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vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Galerie"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Galerie</span> </div> </a> <ul id="toc-Galerie-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Références" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Références"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Références</span> </div> </a> <ul id="toc-Références-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Voir_aussi" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Voir_aussi"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Voir aussi</span> </div> </a> <button aria-controls="toc-Voir_aussi-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Afficher / masquer la sous-section Voir aussi</span> </button> <ul id="toc-Voir_aussi-sublist" class="vector-toc-list"> <li id="toc-Articles_connexes" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Articles_connexes"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.1</span> <span>Articles connexes</span> </div> </a> <ul id="toc-Articles_connexes-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Liens_externes" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Liens_externes"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.2</span> <span>Liens externes</span> </div> </a> <ul id="toc-Liens_externes-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Sommaire" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Basculer la table des matières" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Basculer la table des matières</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Accouplement (biologie)</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Aller à un article dans une autre langue. Disponible en 54 langues." > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-54" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">54 langues</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Paring" title="Paring – afrikaans" lang="af" hreflang="af" data-title="Paring" data-language-autonym="Afrikaans" data-language-local-name="afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-als mw-list-item"><a href="https://als.wikipedia.org/wiki/Begattung" title="Begattung – alémanique" lang="gsw" hreflang="gsw" data-title="Begattung" data-language-autonym="Alemannisch" data-language-local-name="alémanique" class="interlanguage-link-target"><span>Alemannisch</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AA%D8%B2%D8%A7%D9%88%D8%AC" title="تزاوج – arabe" lang="ar" hreflang="ar" data-title="تزاوج" data-language-autonym="العربية" data-language-local-name="arabe" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/C%C3%BCtl%C9%99%C5%9Fm%C9%99" title="Cütləşmə – azerbaïdjanais" lang="az" hreflang="az" data-title="Cütləşmə" data-language-autonym="Azərbaycanca" data-language-local-name="azerbaïdjanais" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D0%BF%D0%B0%D1%80%D0%B0%D0%B2%D0%B0%D0%BD%D0%BD%D0%B5" title="Спараванне – biélorusse" lang="be" hreflang="be" data-title="Спараванне" data-language-autonym="Беларуская" data-language-local-name="biélorusse" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-br mw-list-item"><a href="https://br.wikipedia.org/wiki/Embara%C3%B1" title="Embarañ – breton" lang="br" hreflang="br" data-title="Embarañ" data-language-autonym="Brezhoneg" data-language-local-name="breton" class="interlanguage-link-target"><span>Brezhoneg</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Ritual_d%27aparellament" title="Ritual d'aparellament – catalan" lang="ca" hreflang="ca" data-title="Ritual d'aparellament" data-language-autonym="Català" data-language-local-name="catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Parring" title="Parring – danois" lang="da" hreflang="da" data-title="Parring" data-language-autonym="Dansk" data-language-local-name="danois" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Begattung" title="Begattung – allemand" lang="de" hreflang="de" data-title="Begattung" data-language-autonym="Deutsch" data-language-local-name="allemand" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%96%CE%B5%CF%85%CE%B3%CE%AC%CF%81%CF%89%CE%BC%CE%B1" title="Ζευγάρωμα – grec" lang="el" hreflang="el" data-title="Ζευγάρωμα" data-language-autonym="Ελληνικά" data-language-local-name="grec" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Mating" title="Mating – anglais" lang="en" hreflang="en" data-title="Mating" data-language-autonym="English" data-language-local-name="anglais" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Kopulacio" title="Kopulacio – espéranto" lang="eo" hreflang="eo" data-title="Kopulacio" data-language-autonym="Esperanto" data-language-local-name="espéranto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Apareamiento" title="Apareamiento – espagnol" lang="es" hreflang="es" data-title="Apareamiento" data-language-autonym="Español" data-language-local-name="espagnol" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Kopulatsioon" title="Kopulatsioon – estonien" lang="et" hreflang="et" data-title="Kopulatsioon" data-language-autonym="Eesti" data-language-local-name="estonien" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AC%D9%81%D8%AA%E2%80%8C%DA%AF%DB%8C%D8%B1%DB%8C" title="جفت‌گیری – persan" lang="fa" hreflang="fa" data-title="جفت‌گیری" data-language-autonym="فارسی" data-language-local-name="persan" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Pariutuminen" title="Pariutuminen – finnois" lang="fi" hreflang="fi" data-title="Pariutuminen" data-language-autonym="Suomi" data-language-local-name="finnois" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Apareamento" title="Apareamento – galicien" lang="gl" hreflang="gl" data-title="Apareamento" data-language-autonym="Galego" data-language-local-name="galicien" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%94%D7%96%D7%93%D7%95%D7%95%D7%92%D7%95%D7%AA" title="הזדווגות – hébreu" lang="he" hreflang="he" data-title="הזדווגות" data-language-autonym="עברית" data-language-local-name="hébreu" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Parenje" title="Parenje – croate" lang="hr" hreflang="hr" data-title="Parenje" data-language-autonym="Hrvatski" data-language-local-name="croate" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Kawin" title="Kawin – indonésien" lang="id" hreflang="id" data-title="Kawin" data-language-autonym="Bahasa Indonesia" data-language-local-name="indonésien" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-ilo mw-list-item"><a href="https://ilo.wikipedia.org/wiki/Yot" title="Yot – ilocano" lang="ilo" hreflang="ilo" data-title="Yot" data-language-autonym="Ilokano" data-language-local-name="ilocano" class="interlanguage-link-target"><span>Ilokano</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Accoppiamento_(zoologia)" title="Accoppiamento (zoologia) – italien" lang="it" hreflang="it" data-title="Accoppiamento (zoologia)" data-language-autonym="Italiano" data-language-local-name="italien" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E4%BA%A4%E9%85%8D" title="交配 – japonais" lang="ja" hreflang="ja" data-title="交配" data-language-autonym="日本語" data-language-local-name="japonais" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-jbo mw-list-item"><a href="https://jbo.wikipedia.org/wiki/gletu" title="gletu – lojban" lang="jbo" hreflang="jbo" data-title="gletu" data-language-autonym="La .lojban." data-language-local-name="lojban" class="interlanguage-link-target"><span>La .lojban.</span></a></li><li class="interlanguage-link interwiki-kab mw-list-item"><a href="https://kab.wikipedia.org/wiki/Amercel" title="Amercel – kabyle" lang="kab" hreflang="kab" data-title="Amercel" data-language-autonym="Taqbaylit" data-language-local-name="kabyle" class="interlanguage-link-target"><span>Taqbaylit</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A8%D0%B0%D2%93%D1%8B%D0%BB%D1%8B%D1%81%D1%82%D1%8B%D1%80%D1%83" title="Шағылыстыру – kazakh" lang="kk" hreflang="kk" data-title="Шағылыстыру" data-language-autonym="Қазақша" data-language-local-name="kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EA%B5%90%EB%B0%B0" title="교배 – coréen" lang="ko" hreflang="ko" data-title="교배" data-language-autonym="한국어" data-language-local-name="coréen" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Poravimasis" title="Poravimasis – lituanien" lang="lt" hreflang="lt" data-title="Poravimasis" data-language-autonym="Lietuvių" data-language-local-name="lituanien" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%9F%D0%B0%D1%80%D0%B5%D1%9A%D0%B5" title="Парење – macédonien" lang="mk" hreflang="mk" data-title="Парење" data-language-autonym="Македонски" data-language-local-name="macédonien" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Pengawanan" title="Pengawanan – malais" lang="ms" hreflang="ms" data-title="Pengawanan" data-language-autonym="Bahasa Melayu" data-language-local-name="malais" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-my mw-list-item"><a href="https://my.wikipedia.org/wiki/%E1%80%99%E1%80%AD%E1%80%90%E1%80%BA%E1%80%9C%E1%80%AD%E1%80%AF%E1%80%80%E1%80%BA%E1%80%81%E1%80%BC%E1%80%84%E1%80%BA%E1%80%B8" title="မိတ်လိုက်ခြင်း – birman" lang="my" hreflang="my" data-title="မိတ်လိုက်ခြင်း" data-language-autonym="မြန်မာဘာသာ" data-language-local-name="birman" class="interlanguage-link-target"><span>မြန်မာဘာသာ</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%D9%81%D8%AA_%DA%AF%DB%8C%D8%B1%DB%8C" title="جفت