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<div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Structure in cerebellum that modulates muscle stretch reflexes of proximal limb muscles.</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox"><tbody><tr><th colspan="2" class="infobox-above" style="background-color:lemonchiffon">Interposed nucleus</th></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Details</th></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Identifiers</th></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Latin" title="Latin">Latin</a></th><td class="infobox-data"><i>nucleus interpositus anterior, nucleus interpositus posterior</i></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/NeuroLex" title="NeuroLex">NeuroLex</a> ID</th><td class="infobox-data"><a rel="nofollow" class="external text" href="http://uri.neuinfo.org/nif/nifstd/nlx_anat_20081242">nlx_anat_20081242</a></td></tr><tr><td colspan="2" class="infobox-below"><a href="/wiki/Anatomical_terms_of_neuroanatomy" title="Anatomical terms of neuroanatomy">Anatomical terms of neuroanatomy</a><div style="text-align: right;"><small class="noprint">[<a href="https://www.wikidata.org/wiki/Q6056415" class="extiw" title="d:Q6056415">edit on Wikidata</a>]</small></div></td></tr></tbody></table> <p>The <b>interposed nucleus</b> is the combined paired <a href="/wiki/Globose_nucleus" title="Globose nucleus">globose</a> and <a href="/wiki/Emboliform_nucleus" title="Emboliform nucleus">emboliform nuclei</a>, (<a href="/wiki/Deep_cerebellar_nuclei" title="Deep cerebellar nuclei">deep cerebellar nuclei</a>) on either side of the <a href="/wiki/Cerebellum" title="Cerebellum">cerebellum</a>.<sup id="cite_ref-Purves_1-0" class="reference"><a href="#cite_note-Purves-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Patestas-2016_2-0" class="reference"><a href="#cite_note-Patestas-2016-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> It is located in the <a href="/wiki/Roof_of_the_fourth_ventricle" class="mw-redirect" title="Roof of the fourth ventricle">roof of the fourth ventricle</a>, lateral to the <a href="/wiki/Fastigial_nucleus" title="Fastigial nucleus">fastigial nucleus</a>. The emboliform nucleus is the <i>anterior interposed nucleus</i>, and the globose nucleus is the <i>posterior interposed nucleus</i>.<sup id="cite_ref-Haines_3-0" class="reference"><a href="#cite_note-Haines-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>The interposed nucleus is responsible for coordinating agonist/antagonist muscle pairs, and therefore a lesion in this area causes <a href="/wiki/Tremor" title="Tremor">tremor</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Anatomy">Anatomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Interposed_nucleus&action=edit&section=1" title="Edit section: Anatomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The interposed nucleus is located in the paravermis of the cerebellum.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (June 2024)">citation needed</span></a></i>]</sup> </p><p>The interposed nucleus is smaller than the dentate but larger than the fastigial nucleus.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (June 2024)">citation needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading3"><h3 id="Afferents">Afferents</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Interposed_nucleus&action=edit&section=2" title="Edit section: Afferents"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The interposed nuclei receives Purkine cell terminal afferents from the paravermal cortex of the <a href="/wiki/Spinocerebellum" class="mw-redirect" title="Spinocerebellum">spinocerebellum</a>, as well as collaterals of cerebellar afferents from the <a href="/wiki/Restiform_body" class="mw-redirect" title="Restiform body">restiform body</a> and <a href="/wiki/Ventral_spinocerebellar_tract" class="mw-redirect" title="Ventral spinocerebellar tract">ventral spinocerebellar tract</a>.<sup id="cite_ref-Patestas-2016_2-1" class="reference"><a href="#cite_note-Patestas-2016-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>It receives input from the ipsilateral <a href="/wiki/Posterior_external_arcuate_fibers" class="mw-redirect" title="Posterior external arcuate fibers">posterior external arcuate fibers</a> (cuneocerebellar tract) and the <a href="/wiki/Dorsal_spinocerebellar_tract" class="mw-redirect" title="Dorsal spinocerebellar tract">dorsal spinocerebellar tract</a>, which originate in the <a href="/wiki/Accessory_cuneate_nucleus" title="Accessory cuneate nucleus">accessory cuneate nucleus</a> and the <a href="/wiki/Posterior_thoracic_nucleus" title="Posterior thoracic nucleus">posterior thoracic nucleus</a>, respectively.