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Blending inheritance - Wikipedia

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contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Blending inheritance</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. 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href="https://de.wikipedia.org/wiki/Blending_inheritance" title="Blending inheritance – German" lang="de" hreflang="de" data-title="Blending inheritance" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%88%D8%B1%D8%A7%D8%AB%D8%AA_%D8%A2%D9%85%DB%8C%D8%AE%D8%AA%D9%87" title="وراثت آمیخته – Persian" lang="fa" hreflang="fa" data-title="وراثت آمیخته" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/H%C3%A9r%C3%A9dit%C3%A9_par_m%C3%A9lange" title="Hérédité par mélange – French" lang="fr" hreflang="fr" data-title="Hérédité par mélange" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Herdanza_por_mestura" title="Herdanza por mestura – Galician" lang="gl" hreflang="gl" data-title="Herdanza por mestura" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%98%BC%ED%95%A9_%EC%9C%A0%EC%A0%84" title="혼합 유전 – Korean" lang="ko" hreflang="ko" data-title="혼합 유전" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Adunan_baka" title="Adunan baka – Malay" lang="ms" hreflang="ms" data-title="Adunan baka" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Blending_inheritance" title="Blending inheritance – Dutch" lang="nl" hreflang="nl" data-title="Blending inheritance" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Heran%C3%A7a_por_mistura" title="Herança por mistura – Portuguese" lang="pt" hreflang="pt" data-title="Herança por mistura" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B8%81%E0%B8%B2%E0%B8%A3%E0%B8%96%E0%B9%88%E0%B8%B2%E0%B8%A2%E0%B8%97%E0%B8%AD%E0%B8%94%E0%B8%A5%E0%B8%B1%E0%B8%81%E0%B8%A9%E0%B8%93%E0%B8%B0%E0%B9%81%E0%B8%9A%E0%B8%9A%E0%B8%9C%E0%B8%AA%E0%B8%A1" title="การถ่ายทอดลักษณะแบบผสม – Thai" lang="th" hreflang="th" data-title="การถ่ายทอดลักษณะแบบผสม" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Kar%C4%B1%C5%9Fmal%C4%B1_kal%C4%B1t%C4%B1m" title="Karışmalı kalıtım – Turkish" lang="tr" hreflang="tr" data-title="Karışmalı kalıtım" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E8%9E%8D%E5%90%88%E9%81%97%E4%BC%A0" title="融合遗传 – Chinese" lang="zh" hreflang="zh" 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class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Appearance</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">hide</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> <div id="mw-indicator-good-star" class="mw-indicator"><div class="mw-parser-output"><span typeof="mw:File"><a href="/wiki/Wikipedia:Good_articles*" title="This is a good article. Click here for more information."><img alt="This is a good article. Click here for more information." src="//upload.wikimedia.org/wikipedia/en/thumb/9/94/Symbol_support_vote.svg/19px-Symbol_support_vote.svg.png" decoding="async" width="19" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/94/Symbol_support_vote.svg/29px-Symbol_support_vote.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/94/Symbol_support_vote.svg/39px-Symbol_support_vote.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span></div></div> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Obsolete theory of genetics</div> <p class="mw-empty-elt"> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Blending_inheritance.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Blending_inheritance.svg/440px-Blending_inheritance.svg.png" decoding="async" width="440" height="166" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Blending_inheritance.svg/660px-Blending_inheritance.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Blending_inheritance.svg/880px-Blending_inheritance.svg.png 2x" data-file-width="744" data-file-height="280" /></a><figcaption>Flowers would converge to a single coloration in a few generations if <a href="/wiki/Inheritance_(biology)" class="mw-redirect" title="Inheritance (biology)">inheritance</a> blended the characteristics of the two parents.</figcaption></figure> <p><b>Blending inheritance</b> is an <a href="/wiki/Superseded_scientific_theories" class="mw-redirect" title="Superseded scientific theories">obsolete theory</a> in <a href="/wiki/Biology" title="Biology">biology</a> from the 19th century. The theory is that the progeny <a href="/wiki/Inheritance_(biology)" class="mw-redirect" title="Inheritance (biology)">inherits</a> any characteristic as the average of the parents' values of that characteristic. As an example of this, a crossing of a red flower variety with a white variety of the same <a href="/wiki/Species" title="Species">species</a> would yield pink-flowered offspring. </p><p><a href="/wiki/Charles_Darwin" title="Charles Darwin">Charles Darwin</a>'s theory of inheritance by <a href="/wiki/Pangenesis" title="Pangenesis">pangenesis</a>, with contributions to egg or sperm from every part of the body, implied blending inheritance. His reliance on this mechanism led <a href="/wiki/Fleeming_Jenkin" title="Fleeming Jenkin">Fleeming Jenkin</a> to attack Darwin's theory of <a href="/wiki/Natural_selection" title="Natural selection">natural selection</a> on the grounds that blending inheritance would average out any novel beneficial characteristic before selection had time to act. </p><p>Blending inheritance was discarded with the general acceptance of <a href="/wiki/Particulate_inheritance" title="Particulate inheritance">particulate inheritance</a> during the development of modern <a href="/wiki/Genetics" title="Genetics">genetics</a>, after <abbr title="circa">c.</abbr><span style="white-space:nowrap;">&#8201;1900</span>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="History">History</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Blending_inheritance&amp;action=edit&amp;section=1" title="Edit section: History"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/The_eclipse_of_Darwinism" title="The eclipse of Darwinism">The eclipse of Darwinism</a> and <a href="/wiki/Modern_synthesis_(20th_century)" title="Modern synthesis (20th century)">Modern synthesis (20th century)</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Darwin%27s_Pangenesis.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Darwin%27s_Pangenesis.svg/330px-Darwin%27s_Pangenesis.svg.png" decoding="async" width="330" height="113" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Darwin%27s_Pangenesis.svg/495px-Darwin%27s_Pangenesis.