Cilium | Definition, Function, & Facts

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It captures the prey with its cilia (also shown enlarged). Enzymes in the food vacuoles digest the prey, and contractile vacuoles expel excess water." loading="eager" /> </picture> <button class="magnifying-glass btn btn-circle position-absolute shadow btn-white top-10 right-10" aria-label="Zoom in"> <em class="material-icons link-blue" data-icon="zoom_in"></em> </button> </a> </div> <figcaption class="card-body"> <div class="md-assembly-caption text-muted font-14 font-serif line-clamp"> <span> <a class="gtm-assembly-link md-assembly-title font-weight-bold d-inline font-sans-serif mr-5 media-overlay-link" href="https://cdn.britannica.com/17/62917-050-48EA3253/protozoan-paramecium-swims-search-food-bacteria-prey.jpg" data-href="/media/1/117814/53857"></a> <span><span>The protozoan called paramecium swims freely in its search for food such as bacteria. It captures the prey with its cilia (also shown enlarged). Enzymes in the food vacuoles digest the prey, and contractile vacuoles expel excess water.</span></span> <button class="js-more-btn d-none btn btn-unstyled font-12 bg-white js-content" aria-label="Toggle more/less fact data"> <span class="link-blue">(more)</span> </button> </span> </div> </figcaption> </figure> </div><div class="topic-header"> <div class="d-flex align-items-top justify-content-between"> <div class="d-flex flex-column"> <div> <div> <h1>cilium</h1></div> </div> <div class="topic-identifier font-16 font-md-20">biology</div> </div> </div> <div class="d-none d-sm-flex flex-row"> <div class="mr-10 mb-15"> <button class="ai-ask-button btn border-2 btn-sm js-inline-ai-ask-button btn-outline-red-400 border-red-400"> Ask the Chatbot a Question </button> </div> <div class="d-none d-sm-block md-topic-tools qa-action-buttons mb-15" data-topic-id="117814"> <button class="js-tooltip btn btn-sm btn-outline-blue border pr-10 border-2" > <em class="material-icons md-icon ml-n10 my-n5 mr-5" data-icon="more_vert"></em> More Actions </button> <div class="md-more-popover popover popover-sm p-0 font-14 z-1"> <div> <a class="js-modal btn btn-sm btn-link gtm-topic-tool font-weight-bold btn-link" href="/print/article/117814" target="_blank" > <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="print"></em> Print </a> </div> <div> <button class="js-modal qa-cite-modal-button btn btn-sm btn-link gtm-topic-tool font-weight-bold btn-link" data-modal="[data-topic-id=117814] .md-cite-modal"> <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="verified"></em> Cite </button> <div class="md-cite-modal size-lg d-none"> <div class="md-modal-body"> <div class="h2 font-serif d-flex align-items-center pb-15 border-bottom mb-15"> <em class="material-icons text-blue mr-10">verified</em>Cite </div> <div class="font-serif"> While every effort has been made to follow citation style rules, there may be some discrepancies. Please refer to the appropriate style manual or other sources if you have any questions. </div> <div class="label mt-20 mb-10">Select Citation Style</div> <select class="js-citation-format-select form-select"> <option selected value="mla">MLA</option> <option value="apa">APA</option> <option value="chicago">Chicago Manual of Style</option> </select> <div class="citation font-serif border rounded p-15 mt-20" data-authors="The Editors of Encyclopaedia Britannica" data-title="cilium" data-published-date="26 Dec. 2023" data-url="https://www.britannica.com/science/cilium" > <div class="citation-text"></div> </div> <button class="js-copy-citation-button mt-20 btn btn-xs btn-outline-blue border shadow-sm pr-10" > <em class="material-icons md-icon ml-n10 my-n5 mr-5" data-icon="file_copy"></em> <span class="js-citation-status-text">Copy Citation</span> </button> </div> </div> </div> <div> <button