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Disponible en 47 llengües" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-47" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">47 llengües</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D9%83%D9%88%D9%87%D9%84%D8%B1" title="جورج كوهلر - àrab" lang="ar" hreflang="ar" data-title="جورج كوهلر" data-language-autonym="العربية" data-language-local-name="àrab" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D9%83%D9%88%D9%87%D9%84%D8%B1" title="جورج كوهلر - àrab egipci" lang="arz" hreflang="arz" data-title="جورج كوهلر" data-language-autonym="مصرى" data-language-local-name="àrab egipci" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%DA%98%D8%B1%DA%98_%D8%AC%DB%8C._%D8%A7%D9%81._%DA%A9%D9%88%D9%87%D9%84%D8%B1" title="ژرژ جی. اف. کوهلر - South Azerbaijani" lang="azb" hreflang="azb" data-title="ژرژ جی. اف. کوهلر" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%93%D0%B5%D0%BE%D1%80%D0%B3_%D0%9A%D1%91%D0%BB%D0%B5%D1%80" title="Георг Кёлер - belarús" lang="be" hreflang="be" data-title="Георг Кёлер" data-language-autonym="Беларуская" data-language-local-name="belarús" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%93%D0%B5%D0%BE%D1%80%D0%B3_%D0%9A%D1%8C%D0%BE%D0%BB%D0%B5%D1%80" title="Георг Кьолер - búlgar" lang="bg" hreflang="bg" data-title="Георг Кьолер" data-language-autonym="Български" data-language-local-name="búlgar" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - txec" lang="cs" hreflang="cs" data-title="Georges Köhler" data-language-autonym="Čeština" data-language-local-name="txec" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - alemany" lang="de" hreflang="de" data-title="Georges J. F. Köhler" data-language-autonym="Deutsch" data-language-local-name="alemany" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - anglès" lang="en" hreflang="en" data-title="Georges J. F. Köhler" data-language-autonym="English" data-language-local-name="anglès" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - espanyol" lang="es" hreflang="es" data-title="Georges Köhler" data-language-autonym="Español" data-language-local-name="espanyol" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Georges_Jean_Franz_K%C3%B6hler" title="Georges Jean Franz Köhler - estonià" lang="et" hreflang="et" data-title="Georges Jean Franz Köhler" data-language-autonym="Eesti" data-language-local-name="estonià" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - basc" lang="eu" hreflang="eu" data-title="Georges J. F. Köhler" data-language-autonym="Euskara" data-language-local-name="basc" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DA%98%D8%B1%DA%98_%D8%AC%DB%8C.%D8%A7%D9%81._%DA%A9%D9%88%D9%87%D9%84%D8%B1" title="ژرژ جی.اف. کوهلر - persa" lang="fa" hreflang="fa" data-title="ژرژ جی.اف. کوهلر" data-language-autonym="فارسی" data-language-local-name="persa" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - finès" lang="fi" hreflang="fi" data-title="Georges J. F. Köhler" data-language-autonym="Suomi" data-language-local-name="finès" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - francès" lang="fr" hreflang="fr" data-title="Georges Köhler" data-language-autonym="Français" data-language-local-name="francès" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - irlandès" lang="ga" hreflang="ga" data-title="Georges J. F. Köhler" data-language-autonym="Gaeilge" data-language-local-name="irlandès" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%96%27%D7%95%D7%A8%D7%96%27_%D7%A7%D7%9C%D7%A8" title="ז'ורז' קלר - hebreu" lang="he" hreflang="he" data-title="ז'ורז' קלר" data-language-autonym="עברית" data-language-local-name="hebreu" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - croat" lang="hr" hreflang="hr" data-title="Georges J. F. Köhler" data-language-autonym="Hrvatski" data-language-local-name="croat" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - hongarès" lang="hu" hreflang="hu" data-title="Georges J. F. Köhler" data-language-autonym="Magyar" data-language-local-name="hongarès" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - indonesi" lang="id" hreflang="id" data-title="Georges Köhler" data-language-autonym="Bahasa Indonesia" data-language-local-name="indonesi" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - ido" lang="io" hreflang="io" data-title="Georges J. F. Köhler" data-language-autonym="Ido" data-language-local-name="ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - italià" lang="it" hreflang="it" data-title="Georges