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/54/Darwin%27s_Pangenesis.svg/660px-Darwin%27s_Pangenesis.svg.png 2x" data-file-width="925" data-file-height="317" /></a><figcaption>Diagram of <a href="/wiki/Charles_Darwin" title="Charles Darwin">Charles Darwin</a>'s pangenesis theory. Every part of the body emits tiny particles, <a href="/wiki/Gemmule_(pangenesis)" class="mw-redirect" title="Gemmule (pangenesis)">gemmules</a>, which migrate to the <a href="/wiki/Gonad" title="Gonad">gonads</a> and contribute to the fertilised egg and so to the next generation. The theory implied that changes to the body during an organism's life would be inherited, as proposed in <a href="/wiki/Lamarckism" title="Lamarckism">Lamarckism</a>, and that <a href="/wiki/Inheritance_(biology)" class="mw-redirect" title="Inheritance (biology)">inheritance</a> would be blending.</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Darwin's_pangenesis"><span id="Darwin.27s_pangenesis"></span>Darwin's pangenesis</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Blending_inheritance&amp;action=edit&amp;section=2" title="Edit section: Darwin&#039;s pangenesis"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Pangenesis" title="Pangenesis">Pangenesis</a></div> <p><a href="/wiki/Charles_Darwin" title="Charles Darwin">Charles Darwin</a> developed his theory of <a href="/wiki/Evolution" title="Evolution">evolution</a> by <a href="/wiki/Natural_selection" title="Natural selection">natural selection</a> on the basis of an understanding of <a href="/wiki/Uniformitarianism" title="Uniformitarianism">uniform processes</a> in <a href="/wiki/Geology" title="Geology">geology</a>, acting over very long periods of time on inheritable variation within populations. One of those processes was competition for resources, as <a href="/wiki/Thomas_Malthus" class="mw-redirect" title="Thomas Malthus">Thomas Malthus</a> had indicated, leading to a struggle to survive and to reproduce. Since some individuals would by chance have traits that allowed them to leave more offspring, those traits would tend to increase in the population. Darwin assembled many lines of evidence to show that variation occurred and that <a href="/wiki/Artificial_selection" class="mw-redirect" title="Artificial selection">artificial selection</a> by animal and plant breeding had caused change. All of this demanded a reliable mechanism of <a href="/wiki/Inheritance_(biology)" class="mw-redirect" title="Inheritance (biology)">inheritance</a>.<sup id="cite_ref-Dawkins_1-0" class="reference"><a href="#cite_note-Dawkins-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> </p><p><a href="/wiki/Pangenesis" title="Pangenesis">Pangenesis</a> was Darwin's attempt to provide such a mechanism of inheritance. The idea was that each part of the parent's body emitted tiny particles called <a href="/wiki/Gemmule_(pangenesis)" class="mw-redirect" title="Gemmule (pangenesis)">gemmules</a>, which migrated through the body to contribute to that parent's <a href="/wiki/Gamete" title="Gamete">gametes</a>, their eggs or sperms. The theory had an intuitive appeal, as characteristics of all parts of the body, such as shape of nose, width of shoulders and length of legs are inherited from both the father and the mother. However, it had some serious weaknesses. Firstly, many characteristics can change during an individual's lifetime, and are affected by the environment: <a href="/wiki/Blacksmith" title="Blacksmith">blacksmiths</a> can develop strong arm muscles during their work, so the gemmules from these muscles ought to carry this acquired characteristic. That implies the <a href="/wiki/Lamarckism" title="Lamarckism">Lamarckian</a> <a href="/wiki/Inheritance_of_acquired_characteristics" class="mw-redirect" title="Inheritance of acquired characteristics">inheritance of acquired characteristics</a>. Secondly, the fact that the gemmules were supposed to mix together on <a href="/wiki/Fertilisation" title="Fertilisation">fertilisation</a> implies blending inheritance, namely that the offspring would all be intermediate between the father and the mother in every characteristic. That directly contradicts the observed facts of inheritance, not least that children are usually either male or female rather than all <a href="/wiki/Intersex" title="Intersex">intersex</a>, and that traits such as flower colour often re-emerge after a generation, even when they seem to disappear when two varieties are crossed.<sup id="cite_ref-Dawkins_1-1" class="reference"><a href="#cite_note-Dawkins-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Holterhoff2014_2-0" class="reference"><a href="#cite_note-Holterhoff2014-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Darwin1868_3-0" class="reference"><a href="#cite_note-Darwin1868-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> Darwin was aware of both these objections, and accordingly had strong doubts about blending inheritance, as evidenced in his private correspondence. In a letter to <a href="/wiki/T.H._Huxley" class="mw-redirect" title="T.H. Huxley">T.H. Huxley</a>, dated November 12, 1857, Darwin wrote: </p> <style data-mw-deduplicate="TemplateStyles:r1244412712">.mw-parser-output .templatequote{overflow:hidden;margin:1em 0;padding:0 32px}.mw-parser-output .templatequotecite{line-height:1.5em;text-align:left;margin-top:0}@media(min-width:500px){.mw-parser-output .templatequotecite{padding-left:1.6em}}</style><blockquote class="templatequote"><p>I have lately been inclined to speculate very crudely &amp; indistinctly, that propagation by true fertilisation, will turn out to be a sort of mixture &amp; not true fusion, of two distinct individuals, or rather of innumerable individuals, as each parent has its parents &amp; ancestors:— I can understand on no other view the way in which crossed forms go back to so large an extent to ancestral forms."<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup></p></blockquote> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Blending_Inheritance.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3b/Blending_Inheritance.svg/220px-Blending_Inheritance.svg.png" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3b/Blending_Inheritance.svg/330px-Blending_Inheritance.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3b/Blending_Inheritance.svg/440px-Blending_Inheritance.svg.png 2x" data-file-width="900" data-file-height="675" /></a><figcaption>Blending inheritance leads to the averaging out of every characteristic, which as the engineer <a href="/wiki/Fleeming_Jenkin" title="Fleeming Jenkin">Fleeming Jenkin</a> pointed out, would make natural selection impossible if blending were the mechanism of inheritance.