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div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Allan Bradley</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Professor_Sir_Allan_Bradley,_Founder_%26_Chief_Scientific_Officer_at_the_launch_of_the_Shakespeare_Review_(8740129867).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/eb/Professor_Sir_Allan_Bradley%2C_Founder_%26_Chief_Scientific_Officer_at_the_launch_of_the_Shakespeare_Review_%288740129867%29.jpg/220px-Professor_Sir_Allan_Bradley%2C_Founder_%26_Chief_Scientific_Officer_at_the_launch_of_the_Shakespeare_Review_%288740129867%29.jpg" decoding="async" width="220" height="330" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/eb/Professor_Sir_Allan_Bradley%2C_Founder_%26_Chief_Scientific_Officer_at_the_launch_of_the_Shakespeare_Review_%288740129867%29.jpg/330px-Professor_Sir_Allan_Bradley%2C_Founder_%26_Chief_Scientific_Officer_at_the_launch_of_the_Shakespeare_Review_%288740129867%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/eb/Professor_Sir_Allan_Bradley%2C_Founder_%26_Chief_Scientific_Officer_at_the_launch_of_the_Shakespeare_Review_%288740129867%29.jpg/440px-Professor_Sir_Allan_Bradley%2C_Founder_%26_Chief_Scientific_Officer_at_the_launch_of_the_Shakespeare_Review_%288740129867%29.jpg 2x" data-file-width="3456" data-file-height="5184" /></a></span><div class="infobox-caption">Allan Bradley at the launch of the Shakespeare review, organised by <a href="/wiki/Policy_Exchange" title="Policy Exchange">Policy Exchange</a></div></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">Allan Bradley</div></td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/University_of_Cambridge" title="University of Cambridge">University of Cambridge</a> (BA, MA, PhD)<sup id="cite_ref-whoswho_9-0" class="reference"><a href="#cite_note-whoswho-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup></td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data"><a href="/wiki/Embryonic_stem_cell" title="Embryonic stem cell">Embryonic stem cells</a><sup id="cite_ref-royal_10-0" class="reference"><a href="#cite_note-royal-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style><div class="plainlist"> <ul><li><a href="/wiki/Searle_Scholars_Program" title="Searle Scholars Program">Searle Scholar</a> (1988)</li> <li><a href="/wiki/European_Molecular_Biology_Organization" title="European Molecular Biology Organization">EMBO member</a> (2006)<sup id="cite_ref-membo_1-0" class="reference"><a href="#cite_note-membo-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">FRS</a> (2002)</li></ul> </div></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Genomics" title="Genomics">Mouse genomics</a><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-mice_6-0" class="reference"><a href="#cite_note-mice-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li><a href="/wiki/Baylor_College_of_Medicine" title="Baylor College of Medicine">Baylor College of Medicine</a></li> <li><a href="/wiki/Wellcome_Trust_Sanger_Institute" class="mw-redirect" title="Wellcome Trust Sanger Institute">Wellcome Trust Sanger Institute</a></li> <li><a href="/wiki/Howard_Hughes_Medical_Institute" title="Howard Hughes Medical Institute">Howard Hughes Medical Institute</a></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Thesis" title="Thesis">Thesis</a></th><td class="infobox-data"><i><a rel="nofollow" class="external text" href="https://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.373263">Isolation characterisation and developmental potential of murine embryo-derived stem cells</a></i> <span style="font-size:97%;">(1986)</span></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Doctoral_advisor" title="Doctoral advisor">Doctoral advisor</a></th><td class="infobox-data"><a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a><sup id="cite_ref-wtsi_7-0" class="reference"><a href="#cite_note-wtsi-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Watts2007_8-0" class="reference"><a href="#cite_note-Watts2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></td></tr><tr style="display:none"><td colspan="2"> </td></tr><tr><th scope="row" class="infobox-label">Website</th><td class="infobox-data"><span class="url"><a rel="nofollow" class="external text" href="http://sanger.ac.uk/person/bradley-allan/">sanger<wbr />.ac<wbr />.uk<wbr />/person<wbr />/bradley-allan<wbr />/</a></span></td></tr></tbody></table> <p><b>Allan Bradley</b> <a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">FRS</a> is a British <a href="/wiki/Geneticist" title="Geneticist">geneticist</a> at the <a href="/wiki/Wellcome_Trust_Sanger_Institute" class="mw-redirect" title="Wellcome Trust Sanger Institute">Wellcome Trust Sanger Institute</a>.