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.mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">David Botstein</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Plos_botstein.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Plos_botstein.jpg/220px-Plos_botstein.jpg" decoding="async" width="220" height="192" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Plos_botstein.jpg/330px-Plos_botstein.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Plos_botstein.jpg/440px-Plos_botstein.jpg 2x" data-file-width="1240" data-file-height="1083" /></a></span></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data">September 8, 1942<span style="display:none"> (<span class="dtstart bday">1942-09-08</span>)</span> (age <span class="currentage"></span>82)<br /><div style="display:inline" class="birthplace"><a href="/wiki/Zurich" title="Zurich">Zurich</a>, <a href="/wiki/Switzerland" title="Switzerland">Switzerland</a></div></td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a><br /><a href="/wiki/University_of_Michigan" title="University of Michigan">University of Michigan</a></td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data">Genetic linkage map using restriction fragment length polymorphisms</td></tr><tr><th scope="row" class="infobox-label">Relatives</th><td class="infobox-data"><a href="/wiki/Leon_Botstein" title="Leon Botstein">Leon Botstein</a> (brother)</td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><a href="/wiki/Eli_Lilly_and_Company-Elanco_Research_Award" title="Eli Lilly and Company-Elanco Research Award">Eli Lilly and Company Award in Microbiology</a> (1978)<br /><a href="/wiki/Genetics_Society_of_America_Medal" title="Genetics Society of America Medal">Genetics Society of America Medal</a> (1988)<sup id="cite_ref-genemedal_1-0" class="reference"><a href="#cite_note-genemedal-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><br /><a href="/wiki/William_Allan_Award" title="William Allan Award">Allan Award</a> of the <a href="/wiki/American_Society_of_Human_Genetics" title="American Society of Human Genetics">American Society of Human Genetics</a> (1989)<br /><a href="/wiki/Rosenstiel_Award" title="Rosenstiel Award">Rosenstiel Award</a> (1991)<br /><a href="/wiki/Novartis-Drew_Award" title="Novartis-Drew Award">Novartis-Drew Award</a> (2003)<br /><a href="/wiki/Gruber_Prize_in_Genetics" title="Gruber Prize in Genetics">Gruber Prize in Genetics</a> (2003)<br /><a href="/wiki/Albany_Medical_Center_Prize" title="Albany Medical Center Prize">Albany Medical Center Prize</a> (2010) <br /><a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Breakthrough Prize in Life Sciences</a> (2013) <br /><a href="/wiki/Warren_Alpert_Foundation_Prize" title="Warren Alpert Foundation Prize">Warren Alpert Foundation Prize</a> (2013)<br /><a href="/wiki/Double_Helix_Medal" title="Double Helix Medal">Double Helix Medal</a> (2015)<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><br /><a href="/wiki/Thomas_Hunt_Morgan_Medal" title="Thomas Hunt Morgan Medal">Thomas Hunt Morgan Medal</a> (2020)</td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Genetics" title="Genetics">Genetics</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/MIT" class="mw-redirect" title="MIT">MIT</a><br /><a href="/wiki/Stanford_University" title="Stanford University">Stanford University</a><br /><a href="/wiki/Genentech" title="Genentech">Genentech</a><br /><a href="/wiki/Princeton_University" title="Princeton University">Princeton University</a></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Thesis" title="Thesis">Thesis</a></th><td class="infobox-data"><i><a rel="nofollow" class="external text" href="http://search.proquest.com/docview/302261666">The Synthesis and Maturation of Phage-P22 DNA</a></i> <span style="font-size:97%;">(1967)</span></td></tr><tr><th scope="row" class="infobox-label">Doctoral students</th><td class="infobox-data"><a href="/wiki/Olga_Troyanskaya" title="Olga Troyanskaya">Olga Troyanskaya</a><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> <br /> <a href="/wiki/Fred_Winston" title="Fred Winston">Fred Winston</a><br /><a href="/wiki/Douglas_Koshland" title="Douglas Koshland">Douglas Koshland</a><br /><a href="/wiki/Tim_Stearns" title="Tim Stearns">Tim Stearns</a></td></tr><tr><th scope="row" class="infobox-label">Other notable students</th><td class="infobox-data"><a href="/wiki/Michael_Eisen" title="Michael Eisen">Michael Eisen</a> (postdoc)</td></tr><tr style="display:none"><td colspan="2"> </td></tr><tr><th scope="row" class="infobox-label">Website</th><td class="infobox-data"><span class="url"><a rel="nofollow" class="external text" href="http://www.princeton.edu/genomics/botstein">www<wbr />.princeton<wbr />.edu<wbr />/genomics<wbr />/botstein</a></span><br /><span class="url"><a rel="nofollow" class="external text" href="http://molbio.princeton.edu/faculty/molbio-faculty/96-botstein">molbio<wbr />.princeton<wbr />.edu<wbr />/faculty<wbr />/molbio-faculty<wbr />/96-botstein</a></span></td></tr></tbody></table> <p><b>David Botstein</b> (born September 8, 1942) is an American biologist who is the chief scientific officer of <a href="/wiki/Calico_(company)" title="Calico (company)">Calico</a>. He was the director of the Lewis-Sigler Institute for Integrative Genomics at <a href="/wiki/Princeton_University" title="Princeton University">Princeton University</a><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> from 2003 to 2013, where he remains an Anthony B. Evnin Professor of Genomics. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Education">Education</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=David_Botstein&action=edit&section=1" title="Edit section: Education"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Botstein graduated from the <a href="/wiki/Bronx_High_School_of_Science" title="Bronx High School of Science">Bronx High School of Science</a> in 1959, and <a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a> in 1963. He started his Ph.D. work under <a href="/wiki/Maurice_Sanford_Fox" title="Maurice Sanford Fox">Maurice Sanford Fox</a> at the <a href="/wiki/Massachusetts_Institute_of_Technology" title="Massachusetts Institute of Technology">Massachusetts Institute of Technology</a>, then moved and received a <a href="/wiki/Ph.D." class="mw-redirect" title="Ph.D.">Ph.D.</a> from the <a href="/wiki/University_of_Michigan" title="University of Michigan">University of Michigan</a> in 1967 for work on <a href="/wiki/P22_phage" class="mw-redirect" title="P22 phage">P22 phage</a>.<sup id="cite_ref-botstein_8-0" class="reference"><a href="#cite_note-botstein-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Career">Career</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=David_Botstein&action=edit&section=2" title="Edit section: Career"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Botstein taught at the <a href="/wiki/Massachusetts_Institute_of_Technology" title="Massachusetts Institute of Technology">Massachusetts Institute of Technology</a>, where he became a professor of genetics. Botstein joined <a href="/wiki/Genentech" title="Genentech">Genentech</a>, Inc. in 1987 as vice president – science. In 1990, he became chairman of the Department of Genetics at <a href="/wiki/Stanford_University" title="Stanford University">Stanford University</a>. Botstein was elected to the <a href="/wiki/U.S._National_Academy_of_Sciences" class="mw-redirect" title="U.S. National Academy of Sciences">U.S. National Academy of Sciences</a> in 1981 and to the <a href="/wiki/Institute_of_Medicine" class="mw-redirect" title="Institute of Medicine">Institute of Medicine</a> in 1993. </p><p>Botstein is the director of the Integrated Science Program at <a href="/wiki/Princeton_University" title="Princeton University">Princeton University</a>.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p><p>In 1980, Botstein and his colleagues Ray White, Mark Skolnick, and <a href="/wiki/Ronald_W._Davis" title="Ronald W. Davis">Ronald W. Davis</a> proposed a method<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> for constructing a <a href="/wiki/Genetic_linkage" title="Genetic linkage">genetic linkage</a> map using <a href="/wiki/Restriction_fragment_length_polymorphism" title="Restriction fragment length polymorphism">restriction fragment length polymorphisms</a> that was used in subsequent years to identify several human disease genes including <a href="/wiki/Huntington%27s_disease" title="Huntington's disease">Huntington's</a> and <a href="/wiki/BRCA1" title="BRCA1">BRCA1</a>. Variations of this method were used in the mapping efforts that predated and enabled the sequencing phase of the <a href="/wiki/Human_Genome_Project" title="Human Genome Project">Human Genome Project</a>. </p><p>In 1998, Botstein and his postdoctoral fellow <a href="/wiki/Michael_Eisen" title="Michael Eisen">Michael Eisen</a>, together with graduate student Paul Spellman and colleague <a href="/wiki/Patrick_O._Brown" title="Patrick O. Brown">Patrick Brown</a>, developed a statistical method and graphical interface that is widely used to interpret genomic data including <a href="/wiki/Microarray" title="Microarray">microarray</a> data.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> This approach was refined and applied for diverse applications, including for a molecular classification of heterogenous tumors using gene expression. These efforts included work on discovery of tumor subtypes with <a href="/w/index.php?title=Lou_Staudt&action=edit&redlink=1" class="new" title="Lou Staudt (page does not exist)">Lou Staudt</a>, <a href="/w/index.php?title=Ash_Alizadeh&action=edit&redlink=1" class="new" title="Ash Alizadeh (page does not exist)">Ash Alizadeh</a> and <a href="/wiki/Ronald_Levy_(scientist)" title="Ronald Levy (scientist)">Ronald Levy</a>, yielding a refined classification of <a href="/wiki/Diffuse_large_B-cell_lymphoma" title="Diffuse large B-cell lymphoma">diffuse