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Beschikbaar in 49 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-49" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">49 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A5%D8%AF%D9%88%D8%A7%D8%B1%D8%AF_%D8%AF%D9%88%D9%86%D8%A7%D9%84_%D8%AA%D9%88%D9%85%D8%A7%D8%B3" title="إدوارد دونال توماس – Arabisch" lang="ar" hreflang="ar" data-title="إدوارد دونال توماس" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A7%D8%AF%D9%88%D8%A7%D8%B1%D8%AF_%D8%AF%D9%88%D9%86%D8%A7%D9%84_%D8%AA%D9%88%D9%85%D8%A7%D8%B3" title="ادوارد دونال توماس – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="ادوارد دونال توماس" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%A7%D8%AF%D9%88%D8%A7%D8%B1%D8%AF_%D8%AF%D9%88%D9%86%D8%A7%D9%84_%D8%AA%D9%88%D9%85%D8%A7%D8%B3" title="ادوارد دونال توماس – South Azerbaijani" lang="azb" hreflang="azb" data-title="ادوارد دونال توماس" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%AD%D0%B4%D1%83%D0%B0%D1%80%D0%B4_%D0%94%D0%BE%D0%BD%D0%B0%D0%BB_%D0%A2%D0%BE%D0%BC%D0%B0%D1%81" title="Эдуард Донал Томас – Belarussisch" lang="be" hreflang="be" data-title="Эдуард Донал Томас" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%95%D0%B4%D1%83%D0%B0%D1%80%D0%B4_%D0%94%D0%BE%D0%BD%D0%B0%D0%BB_%D0%A2%D0%BE%D0%BC%D0%B0%D1%81" title="Едуард Донал Томас – Bulgaars" lang="bg" hreflang="bg" data-title="Едуард Донал Томас" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%8F%E0%A6%A1%E0%A6%93%E0%A6%AF%E0%A6%BC%E0%A6%BE%E0%A6%B0%E0%A7%8D%E0%A6%A1_%E0%A6%A1%E0%A7%8B%E0%A6%A8%E0%A6%BE%E0%A6%B2_%E0%A6%A5%E0%A6%AE%E0%A6%BE%E0%A6%B8" title="এডওয়ার্ড ডোনাল থমাস – Bengaals" lang="bn" hreflang="bn" data-title="এডওয়ার্ড ডোনাল থমাস" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Catalaans" lang="ca" hreflang="ca" data-title="Edward Donnall Thomas" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Tsjechisch" lang="cs" hreflang="cs" data-title="E. Donnall Thomas" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Welsh" lang="cy" hreflang="cy" data-title="E. Donnall Thomas" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Deens" lang="da" hreflang="da" data-title="E. Donnall Thomas" data-language-autonym="Dansk" data-language-local-name="Deens" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Duits" lang="de" hreflang="de" data-title="Edward Donnall Thomas" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Engels" lang="en" hreflang="en" data-title="E. Donnall Thomas" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Spaans" lang="es" hreflang="es" data-title="Edward Donnall Thomas" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Estisch" lang="et" hreflang="et" data-title="Edward Donnall Thomas" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Baskisch" lang="eu" hreflang="eu" data-title="Edward Donnall Thomas" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A7%D8%AF%D9%88%D8%A7%D8%B1%D8%AF_%D8%AF%D9%88%D9%86%D8%A7%D9%84_%D8%AA%D9%88%D9%85%D8%A7%D8%B3" title="ادوارد دونال توماس – Perzisch" lang="fa" hreflang="fa" data-title="ادوارد دونال توماس" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Fins" lang="fi" hreflang="fi" data-title="Edward Donnall Thomas" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Frans" lang="fr" hreflang="fr" data-title="Edward Donnall Thomas" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%93%D7%95%D7%A0%D7%9C_%D7%AA%D7%95%D7%9E%D7%90%D7%A1" title="דונל תומאס – Hebreeuws" lang="he" hreflang="he" data-title="דונל תומאס" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Kroatisch" lang="hr" hreflang="hr" data-title="E. Donnall Thomas" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Hongaars" lang="hu" hreflang="hu" data-title="Edward Donnall Thomas" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Donnall_Thomas" title="Donnall Thomas – Indonesisch" lang="id" hreflang="id" data-title="Donnall Thomas" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Ido" lang="io" hreflang="io" data-title="Edward Donnall Thomas" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Italiaans" lang="it" hreflang="it" data-title="Edward