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class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" 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href="https://ca.wikipedia.org/wiki/Wollemia" title="Wollemia – Catalan" lang="ca" hreflang="ca" data-title="Wollemia" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb badge-Q70893996 mw-list-item" title=""><a href="https://ceb.wikipedia.org/wiki/Wollemia" title="Wollemia – Cebuano" lang="ceb" hreflang="ceb" data-title="Wollemia" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs badge-Q70893996 mw-list-item" title=""><a href="https://cs.wikipedia.org/wiki/Wollemie" title="Wollemie – Czech" lang="cs" hreflang="cs" data-title="Wollemie" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/Wollemia" title="Wollemia – Welsh" lang="cy" hreflang="cy" data-title="Wollemia" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Wollemia" title="Wollemia – Danish" lang="da" hreflang="da" data-title="Wollemia" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de badge-Q70894304 mw-list-item" title=""><a href="https://de.wikipedia.org/wiki/Wollemia" title="Wollemia – German" lang="de" hreflang="de" data-title="Wollemia" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Volleemia_(perekond)" title="Volleemia (perekond) – Estonian" lang="et" hreflang="et" data-title="Volleemia (perekond)" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%92%CE%BF%CE%BB%CE%AD%CE%BC%CE%B9%CE%B1" title="Βολέμια – Greek" lang="el" hreflang="el" data-title="Βολέμια" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es badge-Q70893996 mw-list-item" title=""><a href="https://es.wikipedia.org/wiki/Wollemia" title="Wollemia – Spanish" lang="es" hreflang="es" data-title="Wollemia" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo badge-Q70893996 mw-list-item" title=""><a href="https://eo.wikipedia.org/wiki/Wollemia" title="Wollemia – Esperanto" lang="eo" hreflang="eo" data-title="Wollemia" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-fr badge-Q70893996 mw-list-item" title=""><a href="https://fr.wikipedia.org/wiki/Wollemia" title="Wollemia – French" lang="fr" hreflang="fr" data-title="Wollemia" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Wollemia" title="Wollemia – Icelandic" lang="is" hreflang="is" data-title="Wollemia" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-frr mw-list-item"><a href="https://frr.wikipedia.org/wiki/Wollemia" title="Wollemia – Northern Frisian" lang="frr" hreflang="frr" data-title="Wollemia" data-language-autonym="Nordfriisk" data-language-local-name="Northern Frisian" class="interlanguage-link-target"><span>Nordfriisk</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Wollemia" title="Wollemia – Norwegian Nynorsk" lang="nn" hreflang="nn" data-title="Wollemia" data-language-autonym="Norsk nynorsk" data-language-local-name="Norwegian Nynorsk" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%92%D0%BE%D0%BB%D0%BB%D0%B5%D0%BC%D0%B8%D1%8F" title="Воллемия – Russian" lang="ru" hreflang="ru" data-title="Воллемия" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link 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<tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Wakehurst_Place_woodland_Wollemi_pine.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Wakehurst_Place_woodland_Wollemi_pine.jpg/220px-Wakehurst_Place_woodland_Wollemi_pine.jpg" decoding="async" width="220" height="330" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Wakehurst_Place_woodland_Wollemi_pine.jpg/330px-Wakehurst_Place_woodland_Wollemi_pine.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Wakehurst_Place_woodland_Wollemi_pine.jpg/440px-Wakehurst_Place_woodland_Wollemi_pine.jpg 2x" data-file-width="2666" data-file-height="4000" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Mature tree </td></tr> <tr style="color:inherit; text-align: center; background-color: rgb(180,250,180)"> <th colspan="2"><div style="text-align: center"><a href="/wiki/Conservation_status" title="Conservation status">Conservation status</a></div> </th></tr> <tr> <td colspan="2"><div style="text-align: center"><span class="mw-default-size skin-invert-image" typeof="mw:File/Frameless"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4b/Status_iucn3.1_CR.svg/220px-Status_iucn3.1_CR.svg.png" decoding="async" width="220" height="59" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4b/Status_iucn3.1_CR.svg/330px-Status_iucn3.1_CR.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4b/Status_iucn3.1_CR.svg/440px-Status_iucn3.1_CR.svg.png 2x" data-file-width="512" data-file-height="137" /></span></span><br /><a href="/wiki/Critically_endangered_species" class="mw-redirect" title="Critically endangered species">Critically Endangered</a> <small> (<a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN 3.1</a>)<sup id="cite_ref-iucn_status_12_November_2021_1-0" class="reference"><a href="#cite_note-iucn_status_12_November_2021-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></small></div> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(180,250,180)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Wollemia" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Plant" title="Plant">Plantae</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Vascular_plant" title="Vascular plant">Tracheophytes</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Gymnosperm" title="Gymnosperm">Gymnospermae</a> </td></tr> <tr> <td>Division: </td> <td><a href="/wiki/Conifer" title="Conifer">Pinophyta</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Conifer" title="Conifer">Pinopsida</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Araucariales" class="mw-redirect" title="Araucariales">Araucariales</a> </td></tr> <tr> <td>Family: </td> <td><a href="/wiki/Araucariaceae" title="Araucariaceae">Araucariaceae</a> </td></tr> <tr> <td>Genus: </td> <td><a class="mw-selflink selflink"><i>Wollemia</i></a><br /><small>W.G.Jones, K.D.Hill & J.M.Allen</small> </td></tr> <tr> <td>Species: </td> <td><div style="display:inline" class="species"><i><b>W. nobilis</b></i></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(180,250,180)"><a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">Binomial name</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><b><span class="binomial"><span style="font-weight:normal;"></span><i>Wollemia nobilis</i></span></b><br /><div style="font-size: 85%;">W.G.Jones, K.D.Hill & J.M.Allen, 1995</div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Wollemia</b></i> is a genus of <a href="/wiki/Coniferous" class="mw-redirect" title="Coniferous">coniferous</a> trees in the family <a href="/wiki/Araucariaceae" title="Araucariaceae">Araucariaceae</a>, endemic to Australia. It represents only one of three living genera in the family, alongside <i><a href="/wiki/Araucaria" title="Araucaria">Araucaria</a></i> and <i><a href="/wiki/Agathis" title="Agathis">Agathis</a></i> (being more closely related to the latter). The genus has only a single known species, <i><b>Wollemia nobilis,</b></i> commonly known as the <b>Wollemi Pine</b> (though it is not a true pine) which was discovered in 1994 in a temperate <a href="/wiki/Rainforest" title="Rainforest">rainforest</a> wilderness area of the <a href="/wiki/Wollemi_National_Park" title="Wollemi National Park">Wollemi National Park</a> in <a href="/wiki/New_South_Wales" title="New South Wales">New South Wales</a>. It was growing in a remote series of narrow, steep-sided, <a href="/wiki/Sandstone" title="Sandstone">sandstone</a> <a href="/wiki/Gorge" class="mw-redirect" title="Gorge">gorges</a> 150 km (93 mi) north-west of <a href="/wiki/Sydney" title="Sydney">Sydney</a>. The genus is named after the National Park.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>The Wollemi pine is classified as <a href="/wiki/Critically_endangered" class="mw-redirect" title="Critically endangered">critically endangered</a> (CR) on the <a href="/wiki/International_Union_for_Conservation_of_Nature" title="International Union for Conservation of Nature">IUCN</a>'s <a href="/wiki/IUCN_Red_List" title="IUCN Red List">Red List</a>,<sup id="cite_ref-iucn_status_12_November_2021_1-1" class="reference"><a href="#cite_note-iucn_status_12_November_2021-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> and is legally protected in Australia.<sup id="cite_ref-NPWS_3-0" class="reference"><a href="#cite_note-NPWS-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> After it was discovered that the trees could be successfully cloned, new trees were potted up in the Botanic Gardens of Sydney and <a href="/wiki/Australian_Botanic_Garden_Mount_Annan" title="Australian Botanic Garden Mount Annan">Mount Annan</a> and planted as far away as the <a href="/wiki/Humboldt_Botanical_Gardens" title="Humboldt Botanical Gardens">Humboldt Botanical Garden</a> near Eureka, California.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>A Recovery Plan has been drawn up, outlining strategies for the management of this fragile population. The overall objective is to ensure that the species remains viable in the long term.<sup id="cite_ref-NPWS_3-1" class="reference"><a href="#cite_note-NPWS-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Australian prime ministers and foreign affairs ministers have presented Wollemi pines to various dignitaries around the world.<sup id="cite_ref-Brack-2018_5-0" class="reference"><a href="#cite_note-Brack-2018-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>Although often described as a "<a href="/wiki/Living_fossil" title="Living fossil">living fossil</a>", there are no unambiguous fossils of <i>Wollemia</i> and potential fossil records of it have been considered uncertain.<sup id="cite_ref-Seyfullah-2023_6-0" class="reference"><a href="#cite_note-Seyfullah-2023-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Wollemia nobilis</i> is an <a href="/wiki/Evergreen" title="Evergreen">evergreen</a> <a href="/wiki/Tree" title="Tree">tree</a> reaching 25–40 m (82–131 ft) tall. The <a href="/wiki/Bark_(botany)" title="Bark (botany)">bark</a> is very distinctive, dark brown, and knobbly, quoted as resembling the breakfast cereal <a href="/wiki/Coco_Pops" class="mw-redirect" title="Coco Pops">Coco Pops</a>.