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James Whyte Black - Wikipedia
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Beschikbaar in 57 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-57" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">57 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AC%D9%8A%D9%85%D8%B3_%D8%A8%D9%84%D8%A7%D9%83_(%D8%B9%D8%A7%D9%84%D9%85)" title="جيمس بلاك (عالم) – Arabisch" lang="ar" hreflang="ar" data-title="جيمس بلاك (عالم)" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AC%D9%8A%D9%85%D8%B3_%D8%A8%D9%84%D8%A7%D9%83_(%D8%AF%D9%83%D8%AA%D9%88%D8%B1_%D9%85%D9%86_%D8%A7%D9%84%D9%85%D9%85%D9%84%D9%83%D9%87_%D8%A7%D9%84%D9%85%D8%AA%D8%AD%D8%AF%D9%87)" title="جيمس بلاك (دكتور من المملكه المتحده) – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="جيمس بلاك (دكتور من المملكه المتحده)" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Ceyms_Blek" title="Ceyms Blek – Azerbeidzjaans" lang="az" hreflang="az" data-title="Ceyms Blek" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbeidzjaans" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%AC%DB%8C%D9%85%D8%B2_%D8%A8%D9%84%DA%A9" title="جیمز بلک – South Azerbaijani" lang="azb" hreflang="azb" data-title="جیمز بلک" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%94%D0%B6%D1%8D%D0%B9%D0%BC%D1%81_%D0%A3%D0%B0%D0%B9%D1%82_%D0%91%D0%BB%D1%8D%D0%BA" title="Джэймс Уайт Блэк – Belarussisch" lang="be" hreflang="be" data-title="Джэймс Уайт Блэк" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%94%D0%B6%D0%B5%D0%B9%D0%BC%D1%81_%D0%91%D0%BB%D0%B5%D0%BA" title="Джеймс Блек – Bulgaars" lang="bg" hreflang="bg" data-title="Джеймс Блек" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%9C%E0%A7%87%E0%A6%AE%E0%A6%B8_%E0%A6%B9%E0%A7%8B%E0%A6%AF%E0%A6%BC%E0%A6%BE%E0%A6%87%E0%A6%9F_%E0%A6%AC%E0%A7%8D%E0%A6%B2%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%95" title="জেমস হোয়াইট ব্ল্যাক – Bengaals" lang="bn" hreflang="bn" data-title="জেমস হোয়াইট ব্ল্যাক" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Catalaans" lang="ca" hreflang="ca" data-title="James Whyte Black" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D8%AC%DB%95%DB%8C%D9%85%D8%B2_%D8%A8%D9%84%D8%A7%DA%A9" title="جەیمز بلاک – Soranî" lang="ckb" hreflang="ckb" data-title="جەیمز بلاک" data-language-autonym="کوردی" data-language-local-name="Soranî" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/James_W._Black" title="James W. Black – Tsjechisch" lang="cs" hreflang="cs" data-title="James W. Black" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/James_Black" title="James Black – Welsh" lang="cy" hreflang="cy" data-title="James Black" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/James_W._Black" title="James W. Black – Deens" lang="da" hreflang="da" data-title="James W. Black" data-language-autonym="Dansk" data-language-local-name="Deens" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Duits" lang="de" hreflang="de" data-title="James Whyte Black" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist) – Engels" lang="en" hreflang="en" data-title="James Black (pharmacologist)" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/James_Black" title="James Black – Spaans" lang="es" hreflang="es" data-title="James Black" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/James_Black" title="James Black – Estisch" lang="et" hreflang="et" data-title="James Black" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/James_W._Black" title="James W. Black – Baskisch" lang="eu" hreflang="eu" data-title="James W. Black" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AC%DB%8C%D9%85%D8%B2_%D8%A8%D9%84%DA%A9_(%D8%AF%D8%A7%D8%B1%D9%88%D8%B4%D9%86%D8%A7%D8%B3)" title="جیمز بلک (داروشناس) – Perzisch" lang="fa" hreflang="fa" data-title="جیمز بلک (داروشناس)" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/James_W._Black" title="James W. Black – Fins" lang="fi" hreflang="fi" data-title="James W. Black" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Frans" lang="fr" hreflang="fr" data-title="James Whyte Black" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Iers" lang="ga" hreflang="ga" data-title="James Whyte Black" data-language-autonym="Gaeilge" data-language-local-name="Iers" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gd mw-list-item"><a href="https://gd.wikipedia.org/wiki/James_W._Black" title="James W. Black – Schots-Gaelisch" lang="gd" hreflang="gd" data-title="James W. Black" data-language-autonym="Gàidhlig" data-language-local-name="Schots-Gaelisch" class="interlanguage-link-target"><span>Gàidhlig</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Galicisch" lang="gl" hreflang="gl" data-title="James Whyte Black" data-language-autonym="Galego" data-language-local-name="Galicisch" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%92%27%D7%99%D7%99%D7%9E%D7%A1_%D7%95%D7%95._%D7%91%D7%9C%D7%90%D7%A7" title="ג'יימס וו. בלאק – Hebreeuws" lang="he" hreflang="he" data-title="ג'יימס וו. בלאק" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/James_W._Black" title="James W. Black – Kroatisch" lang="hr" hreflang="hr" data-title="James W. Black" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/James_W._Black" title="James W. Black – Hongaars" lang="hu" hreflang="hu" data-title="James W. Black" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Indonesisch" lang="id" hreflang="id" data-title="James Whyte Black" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Ido" lang="io" hreflang="io" data-title="James Whyte Black" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/James_W._Black" title="James W. Black – Italiaans" lang="it" hreflang="it" data-title="James W. Black" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B8%E3%82%A7%E3%83%BC%E3%83%A0%E3%82%B9%E3%83%BB%E3%83%96%E3%83%A9%E3%83%83%E3%82%AF" title="ジェームス・ブラック – Japans" lang="ja" hreflang="ja" data-title="ジェームス・ブラック" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%94%D0%B6%D0%B5%D0%B9%D0%BC%D1%81_%D0%91%D0%BB%D1%8D%D0%BA" title="Джеймс Блэк – Kazachs" lang="kk" hreflang="kk" data-title="Джеймс Блэк" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%A0%9C%EC%9E%84%EC%8A%A4_%ED%99%94%EC%9D%B4%ED%8A%B8_%EB%B8%94%EB%9E%99" title="제임스 화이트 블랙 – Koreaans" lang="ko" hreflang="ko" data-title="제임스 화이트 블랙" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Iacobus_Whyte_Black" title="Iacobus Whyte Black – Latijn" lang="la" hreflang="la" data-title="Iacobus Whyte Black" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-mg mw-list-item"><a href="https://mg.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Malagassisch" lang="mg" hreflang="mg" data-title="James Whyte Black" data-language-autonym="Malagasy" data-language-local-name="Malagassisch" class="interlanguage-link-target"><span>Malagasy</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/James_W._Black" title="James W. Black – Maleis" lang="ms" hreflang="ms" data-title="James W. Black" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%DB%8C%D9%85%D8%B2_%D8%A8%D9%84%DA%A9_(%D8%AF%D8%A7%D8%B1%D9%88%D8%B4%D9%86%D8%A7%D8%B3)" title="جیمز بلک (داروشناس) – Mazanderani" lang="mzn" hreflang="mzn" data-title="جیمز بلک (داروشناس)" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Nedersaksisch" lang="nds" hreflang="nds" data-title="James Whyte Black" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/James_W._Black" title="James W. Black – Noors - Bokmål" lang="nb" hreflang="nb" data-title="James W. Black" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/James_W._Black" title="James W. Black – Occitaans" lang="oc" hreflang="oc" data-title="James