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (June 2024)">citation needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading3"><h3 id="Efferents">Efferents</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Interposed_nucleus&action=edit&section=3" title="Edit section: Efferents"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Afferents from the interposed nuclei leave the cerebellum through <a href="/wiki/Superior_cerebellar_peduncle" title="Superior cerebellar peduncle">superior cerebellar peduncle</a>. They project to:<sup id="cite_ref-Patestas-2016_2-2" class="reference"><a href="#cite_note-Patestas-2016-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <ul><li>contralateral (magnocellular division of) the <b><a href="/wiki/Red_nucleus" title="Red nucleus">red nucleus</a></b> (→ <a href="/wiki/Rubrospinal_tract" title="Rubrospinal tract">rubrospinal tract</a>). The red nucleus is the main target of tne interposed nuclei.</li> <li>ipsilateral <a href="/wiki/Ventral_lateral_nucleus" title="Ventral lateral nucleus"><b>ventral lateral nucleus of thalamus</b></a> (→ <a href="/wiki/Premotor_cortex" title="Premotor cortex">premotor cortex</a> and <a href="/wiki/Primary_motor_cortex" title="Primary motor cortex">primary motor cortex</a> → <a href="/wiki/Lateral_corticospinal_tract" title="Lateral corticospinal tract">lateral corticospinal tract</a>).</li></ul> <p>The rubrospinal and lateral corticospinal tracts are subsequently involved in control of the distal musculature of the extremities. </p> <div class="mw-heading mw-heading2"><h2 id="Function">Function</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Interposed_nucleus&action=edit&section=4" title="Edit section: Function"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The interposed nucleus modulates muscle stretch reflexes of proximal limb muscles, and is also required in delayed <a href="/wiki/Classical_conditioning" title="Classical conditioning">Pavlovian conditioning</a>.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Interposed_nucleus&action=edit&section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-Purves-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-Purves_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation 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a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFPurves2012" class="citation book cs1">Purves, Dale (2012). <i>Neuroscience</i> (5. ed.). Sunderland, Mass: Sinauer. p. 419. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780878936953" title="Special:BookSources/9780878936953"><bdi>9780878936953</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Neuroscience&rft.place=Sunderland%2C+Mass&rft.pages=419&rft.edition=5.&rft.pub=Sinauer&rft.date=2012&rft.isbn=9780878936953&rft.aulast=Purves&rft.aufirst=Dale&rfr_id=info%3Asid%2Fen.wikipedia.org%3AInterposed+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-Patestas-2016-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-Patestas-2016_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Patestas-2016_2-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Patestas-2016_2-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPatestasGartner2016" class="citation book cs1">Patestas, Maria A.; Gartner, Leslie P. (2016). <i>A Textbook of Neuroanatomy</i> (2nd ed.). Hoboken, New Jersey: Wiley-Blackwell. p. 292. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1-118-67746-9" title="Special:BookSources/978-1-118-67746-9"><bdi>978-1-118-67746-9</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=A+Textbook+of+Neuroanatomy&rft.place=Hoboken%2C+New+Jersey&rft.pages=292&rft.edition=2nd&rft.pub=Wiley-Blackwell&rft.date=2016&rft.isbn=978-1-118-67746-9&rft.aulast=Patestas&rft.aufirst=Maria+A.&rft.au=Gartner%2C+Leslie+P.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AInterposed+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-Haines-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-Haines_3-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHaines2018" class="citation book cs1">Haines, Duane (2018). <i>Fundamental neuroscience for basic and clinical applications</i> (Fifth ed.). Philadelphia, PA: Elsevier. p. 396. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780323396325" title="Special:BookSources/9780323396325"><bdi>9780323396325</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Fundamental+neuroscience+for+basic+and+clinical+applications&rft.place=Philadelphia%2C+PA&rft.pages=396&rft.edition=Fifth&rft.pub=Elsevier&rft.date=2018&rft.isbn=9780323396325&rft.aulast=Haines&rft.aufirst=Duane&rfr_id=info%3Asid%2Fen.wikipedia.org%3AInterposed+nucleus" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFClarkZhangLavond1992" class="citation journal cs1">Clark, Robert E.; Zhang, Andrew A.; Lavond, David G. (1992). "Reversible lesions of the cerebellar interpositus nucleus during acquisition and retention of a classically conditioned behavior". <i>Behavioral Neuroscience</i>. <b>106</b> (6): 879–888. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1037%2F0735-7044.106.6.879">10.1037/0735-7044.106.6.879</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/1335267">1335267</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Behavioral+Neuroscience&rft.atitle=Reversible+lesions+of+the+cerebellar+interpositus+nucleus+during+acquisition+and+retention+of+a+classically+conditioned+behavior.&rft.volume=106&rft.issue=6&rft.pages=879-888&rft.date=1992&rft_id=info%3Adoi%2F10.1037%2F0735-7044.106.6.879&rft_id=info%3Apmid%2F1335267&rft.aulast=Clark&rft.aufirst=Robert+E.&rft.au=Zhang%2C+Andrew+A.&rft.au=Lavond%2C+David+G.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AInterposed+nucleus" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Interposed_nucleus&action=edit&section=6" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external free" 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.navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Anatomy_of_the_cerebellum" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Cerebellum" title="Template:Cerebellum"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Cerebellum" title="Template talk:Cerebellum"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Cerebellum" title="Special:EditPage/Template:Cerebellum"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Anatomy_of_the_cerebellum" style="font-size:114%;margin:0 4em"><a href="/wiki/Anatomy_of_the_cerebellum" title="Anatomy of the cerebellum">Anatomy</a> of the <a href="/wiki/Cerebellum" title="Cerebellum">cerebellum</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Surface</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Lobes</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anterior_lobe_of_cerebellum" title="Anterior lobe of cerebellum">Anterior lobe</a></li> <li><a href="/wiki/Posterior_lobe_of_cerebellum" title="Posterior lobe of cerebellum">Posterior lobe</a> <ul><li><a href="/wiki/Horizontal_fissure_of_cerebellum" title="Horizontal fissure of cerebellum">Horizontal fissure</a></li></ul></li> <li><a href="/wiki/Flocculonodular_lobe" title="Flocculonodular lobe">Flocculonodular lobe</a> <ul><li><a href="/wiki/Flocculus_(cerebellar)" class="mw-redirect" title="Flocculus (cerebellar)">Flocculus</a></li> <li><a href="/wiki/Nodule_of_vermis" title="Nodule of vermis">Nodule</a></li></ul></li> <li><a href="/wiki/Primary_fissure_of_cerebellum" title="Primary fissure of cerebellum">Primary fissure</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Medial/lateral</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cerebellar_vermis" title="Cerebellar vermis">Vermis</a>: <i>anterior</i> <ul><li><a href="/wiki/Central_lobule" title="Central lobule">Central lobule</a></li> <li><a href="/wiki/Culmen_(cerebellum)" title="Culmen (cerebellum)">Culmen</a></li> <li><a href="/wiki/Lingula_of_cerebellum" title="Lingula of cerebellum">Lingula</a></li></ul></li> <li><i>posterior</i> <ul><li><a href="/wiki/Folium_vermis" title="Folium vermis">Folium</a></li> <li><a href="/wiki/Tuber_of_vermis" title="Tuber of vermis">Tuber</a></li> <li><a href="/wiki/Uvula_of_cerebellum" title="Uvula of cerebellum">Uvula</a></li></ul></li> <li><a href="/wiki/Vallecula_of_cerebellum" title="Vallecula of cerebellum">Vallecula of cerebellum</a></li></ul> <ul><li><a href="/wiki/Cerebellar_hemisphere" title="Cerebellar hemisphere">Hemisphere</a>: <i>anterior</i> <ul><li><a href="/wiki/Alar_central_lobule" class="mw-redirect" title="Alar central lobule">Alar central lobule</a></li></ul></li> <li><i>posterior</i> <ul><li><a href="/wiki/Biventer_lobule" title="Biventer lobule">Biventer lobule</a></li> <li><a