large B cell lymphomas</a>, and in painting the molecular portraits for refined classification of <a href="/wiki/Breast_cancer_classification" title="Breast cancer classification">breast cancers</a> with <a href="/wiki/Anne-Lise_B%C3%B8rresen-Dale" title="Anne-Lise Børresen-Dale">Anne-Lise Børresen-Dale</a> and Charles Perou. He has subsequently worked on the creation of the influential <a href="/wiki/Gene_Ontology" title="Gene Ontology">Gene Ontology</a><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> with <a href="/wiki/Michael_Ashburner" title="Michael Ashburner">Michael Ashburner</a> and <a href="/wiki/Suzanna_Lewis" title="Suzanna Lewis">Suzanna Lewis</a>. He is one of the founding editors of the journal <i><a href="/wiki/Molecular_Biology_of_the_Cell" title="Molecular Biology of the Cell">Molecular Biology of the Cell</a></i>, along with <a href="/wiki/Erkki_Ruoslahti" title="Erkki Ruoslahti">Erkki Ruoslahti</a> and <a href="/wiki/Keith_Yamamoto" title="Keith Yamamoto">Keith Yamamoto</a>.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>In 2013, Botstein was named chief scientific officer of <a href="/wiki/Google" title="Google">Google</a>'s <a href="/wiki/Anti-aging" class="mw-redirect" title="Anti-aging">anti-aging</a> health startup <a href="/wiki/Calico_(company)" title="Calico (company)">Calico</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Awards">Awards</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=David_Botstein&action=edit&section=3" title="Edit section: Awards"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Botstein has won the <a href="/wiki/Eli_Lilly_and_Company-Elanco_Research_Award" title="Eli Lilly and Company-Elanco Research Award">Eli Lilly and Company Award in Microbiology</a> (1978), the Genetics Society of America Medal (1988, with <a href="/wiki/Ira_Herskowitz" title="Ira Herskowitz">Ira Herskowitz</a>),<sup id="cite_ref-genemedal_1-1" class="reference"><a href="#cite_note-genemedal-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> the <a href="/wiki/William_Allan_Award" title="William Allan Award">Allan Award</a> of the <a href="/wiki/American_Society_of_Human_Genetics" title="American Society of Human Genetics">American Society of Human Genetics</a> (1989, with <a href="/wiki/Raymond_L._White" title="Raymond L. White">Ray White</a>), the <a href="/wiki/Gruber_Prize_in_Genetics" title="Gruber Prize in Genetics">Gruber Prize in Genetics</a> (2003), the <a href="/wiki/Albany_Medical_Center_Prize" title="Albany Medical Center Prize">Albany Medical Center Prize</a> (2010, with <a href="/wiki/Eric_Lander" title="Eric Lander">Eric Lander</a> and <a href="/wiki/Francis_Collins" title="Francis Collins">Francis Collins</a>) and the <a href="/wiki/Dan_David_Prize" title="Dan David Prize">Dan David Prize</a> in 2012. In 2013 he was awarded the $3 million <a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Breakthrough Prize in Life Sciences</a> for his work and in 2020 the <a href="/wiki/Thomas_Hunt_Morgan_Medal" title="Thomas Hunt Morgan Medal">Thomas Hunt Morgan Medal</a> of the <a href="/wiki/Genetics_Society_of_America" title="Genetics Society of America">Genetics Society of America</a>.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> In 2016, <a href="/wiki/Semantic_Scholar" title="Semantic Scholar">Semantic Scholar</a> AI program included Botstein on its list of most top ten most influential biomedical researchers.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Personal">Personal</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=David_Botstein&action=edit&section=4" title="Edit section: Personal"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Botstein is an alumnus of <a href="/wiki/Camp_Rising_Sun_(New_York)" title="Camp Rising Sun (New York)">Camp Rising Sun</a>. He is the brother of the <a href="/wiki/Conducting" title="Conducting">conductor</a> <a href="/wiki/Leon_Botstein" title="Leon Botstein">Leon Botstein</a>. Both of Botstein's parents were physicians. </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=David_Botstein&action=edit&section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-genemedal-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-genemedal_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-genemedal_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFMahowald1988" class="citation journal cs1">Mahowald, A. 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Archived from <a rel="nofollow" class="external text" href="http://www.molbio2.princeton.edu/index.php?option=content&task=view&id=201">the original</a> on November 27, 2006<span class="reference-accessdate">. Retrieved <span class="nowrap">November 8,</span> 2006</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Princeton+University+-+Department+of+Molecular+Biology+-+David+Botstein&rft_id=http%3A%2F%2Fwww.molbio2.princeton.edu%2Findex.php%3Foption%3Dcontent%26task%3Dview%26id%3D201&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span> David Botstein at Princeton Department of Molecular Biology</span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external free" href="https://www.princeton.edu/genomics/botstein/">https://www.princeton.edu/genomics/botstein/</a> Botstein Laboratory Princeton</span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGitschier2006" class="citation journal cs1">Gitschier, J. 