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-scopus_13-0" class="reference"><a href="#cite_note-scopus-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Education">Education</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Allan_Bradley&action=edit&section=1" title="Edit section: Education"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Bradley was educated at the <a href="/wiki/University_of_Cambridge" title="University of Cambridge">University of Cambridge</a> where he earned Bachelor of Arts, Master of Arts and PhD<sup id="cite_ref-bradleyphd_15-0" class="reference"><a href="#cite_note-bradleyphd-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> degrees in genetics from <a href="/wiki/Trinity_College,_Cambridge" title="Trinity College, Cambridge">Trinity College, Cambridge</a> gained while working in the laboratory of <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a>.<sup id="cite_ref-whoswho_9-1" class="reference"><a href="#cite_note-whoswho-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-wtsi_7-1" class="reference"><a href="#cite_note-wtsi-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Career">Career</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Allan_Bradley&action=edit&section=2" title="Edit section: Career"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Following his PhD, Bradley was appointed assistant professor at <a href="/wiki/Baylor_College_of_Medicine" title="Baylor College of Medicine">Baylor College of Medicine</a>, beginning in 1987 where he was also a <a href="/wiki/Searle_Scholars_Program" title="Searle Scholars Program">Searle Scholar</a> in 1988.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Bradley was appointed a <a href="/wiki/Howard_Hughes_Medical_Institute" title="Howard Hughes Medical Institute">Howard Hughes Medical Institute</a> Investigator in 1993 and director of the <a href="/wiki/Wellcome_Trust_Sanger_Institute" class="mw-redirect" title="Wellcome Trust Sanger Institute">Wellcome Trust Sanger Institute</a>, from October 2000 (preceded by <a href="/wiki/John_Sulston" title="John Sulston">John Sulston</a>) to April 2010, succeeded by <a href="/wiki/Michael_Stratton" title="Michael Stratton">Michael Stratton</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Awards_and_honours">Awards and honours</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Allan_Bradley&action=edit&section=3" title="Edit section: Awards and honours"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div><p> Bradley won a 1994 DeBakey Award<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> and was elected a <a href="/wiki/List_of_Fellows_of_the_Royal_Society_elected_in_2002" class="mw-redirect" title="List of Fellows of the Royal Society elected in 2002">Fellow of the Royal Society (FRS) in 2002</a>. His certificate of election reads: <style data-mw-deduplicate="TemplateStyles:r1244412712">.mw-parser-output .templatequote{overflow:hidden;margin:1em 0;padding:0 32px}.mw-parser-output .templatequotecite{line-height:1.5em;text-align:left;margin-top:0}@media(min-width:500px){.mw-parser-output .templatequotecite{padding-left:1.6em}}</style></p><blockquote class="templatequote"><p>"Allan Bradley has made important contributions to the technique of mutation of endogenous genes in mice, an approach that has opened up a new era of research in biology. It is impossible to open an issue of a major journal nowadays without coming across an article that describes the consequences of mutating an endogenous gene in mice. The generation of these mice is based on concepts and techniques that can be traced back to experiments performed and published by Bradley fifteen years ago. In the years since, he has not only used ES cell technology to provide key information on the functions of many genes including several important tumour suppressor genes, but has also continued to improve and develop the techniques, technology, and tools for genetic manipulation in the mouse. Today, mice can be generated with changes as subtle as an alteration in a single nucleotide or as extensive as the deletion of millions of base pairs. These alterations will gain increasing importance in genetic experiments aimed at understanding the function of genes in the mammalian genome in the post genome era."<sup id="cite_ref-royal_10-1" class="reference"><a href="#cite_note-royal-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup></p></blockquote> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Allan_Bradley&action=edit&section=4" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-membo-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-membo_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20150316002020/http://www.embo.org/documents/members/The_EMBO_Pocket_Directory.pdf">"The EMBO Pocket Directory"</a> <span class="cs1-format">(PDF)</span>. European Molecular Biology Organization. Archived from <a rel="nofollow" class="external text" href="http://www.embo.org/documents/members/The_EMBO_Pocket_Directory.pd">the original</a> on 16 March 2015.