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Köhler</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Ga naar een artikel in een andere taal. Beschikbaar in 47 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-47" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">47 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D9%83%D9%88%D9%87%D9%84%D8%B1" title="جورج كوهلر – Arabisch" lang="ar" hreflang="ar" data-title="جورج كوهلر" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%D9%83%D9%88%D9%87%D9%84%D8%B1" title="جورج كوهلر – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="جورج كوهلر" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%DA%98%D8%B1%DA%98_%D8%AC%DB%8C._%D8%A7%D9%81._%DA%A9%D9%88%D9%87%D9%84%D8%B1" title="ژرژ جی. اف. کوهلر – South Azerbaijani" lang="azb" hreflang="azb" data-title="ژرژ جی. اف. کوهلر" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%93%D0%B5%D0%BE%D1%80%D0%B3_%D0%9A%D1%91%D0%BB%D0%B5%D1%80" title="Георг Кёлер – Belarussisch" lang="be" hreflang="be" data-title="Георг Кёлер" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%93%D0%B5%D0%BE%D1%80%D0%B3_%D0%9A%D1%8C%D0%BE%D0%BB%D0%B5%D1%80" title="Георг Кьолер – Bulgaars" lang="bg" hreflang="bg" data-title="Георг Кьолер" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Georges_Jean_Franz_K%C3%B6hler" title="Georges Jean Franz Köhler – Catalaans" lang="ca" hreflang="ca" data-title="Georges Jean Franz Köhler" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Tsjechisch" lang="cs" hreflang="cs" data-title="Georges Köhler" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Duits" lang="de" hreflang="de" data-title="Georges J. F. Köhler" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Engels" lang="en" hreflang="en" data-title="Georges J. F. Köhler" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Spaans" lang="es" hreflang="es" data-title="Georges Köhler" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Georges_Jean_Franz_K%C3%B6hler" title="Georges Jean Franz Köhler – Estisch" lang="et" hreflang="et" data-title="Georges Jean Franz Köhler" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Baskisch" lang="eu" hreflang="eu" data-title="Georges J. F. Köhler" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DA%98%D8%B1%DA%98_%D8%AC%DB%8C.%D8%A7%D9%81._%DA%A9%D9%88%D9%87%D9%84%D8%B1" title="ژرژ جی.اف. کوهلر – Perzisch" lang="fa" hreflang="fa" data-title="ژرژ جی.اف. کوهلر" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Fins" lang="fi" hreflang="fi" data-title="Georges J. F. Köhler" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Frans" lang="fr" hreflang="fr" data-title="Georges Köhler" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Iers" lang="ga" hreflang="ga" data-title="Georges J. F. Köhler" data-language-autonym="Gaeilge" data-language-local-name="Iers" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%96%27%D7%95%D7%A8%D7%96%27_%D7%A7%D7%9C%D7%A8" title="ז'ורז' קלר – Hebreeuws" lang="he" hreflang="he" data-title="ז'ורז' קלר" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Kroatisch" lang="hr" hreflang="hr" data-title="Georges J. F. Köhler" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Hongaars" lang="hu" hreflang="hu" data-title="Georges J. F. Köhler" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Indonesisch" lang="id" hreflang="id" data-title="Georges Köhler" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Ido" lang="io" hreflang="io" data-title="Georges J. F. Köhler" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Italiaans" lang="it" hreflang="it" data-title="Georges Köhler" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B8%E3%83%A7%E3%83%AB%E3%82%B8%E3%83%A5%E3%83%BBJ%E3%83%BBF%E3%83%BB%E3%82%B1%E3%83%BC%E3%83%A9%E3%83%BC" title="ジョルジュ・J・F・ケーラー – Japans" lang="ja" hreflang="ja" data-title="ジョルジュ・J・F・ケーラー" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%96%D0%BE%D1%80%D0%B6_%D0%9A%D1%91%D0%BB%D0%B5%D1%80" title="Жорж Кёлер – Kazachs" lang="kk" hreflang="kk" data-title="Жорж Кёлер" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EA%B2%8C%EC%98%A4%EB%A5%B4%EA%B2%8C%EC%8A%A4_J._F._%EC%BE%B0%EB%9F%AC" title="게오르게스 J. F. 쾰러 – Koreaans" lang="ko" hreflang="ko" data-title="게오르게스 J. F. 