<sup id="cite_ref-woodford2002_7-0" class="reference"><a href="#cite_note-woodford2002-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> The tree <a href="/wiki/Coppice" class="mw-redirect" title="Coppice">coppices</a> readily, and most specimens are multiple-trunked or appear as clumps of trunks thought to derive from old coppice growth, with some consisting of up to 100 stems of differing sizes.<sup id="cite_ref-NPWS_3-2" class="reference"><a href="#cite_note-NPWS-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The branching is unusual in that nearly all the side branches never have further branching. After a few years, each branch either terminates in a <a href="/wiki/Conifer_cone" title="Conifer cone">cone</a> (either male or female) or ceases growth. After this, or when the cone becomes mature, the <a href="/wiki/Branch" title="Branch">branch</a> dies. New branches then arise from dormant buds on the main trunk. Rarely, a side branch will turn erect and develop into a secondary trunk, which then bears a new set of side branches. </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:B5000316-Wollemi_pine_Wollemia_nobilis_.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/36/B5000316-Wollemi_pine_Wollemia_nobilis_.jpg/220px-B5000316-Wollemi_pine_Wollemia_nobilis_.jpg" decoding="async" width="220" height="338" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/36/B5000316-Wollemi_pine_Wollemia_nobilis_.jpg/330px-B5000316-Wollemi_pine_Wollemia_nobilis_.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/36/B5000316-Wollemi_pine_Wollemia_nobilis_.jpg/440px-B5000316-Wollemi_pine_Wollemia_nobilis_.jpg 2x" data-file-width="521" data-file-height="800" /></a><figcaption>A 4-year-old potted Wollemi pine grown from cutting</figcaption></figure> <p>The <a href="/wiki/Leaf" title="Leaf">leaves</a> are flat linear, 3–8 cm (1.2–3.1 in) long and 2–5 mm (0.079–0.197 in) broad. They are arranged spirally on the shoot but twisted at the base to appear in two or four flattened ranks. As the leaves mature, they develop from bright lime-green to a more yellowish-green.<sup id="cite_ref-FF_8-0" class="reference"><a href="#cite_note-FF-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> The seed cones are green, 6–12 cm (2.4–4.7 in) long and 5–10 cm (2.0–3.9 in) in diameter, and mature about 18–20 months after <a href="/wiki/Wind_pollination" class="mw-redirect" title="Wind pollination">wind pollination</a>. They disintegrate at maturity to release the <a href="/wiki/Seed" title="Seed">seeds</a> which are small and brown, thin and papery with a wing around the edge to aid wind-dispersal.<sup id="cite_ref-NPWS_3-3" class="reference"><a href="#cite_note-NPWS-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The male (<a href="/wiki/Pollen" title="Pollen">pollen</a>) cones are slender conic, 5–11 cm (2.0–4.3 in) long and 1–2 cm (0.39–0.79 in) broad and reddish-brown in colour and are lower on the tree than the seed cones.<sup id="cite_ref-NPWS_3-4" class="reference"><a href="#cite_note-NPWS-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Seedlings appear to be slow-growing<sup id="cite_ref-NPWS_3-5" class="reference"><a href="#cite_note-NPWS-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> and mature trees are extremely long-lived; some of the older individuals today are estimated to be between 500 and 1,000 years old.<sup id="cite_ref-FF_8-1" class="reference"><a href="#cite_note-FF-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <ul class="gallery mw-gallery-packed"> <li class="gallerybox" style="width: 98px"> <div class="thumb" style="width: 96px;"><span typeof="mw:File"><a href="/wiki/File:Wollemia_nobilis_apical_shoot_3.jpg" class="mw-file-description" title="Apical shoots"><img alt="Apical shoots" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Wollemia_nobilis_apical_shoot_3.jpg/144px-Wollemia_nobilis_apical_shoot_3.jpg" decoding="async" width="96" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Wollemia_nobilis_apical_shoot_3.jpg/216px-Wollemia_nobilis_apical_shoot_3.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/57/Wollemia_nobilis_apical_shoot_3.jpg/288px-Wollemia_nobilis_apical_shoot_3.jpg 2x" data-file-width="837" data-file-height="1047" /></a></span></div> <div class="gallerytext">Apical shoots</div> </li> <li class="gallerybox" style="width: 162px"> <div class="thumb" style="width: 160px;"><span typeof="mw:File"><a href="/wiki/File:Wollemia_nobilis_Wolemia_szlachetna_2019-07-20_05.jpg" class="mw-file-description" title="Leaves"><img alt="Leaves" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8c/Wollemia_nobilis_Wolemia_szlachetna_2019-07-20_05.jpg/240px-Wollemia_nobilis_Wolemia_szlachetna_2019-07-20_05.jpg" decoding="async" width="160" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8c/Wollemia_nobilis_Wolemia_szlachetna_2019-07-20_05.jpg/360px-Wollemia_nobilis_Wolemia_szlachetna_2019-07-20_05.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8c/Wollemia_nobilis_Wolemia_szlachetna_2019-07-20_05.jpg/480px-Wollemia_nobilis_Wolemia_szlachetna_2019-07-20_05.jpg 2x" data-file-width="4242" data-file-height="3181" /></a></span></div> <div class="gallerytext">Leaves</div> </li> <li class="gallerybox" style="width: 92px"> <div class="thumb" style="width: 90px;"><span typeof="mw:File"><a href="/wiki/File:Wollemia_bark.JPG" class="mw-file-description" title="Bark of W. nobilis"><img alt="Bark of W. nobilis" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/5d/Wollemia_bark.JPG/135px-Wollemia_bark.JPG" decoding="async" width="90" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/5d/Wollemia_bark.JPG/202px-Wollemia_bark.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/5d/Wollemia_bark.JPG/270px-Wollemia_bark.JPG 2x" data-file-width="900" data-file-height="1200" /></a></span></div> <div class="gallerytext">Bark of <i>W. nobilis</i></div> </li> <li class="gallerybox" style="width: 92px"> <div class="thumb" style="width: 90px;"><span typeof="mw:File"><a href="/wiki/File:Wollemia_nobilis_(Wollemi_pine)_seedling.jpg" class="mw-file-description" title="A 4 month old Wollemi pine seedling"><img alt="A 4 month old Wollemi pine seedling" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/68/Wollemia_nobilis_%28Wollemi_pine%29_seedling.jpg/135px-Wollemia_nobilis_%28Wollemi_pine%29_seedling.jpg" decoding="async" width="90" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/68/Wollemia_nobilis_%28Wollemi_pine%29_seedling.jpg/202px-Wollemia_nobilis_%28Wollemi_pine%29_seedling.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/68/Wollemia_nobilis_%28Wollemi_pine%29_seedling.jpg/270px-Wollemia_nobilis_%28Wollemi_pine%29_seedling.jpg 2x" data-file-width="3024" data-file-height="4032" /></a></span></div> <div class="gallerytext">A 4 month old Wollemi pine seedling</div> </li> <li class="gallerybox" style="width: 133.33333333333px"> <div class="thumb" style="width: 131.33333333333px;"><span typeof="mw:File"><a href="/wiki/File:Wollemia_nobilis_cone.jpg" class="mw-file-description" title="Young W. nobilis pollen cone"><img alt="Young W. nobilis pollen cone" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cb/Wollemia_nobilis_cone.jpg/197px-Wollemia_nobilis_cone.jpg" decoding="async" width="132" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cb/Wollemia_nobilis_cone.jpg/295px-Wollemia_nobilis_cone.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cb/Wollemia_nobilis_cone.jpg/394px-Wollemia_nobilis_cone.jpg 2x" data-file-width="2296" data-file-height="2100" /></a></span></div> <div class="gallerytext">Young <i>W. nobilis</i> pollen cone</div> </li> <li class="gallerybox" style="width: 82px"> <div class="thumb" style="width: 80px;"><span typeof="mw:File"><a href="/wiki/File:Wollemia_nobilis_kz7.jpg" class="mw-file-description" title="Female cones"><img alt="Female cones" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/61/Wollemia_nobilis_kz7.jpg/120px-Wollemia_nobilis_kz7.jpg" decoding="async" width="80" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/61/Wollemia_nobilis_kz7.jpg/180px-Wollemia_nobilis_kz7.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/61/Wollemia_nobilis_kz7.jpg/240px-Wollemia_nobilis_kz7.jpg 2x" data-file-width="2912" data-file-height="4368" /></a></span></div> <div class="gallerytext">Female cones</div> </li> <li class="gallerybox" style="width: 82px"> <div class="thumb" style="width: 80px;"><span typeof="mw:File"><a href="/wiki/File:Wollemia_nobilis_at_Kew_Gardens,_July_2016.jpg" class="mw-file-description" title="Specimen at Kew Gardens"><img alt="Specimen at Kew Gardens" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/12/Wollemia_nobilis_at_Kew_Gardens%2C_July_2016.jpg/120px-Wollemia_nobilis_at_Kew_Gardens%2C_July_2016.jpg" decoding="async" width="80" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/12/Wollemia_nobilis_at_Kew_Gardens%2C_July_2016.jpg/180px-Wollemia_nobilis_at_Kew_Gardens%2C_July_2016.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/12/Wollemia_nobilis_at_Kew_Gardens%2C_July_2016.jpg/240px-Wollemia_nobilis_at_Kew_Gardens%2C_July_2016.jpg 2x" data-file-width="3456" data-file-height="5184" /></a></span></div> <div class="gallerytext">Specimen at <a href="/wiki/Kew_Gardens" title="Kew Gardens">Kew Gardens</a></div> </li> </ul> <div class="mw-heading mw-heading2"><h2 id="History">History</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=2" title="Edit section: History"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Discovery">Discovery</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=3" title="Edit section: Discovery"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Wollemia_cones.