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=The+EMBO+Pocket+Directory&rft.pub=European+Molecular+Biology+Organization&rft_id=http%3A%2F%2Fwww.embo.org%2Fdocuments%2Fmembers%2FThe_EMBO_Pocket_Directory.pd&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDonehowerHarveySlagleMcArthur1992" class="citation journal cs1">Donehower, L. A.; Harvey, M.; Slagle, B. L.; McArthur, M. J.; Montgomery Jr, C. A.; Butel, J. S.; Bradley, A. (1992). "Mice deficient for p53 are developmentally normal but susceptible to spontaneous tumours". <i>Nature</i>. <b>356</b> (6366): 215–221. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1992Natur.356..215D">1992Natur.356..215D</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F356215a0">10.1038/356215a0</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/1552940">1552940</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:4348340">4348340</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Mice+deficient+for+p53+are+developmentally+normal+but+susceptible+to+spontaneous+tumours&rft.volume=356&rft.issue=6366&rft.pages=215-221&rft.date=1992&rft_id=info%3Adoi%2F10.1038%2F356215a0&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A4348340%23id-name%3DS2CID&rft_id=info%3Apmid%2F1552940&rft_id=info%3Abibcode%2F1992Natur.356..215D&rft.aulast=Donehower&rft.aufirst=L.+A.&rft.au=Harvey%2C+M.&rft.au=Slagle%2C+B.+L.&rft.au=McArthur%2C+M.+J.&rft.au=Montgomery+Jr%2C+C.+A.&rft.au=Butel%2C+J.+S.&rft.au=Bradley%2C+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDaltonPitts-MeekKeshavFigari1993" class="citation journal cs1">Dalton, D.; Pitts-Meek, S.; Keshav, S.; Figari, I.; Bradley, A.; Stewart, T. (1993). "Multiple defects of immune cell function in mice with disrupted interferon-gamma genes". <i>Science</i>. <b>259</b> (5102): 1739–1742. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1993Sci...259.1739D">1993Sci...259.1739D</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.8456300">10.1126/science.8456300</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/8456300">8456300</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Multiple+defects+of+immune+cell+function+in+mice+with+disrupted+interferon-gamma+genes&rft.volume=259&rft.issue=5102&rft.pages=1739-1742&rft.date=1993&rft_id=info%3Apmid%2F8456300&rft_id=info%3Adoi%2F10.1126%2Fscience.8456300&rft_id=info%3Abibcode%2F1993Sci...259.1739D&rft.aulast=Dalton&rft.aufirst=D.&rft.au=Pitts-Meek%2C+S.&rft.au=Keshav%2C+S.&rft.au=Figari%2C+I.&rft.au=Bradley%2C+A.&rft.au=Stewart%2C+T.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGuoWangBradley2004" class="citation journal cs1">Guo, G.; Wang, W.; Bradley, A. (2004). "Mismatch repair genes identified using genetic screens in Blm-deficient embryonic stem cells". <i>Nature</i>. <b>429</b> (6994): 891–895. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2004Natur.429..891G">2004Natur.429..891G</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fnature02653">10.1038/nature02653</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15215866">15215866</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:2545884">2545884</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Mismatch+repair+genes+identified+using+genetic+screens+in+Blm-deficient+embryonic+stem+cells&rft.volume=429&rft.issue=6994&rft.pages=891-895&rft.date=2004&rft_id=info%3Adoi%2F10.1038%2Fnature02653&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A2545884%23id-name%3DS2CID&rft_id=info%3Apmid%2F15215866&rft_id=info%3Abibcode%2F2004Natur.429..891G&rft.aulast=Guo&rft.aufirst=G.&rft.au=Wang%2C+W.&rft.au=Bradley%2C+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKileHentgesClarkNakamura2003" class="citation journal cs1">Kile, B. T.; Hentges, K. E.; Clark, A. T.; Nakamura, H.; Salinger, A. P.; Liu, B.; Box, N.; Stockton, D. W.; Johnson, R. L.; Behringer, R. R.; Bradley, A.; Justice, M. J. (2003). "Functional genetic analysis of mouse chromosome 11". <i>Nature</i>. <b>425</b> (6953): 81–86. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2003Natur.425...81K">2003Natur.425...81K</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fnature01865">10.1038/nature01865</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/12955145">12955145</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:4409237">4409237</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Functional+genetic+analysis+of+mouse+chromosome+11&rft.volume=425&rft.issue=6953&rft.pages=81-86&rft.date=2003&rft_id=info%3Adoi%2F10.1038%2Fnature01865&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A4409237%23id-name%3DS2CID&rft_id=info%3Apmid%2F12955145&rft_id=info%3Abibcode%2F2003Natur.425...81K&rft.aulast=Kile&rft.aufirst=B.+T.&rft.au=Hentges%2C+K.+E.&rft.au=Clark%2C+A.+T.&rft.au=Nakamura%2C+H.&rft.au=Salinger%2C+A.+P.&rft.au=Liu%2C+B.&rft.au=Box%2C+N.&rft.au=Stockton%2C+D.+W.&rft.au=Johnson%2C+R.+L.&rft.au=Behringer%2C+R.+R.&rft.au=Bradley%2C+A.&rft.au=Justice%2C+M.+J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-mice-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-mice_6-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20100401121618/http://www.sanger.ac.uk/research/projects/mousegenomics">"Mouse genomics – Allan Bradley laboratory"</a>. Wellcome Trust Sanger Institute. Archived from <a rel="nofollow" class="external text" href="http://www.sanger.ac.uk/research/projects/mousegenomics">the original</a> on 1 April 2010.