쾰러" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Georgius_J._F._K%C3%B6hler" title="Georgius J. F. Köhler – Latijn" lang="la" hreflang="la" data-title="Georgius J. F. Köhler" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Maleis" lang="ms" hreflang="ms" data-title="Georges J. F. Köhler" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mt mw-list-item"><a href="https://mt.wikipedia.org/wiki/Georges_J.F._Kohler" title="Georges J.F. Kohler – Maltees" lang="mt" hreflang="mt" data-title="Georges J.F. Kohler" data-language-autonym="Malti" data-language-local-name="Maltees" class="interlanguage-link-target"><span>Malti</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%AC%D9%88%D8%B1%D8%AC_%DA%A9%D9%88%D9%87%D9%84%D8%B1" title="جورج کوهلر – Mazanderani" lang="mzn" hreflang="mzn" data-title="جورج کوهلر" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler – Noors - Bokmål" lang="nb" hreflang="nb" data-title="Georges J.F. Köhler" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Occitaans" lang="oc" hreflang="oc" data-title="Georges J. F. Köhler" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Pools" lang="pl" hreflang="pl" data-title="Georges Köhler" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%D8%A7%D8%B1%D8%AC%D8%B2_%DA%A9%D9%88%DB%81%D9%84%D8%B1" title="جارجز کوہلر – Western Punjabi" lang="pnb" hreflang="pnb" data-title="جارجز کوہلر" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Portugees" lang="pt" hreflang="pt" data-title="Georges Köhler" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Roemeens" lang="ro" hreflang="ro" data-title="Georges J. F. Köhler" data-language-autonym="Română" data-language-local-name="Roemeens" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9A%D1%91%D0%BB%D0%B5%D1%80,_%D0%96%D0%BE%D1%80%D0%B6" title="Кёлер, Жорж – Russisch" lang="ru" hreflang="ru" data-title="Кёлер, Жорж" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Simple English" lang="en-simple" hreflang="en-simple" data-title="Georges J. F. Köhler" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler – Zweeds" lang="sv" hreflang="sv" data-title="Georges J.F. Köhler" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Georges_K%C3%B6hler" title="Georges Köhler – Swahili" lang="sw" hreflang="sw" data-title="Georges Köhler" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler – Turks" lang="tr" hreflang="tr" data-title="Georges J. F. 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font-size:120%;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Nobel_prize_medal.svg" class="mw-file-description" title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  Georges J.F. Köhler </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/17_maart" title="17 maart">17 maart</a> <a href="/wiki/1946" title="1946">1946</a> – <a href="/wiki/1_maart" title="1 maart">1 maart</a> <a href="/wiki/1995" title="1995">1995</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Einweihung_des_Max-Planck-Institutes_f%C3%BCr_Immunbiologie_(1990)_(cropped).jpg" class="mw-file-description"><img alt="Köhler (1990)" src="//upload.wikimedia.org/wikipedia/commons/5/55/Einweihung_des_Max-Planck-Institutes_f%C3%BCr_Immunbiologie_%281990%29_%28cropped%29.jpg" decoding="async" width="120" height="210" class="mw-file-element" data-file-width="107" data-file-height="187" /></a></span> </td></tr> <tr> <td style="text-align:center;" colspan="3"><div style="font-size:90%">Köhler (1990)</div> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><a href="/wiki/Duitsland" title="Duitsland">Duitsland</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/M%C3%BCnchen" title="München">München</a> </td></tr> <tr> <th>Overlijdensplaats </th> <td colspan="2"><a href="/wiki/Freiburg_im_Breisgau" title="Freiburg im Breisgau">Freiburg im Breisgau</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2">Fysiologie of Geneeskunde </td></tr> <tr> <th>Jaar </th> <td colspan="2">1984 </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor onderzoek aan het <a href="/wiki/Immuunsysteem" title="Immuunsysteem">immuunsysteem</a> en de productie van <a href="/wiki/Monoklonale_antistof" title="Monoklonale antistof">monoklonale antistoffen</a> </td></tr> <tr> <th>Samen met </th> <td colspan="2"><a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Kaj Jerne</a><br /><a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Stuart Brown</a><br /><a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Joseph Goldstein</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>Georges Jean Franz Köhler</b> (<a href="/wiki/M%C3%BCnchen" title="München">München</a>, <a href="/wiki/17_maart" title="17 maart">17 maart</a> <a href="/wiki/1946" title="1946">1946</a> – <a href="/wiki/Freiburg_im_Breisgau" title="Freiburg im Breisgau">Freiburg im Breisgau</a>, <a href="/wiki/1_maart" title="1 maart">1 maart</a> <a href="/wiki/1995" title="1995">1995</a>) was een Duits <a