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fc/Wollemia_cones.JPG/220px-Wollemia_cones.JPG" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fc/Wollemia_cones.JPG/330px-Wollemia_cones.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fc/Wollemia_cones.JPG/440px-Wollemia_cones.JPG 2x" data-file-width="2592" data-file-height="1944" /></a><figcaption>Male and female <i>W. nobilis</i> cones</figcaption></figure> <p>The discovery, on or about 10 September 1994, by <a href="/wiki/David_Noble_(canyoner)" title="David Noble (canyoner)">David Noble</a>, Michael Casteleyn, and Tony Zimmerman, occurred only because the group had been systematically exploring the area, looking for new canyons.<sup id="cite_ref-woodford2002_7-1" class="reference"><a href="#cite_note-woodford2002-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> Noble had good botanical knowledge, and quickly recognised the trees as unusual because of the unique bark, and worthy of further investigation. </p><p>He took specimens to work for identification, expecting someone to be able to identify the plants.<sup id="cite_ref-NPR_9-0" class="reference"><a href="#cite_note-NPR-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>a<span class="cite-bracket">]</span></a></sup> His specimens were identified by Wyn Jones, a botanist with National Parks and Jan Allen from the Botanical Gardens. After the identification was made, National Parks then went under a veil of secrecy, with the discoverers not learning the full magnitude of their discovery for about six months. National Parks came close to damaging the stand when a helicopter used to collect cones inadvertently pruned one of the pines with its rotor.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> The species was subsequently named after David Noble.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p><p>The first illustrations of the Wollemi Pine were drawn by David Mackay, a botanical artist and scientific illustrator who was working at the <a href="/wiki/Royal_Botanic_Garden,_Sydney" title="Royal Botanic Garden, Sydney">Royal Botanic Gardens</a> in Sydney when the species was discovered.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>Further study would be needed to establish its relationship to other conifers. The initial suspicion was that it had certain characteristics of the 200 million year-old family <a href="/wiki/Araucariaceae" title="Araucariaceae">Araucariaceae</a>, but was not similar to any living species in the family. Comparison with living and <a href="/wiki/Fossil" title="Fossil">fossilised</a> Araucariaceae proved that it was a member of that family, and it has been placed into a new genus, beside the genera <i><a href="/wiki/Agathis" title="Agathis">Agathis</a></i> and <i><a href="/wiki/Araucaria" title="Araucaria">Araucaria</a></i>. </p><p>Fewer than 60 adult trees are known to be growing wild in four locations, not far apart. It is very difficult to count individuals, as most trees are multi-stemmed and may have a connected root system. <a href="/wiki/Genetic_testing" title="Genetic testing">Genetic testing</a> has revealed that all the specimens are genetically indistinguishable, suggesting that the species has been through a <a href="/wiki/Genetic_bottleneck" class="mw-redirect" title="Genetic bottleneck">genetic bottleneck</a> 10,000–26,000 years ago, in which its population became so low (possibly just one or two individuals) that all <a href="/wiki/Genetic_variability" title="Genetic variability">genetic variability</a> was lost.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Molecular_Ecology_2003_17-0" class="reference"><a href="#cite_note-Molecular_Ecology_2003-17"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Conservation">Conservation</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=4" title="Edit section: Conservation"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading4"><h4 id="Threats">Threats</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=5" title="Edit section: Threats"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In November 2005, wild-growing trees were found to be infected with <i><a href="/wiki/Phytophthora_cinnamomi" title="Phytophthora cinnamomi">Phytophthora cinnamomi</a></i>.<sup id="cite_ref-ABC_Nov_2005_18-0" class="reference"><a href="#cite_note-ABC_Nov_2005-18"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> New South Wales park rangers believe the virulent <a href="/wiki/Water_mould" class="mw-redirect" title="Water mould">water mould</a> was introduced by unauthorised visitors to the site, the location of which is still undisclosed to the public.<sup id="cite_ref-ABC_Nov_2005_18-1" class="reference"><a href="#cite_note-ABC_Nov_2005-18"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>The grove of Wollemia trees was endangered by fire during the <a href="/wiki/2019%E2%80%9320_Australian_bushfire_season#Blue_Mountains_and_Hawkesbury" title="2019–20 Australian bushfire season">2019–2020 Australian bushfire season</a>.<sup id="cite_ref-SMH-hannam2020_19-0" class="reference"><a href="#cite_note-SMH-hannam2020-19"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> They were saved by specialist firefighters from the National Parks and Wildlife Service, supported by the Rural Fire Service who installed an irrigation system as well as dropping <a href="/wiki/Fire_retardant" title="Fire retardant">retardant.</a><sup id="cite_ref-NPR_9-1" class="reference"><a href="#cite_note-NPR-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-SMH-hannam2020_19-1" class="reference"><a href="#cite_note-SMH-hannam2020-19"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Global_involvement_in_conservation">Global involvement in conservation</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=6" title="Edit section: Global involvement in conservation"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Wollemia_Nobilis_Trautmannsdorf_Merano_2018.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/74/Wollemia_Nobilis_Trautmannsdorf_Merano_2018.jpg/220px-Wollemia_Nobilis_Trautmannsdorf_Merano_2018.jpg" decoding="async" width="220" height="329" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/74/Wollemia_Nobilis_Trautmannsdorf_Merano_2018.jpg/330px-Wollemia_Nobilis_Trautmannsdorf_Merano_2018.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/74/Wollemia_Nobilis_Trautmannsdorf_Merano_2018.jpg/440px-Wollemia_Nobilis_Trautmannsdorf_Merano_2018.jpg 2x" data-file-width="2592" data-file-height="3872" /></a><figcaption><i>Wollemia nobilis</i> in the botanical garden of <a href="/wiki/Trauttmansdorff_Castle_Gardens" title="Trauttmansdorff Castle Gardens">Trauttmansdorff Castle Gardens</a> in <a href="/wiki/Merano" title="Merano">Merano</a>, <a href="/wiki/Italy" title="Italy">Italy</a> in 2018</figcaption></figure> <p>"Home gardeners become accidental citizen scientists for Wollemi Pine" was the headline of a 2023 news article reporting results of an unusual conservation strategy underway since 2005.<sup id="cite_ref-physorg-2023_23-0" class="reference"><a href="#cite_note-physorg-2023-23"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> Ten years after safeguarding of rooted branch cuttings had been initiated in botanic gardens around the world, commercial growers were authorized to receive surplus cuttings to propagate for sales to their own customers. This unusual management decision for an endangered plant owed to the "huge public interest in this rare tree" and as an experiment to test whether commercial availability would serve "to protect wild populations from illegal collecting."<sup id="cite_ref-physorg-2023_23-1" class="reference"><a href="#cite_note-physorg-2023-23"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> </p><p>Results were tallied from more than 1,500 people from 31 countries who were growing the cuttings and responded to an online survey created by two plant scientists in Australia.<sup id="cite_ref-offord-2023_24-0" class="reference"><a href="#cite_note-offord-2023-24"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> Results indicated that the species grows well where climate is <a href="/wiki/Temperate_climate" title="Temperate climate">temperate</a> and there is adequate year-round rainfall. <a href="/wiki/Loam" title="Loam">Loamy soils</a> with good drainage proved to be ideal. The team concluded, "This study demonstrates that it is feasible to establish Wollemi pines in many parts of the world and under different climates and cultural regimes, which can help conserve this species in the face of climate change and other threats."<sup id="cite_ref-offord-2023_24-1" class="reference"><a href="#cite_note-offord-2023-24"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Cultivation_and_uses">Cultivation and uses<span class="anchor" id="Cultivation"></span><span class="anchor" id="Uses"></span></h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=7" title="Edit section: Cultivation and uses"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Wollemia_nobilis_sapling.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Wollemia_nobilis_sapling.jpg/200px-Wollemia_nobilis_sapling.jpg" decoding="async" width="200" height="228" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Wollemia_nobilis_sapling.jpg/300px-Wollemia_nobilis_sapling.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/38/Wollemia_nobilis_sapling.jpg/400px-Wollemia_nobilis_sapling.jpg 2x" data-file-width="1360" data-file-height="1553" /></a><figcaption>Cultivated <i>Wollemia nobilis</i></figcaption></figure> <p>A propagation programme made Wollemi pine specimens available to botanical gardens, first in Australia in 2006 and subsequently throughout the world. It may prove to be a valuable tree for ornament, either planted in open ground or for tubs and planters. In Australia, potted native Wollemi pines have been promoted as a Christmas tree.