href="/wiki/Cerebellar_tonsil" title="Cerebellar tonsil">Cerebellar tonsil</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_matter" title="Grey matter">Grey matter</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Deep_cerebellar_nuclei" title="Deep cerebellar nuclei">Deep cerebellar nuclei</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dentate_nucleus" title="Dentate nucleus">Dentate</a></li> <li><i><a class="mw-selflink selflink">interposed</a></i> <ul><li><a href="/wiki/Emboliform_nucleus" title="Emboliform nucleus">Emboliform</a></li> <li><a href="/wiki/Globose_nucleus" title="Globose nucleus">Globose</a></li></ul></li> <li><a href="/wiki/Fastigial_nucleus" title="Fastigial nucleus">Fastigial</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cerebellum" title="Cerebellum">Cerebellar cortex</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Molecular layer</i> <ul><li><a href="/wiki/Stellate_cell" title="Stellate cell">Stellate cell</a></li> <li><a href="/wiki/Basket_cell" title="Basket cell">Basket cell</a></li></ul></li></ul> <ul><li><i>Purkinje cell layer</i> <ul><li><a href="/wiki/Purkinje_cell" title="Purkinje cell">Purkinje cell</a></li> <li><a href="/wiki/Astrocyte#Bergmann_glia" title="Astrocyte">Bergmann glia cell = Golgi epithelial cell</a></li> <li><a href="/wiki/Fa%C3%B1anas_cell" class="mw-redirect" title="Fañanas cell">Fañanas cell</a></li></ul></li></ul> <ul><li><i>Granule cell layer</i> <ul><li><a href="/wiki/Golgi_cell" title="Golgi cell">Golgi cell</a></li> <li><a href="/wiki/Granule_cell" title="Granule cell">Granule cell</a></li> <li><a href="/wiki/Unipolar_brush_cell" title="Unipolar brush cell">Unipolar brush cell</a></li></ul></li></ul> <ul><li>Fibers: <a href="/wiki/Mossy_fiber_(cerebellum)" title="Mossy fiber (cerebellum)">Mossy fibers</a></li> <li><a href="/wiki/Climbing_fiber" title="Climbing fiber">Climbing fiber</a></li> <li><a href="/wiki/Parallel_fiber" class="mw-redirect" title="Parallel fiber">Parallel fiber</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/White_matter" title="White matter">White matter</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Internal</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arbor_vitae_(anatomy)" title="Arbor vitae (anatomy)">Arbor vitae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cerebellar_peduncle" class="mw-redirect" title="Cerebellar peduncle">Peduncles</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Inferior_cerebellar_peduncle" title="Inferior cerebellar peduncle">Inferior</a> (medulla): <a href="/wiki/Dorsal_spinocerebellar_tract" class="mw-redirect" title="Dorsal spinocerebellar tract">Dorsal spinocerebellar tract</a></li> <li><a href="/wiki/Olivocerebellar_tract" title="Olivocerebellar tract">Olivocerebellar tract</a></li> <li><a href="/wiki/Posterior_external_arcuate_fibers" class="mw-redirect" title="Posterior external arcuate fibers">Cuneocerebellar tract</a></li> <li><a href="/wiki/Juxtarestiform_body" title="Juxtarestiform body">Juxtarestiform body</a> (<a href="/wiki/Vestibulocerebellar_tract" title="Vestibulocerebellar tract">Vestibulocerebellar tract</a>)</li> <li><a href="/wiki/Trigeminocerebellar_fibers" title="Trigeminocerebellar fibers">Trigeminocerebellar fibers</a></li></ul> <ul><li><a href="/wiki/Middle_cerebellar_peduncle" title="Middle cerebellar peduncle">Middle</a> (pons): <a href="/wiki/Pontocerebellar_fibers" title="Pontocerebellar fibers">Pontocerebellar fibers</a></li></ul> <ul><li><a href="/wiki/Superior_cerebellar_peduncle" title="Superior cerebellar peduncle">Superior</a> (midbrain): <a href="/wiki/Ventral_spinocerebellar_tract" class="mw-redirect" title="Ventral spinocerebellar tract">Ventral spinocerebellar tract</a></li> <li><a href="/wiki/Dentatothalamic_tract" title="Dentatothalamic tract">Dentatothalamic tract</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Brain_and_spinal_cord:_neural_tracts_and_fasciculi" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Neural_tracts" title="Template:Neural tracts"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Neural_tracts" title="Template talk:Neural tracts"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Neural_tracts" title="Special:EditPage/Template:Neural tracts"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Brain_and_spinal_cord:_neural_tracts_and_fasciculi" style="font-size:114%;margin:0 4em"><a href="/wiki/Human_brain" title="Human brain">Brain</a> and <a href="/wiki/Spinal_cord" title="Spinal cord">spinal