Donnall Thomas" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%A8%E3%83%89%E3%83%AF%E3%83%BC%E3%83%89%E3%83%BB%E3%83%89%E3%83%8A%E3%83%AB%E3%83%BB%E3%83%88%E3%83%BC%E3%83%9E%E3%82%B9" title="エドワード・ドナル・トーマス – Japans" lang="ja" hreflang="ja" data-title="エドワード・ドナル・トーマス" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%AD%D0%B4%D0%B2%D0%B0%D1%80%D0%B4_%D0%94%D0%BE%D0%BD%D0%BD%D0%B0%D0%BB%D0%BB_%D0%A2%D0%BE%D0%BC%D0%B0%D1%81" title="Эдвард Донналл Томас – Kazachs" lang="kk" hreflang="kk" data-title="Эдвард Донналл Томас" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%97%90%EB%93%9C%EC%9B%8C%EB%93%9C_%EB%8F%84%EB%84%90_%ED%86%A0%EB%A8%B8%EC%8A%A4" title="에드워드 도널 토머스 – Koreaans" lang="ko" hreflang="ko" data-title="에드워드 도널 토머스" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Latijn" lang="la" hreflang="la" data-title="E. Donnall Thomas" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%8E%E0%B4%A1%E0%B5%8D%E0%B4%B5%E0%B5%87%E0%B4%A1%E0%B5%8D_%E0%B4%A1%E0%B5%8B%E0%B4%A3%E0%B4%BE%E0%B5%BD_%E0%B4%A4%E0%B5%8B%E0%B4%AE%E0%B4%B8%E0%B5%8D" title="എഡ്വേഡ് ഡോണാൽ തോമസ് – Malayalam" lang="ml" hreflang="ml" data-title="എഡ്വേഡ് ഡോണാൽ തോമസ്" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Maleis" lang="ms" hreflang="ms" data-title="E. Donnall Thomas" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%A7%D8%AF%D9%88%D8%A7%D8%B1%D8%AF_%D8%AF%D9%88%D9%86%D8%A7%D9%84_%D8%AA%D9%88%D9%85%D8%A7%D8%B3" title="ادوارد دونال توماس – Mazanderani" lang="mzn" hreflang="mzn" data-title="ادوارد دونال توماس" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Nedersaksisch" lang="nds" hreflang="nds" data-title="Edward Donnall Thomas" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Noors - Bokmål" lang="nb" hreflang="nb" data-title="E. Donnall Thomas" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Occitaans" lang="oc" hreflang="oc" data-title="E. Donnall Thomas" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Pools" lang="pl" hreflang="pl" data-title="E. Donnall Thomas" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DA%88%D9%88%D9%86%D9%84_%D8%AA%DA%BE%D8%A7%D9%85%D8%B3" title="ڈونل تھامس – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ڈونل تھامس" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Portugees" lang="pt" hreflang="pt" data-title="Edward Donnall Thomas" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A2%D0%BE%D0%BC%D0%B0%D1%81,_%D0%AD%D0%B4%D0%B2%D0%B0%D1%80%D0%B4_%D0%94%D0%BE%D0%BD%D0%BD%D0%B0%D0%BB%D0%BB" title="Томас, Эдвард Донналл – Russisch" lang="ru" hreflang="ru" data-title="Томас, Эдвард Донналл" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Simple English" lang="en-simple" hreflang="en-simple" data-title="E. Donnall Thomas" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas – Zweeds" lang="sv" hreflang="sv" data-title="Edward Donnall Thomas" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Donnall_Thomas" title="Donnall Thomas – Swahili" lang="sw" hreflang="sw" data-title="Donnall Thomas" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Turks" lang="tr" hreflang="tr" data-title="E. Donnall Thomas" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%95%D0%B4%D0%B2%D0%B0%D1%80%D0%B4_%D0%A2%D0%BE%D0%BC%D0%B0%D1%81" title="Едвард Томас – Oekraïens" lang="uk" hreflang="uk" data-title="Едвард Томас" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%A7%DB%8C_%DA%88%D9%88%D9%86%D8%A7%D9%84%DA%88_%D8%AA%DA%BE%D8%A7%D9%85%D8%B3" title="ای ڈونالڈ تھامس – Urdu" lang="ur" hreflang="ur" data-title="ای ڈونالڈ تھامس" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Oezbeeks" lang="uz" hreflang="uz" data-title="E. Donnall Thomas" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Oezbeeks" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Vietnamees" lang="vi" hreflang="vi" data-title="E. Donnall Thomas" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%94%E1%83%93%E1%83%95%E1%83%90%E1%83%A0%E1%83%93_%E1%83%93%E1%83%9D%E1%83%9C%E1%83%90%E1%83%9A_%E1%83%A2%E1%83%9D%E1%83%9B%E1%83%90%E1%83%A1%E1%83%98" title="ედვარდ დონალ ტომასი – Mingreels" lang="xmf" hreflang="xmf" data-title="ედვარდ დონალ ტომასი" data-language-autonym="მარგალური" data-language-local-name="Mingreels" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/E._Donnall_Thomas" title="E. Donnall Thomas – Yoruba" lang="yo" hreflang="yo" data-title="E. 