href="/wiki/Bioloog" class="mw-redirect" title="Bioloog">bioloog</a>. Samen met <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> en <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Kaj Jerne</a> won hij in 1984 de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a> voor onderzoek aan het <a href="/wiki/Immuunsysteem" title="Immuunsysteem">immuunsysteem</a> en de productie van <a href="/wiki/Monoklonale_antistof" title="Monoklonale antistof">monoklonale antistoffen</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Georges_J.F._K%C3%B6hler&veaction=edit&section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Georges_J.F._K%C3%B6hler&action=edit&section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Köhler genoot zijn wetenschappelijke opleiding aan de <a href="/wiki/Universiteit_van_Freiburg" title="Universiteit van Freiburg">Universiteit van Freiburg</a> waar hij in 1974 promoveerde in de biologie. Aansluitend ging hij naar Cambridge om als postdoctoraal student onderzoek te doen onder <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> en waar ze samen de <a href="/w/index.php?title=Hybridoma&action=edit&redlink=1" class="new" title="Hybridoma (de pagina bestaat niet)">hybridomatechniek</a> ontwikkelden.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Met deze techniek lieten ze een <a href="/wiki/Antilichaam" class="mw-redirect" title="Antilichaam">antilichaam</a> producerende <a href="/wiki/B-cel" title="B-cel">B-cel</a> fuseren met een ontspoorde <a href="/w/index.php?title=Myeloomcel&action=edit&redlink=1" class="new" title="Myeloomcel (de pagina bestaat niet)">myeloomcel</a>. Deze samengesmolten cellen (hybridomacellen) waren nu in staat antilichamen te maken van een specifieke soort, namelijk die van de oorspronkelijk B-cel. </p><p>Deze techniek was essentieel om identieke (monoklonale) antilichamen te produceren waarbij slechts één kloon van plasmacellen tot de productie van antistoffen wordt aangezet. Hun doorbraak wordt beschouwd als een van de belangrijkste technieken binnen de biotechnologie en leidde tot een snelle ontwikkeling van verscheidene medicijnen voor de behandeling van onder andere kanker, leukemie en aids. Een deel van dit onderzoek voerde hij uit aan het <a href="/w/index.php?title=Basel_Institut_f%C3%BCr_Immunologie&action=edit&redlink=1" class="new" title="Basel Institut für Immunologie (de pagina bestaat niet)">Basel Institut für Immunologie</a>. </p><p>In 1984 werd Köhler directeur van het <a href="/wiki/Max-Planck-Gesellschaft" title="Max-Planck-Gesellschaft">Max Planck Institute for Immunobiology</a>, waar hij tot zijn dood bleef werken. Hij stierf aan de gevolgen van een <a href="/wiki/Longontsteking" title="Longontsteking">longontsteking</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Externe_link">Externe link</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Georges_J.F._K%C3%B6hler&veaction=edit&section=2" title="Bewerk dit kopje: Externe link" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Georges_J.F._K%C3%B6hler&action=edit&section=2" title="De broncode bewerken van de sectie: Externe link"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><a rel="nofollow" class="external text" href="https://www.nobelprize.org/prizes/medicine/1984/summary/">The Nobel Prize in Physiology or Medicine 1984</a>. NobelPrize.org. Nobel Media AB 2018.</li></ul> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li><span class="citation" id="CITEREFSchmeck_Jr.Fout:_ongeldige_tijd."><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><span style="font-variant:small-caps;">Schmeck Jr., Harold M</span>, <a rel="nofollow" class="external text" href="http://www.nytimes.com/1984/10/16/science/3-immunology-investigators-win-nobel-prize-in-medicine.html">3 Immunology Investigators Win Nobel Prize in Medicine</a>. <a href="/wiki/The_New_York_Times" title="The New York Times">The New York Times</a> (<span class="mw-formatted-date" title="1984-10-16">16 oktober 1984</span>). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150524154041/http://www.nytimes.com/1984/10/16/science/3-immunology-investigators-win-nobel-prize-in-medicine.html">Gearchiveerd</a> op <span class="mw-formatted-date" title="2015-05-24">24 mei 2015</span>. Geraadpleegd op <span class="mw-formatted-date" title="2015-06-15">15 juni 2015</span>. </span></li> <li><span class="citation" id="CITEREFAltmanFout:_ongeldige_tijd."