<sup id="cite_ref-acf_25-0" class="reference"><a href="#cite_note-acf-25"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> It is also proving to be more adaptable and cold-hardy than its restricted <a href="/wiki/Temperate" class="mw-redirect" title="Temperate">temperate</a>-<a href="/wiki/Subtropical" class="mw-redirect" title="Subtropical">subtropical</a>, humid distribution would suggest, tolerating temperatures between −5 and 45 °C (23 and 113 °F), with reports, from Japan and the USA, that it can survive down to −12 °C (10 °F). A grove of Wollemi pines planted in <a href="/wiki/Inverewe_Garden" title="Inverewe Garden">Inverewe Garden</a>, Scotland, believed to be the most northerly location of any successful planting, have survived temperatures of −7 °C (19 °F), recorded in January 2010.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> It also handles both full sun and full shade. Like many other Australian trees, <i>Wollemia</i> is susceptible to the pathogenic water mould <i><a href="/wiki/Phytophthora_cinnamomi" title="Phytophthora cinnamomi">Phytophthora cinnamomi</a></i>, so this may limit its potential as a <a href="/wiki/Timber" class="mw-redirect" title="Timber">timber</a> tree.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> </p><p>The <a href="/wiki/Royal_Botanic_Gardens,_Sydney" class="mw-redirect" title="Royal Botanic Gardens, Sydney">Royal Botanic Gardens</a> in Sydney have published information on how to grow Wollemi pines from seed which has been harvested from helicopters from the forest trees. The majority of seeds that fall from the cone are not viable so need to be sorted to retain the plump and dark ones. These can then be sown on top of seed raising mix and watered. Once the water has drained through the mix, the pot should be placed in a plastic bag and refrigerated for two weeks. After this, the pot should be removed from the plastic bag and placed somewhere warm but not very sunny until the seed germinates (remembering to keep them moist but not wet). This could take several months.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> </p><p>Examples of the species can be viewed at <a href="/wiki/The_Tasmanian_Arboretum" class="mw-redirect" title="The Tasmanian Arboretum">The Tasmanian Arboretum</a> and at Giardini di Villa della Pergola in Alassio, Italy.<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Care">Care</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=8" title="Edit section: Care"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The Wollemi pine is extremely hardy and versatile in cultivation. Despite it being an endangered species, it is easy to grow and requires relatively low maintenance. It will adapt to a diverse range of climatic zones, thriving in full sun to semi shaded outdoor positions. They can be maintained in a pot almost indefinitely, and make good container plants for patios, verandas, and courtyards. Because it tolerates air conditioning, it can also be used as an indoor decorative plant. They require well-drained soil and protection from frost.<sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Pruning">Pruning</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=9" title="Edit section: Pruning"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>When pruning the Wollemi pine, use sterile secateurs at any time of year to retain its compact form. It can be pruned heavily with up to two thirds of the plant size removed. Pruning heavily can be done on the apical growth and the branches. The best time to prune is during the winter months. </p> <div class="mw-heading mw-heading3"><h3 id="Growth_rate">Growth rate</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=10" title="Edit section: Growth rate"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The Wollemi pine has very controlled growth, especially if it is kept in a pot. It may take up to 25 years to reach 6 metres (20 ft) in height. </p> <div class="mw-heading mw-heading2"><h2 id="Phylogeny">Phylogeny</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=11" title="Edit section: Phylogeny"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The genus <i>Wollemia</i> shares morphological characteristics with the genera <i><a href="/wiki/Araucaria" title="Araucaria">Araucaria</a></i> and <i><a href="/wiki/Agathis" title="Agathis">Agathis</a></i>. <i>Wollemia</i> and <i>Araucaria</i> both have closely crowded sessile and amphistomatic (producing stomata on both sides of the leaf) leaves, and <a href="/wiki/Glossary_of_leaf_morphology#aristate" title="Glossary of leaf morphology">aristate</a> bract scales, while <i>Wollemia</i> and <i>Agathis</i> both have fully fused bracts, ovuliferous scales, and winged seeds.<sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> Scrutiny of the fossil record likewise does not clarify <i>Wollemia</i><span class="nowrap" style="padding-left:0.1em;">'s</span> relationship to <i>Araucaria</i> or <i>Agathis</i>, since the former has similarly disparate leaf characters in its adult and juvenile forms, and the latter has similar cone characters.<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> Further, the recent description of several extinct genera within the Araucariaceae points to complex relationships within the family and a significant loss of diversity since the Cretaceous.<sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup> An early study of the <a href="/wiki/RbcL" class="mw-redirect" title="RbcL">rbcL</a> gene sequence places <i>Wollemia</i> in the basal position of the Araucariaceae and as the sister group to <i>Agathis</i> and <i>Araucaria</i>.<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> In contrast, another study of the rbcL sequence shows that <i>Wollemia</i> is the sister group to <i>Agathis</i>, and <i>Araucaria</i> is basal.<sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup> The different outgroup selection and genes used in previous studies are the reasons behind the discrepancy over the groupings of the three genera.<sup id="cite_ref-Liu,_N._2009_37-0" class="reference"><a href="#cite_note-Liu,_N._2009-37"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> Later genetic studies corroborate <i>Wollemia</i>'s placement in the Araucariaceae as sister to <i>Agathis</i> based on data from the 28s rRNA gene,<sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> a combination of rbcL and <a href="/wiki/Maturase_K" title="Maturase K">matK</a> genes,<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> and a comprehensive study encompassing nuclear ribosomal 18S and <a href="/wiki/26S_rRNA" class="mw-redirect" title="26S rRNA">26S rRNA</a>, chloroplast 16S rRNA, rbcL, matK and rps4, and mitochondrial coxl and atp1 genes.<sup id="cite_ref-Liu,_N._2009_37-1" class="reference"><a href="#cite_note-Liu,_N._2009-37"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p><p>Below is the <a href="/wiki/Phylogeny" class="mw-redirect" title="Phylogeny">phylogeny</a> of the <a href="/wiki/Araucariaceae" title="Araucariaceae">Araucariaceae</a> based on the consensus from the most recent <a href="/wiki/Cladistics" title="Cladistics">cladistic analysis</a> of molecular data. It shows the relative positions of <i>Wollemia</i>, <i>Agathis</i>, and <i>Araucaria</i> within the division. </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="line-height:100%;"> <tbody><tr> <td class="clade-label first">  <i><a href="/wiki/Pinales" title="Pinales">Pinales</a></i>   </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p>  <i><a href="/wiki/Araucaria" title="Araucaria">Araucaria</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p>  <i>Wollemia</i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p>  <i><a href="/wiki/Agathis" title="Agathis">Agathis</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading3"><h3 id="Potential_fossil_record">Potential fossil record</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=12" title="Edit section: Potential fossil record"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>There are no fossils that can be assigned definitively to <i>Wollemia. <a href="/wiki/Araucarioides" title="Araucarioides">Araucarioides</a></i> leaves possibly representing <i>Wollemia</i> or a close relative have been reported from the Early <a href="/wiki/Eocene" title="Eocene">Eocene</a> of Tasmania. Some authors have suggested that the fossil pollen genus <i><a href="/w/index.php?title=Dilwynites&action=edit&redlink=1" class="new" title="Dilwynites (page does not exist)">Dilwynites</a></i>, known from the <a href="/wiki/Late_Cretaceous" title="Late Cretaceous">Late Cretaceous</a>-<a href="/wiki/Pliocene" title="Pliocene">Pliocene</a> of Australia, New Zealand, Patagonia and Antarctica is assignable to <i>Wollemia</i>, however, the pollen of <i>Wollemia</i> is highly variable, and its similarity to <i>Dilwynites</i> has been questioned, with <i>Dilwynites</i> also closely resembling the pollen of some species of <i>Agathis</i>. It is therefore possible that <i>Dilwynites</i> pollen represents that of other araucarian conifers as well as possibly also <i>Wollemia</i>.<sup id="cite_ref-Seyfullah-2023_6-1" class="reference"><a href="#cite_note-Seyfullah-2023-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=13" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><i><a href="/wiki/Gasteranthus_extinctus" title="Gasteranthus extinctus">Gasteranthus extinctus</a></i>, a species of plant believed to have gone extinct until it was rediscovered in 2022</li></ul> <div class="mw-heading mw-heading2"><h2 id="Notes">Notes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=14" title="Edit section: Notes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-lower-alpha"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-10">^</a></b></span> <span class="reference-text"> The discovery of the specimen's true identity, based upon knowledge of the literature on fossil remains, is similar to that of <a href="/wiki/Gerard_Krefft#The_Queensland_lungfish_(Neoceratodus_forsteri)" title="Gerard Krefft">Gerard Krefft recognising the Queensland lung fish in January 1870</a>.