cord</a>: <a href="/wiki/Neural_pathway" title="Neural pathway">neural tracts</a> and <a href="/wiki/Nerve_fascicle" title="Nerve fascicle">fasciculi</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Sensory</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Dorsal_column-medial_lemniscus_pathway" class="mw-redirect" title="Dorsal column-medial lemniscus pathway">DCML</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;background: transparent;"><a href="/wiki/Dorsal_column-medial_lemniscus_pathway#First_neuron" class="mw-redirect" title="Dorsal column-medial lemniscus pathway">1°</a>:</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lamellar_corpuscle" class="mw-redirect" title="Lamellar corpuscle">Pacinian corpuscle</a>/<a href="/wiki/Tactile_corpuscle" title="Tactile corpuscle">Meissner's corpuscle</a> → <b><a href="/wiki/Posterior_column" class="mw-redirect" title="Posterior column">Posterior column</a></b> (<a href="/wiki/Gracile_fasciculus" class="mw-redirect" title="Gracile fasciculus">Gracile fasciculus</a>/<a href="/wiki/Cuneate_fasciculus" class="mw-redirect" title="Cuneate fasciculus">Cuneate fasciculus</a>) → <a href="/wiki/Gracile_nucleus" class="mw-redirect" title="Gracile nucleus">Gracile nucleus</a>/<a href="/wiki/Cuneate_nucleus" class="mw-redirect" title="Cuneate nucleus">Cuneate nucleus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: transparent;"><a href="/wiki/Dorsal_column-medial_lemniscus_pathway#Second_neuron" class="mw-redirect" title="Dorsal column-medial lemniscus pathway">2°</a>:</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>→ <i><a href="/wiki/Sensory_decussation" title="Sensory decussation">sensory decussation</a>/arcuate fibers</i> (<a href="/wiki/Posterior_external_arcuate_fibers" class="mw-redirect" title="Posterior external arcuate fibers">Posterior external arcuate fibers</a>, <a href="/wiki/Internal_arcuate_fibers" title="Internal arcuate fibers">Internal arcuate fibers</a>) → <a href="/wiki/Medial_lemniscus" title="Medial lemniscus">Medial lemniscus</a>/<a href="/wiki/Trigeminal_lemniscus" title="Trigeminal lemniscus">Trigeminal lemniscus</a> → <a href="/wiki/Thalamus" title="Thalamus">Thalamus</a> (<a href="/wiki/Ventral_posterolateral_nucleus" title="Ventral posterolateral nucleus">VPL</a>, <a href="/wiki/Ventral_posteromedial_nucleus" title="Ventral posteromedial nucleus">VPM</a>)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background: transparent;"><a href="/wiki/Dorsal_column-medial_lemniscus_pathway#Third_neuron" class="mw-redirect" title="Dorsal column-medial lemniscus pathway">3°</a>:</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>→ <a href="/wiki/Posterior_limb_of_internal_capsule" class="mw-redirect" title="Posterior limb of internal capsule">Posterior limb of internal capsule</a> → <a href="/wiki/Postcentral_gyrus" title="Postcentral gyrus">Postcentral gyrus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Anterolateral_system" class="mw-redirect" title="Anterolateral system">Anterolateral</a>/<br /><a href="/wiki/Pain" title="Pain">pain</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nociception#Neospinothalamic_tract" title="Nociception">Fast</a>/lateral</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>1° (<a href="/wiki/Free_nerve_ending" title="Free nerve ending">Free nerve ending</a> → <a href="/wiki/A_delta_fiber" class="mw-redirect" title="A delta fiber">A delta fiber</a>) → 2° (<a href="/wiki/Anterior_white_commissure" title="Anterior white commissure">Anterior white commissure</a> → <b><a href="/wiki/Lateral_spinothalamic_tract" class="mw-redirect" title="Lateral spinothalamic tract">Lateral</a> and <a href="/wiki/Anterior_spinothalamic_tract" class="mw-redirect" title="Anterior spinothalamic tract">Anterior</a> <a href="/wiki/Spinothalamic_tract" title="Spinothalamic tract">Spinothalamic tract</a></b> → <a href="/wiki/Spinothalamic_tract" title="Spinothalamic tract">Spinal lemniscus</a> → <a href="/wiki/Ventral_posterolateral_nucleus" title="Ventral posterolateral nucleus">VPL</a> of <a href="/wiki/Thalamus" title="Thalamus">Thalamus</a>) → 3° (<a href="/wiki/Postcentral_gyrus" title="Postcentral gyrus">Postcentral gyrus</a>) → 4° (<a href="/wiki/Posterior_parietal_cortex" title="Posterior parietal cortex">Posterior parietal cortex</a>)</li></ul> <p>2° (<b><a href="/wiki/Spinomesencephalic_tract" class="mw-redirect" title="Spinomesencephalic tract">Spinomesencephalic tract</a></b> → <a