Köhler" data-language-autonym="Italiano" data-language-local-name="italià" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B8%E3%83%A7%E3%83%AB%E3%82%B8%E3%83%A5%E3%83%BBJ%E3%83%BBF%E3%83%BB%E3%82%B1%E3%83%BC%E3%83%A9%E3%83%BC" title="ジョルジュ・J・F・ケーラー - japonès" lang="ja" hreflang="ja" data-title="ジョルジュ・J・F・ケーラー" data-language-autonym="日本語" data-language-local-name="japonès" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%96%D0%BE%D1%80%D0%B6_%D0%9A%D1%91%D0%BB%D0%B5%D1%80" title="Жорж Кёлер - kazakh" lang="kk" hreflang="kk" data-title="Жорж Кёлер" data-language-autonym="Қазақша" data-language-local-name="kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EA%B2%8C%EC%98%A4%EB%A5%B4%EA%B2%8C%EC%8A%A4_J._F._%EC%BE%B0%EB%9F%AC" title="게오르게스 J. F. 쾰러 - coreà" lang="ko" hreflang="ko" data-title="게오르게스 J. F. 쾰러" data-language-autonym="한국어" data-language-local-name="coreà" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Georgius_J._F._K%C3%B6hler" title="Georgius J. F. Köhler - llatí" lang="la" hreflang="la" data-title="Georgius J. F. Köhler" data-language-autonym="Latina" data-language-local-name="llatí" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - malai" lang="ms" hreflang="ms" data-title="Georges J. F. Köhler" data-language-autonym="Bahasa Melayu" data-language-local-name="malai" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mt mw-list-item"><a href="https://mt.wikipedia.org/wiki/Georges_J.F._Kohler" title="Georges J.F. Kohler - maltès" lang="mt" hreflang="mt" data-title="Georges J.F. Kohler" data-language-autonym="Malti" data-language-local-name="maltès" class="interlanguage-link-target"><span>Malti</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%DA%A9%D9%88%D9%87%D9%84%D8%B1" title="جورج کوهلر - mazanderani" lang="mzn" hreflang="mzn" data-title="جورج کوهلر" data-language-autonym="مازِرونی" data-language-local-name="mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler - neerlandès" lang="nl" hreflang="nl" data-title="Georges J.F. Köhler" data-language-autonym="Nederlands" data-language-local-name="neerlandès" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler - noruec bokmål" lang="nb" hreflang="nb" data-title="Georges J.F. Köhler" data-language-autonym="Norsk bokmål" data-language-local-name="noruec bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - occità" lang="oc" hreflang="oc" data-title="Georges J. F. Köhler" data-language-autonym="Occitan" data-language-local-name="occità" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - polonès" lang="pl" hreflang="pl" data-title="Georges Köhler" data-language-autonym="Polski" data-language-local-name="polonès" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%D8%A7%D8%B1%D8%AC%D8%B2_%DA%A9%D9%88%DB%81%D9%84%D8%B1" title="جارجز کوہلر - Western Punjabi" lang="pnb" hreflang="pnb" data-title="جارجز کوہلر" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - portuguès" lang="pt" hreflang="pt" data-title="Georges Köhler" data-language-autonym="Português" data-language-local-name="portuguès" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - romanès" lang="ro" hreflang="ro" data-title="Georges J. F. Köhler" data-language-autonym="Română" data-language-local-name="romanès" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9A%D1%91%D0%BB%D0%B5%D1%80,_%D0%96%D0%BE%D1%80%D0%B6" title="Кёлер, Жорж - rus" lang="ru" hreflang="ru" data-title="Кёлер, Жорж" data-language-autonym="Русский" data-language-local-name="rus" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - Simple English" lang="en-simple" hreflang="en-simple" data-title="Georges J. F. Köhler" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler - suec" lang="sv" hreflang="sv" data-title="Georges J.F. Köhler" data-language-autonym="Svenska" data-language-local-name="suec" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler - suahili" lang="sw" hreflang="sw" data-title="Georges Köhler" data-language-autonym="Kiswahili" data-language-local-name="suahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - turc" lang="tr" hreflang="tr" data-title="Georges J. F. Köhler" data-language-autonym="Türkçe" data-language-local-name="turc" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%93%D0%B5%D0%BE%D1%80%D0%B3_%D0%9A%D0%B5%D0%BB%D0%B5%D1%80" title="Георг Келер - ucraïnès" lang="uk" hreflang="uk" data-title="Георг Келер" data-language-autonym="Українська" data-language-local-name="ucraïnès" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%AC%D8%A7%D8%B1%D8%AC%D8%B2_%D8%AC%DB%92_%D8%A7%DB%8C%D9%81_%DA%A9%D9%88%DB%81%D9%84%D8%B1" title="جارجز جے ایف کوہلر - urdú" lang="ur" hreflang="ur" data-title="جارجز جے ایف کوہلر" data-language-autonym="اردو" data-language-local-name="urdú" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - uzbek" lang="uz" hreflang="uz" data-title="Georges J. F. Köhler" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - vietnamita" lang="vi" hreflang="vi" data-title="Georges J. F. Köhler" data-language-autonym="Tiếng Việt" data-language-local-name="vietnamita" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%92%E1%83%94%E1%83%9D%E1%83%A0%E1%83%92_%E1%83%99%E1%83%94%E1%83%9A%E1%83%94%E1%83%A0%E1%83%98" title="გეორგ კელერი - mingrelià" lang="xmf" hreflang="xmf" data-title="გეორგ კელერი" data-language-autonym="მარგალური" data-language-local-name="mingrelià" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler - ioruba" lang="yo" hreflang="yo" data-title="Georges J. F. Köhler" data-language-autonym="Yorùbá" data-language-local-name="ioruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E4%B9%94%E6%B2%BB%C2%B7%E5%85%8B%E5%8B%92" title="乔治·克勒 - xinès" lang="zh" hreflang="zh" data-title="乔治·克勒" data-language-autonym="中文" data-language-local-name="xinès" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q77160#sitelinks-wikipedia" title="Modifica enllaços interlingües" class="wbc-editpage">Modifica els enllaços</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Espais de noms"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet 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wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Georges_K%C3%B6hler" hreflang="en"><span>Commons</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q77160" title="Enllaç a l'element del repositori de dades connectat [g]" accesskey="g"><span>Element a Wikidata</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Eines de la pàgina"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Aparença"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div 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href="mw-data:TemplateStyles:r33733479"> <table class="infobox" style="font-size:90%;width:25em;infobox_bodystyle"><caption class="infobox_titleclass" style="font-weight:bold"><span style="float:left;"><span class="skin-invert" typeof="mw:File"><a href="/wiki/Fitxer:Simpleicons_Interface_user-outline.svg" class="mw-file-description" title="Plantilla:Infotaula persona"><img alt="Plantilla:Infotaula persona" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3f/Simpleicons_Interface_user-outline.svg/20px-Simpleicons_Interface_user-outline.svg.png" decoding="async" width="20" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3f/Simpleicons_Interface_user-outline.svg/30px-Simpleicons_Interface_user-outline.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3f/Simpleicons_Interface_user-outline.svg/40px-Simpleicons_Interface_user-outline.svg.png 2x" data-file-width="485" data-file-height="485" /></a></span></span><span class="fn">Georges Jean Franz Köhler</span></caption><tbody><tr><td colspan="2" class="infobox-full-data infobox-data" style="text-align:center"><span typeof="mw:File"><a href="/wiki/Fitxer:40._Tagung_1990_Mediziner_Portraits;_Prof._K%C3%B6hler_(Freiburg)_-_W134Nr.126381c1_-_Willy_Pragher_(cropped).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/83/40._Tagung_1990_Mediziner_Portraits%3B_Prof._K%C3%B6hler_%28Freiburg%29_-_W134Nr.126381c1_-_Willy_Pragher_%28cropped%29.jpg/188px-40._Tagung_1990_Mediziner_Portraits%3B_Prof._K%C3%B6hler_%28Freiburg%29_-_W134Nr.126381c1_-_Willy_Pragher_%28cropped%29.jpg" decoding="async" width="188" height="300" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/83/40._Tagung_1990_Mediziner_Portraits%3B_Prof._K%C3%B6hler_%28Freiburg%29_-_W134Nr.126381c1_-_Willy_Pragher_%28cropped%29.jpg/282px-40._Tagung_1990_Mediziner_Portraits%3B_Prof._K%C3%B6hler_%28Freiburg%29_-_W134Nr.126381c1_-_Willy_Pragher_%28cropped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/83/40._Tagung_1990_Mediziner_Portraits%3B_Prof._K%C3%B6hler_%28Freiburg%29_-_W134Nr.126381c1_-_Willy_Pragher_%28cropped%29.jpg/376px-40._Tagung_1990_Mediziner_Portraits%3B_Prof._K%C3%B6hler_%28Freiburg%29_-_W134Nr.126381c1_-_Willy_Pragher_%28cropped%29.jpg 2x" data-file-width="423" data-file-height="675" /></a></span><br /> (1990) <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P18" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th colspan="2" class="infobox-header infobox_headerclass" style="text-align:center">Biografia</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Naixement</th><td class="infobox-data" style="text-align:start">17 