CINXE.COM

<!DOCTYPE html> <html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available" lang="en" dir="ltr"> <head> <meta charset="UTF-8"> <title>Richard J. Roberts - Wikipedia</title> <script>(function(){var className="client-js vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available";var cookie=document.cookie.match(/(?:^|; )enwikimwclientpreferences=([^;]+)/);if(cookie){cookie[1].split('%2C').forEach(function(pref){className=className.replace(new RegExp('(^| )'+pref.replace(/-clientpref-\w+$|[^\w-]+/g,'')+'-clientpref-\\w+( |$)'),'$1'+pref+'$2');});}document.documentElement.className=className;}());RLCONF={"wgBreakFrames":false,"wgSeparatorTransformTable":["",""],"wgDigitTransformTable":["",""],"wgDefaultDateFormat":"dmy", "wgMonthNames":["","January","February","March","April","May","June","July","August","September","October","November","December"],"wgRequestId":"b6049bbf-b1e7-4f2a-aabd-918e22706a91","wgCanonicalNamespace":"","wgCanonicalSpecialPageName":false,"wgNamespaceNumber":0,"wgPageName":"Richard_J._Roberts","wgTitle":"Richard J. Roberts","wgCurRevisionId":1254328414,"wgRevisionId":1254328414,"wgArticleId":1700298,"wgIsArticle":true,"wgIsRedirect":false,"wgAction":"view","wgUserName":null,"wgUserGroups":["*"],"wgCategories":["CS1 maint: bot: original URL status unknown","Pages containing links to subscription-only content","All articles with dead external links","Articles with dead external links from August 2016","Articles with short description","Short description is different from Wikidata","Use British English from August 2011","Use dmy dates from December 2020","Articles with hCards","Nobelprize template using Wikidata property P8024","People from Bath, Somerset","Scientists from Derby", "Nobel laureates in Physiology or Medicine","British biochemists","20th-century British biologists","21st-century British biologists","British Nobel laureates","Members of the European Molecular Biology Organization","Alumni of the University of Sheffield","Knights Bachelor","People educated at City of Bath Boys' School","1943 births","Living people","Fellows of the Royal Society","English Nobel laureates","English humanists","English atheists"],"wgPageViewLanguage":"en","wgPageContentLanguage":"en","wgPageContentModel":"wikitext","wgRelevantPageName":"Richard_J._Roberts","wgRelevantArticleId":1700298,"wgIsProbablyEditable":true,"wgRelevantPageIsProbablyEditable":true,"wgRestrictionEdit":[],"wgRestrictionMove":[],"wgNoticeProject":"wikipedia","wgCiteReferencePreviewsActive":false,"wgFlaggedRevsParams":{"tags":{"status":{"levels":1}}},"wgMediaViewerOnClick":true,"wgMediaViewerEnabledByDefault":true,"wgPopupsFlags":0,"wgVisualEditor":{"pageLanguageCode":"en","pageLanguageDir":"ltr", "pageVariantFallbacks":"en"},"wgMFDisplayWikibaseDescriptions":{"search":true,"watchlist":true,"tagline":false,"nearby":true},"wgWMESchemaEditAttemptStepOversample":false,"wgWMEPageLength":20000,"wgRelatedArticlesCompat":[],"wgCentralAuthMobileDomain":false,"wgEditSubmitButtonLabelPublish":true,"wgULSPosition":"interlanguage","wgULSisCompactLinksEnabled":false,"wgVector2022LanguageInHeader":true,"wgULSisLanguageSelectorEmpty":false,"wgWikibaseItemId":"Q309816","wgCheckUserClientHintsHeadersJsApi":["brands","architecture","bitness","fullVersionList","mobile","model","platform","platformVersion"],"GEHomepageSuggestedEditsEnableTopics":true,"wgGETopicsMatchModeEnabled":false,"wgGEStructuredTaskRejectionReasonTextInputEnabled":false,"wgGELevelingUpEnabledForUser":false};RLSTATE={"ext.globalCssJs.user.styles":"ready","site.styles":"ready","user.styles":"ready","ext.globalCssJs.user":"ready","user":"ready","user.options":"loading","ext.cite.styles":"ready","skins.vector.search.codex.styles": "ready","skins.vector.styles":"ready","skins.vector.icons":"ready","jquery.makeCollapsible.styles":"ready","ext.wikimediamessages.styles":"ready","ext.visualEditor.desktopArticleTarget.noscript":"ready","ext.uls.interlanguage":"ready","wikibase.client.init":"ready","ext.wikimediaBadges":"ready"};RLPAGEMODULES=["ext.cite.ux-enhancements","site","mediawiki.page.ready","jquery.makeCollapsible","mediawiki.toc","skins.vector.js","ext.centralNotice.geoIP","ext.centralNotice.startUp","ext.gadget.ReferenceTooltips","ext.gadget.switcher","ext.urlShortener.toolbar","ext.centralauth.centralautologin","mmv.bootstrap","ext.popups","ext.visualEditor.desktopArticleTarget.init","ext.visualEditor.targetLoader","ext.echo.centralauth","ext.eventLogging","ext.wikimediaEvents","ext.navigationTiming","ext.uls.interface","ext.cx.eventlogging.campaigns","ext.cx.uls.quick.actions","wikibase.client.vector-2022","ext.checkUser.clientHints","ext.quicksurveys.init","ext.growthExperiments.SuggestedEditSession", "wikibase.sidebar.tracking"];</script> <script>(RLQ=window.RLQ||[]).push(function(){mw.loader.impl(function(){return["user.options@12s5i",function($,jQuery,require,module){mw.user.tokens.set({"patrolToken":"+\\","watchToken":"+\\","csrfToken":"+\\"}); }];});});</script> <link rel="stylesheet" href="/w/load.php?lang=en&modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cext.wikimediamessages.styles%7Cjquery.makeCollapsible.styles%7Cskins.vector.icons%2Cstyles%7Cskins.vector.search.codex.styles%7Cwikibase.client.init&only=styles&skin=vector-2022"> <script async="" src="/w/load.php?lang=en&modules=startup&only=scripts&raw=1&skin=vector-2022"></script> <meta name="ResourceLoaderDynamicStyles" content=""> <link rel="stylesheet" href="/w/load.php?lang=en&modules=site.styles&only=styles&skin=vector-2022"> <meta name="generator" content="MediaWiki 1.44.0-wmf.4"> <meta name="referrer" content="origin"> <meta name="referrer" content="origin-when-cross-origin"> <meta name="robots" content="max-image-preview:standard"> <meta name="format-detection" content="telephone=no"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/1/19/Roberts%2C_Richard_John_%281943%29.jpg"> <meta property="og:image:width" content="1200"> <meta property="og:image:height" content="1608"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/1/19/Roberts%2C_Richard_John_%281943%29.jpg"> <meta property="og:image:width" content="800"> <meta property="og:image:height" content="1072"> <meta property="og:image:width" content="640"> <meta property="og:image:height" content="858"> <meta name="viewport" content="width=1120"> <meta property="og:title" content="Richard J. Roberts - Wikipedia"> <meta property="og:type" content="website"> <link rel="preconnect" href="//upload.wikimedia.org"> <link rel="alternate" media="only screen and (max-width: 640px)" href="//en.m.wikipedia.org/wiki/Richard_J._Roberts"> <link rel="alternate" type="application/x-wiki" title="Edit this page" href="/w/index.php?title=Richard_J._Roberts&action=edit"> <link rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png"> <link rel="icon" href="/static/favicon/wikipedia.ico"> <link rel="search" type="application/opensearchdescription+xml" href="/w/rest.php/v1/search" title="Wikipedia (en)"> <link rel="EditURI" type="application/rsd+xml" href="//en.wikipedia.org/w/api.php?action=rsd"> <link rel="canonical" href="https://en.wikipedia.org/wiki/Richard_J._Roberts"> <link rel="license" href="https://creativecommons.org/licenses/by-sa/4.0/deed.en"> <link rel="alternate" type="application/atom+xml" title="Wikipedia Atom feed" href="/w/index.php?title=Special:RecentChanges&feed=atom"> <link rel="dns-prefetch" href="//meta.wikimedia.org" /> <link rel="dns-prefetch" href="//login.wikimedia.org"> </head> <body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject mw-editable page-Richard_J_Roberts rootpage-Richard_J_Roberts skin-vector-2022 action-view"><a class="mw-jump-link" href="#bodyContent">Jump to content</a> <div class="vector-header-container"> <header class="vector-header mw-header"> <div class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="Main menu" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">Main menu</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">Main menu</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">hide</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> Navigation </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage-description" class="mw-list-item"><a href="/wiki/Main_Page" title="Visit the main page [z]" accesskey="z"><span>Main page</span></a></li><li id="n-contents" class="mw-list-item"><a href="/wiki/Wikipedia:Contents" title="Guides to browsing Wikipedia"><span>Contents</span></a></li><li id="n-currentevents" class="mw-list-item"><a href="/wiki/Portal:Current_events" title="Articles related to current events"><span>Current events</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/Special:Random" title="Visit a randomly selected article [x]" accesskey="x"><span>Random article</span></a></li><li id="n-aboutsite" class="mw-list-item"><a href="/wiki/Wikipedia:About" title="Learn about Wikipedia and how it works"><span>About Wikipedia</span></a></li><li id="n-contactpage" class="mw-list-item"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us" title="How to contact Wikipedia"><span>Contact us</span></a></li> </ul> </div> </div> <div id="p-interaction" class="vector-menu mw-portlet mw-portlet-interaction" > <div class="vector-menu-heading"> Contribute </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-help" class="mw-list-item"><a href="/wiki/Help:Contents" title="Guidance on how to use and edit Wikipedia"><span>Help</span></a></li><li id="n-introduction" class="mw-list-item"><a href="/wiki/Help:Introduction" title="Learn how to edit Wikipedia"><span>Learn to edit</span></a></li><li id="n-portal" class="mw-list-item"><a href="/wiki/Wikipedia:Community_portal" title="The hub for editors"><span>Community portal</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/Special:RecentChanges" title="A list of recent changes to Wikipedia [r]" accesskey="r"><span>Recent changes</span></a></li><li id="n-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_upload_wizard" title="Add images or other media for use on Wikipedia"><span>Upload file</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/Main_Page" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Wikipedia" src="/static/images/mobile/copyright/wikipedia-wordmark-en.svg" style="width: 7.5em; height: 1.125em;"> <img class="mw-logo-tagline" alt="The Free Encyclopedia" src="/static/images/mobile/copyright/wikipedia-tagline-en.svg" width="117" height="13" style="width: 7.3125em; height: 0.8125em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/Special:Search" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Search Wikipedia [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>Search</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Search Wikipedia" aria-label="Search Wikipedia" autocapitalize="sentences" title="Search Wikipedia [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="Special:Search"> </div> <button class="cdx-button cdx-search-input__end-button">Search</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="Personal tools"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="Change the appearance of the page's font size, width, and color" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="Appearance" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">Appearance</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en" class=""><span>Donate</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:CreateAccount&returnto=Richard+J.