href="/wiki/Superior_colliculus" title="Superior colliculus">Superior colliculus</a> of <a href="/wiki/Midbrain_tectum" class="mw-redirect" title="Midbrain tectum">Midbrain tectum</a>) </p> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nociception#Paleospinothalamic_tract" title="Nociception">Slow</a>/medial</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>1° (<a href="/wiki/Group_C_nerve_fiber" title="Group C nerve fiber">Group C nerve fiber</a> → <b><a href="/wiki/Spinoreticular_tract" title="Spinoreticular tract">Spinoreticular tract</a></b> → <a href="/wiki/Reticular_formation" title="Reticular formation">Reticular formation</a>) → 2° (<a href="/wiki/Medial_dorsal_nucleus" title="Medial dorsal nucleus">MD</a> of <a href="/wiki/Thalamus" title="Thalamus">Thalamus</a>) → 3° (<a href="/wiki/Cingulate_cortex" title="Cingulate cortex">Cingulate cortex</a>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Motor</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pyramidal_tracts" title="Pyramidal tracts">Pyramidal</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>flexion:</i> <a href="/wiki/Primary_motor_cortex" title="Primary motor cortex">Primary motor cortex</a> → <a href="/wiki/Posterior_limb_of_internal_capsule" class="mw-redirect" title="Posterior limb of internal capsule">Posterior limb of internal capsule</a> → <a href="/wiki/Decussation_of_pyramids" class="mw-redirect" title="Decussation of pyramids">Decussation of pyramids</a> → <b><a href="/wiki/Pyramidal_tracts" title="Pyramidal tracts">Corticospinal tract</a> (<a href="/wiki/Lateral_corticospinal_tract" title="Lateral corticospinal tract">Lateral</a>, <a href="/wiki/Anterior_corticospinal_tract" title="Anterior corticospinal tract">Anterior</a>)</b> → <a href="/wiki/Neuromuscular_junction" title="Neuromuscular junction">Neuromuscular junction</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Extrapyramidal_system" title="Extrapyramidal system">Extrapyramidal</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><i>flexion:</i></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Primary_motor_cortex" title="Primary motor cortex">Primary motor cortex</a> → <a href="/wiki/Genu_of_internal_capsule" class="mw-redirect" title="Genu of internal capsule">Genu of internal capsule</a> → <b><a href="/wiki/Corticobulbar_tract" title="Corticobulbar tract">Corticobulbar tract</a></b> → <a href="/wiki/Facial_motor_nucleus" title="Facial motor nucleus">Facial motor nucleus</a> → <a href="/wiki/Facial_muscles" title="Facial muscles">Facial muscles</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><i>flexion:</i></span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Red_nucleus" title="Red nucleus">Red nucleus</a> → <b><a href="/wiki/Rubrospinal_tract" title="Rubrospinal tract">Rubrospinal tract</a></b></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><i>extension:</i></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anatomy_of_the_cerebellum#Phylogenetic_and_functional_divisions" title="Anatomy of the cerebellum"> Vestibulocerebellum</a> → <a href="/wiki/Vestibular_nuclei" title="Vestibular nuclei">Vestibular nuclei</a> → <b><a href="/wiki/Vestibulospinal_tract" title="Vestibulospinal tract">Vestibulospinal tract</a></b></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><i>extension:</i></span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anatomy_of_the_cerebellum#Phylogenetic_and_functional_divisions" title="Anatomy of the cerebellum"> Vestibulocerebellum</a> → <a href="/wiki/Reticular_formation" title="Reticular formation">Reticular formation</a> → <b><a href="/wiki/Reticular_formation#Descending_reticulospinal_tracts" title="Reticular formation">Reticulospinal tract</a></b></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Midbrain_tectum" class="mw-redirect" title="Midbrain tectum">Midbrain tectum</a> → <b><a href="/wiki/Tectospinal_tract" title="Tectospinal tract">Tectospinal tract</a></b> → muscles of neck</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Basal_ganglia" title="Basal ganglia">Basal ganglia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><a href="/wiki/Direct_pathway_of_movement" class="mw-redirect" title="Direct pathway of movement"><i>direct:</i></a></span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em">1° (<a href="/wiki/Motor_cortex" title="Motor cortex">Motor cortex</a> → <a href="/wiki/Striatum" title="Striatum">Striatum</a>) → 2° (<a href="/wiki/Medial_globus_pallidus" class="mw-redirect" title="Medial