abril 1946 <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P569" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span><br /><a href="/wiki/M%C3%BAnic" title="Múnic">Múnic (Alemanya)</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P19" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Mort</th><td class="infobox-data" style="text-align:start">1r març 1995 <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P570" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span> (48 anys)<br /><a href="/wiki/Friburg_de_Brisg%C3%B2via" title="Friburg de Brisgòvia">Friburg de Brisgòvia (Alemanya)</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P20" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Causa de mort</th><td class="infobox-data" style="text-align:start"><a href="/wiki/Infart_de_miocardi" title="Infart de miocardi">infart de miocardi</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P509" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Formació</th><td class="infobox-data" style="text-align:start"><a href="/wiki/Universitat_de_Friburg_de_Brisg%C3%B2via" title="Universitat de Friburg de Brisgòvia">Universitat de Friburg de Brisgòvia</a> <small> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P69" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></small></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Director de tesi</th><td class="infobox-data" style="text-align:start"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1504324" class="extiw" title="d:Special:EntityPage/Q1504324">Fritz Melchers</a> i <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P184" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th colspan="2" class="infobox-header infobox_headerclass" style="text-align:center">Activitat</th></tr><tr><th scope="row" class="infobox-label" style="text-align:left"><span style="white-space:nowrap;">Camp de treball</span></th><td class="infobox-data" style="text-align:start"><a href="/wiki/Immunologia" title="Immunologia">Immunologia</a> i <a href="/wiki/Biologia" title="Biologia">biologia</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P101" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left"><span style="white-space:nowrap;">Lloc de treball </span></th><td class="infobox-data" style="text-align:start"><a href="/wiki/Friburg_de_Brisg%C3%B2via" title="Friburg de Brisgòvia">Friburg de Brisgòvia</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P937" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Ocupació</th><td class="infobox-data" style="text-align:start"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q864503" class="extiw" title="d:Special:EntityPage/Q864503">biòleg</a>, <a href="/wiki/Qu%C3%ADmic" title="Químic">químic</a>, <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q12119633" class="extiw" title="d:Special:EntityPage/Q12119633">immunòleg</a>, <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1622272" class="extiw" title="d:Special:EntityPage/Q1622272">professor d'universitat</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P106" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Ocupador</th><td class="infobox-data" style="text-align:start"><a href="/wiki/Universitat_de_Friburg_de_Brisg%C3%B2via" title="Universitat de Friburg de Brisgòvia">Universitat de Friburg de Brisgòvia</a> <small> </small><br /><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1912076" class="extiw" title="d:Special:EntityPage/Q1912076">Max Planck Institute of Immunobiology and Epigenetics</a> <sup>(en)</sup> <span class="mw-valign-baseline skin-invert" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q1912076?uselang=ca" title="Tradueix"><img alt="Tradueix" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/10px-Noun_Project_label_icon_1116097_cc_mirror.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/15px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/70/Noun_Project_label_icon_1116097_cc_mirror.svg/20px-Noun_Project_label_icon_1116097_cc_mirror.svg.png 2x" data-file-width="158" data-file-height="161" /></a></span> <small> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P108" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></small></td></tr><tr><th scope="row" class="infobox-label" style="text-align:left">Membre de</th><td class="infobox-data" style="text-align:start"><div style="text-align:start;"><a href="/wiki/Akademie_der_Wissenschaften" title="Akademie der Wissenschaften">Acadèmia Alemanya de Ciències Leopoldina</a> <br /><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1376791" class="extiw" title="d:Special:EntityPage/Q1376791">Organització Europea de Biologia Molecular</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P463" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></div></td></tr><tr><th colspan="2" class="infobox-header infobox_headerclass" style="text-align:center">Premis</th></tr><tr><td colspan="2" class="infobox-full-data infobox-data" style="text-align:center;text-align:start"><div style="text-align:start;"><ul><li><small>(1986)</small> <span typeof="mw:File"><span><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9b/DE-BY_Der_Bayerische_Maximiliansorden_f%C3%BCr_Wissenschaft_und_Kunst_BAR.png/30px-DE-BY_Der_Bayerische_Maximiliansorden_f%C3%BCr_Wissenschaft_und_Kunst_BAR.png" decoding="async" width="30" height="8" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9b/DE-BY_Der_Bayerische_Maximiliansorden_f%C3%BCr_Wissenschaft_und_Kunst_BAR.png/45px-DE-BY_Der_Bayerische_Maximiliansorden_f%C3%BCr_Wissenschaft_und_Kunst_BAR.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9b/DE-BY_Der_Bayerische_Maximiliansorden_f%C3%BCr_Wissenschaft_und_Kunst_BAR.png/60px-DE-BY_Der_Bayerische_Maximiliansorden_f%C3%BCr_Wissenschaft_und_Kunst_BAR.png 2x" data-file-width="218" data-file-height="60" /></span></span> <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q565030" class="extiw" title="d:Special:EntityPage/Q565030">Orde bavaresa de Maximilià per la Ciència i les Arts</a></li><li><small>(1985)</small>  <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1046023" class="extiw" title="d:Special:EntityPage/Q1046023">Medalla Carus</a></li><li><small>(1984)</small>  <a href="/wiki/Premi_Albert_Lasker_d%27Investigaci%C3%B3_M%C3%A8dica_B%C3%A0sica" title="Premi Albert Lasker d'Investigació Mèdica Bàsica">Premi Albert Lasker d'Investigació Mèdica Bàsica</a></li><li><small>(1984)</small>  <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q3332215" class="extiw" title="d:Special:EntityPage/Q3332215">Medalla John Scott</a></li><li><small>(1984)</small> <span typeof="mw:File"><span><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/30px-Nobel_prize_medal.svg.png" decoding="async" width="30" height="30" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/45px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/60px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></span></span> <a href="/wiki/Premi_Nobel_de_Fisiologia_o_Medicina" title="Premi Nobel de Fisiologia o Medicina">Premi Nobel de Medicina o Fisiologia</a></li><li><small>(28 maig 1983)</small>  <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q105979196" class="extiw" title="d:Special:EntityPage/Q105979196">doctor honoris causa per la Universitat de Hasselt</a></li><li><small>(1981)</small>  <a href="/wiki/Premi_Internacional_Gairdner_de_Canad%C3%A0" title="Premi Internacional Gairdner de Canadà">Premi Gairdner</a></li><li><small>(1978)</small>  <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1468630" class="extiw" title="d:Special:EntityPage/Q1468630">Premi Heinz Maier-Leibnitz</a></li><li>  <a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1732108" class="extiw" title="d:Special:EntityPage/Q1732108">Premi Memorial Karl Landsteiner</a> <span class="penicon"><span class="mw-valign-baseline" typeof="mw:File"><a href="https://www.wikidata.org/wiki/Q77160?uselang=ca#P166" title="Modifica el valor a Wikidata"><img alt="Modifica el valor a Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/10px-Arbcom_ru_editing.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/15px-Arbcom_ru_editing.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Arbcom_ru_editing.svg/20px-Arbcom_ru_editing.svg.png 2x" data-file-width="600" data-file-height="600" /></a></span></span></li></ul> </div></td></tr></tbody></table> <p><b>Georges Jean Franz Köhler</b> (<a href="/wiki/M%C3%BAnic" title="Múnic">Múnic</a>, <a href="/wiki/Alemanya" title="Alemanya">Alemanya</a> <a href="/wiki/1946" title="1946">1946</a> - <a href="/wiki/Friburg_de_Brisg%C3%B2via" title="Friburg de Brisgòvia">Friburg de Brisgòvia</a> <a href="/wiki/1995" title="1995">1995</a>) fou un <a href="/wiki/Biologia" title="Biologia">biòleg</a> alemany guardonat amb el <a href="/wiki/Premi_Nobel_de_Medicina_o_Fisiologia" class="mw-redirect" title="Premi Nobel de Medicina o Fisiologia">Premi Nobel de Medicina o Fisiologia</a> l'any <a href="/wiki/1984" title="1984">1984</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Biografia">Biografia</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Georges_Jean_Franz_K%C3%B6hler&action=edit&section=1" title="Modifica