</span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=15" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>This article incorporates text from the <a href="/wiki/ARKive" title="ARKive">ARKive</a> fact-file "Wollemia" under the <a href="/wiki/Wikipedia:Text_of_Creative_Commons_Attribution-ShareAlike_3.0_Unported_License" class="mw-redirect" title="Wikipedia:Text of Creative Commons Attribution-ShareAlike 3.0 Unported License">Creative Commons Attribution-ShareAlike 3.0 Unported License</a> and the <a href="/wiki/Wikipedia:Text_of_the_GNU_Free_Documentation_License" title="Wikipedia:Text of the GNU Free Documentation License">GFDL</a>. </p> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239543626"><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-iucn_status_12_November_2021-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-iucn_status_12_November_2021_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-iucn_status_12_November_2021_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output 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Retrieved <span class="nowrap">2018-12-14</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=report&rft.btitle=Wollemia+nobilis+%28Wollemi+Pine%29+Recovery+Plan&rft.pub=New+South+Wales+Department+of+Environment+and+Conservation&rft_id=https%3A%2F%2Fwww.environment.nsw.gov.au%2F-%2Fmedia%2FOEH%2FCorporate-Site%2FDocuments%2FAnimals-and-plants%2FRecovery-plans%2Fwollemi-pine-wollemia-nobilis-recovery-plan-060519.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://pacifichorticulture.org/articles/humboldt-botanical-gardens/">"Pacific Horticulture | Humboldt Botanical Gardens"</a>. <i>Pacific Horticulture</i>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230713180324/https://pacifichorticulture.org/articles/humboldt-botanical-gardens/">Archived</a> from the original on 2023-07-13<span class="reference-accessdate">. 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Retrieved <span class="nowrap">23 January</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Conversation&rft.atitle=Wollemi+pines+are+dinosaur+trees&rft.date=2018-06-15&rft.aulast=Brack&rft.aufirst=Cris&rft_id=https%3A%2F%2Ftheconversation.com%2Fwollemi-pines-are-dinosaur-trees-98160&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-Seyfullah-2023-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-Seyfullah-2023_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Seyfullah-2023_6-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSeyfullahCoiroHofmann2023" class="citation journal cs1">Seyfullah, Leyla J.; Coiro, Mario; Hofmann, Christa-Ch. (February 2023). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.revpalbo.2022.104816">"In situ pollen diversity in the relict conifer Wollemia nobilis"</a>. <i>Review of Palaeobotany and Palynology</i>. <b>309</b>: 104816. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2023RPaPa.30904816S">2023RPaPa.30904816S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.revpalbo.2022.104816">10.1016/j.revpalbo.2022.104816</a></span>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:254332797">254332797</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Review+of+Palaeobotany+and+Palynology&rft.atitle=In+situ+pollen+diversity+in+the+relict+conifer+Wollemia+nobilis&rft.volume=309&rft.pages=104816&rft.date=2023-02&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A254332797%23id-name%3DS2CID&rft_id=info%3Adoi%2F10.1016%2Fj.revpalbo.2022.104816&rft_id=info%3Abibcode%2F2023RPaPa.30904816S&rft.aulast=Seyfullah&rft.aufirst=Leyla+J.&rft.au=Coiro%2C+Mario&rft.au=Hofmann%2C+Christa-Ch.&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252Fj.revpalbo.2022.104816&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-woodford2002-7"><span class="mw-cite-backlink">^ <a href="#cite_ref-woodford2002_7-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-woodford2002_7-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWoodford2002" class="citation book cs1">Woodford, James (2002). <i>The Wollemi Pine: The incredible discovery of a living fossil from the age of the dinosaurs</i>. 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(March 2014)">page needed</span></a></i>]</sup></span> </li> <li id="cite_note-FF-8"><span class="mw-cite-backlink">^ <a href="#cite_ref-FF_8-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-FF_8-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20041013034333/http://www.rbgsyd.gov.au/information_about_plants/wollemi_pine/facts_%26_figures">"wollemi pine/facts & figures"</a>. Royal Botanic Gardens, Sydney. April 2003. Archived from <a rel="nofollow" class="external text" href="http://www.rbgsyd.gov.au/information_about_plants/wollemi_pine/facts_%26_figures">the original</a> on 2004-10-13<span class="reference-accessdate">. Retrieved <span class="nowrap">2011-08-02</span></span> – via <a href="/wiki/ARKive" title="ARKive">ARKive</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=wollemi+pine%2Ffacts+%26+figures&rft.pub=Royal+Botanic+Gardens%2C+Sydney&rft.date=2003-04&rft_id=http%3A%2F%2Fwww.rbgsyd.gov.au%2Finformation_about_plants%2Fwollemi_pine%2Ffacts_%2526_figures&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-NPR-9"><span class="mw-cite-backlink">^ <a href="#cite_ref-NPR_9-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-NPR_9-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWamsley2020" class="citation news cs1">Wamsley, Laurel (16 January 2020). <a rel="nofollow" class="external text" href="https://www.npr.org/2020/01/16/796994699/aussie-firefighters-save-worlds-only-groves-of-prehistoric-wollemi-pines">"Aussie firefighters save world's only groves of prehistoric Wollemi pines"</a>. <a href="/wiki/NPR_News" class="mw-redirect" title="NPR News">NPR News</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230418170006/https://www.npr.org/2020/01/16/796994699/aussie-firefighters-save-worlds-only-groves-of-prehistoric-wollemi-pines">Archived</a> from the original on 2023-04-18<span class="reference-accessdate">. Retrieved <span class="nowrap">2020-01-17</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Aussie+firefighters+save+world%27s+only+groves+of+prehistoric+Wollemi+pines&rft.date=2020-01-16&rft.aulast=Wamsley&rft.aufirst=Laurel&rft_id=https%3A%2F%2Fwww.npr.org%2F2020%2F01%2F16%2F796994699%2Faussie-firefighters-save-worlds-only-groves-of-prehistoric-wollemi-pines&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20050323093506/http://www.rbgsyd.gov.au/information_about_plants/wollemi_pine">"The Wollemi Pine — a very rare discovery"</a>. <a href="/wiki/Royal_Botanic_Gardens,_Sydney" class="mw-redirect" title="Royal Botanic Gardens, Sydney">Royal Botanic Gardens, Sydney</a>. Archived from <a rel="nofollow" class="external text" href="http://www.rbgsyd.gov.au/information_about_plants/wollemi_pine">the original</a> on 2005-03-23<span class="reference-accessdate">. Retrieved <span class="nowrap">2007-02-08</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=The+Wollemi+Pine+%E2%80%94+a+very+rare+discovery&rft.pub=Royal+Botanic+Gardens%2C+Sydney&rft_id=http%3A%2F%2Fwww.rbgsyd.gov.au%2Finformation_about_plants%2Fwollemi_pine&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.anbg.gov.au/gnp/interns-2004/wollemia-nobilis.html">"<i>Wollemia nobilis</i>"</a>. Growing native plants. <i>www.anbg.gov.au</i>. <a href="/wiki/Australian_National_Botanic_Gardens" title="Australian National Botanic Gardens">Australian National Botanic Gardens</a>. <a href="/wiki/Parks_Australia" class="mw-redirect" title="Parks Australia">Parks Australia</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230713180403/https://www.anbg.gov.au/gnp/interns-2004/wollemia-nobilis.html">Archived</a> from the original on 2023-07-13<span class="reference-accessdate">. Retrieved <span class="nowrap">2022-09-17</span></span>. <q>Derivation of the name: <i>Wollemia nobilis</i> ... <i>nobilis</i> – The species epithet is a tribute to <a href="/wiki/David_Noble_(canyoner)" title="David Noble (canyoner)">David Noble</a> who discovered the first stand of Wollemi Pines on an abseiling trip in 1994.</q></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.anbg.gov.au&rft.atitle=Wollemia+nobilis&rft_id=https%3A%2F%2Fwww.anbg.gov.au%2Fgnp%2Finterns-2004%2Fwollemia-nobilis.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBarba2022" class="citation news cs1 cs1-prop-foreign-lang-source">Barba, Eduardo (17 September 2022). <a rel="nofollow" class="external text" href="https://elpais.com/estilo-de-vida/2022-09-17/david-noble-y-el-wollemia-nobilis-la-sorpresa-de-encontrar-un-arbol-desconocido-para-la-ciencia.html">"David Noble y <i>Wollemia nobilis</i>: la sorpresa de encontrar un árbol desconocido para la ciencia"</a>. <i><a href="/wiki/El_Pa%C3%ADs" title="El País">El País</a></i> (in Spanish). Madrid, ES. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230713180426/https://elpais.com/estilo-de-vida/2022-09-17/david-noble-y-el-wollemia-nobilis-la-sorpresa-de-encontrar-un-arbol-desconocido-para-la-ciencia.html">Archived</a> from the original on 2023-07-13<span class="reference-accessdate">. Retrieved <span class="nowrap">2022-09-17</span></span>. <q>En honor de <a href="/wiki/David_Noble_(canyoner)" title="David Noble (canyoner)">David Noble</a> se le dedicó el epíteto de la especie, nobilis.</q></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=El+Pa%C3%ADs&rft.atitle=David+Noble+y+Wollemia+nobilis%3A+la+sorpresa+de+encontrar+un+%C3%A1rbol+desconocido+para+la+ciencia&rft.date=2022-09-17&rft.aulast=Barba&rft.aufirst=Eduardo&rft_id=https%3A%2F%2Felpais.com%2Festilo-de-vida%2F2022-09-17%2Fdavid-noble-y-el-wollemia-nobilis-la-sorpresa-de-encontrar-un-arbol-desconocido-para-la-ciencia.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation news cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20171222104944/http://newengland.focusmag.com.au/david-mackay/">"David Mackay"</a>. <i><span style="font-size:85%;">New England</span> Focus</i>. 