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class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Inhoud</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">naar zijbalk verplaatsen</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">verbergen</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">Top</div> </a> </li> <li id="toc-Levensloop" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Levensloop"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Levensloop</span> </div> </a> <ul id="toc-Levensloop-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Werk" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Werk"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Werk</span> </div> </a> <ul id="toc-Werk-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Boeken_van_Sidney_Brenner" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Boeken_van_Sidney_Brenner"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Boeken van Sidney Brenner</span> </div> </a> <ul id="toc-Boeken_van_Sidney_Brenner-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Externe_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Externe_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Externe links</span> </div> </a> <ul id="toc-Externe_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Inhoud" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Inhoudsopgave omschakelen" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Inhoudsopgave omschakelen</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Sydney Brenner</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Ga naar een artikel in een andere taal. Beschikbaar in 63 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-63" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">63 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Afrikaans" lang="af" hreflang="af" data-title="Sydney Brenner" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-an mw-list-item"><a href="https://an.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Aragonees" lang="an" hreflang="an" data-title="Sydney Brenner" data-language-autonym="Aragonés" data-language-local-name="Aragonees" class="interlanguage-link-target"><span>Aragonés</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B3%D9%8A%D8%AF%D9%86%D9%8A_%D8%A8%D8%B1%D9%8A%D9%86%D8%B1" title="سيدني برينر – Arabisch" lang="ar" hreflang="ar" data-title="سيدني برينر" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%B3%D9%8A%D8%AF%D9%86%D9%89_%D8%A8%D8%B1%D9%8A%D9%86%D8%B1" title="سيدنى برينر – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="سيدنى برينر" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Sidney_Brenner" title="Sidney Brenner – Azerbeidzjaans" lang="az" hreflang="az" data-title="Sidney Brenner" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbeidzjaans" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%B3%DB%8C%D8%AF%D9%86%DB%8C_%D8%A8%D8%B1%D9%86%D8%B1" title="سیدنی برنر – South Azerbaijani" lang="azb" hreflang="azb" data-title="سیدنی برنر" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D1%96%D0%B4%D0%BD%D0%B5%D0%B9_%D0%91%D1%80%D1%8D%D0%BD%D0%B5%D1%80" title="Сідней Брэнер – Belarussisch" lang="be" hreflang="be" data-title="Сідней Брэнер" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A1%D0%B8%D0%B4%D0%BD%D0%B8_%D0%91%D1%80%D0%B5%D0%BD%D1%8A%D1%80" title="Сидни Бренър – Bulgaars" lang="bg" hreflang="bg" data-title="Сидни Бренър" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B8%E0%A6%BF%E0%A6%A1%E0%A6%A8%E0%A6%BF_%E0%A6%AC%E0%A7%8D%E0%A6%B0%E0%A7%87%E0%A6%A8%E0%A6%BE%E0%A6%B0" title="সিডনি ব্রেনার – Bengaals" lang="bn" hreflang="bn" data-title="সিডনি ব্রেনার" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Catalaans" lang="ca" hreflang="ca" data-title="Sydney Brenner" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Tsjechisch" lang="cs" hreflang="cs" data-title="Sydney Brenner" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Deens" lang="da" hreflang="da" data-title="Sydney Brenner" data-language-autonym="Dansk" data-language-local-name="Deens" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Duits" lang="de" hreflang="de" data-title="Sydney Brenner" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Engels" lang="en" hreflang="en" data-title="Sydney Brenner" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Esperanto" lang="eo" hreflang="eo" data-title="Sydney Brenner" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Spaans" lang="es" hreflang="es" data-title="Sydney Brenner" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Estisch" lang="et" hreflang="et" data-title="Sydney Brenner" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Baskisch" lang="eu" hreflang="eu" data-title="Sydney