la secció: Biografia"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Va néixer el <a href="/wiki/17_de_mar%C3%A7" title="17 de març">17 de març</a> de <a href="/wiki/1946" title="1946">1946</a> a la ciutat de <a href="/wiki/M%C3%BAnic" title="Múnic">Múnic</a>, població situada a l'<a href="/wiki/Estats_alemanys" class="mw-redirect" title="Estats alemanys">estat alemany</a> de <a href="/wiki/Baviera" title="Baviera">Baviera</a>. Va estudiar <a href="/wiki/Biologia" title="Biologia">biologia</a> a la <a href="/wiki/Universitat_de_Friburg_de_Brisg%C3%B2via" title="Universitat de Friburg de Brisgòvia">Universitat de Friburg de Brisgòvia</a>, on es va graduar el <a href="/wiki/1971" title="1971">1971</a> i <a href="/wiki/Doctorat" title="Doctorat">doctorar</a> el <a href="/wiki/1974" title="1974">1974</a>. La seva tesi doctoral fou sobre immunologia, amb un treball de recerca que va dur a terme dirigit per Fritz Melchers a l'Institut d'Immunologia de Basilea (Suïssa).<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Durant dos anys fou membre de recerca científica al Laboratori de <a href="/wiki/Biologia_molecular" title="Biologia molecular">biologia molecular</a> a la <a href="/wiki/Universitat_de_Cambridge" title="Universitat de Cambridge">Universitat de Cambridge</a> sota la direcció de <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a>. A partir de <a href="/wiki/1976" title="1976">1976</a> fou membre de l'Institut d'Immunologia de <a href="/wiki/Basilea" title="Basilea">Basilea</a>, on conegué <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Kaj Jerne</a> i el 1984 fou nomenat director de l'<a href="/wiki/Institut_Max_Planck" class="mw-redirect" title="Institut Max Planck">Institut Max Planck</a> d'Immunologia de Friburg, càrrec que va compaginar amb el de catedràtic de la Universitat de Friburg.<sup id="cite_ref-:0_2-0" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>Morí l'<a href="/wiki/1_de_mar%C3%A7" title="1 de març">1 de març</a> de <a href="/wiki/1995" title="1995">1995</a> a la ciutat de <a href="/wiki/Friburg_de_Brisg%C3%B2via" title="Friburg de Brisgòvia">Friburg de Brisgòvia</a>, població situada a l'estat de <a href="/wiki/Baden-W%C3%BCrttemberg" title="Baden-Württemberg">Baden-Württemberg</a>, a conseqüència d'un infart.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Uns mesos abans del seu traspàs se li havia diagnosticat una cardiopatia, però va rebutjar l'ingrés hospitalari per fer-se altres proves i que li implantessin un <a href="/wiki/Stent" title="Stent">stent</a>, si fos necessari, i continuà treballant.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Recerca_científica"><span id="Recerca_cient.C3.ADfica"></span>Recerca científica</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Georges_Jean_Franz_K%C3%B6hler&action=edit&section=2" title="Modifica la secció: Recerca científica"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Al costat de César Milstein aconseguí desenvolupar una nova tècnica <a href="/wiki/Immunologia" title="Immunologia">immunològica</a> que permetia l'obtenció d'<a href="/wiki/Anticossos" class="mw-redirect" title="Anticossos">anticossos</a> purs contra determinats <a href="/wiki/Antigen" title="Antigen">antígens</a>, el que fou anomenat <a href="/wiki/Antic%C3%B2s_monoclonal" title="Anticòs monoclonal">anticòs monoclonal</a>. És una tècnica que permet que les cèl·lules productores d'anticossos en dividir-se generin cèl·lules idèntiques a elles. Van publicar el seu mètode a la revista Nature el 1975 i al final de l'article indicaven: "Aquest tipus de cèl·lules poden cultivar-se in vitro en cultius massius per produir anticossos específics. Cultius d'aquest tipus podrien ser valuosos per al seu ús industrial i en medicina."<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> De fet Avui dia, els anticossos monoclonals són fonamentals en moltes tècniques de <a href="/wiki/Diagn%C3%B2stic" class="mw-redirect" title="Diagnòstic">diagnòstic</a> i en <a href="/wiki/Terap%C3%A8utica" class="mw-redirect" title="Terapèutica">terapèutica</a>.<sup id="cite_ref-:0_2-1" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>L'any <a href="/wiki/1984" title="1984">1984</a>, Köhler compartí el <a href="/wiki/Premi_Nobel_de_Medicina_o_Fisiologia" class="mw-redirect" title="Premi Nobel de Medicina o Fisiologia">Premi Nobel de Medicina o Fisiologia</a> amb Milstein i Jerne, "per les seves teories sobre l'especificitat en el desenvolupament i control del sistema immunitari i el principi per a la producció d'anticossos monoclonals".