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Mouse+genomics+%E2%80%93+Allan+Bradley+laboratory&rft.pub=Wellcome+Trust+Sanger+Institute&rft_id=http%3A%2F%2Fwww.sanger.ac.uk%2Fresearch%2Fprojects%2Fmousegenomics&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-wtsi-7"><span class="mw-cite-backlink">^ <a href="#cite_ref-wtsi_7-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-wtsi_7-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20131113144014/http://www.sanger.ac.uk/research/faculty/abradley/">"Allan Bradley – Wellcome Trust Sanger Institute"</a>. Sanger.ac.uk. Archived from <a rel="nofollow" class="external text" href="http://www.sanger.ac.uk/about/people/biographies/abradley.html">the original</a> on 13 November 2013.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Allan+Bradley+%E2%80%93+Wellcome+Trust+Sanger+Institute&rft.pub=Sanger.ac.uk&rft_id=http%3A%2F%2Fwww.sanger.ac.uk%2Fabout%2Fpeople%2Fbiographies%2Fabradley.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-Watts2007-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-Watts2007_8-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWatts2007" class="citation journal cs1">Watts, Geoff (2007). "Martin Evans: joint winner of 2007 Nobel Prize in medicine". <i>The Lancet</i>. <b>370</b> (9605): 2095. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0140-6736%2807%2961889-3">10.1016/S0140-6736(07)61889-3</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0140-6736">0140-6736</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18156018">18156018</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:7800773">7800773</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Lancet&rft.atitle=Martin+Evans%3A+joint+winner+of+2007+Nobel+Prize+in+medicine&rft.volume=370&rft.issue=9605&rft.pages=2095&rft.date=2007&rft.issn=0140-6736&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A7800773%23id-name%3DS2CID&rft_id=info%3Apmid%2F18156018&rft_id=info%3Adoi%2F10.1016%2FS0140-6736%2807%2961889-3&rft.aulast=Watts&rft.aufirst=Geoff&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-whoswho-9"><span class="mw-cite-backlink">^ <a href="#cite_ref-whoswho_9-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-whoswho_9-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation encyclopaedia cs1"><span class="id-lock-subscription" title="Paid subscription required"><a rel="nofollow" class="external text" href="https://www.ukwhoswho.com/view/article/oupww/whoswho/U43742">"BRADLEY, Prof. Allan"</a></span>. <i><a href="/wiki/Who%27s_Who_(UK)" title="Who's Who (UK)">Who's Who</a></i>. Vol. 2014 (online <a href="/wiki/Oxford_University_Press" title="Oxford University Press">Oxford University Press</a> ed.). A & C Black.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=BRADLEY%2C+Prof.+Allan&rft.btitle=Who%27s+Who&rft.edition=online+Oxford+University+Press&rft.pub=A+%26+C+Black&rft_id=https%3A%2F%2Fwww.ukwhoswho.com%2Fview%2Farticle%2Foupww%2Fwhoswho%2FU43742&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span> <span style="font-size:0.95em; font-size:95%; color: var( --color-subtle, #555 )">(Subscription or <a rel="nofollow" class="external text" href="https://www.ukwhoswho.com/page/subscribe#public">UK public library membership</a> required.)</span></span> </li> <li id="cite_note-royal-10"><span class="mw-cite-backlink">^ <a href="#cite_ref-royal_10-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-royal_10-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20150903180706/https://collections.royalsociety.org/DServe.exe?dsqIni=Dserve.ini&dsqApp=Archive&dsqDb=Catalog&dsqCmd=show.tcl&dsqSearch=(RefNo==%27EC%2F2002%2F01%27)">"Certificate of election EC/2002/01: Allan Bradley"</a>. London: The Royal Society. Archived from <a rel="nofollow" class="external text" href="https://collections.royalsociety.org/DServe.exe?dsqIni=Dserve.ini&dsqApp=Archive&dsqDb=Catalog&dsqCmd=show.tcl&dsqSearch=(RefNo==%27EC%2F2002%2F01%27)">the original</a> on 3 September 2015<span class="reference-accessdate">. Retrieved <span class="nowrap">14 November</span> 2013</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Certificate+of+election+EC%2F2002%2F01%3A+Allan+Bradley&rft.place=London&rft.pub=The+Royal+Society&rft_id=https%3A%2F%2Fcollections.royalsociety.org%2FDServe.exe%3FdsqIni%3DDserve.ini%26dsqApp%3DArchive%26dsqDb%3DCatalog%26dsqCmd%3Dshow.tcl%26dsqSearch%3D%28RefNo%3D%3D%2527EC%252F2002%252F01%2527%29&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20101203010331/http://www.wellcome.ac.uk/News/2010/Features/WTX059873.htm">"Professor Allan Bradley: a decade at Sanger | Wellcome Trust"</a>. Wellcome.ac.uk. 15 June 2010. Archived from <a rel="nofollow" class="external text" href="http://www.wellcome.ac.uk/News/2010/Features/WTX059873.htm">the original</a> on 3 December 2010<span class="reference-accessdate">. Retrieved <span class="nowrap">4 April</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Professor+Allan+Bradley%3A+a+decade+at+Sanger+%26%23124%3B+Wellcome+Trust&rft.pub=Wellcome.ac.uk&rft.date=2010-06-15&rft_id=http%3A%2F%2Fwww.wellcome.ac.uk%2FNews%2F2010%2FFeatures%2FWTX059873.