Brenner" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%DB%8C%D8%AF%D9%86%DB%8C_%D8%A8%D8%B1%D9%86%D8%B1" title="سیدنی برنر – Perzisch" lang="fa" hreflang="fa" data-title="سیدنی برنر" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Fins" lang="fi" hreflang="fi" data-title="Sydney Brenner" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Frans" lang="fr" hreflang="fr" data-title="Sydney Brenner" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Iers" lang="ga" hreflang="ga" data-title="Sydney Brenner" data-language-autonym="Gaeilge" data-language-local-name="Iers" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Galicisch" lang="gl" hreflang="gl" data-title="Sydney Brenner" data-language-autonym="Galego" data-language-local-name="Galicisch" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ha mw-list-item"><a href="https://ha.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Hausa" lang="ha" hreflang="ha" data-title="Sydney Brenner" data-language-autonym="Hausa" data-language-local-name="Hausa" class="interlanguage-link-target"><span>Hausa</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A1%D7%99%D7%93%D7%A0%D7%99_%D7%91%D7%A8%D7%A0%D7%A8" title="סידני ברנר – Hebreeuws" lang="he" hreflang="he" data-title="סידני ברנר" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B8%E0%A4%BF%E0%A4%A1%E0%A4%A8%E0%A5%80_%E0%A4%AC%E0%A5%8D%E0%A4%B0%E0%A5%87%E0%A4%A8%E0%A4%B0" title="सिडनी ब्रेनर – Hindi" lang="hi" hreflang="hi" data-title="सिडनी ब्रेनर" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Kroatisch" lang="hr" hreflang="hr" data-title="Sydney Brenner" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Hongaars" lang="hu" hreflang="hu" data-title="Sydney Brenner" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Indonesisch" lang="id" hreflang="id" data-title="Sydney Brenner" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Ido" lang="io" hreflang="io" data-title="Sydney Brenner" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Italiaans" lang="it" hreflang="it" data-title="Sydney Brenner" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B7%E3%83%89%E3%83%8B%E3%83%BC%E3%83%BB%E3%83%96%E3%83%AC%E3%83%8A%E3%83%BC" title="シドニー・ブレナー – Japans" lang="ja" hreflang="ja" data-title="シドニー・ブレナー" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A1%D0%B8%D0%B4%D0%BD%D0%B5%D0%B9_%D0%91%D1%80%D0%B5%D0%BD%D0%BD%D0%B5%D1%80" title="Сидней Бреннер – Kazachs" lang="kk" hreflang="kk" data-title="Сидней Бреннер" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%8B%9C%EB%93%9C%EB%8B%88_%EB%B8%8C%EB%A0%88%EB%84%88" title="시드니 브레너 – Koreaans" lang="ko" hreflang="ko" data-title="시드니 브레너" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Latijn" lang="la" hreflang="la" data-title="Sydney Brenner" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Sidnijs_Breners" title="Sidnijs Breners – Lets" lang="lv" hreflang="lv" data-title="Sidnijs Breners" data-language-autonym="Latviešu" data-language-local-name="Lets" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%B8%E0%B4%BF%E0%B4%A1%E0%B5%8D%E0%B4%A8%E0%B4%BF_%E0%B4%AC%E0%B5%8D%E0%B4%B0%E0%B5%86%E0%B4%A8%E0%B5%8D%E0%B4%A8%E0%B5%BC" title="സിഡ്നി ബ്രെന്നർ – Malayalam" lang="ml" hreflang="ml" data-title="സിഡ്നി ബ്രെന്നർ" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Maleis" lang="ms" hreflang="ms" data-title="Sydney Brenner" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%B3%DB%8C%D8%AF%D9%86%DB%8C_%D8%A8%D8%B1%D9%86%D8%B1" title="سیدنی برنر – Mazanderani" lang="mzn" hreflang="mzn" data-title="سیدنی برنر" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Nedersaksisch" lang="nds" hreflang="nds" data-title="Sydney Brenner" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Noors - Bokmål" lang="nb" hreflang="nb" data-title="Sydney Brenner" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Occitaans" lang="oc" hreflang="oc" data-title="Sydney Brenner" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Pools" lang="pl" hreflang="pl" data-title="Sydney Brenner" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%B3%DA%88%D9%86%DB%8C_%D8%A8%D8%B1%DB%8C%D9%86%D8%B1" title="سڈنی برینر – Western Punjabi" lang="pnb" hreflang="pnb" data-title="سڈنی برینر" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Portugees" lang="pt" hreflang="pt" data-title="Sydney Brenner" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Roemeens" lang="ro" hreflang="ro" data-title="Sydney Brenner" data-language-autonym="Română" data-language-local-name="Roemeens" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%91%D1%80%D0%B5%D0%BD%D0%BD%D0%B5%D1%80,_%D0%A1%D0%B8%D0%B4%D0%BD%D0%B5%D0%B9" title="Бреннер, Сидней – Russisch" lang="ru" hreflang="ru" data-title="Бреннер, Сидней" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Simple English" lang="en-simple" hreflang="en-simple" data-title="Sydney Brenner" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Sloveens" lang="sl" hreflang="sl" data-title="Sydney Brenner" data-language-autonym="Slovenščina" data-language-local-name="Sloveens" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Albanees" lang="sq" hreflang="sq" data-title="Sydney Brenner" data-language-autonym="Shqip" data-language-local-name="Albanees" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Sidni_Brener" title="Sidni Brener – Servisch" lang="sr" hreflang="sr" data-title="Sidni Brener" data-language-autonym="Српски / srpski" data-language-local-name="Servisch" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Zweeds" lang="sv" hreflang="sv" data-title="Sydney Brenner" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Swahili" lang="sw" hreflang="sw" data-title="Sydney Brenner" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Turks" lang="tr" hreflang="tr" data-title="Sydney Brenner" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%A1%D0%B8%D0%B4%D0%BD%D0%B8_%D0%91%D1%80%D0%B5%D0%BD%D0%BD%D0%B5%D1%80" title="Сидни Бреннер – Tataars" lang="tt" hreflang="tt" data-title="Сидни Бреннер" data-language-autonym="Татарча / tatarça" data-language-local-name="Tataars" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A1%D1%96%D0%B4%D0%BD%D1%96_%D0%91%D1%80%D0%B5%D0%BD%D0%BD%D0%B5%D1%80" title="Сідні Бреннер – Oekraïens" lang="uk" hreflang="uk" data-title="Сідні Бреннер" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%B3%DA%88%D9%86%DB%8C_%D8%A8%D8%B1%DB%8C%D9%86%D8%B1" title="سڈنی برینر – Urdu" lang="ur" hreflang="ur" data-title="سڈنی برینر" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Vietnamees" lang="vi" hreflang="vi" data-title="Sydney Brenner" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E8%A5%BF%E5%BE%B7%E5%B0%BC%C2%B7%E5%B8%83%E4%BC%A6%E7%BA%B3" title="西德尼·布伦纳 – Wuyu" lang="wuu" hreflang="wuu" data-title="西德尼·布伦纳" data-language-autonym="吴语" data-language-local-name="Wuyu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%A1%E1%83%98%E1%83%93%E1%83%9C%E1%83%98_%E1%83%91%E1%83%A0%E1%83%94%E1%83%9C%E1%83%94%E1%83%A0%E1%83%98" title="სიდნი ბრენერი – Mingreels" lang="xmf" hreflang="xmf" data-title="სიდნი ბრენერი" data-language-autonym="მარგალური" data-language-local-name="Mingreels" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Sydney_Brenner" title="Sydney Brenner – Yoruba" lang="yo" hreflang="yo" data-title="Sydney Brenner" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E8%A5%BF%E5%BE%B7%E5%B0%BC%C2%B7%E5%B8%83%E4%BC%A6%E7%BA%B3" title="西德尼·布伦纳 – Chinees" lang="zh" hreflang="zh" data-title="西德尼·布伦纳" data-language-autonym="中文" data-language-local-name="Chinees" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E9%9B%AA%E6%A2%A8%E5%B8%83%E5%80%AB%E7%B4%8D" title="雪梨布倫納 – Kantonees" lang="yue" hreflang="yue" data-title="雪梨布倫納" data-language-autonym="粵語" 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title="Lijst met alle pagina's die naar deze pagina verwijzen [j]" accesskey="j"><span>Links naar deze pagina</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Speciaal:RecenteWijzigingenGelinkt/Sydney_Brenner" rel="nofollow" title="Recente wijzigingen in pagina's waar deze pagina naar verwijst [k]" accesskey="k"><span>Gerelateerde wijzigingen</span></a></li><li id="t-upload" class="mw-list-item"><a href="//commons.wikimedia.org/wiki/Special:UploadWizard?uselang=nl" title="Bestanden uploaden [u]" accesskey="u"><span>Bestand uploaden</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Speciaal:SpecialePaginas" title="Lijst met alle speciale pagina's [q]" accesskey="q"><span>Speciale pagina's</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Sydney_Brenner&oldid=68015129" title="Permanente koppeling naar deze versie van deze pagina"><span>Permanente koppeling</span></a></li><li id="t-info" 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data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Uiterlijk</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">naar zijbalk verplaatsen</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">verbergen</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">Uit Wikipedia, de vrije encyclopedie</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="nl" dir="ltr"><table class="infobox" cellpadding="1" cellspacing="1"> <tbody><tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center; font-size:120%;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Nobel_prize_medal.svg" class="mw-file-description" title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  Sydney Brenner </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/13_januari" title="13 januari">13 januari</a> <a href="/wiki/1927" title="1927">1927</a> – <a href="/wiki/5_april" title="5 april">5 april</a> <a href="/wiki/2019" title="2019">2019</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Sydney_Brenner_OIST_2008_(33208371153)_(cropped).jpg" class="mw-file-description"><img alt="Sydney Brenner in 2008" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/61/Sydney_Brenner_OIST_2008_%2833208371153%29_%28cropped%29.jpg/266px-Sydney_Brenner_OIST_2008_%2833208371153%29_%28cropped%29.jpg" decoding="async" width="266" height="342" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/61/Sydney_Brenner_OIST_2008_%2833208371153%29_%28cropped%29.jpg/399px-Sydney_Brenner_OIST_2008_%2833208371153%29_%28cropped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/61/Sydney_Brenner_OIST_2008_%2833208371153%29_%28cropped%29.jpg/532px-Sydney_Brenner_OIST_2008_%2833208371153%29_%28cropped%29.jpg 2x" data-file-width="1387" data-file-height="1783" /></a></span> </td></tr> <tr> <td style="text-align:center;" colspan="3"><div style="font-size:90%">Sydney Brenner in 2008</div> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Zuid-Afrika" title="Zuid-Afrika">Zuid-Afrika</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/Germiston" title="Germiston">Germiston</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2">Fysiologie of Geneeskunde </td></tr> <tr> <th>Jaar </th> <td colspan="2">2002 </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor hun ontdekkingen betreffende de genetische regulatie van de orgaanontwikkeling en de <a href="/wiki/Apoptose" title="Apoptose">geprogrammeerde celdood</a> </td></tr> <tr> <th>Samen met </th> <td colspan="2"><a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a><br /><a href="/wiki/John_Sulston" class="mw-redirect" title="John Sulston">John Sulston</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a><br /><a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a><br /><a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a><br /><a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>Sydney Brenner</b>, <a href="/wiki/Orde_van_de_Eregezellen_(Verenigd_Koninkrijk)" title="Orde van de Eregezellen (Verenigd Koninkrijk)">CH</a> <a href="/wiki/Royal_Society" title="Royal Society">FRS</a> (<a href="/wiki/Germiston" title="Germiston">Germiston</a>, <a href="/wiki/13_januari" title="13 januari">13 januari</a> <a href="/wiki/1927" title="1927">1927</a> – <a href="/wiki/Singapore" title="Singapore">Singapore</a>, <a href="/wiki/5_april" title="5 april">5 april</a> <a href="/wiki/2019" title="2019">2019</a>) was een Zuid-Afrikaans <a href="/wiki/Bioloog" class="mw-redirect" title="Bioloog">bioloog</a> en <a href="/wiki/Nobelprijs" title="Nobelprijs">Nobelprijswinnaar</a>. In 2002 won hij samen met <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> en <a href="/wiki/John_Sulston" class="mw-redirect" title="John Sulston">John Sulston</a> de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a> voor hun ontdekkingen betreffende de genetische regulatie van de orgaanontwikkeling en de <a href="/wiki/Apoptose" title="Apoptose">geprogrammeerde celdood</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Levensloop">Levensloop</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Sydney_Brenner&veaction=edit&section=1" title="Bewerk dit kopje: Levensloop" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Sydney_Brenner&action=edit&section=1" title="De broncode bewerken van de sectie: Levensloop"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Brenner werd in Zuid-Afrika geboren als zoon van <a href="/wiki/Joden" title="Joden">Joodse</a> immigranten. Zijn vader Morris Brenner kwam in 1910 uit <a href="/wiki/Litouwen" title="Litouwen">Litouwen</a>, en zijn moeder Lena Blacher in 1922 uit <a href="/wiki/Letland" title="Letland">Letland</a>.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Brenner ging naar de Germiston High School en de <a href="/wiki/Universiteit_van_de_Witwatersrand" title="Universiteit van de Witwatersrand">Universiteit van de Witwatersrand</a>. Vervolgens voltooide hij zijn <a href="/wiki/Doctor_of_Philosophy" title="Doctor of Philosophy">PhD</a> aan het <a href="/w/index.php?title=Exeter_College&action=edit&redlink=1" class="new" title="Exeter College (de pagina bestaat niet)">Exeter College</a> in Oxford. Samen met <a href="/wiki/Dorothy_Hodgkin" class="mw-redirect" title="Dorothy Hodgkin">Dorothy Hodgkin</a>, <a href="/w/index.php?title=Leslie_Orgel&action=edit&redlink=1" class="new" title="Leslie Orgel (de pagina bestaat niet)">Leslie Orgel</a>, Jack Dunitz en Beryl Oughton was hij in 