گیری – mazandérani" lang="mzn" hreflang="mzn" data-title="جفت گیری" data-language-autonym="مازِرونی" data-language-local-name="mazandérani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Paren" title="Paren – néerlandais" lang="nl" hreflang="nl" data-title="Paren" data-language-autonym="Nederlands" data-language-local-name="néerlandais" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Paring" title="Paring – norvégien bokmål" lang="nb" hreflang="nb" data-title="Paring" data-language-autonym="Norsk bokmål" data-language-local-name="norvégien bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Kopulacja" title="Kopulacja – polonais" lang="pl" hreflang="pl" data-title="Kopulacja" data-language-autonym="Polski" data-language-local-name="polonais" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/C%C3%B3pula" title="Cópula – portugais" lang="pt" hreflang="pt" data-title="Cópula" data-language-autonym="Português" data-language-local-name="portugais" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/%C3%8Emperechere" title="Împerechere – roumain" lang="ro" hreflang="ro" data-title="Împerechere" data-language-autonym="Română" data-language-local-name="roumain" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A1%D0%BF%D0%B0%D1%80%D0%B8%D0%B2%D0%B0%D0%BD%D0%B8%D0%B5" title="Спаривание – russe" lang="ru" hreflang="ru" data-title="Спаривание" data-language-autonym="Русский" data-language-local-name="russe" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Parenje" title="Parenje – serbo-croate" lang="sh" hreflang="sh" data-title="Parenje" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="serbo-croate" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Mating" title="Mating – Simple English" lang="en-simple" hreflang="en-simple" data-title="Mating" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Kopul%C3%A1cia_(p%C3%A1renie)" title="Kopulácia (párenie) – slovaque" lang="sk" hreflang="sk" data-title="Kopulácia (párenie)" data-language-autonym="Slovenčina" data-language-local-name="slovaque" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Parjenje" title="Parjenje – slovène" lang="sl" hreflang="sl" data-title="Parjenje" data-language-autonym="Slovenščina" data-language-local-name="slovène" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sn mw-list-item"><a href="https://sn.wikipedia.org/wiki/Kushumira" title="Kushumira – shona" lang="sn" hreflang="sn" data-title="Kushumira" data-language-autonym="ChiShona" data-language-local-name="shona" class="interlanguage-link-target"><span>ChiShona</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/%C3%87ift%C3%ABzimi" title="Çiftëzimi – albanais" lang="sq" hreflang="sq" data-title="Çiftëzimi" data-language-autonym="Shqip" data-language-local-name="albanais" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Parenje" title="Parenje – serbe" lang="sr" hreflang="sr" data-title="Parenje" data-language-autonym="Српски / srpski" data-language-local-name="serbe" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Parning" title="Parning – suédois" lang="sv" hreflang="sv" data-title="Parning" data-language-autonym="Svenska" data-language-local-name="suédois" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a href="https://te.wikipedia.org/wiki/%E0%B0%B8%E0%B0%82%E0%B0%AF%E0%B1%8B%E0%B0%97%E0%B0%AE%E0%B1%81" title="సంయోగము – télougou" lang="te" hreflang="te" data-title="సంయోగము" data-language-autonym="తెలుగు" data-language-local-name="télougou" class="interlanguage-link-target"><span>తెలుగు</span></a></li><li class="interlanguage-link interwiki-tl mw-list-item"><a href="https://tl.wikipedia.org/wiki/Pagtatambalan" title="Pagtatambalan – tagalog" lang="tl" hreflang="tl" data-title="Pagtatambalan" data-language-autonym="Tagalog" data-language-local-name="tagalog" class="interlanguage-link-target"><span>Tagalog</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/%C3%87iftle%C5%9Fme" title="Çiftleşme – turc" lang="tr" hreflang="tr" data-title="Çiftleşme" data-language-autonym="Türkçe" data-language-local-name="turc" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A1%D0%BF%D0%B0%D1%80%D0%BE%D0%B2%D1%83%D0%B2%D0%B0%D0%BD%D0%BD%D1%8F" title="Спаровування – ukrainien" lang="uk" hreflang="uk" data-title="Спаровування" data-language-autonym="Українська" data-language-local-name="ukrainien" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Giao_ph%E1%BB%91i" title="Giao phối – vietnamien" lang="vi" hreflang="vi" data-title="Giao phối" data-language-autonym="Tiếng Việt" data-language-local-name="vietnamien" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E4%BA%A4%E9%85%8D" title="交配 – chinois" lang="zh" hreflang="zh" data-title="交配" data-language-autonym="中文" data-language-local-name="chinois" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Kau-ph%C3%B2e" title="Kau-phòe – minnan" lang="nan" hreflang="nan" data-title="Kau-phòe" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E4%BA%A4%E9%85%8D" title="交配 – cantonais" lang="yue" hreflang="yue" data-title="交配" data-language-autonym="粵語" data-language-local-name="cantonais" class="interlanguage-link-target"><span>粵語</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q228395#sitelinks-wikipedia" 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data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Apparence</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">déplacer vers la barre latérale</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">masquer</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">Un article de Wikipédia, l'encyclopédie libre.</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="fr" dir="ltr"><div class="bandeau-container metadata homonymie hatnote"><div class="bandeau-cell bandeau-icone" style="display:table-cell;padding-right:0.5em"><span class="noviewer" typeof="mw:File"><a href="/wiki/Aide:Homonymie" title="Aide:Homonymie"><img alt="Page d’aide sur l’homonymie" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Logo_disambig.svg/20px-Logo_disambig.svg.png" decoding="async" width="20" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Logo_disambig.svg/30px-Logo_disambig.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Logo_disambig.svg/40px-Logo_disambig.svg.png 2x" data-file-width="512" data-file-height="375" /></a></span></div><div class="bandeau-cell" style="display:table-cell;padding-right:0.5em"> <p>Pour les articles homonymes, voir <a href="/wiki/Accouplement" class="mw-disambig" title="Accouplement">Accouplement</a>. </p> </div></div> <div class="bandeau-container metadata homonymie hatnote"><div class="bandeau-cell bandeau-icone" style="display:table-cell;padding-right:0.5em"><span class="noviewer" typeof="mw:File"><a href="/wiki/Aide:Homonymie" title="Aide:Homonymie"><img alt="Page d’aide sur l’homonymie" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Logo_disambig.svg/20px-Logo_disambig.svg.png" decoding="async" width="20" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Logo_disambig.svg/30px-Logo_disambig.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Logo_disambig.svg/40px-Logo_disambig.svg.png 2x" data-file-width="512" data-file-height="375" /></a></span></div><div class="bandeau-cell" style="display:table-cell;padding-right:0.5em"> <p>Pour l'activité elle-même, voir <a href="/wiki/Copulation" title="Copulation">Copulation</a>. </p> </div></div> <div class="infobox_v3 infobox infobox--frwiki noarchive large"><div class="entete" style="background-color:#E1E1E1;color:#000000"><div>Accouplement</div></div><div><div class="images" style="padding:2px 0"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/Fichier:Graphosoma_lineatum_mating_(6).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8f/Graphosoma_lineatum_mating_%286%29.jpg/260px-Graphosoma_lineatum_mating_%286%29.jpg" decoding="async" width="260" height="208" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8f/Graphosoma_lineatum_mating_%286%29.jpg/390px-Graphosoma_lineatum_mating_%286%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8f/Graphosoma_lineatum_mating_%286%29.jpg/520px-Graphosoma_lineatum_mating_%286%29.jpg 2x" data-file-width="1032" data-file-height="825" /></a></span></div></div><table><caption style="color:#000000;text-align:center;background-color:#E1E1E1">Présentation</caption><tbody><tr class=""><th scope="row">Type</th><td class=""><div> <span class="wd_p31">Processus biologique <small>(<a href="https://en.wikipedia.org/wiki/Biological_process" class="extiw" title="en:Biological process"><span class="indicateur-langue" title="Article sur Wikipédia en anglais">en</span></a>)</small><span class="noprint wikidata-linkback skin-invert"><span class="mw-valign-baseline noviewer" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q228395?uselang=fr#P31" title="Voir et modifier les données sur Wikidata"><img alt="Voir et modifier les données sur Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></span></div></td></tr><tr class=""><th scope="row">Partie de</th><td class=""><div> <span class="wd_p361"><a href="/wiki/Sexualit%C3%A9_(reproduction)" title="Sexualité (reproduction)">Reproduction sexuée</a><span class="noprint wikidata-linkback skin-invert"><span class="mw-valign-baseline noviewer" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q228395?uselang=fr#P361" title="Voir