W. Black" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/James_W._Black" title="James W. Black – Pools" lang="pl" hreflang="pl" data-title="James W. Black" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%DB%8C%D9%85%D8%B2_%D8%A8%D9%84%DB%8C%DA%A9" title="جیمز بلیک – Western Punjabi" lang="pnb" hreflang="pnb" data-title="جیمز بلیک" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/James_Black" title="James Black – Portugees" lang="pt" hreflang="pt" data-title="James Black" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/James_W._Black" title="James W. Black – Roemeens" lang="ro" hreflang="ro" data-title="James W. Black" data-language-autonym="Română" data-language-local-name="Roemeens" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%91%D0%BB%D1%8D%D0%BA,_%D0%94%D0%B6%D0%B5%D0%B9%D0%BC%D1%81" title="Блэк, Джеймс – Russisch" lang="ru" hreflang="ru" data-title="Блэк, Джеймс" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/James_Black" title="James Black – Simple English" lang="en-simple" hreflang="en-simple" data-title="James Black" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/James_W._Black" title="James W. Black – Sloveens" lang="sl" hreflang="sl" data-title="James W. Black" data-language-autonym="Slovenščina" data-language-local-name="Sloveens" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/James_Black" title="James Black – Zweeds" lang="sv" hreflang="sv" data-title="James Black" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/James_Black" title="James Black – Swahili" lang="sw" hreflang="sw" data-title="James Black" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/James_W._Black" title="James W. Black – Turks" lang="tr" hreflang="tr" data-title="James W. Black" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D2%96%D0%B5%D0%B9%D0%BC%D1%81_%D0%91%D0%BB%D3%99%D0%BA" title="Җеймс Бләк – Tataars" lang="tt" hreflang="tt" data-title="Җеймс Бләк" data-language-autonym="Татарча / tatarça" data-language-local-name="Tataars" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%94%D0%B6%D0%B5%D0%B9%D0%BC%D1%81_%D0%92%D0%B0%D0%B9%D1%82_%D0%91%D0%BB%D0%B5%D0%BA" title="Джеймс Вайт Блек – Oekraïens" lang="uk" hreflang="uk" data-title="Джеймс Вайт Блек" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%AC%DB%8C%D9%85%D8%B2_%D8%A8%D9%84%DB%8C%DA%A9_(%D8%A7%D8%AF%D9%88%DB%8C%D8%A7%D8%AA%D8%AF%D8%A7%D9%86)" title="جیمز بلیک (ادویاتدان) – Urdu" lang="ur" hreflang="ur" data-title="جیمز بلیک (ادویاتدان)" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/James_W._Black" title="James W. Black – Vietnamees" lang="vi" hreflang="vi" data-title="James W. Black" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-vo mw-list-item"><a href="https://vo.wikipedia.org/wiki/James_Whyte_Black" title="James Whyte Black – Volapük" lang="vo" hreflang="vo" data-title="James Whyte Black" data-language-autonym="Volapük" data-language-local-name="Volapük" class="interlanguage-link-target"><span>Volapük</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%AF%E1%83%94%E1%83%98%E1%83%9B%E1%83%A1_%E1%83%91%E1%83%9A%E1%83%94%E1%83%99%E1%83%98" title="ჯეიმს ბლეკი – Mingreels" lang="xmf" hreflang="xmf" data-title="ჯეიმს ბლეკი" data-language-autonym="მარგალური" data-language-local-name="Mingreels" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/James_W._Black" title="James W. Black – Yoruba" lang="yo" hreflang="yo" data-title="James W. 