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.sanger.ac.uk/about/press/2002/020513.html">"Wellcome Trust Sanger Institute's Director Honoured: Allan Bradley Elected to Royal Society – Wellcome Trust Sanger Institute"</a>. Sanger.ac.uk<span class="reference-accessdate">. Retrieved <span class="nowrap">4 April</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Wellcome+Trust+Sanger+Institute%27s+Director+Honoured%3A+Allan+Bradley+Elected+to+Royal+Society+%E2%80%93+Wellcome+Trust+Sanger+Institute&rft.pub=Sanger.ac.uk&rft_id=http%3A%2F%2Fwww.sanger.ac.uk%2Fabout%2Fpress%2F2002%2F020513.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-scopus-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-scopus_13-0">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.url?authorId=55605072700">Allan Bradley's publications</a> indexed by the <a href="/wiki/Scopus" title="Scopus">Scopus</a> bibliographic database. <span style="font-size:0.95em; font-size:95%; color: var( --color-subtle, #555 )">(subscription required)</span></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKate_Kelland2011" class="citation news cs1">Kate Kelland (12 October 2011). <a rel="nofollow" class="external text" href="http://news.nationalpost.com/2011/10/12/new-stem-cell-method-could-end-need-for-liver-transplants-scientists/#more-99581">"New stem cell method could end need for liver transplants"</a>. <i>National Post</i>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=National+Post&rft.atitle=New+stem+cell+method+could+end+need+for+liver+transplants&rft.date=2011-10-12&rft.au=Kate+Kelland&rft_id=http%3A%2F%2Fnews.nationalpost.com%2F2011%2F10%2F12%2Fnew-stem-cell-method-could-end-need-for-liver-transplants-scientists%2F%23more-99581&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-bradleyphd-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-bradleyphd_15-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBradley1985" class="citation thesis cs1">Bradley, A. (1985). <a rel="nofollow" class="external text" href="https://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.373263"><i>Isolation characterization and developmental potential of murine embryo-derived stem cells</i></a> (PhD thesis). University of Cambridge.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adissertation&rft.title=Isolation+characterization+and+developmental+potential+of+murine+embryo-derived+stem+cells.&rft.degree=PhD&rft.inst=University+of+Cambridge&rft.date=1985&rft.aulast=Bradley&rft.aufirst=A.&rft_id=http%3A%2F%2Fethos.bl.uk%2FOrderDetails.do%3Fuin%3Duk.bl.ethos.373263&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-16">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.searlescholars.net/person/318">"Searle Scholars Program: Allan Bradley (1988)"</a>. Searlescholars.net<span class="reference-accessdate">. Retrieved <span class="nowrap">4 April</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Searle+Scholars+Program%3A+Allan+Bradley+%281988%29&rft.pub=Searlescholars.net&rft_id=http%3A%2F%2Fwww.searlescholars.net%2Fperson%2F318&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://archive.today/20120719182953/http://www.bcm.edu/debakeyawards/Bradley.html">"Bradley – DeBakey Excellence in Research Awards- Baylor College of Medicine, Houston, Texas"</a>. Bcm.edu. Archived from <a rel="nofollow" class="external text" href="http://www.bcm.edu/debakeyawards/Bradley.html">the original</a> on 19 July 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">4 April</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Bradley+%E2%80%93+DeBakey+Excellence+in+Research+Awards-+Baylor+College+of+Medicine%2C+Houston%2C+Texas&rft.pub=Bcm.edu&rft_id=http%3A%2F%2Fwww.bcm.edu%2Fdebakeyawards%2FBradley.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAllan+Bradley" class="Z3988"></span></span> </li> </ol></div></div> <table class="wikitable succession-box noprint" style="margin:0.5em auto; font-size:small;clear:both;"> <tbody><tr> <th colspan="3" style="border-top: 5px solid lightgreen">Non-profit organization positions </th></tr> <tr style="text-align:center;"> <td style="width:30%;" rowspan="1">Preceded by<div style="font-weight: bold"><a href="/wiki/John_Sulston" title="John Sulston">John Sulston</a></div> </td> <td style="width: 40%; text-align: center;" rowspan="1"><b> <a href="/wiki/Director_(business)" title="Director (business)">Director</a> of the <a href="/wiki/Wellcome_Trust_Sanger_Institute" class="mw-redirect" title="Wellcome Trust Sanger Institute">Wellcome Trust Sanger Institute</a> </b><br />2000–2010 </td> <td style="width: 30%; text-align: center;" rowspan="1">Succeeded by<div style="font-weight: bold"><a href="/wiki/Michael_Stratton" title="Michael Stratton">Michael