et modifier les données sur Wikidata"><img alt="Voir et modifier les données sur Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></span></div></td></tr></tbody></table><p class="navbar noprint bordered navigation-not-searchable" style="border-top:1px solid #E1E1E1"><span class="plainlinks" style="text-align:left"><a class="external text" href="https://fr.wikipedia.org/w/index.php?title=Accouplement_%28biologie%29&veaction=edit&section=0">modifier</a> - <a class="external text" href="https://fr.wikipedia.org/w/index.php?title=Accouplement_%28biologie%29&action=edit&section=0">modifier le code</a> - <a href="https://www.wikidata.org/wiki/Q228395" class="extiw" title="d:Q228395">modifier Wikidata</a></span><span style="text-align:right"><span typeof="mw:File"><a href="/wiki/Mod%C3%A8le:Infobox" title="Documentation du modèle"><img alt="Documentation du modèle" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Info_Simple.svg/12px-Info_Simple.svg.png" decoding="async" width="12" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Info_Simple.svg/18px-Info_Simple.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/38/Info_Simple.svg/24px-Info_Simple.svg.png 2x" data-file-width="512" data-file-height="512" /></a></span></span></p></div> <p>L’<b>accouplement</b> est le terme utilisé en <a href="/wiki/Biologie" title="Biologie">biologie</a> pour désigner le rapprochement de deux <a href="/wiki/Individu" title="Individu">individus</a> sexuellement complémentaires aboutissant à une <a href="/wiki/Reproduction_(biologie)" title="Reproduction (biologie)">reproduction sexuée</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Modes_d'accouplement"><span id="Modes_d.27accouplement"></span>Modes d'accouplement</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Accouplement_(biologie)&veaction=edit&section=1" title="Modifier la section : Modes d'accouplement" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Accouplement_(biologie)&action=edit&section=1" title="Modifier le code source de la section : Modes d'accouplement"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Le terme <i>accouplement</i> peut désigner plus largement le <a href="/wiki/Rapport_sexuel" title="Rapport sexuel">rapport sexuel</a> (<a href="/wiki/P%C3%A9n%C3%A9tration_vaginale" title="Pénétration vaginale">rapport vaginal</a> ou <a href="/wiki/Sodomie" title="Sodomie">rapport anal</a>), mais également l'émission simultanée de <a href="/wiki/Gam%C3%A8te" title="Gamète">gamètes</a>, ou encore la fusion de deux individus dans le cas d'<a href="/wiki/Organisme_unicellulaire" title="Organisme unicellulaire">organismes unicellulaires</a>. </p><p>L'accouplement peut avoir lieu à la suite d'un rapprochement des deux individus par des <a href="/wiki/Ph%C3%A9romone" title="Phéromone">phéromones</a> et/ou par une <a href="/wiki/Parade_nuptiale" title="Parade nuptiale">parade nuptiale</a> ou à la suite d'une <a href="/wiki/S%C3%A9lection_sexuelle" title="Sélection sexuelle">sélection sexuelle</a>. </p><p>L'accouplement aboutit généralement à une <a href="/wiki/F%C3%A9condation" title="Fécondation">fécondation</a> interne mais il peut y avoir aussi émission simultanée et rapprochée des gamètes, suivie d'une <a href="/wiki/F%C3%A9condation_externe" title="Fécondation externe">fécondation externe</a> (<a href="/wiki/Amplexus" title="Amplexus">amplexus</a> chez les <a href="/wiki/Amphibia" title="Amphibia">amphibiens</a>). </p> <div class="mw-heading mw-heading2"><h2 id="Galerie">Galerie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Accouplement_(biologie)&veaction=edit&section=2" title="Modifier la section : Galerie" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Accouplement_(biologie)&action=edit&section=2" title="Modifier le code source de la section : Galerie"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul class="gallery mw-gallery-packed"> <li class="gallerybox" style="width: 188.66666666667px"> <div class="thumb" style="width: 186.66666666667px;"><span typeof="mw:File"><a href="/wiki/Fichier:Moscas._Fecundaci%C3%B3n._Bastavales,_Bri%C3%B3n,_090905.jpg" class="mw-file-description" title="Accouplement de mouches domestiques (Musca domestica)."><img alt="Accouplement de mouches domestiques (Musca domestica)." src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Moscas._Fecundaci%C3%B3n._Bastavales%2C_Bri%C3%B3n%2C_090905.jpg/280px-Moscas._Fecundaci%C3%B3n._Bastavales%2C_Bri%C3%B3n%2C_090905.jpg" decoding="async" width="187" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Moscas._Fecundaci%C3%B3n._Bastavales%2C_Bri%C3%B3n%2C_090905.jpg/420px-Moscas._Fecundaci%C3%B3n._Bastavales%2C_Bri%C3%B3n%2C_090905.