+Roberts" title="You are encouraged to create an account and log in; however, it is not mandatory" class=""><span>Create account</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:UserLogin&returnto=Richard+J.+Roberts" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o" class=""><span>Log in</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="Log in and more options" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="Personal tools" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">Personal tools</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="User menu" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en"><span>Donate</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:CreateAccount&returnto=Richard+J.+Roberts" title="You are encouraged to create an account and log in; however, it is not mandatory"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>Create account</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:UserLogin&returnto=Richard+J.+Roberts" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Log in</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Pages for logged out editors <a href="/wiki/Help:Introduction" aria-label="Learn more about editing"><span>learn more</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Special:MyContributions" title="A list of edits made from this IP address [y]" accesskey="y"><span>Contributions</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Special:MyTalk" title="Discussion about edits from this IP address [n]" accesskey="n"><span>Talk</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Contents" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Contents</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">hide</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">(Top)</div> </a> </li> <li id="toc-Early_life_and_education" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Early_life_and_education"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Early life and education</span> </div> </a> <ul id="toc-Early_life_and_education-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Career_and_research" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Career_and_research"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Career and research</span> </div> </a> <ul id="toc-Career_and_research-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Awards_and_honours" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Awards_and_honours"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Awards and honours</span> </div> </a> <ul id="toc-Awards_and_honours-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Richard J. Roberts</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 43 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-43" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">43 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B1%D9%8A%D8%AA%D8%B4%D8%A7%D8%B1%D8%AF_%D8%B1%D9%88%D8%A8%D8%B1%D8%AA%D8%B3" title="ريتشارد روبرتس – Arabic" lang="ar" hreflang="ar" data-title="ريتشارد روبرتس" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%B1%DB%8C%DA%86%D8%A7%D8%B1%D8%AF_%D8%B1%D8%A7%D8%A8%D8%B1%D8%AA%D8%B3" title="ریچارد رابرتس – South Azerbaijani" lang="azb" hreflang="azb" data-title="ریچارد رابرتس" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B0%E0%A6%BF%E0%A6%9A%E0%A6%BE%E0%A6%B0%E0%A7%8D%E0%A6%A1_%E0%A6%9C%E0%A7%87._%E0%A6%B0%E0%A6%AC%E0%A6%BE%E0%A6%B0%E0%A7%8D%E0%A6%9F%E0%A6%B8" title="রিচার্ড জে. রবার্টস – Bangla" lang="bn" hreflang="bn" data-title="রিচার্ড জে. রবার্টস" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A0%D1%8B%D1%87%D0%B0%D1%80%D0%B4_%D0%A0%D0%BE%D0%B1%D0%B5%D1%80%D1%82%D1%81" title="Рычард Робертс – Belarusian" lang="be" hreflang="be" data-title="Рычард Робертс" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Richard_John_Roberts" title="Richard John Roberts – Catalan" lang="ca" hreflang="ca" data-title="Richard John Roberts" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Richard_John_Roberts" title="Richard John Roberts – Czech" lang="cs" hreflang="cs" data-title="Richard John Roberts" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – German" lang="de" hreflang="de" data-title="Richard J. Roberts" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Richard_Roberts" title="Richard Roberts – Estonian" lang="et" hreflang="et" data-title="Richard Roberts" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Richard_John_Roberts" title="Richard John Roberts – Spanish" lang="es" hreflang="es" data-title="Richard John Roberts" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Basque" lang="eu" hreflang="eu" data-title="Richard J. Roberts" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B1%DB%8C%DA%86%D8%A7%D8%B1%D8%AF_%D8%B1%D8%A7%D8%A8%D8%B1%D8%AA%D8%B3" title="ریچارد رابرتس – Persian" lang="fa" hreflang="fa" data-title="ریچارد رابرتس" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Richard_Roberts_(biochimiste)" title="Richard Roberts (biochimiste) – French" lang="fr" hreflang="fr" data-title="Richard Roberts (biochimiste)" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Galician" lang="gl" hreflang="gl" data-title="Richard J. Roberts" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%A6%AC%EC%B2%98%EB%93%9C_J._%EB%A1%9C%EB%B2%84%EC%B8%A0" title="리처드 J. 로버츠 – Korean" lang="ko" hreflang="ko" data-title="리처드 J. 로버츠" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Croatian" lang="hr" hreflang="hr" data-title="Richard J. Roberts" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Richard_John_Roberts" title="Richard John Roberts – Ido" lang="io" hreflang="io" data-title="Richard John Roberts" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Indonesian" lang="id" hreflang="id" data-title="Richard J. Roberts" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Richard_Roberts" title="Richard Roberts – Italian" lang="it" hreflang="it" data-title="Richard Roberts" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A8%D7%99%D7%A6%27%D7%A8%D7%93_%D7%92%27._%D7%A8%D7%95%D7%91%D7%A8%D7%98%D7%A1" title="ריצ'רד ג'. רוברטס – Hebrew" lang="he" hreflang="he" data-title="ריצ'רד ג'. רוברטס" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A0%D0%B8%D1%87%D0%B0%D1%80%D0%B4_%D0%94%D0%B6%D0%BE%D0%BD_%D0%A0%D0%BE%D0%B1%D0%B5%D1%80%D1%82%D1%81" title="Ричард Джон Робертс – Kazakh" lang="kk" hreflang="kk" data-title="Ричард Джон Робертс" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Richard_Roberts" title="Richard Roberts – Swahili" lang="sw" hreflang="sw" data-title="Richard Roberts" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Ricardus_J._Roberts" title="Ricardus J. Roberts – Latin" lang="la" hreflang="la" data-title="Ricardus J. Roberts" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Hungarian" lang="hu" hreflang="hu" data-title="Richard J. Roberts" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%B1%DB%8C%DA%86%D8%A7%D8%B1%D8%AF_%D8%B1%D8%A7%D8%A8%D8%B1%D8%AA%D8%B3" title="ریچارد رابرتس – Mazanderani" lang="mzn" hreflang="mzn" data-title="ریچارد رابرتس" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Malay" lang="ms" hreflang="ms" data-title="Richard J. Roberts" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Richard_John_Roberts" title="Richard John Roberts – Dutch" lang="nl" hreflang="nl" data-title="Richard John Roberts" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%AA%E3%83%81%E3%83%A3%E3%83%BC%E3%83%89%E3%83%BB%E3%83%AD%E3%83%90%E3%83%BC%E3%83%84" title="リチャード・ロバーツ – Japanese" lang="ja" hreflang="ja" data-title="リチャード・ロバーツ" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Richard J. Roberts" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Occitan" lang="oc" hreflang="oc" data-title="Richard J. Roberts" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%B1%DB%8C%DA%86%D8%B1%DA%88_%D8%B1%D8%A7%D8%A8%D8%B1%D9%B9%D8%B3" title="ریچرڈ رابرٹس – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ریچرڈ رابرٹس" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Richard_John_Roberts" title="Richard John Roberts – Low German" lang="nds" hreflang="nds" data-title="Richard John Roberts" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Polish" lang="pl" hreflang="pl" data-title="Richard J. Roberts" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Richard_Roberts" title="Richard Roberts – Portuguese" lang="pt" hreflang="pt" data-title="Richard Roberts" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A0%D0%BE%D0%B1%D0%B5%D1%80%D1%82%D1%81,_%D0%A0%D0%B8%D1%87%D0%B0%D1%80%D0%B4_%D0%94%D0%B6%D0%BE%D0%BD" title="Робертс, Ричард Джон – Russian" lang="ru" hreflang="ru" data-title="Робертс, Ричард Джон" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Richard_Roberts" title="Richard Roberts – Simple English" lang="en-simple" hreflang="en-simple" data-title="Richard Roberts" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Richard_Roberts" title="Richard Roberts – Finnish" lang="fi" hreflang="fi" data-title="Richard Roberts" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Swedish" lang="sv" hreflang="sv" data-title="Richard J. Roberts" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%A0%D0%B8%D1%87%D0%B0%D1%80%D0%B4_%D0%A0%D0%BE%D0%B1%D0%B5%D1%80%D1%82%D1%81" title="Ричард Робертс – Tatar" lang="tt" hreflang="tt" data-title="Ричард Робертс" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Turkish" lang="tr" hreflang="tr" data-title="Richard J. Roberts" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A0%D1%96%D1%87%D0%B0%D1%80%D0%B4_%D0%A0%D0%BE%D0%B1%D0%B5%D1%80%D1%82%D1%81" title="Річард Робертс – Ukrainian" lang="uk" hreflang="uk" data-title="Річард Робертс" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%B1%DA%86%D8%B1%DA%88_%D8%AC%DB%92_%D8%B1%D8%A7%D8%A8%D8%B1%D9%B9" title="رچرڈ جے رابرٹ – Urdu" lang="ur" hreflang="ur" data-title="رچرڈ جے رابرٹ" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Richard_J._Roberts" title="Richard J. Roberts – Yoruba" lang="yo" hreflang="yo" data-title="Richard J. Roberts" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E7%90%86%E5%AF%9F%C2%B7%E7%BE%85%E4%BC%AF%E8%8C%A8" title="理察·羅伯茨 – Chinese" lang="zh" hreflang="zh" data-title="理察·羅伯茨" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q309816#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Richard_J._Roberts" title="View the content page [c]" accesskey="c"><span>Article</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Talk:Richard_J._Roberts" rel="discussion" title="Discuss improvements to the content page [t]" accesskey="t"><span>Talk</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Change language variant" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">English</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Views"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Richard_J._Roberts"><span>Read</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Richard_J._Roberts&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Richard_J._Roberts&action=history" title="Past revisions of this page [h]" accesskey="h"><span>View history</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Tools" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Tools</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Tools</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">hide</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="More options" > <div class="vector-menu-heading"> Actions </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Richard_J._Roberts"><span>Read</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Richard_J._Roberts&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Richard_J._Roberts&action=history"><span>View history</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> General </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Special:WhatLinksHere/Richard_J._Roberts" title="List of all English Wikipedia pages containing links to this page [j]" accesskey="j"><span>What links here</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Special:RecentChangesLinked/Richard_J._Roberts" rel="nofollow" title="Recent changes in pages linked from this page [k]" accesskey="k"><span>Related changes</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_Upload_Wizard" title="Upload files [u]" accesskey="u"><span>Upload file</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Special:SpecialPages" title="A list of all special pages [q]" accesskey="q"><span>Special pages</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Richard_J._Roberts&oldid=1254328414" title="Permanent link to this revision of this page"><span>Permanent link</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Richard_J._Roberts&action=info" title="More information about this page"><span>Page information</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Special:CiteThisPage&page=Richard_J._Roberts&id=1254328414&wpFormIdentifier=titleform" title="Information on how to cite this page"><span>Cite this page</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Special:UrlShortener&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FRichard_J._Roberts"><span>Get shortened URL</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Special:QrCode&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FRichard_J._Roberts"><span>Download QR code</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Print/export </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Special:DownloadAsPdf&page=Richard_J._Roberts&action=show-download-screen" title="Download this page as a PDF file"><span>Download as PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Richard_J._Roberts&printable=yes" title="Printable version of this page [p]" accesskey="p"><span>Printable version</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> In other projects </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Richard_John_Roberts" hreflang="en"><span>Wikimedia Commons</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q309816" title="Structured data on this page hosted by Wikidata [g]" accesskey="g"><span>Wikidata item</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Appearance</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">hide</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">British biochemist (born 1943)</div> <p class="mw-empty-elt"> </p> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="honorific-prefix" style="font-size: 77%; font-weight: normal;"><a href="/wiki/Sir" title="Sir">Sir</a></div><div class="fn">Richard J. Roberts</div><div class="honorific-suffix" style="font-size: 77%; font-weight: normal;"><span class="nobold noexcerpt nowraplinks" style="font-size:;"><span style="font-size: 100%;"><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">FRS</a></span></span></div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Roberts,_Richard_John_(1943).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/19/Roberts%2C_Richard_John_%281943%29.jpg/220px-Roberts%2C_Richard_John_%281943%29.jpg" decoding="async" width="220" height="295" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/19/Roberts%2C_Richard_John_%281943%29.jpg/330px-Roberts%2C_Richard_John_%281943%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/19/Roberts%2C_Richard_John_%281943%29.jpg/440px-Roberts%2C_Richard_John_%281943%29.jpg 2x" data-file-width="509" data-file-height="682" /></a></span><div class="infobox-caption">Roberts in 2007</div></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">Richard John Roberts</div><br /><span style="display:none"> (<span class="bday">1943-09-06</span>) </span>6 September 1943<span class="noprint ForceAgeToShow"> (age 81)</span><sup id="cite_ref-whoswho_3-2" class="reference"><a href="#cite_note-whoswho-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><br /><div style="display:inline" class="birthplace"><a href="/wiki/Derby" title="Derby">Derby</a>, England</div></td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/University_of_Sheffield" title="University of Sheffield">University of Sheffield</a> (<a href="/wiki/BSc" class="mw-redirect" title="BSc">BSc</a>, <a href="/wiki/PhD" class="mw-redirect" title="PhD">PhD</a>)</td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style><div class="plainlist"> <ul><li><a href="/wiki/Alternative_splicing" title="Alternative splicing">Alternative splicing</a></li> <li>Work on <a href="/wiki/Intron" title="Intron">introns</a></li> <li><a href="/wiki/Restriction_enzyme" title="Restriction enzyme">Restriction endonucleases</a><sup id="cite_ref-Roberts2003_7-0" class="reference"><a href="#cite_note-Roberts2003-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/DNA_methylation" title="DNA methylation">DNA methylation</a></li> <li><a href="/wiki/Computational_biology" title="Computational biology">Computational molecular biology</a><sup id="cite_ref-whoswho_3-3" class="reference"><a href="#cite_note-whoswho-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> (1993)<sup id="cite_ref-nobel1_1-0" class="reference"><a href="#cite_note-nobel1-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-nobel4_2-0" class="reference"><a href="#cite_note-nobel4-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">FRS</a> (1995)<sup id="cite_ref-whoswho_3-0" class="reference"><a href="#cite_note-whoswho-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-frs_4-0" class="reference"><a href="#cite_note-frs-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/EMBO_Membership" title="EMBO Membership">EMBO Membership</a> (1995)<sup id="cite_ref-membo_5-0" class="reference"><a href="#cite_note-membo-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Knight_Bachelor" title="Knight Bachelor">Knight Bachelor</a> (2008)<sup id="cite_ref-whoswho_3-1" class="reference"><a href="#cite_note-whoswho-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Lomonosov_Gold_Medal" title="Lomonosov Gold Medal">Lomonosov Gold Medal</a> (2021)<sup id="cite_ref-Lomonosov_Gold_Medal_2021_6-0" class="reference"><a href="#cite_note-Lomonosov_Gold_Medal_2021-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></li></ul> </div></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Molecular_biology" title="Molecular biology">Molecular biology</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li><a href="/wiki/University_of_Sheffield" title="University of Sheffield">University of Sheffield</a></li> <li><a href="/wiki/New_England_Biolabs" title="New England Biolabs">New England Biolabs</a></li> <li><a href="/wiki/Cold_Spring_Harbor_Laboratory" title="Cold Spring Harbor Laboratory">Cold Spring Harbor Laboratory</a></li> <li><a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a></li></ul> </div></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Thesis" title="Thesis">Thesis</a></th><td class="infobox-data"><i><a rel="nofollow" class="external text" href="http://copac.jisc.ac.uk/id/36827249?style=html">Phytochemical studies involving neoflavanoids and isoflavanoids</a></i> <span style="font-size:97%;">(1969)</span></td></tr><tr style="display:none"><td colspan="2"> </td></tr><tr><th scope="row" class="infobox-label">Website</th><td class="infobox-data"><span class="url"><a rel="nofollow" class="external text" href="http://nobelprize.org/nobel_prizes/medicine/laureates/1993/roberts-bio.html">nobelprize<wbr />.org<wbr />/nobel<wbr />_prizes<wbr />/medicine<wbr />/laureates<wbr />/1993<wbr />/roberts-bio<wbr />.html</a></span></td></tr></tbody></table> <p><b>Sir Richard John Roberts</b> <span class="nobold noexcerpt nowraplinks" style="font-size:;"><span style="font-size: 85%;"><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">FRS</a></span></span><sup id="cite_ref-frs_4-1" class="reference"><a href="#cite_note-frs-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> (born 6 September 1943) is a British <a href="/wiki/Biochemist" title="Biochemist">biochemist</a> and <a href="/wiki/Molecular_biology" title="Molecular biology">molecular biologist</a>. He was awarded the 1993 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> with <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Allen Sharp</a> for the discovery of <a href="/wiki/Intron" title="Intron">introns</a> in <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotic</a> <a href="/wiki/DNA" title="DNA">DNA</a> and the mechanism of <a href="/wiki/Genetic_engineering" title="Genetic engineering">gene-splicing</a>. He currently works at <a href="/wiki/New_England_Biolabs" title="New England Biolabs">New England Biolabs</a>.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-scopus_10-0" class="reference"><a href="#cite_note-scopus-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Early_life_and_education">Early life and education</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Richard_J._Roberts&action=edit&section=1" title="Edit section: Early life and education"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Roberts was born in <a href="/wiki/Derby" title="Derby">Derby</a>, the son of Edna (Allsop) and John Roberts, an auto mechanic.<sup id="cite_ref-nobelbio_11-0" class="reference"><a href="#cite_note-nobelbio-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> When he was four, Roberts' family moved to <a href="/wiki/Bath,_Somerset" title="Bath, Somerset">Bath</a>. In Bath, he attended <a href="/wiki/City_of_Bath_Boys%27_School" class="mw-redirect" title="City of Bath Boys' School">City of Bath Boys' School</a>.<sup id="cite_ref-nobelbio_11-1" class="reference"><a href="#cite_note-nobelbio-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> As a child he at first wanted to be a <a href="/wiki/Detective" title="Detective">detective</a> and then, when given a <a href="/wiki/Chemistry_set" title="Chemistry set">chemistry set</a>, a <a href="/wiki/Chemist" title="Chemist">chemist</a>. In 1965 he graduated from the <a href="/wiki/University_of_Sheffield" title="University of Sheffield">University of Sheffield</a> with a <a href="/wiki/Bachelor_of_Science" title="Bachelor of Science">Bachelor of Science</a> degree in <a href="/wiki/Chemistry" title="Chemistry">chemistry</a> followed by a <a href="/wiki/PhD" class="mw-redirect" title="PhD">PhD</a> in 1969.<sup id="cite_ref-whoswho_3-4" class="reference"><a href="#cite_note-whoswho-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> His thesis involved <a href="/wiki/Phytochemical" title="Phytochemical">phytochemical</a> studies of <a href="/wiki/Neoflavonoid" title="Neoflavonoid">neoflavonoids</a> and <a href="/wiki/Isoflavonoid" title="Isoflavonoid">isoflavonoids</a>.