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Retrieved <span class="nowrap">2017-12-20</span></span> – via newengland.focusmag.com.au.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=%3Cspan+style%3D%22font-size%3A85%25%3B%22%3ENew+England%3C%2Fspan%3E+Focus&rft.atitle=David+Mackay&rft.date=2009-02-08&rft_id=http%3A%2F%2Fnewengland.focusmag.com.au%2Fdavid-mackay%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-15">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFStevensonRamakrishnanAlvesCoelho2023" class="citation report cs1">Stevenson, Dennis Wm.; Ramakrishnan, Srividya; Alves, Cristiane de Santis; Coelho, Laís Araujo; Kramer, Melissa; Goodwin, Sara; et al. 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Jager, Muriel; Deutsch, Jean; Broutin, Jean; Masselot, Monique (May 1998). <a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F2446539">"Phylogenetic relationships of conifers inferred from partial 28S rRNA gene sequences"</a>. <i>American Journal of Botany</i>. <b>85</b> (5): 688. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F2446539">10.2307/2446539</a></span>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/2446539">2446539</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21684951">21684951</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Journal+of+Botany&rft.atitle=Phylogenetic+relationships+of+conifers+inferred+from+partial+28S+rRNA+gene+sequences&rft.volume=85&rft.issue=5&rft.pages=688&rft.date=1998-05&rft_id=info%3Apmid%2F21684951&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F2446539%23id-name%3DJSTOR&rft_id=info%3Adoi%2F10.2307%2F2446539&rft.aulast=Stefanovi%C4%87&rft.aufirst=Sa%C5%A1a&rft.au=Jager%2C+Muriel&rft.au=Deutsch%2C+Jean&rft.au=Broutin%2C+Jean&rft.au=Masselot%2C+Monique&rft_id=https%3A%2F%2Fdoi.org%2F10.2307%252F2446539&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> <li id="cite_note-39"><span class="mw-cite-backlink"><b><a href="#cite_ref-39">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFQuinnPriceGadek2002" class="citation journal cs1">Quinn, C.J.; Price, R.A.; Gadek, P.A. 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"Familial concepts and relationships in the conifer, based on <i>rbcL</i> and <i>matK</i> sequence comparisons". <i>Kew Bulletin</i>. <b>57</b> (3): 513–531. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2002KewBu..57..513Q">2002KewBu..57..513Q</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2307%2F4110984">10.2307/4110984</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/4110984">4110984</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:83816639">83816639</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Kew+Bulletin&rft.atitle=Familial+concepts+and+relationships+in+the+conifer%2C+based+on+rbcL+and+matK+sequence+comparisons&rft.volume=57&rft.issue=3&rft.pages=513-531&rft.date=2002&rft_id=info%3Adoi%2F10.2307%2F4110984&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A83816639%23id-name%3DS2CID&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F4110984%23id-name%3DJSTOR&rft_id=info%3Abibcode%2F2002KewBu..57..513Q&rft.aulast=Quinn&rft.aufirst=C.J.&rft.au=Price%2C+R.A.&rft.au=Gadek%2C+P.A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Wollemia&action=edit&section=16" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; font-style: italic;"><a href="https://commons.wikimedia.org/wiki/Category:Wollemia" class="extiw" title="commons:Category:Wollemia">Wollemia</a></span>.</div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1237033735"><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/34px-Wikispecies-logo.svg.png" decoding="async" width="34" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/51px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/68px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></span></span></div> <div class="side-box-text plainlist"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a> has information related to <i><b><a href="https://species.wikimedia.org/wiki/Special:Search/Wollemia" class="extiw" title="wikispecies:Special:Search/Wollemia">Wollemia</a></b></i>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="https://web.archive.org/web/*/www.arkive.org/"><i>Wollemia nobilis</i></a> media from <a href="/wiki/ARKive" title="ARKive">ARKive</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q12981878#P2833" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFThomas,_P.2011" class="citation journal cs1">Thomas, P. (2011). <a rel="nofollow" class="external text" href="https://www.iucnredlist.org/species/34926/9898196">"<i>Wollemia nobilis</i>"</a>. <i><a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN Red List of Threatened Species</a></i>. <b>2011</b>: e.T34926A9898196. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.2305%2FIUCN.UK.2011-2.RLTS.T34926A9898196.en">10.2305/IUCN.UK.2011-2.RLTS.T34926A9898196.en</a></span><span class="reference-accessdate">. Retrieved <span class="nowrap">12 November</span> 2021</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=IUCN+Red+List+of+Threatened+Species&rft.atitle=Wollemia+nobilis&rft.volume=2011&rft.pages=e.T34926A9898196&rft.date=2011&rft_id=info%3Adoi%2F10.2305%2FIUCN.UK.2011-2.RLTS.T34926A9898196.en&rft.au=Thomas%2C+P.&rft_id=https%3A%2F%2Fwww.iucnredlist.org%2Fspecies%2F34926%2F9898196&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span> Listed as Critically Endangered (CR D v2.3)</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20050323093506/http://www.rbgsyd.gov.au/information_about_plants/wollemi_pine">"The Wollemi Pine – a very rare discovery"</a>. <a href="/wiki/Royal_Botanic_Gardens,_Sydney" class="mw-redirect" title="Royal Botanic Gardens, Sydney">Royal Botanic Gardens, Sydney</a>. Archived from <a rel="nofollow" class="external text" href="http://www.rbgsyd.gov.au/information_about_plants/wollemi_pine">the original</a> on 2005-03-23<span class="reference-accessdate">. Retrieved <span class="nowrap">2007-02-08</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=The+Wollemi+Pine+%E2%80%93+a+very+rare+discovery&rft.pub=Royal+Botanic+Gardens%2C+Sydney&rft_id=http%3A%2F%2Fwww.rbgsyd.gov.au%2Finformation_about_plants%2Fwollemi_pine&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span> (includes facts and figures, ecology, biology)</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20050525063832/http://www.rbgkew.org.uk/plants/wollemipine.html">Royal Botanic Gardens at Kew's web page about the "Wollemi pine"</a></li> <li><a rel="nofollow" class="external text" href="http://wollemipine.com/">WollemiPine.com</a></li> <li><a rel="nofollow" class="external text" href="http://www.conifers.org/ar/wo/index.html">Wollemia nobilis</a> at the Gymnosperm Database</li> <li><a rel="nofollow" class="external text" href="http://news.bbc.co.uk/1/hi/england/london/4531805.stm">BBC News item</a> 10 May 2005</li> <li><a rel="nofollow" class="external text" href="http://news.bbc.co.uk/1/hi/england/cornwall/6201616.stm">BBC News – 'Dinosaur trees' heavily guarded</a> – 02/12/06</li> <li><a rel="nofollow" class="external text" href="http://www.abc.net.au/gardening/stories/s1471653.htm">ABC-TV Gardening Fact Sheet</a></li> <li><a rel="nofollow" class="external text" href="http://www.abc.net.au/catalyst/stories/s1371627.htm">ABC-TV Science visits Wollemi Pines in the wild</a> 19 May 2005</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20070905075700/http://www.nationalparks.nsw.gov.au/PDFs/_Recovery_Plan_Wollemi_Pine.pdf">Wollemia nobilis (Wollemi Pine) Recovery Plan</a>, January 2007</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWarren2007" class="citation news cs1">Warren, Matthew (16 April 2007). <a rel="nofollow" class="external text" href="http://www.theaustralian.com.au/news/nation/biologist-takes-axe-to-the-myth-of-wollemi/story-e6frg6nf-1111113340564">"Biologist takes axe to the 'myth' of Wollemi"</a>. <i>The Australian</i><span class="reference-accessdate">. Retrieved <span class="nowrap">March 4,</span> 2014</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Australian&rft.atitle=Biologist+takes+axe+to+the+%27myth%27+of+Wollemi&rft.date=2007-04-16&rft.aulast=Warren&rft.aufirst=Matthew&rft_id=http%3A%2F%2Fwww.theaustralian.com.au%2Fnews%2Fnation%2Fbiologist-takes-axe-to-the-myth-of-wollemi%2Fstory-e6frg6nf-1111113340564&rfr_id=info%3Asid%2Fen.wikipedia.org%3AWollemia" class="Z3988"></span></li> <li><a rel="nofollow" class="external text" href="http://www.abc.net.au/rn/scienceshow/stories/2007/1902727.htm">The Wollemi Pine</a> Transcript of interview on <i>The Science Show</i> (April 2007) with <a href="/wiki/Timothy_John_Entwisle" title="Timothy John Entwisle">Tim Entwisle</a>, then director of the Royal Botanic Gardens in Sydney.</li> <li><a rel="nofollow" class="external text" href="http://www.monumentaltrees.com/en/content/westonbirt/#wollemia">Images and information about the Wollemi Pine in Westonbirt Arboretum</a></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20121023110403/http://press.nationalgeographic.com/2006/09/19/presentsamajorbotanicalfindofthecenturylivingfossilwollemipineavailableforthefirsttimeinnorthamerica/">Wollemi Pine available for first time in North America</a> from National Geographic.