1953 een van de eersten die het model van de structuur van <a href="/wiki/Desoxyribonucle%C3%AFnezuur" class="mw-redirect" title="Desoxyribonucleïnezuur">DNA</a> te zien kregen, gemaakt door <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> en <a href="/wiki/James_Watson" title="James Watson">James Watson</a>. Op dat moment was hij werkzaam bij de faculteit scheikunde van de <a href="/wiki/Universiteit_van_Oxford" title="Universiteit van Oxford">Universiteit van Oxford</a>. Brenner werkte later met Crick in het <a href="/wiki/Cavendish-laboratorium" title="Cavendish-laboratorium">Cavendish-laboratorium</a> en het Laboratory of Molecular Biology in <a href="/wiki/Cambridge_(Verenigd_Koninkrijk)" title="Cambridge (Verenigd Koninkrijk)">Cambridge</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Sydney_Brenner&veaction=edit&section=2" title="Bewerk dit kopje: Werk" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Sydney_Brenner&action=edit&section=2" title="De broncode bewerken van de sectie: Werk"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Brenner deed in de jaren 60 fundamentele ontdekkingen op het gebied van <a href="/wiki/Moleculaire_biologie" title="Moleculaire biologie">moleculaire biologie</a>, die toen in opkomst was. Zo was hij betrokken bij de identificatie van <a href="/wiki/Messenger_RNA" class="mw-redirect" title="Messenger RNA">messenger RNA</a> (mRNA), en ontdekte hij met Crick in 1961 <a href="/wiki/Frameshiftmutatie" title="Frameshiftmutatie">frameshiftmutaties</a>. Dit onderzoek gaf al deels inzicht in de <a href="/wiki/Genetische_code" title="Genetische code">genetische code</a>. Brenner focuste zich vervolgens op het gebruik van <i><a href="/wiki/Caenorhabditis_elegans" title="Caenorhabditis elegans">Caenorhabditis elegans</a></i> als een <a href="/wiki/Modelorganisme" title="Modelorganisme">modelorganisme</a> voor het onderzoek van <a href="/wiki/Ontwikkelingsbiologie" title="Ontwikkelingsbiologie">ontwikkeling</a> bij dieren. Brenner koos dit 1 millimeter lange, doorzichtige <a href="/wiki/Rondwormen" title="Rondwormen">rondwormpje</a> vooral omdat hij erg geschikt was voor genetische analyses, en snel in grote hoeveelheden gekweekt kon worden. De titel van zijn Nobelspeech, "Nature's Gift to Science", is een verwijzing naar dit gegeven.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>Brenner richtte in 1996 het <a href="/w/index.php?title=Molecular_Sciences_Institute&action=edit&redlink=1" class="new" title="Molecular Sciences Institute (de pagina bestaat niet)">Molecular Sciences Institute</a> op in <a href="/wiki/Berkeley_(Californi%C3%AB)" title="Berkeley (Californië)">Berkeley</a> en werd in 2000 <i>Distinguished Research Professor</i> aan het <a href="/w/index.php?title=Salk_Institute_for_Biological_Studies&action=edit&redlink=1" class="new" title="Salk Institute for Biological Studies (de pagina bestaat niet)">Salk Institute for Biological Studies</a>. Hij verbond zich aan het <a href="/w/index.php?title=Institute_of_Molecular_and_Cell_Biology&action=edit&redlink=1" class="new" title="Institute of Molecular and Cell Biology (de pagina bestaat niet)">Institute of Molecular and Cell Biology</a> in Singapore, het Okinawa Institute of Science and Technology, de <a href="/w/index.php?title=Janelia_Research_Campus&action=edit&redlink=1" class="new" title="Janelia Research Campus (de pagina bestaat niet)">Janelia Research Campus</a> en het <a href="/w/index.php?title=Howard_Hughes_Medical_Institute&action=edit&redlink=1" class="new" title="Howard Hughes Medical Institute (de pagina bestaat niet)">Howard Hughes Medical Institute</a>. Hij was lid van de Raad van Wetenschappelijk Bestuur van het <a href="/w/index.php?title=The_Scripps_Research_Institute&action=edit&redlink=1" class="new" title="The Scripps Research Institute (de pagina bestaat niet)">The Scripps Research Institute</a>. Jarenlang schreef hij columns voor het tijdschrift <i>Current Biology</i>. </p><p>Op 11 oktober 2006 won Brenner de National Science and Technology Medal van <a href="/w/index.php?title=A*STAR&action=edit&redlink=1" class="new" title="A*STAR (de pagina bestaat niet)">A*STAR</a> (Agency for Science, Technology and Research) te Singapore.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Hij overleed op 92-jarige leeftijd in <a href="/wiki/Singapore" title="Singapore">Singapore</a>, alwaar hij werkzaam was voor A*STAR.