<sup id="cite_ref-robertsphd_12-0" class="reference"><a href="#cite_note-robertsphd-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ORCID_13-0" class="reference"><a href="#cite_note-ORCID-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Career_and_research">Career and research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Richard_J._Roberts&action=edit&section=2" title="Edit section: Career and research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>During 1969–1972, he did <a href="/wiki/Postdoctoral_research" class="mw-redirect" title="Postdoctoral research">postdoctoral research</a> at <a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a>.<sup id="cite_ref-nobelbio_11-2" class="reference"><a href="#cite_note-nobelbio-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> before moving to <a href="/wiki/Cold_Spring_Harbor_Laboratory" title="Cold Spring Harbor Laboratory">Cold Spring Harbor Laboratory</a>,<sup id="cite_ref-adeno_14-0" class="reference"><a href="#cite_note-adeno-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> where he was hired by <a href="/wiki/James_Dewey_Watson" class="mw-redirect" title="James Dewey Watson">James Dewey Watson</a>, a co-discoverer of the structure of <a href="/wiki/DNA" title="DNA">DNA</a> and a fellow Nobel laureate. In this period he also visited the <a href="/wiki/MRC_Laboratory_of_Molecular_Biology" title="MRC Laboratory of Molecular Biology">MRC Laboratory of Molecular Biology</a> for the first time, working alongside <a href="/wiki/Frederick_Sanger" title="Frederick Sanger">Fred Sanger</a>.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> In 1977, he published his discovery of RNA splicing.<sup id="cite_ref-adeno_14-1" class="reference"><a href="#cite_note-adeno-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> In 1992, he moved to <a href="/wiki/New_England_Biolabs" title="New England Biolabs">New England Biolabs</a>.<sup id="cite_ref-nobelbio_11-3" class="reference"><a href="#cite_note-nobelbio-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> The following year, he shared a Nobel Prize with his former colleague at Cold Spring Harbor <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Allen Sharp</a>.<sup id="cite_ref-KlimasauskasKumar1994_16-0" class="reference"><a href="#cite_note-KlimasauskasKumar1994-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p><p>Roberts's discovery of the <a href="/wiki/Alternative_splicing" title="Alternative splicing">alternative splicing</a> of genes, in particular, has had a profound impact on the study and applications of molecular biology.<sup id="cite_ref-frs_4-2" class="reference"><a href="#cite_note-frs-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> The realisation that individual genes could exist as separate, disconnected segments within longer strands of DNA first arose in his 1977 study of <a href="/wiki/Adenovirus" class="mw-redirect" title="Adenovirus">adenovirus</a>,<sup id="cite_ref-adeno_14-2" class="reference"><a href="#cite_note-adeno-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> one of the viruses responsible for causing the common cold. Robert's research in this field resulted in a fundamental shift in our understanding of <a href="/wiki/Genetics" title="Genetics">genetics</a>, and has led to the discovery of split genes in higher organisms, including human beings.<sup id="cite_ref-frs_4-3" class="reference"><a href="#cite_note-frs-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-scopus_10-1" class="reference"><a href="#cite_note-scopus-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Awards_and_honours">Awards and honours</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Richard_J._Roberts&action=edit&section=3" title="Edit section: Awards and honours"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In 1992, Roberts received an <a href="/wiki/Honorary_degree" title="Honorary degree">honorary doctorate</a> from the Faculty of Medicine at <a href="/wiki/Uppsala_University" title="Uppsala University">Uppsala University</a>, <a href="/wiki/Sweden" title="Sweden">Sweden</a>.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> After becoming a Nobel laureate in 1993 he was awarded an Honorary Degree (Doctor of Science) by the <a href="/wiki/University_of_Bath" title="University of Bath">University of Bath</a> in 1994.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> Roberts also received the Golden Plate Award of the <a href="/wiki/Academy_of_Achievement" title="Academy of Achievement">American Academy of Achievement</a> in 1994.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> In 2021 he was awarded the <a href="/wiki/Lomonosov_Gold_Medal" title="Lomonosov Gold Medal">Lomonosov Gold Medal</a> of the Russian Academy of Sciences.<sup id="cite_ref-Lomonosov_Gold_Medal_2021_6-1" class="reference"><a href="#cite_note-Lomonosov_Gold_Medal_2021-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>Roberts was elected a <a href="/wiki/List_of_Fellows_of_the_Royal_Society_elected_in_1995" class="mw-redirect" title="List of Fellows of the Royal Society elected in 1995">Fellow of the Royal Society (FRS) in 1995</a><sup id="cite_ref-frs_4-4" class="reference"><a href="#cite_note-frs-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> and a <a href="/wiki/EMBO_Member" class="mw-redirect" title="EMBO Member">member of the European Molecular Biology Organization (EMBO)</a> in the same year.<sup id="cite_ref-membo_5-1" class="reference"><a href="#cite_note-membo-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> In 2005, a multimillion-pound expansion to the chemistry department at the <a href="/wiki/University_of_Sheffield" title="University of Sheffield">University of Sheffield</a>, where he had been a student, was named after him. A refurbished science department at <a href="/wiki/Beechen_Cliff_School" title="Beechen Cliff School">Beechen Cliff School</a> (previously City of Bath Boys' School) was also named after Roberts, who had donated a substantial sum of his Nobel prize winnings to the school.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> </p><p>Roberts is an atheist and was one of the signers of the <a href="/wiki/Humanist_Manifesto_II" title="Humanist Manifesto II">Humanist Manifesto</a>.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> He was <a href="/wiki/Knight_Bachelor" title="Knight Bachelor">knighted</a> in the 2008 Birthday Honours.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p><p>Roberts is a member of the Advisory Board of <a href="/wiki/Patient_Innovation" title="Patient Innovation">Patient Innovation</a>,<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> a nonprofit, international, multilingual, free venue for patients and caregivers of any disease to share their innovations. </p><p>Roberts has been a <a href="/wiki/Keynote" title="Keynote">keynote</a> speaker at the Congress of Future Medical Leaders (2014, 2015, 2016, 2020).<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p><p>He also is the chairman of The Laureate Science Alliance, a non-profit supporting research worldwide. </p><p>In 2016, Roberts and other Nobelists composed and signed a "Laureates Letter Supporting Precision Agriculture (GMOs)" addressed to the leaders of Greenpeace, the United Nations and global governments and Sir Roberts has advocated for <a href="/wiki/Genetically_Modified_Organisms" class="mw-redirect" title="Genetically Modified Organisms">Genetically Modified Organisms</a> (GMOs) in general and <a href="/wiki/Golden_Rice" class="mw-redirect" title="Golden Rice">Golden Rice</a> in particular to advance health in developing countries, noting the high safety record of GM foods.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Richard_J._Roberts&action=edit&section=4" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><div class="side-box metadata side-box-right"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/40px-Scholia_logo.svg.png" decoding="async" width="40" height="39" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/60px-Scholia_logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Scholia_logo.svg/80px-Scholia_logo.svg.png 2x" data-file-width="107" data-file-height="104" /></span></span></div> <div class="side-box-text plainlist"><a href="https://www.wikidata.org/wiki/Wikidata:Scholia" class="extiw" title="d:Wikidata:Scholia">Scholia</a> has an <i>author</i> profile for <i><b><a href="https://iw.toolforge.org/scholia/author/Q309816" class="extiw" title="toolforge:scholia/author/Q309816">Richard J. Roberts</a></b></i>.</div></div> </div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-nobel1-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-nobel1_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFShampoKyle2003" class="citation journal cs1">Shampo, M. A.; Kyle, R. A. (2003). <a rel="nofollow" class="external text" href="https://doi.org/10.4065%2F78.2.132">"Richard J. Roberts—Nobel Laureate for Discovery of Split Genes"</a>. <i>Mayo Clinic Proceedings</i>. <b>78</b> (2): 132. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.4065%2F78.2.132">10.4065/78.2.132</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/12583523">12583523</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Mayo+Clinic+Proceedings&rft.atitle=Richard+J.+Roberts%E2%80%94Nobel+Laureate+for+Discovery+of+Split+Genes&rft.volume=78&rft.issue=2&rft.pages=132&rft.date=2003&rft_id=info%3Adoi%2F10.4065%2F78.2.132&rft_id=info%3Apmid%2F12583523&rft.aulast=Shampo&rft.aufirst=M.+A.&rft.au=Kyle%2C+R.+A.&rft_id=https%3A%2F%2Fdoi.org%2F10.4065%252F78.2.132&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-nobel4-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-nobel4_2-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCarr1993" class="citation journal cs1">Carr, Kimberly (1993). <a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F365597a0">"Nobel goes to discoverers of 'split genes'<span class="cs1-kern-right"></span>"</a>. <i>Nature</i>. <b>365</b> (6447): 597. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1993Natur.365..597C">1993Natur.365..597C</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F365597a0">10.1038/365597a0</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/8413620">8413620</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Nobel+goes+to+discoverers+of+%27split+genes%27&rft.volume=365&rft.issue=6447&rft.pages=597&rft.date=1993&rft_id=info%3Apmid%2F8413620&rft_id=info%3Adoi%2F10.1038%2F365597a0&rft_id=info%3Abibcode%2F1993Natur.365..597C&rft.aulast=Carr&rft.aufirst=Kimberly&rft_id=https%3A%2F%2Fdoi.org%2F10.1038%252F365597a0&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-whoswho-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-whoswho_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-whoswho_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-whoswho_3-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-whoswho_3-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-whoswho_3-4"><sup><i><b>e</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation encyclopaedia cs1"><span class="id-lock-subscription" title="Paid subscription required"><a rel="nofollow" class="external text" href="https://www.ukwhoswho.com/view/article/oupww/whoswho/U32757">"ROBERTS,Sir Richard (John)"</a></span>. <i><a href="/wiki/Who%27s_Who_(UK)" title="Who's Who (UK)">Who's Who</a></i>. Vol. 1995 (online <a href="/wiki/Oxford_University_Press" title="Oxford University Press">Oxford University Press</a> ed.). A & C Black.