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Archived from <a rel="nofollow" class="external text" href="http://www.dailyprincetonian.com/archives/2005/10/18/news/13503.shtml">the original</a> on October 20, 2006<span class="reference-accessdate">. Retrieved <span class="nowrap">October 19,</span> 2006</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=The+Daily+Princetonian+-+Mapping+the+path+of+genetics&rft_id=http%3A%2F%2Fwww.dailyprincetonian.com%2Farchives%2F2005%2F10%2F18%2Fnews%2F13503.shtml&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span> The Daily Princetonian – Mapping the path of genetics</span> </li> <li id="cite_note-botstein-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-botstein_8-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBotstein1967" class="citation thesis cs1"><a class="mw-selflink selflink">Botstein, David</a> (1967). <i>The Synthesis and Maturation of Phage-P22 DNA</i> (PhD thesis). University of Michigan. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ProQuest" title="ProQuest">ProQuest</a> <a rel="nofollow" class="external text" href="https://search.proquest.com/docview/302261666">302261666</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adissertation&rft.title=The+Synthesis+and+Maturation+of+Phage-P22+DNA&rft.degree=PhD&rft.inst=University+of+Michigan&rft.date=1967&rft.aulast=Botstein&rft.aufirst=David&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFThean" class="citation web cs1">Thean, Tara. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20110830154320/http://www.dailyprincetonian.com/2009/10/12/24100/">"Integrated Science Pays Off for Graduates"</a>. <i>The Daily Princetonian</i>. 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(1980). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1686077">"Construction of a genetic linkage map in man using restriction fragment length polymorphisms"</a>. <i>American Journal of Human Genetics</i>. <b>32</b> (3): 314–331. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1686077">1686077</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/6247908">6247908</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Journal+of+Human+Genetics&rft.atitle=Construction+of+a+genetic+linkage+map+in+man+using+restriction+fragment+length+polymorphisms&rft.volume=32&rft.issue=3&rft.pages=314-331&rft.date=1980&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1686077%23id-name%3DPMC&rft_id=info%3Apmid%2F6247908&rft.aulast=Botstein&rft.aufirst=D.&rft.au=White%2C+R.&rft.au=Skolnick%2C+M.&rft.au=Davis%2C+R.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC1686077&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFEisenSpellmanBrownBotstein1998" class="citation journal cs1"><a href="/wiki/Michael_Eisen" title="Michael Eisen">Eisen, M.</a>; Spellman, P.; <a href="/wiki/Patrick_O._Brown" title="Patrick O. Brown">Brown, P.</a>; <a class="mw-selflink selflink">Botstein, D.</a> (1998). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC24541">"Cluster analysis and display of genome-wide expression patterns"</a>. <i>Proceedings of the National Academy of Sciences of the United States of America</i>. <b>95</b> (25): 14863–14868. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1998PNAS...9514863E">1998PNAS...9514863E</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.95.25.14863">10.1073/pnas.95.25.14863</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC24541">24541</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9843981">9843981</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences+of+the+United+States+of+America&rft.atitle=Cluster+analysis+and+display+of+genome-wide+expression+patterns&rft.volume=95&rft.issue=25&rft.pages=14863-14868&rft.date=1998&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC24541%23id-name%3DPMC&rft_id=info%3Apmid%2F9843981&rft_id=info%3Adoi%2F10.1073%2Fpnas.95.25.14863&rft_id=info%3Abibcode%2F1998PNAS...9514863E&rft.aulast=Eisen&rft.aufirst=M.&rft.au=Spellman%2C+P.&rft.au=Brown%2C+P.&rft.au=Botstein%2C+D.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC24541&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBotsteinCherryAshburnerBall2000" class="citation journal cs1"><a class="mw-selflink selflink">Botstein, D.</a>; Cherry, J. 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The Gene Ontology Consortium"</a>. <i><a href="/wiki/Nature_Genetics" title="Nature Genetics">Nature Genetics</a></i>. <b>25</b> (1): 25–29. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F75556">10.1038/75556</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3037419">3037419</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/10802651">10802651</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature+Genetics&rft.atitle=Gene+ontology%3A+Tool+for+the+unification+of+biology.+The+Gene+Ontology+Consortium&rft.volume=25&rft.issue=1&rft.pages=25-29&rft.date=2000&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3037419%23id-name%3DPMC&rft_id=info%3Apmid%2F10802651&rft_id=info%3Adoi%2F10.1038%2F75556&rft.aulast=Botstein&rft.aufirst=D.&rft.au=Cherry%2C+J.+M.&rft.au=Ashburner%2C+M.&rft.au=Ball%2C+C.+A.&rft.au=Blake%2C+J.+A.&rft.au=Butler%2C+H.&rft.au=Davis%2C+A.+P.&rft.au=Dolinski%2C+K.&rft.au=Dwight%2C+S.+S.&rft.au=Eppig%2C+J.+T.&rft.au=Harris%2C+M.+A.&rft.au=Hill%2C+D.+P.&rft.au=Issel-Tarver%2C+L.&rft.au=Kasarskis%2C+A.&rft.au=Lewis%2C+S.&rft.au=Matese%2C+J.+C.&rft.au=Richardson%2C+J.+E.&rft.au=Ringwald%2C+M.&rft.au=Rubin%2C+G.+M.&rft.au=Sherlock%2C+G.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3037419&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span> <span style="position:relative; top: -2px;"><span typeof="mw:File"><a href="/wiki/Open_access" title="open access publication – free to read"><img alt="Open access icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Open_Access_logo_PLoS_transparent.svg/9px-Open_Access_logo_PLoS_transparent.svg.png" decoding="async" width="9" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Open_Access_logo_PLoS_transparent.svg/14px-Open_Access_logo_PLoS_transparent.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/77/Open_Access_logo_PLoS_transparent.svg/18px-Open_Access_logo_PLoS_transparent.svg.png 2x" data-file-width="640" data-file-height="1000" /></a></span></span></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.molbiolcell.org/site/misc/edboard.xhtml">"MBC Editorial Board"</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=MBC+Editorial+Board&rft_id=http%3A%2F%2Fwww.molbiolcell.org%2Fsite%2Fmisc%2Fedboard.xhtml&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://genestogenomes.org/congratulations-to-the-recipients-of-the-2020-gsa-awards/">"Congratulations to the recipients of the 2020 GSA Awards!"</a>. 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Computer program guided by artificial intelligence says it knows"</a>. <i>Science | AAAS</i><span class="reference-accessdate">. Retrieved <span class="nowrap">September 22,</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Science+%7C+AAAS&rft.atitle=Who%27s+the+most+influential+biomedical+scientist%3F+Computer+program+guided+by+artificial+intelligence+says+it+knows&rft.date=2017-10-17&rft.aulast=Singh&rft.aufirst=Dalmeet&rft_id=https%3A%2F%2Fwww.science.org%2Fcontent%2Farticle%2Fwho-s-most-influential-biomedical-scientist-computer-program-guided-artificial&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADavid+Botstein" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=David_Botstein&action=edit&section=6" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://www.ibiology.org/science-and-society/integrated-science-curriculum-princeton/">David Botstein's Talk: "An Integrated Science Curriculum"</a></li> <li><a rel="nofollow" class="external text" href="https://www.ibiology.org/genetics-and-gene-regulation/fruits-genome-sequences/">David Botstein iBiology Seminar: "Fruits of the Genome Sequence"</a></li> <li><a rel="nofollow" class="external text" href="https://archivesspace.mit.edu/repositories/2/resources/790">David Botstein personal archives</a>, MC-0227. Massachusetts Institute of Technology, Department of Distinctive Collections, Cambridge, Massachusetts.</li></ul> <table class="wikitable succession-box noprint" style="margin:0.5em auto; font-size:small;clear:both;"> <tbody><tr> <th colspan="3" style="border-top: 5px solid #DAA520;">Academic offices </th></tr> <tr style="text-align:center;"> <td style="width:30%;" rowspan="1">Preceded by<div style="font-weight: bold"><a href="/wiki/Shirley_Tilghman" class="mw-redirect" title="Shirley Tilghman">Shirley Tilghman</a></div> </td> <td style="width: 40%; text-align: center;" rowspan="1"><b> Director <br /> Lewis-Sigler Institute for Integrative Genomics<br />Princeton University </b><br />2003-2015 </td> <td style="width: 30%; text-align: center;" rowspan="1">Succeeded by<div style="font-weight: bold"><a href="/wiki/Michael_Levine_(biologist)" title="Michael Levine (biologist)">Michael Levine</a></div> </td></tr> </tbody></table> <div class="navbox-styles"><style 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abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Breakthrough_Prize_laureates" title="Template:Breakthrough Prize laureates"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Breakthrough_Prize_laureates" title="Template talk:Breakthrough Prize laureates"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Breakthrough_Prize_laureates" title="Special:EditPage/Template:Breakthrough Prize laureates"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Breakthrough_Prize_laureates" style="font-size:114%;margin:0 