</li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol 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.navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Classification_of_Acrogymnospermae_(living_Gymnosperms)" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background: #C3EEC3;;background: #90EE90;"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" 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href="/wiki/Template_talk:Acrogymnospermae_classification" title="Template talk:Acrogymnospermae classification"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Acrogymnospermae_classification" title="Special:EditPage/Template:Acrogymnospermae classification"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Classification_of_Acrogymnospermae_(living_Gymnosperms)" style="font-size:114%;margin:0 4em">Classification of <a href="/wiki/Acrogymnospermae" class="mw-redirect" title="Acrogymnospermae">Acrogymnospermae</a> (living <a href="/wiki/Gymnosperm" title="Gymnosperm">Gymnosperms</a>)</div></th></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: #C3EEC3;"><div> <ul><li>Supergroup <a href="/wiki/Template:Plant_classification" title="Template:Plant classification">Plant</a>:</li> <li><a href="/wiki/Template:Bryophyta" title="Template:Bryophyta">Bryophyta</a></li> <li><a href="/wiki/Template:Marchantiophyta" title="Template:Marchantiophyta">Marchantiophyta</a></li> <li><a href="/wiki/Template:Fern_classification" title="Template:Fern classification">Polypodiophyta</a></li> <li><a href="/wiki/Template:Acrogymnospermae_classification" title="Template:Acrogymnospermae classification">Acrogymnospermae</a></li> <li><a href="/wiki/Template:Angiosperm_families" title="Template:Angiosperm families">Angiospermae</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.0em; background: #D4EED4;"><a href="/wiki/Ginkgoidae" title="Ginkgoidae">Ginkgoidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ginkgoales" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ginkgoales" title="Ginkgoales">Ginkgoales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ginkgoaceae" title="Ginkgoaceae">Ginkgoaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ginkgo" title="Ginkgo">Ginkgo</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.0em; background: #D4EED4;"><a href="/wiki/Cycadidae" title="Cycadidae">Cycadidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cycadales" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cycadales" class="mw-redirect" title="Cycadales">Cycadales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cycadaceae" class="mw-redirect" title="Cycadaceae">Cycadaceae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cycas" title="Cycas">Cycas</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Zamiaceae" title="Zamiaceae">Zamiaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Diooideae</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dioon" title="Dioon">Dioon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Zamioideae</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Macrozamia" title="Macrozamia">Macrozamia</a></i></li> <li><i><a href="/wiki/Lepidozamia" title="Lepidozamia">Lepidozamia</a></i></li> <li><i><a href="/wiki/Encephalartos" title="Encephalartos">Encephalartos</a></i></li> <li><i><a href="/wiki/Bowenia" title="Bowenia">Bowenia</a></i></li> <li><i><a href="/wiki/Ceratozamia" title="Ceratozamia">Ceratozamia</a></i></li> <li><i><a href="/wiki/Stangeria" title="Stangeria">Stangeria</a></i></li> <li><i><a href="/wiki/Zamia" title="Zamia">Zamia</a></i></li> <li><i><a href="/wiki/Microcycas" title="Microcycas">Microcycas</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1.0em; background: #D4EED4;"><a href="/wiki/Pinidae" title="Pinidae">Pinidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Gnetales" class="mw-redirect" title="Gnetales">Gnetales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ephedraceae" title="Ephedraceae">Ephedraceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ephedra_(plant)" title="Ephedra (plant)">Ephedra</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Gnetaceae" class="mw-redirect" title="Gnetaceae">Gnetaceae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Gnetum" title="Gnetum">Gnetum</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Welwitschiaceae" title="Welwitschiaceae">Welwitschiaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Welwitschia" title="Welwitschia">Welwitschia</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pinales" title="Pinales">Pinales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pinaceae" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pinaceae" title="Pinaceae">Pinaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Abietoideae</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cedrus" title="Cedrus">Cedrus</a></i></li> <li><i><a href="/wiki/Pseudolarix" title="Pseudolarix">Pseudolarix</a></i></li> <li><i><a href="/wiki/Nothotsuga" title="Nothotsuga">Nothotsuga</a></i></li> <li><i><a href="/wiki/Tsuga" title="Tsuga">Tsuga</a></i></li> <li><i><a href="/wiki/Keteleeria" title="Keteleeria">Keteleeria</a></i></li> <li><i><a href="/wiki/Abies" class="mw-redirect" title="Abies">Abies</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Pinoideae</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Pseudotsuga" title="Pseudotsuga">Pseudotsuga</a></i></li> <li><i><a href="/wiki/Larix" class="mw-redirect" title="Larix">Larix</a></i></li> <li><i><a href="/wiki/Cathaya" title="Cathaya">Cathaya</a></i></li> <li><i><a href="/wiki/Picea" class="mw-redirect" title="Picea">Picea</a></i></li> <li><i><a href="/wiki/Pinus" class="mw-redirect" title="Pinus">Pinus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Araucariales" class="mw-redirect" title="Araucariales">Araucariales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Araucariaceae" title="Araucariaceae">Araucariaceae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a class="mw-selflink selflink">Wollemia</a></i></li> <li><i><a href="/wiki/Agathis" title="Agathis">Agathis</a></i></li> <li><i><a href="/wiki/Araucaria" title="Araucaria">Araucaria</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Podocarpaceae" title="Podocarpaceae">Podocarpaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Phyllocladoideae</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Halocarpus" title="Halocarpus">Halocarpus</a></i></li> <li><i><a href="/wiki/Pectinopitys" title="Pectinopitys">Pectinopitys</a></i></li> <li><i><a href="/wiki/Sundacarpus" title="Sundacarpus">Sundacarpus</a></i></li> <li><i><a href="/wiki/Prumnopitys" title="Prumnopitys">Prumnopitys</a></i></li> <li><i><a href="/wiki/Parasitaxus" title="Parasitaxus">Parasitaxus</a></i></li> <li><i><a href="/wiki/Lagarostrobos" title="Lagarostrobos">Lagarostrobos</a></i></li> <li><i><a href="/wiki/Manoao" title="Manoao">Manoao</a></i></li> <li><i><a href="/wiki/Lepidothamnus" title="Lepidothamnus">Lepidothamnus</a></i></li> <li><i><a href="/wiki/Phyllocladus" title="Phyllocladus">Phyllocladus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Podocarpoideae</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Saxegothaea" title="Saxegothaea">Saxegothaea</a></i></li> <li><i><a href="/wiki/Microcachrys" title="Microcachrys">Microcachrys</a></i></li> <li><i><a href="/wiki/Pherosphaera" title="Pherosphaera">Pherosphaera</a></i></li> <li><i><a href="/wiki/Acmopyle" title="Acmopyle">Acmopyle</a></i></li> <li><i><a href="/wiki/Dacrycarpus" title="Dacrycarpus">Dacrycarpus</a></i></li> <li><i><a href="/wiki/Falcatifolium" title="Falcatifolium">Falcatifolium</a></i></li> <li><i><a href="/wiki/Dacrydium" title="Dacrydium">Dacrydium</a></i></li> <li><i><a href="/wiki/Retrophyllum" title="Retrophyllum">Retrophyllum</a></i></li> <li><i><a href="/wiki/Afrocarpus" title="Afrocarpus">Afrocarpus</a></i></li> <li><i><a href="/wiki/Nageia" title="Nageia">Nageia</a></i></li> <li><i><a href="/wiki/Podocarpus" title="Podocarpus">Podocarpus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cupressales" class="mw-redirect" title="Cupressales">Cupressales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sciadopityaceae" title="Sciadopityaceae">Sciadopityaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Sciadopitys" title="Sciadopitys">Sciadopitys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Taxaceae" title="Taxaceae">Taxaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Cephalotaxeae</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cephalotaxus" title="Cephalotaxus">Cephalotaxus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Taxoideae</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amentotaxus" title="Amentotaxus">Amentotaxus</a></i></li> <li><i><a href="/wiki/Torreya" title="Torreya">Torreya</a></i></li> <li><i><a href="/wiki/Austrotaxus" title="Austrotaxus">Austrotaxus</a></i></li> <li><i><a href="/wiki/Pseudotaxus" title="Pseudotaxus">Pseudotaxus</a></i></li> <li><i><a href="/wiki/Taxus" title="Taxus">Taxus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cupressaceae" title="Cupressaceae">Cupressaceae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cunninghamioideae" class="mw-redirect" title="Cunninghamioideae">Cunninghamioideae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cunninghamia" title="Cunninghamia">Cunninghamia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Taiwanioideae" class="mw-redirect" title="Taiwanioideae">Taiwanioideae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Taiwania" title="Taiwania">Taiwania</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Athrotaxidoideae" class="mw-redirect" title="Athrotaxidoideae">Athrotaxidoideae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Athrotaxis" title="Athrotaxis">Athrotaxis</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sequoioideae" title="Sequoioideae">Sequoioideae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Metasequoia" title="Metasequoia">Metasequoia</a></i></li> <li><i><a href="/wiki/Sequoiadendron" title="Sequoiadendron">Sequoiadendron</a></i></li> <li><i><a href="/wiki/Sequoia_(genus)" title="Sequoia (genus)">Sequoia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Taxodioideae" title="Taxodioideae">Taxodioideae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cryptomeria" title="Cryptomeria">Cryptomeria</a></i></li> <li><i><a href="/wiki/Glyptostrobus" title="Glyptostrobus">Glyptostrobus</a></i></li> <li><i><a href="/wiki/Taxodium" title="Taxodium">Taxodium</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Callitroideae" class="mw-redirect" title="Callitroideae">Callitroideae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Papuacedrus" title="Papuacedrus">Papuacedrus</a></i></li> <li><i><a href="/wiki/Austrocedrus" title="Austrocedrus">Austrocedrus</a></i></li> <li><i><a