<sup id="cite_ref-Dawkins_1-2" class="reference"><a href="#cite_note-Dawkins-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-FOOTNOTEBowler198323–26,_196–253_5-0" class="reference"><a href="#cite_note-FOOTNOTEBowler198323–26,_196–253-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-FOOTNOTELarson2004120–124_6-0" class="reference"><a href="#cite_note-FOOTNOTELarson2004120–124-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup></figcaption></figure> <p>In a letter to <a href="/wiki/Alfred_Wallace" class="mw-redirect" title="Alfred Wallace">Alfred Wallace</a>, dated February 6, 1866, Darwin mentioned conducting hybridization experiments with pea plants, not unlike those done by <a href="/wiki/Gregor_Mendel" title="Gregor Mendel">Gregor Mendel</a>, and like him obtaining <a href="/wiki/Mendel%27s_law_of_segregation" class="mw-redirect" title="Mendel&#39;s law of segregation">segregating</a> (unblended) varieties, effectively disproving his theory of pangenesis with blending: </p> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1244412712"><blockquote class="templatequote"><p>I do not think you understand what I mean by the non-blending of certain varieties. It does not refer to fertility; an instance I will explain. I crossed the Painted Lady and Purple <a href="/wiki/Sweetpea" class="mw-redirect" title="Sweetpea">sweetpeas</a>, which are very differently coloured varieties, and got, even out of the same pod, both varieties perfect but not intermediate. Something of this kind I should think must occur at least with your butterflies &amp; the three forms of <i><a href="/wiki/Lythrum" title="Lythrum">Lythrum</a></i>; tho’ those cases are in appearance so wonderful. I do not know that they are really more so than every female in the world producing distinct male and female offspring...<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup></p></blockquote> <p>Blending inheritance was also clearly incompatible with Darwin's theory of evolution by natural selection. The engineer <a href="/wiki/Fleeming_Jenkin" title="Fleeming Jenkin">Fleeming Jenkin</a> used this to attack natural selection in his 1867 review of Darwin's <i><a href="/wiki/On_the_Origin_of_Species" title="On the Origin of Species">On the Origin of Species</a></i>. Jenkin noted that if inheritance were by blending, any beneficial trait that might arise in a lineage would have "blended away" long before natural selection had time to act.<sup id="cite_ref-Dawkins_1-3" class="reference"><a href="#cite_note-Dawkins-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-FOOTNOTEBowler198323–26,_196–253_5-1" class="reference"><a href="#cite_note-FOOTNOTEBowler198323–26,_196–253-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-FOOTNOTELarson2004120–124_6-1" class="reference"><a href="#cite_note-FOOTNOTELarson2004120–124-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> The evolutionary biologist <a href="/wiki/Richard_Dawkins" title="Richard Dawkins">Richard Dawkins</a> commented that blending inheritance was observably wrong, as it implied that every generation would be more uniform than the one before, and that Darwin should have said as much to Jenkin. The problem was not with natural selection, but with blending, and in Dawkins's view, Darwin should have settled for saying that the mechanism of inheritance was unknown, but certainly non-blending.<sup id="cite_ref-Dawkins_1-4" class="reference"><a href="#cite_note-Dawkins-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Replacement_by_Mendelian_inheritance">Replacement by Mendelian inheritance</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Blending_inheritance&amp;action=edit&amp;section=3" title="Edit section: Replacement by Mendelian inheritance"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Punnett_square_mendel_flowers.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Punnett_square_mendel_flowers.svg/220px-Punnett_square_mendel_flowers.svg.png" decoding="async" width="220" height="220" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Punnett_square_mendel_flowers.svg/330px-Punnett_square_mendel_flowers.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/17/Punnett_square_mendel_flowers.svg/440px-Punnett_square_mendel_flowers.svg.png 2x" data-file-width="550" data-file-height="550" /></a><figcaption>A <a href="/wiki/Punnett_square" title="Punnett square">Punnett square</a> for one of Mendel's pea plant experiments - self-fertilization of the F1 generation, shows that inheritance is <a href="/wiki/Particulate_inheritance" title="Particulate inheritance">particulate</a>, not blending.</figcaption></figure> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Mendelian_inheritance" title="Mendelian inheritance">Mendelian inheritance</a> and <a href="/wiki/Germ_plasm" title="Germ plasm">Germ plasm</a></div> <p>Blending inheritance was dismissed by the eventual widespread acceptance, after his death, of <a href="/wiki/Gregor_Mendel" title="Gregor Mendel">Gregor Mendel</a>'s theory of <a href="/wiki/Particulate_inheritance" title="Particulate inheritance">particulate inheritance</a>, which he had presented in <i><a href="/wiki/Experiments_on_Plant_Hybridization" title="Experiments on Plant Hybridization">Experiments on Plant Hybridization</a></i> (1865).<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> In 1892, <a href="/wiki/August_Weismann" title="August Weismann">August Weismann</a> set out the idea of a hereditary material, which he called the <a href="/wiki/Germ_plasm" title="Germ plasm">germ plasm</a>, confined to the gonads and independent of the rest of the body (the <a href="/wiki/Somatic_(biology)" title="Somatic (biology)">soma</a>). In Weismann's view, the germ plasm formed the body, but the body did not influence the germ plasm, except indirectly by natural selection. This contradicted both Darwin's pangenesis and Lamarckian inheritance.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-FOOTNOTEBowler1989247–253,_257_11-0" class="reference"><a href="#cite_note-FOOTNOTEBowler1989247–253,_257-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> Mendel's work was rediscovered in 1900 by the geneticist <a href="/wiki/Hugo_de_Vries" title="Hugo de Vries">Hugo de Vries</a> and others, soon confirmed that same year by experiments by <a href="/wiki/William_Bateson" title="William Bateson">William Bateson</a>.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> Mendelian inheritance with segregating, particulate <a href="/wiki/Allele" title="Allele">alleles</a> came to be understood as the explanation for both discrete and continuously varying characteristics.<sup id="cite_ref-FOOTNOTELarson2004157–166,_221–243_13-0" class="reference"><a href="#cite_note-FOOTNOTELarson2004157–166,_221–243-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-FOOTNOTEBowler1989275–276_14-0" class="reference"><a href="#cite_note-FOOTNOTEBowler1989275–276-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">&#91;</span>a<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Blending_inheritance&amp;action=edit&amp;section=4" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Aristotle%27s_biology" title="Aristotle&#39;s biology">Aristotle's biology</a></li> <li><a href="/wiki/Incomplete_dominance" class="mw-redirect" title="Incomplete dominance">Incomplete dominance</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="Notes">Notes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Blending_inheritance&amp;action=edit&amp;section=5" title="Edit section: Notes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-lower-alpha"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-16">^</a></b></span> <span class="reference-text">Blending inheritance has sometimes been wrongly identified with the continuous inheritance advocated by the <a href="/wiki/Biometrics" title="Biometrics">biometricians</a> at the start of the 20th century. There were indeed disagreements between them and the early Mendelians, but these did not concern blending.