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Moscas._Fecundaci%C3%B3n._Bastavales%2C_Bri%C3%B3n%2C_090905.jpg/560px-Moscas._Fecundaci%C3%B3n._Bastavales%2C_Bri%C3%B3n%2C_090905.jpg 2x" data-file-width="2599" data-file-height="1671" /></a></span></div> <div class="gallerytext">Accouplement de <a href="/wiki/Mouche_domestique" title="Mouche domestique">mouches domestiques</a> (<i>Musca domestica</i>).</div> </li> <li class="gallerybox" style="width: 162px"> <div class="thumb" style="width: 160px;"><span typeof="mw:File"><a href="/wiki/Fichier:AccouplementInsectes.jpg" class="mw-file-description" title="Accouplement de moustiques (Culicidae)."><img alt="Accouplement de moustiques (Culicidae)." src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cb/AccouplementInsectes.jpg/240px-AccouplementInsectes.jpg" decoding="async" width="160" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cb/AccouplementInsectes.jpg/360px-AccouplementInsectes.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cb/AccouplementInsectes.jpg/480px-AccouplementInsectes.jpg 2x" data-file-width="3968" data-file-height="2976" /></a></span></div> <div class="gallerytext">Accouplement de <a href="/wiki/Moustique" title="Moustique">moustiques</a> (<i>Culicidae</i>).</div> </li> <li class="gallerybox" style="width: 162px"> <div class="thumb" style="width: 160px;"><span typeof="mw:File"><span><video id="mwe_player_0" poster="//upload.wikimedia.org/wikipedia/commons/thumb/4/4c/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv/240px--Accouplement_de_bourdons_%28Bombus_sp.%29.ogv.jpg" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="160" height="120" data-durationhint="41" data-mwtitle="Accouplement_de_bourdons_(Bombus_sp.).ogv" data-mwprovider="wikimediacommons"><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/4/4c/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv.480p.vp9.webm" type="video/webm; codecs="vp9, opus"" data-transcodekey="480p.vp9.webm" data-width="640" data-height="480" /><source src="//upload.wikimedia.org/wikipedia/commons/4/4c/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv" type="video/ogg; codecs="theora, vorbis"" data-width="640" data-height="480" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/4/4c/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv.144p.mjpeg.mov" type="video/quicktime" data-transcodekey="144p.mjpeg.mov" data-width="192" data-height="144" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/4/4c/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv.240p.vp9.webm" type="video/webm; codecs="vp9, opus"" data-transcodekey="240p.vp9.webm" data-width="320" data-height="240" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/4/4c/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv.360p.webm" type="video/webm; codecs="vp8, vorbis"" data-transcodekey="360p.webm" data-width="480" data-height="360" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/4/4c/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv/Accouplement_de_bourdons_%28Bombus_sp.%29.ogv.360p.vp9.webm" type="video/webm; codecs="vp9, opus"" data-transcodekey="360p.vp9.webm" data-width="480" data-height="360" /></video></span></span></div> <div class="gallerytext">Accouplement de bourdons du genre <i><a href="/wiki/Bombus" title="Bombus">Bombus</a></i>.</div> </li> <li class="gallerybox" style="width: 92px"> <div class="thumb" style="width: 90px;"><span typeof="mw:File"><a href="/wiki/Fichier:Joined_moths.JPG" class="mw-file-description" title="Accouplement de Sphinx du peuplier (Laothoe populi)."><img alt="Accouplement de Sphinx du peuplier (Laothoe populi)." src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e3/Joined_moths.JPG/135px-Joined_moths.JPG" decoding="async" width="90" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e3/Joined_moths.JPG/202px-Joined_moths.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e3/Joined_moths.JPG/270px-Joined_moths.JPG 2x" data-file-width="2304" data-file-height="3072" /></a></span></div> <div class="gallerytext">Accouplement de Sphinx du peuplier (<i><a href="/wiki/Laothoe_populi" title="Laothoe populi">Laothoe populi</a></i>).</div> </li> <li class="gallerybox" style="width: 170px"> <div class="thumb" style="width: 168px;"><span typeof="mw:File"><a href="/wiki/Fichier:Bufo_Bufo.jpg" class="mw-file-description" title="Amplexus chez le crapaud commun (Bufo bufo)."><img alt="Amplexus chez le crapaud commun (Bufo bufo)." src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d9/Bufo_Bufo.jpg/252px-Bufo_Bufo.jpg" decoding="async" width="168" height="120" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d9/Bufo_Bufo.jpg/378px-Bufo_Bufo.