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href="/wiki/Bestand:Nobel_prize_medal.svg" class="mw-file-description" title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  E. Donnall Thomas </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/15_maart" title="15 maart">15 maart</a> <a href="/wiki/1920" title="1920">1920</a> – <a href="/wiki/20_oktober" title="20 oktober">20 oktober</a> <a href="/wiki/2012" title="2012">2012</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Edward_Donnall_%22Don%22_Thomas.jpg" class="mw-file-description"><img alt="Edward Donnall Thomas" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cc/Edward_Donnall_%22Don%22_Thomas.jpg/266px-Edward_Donnall_%22Don%22_Thomas.jpg" decoding="async" width="266" height="273" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/c/cc/Edward_Donnall_%22Don%22_Thomas.jpg 1.5x" data-file-width="367" data-file-height="377" /></a></span> </td></tr> <tr> <td style="text-align:center;" colspan="3"><div style="font-size:90%">Edward Donnall Thomas</div> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Verenigde_Staten" title="Verenigde Staten">Verenigde Staten</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/Mart_(Texas)" title="Mart (Texas)">Mart (Texas)</a> </td></tr> <tr> <th>Overlijdensplaats </th> <td colspan="2"><a href="/wiki/Seattle" title="Seattle">Seattle</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2">Fysiologie of Geneeskunde </td></tr> <tr> <th>Jaar </th> <td colspan="2">1990 </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor onderzoek aan orgaan- en celtransplantatie </td></tr> <tr> <th>Samen met </th> <td colspan="2"><a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Joseph Murray</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">John Michael Bishop</a><br /><a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a><br /><a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>Edward Donnall (Don) Thomas</b> (<a href="/wiki/Mart_(Texas)" title="Mart (Texas)">Mart (Texas)</a>, <a href="/wiki/15_maart" title="15 maart">15 maart</a> <a href="/wiki/1920" title="1920">1920</a> – <a href="/wiki/Seattle" title="Seattle">Seattle</a>, <a href="/wiki/20_oktober" title="20 oktober">20 oktober</a> <a href="/wiki/2012" title="2012">2012</a>) was een <a href="/wiki/Verenigde_Staten" title="Verenigde Staten">Amerikaans</a> <a href="/wiki/Arts" title="Arts">arts</a> en <a href="/wiki/Professor" title="Professor">professor</a> aan de <a href="/wiki/Universiteit_van_Washington" title="Universiteit van Washington">Universiteit van Washington</a>. In 1990 won hij samen met <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Joseph Murray</a> de <a href="/wiki/Nobelprijs_voor_de_Fysiologie_of_Geneeskunde" class="mw-redirect" title="Nobelprijs voor de Fysiologie of Geneeskunde">Nobelprijs voor de Fysiologie of Geneeskunde</a> voor hun onderzoek en ontwikkeling aan <a href="/wiki/Orgaantransplantatie" class="mw-redirect" title="Orgaantransplantatie">orgaan</a>- en <a href="/w/index.php?title=Celtransplantatie&action=edit&redlink=1" class="new" title="Celtransplantatie (de pagina bestaat niet)">celtransplantatie</a>. Thomas ontwierp <a href="/wiki/Beenmerg" title="Beenmerg">beenmergtransplantatie</a>, wat als genezing kan dienen voor <a href="/wiki/Leukemie" title="Leukemie">leukemie</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edward_Donnall_Thomas&veaction=edit&section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Edward_Donnall_Thomas&action=edit&section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>De Texaan Thomas was de zoon van de arts Edward E. Thomas en Angie Hill Donnall. Hij studeerde <a href="/wiki/Scheikunde" title="Scheikunde">scheikunde</a> en <a href="/wiki/Chemische_technologie" title="Chemische