<sup id="cite_ref-Porter2014_15-0" class="reference"><a href="#cite_note-Porter2014-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Blending_inheritance&amp;action=edit&amp;section=6" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239543626"><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-Dawkins-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Dawkins_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Dawkins_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Dawkins_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Dawkins_1-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Dawkins_1-4"><sup><i><b>e</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFDawkins,_Richard2003" class="citation news cs1"><a href="/wiki/Richard_Dawkins" title="Richard Dawkins">Dawkins, Richard</a> (8 February 2003). <a rel="nofollow" class="external text" href="https://www.theguardian.com/books/2003/feb/08/peopleinscience.evolution">"An early flowering of genetics"</a>. <a href="/wiki/The_Guardian" title="The Guardian">The Guardian</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.atitle=An+early+flowering+of+genetics&amp;rft.date=2003-02-08&amp;rft.au=Dawkins%2C+Richard&amp;rft_id=https%3A%2F%2Fwww.theguardian.com%2Fbooks%2F2003%2Ffeb%2F08%2Fpeopleinscience.evolution&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> <li id="cite_note-Holterhoff2014-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-Holterhoff2014_2-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHolterhoff,_Kate2014" class="citation journal cs1">Holterhoff, Kate (2014). "The History and Reception of Charles Darwin's Hypothesis of Pangenesis". <i>Journal of the History of Biology</i>. <b>47</b> (4): 661–695. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2Fs10739-014-9377-0">10.1007/s10739-014-9377-0</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/24570302">24570302</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:254544807">254544807</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Journal+of+the+History+of+Biology&amp;rft.atitle=The+History+and+Reception+of+Charles+Darwin%27s+Hypothesis+of+Pangenesis&amp;rft.volume=47&amp;rft.issue=4&amp;rft.pages=661-695&amp;rft.date=2014&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A254544807%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F24570302&amp;rft_id=info%3Adoi%2F10.1007%2Fs10739-014-9377-0&amp;rft.au=Holterhoff%2C+Kate&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> <li id="cite_note-Darwin1868-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-Darwin1868_3-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDarwin1868" class="citation book cs1">Darwin, Charles (1868). <a rel="nofollow" class="external text" href="http://darwin-online.org.uk/content/frameset?itemID=F880.1&amp;viewtype=text&amp;pageseq=1"><i>The variation of animals and plants under domestication</i></a>. London: John Murray. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/1-4191-8660-4" title="Special:BookSources/1-4191-8660-4"><bdi>1-4191-8660-4</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=The+variation+of+animals+and+plants+under+domestication&amp;rft.place=London&amp;rft.pub=John+Murray&amp;rft.date=1868&amp;rft.isbn=1-4191-8660-4&amp;rft.aulast=Darwin&amp;rft.aufirst=Charles&amp;rft_id=http%3A%2F%2Fdarwin-online.org.uk%2Fcontent%2Fframeset%3FitemID%3DF880.1%26viewtype%3Dtext%26pageseq%3D1&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.darwinproject.ac.uk/entry-2166">Darwin Correspondence Project, Letter #2166 -- Darwin C. R. to Huxley, T. H. dated before 12 Nov 1857</a></span> </li> <li id="cite_note-FOOTNOTEBowler198323–26,_196–253-5"><span class="mw-cite-backlink">^ <a href="#cite_ref-FOOTNOTEBowler198323–26,_196–253_5-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-FOOTNOTEBowler198323–26,_196–253_5-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><a href="#CITEREFBowler1983">Bowler 1983</a>, pp.&#160;23–26, 196–253.</span> </li> <li id="cite_note-FOOTNOTELarson2004120–124-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-FOOTNOTELarson2004120–124_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-FOOTNOTELarson2004120–124_6-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><a href="#CITEREFLarson2004">Larson 2004</a>, pp.&#160;120–124.</span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.darwinproject.ac.uk/entry-4989">Darwin Correspondence Project, Letter #4989 -- Darwin, C. R. to Wallace A. R. dated 6 Feb 1866</a></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMendel,_J._G.1866" class="citation journal cs1"><a href="/wiki/Gregor_Mendel" title="Gregor Mendel">Mendel, J. G.</a> (1866). <a rel="nofollow" class="external text" href="https://www.biodiversitylibrary.org/item/124139#page/133/mode/1up">"Versuche über Pflanzenhybriden"</a>. <i>Verhandlungen des Naturforschenden Vereines in Brünn</i>. <b>4</b> (1865): 3–47.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Verhandlungen+des+Naturforschenden+Vereines+in+Br%C3%BCnn&amp;rft.atitle=Versuche+%C3%BCber+Pflanzenhybriden&amp;rft.volume=4&amp;rft.issue=1865&amp;rft.pages=3-47&amp;rft.date=1866&amp;rft.au=Mendel%2C+J.+G.&amp;rft_id=https%3A%2F%2Fwww.biodiversitylibrary.org%2Fitem%2F124139%23page%2F133%2Fmode%2F1up&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDrueryBateson1901" class="citation journal cs1">Druery, C. T.; Bateson, William (1901). <a rel="nofollow" class="external text" href="http://www.esp.org/foundations/genetics/classical/gm-65.pdf">"Experiments in plant hybridization"</a> <span class="cs1-format">(PDF)</span>. <i>Journal of the Royal Horticultural Society</i>. <b>26</b>: 1–32.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Journal+of+the+Royal+Horticultural+Society&amp;rft.atitle=Experiments+in+plant+hybridization&amp;rft.volume=26&amp;rft.pages=1-32&amp;rft.date=1901&amp;rft.aulast=Druery&amp;rft.aufirst=C.+T.&amp;rft.au=Bateson%2C+William&amp;rft_id=http%3A%2F%2Fwww.esp.org%2Ffoundations%2Fgenetics%2Fclassical%2Fgm-65.pdf&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> <li id="cite_note-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-10">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWeismann,_August1892" class="citation book cs1"><a href="/wiki/August_Weismann" title="August Weismann">Weismann, August</a> (1892). <a rel="nofollow" class="external text" href="https://archive.org/details/bub_gb_f5U-AAAAYAAJ"><i>Das Keimplasma: eine Theorie der Vererbung</i></a>. Jena: Fischer.