Stratton</a></div> </td></tr> </tbody></table> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol 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title="Gurdon Institute">Gurdon Institute</a></li> <li>Sainsbury Wellcome Centre for Neural Circuits and Behaviour</li> <li><a href="/wiki/Science_Learning_Centres" title="Science Learning Centres">Science Learning Centres</a></li> <li><a href="/wiki/Wellcome_Trust_Centre_for_Cell-Matrix_Research" title="Wellcome Trust Centre for Cell-Matrix Research">WTC for Cell-Matrix Research</a></li> <li><a href="/wiki/Wellcome_Trust_Centre_for_Gene_Regulation_and_Expression" class="mw-redirect" title="Wellcome Trust Centre for Gene Regulation and Expression">WTC for Gene Regulation and Expression</a></li> <li><a href="/wiki/Wellcome_Trust_Centre_for_Human_Genetics" class="mw-redirect" title="Wellcome Trust Centre for Human Genetics">WTC for Human Genetics</a></li> <li>WTC for Mitochondrial Research</li> <li>WTC for Molecular Parasitology</li> <li><a href="/wiki/Wellcome_Trust_Centre_for_Neuroimaging" class="mw-redirect" title="Wellcome Trust Centre for Neuroimaging">WTC for Neuroimaging</a></li> <li><a href="/wiki/Wellcome_Trust_Centre_for_Stem_Cell_Research" class="mw-redirect" title="Wellcome Trust Centre for Stem Cell Research">WTC for Stem Cell Research</a></li> <li><a href="/wiki/Wellcome_Sanger_Institute" title="Wellcome Sanger Institute">Wellcome Sanger Institute</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:9em;text-align: left;background-color: #eee;">Former</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Wellcome_Trust_Centre_for_the_History_of_Medicine" title="Wellcome Trust Centre for the History of Medicine">Wellcome Trust Centre for the History of Medicine</a></li> <li><a href="/wiki/Wellcome_Research_Laboratories" title="Wellcome Research Laboratories">Wellcome Research Laboratories</a></li></ul> </div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="7" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Wellcome.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/83/Wellcome.jpg/180px-Wellcome.jpg" decoding="async" width="180" height="135" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/83/Wellcome.jpg/270px-Wellcome.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/83/Wellcome.jpg/360px-Wellcome.jpg 2x" data-file-width="2048" data-file-height="1536" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;">Projects and facilities</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/1000_Genomes_Project" title="1000 Genomes Project">1000 Genomes Project</a></li> <li><i><a href="/wiki/Big_Picture_(magazine)" title="Big Picture (magazine)">Big Picture</a></i></li> <li><a href="/wiki/Cambridge_Biomedical_Campus" title="Cambridge Biomedical Campus">Cambridge Biomedical Campus</a></li> <li><a href="/wiki/Cancer_Genome_Project" title="Cancer Genome Project">Cancer Genome Project</a></li> <li><a href="/wiki/ChEMBL" title="ChEMBL">ChEMBL</a></li> <li><a href="/wiki/Coalition_for_Epidemic_Preparedness_Innovations" title="Coalition for Epidemic Preparedness Innovations">Coalition for Epidemic Preparedness Innovations</a></li> <li><a href="/wiki/COSMIC_cancer_database" title="COSMIC cancer database">COSMIC cancer database</a></li> <li><a href="/wiki/DECIPHER" title="DECIPHER">DECIPHER</a></li> <li><a href="/wiki/Diamond_Light_Source" title="Diamond Light Source">Diamond Light Source</a></li> <li><i><a href="/wiki/ELife" title="ELife">eLife</a></i></li> <li><a href="/wiki/Ensembl" class="mw-redirect" title="Ensembl">Ensembl</a></li> <li><a href="/wiki/Farmcare" title="Farmcare">Farmcare</a></li> <li><a href="/wiki/Genome_Reference_Consortium" title="Genome Reference Consortium">Genome Reference Consortium</a></li> <li><a href="/wiki/Human_Genome_Project" title="Human Genome Project">Human Genome Project</a></li> <li><a href="/w/index.php?title=Malawi_Liverpool_Wellcome_Trust&action=edit&redlink=1" class="new" title="Malawi Liverpool Wellcome Trust (page does not exist)">Malawi Liverpool Wellcome Trust</a></li> <li><a href="/wiki/MEROPS" title="MEROPS">MEROPS</a></li> <li><a href="/wiki/Pfam" title="Pfam">Pfam</a></li> <li><a href="/wiki/Rfam" title="Rfam">Rfam</a></li> <li><a href="/wiki/UK_Biobank" title="UK Biobank">UK Biobank</a></li> <li><a href="/wiki/Wellcome_Collection" title="Wellcome Collection">Wellcome Collection</a></li> <li><a href="/wiki/Wellcome_Genome_Campus" title="Wellcome Genome Campus">Wellcome Genome Campus</a></li> <li><a href="/wiki/Wellcome_Library" title="Wellcome Library">Wellcome Library</a></li> <li><a href="/wiki/WormBase" title="WormBase">WormBase</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;">Board of governors</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Eliza_Manningham-Buller" title="Eliza Manningham-Buller">Eliza Manningham-Buller</a></li> <li><a href="/wiki/Michael_Ferguson_(biochemist)" title="Michael Ferguson (biochemist)">Michael Ferguson</a></li> <li><a href="/wiki/Tobias_Bonhoeffer" title="Tobias Bonhoeffer">Tobias