technologie">chemische technologie</a> aan de <a href="/wiki/Universiteit_van_Texas_in_Austin" title="Universiteit van Texas in Austin">Universiteit van Texas in Austin</a>. In 1941 haalde hij zijn <a href="/wiki/Bachelor_of_Arts" title="Bachelor of Arts">Bachelor of Arts</a> en in 1943 zijn <a href="/wiki/Master_(graad)" title="Master (graad)">Master</a>. Hij trouwde met Dorothy (Dottie) Martin. Samen kregen ze drie kinderen. </p><p>In 1943 ging Thomas naar <a href="/wiki/Harvard_Medical_School" title="Harvard Medical School">Harvard Medical School</a>, alwaar hij in 1946 zijn <a href="/wiki/Doctor_of_Medicine" title="Doctor of Medicine">Doctor of Medicine</a> behaalde. Aan Harvard raakte hij geïnteresseerd in <a href="/wiki/Leukemie" title="Leukemie">leukemie</a> en <a href="/wiki/Beenmerg" title="Beenmerg">beenmerg</a>. Na zijn afstuderen diende hij twee jaar in het leger waarna hij terugkeerde naar Boston om verder onderzoek te doen. In 1955 werd hij geneesheer aan het Mary Imogene Bassett Hospital (huidige Bassett Medical Center) in <a href="/wiki/Cooperstown_(New_York)" title="Cooperstown (New York)">Cooperstown</a>. In 1963 verhuisde hij naar Seattle waar hij hoogleraar werd aan de <a href="/wiki/Universiteit_van_Washington" title="Universiteit van Washington">Universiteit van Washington</a> Medical School. </p> <div class="mw-heading mw-heading2"><h2 id="Beenmergtransplantatie">Beenmergtransplantatie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Edward_Donnall_Thomas&veaction=edit&section=2" title="Bewerk dit kopje: Beenmergtransplantatie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Edward_Donnall_Thomas&action=edit&section=2" title="De broncode bewerken van de sectie: Beenmergtransplantatie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Direct na zijn aankomst in Cooperstown raakte Thomas als jong onderzoeker betrokken bij experimentele transplantaties van beenmerg op honden om de technische problemen uit te werken. Hij vermoedde dat vervanging van het beenmerg van patiënten met dat van een gezonde donor leukemie kon genezen. Rond 1957 voerde zijn team de eerste beenmergtransplantatie uit op zes patiënten nadat hij eerst hun eigen abnormale beenmerg had vernietigd met straling. De resultaten waren teleurstellend. Geen van de patiënten leefden langer dan honderd dagen door complicaties en doordat het lichaam het getransplanteerde beenmerg afstootte of door <a href="/wiki/Graft-versus-hostreactie" title="Graft-versus-hostreactie">graft-versus-hostreactie</a> waarbij de donorcellen het gastweefsel aanvallen. Alleen bij identieke tweelingen traden geen afstotingsverschijnselen op. </p><p>In 1969, na jaren van onderzoek naar <a href="/wiki/Histocompatibiliteit" class="mw-redirect" title="Histocompatibiliteit">histocompatibiliteit</a> (weefselafstoting) en de ontwikkeling van geperfectioneerde <a href="/wiki/Antibiotica" class="mw-redirect" title="Antibiotica">antibiotica</a> om infecties te onderdrukken, voerde Thomas de eerste succesvolle beenmergtransplantatie uit waarbij de donor en ontvanger geen eeneiige tweeling was. Gedurende de jaren zeventig ontwikkelde en verbeterde hij de behandelprocedure van leukemiepatiënten. In 1990 won hij hiervoor de Nobelprijs voor de Fysiologie en Geneeskunde, samen met <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Joseph Murray</a> die soortgelijke problemen overwon bij orgaantransplantaties. Hij geldbedrag van zijn Nobelprijs, zo'n 350.000 dollar, doneerde Thomas aan het Fred Hutchinson Cancer Research Center waar hij tientallen jaren werkzaam was geweest. In hetzelfde jaar, 1990, werd Thomas ook onderscheiden met de <a href="/wiki/National_Medal_of_Science" title="National Medal of Science">National Medal of Science</a>. </p> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> Fred Hutchinson Research Center: <a rel="nofollow" class="external text" href="http://www.fhcrc.org/research/nobel/thomas/dondottie.html">Nobel Prize: The Don and Dottie Story</a></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><a rel="nofollow" class="external