4em"><a href="/wiki/Breakthrough_Prize" title="Breakthrough Prize">Breakthrough Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Breakthrough_Prize_in_Mathematics" title="Breakthrough Prize in Mathematics">Mathematics</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Simon_Donaldson" title="Simon Donaldson">Simon Donaldson</a>, <a href="/wiki/Maxim_Kontsevich" title="Maxim Kontsevich">Maxim Kontsevich</a>, <a href="/wiki/Jacob_Lurie" title="Jacob Lurie">Jacob Lurie</a>, <a href="/wiki/Terence_Tao" title="Terence Tao">Terence Tao</a> and <a href="/wiki/Richard_Taylor_(mathematician)" title="Richard Taylor (mathematician)">Richard Taylor</a> (2015)</li> <li><a href="/wiki/Ian_Agol" title="Ian Agol">Ian Agol</a> (2016)</li> <li><a href="/wiki/Jean_Bourgain" title="Jean Bourgain">Jean Bourgain</a> (2017)</li> <li><a href="/wiki/Christopher_Hacon" title="Christopher Hacon">Christopher Hacon</a>, <a href="/wiki/James_McKernan" title="James McKernan">James McKernan</a> (2018)</li> <li><a href="/wiki/Vincent_Lafforgue" title="Vincent Lafforgue">Vincent Lafforgue</a> (2019)</li> <li><a href="/wiki/Alex_Eskin" title="Alex Eskin">Alex Eskin</a> (2020)</li> <li><a href="/wiki/Martin_Hairer" title="Martin Hairer">Martin Hairer</a> (2021)</li> <li><a href="/wiki/Takuro_Mochizuki" class="mw-redirect" title="Takuro Mochizuki">Takuro Mochizuki</a> (2022)</li> <li><a href="/wiki/Daniel_A._Spielman" class="mw-redirect" title="Daniel A. Spielman">Daniel A. Spielman</a> (2023)</li> <li><a href="/wiki/Simon_Brendle" title="Simon Brendle">Simon Brendle</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Breakthrough_Prize_in_Fundamental_Physics" title="Breakthrough Prize in Fundamental Physics">Fundamental<br />physics</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nima_Arkani-Hamed" title="Nima Arkani-Hamed">Nima Arkani-Hamed</a>, <a href="/wiki/Alan_Guth" title="Alan Guth">Alan Guth</a>, <a href="/wiki/Alexei_Kitaev" title="Alexei Kitaev">Alexei Kitaev</a>, <a href="/wiki/Maxim_Kontsevich" title="Maxim Kontsevich">Maxim Kontsevich</a>, <a href="/wiki/Andrei_Linde" title="Andrei Linde">Andrei Linde</a>, <a href="/wiki/Juan_Mart%C3%ADn_Maldacena" class="mw-redirect" title="Juan Martín Maldacena">Juan Maldacena</a>, <a href="/wiki/Nathan_Seiberg" title="Nathan Seiberg">Nathan Seiberg</a>, <a href="/wiki/Ashoke_Sen" title="Ashoke Sen">Ashoke Sen</a>, <a href="/wiki/Edward_Witten" title="Edward Witten">Edward Witten</a> (2012)</li> <li><i>Special</i>: <a href="/wiki/Stephen_Hawking" title="Stephen Hawking">Stephen Hawking</a>, <a href="/wiki/Peter_Jenni" title="Peter Jenni">Peter Jenni</a>, <a href="/wiki/Fabiola_Gianotti" title="Fabiola Gianotti">Fabiola Gianotti</a> (ATLAS), <a href="/wiki/Michel_Della_Negra" title="Michel Della Negra">Michel Della Negra</a>, <a href="/wiki/Tejinder_Virdee" title="Tejinder Virdee">Tejinder Virdee</a>, <a href="/wiki/Guido_Tonelli" title="Guido Tonelli">Guido Tonelli</a>, <a href="/wiki/Joseph_Incandela" title="Joseph Incandela">Joseph Incandela</a> (CMS) and <a href="/wiki/Lyn_Evans" title="Lyn Evans">Lyn Evans</a> (LHC) (2013)</li> <li><a href="/wiki/Alexander_Markovich_Polyakov" title="Alexander Markovich Polyakov">Alexander Polyakov</a> (2013)</li> <li><a href="/wiki/Michael_Green_(physicist)" title="Michael Green (physicist)">Michael Green</a> and <a href="/wiki/John_Henry_Schwarz" title="John Henry Schwarz">John Henry Schwarz</a> (2014)</li> <li><a href="/wiki/Saul_Perlmutter" title="Saul Perlmutter">Saul Perlmutter</a> and members of the <a href="/wiki/Supernova_Cosmology_Project" title="Supernova Cosmology Project">Supernova Cosmology Project</a>; <a href="/wiki/Brian_Schmidt" title="Brian Schmidt">Brian Schmidt</a>, <a href="/wiki/Adam_Riess" title="Adam Riess">Adam Riess</a> and members of the <a href="/wiki/High-Z_Supernova_Search_Team" title="High-Z Supernova Search Team">High-Z Supernova Team</a> (2015)</li> <li><i>Special</i>: <a href="/wiki/Ronald_Drever" title="Ronald Drever">Ronald Drever</a>, <a href="/wiki/Kip_Thorne" title="Kip Thorne">Kip Thorne</a>, <a href="/wiki/Rainer_Weiss" title="Rainer Weiss">Rainer Weiss</a> and contributors to <a href="/wiki/LIGO" title="LIGO">LIGO</a> project (2016)</li> <li><a href="/wiki/Wang_Yifang" title="Wang Yifang">Yifang Wang</a>, <a href="/wiki/Kam-Biu_Luk" title="Kam-Biu Luk">Kam-Biu Luk</a> and the <a href="/wiki/Daya_Bay_Reactor_Neutrino_Experiment" title="Daya Bay Reactor Neutrino Experiment">Daya Bay team</a>, <a href="/wiki/Atsuto_Suzuki" title="Atsuto Suzuki">Atsuto Suzuki</a> and the <a href="/wiki/Kamioka_Liquid_Scintillator_Antineutrino_Detector" title="Kamioka Liquid Scintillator Antineutrino Detector">KamLAND</a> team, <a href="/wiki/K%C5%8Dichir%C5%8D_Nishikawa" title="Kōichirō Nishikawa">Kōichirō Nishikawa</a> and the <a href="/wiki/K2K_experiment" title="K2K experiment">K2K</a> / <a href="/wiki/T2K_experiment" title="T2K experiment">T2K</a> team, <a href="/wiki/Arthur_B._McDonald" title="Arthur