Bonhoeffer</a></li> <li>Alan Brown</li> <li><a href="https://www.wikidata.org/wiki/Q24716612" class="extiw" title="d:Q24716612">William Burns</a></li> <li><a href="/wiki/Bryan_Grenfell" title="Bryan Grenfell">Bryan Grenfell</a></li> <li><a href="/wiki/Naguib_Kheraj" title="Naguib Kheraj">Naguib Kheraj</a></li> <li><a href="/wiki/Fiona_Powrie" title="Fiona Powrie">Fiona Powrie</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;">Executive leadership team</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Jeremy_Farrar" title="Jeremy Farrar">Jeremy Farrar</a></li> <li>Chris Bird</li> <li>Stephen Caddick</li> <li>Simon Chaplin</li> <li>Alyson Fox</li> <li>Peter Pereira Gray</li> <li>Mark Henderson</li> <li>Chonnettia Jones</li> <li>Tim Livett</li> <li>Nick Moakes</li> <li>Kathy Poole</li> <li>Jim Smith</li> <li>James Thomas</li> <li>Ed Whiting</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;">Former directors</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Peter_Williams_(physician)" title="Peter Williams (physician)">Peter Williams</a> (1965–1991)</li> <li><a href="/wiki/Bridget_Ogilvie" title="Bridget Ogilvie">Bridget Ogilvie</a> (1991–1998)</li> <li><a href="/wiki/Michael_Dexter" title="Michael Dexter">Michael Dexter</a> (1998–2008)</li> <li><a href="/wiki/Mark_Walport" title="Mark Walport">Mark Walport</a> (2003–2013)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;">Other key people</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Damon_Buffini" title="Damon Buffini">Damon Buffini</a></li> <li><a href="/wiki/William_Castell" title="William Castell">William Castell</a></li> <li><a href="/wiki/Dominic_Cadbury" title="Dominic Cadbury">Dominic Cadbury</a></li> <li><a href="/wiki/Harold_Cook_(medical_historian)" class="mw-redirect" title="Harold Cook (medical historian)">Harold Cook</a></li> <li><a href="/wiki/Kay_Davies" title="Kay Davies">Kay Davies</a></li> <li><a href="/wiki/Oliver_Franks,_Baron_Franks" title="Oliver Franks, Baron Franks">Oliver Franks</a></li> <li><a href="/wiki/Roger_Gibbs" title="Roger Gibbs">Roger Gibbs</a></li> <li><a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a></li> <li><a href="/wiki/Richard_Hynes" title="Richard Hynes">Richard Hynes</a></li> <li><a href="/wiki/Anne_Mandall_Johnson" title="Anne Mandall Johnson">Anne Johnson</a></li> <li><a href="/wiki/Roy_Porter" title="Roy Porter">Roy Porter</a></li> <li><a href="https://www.wikidata.org/wiki/Q21165217" class="extiw" title="d:Q21165217">Peter Rigby</a></li> <li><a href="/wiki/David_Steel_(businessman)" title="David Steel (businessman)">David Steel</a></li> <li><a href="/wiki/David_Ian_Stuart" class="mw-redirect" title="David Ian Stuart">David Stuart</a></li> <li><a href="/wiki/John_Sulston" title="John Sulston">John Sulston</a></li> <li><a href="/wiki/Henry_Wellcome" title="Henry Wellcome">Henry Wellcome</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;">Awards and fellowships</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Capital Awards</li> <li>Collaborative Awards in Science</li> <li>Investigator Awards in Science</li> <li>Institutional Strategic Support Fund</li> <li>Science Strategic Award</li> <li>Sir Henry Dale <a href="/wiki/Fellow" title="Fellow">Fellowship</a></li> <li>Sir Henry Wellcome <a href="/wiki/Postdoctoral_researcher" title="Postdoctoral researcher">Postdoctoral</a> Fellowship</li> <li><a href="/wiki/Wellcome_Book_Prize" title="Wellcome Book Prize">Wellcome Book Prize</a></li> <li>Wellcome Image Awards</li> <li>Wellcome Trust Centre</li> <li><a href="/wiki/Wellcome_Trust_Principal_Research_Fellow" title="Wellcome Trust Principal Research Fellow">Wellcome Trust Principal Research Fellow</a></li> <li>Wellcome Trust Senior Research Fellow</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <b><a href="/wiki/Category:Wellcome_Trust" title="Category:Wellcome Trust">Category</a></b></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Fellows_of_the_Royal_Society_elected_in_2002" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:FRS_2002" title="Template:FRS 2002"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:FRS_2002" title="Template talk:FRS 2002"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:FRS_2002" title="Special:EditPage/Template:FRS 2002"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Fellows_of_the_Royal_Society_elected_in_2002" style="font-size:114%;margin:0 4em"><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">Fellows</a> of the <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a> elected <a href="/wiki/List_of_fellows_of_the_Royal_Society_elected_in_2002" title="List of fellows of the Royal Society elected in 2002">in 2002</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Fellows</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Allan Bradley</a></li> <li><a href="/wiki/Robin_Carrell" title="Robin Carrell">Robin Carrell</a></li> <li><a href="/wiki/Mick_Crawley" title="Mick Crawley">Michael John Crawley</a></li> <li><a href="/wiki/Stuart_Cull-Candy" title="Stuart