text" href="http://nobelprize.org/medicine/laureates/1990/">Nobel Prize in Physiology or Medicine, 1990</a></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Nederlands">(<span>nl</span>) </span> <a rel="nofollow" class="external text" href="https://www.nu.nl/wetenschap/2938651/pionier-beenmergtransplantatie-overleden.html">Pionier beenmergtransplantatie overleden</a></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span> <a rel="nofollow" class="external text" href="http://seattletimes.com/html/businesstechnology/2019485178_thomasobit21m.html">Overlijdensbericht van Edward Donnall Thomas</a></li> <li><span class="citation" id="CITEREF21_oktober_2012"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><span style="font-variant:small-caps;">Denise Gellene</span>, <a rel="nofollow" class="external text" href="http://www.nytimes.com/2012/10/22/science/e-donnall-thomas-furthered-bone-marrow-transplants-dies.html">E. Donnall Thomas, Who Advanced Bone Marrow Transplants, Dies at 92</a>. The New York Times (<span class="mw-formatted-date" title="2012-10-21">21 oktober 2012</span>). Geraadpleegd op <span class="mw-formatted-date" title="2014-07-02">2 juli 2014</span>. </span></li> <li><span style="font-variant:small-caps;">Hanson, Betsy</span> (1995). "E. Donnall Thomas". <i>Notable Twentieth-Century Scientists</i>.  Detroit: Gale Research Inc.</li></ul> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output .navigatie-afb-rechts{position:absolute}.mw-parser-output .navigatie-afb-rechts{right:2px}.mw-parser-output .navigatie-afb-groot{float:right;padding-left:0.5em}.mw-parser-output .navigatie-titel{background-color:#ddeef8;padding:2px 5.5em;font-weight:bold}.mw-parser-output .navigatie-inhoud{padding:0.5em}.mw-parser-output .navigatie-inhoud p:first-child{margin:0}.mw-parser-output .navigatie div[style*="background-color"],.mw-parser-output .navigatie 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data-file-width="20" data-file-height="20" /></a></span></span></div></div> <div id="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" class="navigatie-titel">Winnaars van de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a></div> <div class="navigatie-inhoud mw-collapsible-content"> <p>1901: <a href="/wiki/Emil_Adolf_von_Behring" title="Emil Adolf von Behring">Behring</a> · 1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> · 1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> · 1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Pavlov</a> · 1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> · 1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a>, <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Ramón y Cajal</a> · 1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a class="mw-selflink selflink">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1&useformat=desktop" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Overgenomen van "<a dir="ltr" href="https://nl.wikipedia.org/w/index.php?title=Edward_Donnall_Thomas&oldid=64578034">https://nl.wikipedia.org/w/index.php?title=Edward_Donnall_Thomas&oldid=64578034</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Categorie:Alles" title="Categorie:Alles">Categorieën</a>: <ul><li><a href="/wiki/Categorie:Amerikaans_medicus" title="Categorie:Amerikaans medicus">Amerikaans medicus</a></li><li><a href="/wiki/Categorie:Winnaar_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Categorie:Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde">Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde</a></li><li><a href="/wiki/Categorie:20e-eeuws_medicus" title="Categorie:20e-eeuws medicus">20e-eeuws medicus</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> Deze pagina is voor het laatst bewerkt op 26 jun 2023 om 18:35.</li> <li id="footer-info-copyright">De tekst is beschikbaar onder de licentie <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/deed.nl">Creative Commons Naamsvermelding/Gelijk delen</a>, er kunnen aanvullende voorwaarden van toepassing zijn. 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