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Boeken_van_Sidney_Brenner">Boeken van Sidney Brenner</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Sydney_Brenner&veaction=edit&section=3" title="Bewerk dit kopje: Boeken van Sidney Brenner" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Sydney_Brenner&action=edit&section=3" title="De broncode bewerken van de sectie: Boeken van Sidney Brenner"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><i>Loose End</i> : Collection of Loose Ends/False Starts columns by 'Uncle Syd.' from January 1994 to December 2000 (Current Biology, 1997) <a href="/wiki/Speciaal:Boekbronnen/1859223257" class="internal mw-magiclink-isbn">ISBN 1-85922-325-7</a>.</li> <li><i>My Life in Science</i>, with Lewis Wolpert, edited by Errol C. Friedberg and Eleanor Lawrence, BioMed Central 2001, 199pp <a href="/wiki/Speciaal:Boekbronnen/0954027809" class="internal mw-magiclink-isbn">ISBN 0-9540278-0-9</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="Externe_links">Externe links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Sydney_Brenner&veaction=edit&section=4" title="Bewerk dit kopje: Externe links" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Sydney_Brenner&action=edit&section=4" title="De broncode bewerken van de sectie: Externe links"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="http://webofstories.com/gl/sydney.brenner">Sydney Brenner vertelt zijn levensverhaal op Web of Stories</a></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/2002/summary/">The Nobel Prize in Physiology or Medicine 2002</a>. NobelPrize.org. Nobel Media AB 2018.</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20051228122848/http://elegans.swmed.edu/Sydney.html">Sydney Brenner en rondwormen</a></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20120416025158/http://www.alanmacfarlane.com/ancestors/brenner.htm">Sydney Brenner interviewed by Alan Macfarlane, 23 August 2007 (film)</a></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="http://www.laskerfoundation.org/new-noteworthy/articles/sydney-brenner-genes-tell-us-about-past/">Video: Sydney Brenner's seminar "What Genomes Can Tell Us About the Past"</a></li></ul> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li>Dit artikel of een eerdere versie ervan is een (gedeeltelijke) vertaling van het artikel <span class="plainlinks"><a class="external text" href="https://en.wikipedia.org/wiki/Sydney_Brenner?oldid=">Sydney Brenner</a> op de Engelstalige Wikipedia, dat onder de licentie <a rel="nofollow" class="external text" href="https://creativecommons.org/licenses/by-sa/3.0/deed.nl">Creative Commons Naamsvermelding/Gelijk delen</a> valt. Zie de <a class="external text" href="https://en.wikipedia.org/wiki/Sydney_Brenner?action=history">bewerkingsgeschiedenis</a></span> aldaar.<style data-mw-deduplicate="TemplateStyles:r66936091">.mw-parser-output .error-bronvermelding{color:#ff0000;font-weight:bold}.mw-parser-output .error-bronvermelding span{font-size:larger}</style></li> <li><i>The Science Times Book of the Brain</i>, redacteur Nicholas Wade, 1998, The Lyons Press</li> <li>Judson, H. F., <i>The Eighth Day of Creation</i> (1979), p. 10-11</li> <li><span class="citation" id="CITEREFC.D._Dijkstra2002"><span style="font-variant:small-caps;">C.D. Dijkstra</span> (2002). <cite>Nobelprijs Fysiologie of Geneeskunde 2002 voor onderzoek naar de genetische regulatie van orgaanontwikkeling en geprogrammeerde celdood</cite>. <i>Nederlands Tijdschrift voor Geneeskunde</i> <b>146</b> (52): 2525-2527. </span></li></ul> <div class="reflist" style="list-style-type: decimal;"><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><span class="citation"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/2002/brenner/auto-biography/">Sydney Brenner – Biographical</a>. NobelPrize.org. Nobel Media AB 2018.. Geraadpleegd op <span class="mw-formatted-date" title="2008-09-28">28 september 2008</span>. </span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF2002"><span style="font-variant:small-caps;">Sydney Brenner</span>, <a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/2002/brenner/lecture/">Nobel Lecture: Nature's Gift to Science</a> (video & pdf). NobelPrize.org. Nobel Media AB 2018. (<span class="mw-formatted-date" title="2002-12-08">8 december 2002</span>). Geraadpleegd op <span class="mw-formatted-date" title="2008-09-28">28 september 2008</span>. </span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20070420201534/http://www.a-star.edu.sg/astar/about/action/award_NSTM_citation.do?id=0e39f93f71TO&awardId=0e29810124jK">A*STAR Corporate Site - Awards - NSTM - Winner</a></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><span class="citation" id="CITEREF2019"><span style="font-variant:small-caps;">Nicholas Wade</span>, <a rel="nofollow" class="external text" href="https://www.nytimes.com/2019/04/05/obituaries/sydney-brenner-dead.html">Sydney Brenner, a Decipherer of the Genetic Code, Is Dead at 92</a>. The New York Times (<span