Cull-Candy">Stuart Cull-Candy</a></li> <li><a href="/wiki/John_Dainton" title="John Dainton">John Dainton</a></li> <li><i><a href="https://www.wikidata.org/wiki/Q21165803" class="extiw" title="d:Q21165803">Roger John Davis</a></i></li> <li><a href="/wiki/Anne_Dell" title="Anne Dell">Anne Dell</a></li> <li><a href="/wiki/David_Dolphin" title="David Dolphin">David Henry Dolphin</a></li> <li><a href="/wiki/David_Fowler_(physicist)" title="David Fowler (physicist)">David Fowler</a></li> <li><a href="/wiki/Steve_Furber" title="Steve Furber">Steve Furber</a></li> <li><a href="/wiki/Graham_Goodwin" title="Graham Goodwin">Graham Goodwin</a></li> <li><a href="/wiki/Jean-Pierre_Hansen" title="Jean-Pierre Hansen">Jean-Pierre Hansen</a></li> <li><a href="/wiki/Nicholas_Hastie" title="Nicholas Hastie">Nicholas Hastie</a></li> <li><a href="/wiki/Christopher_Hawkesworth" title="Christopher Hawkesworth">Christopher Hawkesworth</a></li> <li><a href="/wiki/Judith_Howard" title="Judith Howard">Judith Howard</a></li> <li><a href="/wiki/Philip_Ingham" title="Philip Ingham">Philip Ingham</a></li> <li><a href="/wiki/David_Ish-Horowicz" title="David Ish-Horowicz">David Ish-Horowicz</a></li> <li><a href="/wiki/James_A._Jackson" title="James A. Jackson">James A. Jackson</a></li> <li><i><a href="https://www.wikidata.org/wiki/Q21165436" class="extiw" title="d:Q21165436">Bruce Ernest Kemp</a></i></li> <li><i><a href="https://www.wikidata.org/wiki/Q21166265" class="extiw" title="d:Q21166265">John Vincent Kilmartin</a></i></li> <li><i><a href="https://www.wikidata.org/wiki/Q21165856" class="extiw" title="d:Q21165856">David Malcolm James Lilley</a></i></li> <li><a href="/wiki/Terry_Lyons_(mathematician)" title="Terry Lyons (mathematician)">Terry Lyons</a></li> <li><a href="/wiki/Georgina_Mace" title="Georgina Mace">Georgina Mace</a></li> <li><a href="/wiki/John_McCanny" title="John McCanny">John McCanny</a></li> <li><i><a href="https://www.wikidata.org/wiki/Q21165418" class="extiw" title="d:Q21165418">Brian Cecil Joseph Moore</a></i></li> <li><a href="/wiki/David_Parker_(chemist)" title="David Parker (chemist)">David Parker</a></li> <li><a href="/wiki/Martyn_Poliakoff" title="Martyn Poliakoff">Martyn Poliakoff</a></li> <li><a href="/wiki/Eric_Priest" title="Eric Priest">Eric Priest</a></li> <li><a href="/wiki/Terence_Quinn" title="Terence Quinn">Terence Quinn</a></li> <li><a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a></li> <li><a href="/wiki/Mary_Rees" title="Mary Rees">Mary Rees</a></li> <li><a href="/wiki/Miles_Reid" title="Miles Reid">Miles Reid</a></li> <li><a href="/wiki/David_William_Rhind" title="David William Rhind">David William Rhind</a></li> <li><a href="/wiki/Thomas_Maurice_Rice" title="Thomas Maurice Rice">Thomas Maurice Rice</a></li> <li><a href="/wiki/Roy_Sambles" title="Roy Sambles">Roy Sambles</a></li> <li><a href="/wiki/Peter_Sarnak" title="Peter Sarnak">Peter Sarnak</a></li> <li><a href="/wiki/Tony_Sinclair_(biologist)" title="Tony Sinclair (biologist)">Tony Sinclair</a></li> <li><i><a href="https://www.wikidata.org/wiki/Q507502" class="extiw" title="d:Q507502">Andrew Benjamin Smith</a></i></li> <li><i><a href="https://www.wikidata.org/wiki/Q21165231" class="extiw" title="d:Q21165231">Anthony John Stace</a></i></li> <li><a href="/wiki/Nicholas_Strausfeld" title="Nicholas Strausfeld">Nicholas Strausfeld</a></li> <li><a href="/wiki/Mark_Welland" title="Mark Welland">Mark Welland</a></li> <li><a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Ian Wilmut</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Foreign</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Claude_All%C3%A8gre" title="Claude Allègre">Claude Allègre</a></li> <li><a href="/wiki/Per_Andersen" title="Per Andersen">Per Andersen</a></li> <li><a href="/wiki/Hubert_Markl" title="Hubert Markl">Hubert Markl</a></li> <li><a href="/wiki/Alexander_Pines" title="Alexander Pines">Alexander Pines</a></li> <li><a href="/wiki/Peter_H._Raven" title="Peter H. Raven">Peter H. Raven</a></li> <li><a href="/wiki/Carl_Wunsch" title="Carl Wunsch">Carl Wunsch</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/List_of_fellows_of_the_Royal_Society" title="List of fellows of the Royal Society">List of fellows of the Royal Society</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q4730491#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q4730491#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q4730491#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">International</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/41467379">VIAF</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/no98130852">United States</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Academics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://orcid.org/0000-0002-2349-8839">ORCID</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.uri?authorId=57098086000">Scopus</a></span></li></ul></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; 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