globus pallidus">GPi</a>) → 3° (<a href="/wiki/Lenticular_fasciculus" title="Lenticular fasciculus">Lenticular fasciculus</a>/<a href="/wiki/Ansa_lenticularis" title="Ansa lenticularis">Ansa lenticularis</a> → <a href="/wiki/Thalamic_fasciculus" title="Thalamic fasciculus">Thalamic fasciculus</a> → <a href="/wiki/Ventral_lateral_nucleus" title="Ventral lateral nucleus">VL</a> of <a href="/wiki/Thalamus" title="Thalamus">Thalamus</a>) → 4° (<a href="/wiki/Thalamocortical_radiations" title="Thalamocortical radiations">Thalamocortical radiations</a> → <a href="/wiki/Supplementary_motor_area" title="Supplementary motor area">Supplementary motor area</a>) → 5° (<a href="/wiki/Motor_cortex" title="Motor cortex">Motor cortex</a>)</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><a href="/wiki/Indirect_pathway_of_movement" class="mw-redirect" title="Indirect pathway of movement"><i>indirect:</i></a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em">1° (<a href="/wiki/Motor_cortex" title="Motor cortex">Motor cortex</a> → <a href="/wiki/Striatum" title="Striatum">Striatum</a>) → 2° (<a href="/wiki/Lateral_globus_pallidus" class="mw-redirect" title="Lateral globus pallidus">GPe</a>) → 3° (<a href="/wiki/Subthalamic_fasciculus" title="Subthalamic fasciculus">Subthalamic fasciculus</a> → <a href="/wiki/Subthalamic_nucleus" title="Subthalamic nucleus">Subthalamic nucleus</a>) → 4° (<a href="/wiki/Subthalamic_fasciculus" title="Subthalamic fasciculus">Subthalamic fasciculus</a> → <a href="/wiki/Medial_globus_pallidus" class="mw-redirect" title="Medial globus pallidus">GPi</a>) → 5° (<a href="/wiki/Lenticular_fasciculus" title="Lenticular fasciculus">Lenticular fasciculus</a>/<a href="/wiki/Ansa_lenticularis" title="Ansa lenticularis">Ansa lenticularis</a> → <a href="/wiki/Thalamic_fasciculus" title="Thalamic fasciculus">Thalamic fasciculus</a> → <a href="/wiki/Ventral_lateral_nucleus" title="Ventral lateral nucleus">VL</a> of <a href="/wiki/Thalamus" title="Thalamus">Thalamus</a>) → 6° (<a href="/wiki/Thalamocortical_radiations" title="Thalamocortical radiations">Thalamocortical radiations</a> → <a href="/wiki/Supplementary_motor_area" title="Supplementary motor area">Supplementary motor area</a>) → 7° (<a href="/wiki/Motor_cortex" title="Motor cortex">Motor cortex</a>)</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><span class="nobold"><i><a href="/wiki/Nigrostriatal_pathway" title="Nigrostriatal pathway">nigrostriatal pathway</a>:</i></span></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Substantia_nigra_pars_compacta" class="mw-redirect" title="Substantia nigra pars compacta">Pars compacta</a> → <a href="/wiki/Striatum" title="Striatum">Striatum</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cerebellum" title="Cerebellum">Cerebellar</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Afferent</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Vestibular_nuclei" title="Vestibular nuclei">Vestibular nuclei</a> → <b><a href="/wiki/Vestibulocerebellar_tract" title="Vestibulocerebellar tract">Vestibulocerebellar tract</a></b> → <a href="/wiki/Inferior_cerebellar_peduncle" title="Inferior cerebellar peduncle">ICP</a> → <a href="/wiki/Cerebellum" title="Cerebellum">Cerebellum</a> → <a href="/wiki/Cerebellum_granule_cell" class="mw-redirect" title="Cerebellum granule cell">Granule cell</a></li></ul> <ul><li><a href="/wiki/Pontine_nuclei" title="Pontine nuclei">Pontine nuclei</a> → <b><a href="/wiki/Pontocerebellar_fibers" title="Pontocerebellar fibers">Pontocerebellar fibers</a></b> → <a href="/wiki/Middle_cerebellar_peduncle" title="Middle cerebellar peduncle">MCP</a> → <a href="/wiki/Deep_cerebellar_nuclei" title="Deep cerebellar nuclei">Deep cerebellar nuclei</a> → <a href="/wiki/Cerebellum_granule_cell" class="mw-redirect" title="Cerebellum granule cell">Granule cell</a></li></ul> <ul><li><a href="/wiki/Inferior_olivary_nucleus" title="Inferior olivary nucleus">Inferior olivary nucleus</a> → <b><a href="/wiki/Olivocerebellar_tract" title="Olivocerebellar tract">Olivocerebellar tract</a></b> → <a href="/wiki/Inferior_cerebellar_peduncle" title="Inferior cerebellar peduncle">ICP</a> → <a href="/wiki/Cerebellar_hemisphere" title="Cerebellar hemisphere">Hemisphere</a> → <a href="/wiki/Purkinje_cell" title="Purkinje cell">Purkinje cell</a> → <a href="/wiki/Deep_cerebellar_nuclei" title="Deep cerebellar nuclei">Deep cerebellar