class="js-share-modal-button js-modal btn btn-sm btn-link gtm-topic-tool font-weight-bold btn-link" data-modal="[data-topic-id=117814] .md-share-modal"> <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="share"></em> Share </button> <div class="md-share-modal size-lg d-none qa-share-modal"> <div class="md-modal-body"> <div class="h2 font-serif d-flex align-items-center pb-15 border-bottom"> <em class="material-icons text-blue mr-10" data-icon="share"></em> Share </div> <div class="label my-20">Share to social media</div> <div class="md-social-toolbar-circle d-flex align-items-start inverted" data-value="share" title="cilium" data-url="https://www.britannica.com/science/cilium" > <a class="social-icon facebook justify-content-center d-flex align-items-center align-self-center" data-provider="facebook" href="https://www.facebook.com/BRITANNICA/" target="_blank" rel="noopener"><span>Facebook</span></a> <a class="social-icon x justify-content-center d-flex align-items-center align-self-center" data-provider="x" href="https://x.com/britannica" target="_blank" rel="noopener"><span>X</span></a> </div> <div class="label pt-20 mt-20 mb-5 border-top">URL</div> <a class="font-serif text-truncate d-inline-block" href="https://www.britannica.com/science/cilium">https://www.britannica.com/science/cilium</a> </div> </div> </div> <div> <button class="js-feedback-modal-button js-modal btn btn-sm btn-link gtm-topic-tool font-weight-bold btn-link" data-modal=".md-feedback-modal"> <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="message"></em> Feedback </button> </div> <div> <button class="qa-external-website-modal-button js-modal btn btn-sm btn-link gtm-topic-tool font-weight-bold btn-link" data-modal="[data-topic-id=117814] .md-websites-modal"> <em class="material-icons md-icon ml-n10 mr-5" data-icon="link"></em> External Websites </button> </div> </div> <div class="md-feedback-modal size-lg d-none"> <div class="md-modal-body"> <div class="h2 font-serif pb-15 border-bottom"> Feedback </div> <form method="post" action="/submission/feedback/117814"> <div class="my-20"> Corrections? 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Let us know if you have suggestions to improve this article (requires login). </div> <div class="type-menu"> <label for="feedback-type" class="label mb-10">Feedback Type</label> <select id="feedback-type" class="form-select mb-30" name="feedbackTypeId" required> <option value="" selected="selected">Select a type (Required)</option> <option value="1">Factual Correction</option> <option value="2">Spelling/Grammar Correction</option> <option value="3">Link Correction</option> <option value="4">Additional Information</option> <option value="5">Other</option> </select> </div> <label for="feedback" class="label mb-10">Your Feedback</label> <textarea id="feedback" class="form-control mb-30" name="feedback" maxlength="3000" rows="7" required></textarea> <button class="btn btn-blue" type="submit">Submit Feedback</button> </form> <div class="success-messaging d-none mt-30"> <div class="title">Thank you for your feedback</div> <p>Our editors will review what you鈥檝e submitted and determine whether to revise the article.</p> </div> </div> </div> <div class="md-websites-modal size-lg d-none"> <div class="md-modal-body"> <div class="h2 font-serif pb-15 border-bottom font-weight-bold"> External Websites </div> <div class="pb-20"> <ul class="list-unstyled mt-20 lh-lg"> <li><a class="external" href="http://www.ucmp.berkeley.edu/protista/ciliata.html" target="_blank" rel="noopener ">University of California, Berkeley: Museum of Paleontology - Introduction to the Ciliata</a></li> </ul> </div> </div> </div> </div> </div> <div class="toc-header-marker"></div> <button class="ai-ask-button btn border-2 js-header-ai-ask-button d-none btn-sm btn-outline-red-400 border-red-400 mr-0 mr-lg-10 