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Das+Keimplasma%3A+eine+Theorie+der+Vererbung&amp;rft.place=Jena&amp;rft.pub=Fischer&amp;rft.date=1892&amp;rft.au=Weismann%2C+August&amp;rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fbub_gb_f5U-AAAAYAAJ&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> <li id="cite_note-FOOTNOTEBowler1989247–253,_257-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEBowler1989247–253,_257_11-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFBowler1989">Bowler 1989</a>, pp.&#160;247–253, 257.</span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAmbrose" class="citation web cs1">Ambrose, Mike. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20160614210558/https://www.jic.ac.uk/germplas/PISUM/ZGS4F.HTM">"Mendel's Peas"</a>. Norwich, UK: Germplasm Resources Unit, <a href="/wiki/John_Innes_Centre" title="John Innes Centre">John Innes Centre</a>. Archived from <a rel="nofollow" class="external text" href="https://www.jic.ac.uk/germplas/PISUM/ZGS4F.HTM">the original</a> on 14 June 2016<span class="reference-accessdate">. Retrieved <span class="nowrap">13 December</span> 2017</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=unknown&amp;rft.btitle=Mendel%27s+Peas&amp;rft.place=Norwich%2C+UK&amp;rft.pub=Germplasm+Resources+Unit%2C+John+Innes+Centre&amp;rft.aulast=Ambrose&amp;rft.aufirst=Mike&amp;rft_id=https%3A%2F%2Fwww.jic.ac.uk%2Fgermplas%2FPISUM%2FZGS4F.HTM&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> <li id="cite_note-FOOTNOTELarson2004157–166,_221–243-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTELarson2004157–166,_221–243_13-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFLarson2004">Larson 2004</a>, pp.&#160;157–166, 221–243.</span> </li> <li id="cite_note-FOOTNOTEBowler1989275–276-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-FOOTNOTEBowler1989275–276_14-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFBowler1989">Bowler 1989</a>, pp.&#160;275–276.</span> </li> <li id="cite_note-Porter2014-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-Porter2014_15-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPorter2014" class="citation journal cs1">Porter, Theodore M. (2014). "The curious case of blending inheritance". <i>Studies in History and Philosophy of Science Part C: Studies in History and Philosophy of Biological and Biomedical Sciences</i>. <b>46</b>: 125–132. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.shpsc.2014.02.003">10.1016/j.shpsc.2014.02.003</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/24803228">24803228</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Studies+in+History+and+Philosophy+of+Science+Part+C%3A+Studies+in+History+and+Philosophy+of+Biological+and+Biomedical+Sciences&amp;rft.atitle=The+curious+case+of+blending+inheritance&amp;rft.volume=46&amp;rft.pages=125-132&amp;rft.date=2014&amp;rft_id=info%3Adoi%2F10.1016%2Fj.shpsc.2014.02.003&amp;rft_id=info%3Apmid%2F24803228&amp;rft.aulast=Porter&amp;rft.aufirst=Theodore+M.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Sources">Sources</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Blending_inheritance&amp;action=edit&amp;section=7" title="Edit section: Sources"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBowler1983" class="citation book cs1"><a href="/wiki/Peter_J._Bowler" title="Peter J. Bowler">Bowler, Peter J.</a> (1983). <i>The Eclipse of Darwinism: anti-Darwinian evolutionary theories in the decades around 1900</i>. <a href="/wiki/Johns_Hopkins_University_Press" title="Johns Hopkins University Press">Johns Hopkins University Press</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-8018-4391-4" title="Special:BookSources/978-0-8018-4391-4"><bdi>978-0-8018-4391-4</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=The+Eclipse+of+Darwinism%3A+anti-Darwinian+evolutionary+theories+in+the+decades+around+1900&amp;rft.pub=Johns+Hopkins+University+Press&amp;rft.date=1983&amp;rft.isbn=978-0-8018-4391-4&amp;rft.aulast=Bowler&amp;rft.aufirst=Peter+J.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBowler1989" class="citation book cs1"><a href="/wiki/Peter_J._Bowler" title="Peter J. Bowler">Bowler, Peter J.</a> (1989). <span class="id-lock-registration" title="Free registration required"><a rel="nofollow" class="external text" href="https://archive.org/details/evolutionhistory0000bowl"><i>Evolution: The History of an Idea</i></a></span> (2nd revised&#160;ed.). Berkeley, CA: <a href="/wiki/University_of_California_Press" title="University of California Press">University of California Press</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-520-06386-0" title="Special:BookSources/978-0-520-06386-0"><bdi>978-0-520-06386-0</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Evolution%3A+The+History+of+an+Idea&amp;rft.place=Berkeley%2C+CA&amp;rft.edition=2nd+revised&amp;rft.pub=University+of+California+Press&amp;rft.date=1989&amp;rft.isbn=978-0-520-06386-0&amp;rft.aulast=Bowler&amp;rft.aufirst=Peter+J.&amp;rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fevolutionhistory0000bowl&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLarson2004" class="citation book cs1"><a href="/wiki/Edward_Larson" class="mw-redirect" title="Edward Larson">Larson, Edward J.</a> (2004). <span class="id-lock-registration" title="Free registration required"><a rel="nofollow" class="external text" href="https://archive.org/details/evolutionremarka00lars"><i>Evolution:The Remarkable History of Scientific Theory</i></a></span>. Modern Library. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/0-679-64288-9" title="Special:BookSources/0-679-64288-9"><bdi>0-679-64288-9</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Evolution%3AThe+Remarkable+History+of+Scientific+Theory&amp;rft.pub=Modern+Library&amp;rft.date=2004&amp;rft.isbn=0-679-64288-9&amp;rft.aulast=Larson&amp;rft.aufirst=Edward+J.