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=ROBERTS%2CSir+Richard+%28John%29&rft.btitle=Who%27s+Who&rft.edition=online+Oxford+University+Press&rft.pub=A+%26+C+Black&rft_id=https%3A%2F%2Fwww.ukwhoswho.com%2Fview%2Farticle%2Foupww%2Fwhoswho%2FU32757&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span> <span style="font-size:0.95em; font-size:95%; color: var( --color-subtle, #555 )">(Subscription or <a rel="nofollow" class="external text" href="https://www.ukwhoswho.com/page/subscribe#public">UK public library membership</a> required.)</span></span> </li> <li id="cite_note-frs-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-frs_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-frs_4-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-frs_4-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-frs_4-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-frs_4-4"><sup><i><b>e</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAnon1995" class="citation web cs1">Anon (1995). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20151006213105/https://royalsociety.org/people/richard-roberts-12188/">"Sir Richard Roberts FRS"</a>. London: <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a>. Archived from <a rel="nofollow" class="external text" href="https://royalsociety.org/people/richard-roberts-12188/">the original</a> on 6 October 2015.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Sir+Richard+Roberts+FRS&rft.place=London&rft.pub=Royal+Society&rft.date=1995&rft.au=Anon&rft_id=https%3A%2F%2Froyalsociety.org%2Fpeople%2Frichard-roberts-12188%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span> One or more of the preceding sentences incorporates text from the royalsociety.org website where: <style data-mw-deduplicate="TemplateStyles:r1244412712">.mw-parser-output .templatequote{overflow:hidden;margin:1em 0;padding:0 32px}.mw-parser-output .templatequotecite{line-height:1.5em;text-align:left;margin-top:0}@media(min-width:500px){.mw-parser-output .templatequotecite{padding-left:1.6em}}</style><blockquote class="templatequote"><p>"All text published under the heading 'Biography' on Fellow profile pages is available under <a href="/wiki/Creative_Commons_license" title="Creative Commons license">Creative Commons Attribution 4.0 International License</a>.” --<link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20150925220834/https://royalsociety.org/about-us/terms-conditions-policies/">"Royal Society Terms, conditions and policies"</a>. Archived from the original on 25 September 2015<span class="reference-accessdate">. Retrieved <span class="nowrap">9 March</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Royal+Society+Terms%2C+conditions+and+policies&rft_id=https%3A%2F%2Froyalsociety.org%2Fabout-us%2Fterms-conditions-policies%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_web" title="Template:Cite web">cite web</a>}}</code>: CS1 maint: bot: original URL status unknown (<a href="/wiki/Category:CS1_maint:_bot:_original_URL_status_unknown" title="Category:CS1 maint: bot: original URL status unknown">link</a>)</span></p></blockquote></span> </li> <li id="cite_note-membo-5"><span class="mw-cite-backlink">^ <a href="#cite_ref-membo_5-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-membo_5-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAnon2016" class="citation web cs1">Anon (2016). <a rel="nofollow" class="external text" href="http://people.embo.org/profile/richard-j-roberts">"Richard J. Roberts EMBO profile"</a>. <i>people.embo.org</i>. Heidelberg: <a href="/wiki/European_Molecular_Biology_Organization" title="European Molecular Biology Organization">European Molecular Biology Organization</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=people.embo.org&rft.atitle=Richard+J.+Roberts+EMBO+profile&rft.date=2016&rft.au=Anon&rft_id=http%3A%2F%2Fpeople.embo.org%2Fprofile%2Frichard-j-roberts&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-Lomonosov_Gold_Medal_2021-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-Lomonosov_Gold_Medal_2021_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Lomonosov_Gold_Medal_2021_6-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.ras.ru/news/news_release.aspx?ID=72609491-fb8d-4235-97a2-0853af546268">Lomonosov Gold Medal 2021</a>(in Russian)</span> </li> <li id="cite_note-Roberts2003-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-Roberts2003_7-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRoberts2003" class="citation journal cs1">Roberts, Richard John (2003). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC152790">"A nomenclature for restriction enzymes, DNA methyltransferases, homing endonucleases and their genes"</a>. <i><a href="/wiki/Nucleic_Acids_Research" title="Nucleic Acids Research">Nucleic Acids Research</a></i>. <b>31</b> (7): 1805–1812. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fnar%2Fgkg274">10.1093/nar/gkg274</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC152790">152790</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/12654995">12654995</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nucleic+Acids+Research&rft.atitle=A+nomenclature+for+restriction+enzymes%2C+DNA+methyltransferases%2C+homing+endonucleases+and+their+genes&rft.volume=31&rft.issue=7&rft.pages=1805-1812&rft.date=2003&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC152790%23id-name%3DPMC&rft_id=info%3Apmid%2F12654995&rft_id=info%3Adoi%2F10.1093%2Fnar%2Fgkg274&rft.aulast=Roberts&rft.aufirst=Richard+John&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC152790&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span> <span style="position:relative; top: -2px;"><span typeof="mw:File"><a href="/wiki/Open_access" title="open access publication – free to read"><img alt="Open access icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Open_Access_logo_PLoS_transparent.svg/9px-Open_Access_logo_PLoS_transparent.svg.png" decoding="async" width="9" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Open_Access_logo_PLoS_transparent.svg/14px-Open_Access_logo_PLoS_transparent.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/77/Open_Access_logo_PLoS_transparent.svg/18px-Open_Access_logo_PLoS_transparent.svg.png 2x" data-file-width="640" data-file-height="1000" /></a></span></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRobertsChangHuRachlin2010" class="citation journal cs1"><a class="mw-selflink selflink">Roberts, R. J.</a>; Chang, Y. -C.; Hu, Z.; Rachlin, J. N.; Anton, B. P.; Pokrzywa, R. M.; Choi, H. -P.; Faller, L. L.; Guleria, J.; Housman, G.; Klitgord, N.; Mazumdar, V.; McGettrick, M. G.; Osmani, L.; Swaminathan, R.; Tao, K. R.; Letovsky, S.; Vitkup, D.; Segrè, D.; Salzberg, S. L.; Delisi, C.; Steffen, M.; Kasif, S. (2010). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3013729">"COMBREX: A project to accelerate the functional annotation of prokaryotic genomes"</a>. <i><a href="/wiki/Nucleic_Acids_Research" title="Nucleic Acids Research">Nucleic Acids Research</a></i>. <b>39</b> (Database issue): D11–D14. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fnar%2Fgkq1168">10.1093/nar/gkq1168</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3013729">3013729</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21097892">21097892</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nucleic+Acids+Research&rft.atitle=COMBREX%3A+A+project+to+accelerate+the+functional+annotation+of+prokaryotic+genomes&rft.volume=39&rft.issue=Database+issue&rft.pages=D11-D14&rft.date=2010&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3013729%23id-name%3DPMC&rft_id=info%3Apmid%2F21097892&rft_id=info%3Adoi%2F10.1093%2Fnar%2Fgkq1168&rft.aulast=Roberts&rft.aufirst=R.+J.&rft.au=Chang%2C+Y.+-C.&rft.au=Hu%2C+Z.&rft.au=Rachlin%2C+J.+N.&rft.au=Anton%2C+B.+P.&rft.au=Pokrzywa%2C+R.+M.&rft.au=Choi%2C+H.+-P.&rft.au=Faller%2C+L.+L.&rft.au=Guleria%2C+J.&rft.au=Housman%2C+G.&rft.au=Klitgord%2C+N.&rft.au=Mazumdar%2C+V.&rft.au=McGettrick%2C+M.+G.&rft.au=Osmani%2C+L.&rft.au=Swaminathan%2C+R.&rft.au=Tao%2C+K.+R.&rft.au=Letovsky%2C+S.&rft.au=Vitkup%2C+D.&rft.au=Segr%C3%A8%2C+D.&rft.au=Salzberg%2C+S.+L.&rft.au=Delisi%2C+C.&rft.au=Steffen%2C+M.&rft.au=Kasif%2C+S.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3013729&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRobertsVarmusAshburnerBrown2001" class="citation journal cs1"><a class="mw-selflink selflink">Roberts, R. J.</a>; <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus, H. E.</a>; <a href="/wiki/Michael_Ashburner" title="Michael Ashburner">Ashburner, M.</a>; <a href="/wiki/Patrick_O._Brown" title="Patrick O. Brown">Brown, P. O.</a>; <a href="/wiki/Michael_Eisen" title="Michael Eisen">Eisen, M. B.</a>; <a href="/wiki/Chaitan_Khosla" title="Chaitan Khosla">Khosla, C.</a>; Kirschner, M.; <a href="/wiki/Roel_Nusse" title="Roel Nusse">Nusse, R.</a>; Scott, M. (2001). "Information Access: Building A <i>GenBank</i> of the Published Literature". <i>Science</i>. <b>291</b> (5512): 2318–9. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1060273">10.1126/science.1060273</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11269300">11269300</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:34395787">34395787</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Information+Access%3A+Building+A+GenBank+of+the+Published+Literature&rft.volume=291&rft.issue=5512&rft.pages=2318-9&rft.date=2001&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A34395787%23id-name%3DS2CID&rft_id=info%3Apmid%2F11269300&rft_id=info%3Adoi%2F10.1126%2Fscience.1060273&rft.aulast=Roberts&rft.aufirst=R.+J.&rft.au=Varmus%2C+H.+E.&rft.au=Ashburner%2C+M.&rft.au=Brown%2C+P.+O.&rft.au=Eisen%2C+M.+B.&rft.au=Khosla%2C+C.&rft.au=Kirschner%2C+M.&rft.au=Nusse%2C+R.&rft.au=Scott%2C+M.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-scopus-10"><span class="mw-cite-backlink">^ <a href="#cite_ref-scopus_10-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-scopus_10-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.url?authorId=7404281937">Richard J. Roberts's publications</a> indexed by the <a href="/wiki/Scopus" title="Scopus">Scopus</a> bibliographic database. <span style="font-size:0.95em; font-size:95%; color: var( --color-subtle, #555 )">(subscription required)</span></span> </li> <li id="cite_note-nobelbio-11"><span class="mw-cite-backlink">^ <a href="#cite_ref-nobelbio_11-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-nobelbio_11-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-nobelbio_11-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-nobelbio_11-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAnon1993" class="citation web cs1">Anon (1993). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20160304003141/http://www.nobelprize.org/nobel_prizes/medicine/laureates/1993/roberts-bio.html">"Richard J. Roberts - Biographical"</a>. Nobel Foundation. Archived from <a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/1993/roberts-bio.html">the original</a> on 4 March 2016<span class="reference-accessdate">. Retrieved <span class="nowrap">28 February</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Richard+J.+Roberts+-+Biographical&rft.pub=Nobel+Foundation&rft.date=1993&rft.au=Anon&rft_id=https%3A%2F%2Fwww.nobelprize.org%2Fnobel_prizes%2Fmedicine%2Flaureates%2F1993%2Froberts-bio.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-robertsphd-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-robertsphd_12-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRoberts1969" class="citation thesis cs1">Roberts, Richard John (1969). <a rel="nofollow" class="external text" href="http://copac.jisc.ac.uk/id/36827249?style=html"><i>Phytochemical studies involving neoflavanoids and isoflavanoids</i></a>. <i>copac.jisc.ac.uk</i> (PhD thesis). University of Sheffield. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/270832404">270832404</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adissertation&rft.title=Phytochemical+studies+involving+neoflavanoids+and+isoflavanoids&rft.degree=PhD&rft.inst=University+of+Sheffield&rft.date=1969&rft_id=info%3Aoclcnum%2F270832404&rft.aulast=Roberts&rft.aufirst=Richard+John&rft_id=http%3A%2F%2Fcopac.jisc.ac.uk%2Fid%2F36827249%3Fstyle%3Dhtml&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-ORCID-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-ORCID_13-0">^</a></b></span> <span class="reference-text">Richard J. Roberts <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ORCID_(identifier)" class="mw-redirect" title="ORCID (identifier)">ORCID</a> <a rel="nofollow" class="external text" href="https://orcid.org/0000-0002-4348-0169">0000-0002-4348-0169</a></span> </li> <li id="cite_note-adeno-14"><span class="mw-cite-backlink">^ <a href="#cite_ref-adeno_14-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-adeno_14-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-adeno_14-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFChowGelinasBrokerRoberts1977" class="citation journal cs1">Chow, Louise T.; Gelinas, Richard E.; Broker, Thomas R.; Roberts, Richard J. (1977). "An amazing sequence arrangement at the 5' ends of adenovirus 2 messenger RNA". <i>Cell</i>. <b>12</b> (1): 1–8. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2F0092-8674%2877%2990180-5">10.1016/0092-8674(77)90180-5</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/902310">902310</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:2099968">2099968</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cell&rft.atitle=An+amazing+sequence+arrangement+at+the+5%27+ends+of+adenovirus+2+messenger+RNA&rft.volume=12&rft.issue=1&rft.pages=1-8&rft.date=1977&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A2099968%23id-name%3DS2CID&rft_id=info%3Apmid%2F902310&rft_id=info%3Adoi%2F10.1016%2F0092-8674%2877%2990180-5&rft.aulast=Chow&rft.aufirst=Louise+T.&rft.au=Gelinas%2C+Richard+E.&rft.au=Broker%2C+Thomas+R.&rft.au=Roberts%2C+Richard+J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-15">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/1993/roberts/biographical/">"The Nobel Prize in Physiology or Medicine 1993"</a>. <i>NobelPrize.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">14 July</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=NobelPrize.org&rft.atitle=The+Nobel+Prize+in+Physiology+or+Medicine+1993&rft_id=https%3A%2F%2Fwww.nobelprize.org%2Fprizes%2Fmedicine%2F1993%2Froberts%2Fbiographical%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-KlimasauskasKumar1994-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-KlimasauskasKumar1994_16-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKlimasauskasKumarRobertsCheng1994" class="citation journal cs1">Klimasauskas, Saulius; Kumar, Sanjay; Roberts, Richard J.; Cheng, Xiaodong (1994). "Hhal methyltransferase flips its target base out of the DNA helix". <i>Cell</i>. <b>76</b> (2): 357–369. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2F0092-8674%2894%2990342-5">10.1016/0092-8674(94)90342-5</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/8293469">8293469</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:23161543">23161543</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cell&rft.atitle=Hhal+methyltransferase+flips+its+target+base+out+of+the+DNA+helix&rft.volume=76&rft.issue=2&rft.pages=357-369&rft.date=1994&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A23161543%23id-name%3DS2CID&rft_id=info%3Apmid%2F8293469&rft_id=info%3Adoi%2F10.1016%2F0092-8674%2894%2990342-5&rft.aulast=Klimasauskas&rft.aufirst=Saulius&rft.au=Kumar%2C+Sanjay&rft.au=Roberts%2C+Richard+J.&rft.au=Cheng%2C+Xiaodong&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.uu.se/en/about-uu/traditions/prizes/honorary-doctorates/">"Honorary doctorates - Uppsala University, Sweden"</a>. <i>Uu.se</i>. 17 August 2016<span class="reference-accessdate">. Retrieved <span class="nowrap">28 August</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Uu.se&rft.atitle=Honorary+doctorates+-+Uppsala+University%2C+Sweden&rft.date=2016-08-17&rft_id=http%3A%2F%2Fwww.uu.se%2Fen%2Fabout-uu%2Ftraditions%2Fprizes%2Fhonorary-doctorates%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-18"><span class="mw-cite-backlink"><b><a href="#cite_ref-18">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20151219000643/http://www.bath.ac.uk/ceremonies/hongrads/">"Honorary Graduates 1989 to present"</a>. <i>bath.ac.uk</i>. <a href="/wiki/University_of_Bath" title="University of Bath">University of Bath</a>. Archived from <a rel="nofollow" class="external text" href="http://www.bath.ac.uk/ceremonies/hongrads/">the original</a> on 19 December 2015<span class="reference-accessdate">. Retrieved <span class="nowrap">18 February</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=bath.ac.uk&rft.atitle=Honorary+Graduates+1989+to+present&rft_id=http%3A%2F%2Fwww.bath.ac.uk%2Fceremonies%2Fhongrads%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-19"><span class="mw-cite-backlink"><b><a href="#cite_ref-19">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://achievement.org/our-history/golden-plate-awards/#science-exploration">"Golden Plate Awardees of the American Academy of Achievement"</a>. <i>achievement.org</i>. <a href="/wiki/American_Academy_of_Achievement" class="mw-redirect" title="American Academy of Achievement">American Academy of Achievement</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=achievement.org&rft.atitle=Golden+Plate+Awardees+of+the+American+Academy+of+Achievement&rft_id=https%3A%2F%2Fachievement.org%2Four-history%2Fgolden-plate-awards%2F%23science-exploration&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-20"><span class="mw-cite-backlink"><b><a href="#cite_ref-20">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://support.beechencliff.org.uk/prospect/prospectus3.html">"Beechen Cliff School website"</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Beechen+Cliff+School+website&rft_id=http%3A%2F%2Fsupport.beechencliff.org.uk%2Fprospect%2Fprospectus3.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span> <sup class="noprint Inline-Template"><span style="white-space: nowrap;">[<i><a href="/wiki/Wikipedia:Link_rot" title="Wikipedia:Link rot"><span title=" Dead link tagged August 2016">dead link</span></a></i><span style="visibility:hidden; color:transparent; padding-left:2px">‍</span>]</span></sup></span> </li> <li id="cite_note-21"><span class="mw-cite-backlink"><b><a href="#cite_ref-21">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20121020110719/http://www.americanhumanist.org/humanism/Humanist_Manifesto_II">"Humanist Manifesto II"</a>. American Humanist Association. Archived from <a rel="nofollow" class="external text" href="http://www.americanhumanist.org/Humanism/Humanist_Manifesto_II">the original</a> on 20 October 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">4 October</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Humanist+Manifesto+II&rft.pub=American+Humanist+Association&rft_id=http%3A%2F%2Fwww.americanhumanist.org%2FHumanism%2FHumanist_Manifesto_II&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-22"><span class="mw-cite-backlink"><b><a href="#cite_ref-22">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFReville2006" class="citation news cs1">Reville, William (2006). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20160305035023/http://www.irishtimes.com/news/a-bright-journey-to-atheism-or-a-road-that-ignores-all-the-signs-1.1040886">"A bright journey to atheism, or a road that ignores all the signs?"</a>. <i><a href="/wiki/The_Irish_Times" title="The Irish Times">The Irish Times</a></i>. Dublin. Archived from <a rel="nofollow" class="external text" href="https://www.irishtimes.com/news/a-bright-journey-to-atheism-or-a-road-that-ignores-all-the-signs-1.1040886">the original</a> on 5 March 2016.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Irish+Times&rft.atitle=A+bright+journey+to+atheism%2C+or+a+road+that+ignores+all+the+signs%3F&rft.date=2006&rft.aulast=Reville&rft.aufirst=William&rft_id=https%3A%2F%2Fwww.irishtimes.com%2Fnews%2Fa-bright-journey-to-atheism-or-a-road-that-ignores-all-the-signs-1.1040886&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-23"><span class="mw-cite-backlink"><b><a href="#cite_ref-23">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation news cs1"><a rel="nofollow" class="external text" href="https://www.independent.co.uk/news/uk/home-news/queens-birthday-honours-full-list-846831.html">"Queen's Birthday Honours: Full list | the Independent"</a>. <i>The Independent</i>. 13 June 2008.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Independent&rft.atitle=Queen%27s+Birthday+Honours%3A+Full+list+%26%23124%3B+the+Independent&rft.date=2008-06-13&rft_id=https%3A%2F%2Fwww.independent.co.uk%2Fnews%2Fuk%2Fhome-news%2Fqueens-birthday-honours-full-list-846831.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-24"><span class="mw-cite-backlink"><b><a href="#cite_ref-24">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://patient-innovation.com">Patient Innovation</a></span> </li> <li id="cite_note-25"><span class="mw-cite-backlink"><b><a href="#cite_ref-25">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.futuredocs.com/speakers/">"Speakers"</a>. <i>Congress of Future Medical Leaders</i><span class="reference-accessdate">. Retrieved <span class="nowrap">31 March</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Congress+of+Future+Medical+Leaders&rft.atitle=Speakers&rft_id=https%3A%2F%2Fwww.futuredocs.com%2Fspeakers%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-26"><span class="mw-cite-backlink"><b><a href="#cite_ref-26">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKeith_Edmisten2018" class="citation web cs1">Keith Edmisten (9 July 2018). <a rel="nofollow" class="external text" href="https://agbiotech.ces.ncsu.edu/2018/07/gmos-are-vital-against-hunger-says-nobel-laureate-sir-richard-j-roberts/">"<span class="cs1-kern-left"></span>"GMOs Are Vital Against Hunger," Says Nobel Laureate Sir Richard J. Roberts"</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=%22GMOs+Are+Vital+Against+Hunger%2C%22+Says+Nobel+Laureate+Sir+Richard+J.+Roberts&rft.date=2018-07-09&rft.au=Keith+Edmisten&rft_id=https%3A%2F%2Fagbiotech.ces.ncsu.edu%2F2018%2F07%2Fgmos-are-vital-against-hunger-says-nobel-laureate-sir-richard-j-roberts%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> <li id="cite_note-27"><span class="mw-cite-backlink"><b><a href="#cite_ref-27">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFEd_Regis2019" class="citation book cs1">Ed Regis (8 October 2019). <i>Golden Rice: The Imperiled Birth of a GMO Superfood</i>. Johns Hopkins University Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1421433035" title="Special:BookSources/978-1421433035"><bdi>978-1421433035</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Golden+Rice%3A+The+Imperiled+Birth+of+a+GMO+Superfood&rft.pub=Johns+Hopkins+University+Press&rft.date=2019-10-08&rft.isbn=978-1421433035&rft.au=Ed+Regis&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARichard+J.