class="mw-formatted-date" title="2019-04-05">5 april 2019</span>). </span></span> </li> </ol></div></div> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output .navigatie-afb-rechts{position:absolute}.mw-parser-output .navigatie-afb-rechts{right:2px}.mw-parser-output .navigatie-afb-groot{float:right;padding-left:0.5em}.mw-parser-output .navigatie-titel{background-color:#ddeef8;padding:2px 5.5em;font-weight:bold}.mw-parser-output .navigatie-inhoud{padding:0.5em}.mw-parser-output .navigatie-inhoud p:first-child{margin:0}.mw-parser-output .navigatie div[style*="background-color"],.mw-parser-output .navigatie div[style*="background"]{color:inherit}@media screen{html.skin-theme-clientpref-night .mw-parser-output .navigatie-titel{background-color:var(--background-color-interactive,#eaecf0)!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navigatie-titel{background-color:var(--background-color-interactive,#eaecf0)!important}}</style><div class="navigatie mw-collapsible mw-collapsed" role="navigation" aria-labelledby="Bibliografische informatie" data-collapsetext="Inklappen" data-expandtext="Uitklappen"><div id="Bibliografische_informatie" class="navigatie-titel"><a href="/wiki/Wikipedia:Bibliografische_informatie" title="Wikipedia:Bibliografische informatie">Bibliografische informatie</a></div><div class="navigatie-inhoud mw-collapsible-content plainlist ta-left"><ul style="column-width:36em"><li><b><a href="/wiki/Bibsys" class="mw-redirect" title="Bibsys">Bibsys</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://authority.bibsys.no/authority/rest/authorities/html/98050577">98050577</a></span></li><li><b><a href="/wiki/Biblioteca_Nacional_de_Espa%C3%B1a" title="Biblioteca Nacional de España">Biblioteca Nacional de España</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://catalogo.bne.es/uhtbin/authoritybrowse.cgi?action=display&authority_id=XX1761005">XX1761005</a></span></li><li><b><a href="/wiki/Biblioth%C3%A8que_nationale_de_France" title="Bibliothèque nationale de France">Bibliothèque nationale de France</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb13522242t">cb13522242t</a> <a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb13522242t">(data)</a></span></li><li><b><a href="/wiki/Cat%C3%A0leg_d%27autoritats_de_noms_i_t%C3%ADtols_de_Catalunya" title="Catàleg d'autoritats de noms i títols de Catalunya">Catàleg d'autoritats de noms i títols de Catalunya</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://cantic.bnc.cat/registre/981058529803306706">981058529803306706</a></span></li><li><b><a href="/wiki/CiNii" title="CiNii">CiNii</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://ci.nii.ac.jp/author/DA04993900?l=en">DA04993900</a></span></li><li><b><a href="/wiki/Gemeinsame_Normdatei" title="Gemeinsame Normdatei">Gemeinsame Normdatei</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/128411848">128411848</a></span></li><li><b><a href="/wiki/International_Standard_Name_Identifier" title="International Standard Name Identifier">International Standard Name Identifier</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://isni.org/isni/0000000110743762">0000 0001 1074 3762</a></span></li><li><b><a href="/wiki/Library_of_Congress_Control_Number" title="Library of Congress Control Number">Library of Congress Control Number</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/names/n81037896">n81037896</a></span></li><li><b><a href="/wiki/Mathematics_Genealogy_Project" title="Mathematics Genealogy Project">Mathematics Genealogy Project</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://genealogy.math.ndsu.nodak.edu/id.php?id=261921">261921</a></span></li><li><b><a href="/wiki/Bibliotheek_van_het_Japanse_parlement" title="Bibliotheek van het Japanse parlement">Bibliotheek van het Japanse parlement</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00986898">00986898</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Tsjechi%C3%AB" title="Nationale Bibliotheek van Tsjechië">Nationale Bibliotheek van Tsjechië</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=nlk20030145230&CON_LNG=ENG">nlk20030145230</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Isra%C3%ABl" title="Nationale Bibliotheek van Israël">Nationale Bibliotheek van Israël</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&local_base=NLX10&find_code=UID&request=987007259058505171">987007259058505171</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Polen" title="Nationale Bibliotheek van Polen">Nationale Bibliotheek van Polen</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://dbn.bn.org.pl/descriptor-details/a0000003021712">a0000003021712</a></span></li><li><b><a href="https://www.wikidata.org/wiki/Property:P1006" class="extiw" title="d:Property:P1006">Nederlandse