><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><span style="font-variant:small-caps;">Altman, Lawrence K</span>, <a rel="nofollow" class="external text" href="http://www.nytimes.com/1984/10/16/science/a-discovery-and-its-impact-nine-years-of-excitement.html">A Discovery and its Impact: Nine Years of Excitement</a>. <a href="/wiki/The_New_York_Times" title="The New York Times">The New York Times</a> (<span class="mw-formatted-date" title="1984-10-16">16 oktober 1984</span>). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150524154041/http://www.nytimes.com/1984/10/16/science/a-discovery-and-its-impact-nine-years-of-excitement.html">Gearchiveerd</a> op <span class="mw-formatted-date" title="2015-05-24">24 mei 2015</span>. Geraadpleegd op <span class="mw-formatted-date" title="2015-06-15">15 juni 2015</span>. </span></li> <li><span class="citation" id="CITEREFWadeFout:_ongeldige_tijd."><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><span style="font-variant:small-caps;">Wade, Nicholas</span>, <a rel="nofollow" class="external text" href="http://www.nytimes.com/1995/03/04/obituaries/georges-kohler-48-medicine-nobel-winner.html">Georges Kohler, 48, Medicine Nobel Winner</a>. <a href="/wiki/The_New_York_Times" title="The New York Times">The New York Times</a> (<span class="mw-formatted-date" title="1995-03-04">4 maart 1995</span>). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150222164135/http://www.nytimes.com/1995/03/04/obituaries/georges-kohler-48-medicine-nobel-winner.html">Gearchiveerd</a> op <span class="mw-formatted-date" title="2015-02-22">22 februari 2015</span>. Geraadpleegd op <span class="mw-formatted-date" title="2015-02-22">22 februari 2015</span>. </span></li></ul> <div class="reflist" style="list-style-type: decimal;"><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREFG._Köhler,_C._Milstein1975"><span style="font-variant:small-caps;">G. Köhler, C. Milstein</span> (1975). <cite>Continuous cultures of fused cells secreting antibody of predefined specificity</cite>. <i>Nature</i> <b>256</b>: 495-497. <a href="/wiki/Digital_object_identifier" title="Digital object identifier">DOI</a>: <a rel="nofollow" class="external text" href="https://doi.org/10.1038/256495a0">10.1038/256495a0</a>. </span></span> </li> </ol></div></div> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output .navigatie-afb-rechts{position:absolute}.mw-parser-output .navigatie-afb-rechts{right:2px}.mw-parser-output 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href="/wiki/Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Sjabloon:Navigatie winnaars van de Nobelprijs voor Fysiologie of Geneeskunde"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/16px-OOjs_UI_icon_article-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/24px-OOjs_UI_icon_article-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/32px-OOjs_UI_icon_article-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> · <span class="ui-state-disabled" typeof="mw:File"><span title="Overleg sjabloon (de pagina bestaat niet)"><img alt="Overleg sjabloon (de pagina bestaat niet)" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/16px-OOjs_UI_icon_speechBubbles-ltr.svg.png" decoding="async" 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title="Santiago Ramón y Cajal">Ramón y Cajal</a> · 1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a class="mw-selflink selflink">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Overgenomen van "<a dir="ltr" href="https://nl.wikipedia.org/w/index.php?title=Georges_J.F._Köhler&oldid=64702902">https://nl.wikipedia.org/w/index.php?title=Georges_J.F._Köhler&oldid=64702902</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Categorie:Alles" title="Categorie:Alles">Categorieën</a>: <ul><li><a href="/wiki/Categorie:Duits_bioloog" title="Categorie:Duits bioloog">Duits bioloog</a></li><li><a href="/wiki/Categorie:Winnaar_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Categorie:Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde">Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde</a></li><li><a href="/wiki/Categorie:20e-eeuws_medicus" title="Categorie:20e-eeuws medicus">20e-eeuws medicus</a></li><li><a href="/wiki/Categorie:20e-eeuws_bioloog" title="Categorie:20e-eeuws bioloog">20e-eeuws bioloog</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> Deze pagina is voor het laatst bewerkt op 14 jul 2023 om 21:30.</li> <li id="footer-info-copyright">De tekst is beschikbaar onder de licentie <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/deed.nl">Creative Commons Naamsvermelding/Gelijk delen</a>, er kunnen aanvullende voorwaarden van toepassing zijn. 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