+Roberts" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Richard_J._Roberts&action=edit&section=5" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/448">Richard J. Roberts</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q309816#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> including the Nobel Lecture <i>An Amazing Distortion in DNA Induced by a Methyltransferase</i></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist dd:last-child::after,.mw-parser-output .hlist dt:last-child::after,.mw-parser-output .hlist li:last-child::after{content:none}.mw-parser-output .hlist dd dd:first-child::before,.mw-parser-output .hlist dd dt:first-child::before,.mw-parser-output .hlist dd li:first-child::before,.mw-parser-output .hlist dt dd:first-child::before,.mw-parser-output .hlist dt dt:first-child::before,.mw-parser-output .hlist dt li:first-child::before,.mw-parser-output .hlist li dd:first-child::before,.mw-parser-output .hlist li dt:first-child::before,.mw-parser-output .hlist li li:first-child::before{content:" (";font-weight:normal}.mw-parser-output .hlist dd dd:last-child::after,.mw-parser-output .hlist dd dt:last-child::after,.mw-parser-output .hlist dd li:last-child::after,.mw-parser-output .hlist dt dd:last-child::after,.mw-parser-output .hlist dt dt:last-child::after,.mw-parser-output .hlist dt li:last-child::after,.mw-parser-output .hlist li dd:last-child::after,.mw-parser-output .hlist li dt:last-child::after,.mw-parser-output .hlist li li:last-child::after{content:")";font-weight:normal}.mw-parser-output .hlist ol{counter-reset:listitem}.mw-parser-output .hlist ol>li{counter-increment:listitem}.mw-parser-output .hlist ol>li::before{content:" "counter(listitem)"\a0 "}.mw-parser-output .hlist dd ol>li:first-child::before,.mw-parser-output .hlist dt ol>li:first-child::before,.mw-parser-output .hlist li ol>li:first-child::before{content:" ("counter(listitem)"\a0 "}</style><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output .navbox-list-with-group{text-align:left;border-left-width:2px;border-left-style:solid}.mw-parser-output tr+tr>.navbox-abovebelow,.mw-parser-output tr+tr>.navbox-group,.mw-parser-output tr+tr>.navbox-image,.mw-parser-output tr+tr>.navbox-list{border-top:2px solid #fdfdfd}.mw-parser-output .navbox-title{background-color:#ccf}.mw-parser-output .navbox-abovebelow,.mw-parser-output .navbox-group,.mw-parser-output .navbox-subgroup .navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Fellows_of_the_Royal_Society_elected_in_1995" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:FRS_1995" title="Template:FRS 1995"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:FRS_1995" title="Template talk:FRS 1995"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:FRS_1995" title="Special:EditPage/Template:FRS 1995"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Fellows_of_the_Royal_Society_elected_in_1995" style="font-size:114%;margin:0 4em"><a href="/wiki/Fellow_of_the_Royal_Society" title="Fellow of the Royal Society">Fellows</a> of the <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a> elected <a href="/wiki/List_of_Fellows_of_the_Royal_Society_elected_in_1995" class="mw-redirect" title="List of Fellows of the Royal Society elected in 1995">in 1995</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Fellows</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li> <a href="/wiki/Martin_A._Bennett" title="Martin A. Bennett">Martin A. Bennett</a></li> <li> <a href="/wiki/Iain_Donald_Campbell" title="Iain Donald Campbell">Iain Campbell</a></li> <li> <a href="/wiki/Johnson_Cann" title="Johnson Cann">Johnson Cann</a></li> <li> <a href="/wiki/Keith_Chater" title="Keith Chater">Keith Chater</a></li> <li> <a href="/wiki/Ed_Corrigan" title="Ed Corrigan">Ed Corrigan</a></li> <li> <a href="/wiki/E._Brian_Davies" title="E. Brian Davies">E. Brian Davies</a></li> <li> <a href="/wiki/Graham_Dixon-Lewis" title="Graham Dixon-Lewis">Graham Dixon-Lewis</a></li> <li> <a href="/wiki/Richard_Ellis_(astronomer)" title="Richard Ellis (astronomer)">Richard Ellis</a></li> <li> <a href="/wiki/Graham_Farquhar" title="Graham Farquhar">Graham Farquhar</a></li> <li> <a href="/wiki/Jeffrey_Harborne" title="Jeffrey Harborne">Jeffrey Harborne</a></li> <li> <a href="/wiki/Julia_Higgins" title="Julia Higgins">Julia Higgins</a></li> <li> <a href="https://www.wikidata.org/wiki/Q21166333" class="extiw" title="d:Q21166333">Jonathan Charles Howard</a></li> <li> <a href="/wiki/Antony_Jameson" title="Antony Jameson">Antony Jameson</a></li> <li> <a href="https://www.wikidata.org/wiki/Q21165176" class="extiw" title="d:Q21165176">Jack Henri Kaplan</a></li> <li> <a href="/wiki/Gurdev_Khush" title="Gurdev Khush">Gurdev Khush</a></li> <li> <a href="/wiki/Anthony_Ledwith" title="Anthony Ledwith">Anthony Ledwith</a> </li> <li> <a href="/wiki/Frank_Matthews_Leslie" title="Frank Matthews Leslie">Frank Matthews Leslie</a></li> <li> <a href="/wiki/Robin_Marshall" title="Robin Marshall">Robin Marshall</a></li> <li> <a href="/wiki/Christopher_Marshall_(doctor)" title="Christopher Marshall (doctor)">Christopher John Marshall</a></li> <li> <a href="/wiki/Paul_Mason_(meteorologist)" title="Paul Mason (meteorologist)">Paul Mason</a></li> <li> <a href="/wiki/David_A._B._Miller" title="David A. B. Miller">David Miller</a></li> <li> <a href="/wiki/Richard_Alan_North" title="Richard Alan North">Richard Alan North</a></li> <li> <a href="/wiki/Tim_Pedley" title="Tim Pedley">Tim Pedley</a></li> <li> <a href="https://www.wikidata.org/wiki/Q21166451" class="extiw" title="d:Q21166451">Geoffrey James Pert</a></li> <li> <a href="/wiki/Keith_Peters_(physician)" title="Keith Peters (physician)">Keith Peters</a></li> <li> <a href="https://www.wikidata.org/wiki/Q21165499" class="extiw" title="d:Q21165499">Jeremy David Pickett-Heaps</a></li> <li> <a class="mw-selflink selflink">Richard J. Roberts</a></li> <li> <a href="/wiki/Joseph_Rotblat" title="Joseph Rotblat">Joseph Rotblat</a></li> <li> <a href="/wiki/Jeremy_Sanders" title="Jeremy Sanders">Jeremy Sanders</a></li> <li> <a href="/wiki/Robert_Malcolm_Simmons" title="Robert Malcolm Simmons">Robert Malcolm Simmons</a></li> <li> <a href="/wiki/Ian_William_Murison_Smith" title="Ian William Murison Smith">Ian William Murison Smith</a></li> <li> <a href="/wiki/Peter_Sneath" title="Peter Sneath">Peter Sneath</a></li> <li> <a href="https://www.wikidata.org/wiki/Q21166522" class="extiw" title="d:Q21166522">Laszlo Solymar</a></li> <li> <a href="/wiki/Roger_Tayler" title="Roger Tayler">Roger Tayler</a></li> <li> <a href="/wiki/Richard_Taylor_(mathematician)" title="Richard Taylor (mathematician)">Richard Taylor</a></li> <li> <a href="/wiki/Shirley_M._Tilghman" title="Shirley M. Tilghman">Shirley M. Tilghman</a></li> <li> <a href="/wiki/John_E._Walker" title="John E. Walker">John E. Walker</a></li> <li> <a href="/wiki/Stephen_C._West" title="Stephen C. West">Stephen C. West</a></li> <li> <a href="/wiki/Colin_Windsor" title="Colin Windsor">Colin Windsor</a></li> <li> <a href="/wiki/Andrew_Wyllie_(pathologist)" title="Andrew Wyllie (pathologist)">Andrew Wyllie</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Foreign</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li> <a href="/wiki/Ugo_Fano" title="Ugo Fano">Ugo Fano</a></li> <li> <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a></li> <li> <a href="/wiki/Salome_Gluecksohn-Waelsch" title="Salome Gluecksohn-Waelsch">Salome Gluecksohn-Waelsch</a></li> <li> <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></li> <li> <a href="/wiki/Calvin_Quate" title="Calvin Quate">Calvin Quate</a></li> <li> <a href="/wiki/John_Archibald_Wheeler" title="John Archibald Wheeler">John Archibald Wheeler</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1239334494">@media screen{html.skin-theme-clientpref-night .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"></div><div role="navigation" class="navbox" aria-labelledby="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Template:Nobel Prize in Physiology or Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Nobel_Prize_in_Physiology_or_Medicine" title="Template talk:Nobel Prize in Physiology or Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Special:EditPage/Template:Nobel Prize in Physiology or Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Physiology_or_Medicine" title="List of Nobel laureates in Physiology or Medicine">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_Macleod_(physiologist)" title="John Macleod (physiologist)">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1925</span></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Krebs_(biochemist)" title="Hans Krebs (biochemist)">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Beadle" title="George Beadle">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_MacLeod_Cormack" title="Allan MacLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a class="mw-selflink selflink">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards_(physiologist)" title="Robert Edwards (physiologist)">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>posthumously</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe_(neuroscientist)" title="John O'Keefe (neuroscientist)">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap"><a href="/wiki/2020_Nobel_Prize_in_Physiology_or_Medicine" title="2020 Nobel Prize in Physiology or Medicine">2020</a>: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton_(virologist)" class="mw-redirect" title="Michael Houghton (virologist)">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap"><a href="/wiki/2021_Nobel_Prize_in_Physiology_or_Medicine" title="2021 Nobel Prize in Physiology or Medicine">2021</a>: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap"><a href="/wiki/2022_Nobel_Prize_in_Physiology_or_Medicine" title="2022 Nobel Prize in Physiology or Medicine">2022</a>: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap"><a href="/wiki/2023_Nobel_Prize_in_Physiology_or_Medicine" title="2023 Nobel Prize in Physiology or Medicine">2023</a>: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="1993_Nobel_Prize_laureates" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:1993_Nobel_Prize_winners" title="Template:1993 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:1993_Nobel_Prize_winners" title="Template talk:1993 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:1993_Nobel_Prize_winners" title="Special:EditPage/Template:1993 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="1993_Nobel_Prize_laureates" style="font-size:114%;margin:0 4em">1993 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Kary_Mullis" title="Kary Mullis">Kary B. Mullis</a> (United States)</li> <li><a href="/wiki/Michael_Smith_(chemist)" title="Michael Smith (chemist)">Michael Smith</a> (Canada)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/1993_Nobel_Prize_in_Literature" title="1993 Nobel Prize in Literature">1993</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><a href="/wiki/Toni_Morrison" title="Toni Morrison">Toni Morrison</a> (United States)</div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nelson_Mandela" title="Nelson Mandela">Nelson Mandela</a> (South Africa)</li> <li><a href="/wiki/F._W._de_Klerk" title="F. W. de Klerk">Frederik Willem de Klerk</a> (South Africa)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Russell_Alan_Hulse" title="Russell Alan Hulse">Russell Alan Hulse</a> (United States)</li> <li><a href="/wiki/Joseph_Hooton_Taylor_Jr." title="Joseph Hooton Taylor Jr.">Joseph Hooton Taylor Jr.