ml-5 ml-sm-10 ml-lg-0 p-10"> Ask the Chatbot a Question </button> <div class="caption alternate-titles">Also known as: cilia, ciliums, undulipodia</div> <div class="md-byline module-spacing "> <div class="font-serif font-12"> <span class="written-by text-gray-700"> Written and fact-checked by </span> <div class="editor-popover popover p-0"> <a class="d-block p-20 gtm-byline font-12 byline-contributor" href="/editor/The-Editors-of-Encyclopaedia-Britannica/4419" > <div class="editor-title font-16 font-weight-bold">The Editors of Encyclopaedia Britannica</div> <div class="editor-description font-12 font-serif mt-5 text-black">Encyclopaedia Britannica's editors oversee subject areas in which they have extensive knowledge, whether from years of experience gained by working on that content or via study for an advanced degree. They write new content and verify and edit content received from contributors.</div> </a> <div data-popper-arrow></div> </div> <span class="btn btn-link editor-link p-0 qa-byline-link gtm-byline font-12 byline-contributor text-decoration-underline"> The Editors of Encyclopaedia Britannica</span></div> <div class="last-updated font-12 font-serif"> <a class="byline-edit-history" href="https://www.britannica.com/science/cilium/additional-info#history" rel="nofollow">Article History</a> </div></div> </div> <button class="d-flex d-lg-none btn btn-outline-blue border rounded-sm shadow-sm mobile-toc-button gtm-mobile-toc-inline-button d-none d-sm-block js-sections-inline-button module-spacing btn d-lg-none"> <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="toc"></em> Table of Contents </button> <div class="d-flex d-sm-none flex-row"> <button class="d-flex d-lg-none btn btn-outline-blue border rounded-sm shadow-sm mobile-toc-button gtm-mobile-toc-inline-button js-sections-inline-button module-spacing"> <em class="material-icons mr-5 ml-n10 my-n5 md-icon" data-icon="toc"></em> Table of Contents </button> <button class="ai-ask-button btn border-2 ai-ask-button btn border-2 module-spacing btn-sm js-inline-ai-ask-button btn-outline-red-400 border-red-400 p-10 ml-5"> Ask the Chatbot a Question </button> </div> <div class="js-qf-module qf-module px-40 px-sm-20 py-15 mx-auto module-spacing font-14 bg-gray-50 rounded"> <div class="facts-list mt-10"> <div class=""> <div class="js-fact mb-10 line-clamp clamp-3"> <dl> <dt>Plural: </dt> <dd>cilia</dd> </dl> <button class="js-more-btn d-none btn btn-unstyled font-12 bg-gray-50" aria-label="Toggle more/less fact data"> <em class="js-content link-blue">(Show more)</em> </button> </div> </div> <div class=""> <div class="js-fact mb-10 line-clamp clamp-3"> <dl> <dt>Related Topics: </dt> <dd><a href="/science/kinocilium" topicid="318825">kinocilium</a></dd> <dd><a href="/science/membranelle" topicid="374297">membranelle</a></dd> <dd><a href="/science/axoneme" topicid="46366">axoneme</a></dd> <dd><a href="/science/metachronal-wave" topicid="377401">metachronal wave</a></dd> <dd><a href="/science/kinety" topicid="318224">kinety</a></dd> </dl> <button class="js-more-btn d-none btn btn-unstyled font-12 bg-gray-50" aria-label="Toggle more/less fact data"> <em class="js-content link-blue">(Show more)</em> </button> </div> <div class="text-center"> <a class="btn btn-sm btn-link p-0" href="/facts/cilium"> See all related content </a> </div> </div> </div> </div><span class="marker before-article"></span><section data-level="1" id="ref1"><span class="marker PREMOD1 mod-inline"></span><p class="topic-paragraph"><strong><span id="ref266121"></span>cilium</strong>, short eyelashlike filament that is numerous on <a href="https://www.britannica.com/science/tissue" class="md-crosslink autoxref " data-show-preview="true">tissue</a> cells of most animals and provides the means for <span id="ref266122"></span><a href="https://www.britannica.com/topic/locomotion" class="md-crosslink " data-show-preview="true">locomotion</a> of protozoans of the phylum <a href="https://www.britannica.com/science/ciliate" class="md-crosslink autoxref " data-show-preview="true">Ciliophora</a>. Cilia may be fused in short <a class="md-dictionary-link md-dictionary-tt-off eb" data-term="transverse" href="https://www.britannica.com/dictionary/transverse" data-type="EB">transverse</a> rows to form membranelles or in tufts to form cirri. Capable of beating in unison, cilia move mammalian ova through oviducts, generate water currents to carry food and oxygen past the gills of clams, carry food through the digestive systems of snails, circulate <a href="https://www.britannica.com/science/cerebrospinal-fluid" class="md-crosslink autoxref " data-show-preview="true">cerebrospinal fluid</a> of animals, and clean debris from the respiratory systems of mammals. In modified form, cilia trigger the discharge of stinging devices in jellyfish and give rise to the light-sensitive rods of the mammalian retina and the odour-detecting units of olfactory neurons.</p><div class="one-good-fact-module"> </div><span class="marker MOD1 mod-inline"></span><span class="marker PREMOD2 mod-inline"></span><div class="assemblies"><div class="w-100"><figure class="md-assembly m-0 mb-md-0 card card-borderless print-false" data-assembly-id="16442" data-asm-type="video"><div class="md-assembly-wrapper card-media " data-type="video" video-id="22035"><a data-id="22035" class="gtm-assembly-link d-flex justify-content-center" style="--aspect-ratio: 16/9" href="/video/cells-human-respiratory-tract/-16442"><img src="https://cdn.britannica.com/35/22035-138-8A80598D/cells-human-respiratory-tract.jpg?w=800&h=450&c=crop" alt="Observe ciliated cells throughout the human respiratory system" loading="lazy"><script type="application/json"> { "sources": [ { "file" : "//content.jwplatform.com/manifests/4wV82Bh0.m3u8" } ], "image": "https://cdn.britannica.com/35/22035-138-8A80598D/cells-human-respiratory-tract.jpg" ,"tracks": [ { "file" : "//assets-jpcust.jwpsrv.com/tracks/lj7RJLF5", "label": "English" } ] ,"adfile": "//content.jwplatform.com/manifests/XKpU7CSc.m3u8" } </script><div class="btn btn-xl btn-white btn-circle position-absolute shadow" style="top: 50%; transform: translateY(-50%)"><em class="material-icons" data-icon="play_arrow"></em></div></a></div><figcaption class="card-body"><div class="md-assembly-caption text-muted font-14 font-serif line-clamp"><span><span class="md-assembly-title font-weight-bold mr-5 d-inline font-sans-serif md-video-caption" video-control="22035">Observe ciliated cells throughout the human respiratory system</span><span>Ciliated cells of the human respiratory tract</span><button class="js-more-btn d-none btn btn-unstyled font-12 bg-white js-content" aria-label="Toggle more/less fact data"><span class="link-blue">(more)</span></button></span></div><a class="font-14 mt-10 d-inline-block" href="/science/cilium/images-videos">See all videos for this article</a></figcaption></figure></div></div><p class="topic-paragraph">A cilium, like a <a href="https://www.britannica.com/science/flagellum" class="md-crosslink " data-show-preview="true">flagellum</a>, is composed of a central core (the <span id="ref266123"></span><a href="https://www.britannica.com/science/axoneme" class="md-crosslink ">axoneme</a>), which contains two central microtubules that are surrounded by an outer ring of nine pairs of microtubules. The outer ring of microtubules is surrounded by a <a href="https://www.britannica.com/science/membrane-biology" class="md-crosslink autoxref " data-show-preview="true">membrane</a> that is <a class="md-dictionary-link md-dictionary-tt-off eb" data-term="continuous" href="https://www.britannica.com/dictionary/continuous" data-type="EB">continuous</a> with the <a