&amp;rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fevolutionremarka00lars&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3ABlending+inheritance" class="Z3988"></span></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist 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td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Evolutionary_biology" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Evolution" title="Template:Evolution"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Evolution" title="Template talk:Evolution"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Evolution" title="Special:EditPage/Template:Evolution"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Evolutionary_biology" style="font-size:114%;margin:0 4em"><a href="/wiki/Evolutionary_biology" title="Evolutionary biology">Evolutionary biology</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><a href="/wiki/Introduction_to_evolution" title="Introduction to evolution">Introduction</a></li> <li><a href="/wiki/Outline_of_evolution" title="Outline of evolution">Outline</a></li> <li><a href="/wiki/Timeline_of_the_evolutionary_history_of_life" title="Timeline of the evolutionary history of life">Timeline of evolution</a></li> <li><a href="/wiki/History_of_life" title="History of life">History of life</a></li> <li><a href="/wiki/Index_of_evolutionary_biology_articles" title="Index of evolutionary biology articles">Index</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Evolution" title="Evolution">Evolution</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></li> <li><a href="/wiki/Adaptation" title="Adaptation">Adaptation</a></li> <li><a href="/wiki/Adaptive_radiation" title="Adaptive radiation">Adaptive radiation</a></li> <li><a href="/wiki/Altruism_(biology)" title="Altruism (biology)">Altruism</a> <ul><li><a href="/wiki/Cheating_(biology)" title="Cheating (biology)">Cheating</a></li> <li><a href="/wiki/Reciprocal_altruism" title="Reciprocal altruism">Reciprocal</a></li></ul></li> <li><a href="/wiki/Baldwin_effect" title="Baldwin effect">Baldwin effect</a></li> <li><a href="/wiki/Cladistics" title="Cladistics">Cladistics</a></li> <li><a href="/wiki/Coevolution" title="Coevolution">Coevolution</a> <ul><li><a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">Mutualism</a></li></ul></li> <li><a href="/wiki/Common_descent" title="Common descent">Common descent</a></li> <li><a href="/wiki/Convergent_evolution" title="Convergent evolution">Convergence</a></li> <li><a href="/wiki/Divergent_evolution" title="Divergent evolution">Divergence</a></li> <li><a href="/wiki/Earliest_known_life_forms" title="Earliest known life forms">Earliest known life forms</a></li> <li><a href="/wiki/Evidence_of_common_descent" title="Evidence of common descent">Evidence of evolution</a></li> <li><a href="/wiki/Evolutionary_arms_race" title="Evolutionary arms race">Evolutionary arms race</a></li> <li><a href="/wiki/Evolutionary_pressure" title="Evolutionary pressure">Evolutionary pressure</a></li> <li><a href="/wiki/Exaptation" title="Exaptation">Exaptation</a></li> <li><a href="/wiki/Extinction" title="Extinction">Extinction</a> <ul><li><a href="/wiki/Extinction_event" title="Extinction event">Event</a></li></ul></li> <li><a href="/wiki/Homology_(biology)" title="Homology (biology)">Homology</a></li> <li><a href="/wiki/Last_universal_common_ancestor" title="Last universal common ancestor">Last universal common ancestor</a></li> <li><a href="/wiki/Macroevolution" title="Macroevolution">Macroevolution</a></li> <li><a href="/wiki/Microevolution" title="Microevolution">Microevolution</a></li> <li><a href="/wiki/Evolutionary_mismatch" title="Evolutionary mismatch">Mismatch</a></li> <li><a href="/wiki/Nonadaptive_radiation" title="Nonadaptive radiation">Non-adaptive radiation</a></li> <li><a href="/wiki/Abiogenesis" title="Abiogenesis">Origin of life</a></li> <li><a href="/wiki/Panspermia" title="Panspermia">Panspermia</a></li> <li><a href="/wiki/Parallel_evolution" title="Parallel evolution">Parallel evolution</a></li> <li><a href="/wiki/Signalling_theory" title="Signalling theory">Signalling theory</a> <ul><li><a href="/wiki/Handicap_principle" title="Handicap principle">Handicap principle</a></li></ul></li> <li><a href="/wiki/Speciation" title="Speciation">Speciation</a> <ul><li><a href="/wiki/Species" title="Species">Species</a></li> <li><a href="/wiki/Species_complex" title="Species complex">Species complex</a></li></ul></li> <li><a href="/wiki/Evolutionary_taxonomy" title="Evolutionary taxonomy">Taxonomy</a></li> <li><a href="/wiki/Unit_of_selection" title="Unit of selection">Unit of selection</a> <ul><li><a href="/wiki/Gene-centered_view_of_evolution" title="Gene-centered view of evolution">Gene-centered view of evolution</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Population_genetics" title="Population genetics">Population<br />genetics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Selective_breeding" title="Selective breeding">Artificial selection</a></li> <li><a href="/wiki/Biodiversity" title="Biodiversity">Biodiversity</a></li> <li><a href="/wiki/Evolutionarily_stable_strategy" title="Evolutionarily stable strategy">Evolutionarily stable strategy</a></li> <li><a href="/wiki/Fisher%27s_principle" title="Fisher&#39;s principle">Fisher's principle</a></li> <li><a href="/wiki/Fitness_(biology)" title="Fitness (biology)">Fitness</a> <ul><li><a href="/wiki/Inclusive_fitness" title="Inclusive fitness">Inclusive</a></li></ul></li> <li><a href="/wiki/Gene_flow" title="Gene flow">Gene flow</a></li> <li><a href="/wiki/Genetic_drift" title="Genetic drift">Genetic drift</a></li> <li><a href="/wiki/Kin_selection" title="Kin selection">Kin selection</a> <ul><li><a href="/wiki/Parental_investment" title="Parental investment">Parental investment</a></li> <li><a href="/wiki/Parent%E2%80%93offspring_conflict" title="Parent–offspring conflict">Parent–offspring conflict</a></li></ul></li> <li><a href="/wiki/Mutation" title="Mutation">Mutation</a></li> <li><a href="/wiki/Population" title="Population">Population</a></li> <li><a href="/wiki/Natural_selection" title="Natural selection">Natural selection</a></li> <li><a href="/wiki/Sexual_dimorphism" title="Sexual dimorphism">Sexual dimorphism</a></li> <li><a href="/wiki/Sexual_selection" title="Sexual selection">Sexual selection</a> <ul><li><a href="/wiki/Sexual_selection_in_flowering_plants" title="Sexual selection in flowering plants">Flowering plants</a></li> <li><a href="/wiki/Sexual_selection_in_fungi" title="Sexual selection in fungi">Fungi</a></li> <li><a href="/wiki/Mate_choice" title="Mate choice">Mate choice</a></li></ul></li> <li><a href="/wiki/Social_selection" title="Social selection">Social selection</a></li> <li><a href="/wiki/Trivers%E2%80%93Willard_hypothesis" title="Trivers–Willard hypothesis">Trivers–Willard hypothesis</a></li> <li><a href="/wiki/Genetic_variation" title="Genetic variation">Variation</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Development</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Canalisation_(genetics)" title="Canalisation (genetics)">Canalisation</a></li> <li><a href="/wiki/Evolutionary_developmental_biology" title="Evolutionary developmental biology">Evolutionary developmental biology</a></li> <li><a href="/wiki/Genetic_assimilation" title="Genetic assimilation">Genetic assimilation</a></li> <li><a href="/wiki/Inversion_(evolutionary_biology)" title="Inversion (evolutionary biology)">Inversion</a></li> <li><a href="/wiki/Modularity_(biology)" title="Modularity (biology)">Modularity</a></li> <li><a href="/wiki/Phenotypic_plasticity" title="Phenotypic plasticity">Phenotypic plasticity</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Of <a href="/wiki/Taxon" title="Taxon">taxa</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolution_of_bacteria" title="Evolution