiidae <small>(false flower beetles)</small></a></li> <li><a href="/wiki/Stenotrachelidae" title="Stenotrachelidae">Stenotrachelidae <small>(false longhorn beetles)</small></a></li> <li><a href="/wiki/Synchroidae" title="Synchroidae">Synchroidae <small>(synchroa bark beetles)</small></a></li> <li><a href="/wiki/Darkling_beetle" title="Darkling beetle">Tenebrionidae <small>(darkling beetles)</small></a></li> <li><a href="/wiki/Tetratomidae" title="Tetratomidae">Tetratomidae <small>(polypore fungus beetles)</small></a></li> <li><a href="/wiki/Trictenotomidae" title="Trictenotomidae">Trictenotomidae</a></li> <li><a href="/wiki/Ulodidae" title="Ulodidae">Ulodidae</a></li> <li><a href="/wiki/Zopheridae" title="Zopheridae">Zopheridae <small>(ironclad beetles, cylindrical bark beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Elateriformia" title="Elateriformia">Elateriformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Buprestoidea" title="Buprestoidea">Buprestoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Buprestidae" title="Buprestidae">Buprestidae <small>(jewel beetles, or metallic wood-boring beetles)</small></a></li> <li><a href="/wiki/Schizopodidae" title="Schizopodidae">Schizopodidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Byrrhoidea" title="Byrrhoidea">Byrrhoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Byrrhidae" title="Byrrhidae">Byrrhidae <small>(pill beetles)</small></a></li> <li><a href="/wiki/Callirhipidae" title="Callirhipidae">Callirhipidae <small>(cedar beetles)</small></a></li> <li><a href="/wiki/Chelonariidae" title="Chelonariidae">Chelonariidae <small>(turtle beetles)</small></a></li> <li><a href="/wiki/Cneoglossidae" title="Cneoglossidae">Cneoglossidae</a></li> <li><a href="/wiki/Dryopidae" title="Dryopidae">Dryopidae <small>(long-toed water beetles)</small></a></li> <li><a href="/wiki/Elmidae" title="Elmidae">Elmidae <small>(riffle beetles)</small></a></li> <li><a href="/wiki/Eulichadidae" title="Eulichadidae">Eulichadidae <small>(forest stream beetles)</small></a></li> <li><a href="/wiki/Heteroceridae" title="Heteroceridae">Heteroceridae <small>(variegated mud-loving beetles)</small></a></li> <li><a href="/wiki/Limnichidae" title="Limnichidae">Limnichidae <small>(minute mud beetles)</small></a></li> <li><a href="/wiki/Lutrochidae" class="mw-redirect" title="Lutrochidae">Lutrochidae <small>(travertine beetles)</small></a></li> <li><a href="/wiki/Water-penny_beetles" class="mw-redirect" title="Water-penny beetles">Psephenidae <small>(water-penny beetles)</small></a></li> <li><a href="/wiki/Ptilodactylidae" title="Ptilodactylidae">Ptilodactylidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Dascilloidea" title="Dascilloidea">Dascilloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dascillidae" title="Dascillidae">Dascillidae <small>(soft bodied plant beetles)</small></a></li> <li><a href="/wiki/Rhipiceridae" title="Rhipiceridae">Rhipiceridae <small>(cicada beetle, cicada parasite beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Elateroidea" title="Elateroidea">Elateroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Artematopodidae" title="Artematopodidae">Artematopodidae <small>(soft-bodied plant beetles)</small></a></li> <li><a href="/wiki/Brachypsectridae" title="Brachypsectridae">Brachypsectridae <small>(Texas beetles)</small></a></li> <li><a href="/wiki/Soldier_beetle" title="Soldier beetle">Cantharidae <small>(soldier beetles)</small></a></li> <li><a href="/wiki/Cerophytidae" title="Cerophytidae">Cerophytidae <small>(rare click beetles)</small></a></li> <li><a href="/wiki/Click_beetle" title="Click beetle">Elateridae <small>(click beetles)</small></a></li> <li><a href="/wiki/Eucnemidae" title="Eucnemidae">Eucnemidae <small>(false click beetles)</small></a></li> <li><a href="/wiki/Jurasaidae" title="Jurasaidae">Jurasaidae</a></li> <li><a href="/wiki/Firefly" title="Firefly">Lampyridae <small>(fireflies)</small></a></li> <li><a href="/wiki/Lycidae" title="Lycidae">Lycidae <small>(net-winged beetles)</small></a></li> <li><a href="/wiki/Omethidae" title="Omethidae">Omethidae <small>(false fireflies, long-lipped beetles)</small></a></li> <li><a href="/wiki/Phengodidae" title="Phengodidae">Phengodidae <small>(glowworm beetles)</small></a></li> <li><a href="/wiki/Rhagophthalmidae" title="Rhagophthalmidae">Rhagophthalmidae</a></li> <li><a href="/wiki/Sinopyrophoridae" class="mw-redirect" title="Sinopyrophoridae">Sinopyrophoridae</a></li> <li><a href="/wiki/Throscidae" title="Throscidae">Throscidae <small>(false metallic wood-boring beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipidae <small>(<i>Rhinorhipus tamborinensis</i>)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scirtoidea" title="Scirtoidea">Scirtoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Clambidae" title="Clambidae">Clambidae</a></li> <li><a href="/wiki/Declinia" title="Declinia">Decliniidae <small>(<i>Declinia relicta</i>)</small></a></li> <li><a href="/wiki/Eucinetidae" title="Eucinetidae">Eucinetidae <small>(plate-thigh beetles)</small></a></li> <li><a href="/wiki/Scirtidae" title="Scirtidae">Scirtidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Belohina" title="Belohina">Belohinidae <small>(<i>Belohina inexpectata</i>)</small></a></li> <li><a href="/wiki/Bolboceratidae" title="Bolboceratidae">Bolboceratidae</a></li> <li><a href="/wiki/False_stag_beetle" title="False stag beetle">Diphyllostomatidae <small>(false stag beetles)</small></a></li> <li><a