href="https://www.britannica.com/science/cell-biology" class="md-crosslink autoxref " data-show-preview="true">cell</a> membrane; ciliary outgrowth is controlled by the <span id="ref266124"></span><a href="https://www.britannica.com/science/kinetosome" class="md-crosslink ">basal body</a> that is located just inside the cell surface at the base of the cilium. Beneath the surface of some cells there is a network of fibrous rootlets or microtubular bundles that may provide support for the <a href="https://www.britannica.com/science/epithelium" class="md-crosslink autoxref " data-show-preview="true">epithelium</a> or coordinate ciliary beating.</p><span class="marker MOD2 mod-inline"></span></section> <span class="md-signature"><a href="/editor/The-Editors-of-Encyclopaedia-Britannica/4419">The Editors of Encyclopaedia Britannica</a></span> <span class="md-signature">This article was most recently revised and updated by <a href="/editor/Adam-Augustyn/6394">Adam Augustyn</a>.</span><span class="marker end-of-content"></span><span class="marker after-article"></span></div> <div id="chatbot-root"></div> </div> </div> </div> <div class="ai-dialog-placeholder"></div> </div> </div> <aside class="col-md-da-320"></aside> </div> </div> </div> </div> </article></div> </div></div> </div> </main> <div id="md-footer"></div> <noscript><iframe src="//www.googletagmanager.com/ns.html?id=GTM-5W6NC8" height="0" width="0" style="display:none;visibility:hidden"></iframe></noscript> <script type="text/javascript" id="_informizely_script_tag"> var IzWidget = IzWidget || {}; (function (d) { var scriptElement = d.createElement('script'); scriptElement.type = 'text/javascript'; scriptElement.async = true; scriptElement.src = "https://insitez.blob.core.windows.net/site/f780f33e-a610-4ac2-af81-3eb184037547.js"; var node = d.getElementById('_informizely_script_tag'); node.parentNode.insertBefore(scriptElement, node); } )(document); </script>  <script> window.ap3c = window.ap3c || {}; var ap3c = window.ap3c; ap3c.cmd = ap3c.cmd || []; ap3c.cmd.push(function() { ap3c.init('ZO4siT4cLwnykPnzZWJtd3Byb2Q', 'https://engage.email.britannica.com/'); ap3c.track({v: 0}); }); ap3c.activity = function(act) { ap3c.act = (ap3c.act || []); ap3c.act.push(act); }; var s, t; s = document.createElement('script'); s.type = 'text/javascript'; s.src = "https://engage.email.britannica.com/app.js"; t = document.getElementsByTagName('script')[0]; t.parentNode.insertBefore(s, t); </script> <script class="marketing-page-info" type="application/json"> {"pageType":"Topic","templateName":"DESKTOP","pageNumber":1,"pagesTotal":1,"pageId":117814,"pageLength":214,"initialLoad":true,"lastPageOfScroll":false} </script> <script class="marketing-content-info" type="application/json"> [] </script> <script src="https://cdn.britannica.com/mendel-resources/3-130/js/libs/jquery-3.5.0.min.js?v=3.130.14"></script> <script type="text/javascript" data-type="Init Mendel Code Splitting"> (function() { $.ajax({ dataType: 'script', cache: true, url: 'https://cdn.britannica.com/mendel-resources/3-130/dist/topic-page.js?v=3.130.14' }); })(); </script> <script class="analytics-metadata" type="application/json"> {"leg":"C","adLeg":"C","userType":"ANONYMOUS","pageType":"Topic","pageSubtype":null,"articleTemplateType":"SMALL","gisted":false,"pageNumber":1,"hasSummarizeButton":false,"hasAskButton":false} </script> <script type="text/javascript"> EBStat={accountId:-1,hostnameOverride:'webstats.eb.com',domain:'www.britannica.com', json:''}; </script> <script type="text/javascript"> ( function() { $.ajax( { dataType: 'script', cache: true, url: '//www.britannica.com/webstats/mendelstats.js?v=1' } ) .done( function() { try {writeStat(null,EBStat);} catch(err){} } ); })(); </script> <div id="bc-fixed-dialogue"></div> </body> </html>

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