nuclei</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Efferent</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dentate_nucleus" title="Dentate nucleus">Dentate nucleus</a> in <a href="/wiki/Cerebellar_hemisphere" title="Cerebellar hemisphere">Lateral hemisphere/pontocerebellum</a> → <a href="/wiki/Superior_cerebellar_peduncle" title="Superior cerebellar peduncle">SCP</a> → <b><a href="/wiki/Dentatothalamic_tract" title="Dentatothalamic tract">Dentatothalamic tract</a></b> → <a href="/wiki/Thalamus" title="Thalamus">Thalamus</a> (<a href="/wiki/Ventral_lateral_nucleus" title="Ventral lateral nucleus">VL</a>) → <a href="/wiki/Motor_cortex" title="Motor cortex">Motor cortex</a></li></ul> <ul><li><a class="mw-selflink selflink">Interposed nucleus</a> in <a href="/wiki/Cerebellar_hemisphere" title="Cerebellar hemisphere">Intermediate hemisphere/spinocerebellum</a> → <a href="/wiki/Superior_cerebellar_peduncle" title="Superior cerebellar peduncle">SCP</a> → <a href="/wiki/Reticular_formation" title="Reticular formation">Reticular formation</a>, or → <b><a href="/wiki/Cerebellothalamic_tract" title="Cerebellothalamic tract">Cerebellothalamic tract</a></b> → <a href="/wiki/Red_nucleus" title="Red nucleus">Red nucleus</a> → <a href="/wiki/Thalamus" title="Thalamus">Thalamus</a> (<a href="/wiki/Ventral_lateral_nucleus" title="Ventral lateral nucleus">VL</a>) → <a href="/wiki/Motor_cortex" title="Motor cortex">Motor cortex</a></li></ul> <ul><li><a href="/wiki/Fastigial_nucleus" title="Fastigial nucleus">Fastigial nucleus</a> in <a href="/wiki/Flocculonodular_lobe" title="Flocculonodular lobe">Flocculonodular lobe/vestibulocerebellum</a> → <b><a href="/wiki/Vestibulocerebellar_tract" title="Vestibulocerebellar tract">Vestibulocerebellar tract</a></b> → <a href="/wiki/Vestibular_nuclei" title="Vestibular nuclei">Vestibular nuclei</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Bidirectional:<br /><a href="/wiki/Spinocerebellar_tract" class="mw-redirect" title="Spinocerebellar tract">Spinocerebellar</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Proprioception#Conscious_and_unconscious_proprioception" title="Proprioception">Unconscious<br /> proprioception</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lower_limb" class="mw-redirect" title="Lower limb">lower limb</a> → 1° (<a href="/wiki/Muscle_spindle" title="Muscle spindle">muscle spindles</a> → <a href="/wiki/Dorsal_root_ganglion" title="Dorsal root ganglion">DRG</a>) → 2° (<a href="/wiki/Posterior_thoracic_nucleus" title="Posterior thoracic nucleus">Posterior thoracic nucleus</a> → <b><a href="/wiki/Dorsal_spinocerebellar_tract" class="mw-redirect" title="Dorsal spinocerebellar tract">Dorsal/posterior spinocerebellar tract</a></b> → <a href="/wiki/Inferior_cerebellar_peduncle" title="Inferior cerebellar peduncle">ICP</a> → <a href="/wiki/Cerebellar_vermis" title="Cerebellar vermis">Cerebellar vermis</a>)</li></ul> <ul><li><a href="/wiki/Upper_limb" title="Upper limb">upper limb</a> → 1° (<a href="/wiki/Muscle_spindle" title="Muscle spindle">muscle spindles</a> → <a href="/wiki/Dorsal_root_ganglion" title="Dorsal root ganglion">DRG</a>) → 2° (<a href="/wiki/Accessory_cuneate_nucleus" title="Accessory cuneate nucleus">Accessory cuneate nucleus</a> → <b><a href="/wiki/Posterior_external_arcuate_fibers" class="mw-redirect" title="Posterior external arcuate fibers">Cuneocerebellar tract</a></b> → <a href="/wiki/Inferior_cerebellar_peduncle" title="Inferior cerebellar peduncle">ICP</a> → <a href="/wiki/Anterior_lobe_of_cerebellum" title="Anterior lobe of cerebellum">Anterior lobe of cerebellum</a>)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Reflex_arc" title="Reflex arc">Reflex arc</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lower_limb" class="mw-redirect" title="Lower limb">lower limb</a> → 1° (<a href="/wiki/Golgi_tendon_organ" title="Golgi tendon organ">Golgi tendon organ</a>) → 2° (<b><a href="/wiki/Ventral_spinocerebellar_tract" class="mw-redirect" title="Ventral spinocerebellar tract">Ventral/anterior spinocerebellar tract</a></b>→ <a href="/wiki/Superior_cerebellar_peduncle" title="Superior cerebellar peduncle">SCP</a> → <a href="/wiki/Cerebellar_vermis" title="Cerebellar vermis">Cerebellar vermis</a>)</li></ul> <ul><li><a href="/wiki/Upper_limb" title="Upper limb">upper limb</a> → 1° (<a href="/wiki/Golgi_tendon_organ" title="Golgi tendon organ">Golgi tendon organ</a>) → 2° (<b><a href="/wiki/Rostral_spinocerebellar_tract" 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