<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Reconeixements">Reconeixements</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Georges_Jean_Franz_K%C3%B6hler&action=edit&section=3" title="Modifica la secció: Reconeixements"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>En honor seu s'anomenà l'<a href="/wiki/Asteroide" title="Asteroide">asteroide</a> <a href="/w/index.php?title=(11775)_K%C3%B6hler&action=edit&redlink=1" class="new" title="(11775) Köhler (encara no existeix)">(11775) Köhler</a> descobert el <a href="/wiki/16_d%27octubre" title="16 d'octubre">16 d'octubre</a> de <a href="/wiki/1977" title="1977">1977</a> per <a href="/wiki/Cornelis_Johannes_van_Houten" title="Cornelis Johannes van Houten">C. J. van Houten</a>, <a href="/wiki/Ingrid_van_Houten-Groeneveld" title="Ingrid van Houten-Groeneveld">I. van Houten-Groeneveld</a> i <a href="/wiki/Tom_Gehrels" title="Tom Gehrels">Tom Gehrels</a>.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>Des de 1998, la Societat Alemanya d'Immunologia concedeix anualment el Premi Georges Köhler.<sup id="cite_ref-:0_2-2" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Referències"><span id="Refer.C3.A8ncies"></span>Referències</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Georges_Jean_Franz_K%C3%B6hler&action=edit&section=4" title="Modifica la secció: Referències"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="reflist {{#if: | references-column-count references-column-count-{{{col}}}" style="list-style-type: decimal;"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><span class="citation" style="font-style:normal">«<a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/1984/kohler/biographical/">The Nobel Prize in Physiology or Medicine 1984. Georges J.F. Köhler. Biographical</a>» (en anglès). [Consulta: 17 març 2021].</span></span> </li> <li id="cite_note-:0-2"><span class="mw-cite-backlink">↑ <sup><a href="#cite_ref-:0_2-0">2,0</a></sup> <sup><a href="#cite_ref-:0_2-1">2,1</a></sup> <sup><a href="#cite_ref-:0_2-2">2,2</a></sup></span> <span class="reference-text"><span class="citation" style="font-style:normal">«<a rel="nofollow" class="external text" href="https://uni-freiburg.de/university/university-at-a-glance/outstanding-achievements/nobel-prize/georges-koehler/">Georges Köhler</a>» (en anglès).  University of Freiburg. [Consulta: 17 març 2021].</span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"><span class="citation" style="font-style:normal">«<a rel="nofollow" class="external text" href="https://www.aai.org/About/History/Notable-Members/Nobel-Laureates/GeorgesJFKohler">Georges J. F. Köhler, Ph.D.</a>» (en anglès).  The American Association of Immunologists. [Consulta: 17 març 2021].</span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><span class="citation" style="font-style:normal" id="CITEREFAnderssonMelchers1995"><span style="font-variant: small-caps;">Andersson</span>, J.; <span style="font-variant: small-caps;">Melchers</span>, F. «<a rel="nofollow" class="external text" href="https://onlinelibrary.wiley.com/doi/abs/10.1111/j.1365-3083.1995.tb03618.x">Obituary: Georges Köhler 1946-1995</a>» (en anglès). <i>Scandinavian Journal of Immunology</i>, 42, 1, 1995, pàg. 1–2. <a href="/wiki/DOI" title="DOI">DOI</a>: <a rel="nofollow" class="external text" href="https://dx.doi.org/10.1111%2Fj.1365-3083.1995.tb03618.x">10.1111/j.1365-3083.1995.tb03618.x</a>. <a href="/wiki/ISSN" title="ISSN">ISSN</a>: <a rel="nofollow" class="external text" href="//www.worldcat.org/issn/1365-3083">1365-3083</a>.</span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><a href="#cite_ref-5">↑</a></span> <span class="reference-text"><span class="citation book" style="font-style:normal" id="CITEREFEichmann2005"><span style="font-variant: small-caps;">Eichmann</span>, Klaus. «Köhler's death». A: <a rel="nofollow" class="external text" href="https://link.springer.com/book/10.1007/3-7643-7413-6#toc"><i>Köhler's Invention</i></a> (en anglès).  Birkhäuser Basel, 2005, p. 169-172. <span style="font-size:90%;"><a href="/wiki/DOI" title="DOI">DOI</a> <a rel="nofollow" class="external text" href="https://dx.doi.org/10.1007/3-7643-7413-6_15">10.1007/3-7643-7413-6_15</a></span>.