of bacteria">Bacteria</a></li> <li><a href="/wiki/Evolution_of_birds" title="Evolution of birds">Birds</a> <ul><li><a href="/wiki/Origin_of_birds" title="Origin of birds">origin</a></li></ul></li> <li><a href="/wiki/Evolution_of_brachiopods" title="Evolution of brachiopods">Brachiopods</a></li> <li><a href="/wiki/Evolution_of_molluscs" title="Evolution of molluscs">Molluscs</a> <ul><li><a href="/wiki/Evolution_of_cephalopods" title="Evolution of cephalopods">Cephalopods</a></li></ul></li> <li><a href="/wiki/Dinosaur#Evolutionary_history" title="Dinosaur">Dinosaurs</a></li> <li><a href="/wiki/Evolution_of_fish" title="Evolution of fish">Fish</a></li> <li><a href="/wiki/Evolution_of_fungi" title="Evolution of fungi">Fungi</a></li> <li><a href="/wiki/Evolution_of_insects" title="Evolution of insects">Insects</a> <ul><li><a href="/wiki/Evolution_of_butterflies" title="Evolution of butterflies">butterflies</a></li></ul></li> <li><a href="/wiki/History_of_life" title="History of life">Life</a></li> <li><a href="/wiki/Evolution_of_mammals" title="Evolution of mammals">Mammals</a> <ul><li><a href="/wiki/Cat_gap" title="Cat gap">cats</a></li> <li><a href="/wiki/Canidae#Evolution" title="Canidae">canids</a> <ul><li><a href="/wiki/Evolution_of_the_wolf" title="Evolution of the wolf">wolves</a></li> <li><a href="/wiki/Domestication_of_the_dog" title="Domestication of the dog">dogs</a></li></ul></li> <li><a href="/wiki/Hyena#Evolution" title="Hyena">hyenas</a></li> <li><a href="/wiki/Evolution_of_cetaceans" title="Evolution of cetaceans">dolphins and whales</a></li> <li><a href="/wiki/Evolution_of_the_horse" title="Evolution of the horse">horses</a></li> <li><a href="/wiki/Evolution_of_Macropodidae" title="Evolution of Macropodidae">Kangaroos</a></li> <li><a href="/wiki/Evolution_of_primates" title="Evolution of primates">primates</a> <ul><li><a href="/wiki/Human_evolution" title="Human evolution">humans</a></li> <li><a href="/wiki/Evolution_of_lemurs" title="Evolution of lemurs">lemurs</a></li></ul></li> <li><a href="/wiki/Evolution_of_sirenians" title="Evolution of sirenians">sea cows</a></li></ul></li> <li><a href="/wiki/Evolutionary_history_of_plants" title="Evolutionary history of plants">Plants</a> <ul><li><a href="/wiki/Pollinator-mediated_selection" title="Pollinator-mediated selection">pollinator-mediated</a></li></ul></li> <li><a href="/wiki/Evolution_of_reptiles" title="Evolution of reptiles">Reptiles</a></li> <li><a href="/wiki/Evolution_of_spiders" title="Evolution of spiders">Spiders</a></li> <li><a href="/wiki/Evolution_of_tetrapods" title="Evolution of tetrapods">Tetrapods</a></li> <li><a href="/wiki/Viral_evolution" title="Viral evolution">Viruses</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Of <a href="/wiki/Organ_(anatomy)" class="mw-redirect" title="Organ (anatomy)">organs</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolution_of_cells" title="Evolution of cells">Cell</a></li> <li><a href="/wiki/Models_of_DNA_evolution" title="Models of DNA evolution">DNA</a></li> <li><a href="/wiki/Evolution_of_flagella" title="Evolution of flagella">Flagella</a></li> <li><a href="/wiki/Eukaryote#Origin_of_eukaryotes" title="Eukaryote">Eukaryotes</a> <ul><li><a href="/wiki/Symbiogenesis" title="Symbiogenesis">symbiogenesis</a></li> <li><a href="/wiki/Chromosome#Eukaryotes" title="Chromosome">chromosome</a></li> <li><a href="/wiki/Endomembrane_system#Evolution" title="Endomembrane system">endomembrane system</a></li> <li><a href="/wiki/Mitochondrion#Origin_and_evolution" title="Mitochondrion">mitochondria</a></li> <li><a href="/wiki/Cell_nucleus#Evolution" title="Cell nucleus">nucleus</a></li> <li><a href="/wiki/Plastid#Origin" title="Plastid">plastids</a></li></ul></li> <li>In animals <ul><li><a href="/wiki/Evolution_of_the_eye" title="Evolution of the eye">eye</a></li> <li><a href="/wiki/Hair#Evolution" title="Hair">hair</a></li> <li><a href="/wiki/Evolution_of_mammalian_auditory_ossicles" title="Evolution of mammalian auditory ossicles">auditory ossicle</a></li> <li><a href="/wiki/Evolution_of_nervous_systems" title="Evolution of nervous systems">nervous system</a></li> <li><a href="/wiki/Evolution_of_the_brain" title="Evolution of the brain">brain</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Of <a href="/wiki/Biological_process" title="Biological process">processes</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolution_of_ageing" title="Evolution of ageing">Aging</a> <ul><li><a href="/wiki/Death#Evolution_of_aging_and_mortality" title="Death">Death</a></li> <li><a href="/wiki/Programmed_cell_death#Evolutionary_origin_of_mitochondrial_apoptosis" title="Programmed cell death">Programmed cell death</a></li></ul></li> <li><a href="/wiki/Origin_of_avian_flight" title="Origin of avian flight">Avian flight</a></li> <li><a href="/wiki/Evolution_of_biological_complexity" title="Evolution of biological complexity">Biological complexity</a></li> <li><a href="/wiki/Cooperation_(evolution)" title="Cooperation (evolution)">Cooperation</a></li> <li><a href="/wiki/Evolution_of_color_vision" title="Evolution of color vision">Color vision</a> <ul><li><a href="/wiki/Evolution_of_color_vision_in_primates" title="Evolution of color vision in primates">in primates</a></li></ul></li> <li><a href="/wiki/Evolution_of_emotion" title="Evolution of emotion">Emotion</a></li> <li><a href="/wiki/Empathy#Evolution_across_species" title="Empathy">Empathy</a></li> <li><a href="/wiki/Evolutionary_ethics" title="Evolutionary ethics">Ethics</a></li> <li><a href="/wiki/Evolution_of_eusociality" title="Evolution of eusociality">Eusociality</a></li> <li><a href="/wiki/Immune_system#Evolution_and_other_mechanisms" title="Immune system">Immune system</a></li> <li><a href="/wiki/Metabolism#Evolution" title="Metabolism">Metabolism</a></li> <li><a href="/wiki/Monogamy_in_animals" title="Monogamy in animals">Monogamy</a></li> <li><a href="/wiki/Evolution_of_morality" title="Evolution of morality">Morality</a></li> <li><a href="/wiki/Mosaic_evolution" title="Mosaic evolution">Mosaic evolution</a></li> <li><a href="/wiki/Multicellular_organism#Evolutionary_history" title="Multicellular organism">Multicellularity</a></li> <li><a href="/wiki/Evolution_of_sexual_reproduction" title="Evolution of sexual reproduction">Sexual reproduction</a> <ul><li><a href="/wiki/Anisogamy#Evolution" title="Anisogamy">Gamete differentiation/sexes</a></li> <li><a href="/wiki/Biological_life_cycle" title="Biological life cycle">Life cycles/nuclear phases</a></li> <li><a href="/wiki/Mating_type" title="Mating type">Mating types</a></li> <li><a href="/wiki/Origin_and_function_of_meiosis" title="Origin and function of meiosis">Meiosis</a></li> <li><a href="/wiki/Sex-determination_system#Evolution" title="Sex-determination system">Sex-determination</a></li></ul></li> <li><a href="/wiki/Evolution_of_snake_venom" title="Evolution