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title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  James Whyte Black </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/14_juli" title="14 juli">14 juli</a> <a href="/wiki/1924" title="1924">1924</a> – <a href="/wiki/22_maart" title="22 maart">22 maart</a> <a href="/wiki/2010" title="2010">2010</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:James_Black_(pharmacologist).jpg" class="mw-file-description"><img alt="James Whyte Black" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/69/James_Black_%28pharmacologist%29.jpg/266px-James_Black_%28pharmacologist%29.jpg" decoding="async" width="266" height="235" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/69/James_Black_%28pharmacologist%29.jpg/399px-James_Black_%28pharmacologist%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/69/James_Black_%28pharmacologist%29.jpg/532px-James_Black_%28pharmacologist%29.jpg 2x" data-file-width="4065" data-file-height="3592" /></a></span> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Schotland" title="Schotland">Schotland</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/w/index.php?title=Uddingston&action=edit&redlink=1" class="new" title="Uddingston (de pagina bestaat niet)">Uddingston</a> </td></tr> <tr> <th>Overlijdensplaats </th> <td colspan="2">Londen </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2">Fysiologie of Geneeskunde </td></tr> <tr> <th>Jaar </th> <td colspan="2">1988 </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor hun ontdekkingen van belangrijke principes voor de <a href="/wiki/Farmacologie" title="Farmacologie">farmacologie</a> </td></tr> <tr> <th>Samen met </th> <td colspan="2"><a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Gertrude Elion</a><br /><a href="/wiki/George_Hitchings" class="mw-redirect" title="George Hitchings">George Hitchings</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">John Michael Bishop</a><br /><a href="/wiki/Harold_Varmus" class="mw-redirect" title="Harold Varmus">Harold Varmus</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>James Whyte Black</b> (<a href="/w/index.php?title=Uddingston&action=edit&redlink=1" class="new" title="Uddingston (de pagina bestaat niet)">Uddingston</a>, <a href="/wiki/14_juli" title="14 juli">14 juli</a> <a href="/wiki/1924" title="1924">1924</a> – <a href="/wiki/Londen" title="Londen">Londen</a>, <a href="/wiki/22_maart" title="22 maart">22 maart</a> <a href="/wiki/2010" title="2010">2010</a>) was een <a href="/wiki/Schotland" title="Schotland">Schotse</a> farmacoloog, die de <a href="/wiki/B%C3%A8tablokkers" title="Bètablokkers">bètablokker</a> <a href="/wiki/Propranolol" title="Propranolol">propranolol</a> en <a href="/wiki/H2-receptorantagonist" title="H2-receptorantagonist">H2-receptorantagonist</a> <a href="/wiki/Cimetidine" title="Cimetidine">cimetidine</a> voor het eerst heeft gesynthetiseerd. Hij kreeg voor deze uitvindingen in 1988 de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a>.<sup id="cite_ref-nobel_1-0" class="reference"><a href="#cite_note-nobel-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=James_Whyte_Black&veaction=edit&section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=James_Whyte_Black&action=edit&section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Black was de jongste van vier zonen van een Schotse mijnwerker die zich had opgewekt tot mijnbouwingenieur. Blacks academische carrière begon aan de <a href="/wiki/Universiteit_van_St_Andrews" title="Universiteit van St Andrews">universiteit van St Andrews</a>, waar hij geneeskunde studeerde. Hij maakte deel uit van de vakgroep fysiologie, voordat hij een positie als docent aan de <a href="/w/index.php?title=Universiteit_van_Malaya&action=edit&redlink=1" class="new" title="Universiteit van Malaya (de pagina bestaat niet)">universiteit van Malaya</a> in <a href="/wiki/Singapore" title="Singapore">Singapore</a> aannam. In 1950 richtte hij de vakgroep fysiologie aan de universiteit van Glasgow (dierenartsenschool) op. Tijdens zijn loopbaan werkte hij zowel in de wetenschap als in het bedrijfsleven. </p><p>Hij werkte bij <a href="/wiki/ICI" title="ICI">ICI Pharmaceuticals</a> (1958-1964), <a