</span><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=K%C3%B6hler%27s+Invention&rft.atitle=K%C3%B6hler%27s+death&rft.aulast=Eichmann&rft.aufirst=Klaus&rft.date=2005&rft.pub=Birkh%C3%A4user+Basel&rft.pages=169-172&rft_id=info:doi/10.1007%2F3-7643-7413-6_15&rft_id=https%3A%2F%2Flink.springer.com%2Fbook%2F10.1007%2F3-7643-7413-6%23toc"><span style="display: none;"> </span></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><a href="#cite_ref-6">↑</a></span> <span class="reference-text"><span class="citation" style="font-style:normal" id="CITEREFKöhlerMilstein1975"><span style="font-variant: small-caps;">Köhler</span>, G.; <span style="font-variant: small-caps;">Milstein</span>, C. «<a rel="nofollow" class="external text" href="https://lyneslab.uconn.edu/wp-content/uploads/sites/2599/2020/09/Kohler-and-Milstein-.pdf">Continuous cultures of fused cells secreting antibody of predefined specifity</a>» (pdf) (en anglès). <i>Nature</i>, 256, 1975, pàg. 495-497 [Consulta: 17 març 2021].</span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><a href="#cite_ref-7">↑</a></span> <span class="reference-text"><span class="citation" style="font-style:normal">«<a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/1984/summary/">The Nobel Prize in Physiology or Medicine 1984. Summary</a>» (en anglès). [Consulta: 17 març 2021].</span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><a href="#cite_ref-8">↑</a></span> <span class="reference-text"><span class="citation" style="font-style:normal">«<a rel="nofollow" class="external text" href="https://ssd.jpl.nasa.gov/sbdb.cgi?sstr=11775">11775 Kohler (3224 T-3)</a>» (en anglès).  Jet Propulsion Laboratoy Small-Body Database Browser. [Consulta: 17 març 2021].</span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Enllaços_externs"><span id="Enlla.C3.A7os_externs"></span>Enllaços externs</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Georges_Jean_Franz_K%C3%B6hler&action=edit&section=5" title="Modifica la secció: Enllaços externs"><span>modifica</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><span class="citation" style="font-style:normal">«<a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/430">Georges Jean Franz Köhler</a>» (en anglès). <i>The Nobel Prize</i>.  The Nobel Foundation.</span></li></ul> <div role="navigation" class="navbox" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Control_d%27autoritats" title="Control d'autoritats">Registres d'autoritat</a></th><td class="navbox-list navbox-odd" style="text-align:left;border-left-width:2px;border-left-style:solid;width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a rel="nofollow" class="external text" href="http://www.worldcat.org/identities/lccn-n2005-035031">WorldCat</a></li> <li><a href="/wiki/Gemeinsame_Normdatei" title="Gemeinsame Normdatei">GND</a> <span class="uid"> (<a rel="nofollow" class="external text" href="http://d-nb.info/gnd/124564127">1</a>)</span></li> <li><a href="/wiki/LCCN" class="mw-redirect" title="LCCN">LCCN</a> <span class="uid"> (<a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/names/n2005035031">1</a>)</span></li> <li><a href="/wiki/VIAF" class="mw-redirect" title="VIAF">VIAF</a> <span class="uid"> (<a rel="nofollow" class="external text" href="http://viaf.org/viaf/9254166">1</a>)</span></li> <li><a href="/wiki/ISNI" class="mw-redirect" title="ISNI">ISNI</a> <span class="uid"> (<a rel="nofollow" class="external text" href="http://isni.org/isni/0000000109531594">1</a>)</span></li> <li><a href="/wiki/SUDOC" class="mw-redirect" title="SUDOC">SUDOC</a> <span class="uid"> (<a rel="nofollow" class="external text" href="http://www.idref.fr/09949499X">1</a>)</span></li> <li><a href="/wiki/Biblioteca_Nacional_de_la_Rep%C3%BAblica_Txeca" title="Biblioteca Nacional de la República Txeca">NKC</a> <span class="uid"> (<a rel="nofollow" class="external text" href="http://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=xx0226128&CON_LNG=ENG">1</a>)</span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Bases d'informació</th><td class="navbox-list navbox-even" style="text-align:left;border-left-width:2px;border-left-style:solid;width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/GEC" class="mw-redirect" title="GEC">GEC</a> <span class="uid"> (<a rel="nofollow" class="external text" href="https://www.enciclopedia.cat/gran-enciclopedia-catalana/georges-j.f.-kohler">1</a>)</span></li> <li><a href="/wiki/Diccionari_Hist%C3%B2ric_de_Su%C3%AFssa" title="Diccionari Històric de Suïssa">HDS</a> <span class="uid"> (<a rel="nofollow" class="external text" href="http://www.hls-dhs-dss.ch/textes/f/F046541.php">1</a>)</span></li></ul> </div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; 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