Thesaurus van Auteursnamen</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p236144499">236144499</a></span></li><li><b><a href="/wiki/R%C3%89RO" title="RÉRO">Réseau des bibliothèques de Suisse occidentale</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://data.rero.ch/02-A003041355">02-A003041355</a></span></li><li><b><a href="/wiki/LIBRIS" title="LIBRIS">LIBRIS</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://libris.kb.se/auth/240037">240037</a></span></li><li><b><a href="/wiki/SNAC" title="SNAC">SNAC</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://snaccooperative.org/ark:/99166/w6c37d8g">w6c37d8g</a></span></li><li><b><a href="/wiki/Syst%C3%A8me_universitaire_de_documentation" title="Système universitaire de documentation">Système universitaire de documentation</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/031611338">031611338</a></span></li><li><b><a href="/wiki/Virtual_International_Authority_File" title="Virtual International Authority File">Virtual International Authority File</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/84854329">84854329</a></span></li><li><b></b> <b><a href="/wiki/WorldCat" title="WorldCat">WorldCat</a></b> (via VIAF): <a rel="nofollow" class="external text" href="https://www.worldcat.org/identities/containsVIAFID/84854329">84854329</a></li></ul></div></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67837862"> <div class="navigatie mw-collapsible mw-collapsed" role="navigation" aria-labelledby="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" data-expandtext="Uitklappen" data-collapsetext="Inklappen"> <div class="navigatie-afb-links noviewer"><div class="navigatie-bewerken show-autoconfirmed nomobile"><span class="nowrap skin-invert-image"><span typeof="mw:File"><a href="/wiki/Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Sjabloon:Navigatie winnaars van de Nobelprijs voor Fysiologie of Geneeskunde"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/16px-OOjs_UI_icon_article-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/24px-OOjs_UI_icon_article-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/32px-OOjs_UI_icon_article-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> · <span class="ui-state-disabled" typeof="mw:File"><span title="Overleg sjabloon (de pagina bestaat niet)"><img alt="Overleg sjabloon (de pagina bestaat niet)" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/16px-OOjs_UI_icon_speechBubbles-ltr.svg.png" decoding="async" 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title="Santiago Ramón y Cajal">Ramón y Cajal</a> · 1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a class="mw-selflink selflink">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Overgenomen van "<a dir="ltr" href="https://nl.wikipedia.org/w/index.php?title=Sydney_Brenner&oldid=68015129">https://nl.wikipedia.org/w/index.php?title=Sydney_Brenner&oldid=68015129</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Categorie:Alles" title="Categorie:Alles">Categorieën</a>: <ul><li><a href="/wiki/Categorie:Geneticus" title="Categorie:Geneticus">Geneticus</a></li><li><a href="/wiki/Categorie:Winnaar_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Categorie:Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde">Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde</a></li><li><a href="/wiki/Categorie:Zuid-Afrikaans_bioloog" title="Categorie:Zuid-Afrikaans bioloog">Zuid-Afrikaans bioloog</a></li><li><a href="/wiki/Categorie:Zuid-Afrikaans_hoogleraar" title="Categorie:Zuid-Afrikaans hoogleraar">Zuid-Afrikaans hoogleraar</a></li><li><a href="/wiki/Categorie:Lid_van_de_Acad%C3%A9mie_des_sciences" title="Categorie:Lid van de Académie des sciences">Lid van de Académie des sciences</a></li><li><a href="/wiki/Categorie:20e-eeuws_medicus" title="Categorie:20e-eeuws medicus">20e-eeuws medicus</a></li><li><a href="/wiki/Categorie:20e-eeuws_bioloog" title="Categorie:20e-eeuws bioloog">20e-eeuws bioloog</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Verborgen categorie: <ul><li><a href="/wiki/Categorie:Wikipedia:Pagina%27s_die_ISBN_magische_links_gebruiken" title="Categorie:Wikipedia:Pagina's die ISBN magische links gebruiken">Wikipedia:Pagina's die ISBN magische links gebruiken</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> Deze pagina is voor het laatst bewerkt op 25 aug 2024 om 08:16.</li> <li id="footer-info-copyright">De tekst is beschikbaar onder de licentie <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/deed.nl">Creative Commons Naamsvermelding/Gelijk delen</a>, er kunnen aanvullende voorwaarden van toepassing zijn. 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