href="/wiki/Pilgerodendron" title="Pilgerodendron">Pilgerodendron</a></i></li> <li><i><a href="/wiki/Libocedrus" title="Libocedrus">Libocedrus</a></i></li> <li><i><a href="/wiki/Diselma" title="Diselma">Diselma</a></i></li> <li><i><a href="/wiki/Fitzroya" title="Fitzroya">Fitzroya</a></i></li> <li><i><a href="/wiki/Widdringtonia" title="Widdringtonia">Widdringtonia</a></i></li> <li><i><a href="/wiki/Neocallitropsis" class="mw-redirect" title="Neocallitropsis">Neocallitropsis</a></i></li> <li><i><a href="/wiki/Callitris" title="Callitris">Callitris</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cupressoideae" class="mw-redirect" title="Cupressoideae">Cupressoideae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Thujopsis" title="Thujopsis">Thujopsis</a></i></li> <li><i><a href="/wiki/Thuja" title="Thuja">Thuja</a></i></li> <li><i><a href="/wiki/Chamaecyparis" title="Chamaecyparis">Chamaecyparis</a></i></li> <li><i><a href="/wiki/Tetraclinis" title="Tetraclinis">Tetraclinis</a></i></li> <li><i><a href="/wiki/Microbiota" title="Microbiota">Microbiota</a></i></li> <li><i><a href="/wiki/Platycladus" title="Platycladus">Platycladus</a></i></li> <li><i><a href="/wiki/Calocedrus" title="Calocedrus">Calocedrus</a></i></li> <li><i><a href="/wiki/Cupressus" title="Cupressus">Cupressus</a></i></li> <li><i><a href="/wiki/Xanthocyparis" title="Xanthocyparis">Xanthocyparis</a></i></li> <li><i><a href="/wiki/Hesperocyparis" title="Hesperocyparis">Hesperocyparis</a></i></li> <li><i><a href="/wiki/Juniperus" class="mw-redirect" title="Juniperus">Juniperus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Wollemia</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q12981878" class="extiw" title="wikidata:Q12981878">Q12981878</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Wollemia" class="extiw" title="wikispecies:Wollemia">Wollemia</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Plant_Name_Index" title="Australian Plant Name Index">APNI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://id.biodiversity.org.au/name/apni/154750">154750</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/886H">886H</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/14999">14999</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/EPPO_Code" title="EPPO Code">EPPO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://gd.eppo.int/taxon/1WLMG">1WLMG</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Flora_of_Australia" title="Flora of Australia">FoAO2</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://profiles.ala.org.au/opus/foa/profile/Wollemia">Wollemia</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/2685084">2685084</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Germplasm_Resources_Information_Network" title="Germplasm Resources Information Network">GRIN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://npgsweb.ars-grin.gov/gringlobal/taxonomygenus.aspx?id=17622">17622</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/49383">49383</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/International_Plant_Names_Index" title="International Plant Names Index">IPNI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ipni.org/n/986235-1">986235-1</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=1298175">1298175</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Biodiversity_Network" title="National Biodiversity Network">NBN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://data.nbn.org.uk/Taxa/NHMSYS0021196592">NHMSYS0021196592</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=56997">56997</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=1020082">1020082</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Plants_of_the_World_Online" title="Plants of the World Online">POWO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://powo.science.kew.org/taxon/urn%3Alsid%3Aipni.org%3Anames%3A986235-1">urn:lsid:ipni.org:names:986235-1</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Tropicos" title="Tropicos">Tropicos</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://legacy.tropicos.org/Name/50091211">50091211</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Flora_Online" title="World Flora Online">WFO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://list.worldfloraonline.org/wfo-4000040804">wfo-4000040804</a></span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Wollemia nobilis</i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q190510" class="extiw" title="wikidata:Q190510">Q190510</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Wollemia_nobilis" class="extiw" title="wikispecies:Wollemia nobilis">Wollemia nobilis</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Plant_Name_Index" title="Australian Plant Name Index">APNI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://id.biodiversity.org.au/name/apni/154751">154751</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/ARKive" title="ARKive">ARKive</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://web.archive.org/web/*/http://www.arkive.org/wd/wollemia-nobilis/">wollemia-nobilis</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/5C32P">5C32P</a></span></span></li> <li><span style="white-space:nowrap;">Conifers.org: <span class="uid"><a rel="nofollow" class="external text" href="https://conifers.org/ar/Wollemia.php">Wollemia</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/128312">128312</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/EPPO_Code" title="EPPO Code">EPPO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://gd.eppo.int/taxon/WLMNO">WLMNO</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Flora_of_Australia" title="Flora of Australia">FoAO2</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://profiles.ala.org.au/opus/foa/profile/Wollemia%20nobilis">Wollemia nobilis</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/2685085">2685085</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Germplasm_Resources_Information_Network" title="Germplasm Resources Information Network">GRIN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://npgsweb.ars-grin.gov/gringlobal/taxonomydetail.aspx?id=416628">416628</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/49381">49381</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/International_Plant_Names_Index" title="International Plant Names Index">IPNI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ipni.org/n/986236-1">986236-1</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=11442459">11442459</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Invasive_Species_Compendium" title="Invasive Species Compendium">ISC</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.cabi.org/isc/datasheet/56830">56830</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://apiv3.iucnredlist.org/api/v3/taxonredirect/34926">34926</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Missouri_Botanical_Garden" title="Missouri Botanical Garden">MoBotPF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.missouribotanicalgarden.org/PlantFinder/PlantFinderDetails.aspx?taxonid=291688">291688</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Biodiversity_Network" title="National Biodiversity Network">NBN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://data.nbn.org.uk/Taxa/NHMSYS0021196598">NHMSYS0021196598</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=56998">56998</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Observation.org" title="Observation.org">Observation.org</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://observation.org/species/438571/">438571</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=625420">625420</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/The_Plant_List" title="The Plant List">Plant List</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.theplantlist.org/tpl1.1/record/kew-214863">kew-214863</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Plants_of_the_World_Online" title="Plants of the World Online">POWO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://powo.science.kew.org/taxon/urn%3Alsid%3Aipni.org%3Anames%3A986236-1">urn:lsid:ipni.org:names:986236-1</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Royal_Horticultural_Society" title="Royal Horticultural Society">RHS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.rhs.org.uk/Plants/134274/wd/Details">134274</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Species_Profile_and_Threats_Database" class="mw-redirect" title="Species Profile and Threats Database">SPRAT</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.environment.gov.au/cgi-bin/sprat/public/publicspecies.pl?taxon_id=64545">64545</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Tropicos" title="Tropicos">Tropicos</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://legacy.tropicos.org/Name/50091212">50091212</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Flora_Online" title="World Flora Online">WFO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://list.worldfloraonline.org/wfo-0000334493">wfo-0000334493</a></span></span></li></ul> </div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Wollemia&oldid=1258970270">https://en.wikipedia.org/w/index.php?title=Wollemia&oldid=1258970270</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:IUCN_Red_List_critically_endangered_species" title="Category:IUCN Red List critically endangered species">IUCN Red List critically endangered species</a></li><li><a href="/wiki/Category:Araucariaceae" title="Category:Araucariaceae">Araucariaceae</a></li><li><a href="/wiki/Category:Monotypic_conifer_genera" title="Category:Monotypic conifer genera">Monotypic conifer genera</a></li><li><a href="/wiki/Category:Critically_endangered_flora_of_Australia" title="Category:Critically endangered flora of Australia">Critically endangered flora of Australia</a></li><li><a href="/wiki/Category:Flora_of_New_South_Wales" 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