href="/w/index.php?title=Smith,_Kline_and_French&action=edit&redlink=1" class="new" title="Smith, Kline and French (de pagina bestaat niet)">Smith, Kline and French</a> (1964-1973) en de <a href="/w/index.php?title=Wellcome_Foundation&action=edit&redlink=1" class="new" title="Wellcome Foundation (de pagina bestaat niet)">Wellcome Foundation</a> (1978-1984) en werd in 1973 aangesteld als hoogleraar in de <a href="/wiki/Farmacologie" title="Farmacologie">farmacologie</a> aan het <a href="/wiki/University_College_London" title="University College London">University College London</a> (tot 1978). Van 1984 tot 1992 was hij professor aan het <a href="/wiki/King%27s_College_London" title="King's College London">King's College London</a>. Na zijn pensionering in 1992 was hij tot 2006 Chancellor aan de Dundee-universiteit. </p><p>James Black was sinds 1944 gehuwd met Hillery Vaughen, met wie hij een dochter, Stefanie, kreeg. Na haar overlijden in 1989 hertrouwde hij vijf jaar later met Rona McLeod Mackie. Hij overleed op 85-jarige leeftijd in Londen. </p> <div class="mw-heading mw-heading2"><h2 id="Erkenning">Erkenning</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=James_Whyte_Black&veaction=edit&section=2" title="Bewerk dit kopje: Erkenning" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=James_Whyte_Black&action=edit&section=2" title="De broncode bewerken van de sectie: Erkenning"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In 1976 werd hij lid van de <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a> en in hetzelfde jaar kreeg hij de <a href="/wiki/Lasker-DeBakey_Clinical_Medical_Research_Award" title="Lasker-DeBakey Clinical Medical Research Award">Lasker-DeBakey Clinical Medical Research Award</a>.<sup id="cite_ref-lasker_2-0" class="reference"><a href="#cite_note-lasker-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> In 1979 werd hem de <a href="/wiki/Baillet_Latour_Gezondheidsprijs" title="Baillet Latour Gezondheidsprijs">Baillet Latour Gezondheidsprijs</a> toegekend. Hij werd geridderd in 1981 en sindsdien mag hij als <i><a href="/wiki/Sir_(titel)" title="Sir (titel)">sir</a></i> benoemd worden. In 2000 werd hij door <a href="/wiki/Elizabeth_II_van_het_Verenigd_Koninkrijk" title="Elizabeth II van het Verenigd Koninkrijk">Elizabeth II</a> benoemd tot lid van de <a href="/wiki/Order_of_Merit_(Verenigd_Koninkrijk)" title="Order of Merit (Verenigd Koninkrijk)">Order of Merit</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=James_Whyte_Black&veaction=edit&section=3" title="Bewerk dit kopje: Werk" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=James_Whyte_Black&action=edit&section=3" title="De broncode bewerken van de sectie: Werk"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Black droeg bij aan de basis-wetenschappelijke en klinische kennis op het gebied van de <a href="/wiki/Cardiologie" title="Cardiologie">cardiologie</a>. In het onderzoek dat leidde tot propranolol kwam Black een scriptie uit 1948 tegen van de Amerikaanse wetenschapper <a href="/w/index.php?title=Raymond_Ahlquist&action=edit&redlink=1" class="new" title="Raymond Ahlquist (de pagina bestaat niet)">Raymond Ahlquist</a>, die stelde dat er twee verschillende receptoren, alfa- en bètareceptoren, zijn die een verschillend respons veroorzaakten op de hartspier. Black focuste zich op de bètareceptoren, eiwitmoleculen op het hart die door de stresshormonen <a href="/wiki/Adrenaline" title="Adrenaline">adrenaline</a> en <a href="/wiki/Noradrenaline" title="Noradrenaline">noradrenaline</a> worden gebonden om het hart sneller te laten werken. Black onderzocht diverse chemische stoffen om deze receptoren te blokkeren zonder ze te stimuleren met als resultaat de ontdekking van de bètablokker propranolol. </p><p>De ontdekking van propranolol midden jaren vijftig revolutioneerde de behandeling van <i><a href="/wiki/Angina_pectoris" title="Angina pectoris">angina pectoris</a></i> (hartkramp) en andere hartziekten, zoals hoge bloeddruk en hartritmestoornissen. Het wordt als een van de belangrijkste ontdekkingen van de 20e eeuw op medisch gebied beschouwd.