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d9/Bufo_Bufo.jpg/503px-Bufo_Bufo.jpg 2x" data-file-width="1286" data-file-height="920" /></a></span></div> <div class="gallerytext"><a href="/wiki/Amplexus" title="Amplexus">Amplexus</a> chez le <a href="/wiki/Crapaud_commun" title="Crapaud commun">crapaud commun</a> (<i>Bufo bufo</i>).</div> </li> </ul> <div class="mw-heading mw-heading2"><h2 id="Références"><span id="R.C3.A9f.C3.A9rences"></span>Références</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Accouplement_(biologie)&veaction=edit&section=3" title="Modifier la section : Références" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Accouplement_(biologie)&action=edit&section=3" title="Modifier le code source de la section : Références"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="references-small decimal" style=""> </div> <div class="mw-heading mw-heading2"><h2 id="Voir_aussi">Voir aussi</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Accouplement_(biologie)&veaction=edit&section=4" title="Modifier la section : Voir aussi" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Accouplement_(biologie)&action=edit&section=4" title="Modifier le code source de la section : Voir aussi"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Articles_connexes">Articles connexes</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Accouplement_(biologie)&veaction=edit&section=5" title="Modifier la section : Articles connexes" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Accouplement_(biologie)&action=edit&section=5" title="Modifier le code source de la section : Articles connexes"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Reproduction_(biologie)" title="Reproduction (biologie)">Reproduction (biologie)</a></li> <li><a href="/wiki/Copulation" title="Copulation">Copulation</a></li></ul> <div class="mw-heading mw-heading3"><h3 id="Liens_externes">Liens externes</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Accouplement_(biologie)&veaction=edit&section=6" title="Modifier la section : Liens externes" class="mw-editsection-visualeditor"><span>modifier</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Accouplement_(biologie)&action=edit&section=6" title="Modifier le code source de la section : Liens externes"><span>modifier le code</span></a><span class="mw-editsection-bracket">]</span></span></div> <p class="mw-empty-elt"> </p> <ul><li class="mw-empty-elt"></li> <li class="mw-empty-elt"></li> <li><div class="liste-horizontale"><span class="wd_identifiers">Notices dans des dictionnaires ou encyclopédies généralistes<span class="noprint wikidata-linkback skin-invert"><span class="mw-valign-baseline noviewer" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q228395?uselang=fr#identifiers" title="Voir et modifier les données sur Wikidata"><img alt="Voir et modifier les données sur Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></span> : <ul><li><a rel="nofollow" class="external text" href="https://www.britannica.com/topic/mating"><i>Britannica</i></a></li> <li><a rel="nofollow" class="external text" href="https://denstoredanske.lex.dk//parring/"><i>Den Store Danske Encyklopædi</i></a></li> <li><a rel="nofollow" class="external text" href="https://www.enciclopedia.cat/EC-GEC-0171974.xml"><i>Gran Enciclopèdia Catalana</i></a></li> </ul></div></li> <li><div class="liste-horizontale"><span class="wd_identifiers"><a href="/wiki/Autorit%C3%A9_(sciences_de_l%27information)" title="Autorité (sciences de l'information)">Notices d'autorité</a><span class="noprint wikidata-linkback skin-invert"><span class="mw-valign-baseline noviewer" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q228395?uselang=fr#identifiers" title="Voir et modifier les données sur Wikidata"><img alt="Voir et modifier les données sur Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/10px-Blue_pencil.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/15px-Blue_pencil.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/73/Blue_pencil.svg/20px-Blue_pencil.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></span> : <ul><li><span class="nowrap uid noarchive"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb12049400j">BnF</a></span> (<span class="nowrap uid noarchive"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb12049400j">données</a></span>)</li> <li><span class="nowrap uid noarchive"><a rel="nofollow" class="external text" href="http://d-nb.info/gnd/4139614-5">GND</a></span></li> </ul></div></li></ul> <ul id="bandeau-portail" class="bandeau-portail"><li><span class="bandeau-portail-element"><span class="bandeau-portail-icone"><span class="noviewer skin-invert-image" typeof="mw:File"><a 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