</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Richard J. Roberts</a> (United Kingdom)</li> <li><a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip A. Sharp</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economic Sciences</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Robert_Fogel" title="Robert Fogel">Robert Fogel</a> (United States)</li> <li><a href="/wiki/Douglass_North" title="Douglass North">Douglass North</a> (United States)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/Template:1988_Nobel_Prize_winners" title="Template:1988 Nobel Prize winners">1988</a></dd> <dd><a href="/wiki/Template:1989_Nobel_Prize_winners" title="Template:1989 Nobel Prize winners">1989</a></dd> <dd><a href="/wiki/Template:1990_Nobel_Prize_winners" title="Template:1990 Nobel Prize winners">1990</a></dd> <dd><a href="/wiki/Template:1991_Nobel_Prize_winners" title="Template:1991 Nobel Prize winners">1991</a></dd> <dd><a href="/wiki/Template:1992_Nobel_Prize_winners" title="Template:1992 Nobel Prize winners">1992</a></dd> <dd><b>1993 </b></dd> <dd><a href="/wiki/Template:1994_Nobel_Prize_winners" title="Template:1994 Nobel Prize winners">1994</a></dd> <dd><a href="/wiki/Template:1995_Nobel_Prize_winners" title="Template:1995 Nobel Prize winners">1995</a></dd> <dd><a href="/wiki/Template:1996_Nobel_Prize_winners" title="Template:1996 Nobel Prize winners">1996</a></dd> <dd><a href="/wiki/Template:1997_Nobel_Prize_winners" title="Template:1997 Nobel Prize winners">1997</a></dd> <dd><a href="/wiki/Template:1998_Nobel_Prize_winners" title="Template:1998 Nobel Prize winners">1998</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q309816#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q309816#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q309816#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">International</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/000000008357531X">ISNI</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/12459768">VIAF</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/1172887764">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n2020186313">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb134903704">France</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb134903704">BnF data</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p069413231">Netherlands</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://authority.bibsys.no/authority/rest/authorities/html/90248465">Norway</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&local_base=NLX10&find_code=UID&request=987007452710205171">Israel</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Academics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://orcid.org/0000-0002-4348-0169">ORCID</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.uri?authorId=7404281937">Scopus</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://dblp.org/pid/76/1661">DBLP</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/085881813">IdRef</a></span></li></ul></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Richard_J._Roberts&oldid=1254328414">https://en.wikipedia.org/w/index.php?title=Richard_J._Roberts&oldid=1254328414</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:People_from_Bath,_Somerset" title="Category:People from Bath, Somerset">People from Bath, Somerset</a></li><li><a href="/wiki/Category:Scientists_from_Derby" title="Category:Scientists from Derby">Scientists from Derby</a></li><li><a href="/wiki/Category:Nobel_laureates_in_Physiology_or_Medicine" title="Category:Nobel laureates in Physiology or Medicine">Nobel laureates in Physiology or Medicine</a></li><li><a href="/wiki/Category:British_biochemists" title="Category:British biochemists">British biochemists</a></li><li><a href="/wiki/Category:20th-century_British_biologists" title="Category:20th-century British biologists">20th-century British biologists</a></li><li><a href="/wiki/Category:21st-century_British_biologists" title="Category:21st-century British biologists">21st-century British biologists</a></li><li><a href="/wiki/Category:British_Nobel_laureates" title="Category:British Nobel laureates">British Nobel laureates</a></li><li><a href="/wiki/Category:Members_of_the_European_Molecular_Biology_Organization" title="Category:Members of the European Molecular Biology Organization">Members of the European Molecular Biology Organization</a></li><li><a href="/wiki/Category:Alumni_of_the_University_of_Sheffield" title="Category:Alumni of the University of Sheffield">Alumni of the University of Sheffield</a></li><li><a href="/wiki/Category:Knights_Bachelor" title="Category:Knights Bachelor">Knights Bachelor</a></li><li><a href="/wiki/Category:People_educated_at_City_of_Bath_Boys%27_School" title="Category:People educated at City of Bath Boys' School">People educated at City of Bath Boys' School</a></li><li><a href="/wiki/Category:1943_births" title="Category:1943 births">1943 births</a></li><li><a href="/wiki/Category:Living_people" title="Category:Living people">Living people</a></li><li><a href="/wiki/Category:Fellows_of_the_Royal_Society" title="Category:Fellows of the Royal Society">Fellows of the Royal Society</a></li><li><a href="/wiki/Category:English_Nobel_laureates" title="Category:English Nobel laureates">English Nobel laureates</a></li><li><a href="/wiki/Category:English_humanists" title="Category:English humanists">English humanists</a></li><li><a href="/wiki/Category:English_atheists" title="Category:English atheists">English atheists</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Hidden categories: <ul><li><a href="/wiki/Category:CS1_maint:_bot:_original_URL_status_unknown" title="Category:CS1 maint: bot: original URL status unknown">CS1 maint: bot: original URL status unknown</a></li><li><a href="/wiki/Category:Pages_containing_links_to_subscription-only_content" title="Category:Pages containing links to subscription-only content">Pages containing links to subscription-only content</a></li><li><a href="/wiki/Category:All_articles_with_dead_external_links" title="Category:All articles with dead external links">All articles with dead external links</a></li><li><a href="/wiki/Category:Articles_with_dead_external_links_from_August_2016" title="Category:Articles with dead external links from August 2016">Articles with dead external links from August 2016</a></li><li><a href="/wiki/Category:Articles_with_short_description" title="Category:Articles with short description">Articles with short description</a></li><li><a href="/wiki/Category:Short_description_is_different_from_Wikidata" title="Category:Short description is different from Wikidata">Short description is different from Wikidata</a></li><li><a href="/wiki/Category:Use_British_English_from_August_2011" title="Category:Use British English from August 2011">Use British English from August 2011</a></li><li><a href="/wiki/Category:Use_dmy_dates_from_December_2020" title="Category:Use dmy dates from December 2020">Use dmy dates from December 2020</a></li><li><a href="/wiki/Category:Articles_with_hCards" title="Category:Articles with hCards">Articles with hCards</a></li><li><a href="/wiki/Category:Nobelprize_template_using_Wikidata_property_P8024" title="Category:Nobelprize template using Wikidata property P8024">Nobelprize template using Wikidata property P8024</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> This page was last edited on 30 October 2024, at 13:32<span class="anonymous-show"> (UTC)</span>.</li> <li id="footer-info-copyright">Text is available under the <a href="/wiki/Wikipedia:Text_of_the_Creative_Commons_Attribution-ShareAlike_4.0_International_License" title="Wikipedia:Text of the Creative Commons Attribution-ShareAlike 4.0 International License">Creative Commons Attribution-ShareAlike 4.0 License</a>; additional terms may apply. By using this site, you agree to the <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Terms_of_Use" class="extiw" title="foundation:Special:MyLanguage/Policy:Terms of Use">Terms of Use</a> and <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy" class="extiw" title="foundation:Special:MyLanguage/Policy:Privacy policy">Privacy Policy</a>. Wikipedia® is a registered trademark of the <a rel="nofollow" class="external text" href="https://wikimediafoundation.org/">Wikimedia Foundation, Inc.</a>, a non-profit organization.</li> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy">Privacy policy</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia:About">About Wikipedia</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:General_disclaimer">Disclaimers</a></li> <li id="footer-places-contact"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us">Contact Wikipedia</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">Code of Conduct</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">Developers</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/en.wikipedia.org">Statistics</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookie statement</a></li> <li id="footer-places-mobileview"><a href="//en.m.wikipedia.org/w/index.php?title=Richard_J._Roberts&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobile view</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-694cf4987f-cvmxh","wgBackendResponseTime":151,"wgPageParseReport":{"limitreport":{"cputime":"0.778","walltime":"1.036","ppvisitednodes":{"value":5527,"limit":1000000},"postexpandincludesize":{"value":147842,"limit":2097152},"templateargumentsize":{"value":10601,"limit":2097152},"expansiondepth":{"value":17,"limit":100},"expensivefunctioncount":{"value":9,"limit":500},"unstrip-depth":{"value":1,"limit":20},"unstrip-size":{"value":136251,"limit":5000000},"entityaccesscount":{"value":1,"limit":400},"timingprofile":["100.00% 876.831 1 -total"," 37.29% 326.972 1 Template:Reflist"," 25.52% 223.786 1 Template:Infobox_scientist"," 15.37% 134.756 7 Template:Cite_journal"," 11.53% 101.098 2 Template:Navbox"," 9.33% 81.826 1 Template:FRS_1995"," 8.80% 77.167 1 Template:Short_description"," 6.78% 59.489 1 Template:Birth_date_and_age"," 5.60% 49.139 12 Template:Cite_web"," 5.58% 48.929 1 Template:Dead_link"]},"scribunto":{"limitreport-timeusage":{"value":"0.450","limit":"10.000"},"limitreport-memusage":{"value":8496646,"limit":52428800}},"cachereport":{"origin":"mw-api-int.codfw.canary-6594497df4-s24xp","timestamp":"20241125050407","ttl":2592000,"transientcontent":false}}});});</script> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"Richard J. Roberts","url":"https:\/\/en.wikipedia.org\/wiki\/Richard_J._Roberts","sameAs":"http:\/\/www.wikidata.org\/entity\/Q309816","mainEntity":"http:\/\/www.wikidata.org\/entity\/Q309816","author":{"@type":"Organization","name":"Contributors to Wikimedia projects"},"publisher":{"@type":"Organization","name":"Wikimedia Foundation, Inc.","logo":{"@type":"ImageObject","url":"https:\/\/www.wikimedia.org\/static\/images\/wmf-hor-googpub.png"}},"datePublished":"2005-04-06T13:05:28Z","dateModified":"2024-10-30T13:32:27Z","image":"https:\/\/upload.wikimedia.org\/wikipedia\/commons\/1\/19\/Roberts%2C_Richard_John_%281943%29.jpg","headline":"English biochemist and molecular biologist"}</script> </body> </html>