<sup id="cite_ref-propranolol_3-0" class="reference"><a href="#cite_note-propranolol-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-tagamet_4-0" class="reference"><a href="#cite_note-tagamet-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-scotpaper_5-0" class="reference"><a href="#cite_note-scotpaper-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>Ook ontwikkelde hij <a href="/wiki/Cimetidine" title="Cimetidine">cimetidine</a>, een H2-receptorantagonist die de aanmaak van <a href="/wiki/Maagzuur" title="Maagzuur">zuur</a> in de maag verminderd en daarom gebruikt wordt voor de behandeling van <a href="/wiki/Zuurbrand" class="mw-redirect" title="Zuurbrand">zuurbranden</a> en <a href="/wiki/Maagzweer" title="Maagzweer">maagzweren</a>. Pas nadat in de jaren 1980 twee australische onderzoekers, <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> en <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a>, ontdekten dat een bacteriële infectie van de <i><a href="/wiki/Helicobacter_pylori" title="Helicobacter pylori">Helicobacter pylori</a></i> de oorzaak was van maagzweren in de behandeling ervan overgenomen door antibiotica. </p> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li><span class="citation" id="CITEREFAltmanFout:_ongeldige_tijd."><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><span style="font-variant:small-caps;">Altman, Lawrence K.</span>, <a rel="nofollow" class="external text" href="http://www.nytimes.com/2010/03/23/health/23black.html">Dr. James Black, Pharmacologist Who Discovered Beta Blockers, Dies at 85</a>. The New York Times (<span class="mw-formatted-date" title="2010-03-22">22 maart 2010</span>). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150403021850/http://www.nytimes.com/2010/03/23/health/23black.html">Gearchiveerd</a> op <span class="mw-formatted-date" title="2015-04-03">3 april 2015</span>. Geraadpleegd op <span class="mw-formatted-date" title="2015-03-08">8 maart 2015</span>. </span></li> <li><span style="font-variant:small-caps;">Fricker, Janet</span> (2010). "Bètablokkers". <i>1001 uitvindingen die onze wereld veranderd hebben</i>.  Red. Jack Challoner. Libero, Kerkdriel. p. 736. <span class="ISBN"><a href="/wiki/Speciaal:Boekbronnen/978-90-5764-460-3" title="Speciaal:Boekbronnen/978-90-5764-460-3">ISBN 978-90-5764-460-3 </a></span>.</li> <li><span style="font-variant:small-caps;">George, Karyn Hede</span> (1995). "James Black". <i>Notable Twentieth-Century Scientists</i>.  Detroit: Gale Research Inc. <span class="ISBN"><a href="/wiki/Speciaal:Boekbronnen/978-0810391819" title="Speciaal:Boekbronnen/978-0810391819">ISBN 978-0810391819 </a></span>.</li></ul> <div class="reflist" style="list-style-type: decimal;"><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-nobel-1"><span class="mw-cite-backlink"><a href="#cite_ref-nobel_1-0">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF1988"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/1988/black/biographical/">Sir James W. Black – Biographical</a>. NobelPrize.org (1988). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230328160935/https://www.nobelprize.org/prizes/medicine/1988/black/biographical/">Gearchiveerd</a> op <span class="mw-formatted-date" title="2023-03-28">28 maart 2023</span>. </span></span> </li> <li id="cite_note-lasker-2"><span class="mw-cite-backlink"><a href="#cite_ref-lasker_2-0">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF1976"><a rel="nofollow" class="external text" href="http://www.laskerfoundation.org/awards/1976clinical.htm">1976 winners: Albert Lasker Award for Clinical Medical Research</a>. Lasker Medical Research Network (1976). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150919100525/http://www.laskerfoundation.org/awards/1976clinical.htm">Gearchiveerd</a> op <span class="mw-formatted-date" title="2015-09-19">19 september 2015</span>. </span></span> </li> <li id="cite_note-propranolol-3"><span class="mw-cite-backlink"><a href="#cite_ref-propranolol_3-0">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF1997"><span style="font-variant:small-caps;">Melanie Patricia Stapleton</span>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/9456487">Sir James Black and Propranolol</a>. Texas Heart Institute Journal (1997). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20190606050657/https://www.ncbi.nlm.nih.gov/pubmed/9456487">Gearchiveerd</a> op <span class="mw-formatted-date" title="2019-06-06">6 juni 2019</span>. </span></span> </li> <li id="cite_note-tagamet-4"><span class="mw-cite-backlink"><a href="#cite_ref-tagamet_4-0">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF2005"><a rel="nofollow" class="external text" href="https://pubs.acs.org/cen/coverstory/83/8325/8325tagamet.html">"anTAGonist" and "ciMETidine"</a>. American Chemical Society (2005). Geraadpleegd op <span class="mw-formatted-date" title="2005-12-25">25 december 2005</span>. </span></span> </li> <li id="cite_note-scotpaper-5"><span class="mw-cite-backlink"><a href="#cite_ref-scotpaper_5-0">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF2004"><span style="font-variant:small-caps;">Alan Taylor</span>, <a rel="nofollow" class="external text" href="http://findarticles.com/p/articles/mi_qn4156/is_20040125/ai_n9627859">Or is this our national hero?</a>. Sunday Herald (2004). Geraadpleegd op <span class="mw-formatted-date" title="2004-01-25">25 januari 2004</span>. </span></span> </li> </ol></div></div> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output .navigatie-afb-rechts{position:absolute}.mw-parser-output .navigatie-afb-rechts{right:2px}.mw-parser-output .navigatie-afb-groot{float:right;padding-left:0.5em}.mw-parser-output .navigatie-titel{background-color:#ddeef8;padding:2px 5.5em;font-weight:bold}.mw-parser-output .navigatie-inhoud{padding:0.5em}.mw-parser-output .navigatie-inhoud 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title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a class="mw-selflink selflink">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div> <!-- NewPP limit report Parsed by mw‐web.eqiad.main‐5c8dd59f85‐45gjr Cached time: 20241211130942 Cache expiry: 2592000 Reduced expiry: false Complications: [] CPU time usage: 0.181 seconds Real time usage: 0.490 seconds Preprocessor visited node count: 3804/1000000 Post‐expand include size: 39352/2097152 bytes Template argument size: 17745/2097152 bytes Highest expansion depth: 14/100 Expensive parser function count: 2/500 Unstrip recursion depth: 0/20 Unstrip post‐expand size: 5605/5000000 bytes Lua time usage: 0.008/10.000 seconds Lua memory usage: 952559/52428800 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 178.370 1 -total 44.63% 79.599 1 Sjabloon:Infobox_Nobelprijswinnaar 42.44% 75.692 1 Sjabloon:Infobox_generiek 41.12% 73.342 1 Sjabloon:Appendix 27.09% 48.314 6 Sjabloon:Citeer_web 21.23% 37.866 1 Sjabloon:Wikidata_afbeelding 14.99% 26.735 1 Sjabloon:References 11.79% 21.032 1 Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_de_Fysiologie_of_Geneeskunde 8.82% 15.737 1 Sjabloon:Navigatie_uitklapbaar 8.61% 15.359 1 Sjabloon:PAGENAMEBASE --> <!-- Saved in parser cache with key nlwiki:pcache:976823:|#|:idhash:dateformat=default and timestamp 20241211130942 and revision id 68459665. 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