href="/wiki/Geotrupidae" title="Geotrupidae">Geotrupidae <small>(dor beetles)</small></a></li> <li><a href="/wiki/Glaphyridae" title="Glaphyridae">Glaphyridae <small>(bumble bee scarab beetles)</small></a></li> <li><a href="/wiki/Glaresis" title="Glaresis">Glaresidae <small>(enigmatic scarab beetles)</small></a></li> <li><a href="/wiki/Hybosoridae" title="Hybosoridae">Hybosoridae <small>(scavenger scarab beetles)</small></a></li> <li><a href="/wiki/Stag_beetle" title="Stag beetle">Lucanidae <small>(stag beetles)</small></a></li> <li><a href="/wiki/Ochodaeidae" title="Ochodaeidae">Ochodaeidae <small>(sand-loving scarab beetles)</small></a></li> <li><a href="/wiki/Passalidae" title="Passalidae">Passalidae <small>(betsy beetles)</small></a></li> <li><a href="/wiki/Rain_beetle" title="Rain beetle">Pleocomidae <small>(rain beetles)</small></a></li> <li><a href="/wiki/Scarabaeidae" title="Scarabaeidae">Scarabaeidae <small>(scarabs)</small></a></li> <li><a href="/wiki/Trogidae" title="Trogidae">Trogidae <small>(hide beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Staphyliniformia" title="Staphyliniformia">Staphyliniformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Histeroidea" title="Histeroidea">Histeroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Histeridae" title="Histeridae">Histeridae <small>(clown beetles)</small></a></li> <li><a href="/wiki/Sphaerites" title="Sphaerites">Sphaeritidae <small>(false clown beetles)</small></a></li> <li><a href="/wiki/Syntelia" title="Syntelia">Synteliidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Hydrophiloidea" title="Hydrophiloidea">Hydrophiloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Epimetopidae" title="Epimetopidae">Epimetopidae</a></li> <li><a href="/wiki/Georissus" title="Georissus">Georissidae <small>(minute mud-loving beetles)</small></a></li> <li><a href="/wiki/Helophorus" title="Helophorus">Helophoridae</a></li> <li><a href="/wiki/Hydrochidae" class="mw-redirect" title="Hydrochidae">Hydrochidae</a></li> <li><a href="/wiki/Hydrophilidae" title="Hydrophilidae">Hydrophilidae <small>(water scavenger beetles)</small></a></li> <li><a href="/wiki/Spercheidae" class="mw-redirect" title="Spercheidae">Spercheidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Staphylinoidea" title="Staphylinoidea">Staphylinoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agyrtidae" title="Agyrtidae">Agyrtidae <small>(primitive carrion beetles)</small></a></li> <li><a href="/wiki/Hydraenidae" title="Hydraenidae">Hydraenidae</a></li> <li><a href="/wiki/Leiodidae" title="Leiodidae">Leiodidae <small>(round fungus beetles)</small></a></li> <li><a href="/wiki/Ptiliidae" title="Ptiliidae">Ptiliidae <small>(feather-winged beetles)</small></a></li> <li><a href="/wiki/Silphidae" title="Silphidae">Silphidae <small>(carrion beetles)</small></a></li> <li><a href="/wiki/Rove_beetle" title="Rove beetle">Staphylinidae <small>(rove beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/List_of_subgroups_of_the_order_Coleoptera" title="List of subgroups of the order Coleoptera">List of subgroups of the order Coleoptera</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Polyphaga</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q147877" class="extiw" title="wikidata:Q147877">Q147877</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Polyphaga_(Coleoptera)" class="extiw" title="wikispecies:Polyphaga (Coleoptera)">Polyphaga (Coleoptera)</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Animal_Diversity_Web" title="Animal Diversity Web">ADW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://animaldiversity.org/accounts/Polyphaga/">Polyphaga</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Faunal_Directory" title="Australian Faunal Directory">AFD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://biodiversity.org.au/afd/taxa/Polyphaga">Polyphaga</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/BugGuide" title="BugGuide">BugGuide</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://bugguide.net/node/view/40606">40606</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/46837289">46837289</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:faunaeur.org:taxname:11014">11014</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea (new)</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://fauna-eu.org/cdm_dataportal/taxon/61b64244-2103-40e4-b029-1eb349ac7565">61b64244-2103-40e4-b029-1eb349ac7565</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/71130">71130</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=112747">112747</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Biodiversity_Network" title="National Biodiversity Network">NBN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://data.nbn.org.uk/Taxa/NHMSYS0001719665">NHMSYS0001719665</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=41084">41084</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/077114ee-5104-4a42-96ed-7558691f819e">077114ee-5104-4a42-96ed-7558691f819e</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=684689">684689</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=69199">69199</a></span></span></li> <li><span style="white-space:nowrap;">PPE: <span class="uid"><a rel="nofollow" class="external text" href="https://bladmineerders.nl/parasites/animalia/arthropoda/insecta/coleoptera/polyphaga/">polyphaga</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div 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