of snake venom">Snake venom</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tempo_and_Mode_in_Evolution" title="Tempo and Mode in Evolution">Tempo and modes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Phyletic_gradualism" title="Phyletic gradualism">Gradualism</a>/<a href="/wiki/Punctuated_equilibrium" title="Punctuated equilibrium">Punctuated equilibrium</a>/<a href="/wiki/Saltation_(biology)" title="Saltation (biology)">Saltationism</a></li> <li><a href="/wiki/Point_mutation" title="Point mutation">Micromutation</a>/<a href="/wiki/Mutationism#Later_mutationist_theories" title="Mutationism">Macromutation</a></li> <li><a href="/wiki/Uniformitarianism" title="Uniformitarianism">Uniformitarianism</a>/<a href="/wiki/Catastrophism" title="Catastrophism">Catastrophism</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Speciation" title="Speciation">Speciation</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Allopatric_speciation" title="Allopatric speciation">Allopatric</a></li> <li><a href="/wiki/Anagenesis" title="Anagenesis">Anagenesis</a></li> <li><a href="/wiki/Catagenesis_(biology)" title="Catagenesis (biology)">Catagenesis</a></li> <li><a href="/wiki/Cladogenesis" title="Cladogenesis">Cladogenesis</a></li> <li><a href="/wiki/Cospeciation" title="Cospeciation">Cospeciation</a></li> <li><a href="/wiki/Ecological_speciation" title="Ecological speciation">Ecological</a></li> <li><a href="/wiki/Hybrid_speciation" title="Hybrid speciation">Hybrid</a></li> <li><a href="/wiki/Nonecological_speciation" title="Nonecological speciation">Non-ecological</a></li> <li><a href="/wiki/Parapatric_speciation" title="Parapatric speciation">Parapatric</a></li> <li><a href="/wiki/Peripatric_speciation" title="Peripatric speciation">Peripatric</a></li> <li><a href="/wiki/Reinforcement_(speciation)" title="Reinforcement (speciation)">Reinforcement</a></li> <li><a href="/wiki/Sympatric_speciation" title="Sympatric speciation">Sympatric</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/History_of_evolutionary_thought" title="History of evolutionary thought">History</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Evolutionary_ideas_of_the_Renaissance_and_Enlightenment" title="Evolutionary ideas of the Renaissance and Enlightenment">Renaissance and Enlightenment</a></li> <li><a href="/wiki/Transmutation_of_species" title="Transmutation of species">Transmutation of species</a></li> <li><a href="/wiki/David_Hume" title="David Hume">David Hume</a> <ul><li><i><a href="/wiki/Dialogues_Concerning_Natural_Religion" title="Dialogues Concerning Natural Religion">Dialogues Concerning Natural Religion</a></i></li></ul></li> <li><a href="/wiki/Charles_Darwin" title="Charles Darwin">Charles Darwin</a> <ul><li><i><a href="/wiki/On_the_Origin_of_Species" title="On the Origin of Species">On the Origin of Species</a></i></li></ul></li> <li><a href="/wiki/History_of_paleontology" title="History of paleontology">History of paleontology</a></li> <li><a href="/wiki/Transitional_fossil" title="Transitional fossil">Transitional fossil</a></li> <li><a class="mw-selflink selflink">Blending inheritance</a></li> <li><a href="/wiki/Mendelian_inheritance" title="Mendelian inheritance">Mendelian inheritance</a></li> <li><a href="/wiki/The_eclipse_of_Darwinism" title="The eclipse of Darwinism">The eclipse of Darwinism</a></li> <li><a href="/wiki/Neo-Darwinism" title="Neo-Darwinism">Neo-Darwinism</a></li> <li><a href="/wiki/Modern_synthesis_(20th_century)" title="Modern synthesis (20th century)">Modern synthesis</a></li> <li><a href="/wiki/History_of_molecular_evolution" title="History of molecular evolution">History of molecular evolution</a></li> <li><a href="/wiki/Extended_evolutionary_synthesis" title="Extended evolutionary synthesis">Extended evolutionary synthesis</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Philosophy</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Darwinism" title="Darwinism">Darwinism</a></li> <li><a href="/wiki/Alternatives_to_Darwinian_evolution" title="Alternatives to Darwinian evolution">Alternatives</a> <ul><li><a href="/wiki/Catastrophism" title="Catastrophism">Catastrophism</a></li> <li><a href="/wiki/Lamarckism" title="Lamarckism">Lamarckism</a></li> <li><a href="/wiki/Orthogenesis" title="Orthogenesis">Orthogenesis</a></li> <li><a href="/wiki/Mutationism" title="Mutationism">Mutationism</a></li> <li><a href="/wiki/Saltation_(biology)" title="Saltation (biology)">Saltationism</a></li> <li><a href="/wiki/Structuralism_(biology)" title="Structuralism (biology)">Structuralism</a> <ul><li><a href="/wiki/Spandrel_(biology)" title="Spandrel (biology)">Spandrel</a></li></ul></li> <li><a href="/wiki/Theistic_evolution" title="Theistic evolution">Theistic</a></li> <li><a href="/wiki/Vitalism" title="Vitalism">Vitalism</a></li></ul></li> <li><a href="/wiki/Teleology_in_biology" title="Teleology in biology">Teleology in biology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Related</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biogeography" title="Biogeography">Biogeography</a></li> <li><a href="/wiki/Ecological_genetics" title="Ecological genetics">Ecological genetics</a></li> <li><a href="/wiki/Evolutionary_medicine" title="Evolutionary medicine">Evolutionary medicine</a></li> <li><a href="/wiki/Group_selection" title="Group selection">Group selection</a> <ul><li><a href="/wiki/Cultural_evolution" title="Cultural evolution">Cultural evolution</a></li> <li><a href="/wiki/Cultural_group_selection" title="Cultural group selection">Cultural group selection</a></li> <li><a href="/wiki/Dual_inheritance_theory" title="Dual inheritance theory">Dual inheritance theory</a></li></ul></li> <li><a href="/wiki/Hologenome_theory_of_evolution" title="Hologenome theory of evolution">Hologenome theory of evolution</a></li> <li><a href="/wiki/Missing_heritability_problem" title="Missing heritability problem">Missing heritability problem</a></li> <li><a href="/wiki/Molecular_evolution" title="Molecular evolution">Molecular evolution</a></li> <li><a href="/wiki/Astrobiology" title="Astrobiology">Astrobiology</a></li> <li><a href="/wiki/Phylogenetics" title="Phylogenetics">Phylogenetics</a> <ul><li><a href="/wiki/Phylogenetic_tree" title="Phylogenetic tree">Tree</a></li></ul></li> <li><a href="/wiki/Polymorphism_(biology)" title="Polymorphism (biology)">Polymorphism</a></li> <li><a href="/wiki/Protocell" title="Protocell">Protocell</a></li> <li><a href="/wiki/Systematics" title="Systematics">Systematics</a></li> <li><a href="/wiki/Transgenerational_epigenetic_inheritance" title="Transgenerational epigenetic inheritance">Transgenerational epigenetic inheritance</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Evolutionary_biology" title="Category:Evolutionary biology">Category</a></li> <li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" 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