B. McDonald">Arthur B. McDonald</a> and the <a href="/wiki/Sudbury_Neutrino_Observatory" title="Sudbury Neutrino Observatory">Sudbury Neutrino Observatory</a> team, <a href="/wiki/Takaaki_Kajita" title="Takaaki Kajita">Takaaki Kajita</a> and <a href="/wiki/Y%C5%8Dichir%C5%8D_Suzuki" title="Yōichirō Suzuki">Yōichirō Suzuki</a> and the <a href="/wiki/Super-Kamiokande" title="Super-Kamiokande">Super-Kamiokande</a> team (2016)</li> <li><a href="/wiki/Joseph_Polchinski" title="Joseph Polchinski">Joseph Polchinski</a>, <a href="/wiki/Andrew_Strominger" title="Andrew Strominger">Andrew Strominger</a>, <a href="/wiki/Cumrun_Vafa" title="Cumrun Vafa">Cumrun Vafa</a> (2017)</li> <li><a href="/wiki/Charles_L._Bennett" title="Charles L. Bennett">Charles L. Bennett</a>, <a href="/wiki/Gary_Hinshaw" title="Gary Hinshaw">Gary Hinshaw</a>, <a href="/wiki/Norman_Jarosik" title="Norman Jarosik">Norman Jarosik</a>, <a href="/wiki/Lyman_Page" title="Lyman Page">Lyman Page Jr.</a>, <a href="/wiki/David_Spergel" title="David Spergel">David Spergel</a> (2018)</li> <li><i>Special</i>: <a href="/wiki/Jocelyn_Bell_Burnell" title="Jocelyn Bell Burnell">Jocelyn Bell Burnell</a> (2018)</li> <li><a href="/wiki/Charles_L._Kane" title="Charles L. Kane">Charles Kane</a> and <a href="/wiki/Eugene_J._Mele" title="Eugene J. Mele">Eugene Mele</a> (2019)</li> <li><i>Special</i>: <a href="/wiki/Sergio_Ferrara" title="Sergio Ferrara">Sergio Ferrara</a>, <a href="/wiki/Daniel_Z._Freedman" title="Daniel Z. Freedman">Daniel Z. Freedman</a>, <a href="/wiki/Peter_van_Nieuwenhuizen" title="Peter van Nieuwenhuizen">Peter van Nieuwenhuizen</a> (2019)</li> <li><a href="/wiki/The_Event_Horizon_Telescope_Collaboration" class="mw-redirect" title="The Event Horizon Telescope Collaboration">The Event Horizon Telescope Collaboration</a> (2020)</li> <li><a href="/wiki/Eric_Adelberger" class="mw-redirect" title="Eric Adelberger">Eric Adelberger</a>, <a href="/wiki/Jens_H._Gundlach" title="Jens H. Gundlach">Jens H. Gundlach</a> and <a href="/wiki/Blayne_Heckel" title="Blayne Heckel">Blayne Heckel</a> (2021)</li> <li><i>Special</i>: <a href="/wiki/Steven_Weinberg" title="Steven Weinberg">Steven Weinberg</a> (2021)</li> <li><a href="/wiki/Hidetoshi_Katori" title="Hidetoshi Katori">Hidetoshi Katori</a> and <a href="/wiki/Jun_Ye" title="Jun Ye">Jun Ye</a> (2022)</li> <li><a href="/wiki/Charles_H._Bennett_(physicist)" title="Charles H. Bennett (physicist)">Charles H. Bennett</a>, <a href="/wiki/Gilles_Brassard" title="Gilles Brassard">Gilles Brassard</a>, <a href="/wiki/David_Deutsch" title="David Deutsch">David Deutsch</a>, <a href="/wiki/Peter_W._Shor" class="mw-redirect" title="Peter W. Shor">Peter W. Shor</a> (2023)</li> <li><a href="/wiki/John_Cardy" title="John Cardy">John Cardy</a> and <a href="/wiki/Alexander_Zamolodchikov" title="Alexander Zamolodchikov">Alexander Zamolodchikov</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Breakthrough_Prize_in_Life_Sciences" title="Breakthrough Prize in Life Sciences">Life sciences</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cornelia_Bargmann" title="Cornelia Bargmann">Cornelia Bargmann</a>, <a class="mw-selflink selflink">David Botstein</a>, <a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a>, <a href="/wiki/Hans_Clevers" title="Hans Clevers">Hans Clevers</a>, <a href="/wiki/Titia_de_Lange" title="Titia de Lange">Titia de Lange</a>, <a href="/wiki/Napoleone_Ferrara" title="Napoleone Ferrara">Napoleone Ferrara</a>, <a href="/wiki/Eric_Lander" title="Eric Lander">Eric Lander</a>, <a href="/wiki/Charles_Sawyers" title="Charles Sawyers">Charles Sawyers</a>, <a href="/wiki/Robert_Weinberg_(biologist)" title="Robert Weinberg (biologist)">Robert Weinberg</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> and <a href="/wiki/Bert_Vogelstein" title="Bert Vogelstein">Bert Vogelstein</a> (2013)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a>, <a href="/wiki/Mahlon_DeLong" title="Mahlon DeLong">Mahlon DeLong</a>, <a href="/wiki/Michael_N._Hall" title="Michael N. Hall">Michael N. Hall</a>, <a href="/wiki/Robert_S._Langer" title="Robert S. Langer">Robert S. Langer</a>, <a href="/wiki/Richard_P._Lifton" title="Richard P. Lifton">Richard P. Lifton</a> and <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a> (2014)</li> <li><a href="/wiki/Alim_Louis_Benabid" title="Alim Louis Benabid">Alim Louis Benabid</a>, <a href="/wiki/Charles_David_Allis" title="Charles David Allis">Charles David Allis</a>, <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a>, <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> and <a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> (2015)</li> <li><a href="/wiki/Edward_Boyden" title="Edward Boyden">Edward Boyden</a>, <a href="/wiki/Karl_Deisseroth" title="Karl Deisseroth">Karl Deisseroth</a>, <a href="/wiki/John_Hardy_(geneticist)" title="John Hardy (geneticist)">John Hardy</a>, <a href="/wiki/Helen_Hobbs" title="Helen Hobbs">Helen Hobbs</a> and <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a> (2016)</li> <li><a href="/wiki/Stephen_Elledge" title="Stephen Elledge">Stephen J. Elledge</a>, <a href="/wiki/Harry_F._Noller" title="Harry F. Noller">Harry F. Noller</a>, <a href="/wiki/Roel_Nusse" title="Roel Nusse">Roeland Nusse</a>, <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a>, <a href="/wiki/Huda_Zoghbi" title="Huda Zoghbi">Huda Zoghbi</a> (2017)</li> <li><a href="/wiki/Joanne_Chory" title="Joanne Chory">Joanne Chory</a>, <a href="/wiki/Peter_Walter" title="Peter Walter">Peter Walter</a>, <a href="/wiki/Kazutoshi_Mori" title="Kazutoshi Mori">Kazutoshi Mori</a>, <a href="/wiki/Kim_Nasmyth" title="Kim Nasmyth">Kim Nasmyth</a>, <a href="/wiki/Don_W._Cleveland" title="Don W. Cleveland">Don W. Cleveland</a> (2018)</li> <li><a href="/wiki/C._Frank_Bennett" title="C. Frank Bennett">C. Frank Bennett</a> and <a href="/wiki/Adrian_R._Krainer" class="mw-redirect" title="Adrian R. Krainer">Adrian R. Krainer</a>, <a href="/wiki/Angelika_Amon" title="Angelika Amon">Angelika Amon</a>, <a href="/wiki/Xiaowei_Zhuang" title="Xiaowei Zhuang">Xiaowei Zhuang</a>, <a href="/wiki/Zhijian_Chen" title="Zhijian Chen">Zhijian Chen</a> (2019)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a>, <a href="/wiki/Franz-Ulrich_Hartl" title="Franz-Ulrich Hartl">Franz-Ulrich Hartl</a>, <a href="/wiki/Arthur_L._Horwich" title="Arthur L. Horwich">Arthur L. Horwich</a>, <a href="/wiki/David_Julius" title="David Julius">David Julius</a>, <a href="/wiki/Virginia_Man-Yee_Lee" title="Virginia Man-Yee Lee">Virginia Man-Yee Lee</a> (2020)</li> <li><a href="/wiki/David_Baker_(biochemist)" title="David Baker (biochemist)">David Baker</a>, <a href="/wiki/Catherine_Dulac" title="Catherine Dulac">Catherine Dulac</a>, <a href="/wiki/Dennis_Lo" title="Dennis Lo">Dennis Lo</a>, <a href="/w/index.php?title=Richard_J._Youle&action=edit&redlink=1" class="new" title="Richard J. Youle (page does not exist)">Richard J. Youle</a><span class="noprint" style="font-size:85%; font-style: normal;"> [<a href="https://de.wikipedia.org/wiki/Richard_J._Youle" class="extiw" title="de:Richard J. Youle">de</a>]</span> (2021)</li> <li><a href="/wiki/Jeffery_W._Kelly" title="Jeffery W. Kelly">Jeffery W. Kelly</a>, <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a>, <a href="/wiki/Shankar_Balasubramanian" title="Shankar Balasubramanian">Shankar Balasubramanian</a>, <a href="/wiki/David_Klenerman" title="David Klenerman">David Klenerman</a> and <a href="/wiki/Pascal_Mayer" title="Pascal Mayer">Pascal Mayer</a> (2022)</li> <li><a href="/wiki/Clifford_P._Brangwynne" title="Clifford P. Brangwynne">Clifford P. Brangwynne</a>, <a href="/wiki/Anthony_A._Hyman" title="Anthony A. Hyman">Anthony A. Hyman</a>, <a href="/wiki/Demis_Hassabis" title="Demis Hassabis">Demis Hassabis</a>, <a href="/wiki/John_M._Jumper" title="John M. Jumper">John Jumper</a>, <a href="/wiki/Emmanuel_Mignot" title="Emmanuel Mignot">Emmanuel Mignot</a>, <a href="/wiki/Masashi_Yanagisawa" title="Masashi Yanagisawa">Masashi Yanagisawa</a> (2023)</li> <li><a href="/wiki/Carl_June" class="mw-redirect" title="Carl June">Carl June</a>, <a href="/wiki/Michel_Sadelain" title="Michel Sadelain">Michel Sadelain</a>, <a href="/wiki/Sabine_Hadida" title="Sabine Hadida">Sabine Hadida</a>, <a href="/wiki/Paul_Negulescu" title="Paul Negulescu">Paul Negulescu</a>, <a href="/w/index.php?title=Fredrick_Van_Goor&action=edit&redlink=1" class="new" title="Fredrick Van Goor (page does not exist)">Fredrick Van Goor</a>, <a href="/w/index.php?title=Thomas_Gasser&action=edit&redlink=1" class="new" title="Thomas Gasser (page does not exist)">Thomas Gasser</a>, <a href="/wiki/Ellen_Sidransky" title="Ellen Sidransky">Ellen Sidransky</a> and <a href="/wiki/Andrew_Singleton" title="Andrew Singleton">Andrew Singleton</a> (2024)</li></ul> 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style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/0000000364045980">ISNI</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/225278991">VIAF</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://id.worldcat.org/fast/61918/">FAST</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.oclc.org/worldcat/entity/E39PBJhGYCchdRrqbMptDbRFrq">WorldCat</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n80130585">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb122798985">France</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb122798985">BnF data</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=mzk2003195006&CON_LNG=ENG">Czech Republic</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p068703481">Netherlands</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://authority.bibsys.no/authority/rest/authorities/html/90137752">Norway</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://kopkatalogs.lv/F?func=direct&local_base=lnc10&doc_number=000177471&P_CON_LNG=ENG">Latvia</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" 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