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js-swp-download-button" data-signup-modal="{"location":"swp-splash-paper-cover","attachmentId":89458184,"attachmentType":"pdf"}"><img alt="First page of “Retinotopy and color sensitivity in human visual cortical area V8”" class="ds-work-cover--cover-thumbnail" src="https://0.academia-photos.com/attachment_thumbnails/89458184/mini_magick20220810-1-1j6p3j3.png?1660152113" /><img alt="PDF Icon" class="ds-work-cover--file-icon" src="//a.academia-assets.com/assets/single_work_splash/adobe.icon-574afd46eb6b03a77a153a647fb47e30546f9215c0ee6a25df597a779717f9ef.svg" /><div class="ds-work-cover--hover-container"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">download</span><p>Download Free PDF</p></div><div class="ds-work-cover--ribbon-container">Download Free PDF</div><div class="ds-work-cover--ribbon-triangle"></div></button></div></div></div><div class="ds-work-card--work-information"><h1 class="ds-work-card--work-title">Retinotopy and color sensitivity in human visual cortical area V8</h1><div class="ds-work-card--work-authors ds-work-card--detail"><a class="ds-work-card--author js-wsj-grid-card-author ds2-5-body-md ds2-5-body-link" data-author-id="36412689" href="https://independent.academia.edu/RTootell"><img alt="Profile image of R. Tootell" class="ds-work-card--author-avatar" src="//a.academia-assets.com/images/s65_no_pic.png" />R. Tootell</a></div><p class="ds-work-card--detail ds2-5-body-sm">1998, Nature Neuroscience</p><div class="ds-work-card--button-container"><button class="ds2-5-button js-swp-download-button" data-signup-modal="{"location":"continue-reading-button--work-card","attachmentId":89458184,"attachmentType":"pdf","workUrl":"https://www.academia.edu/84429963/Retinotopy_and_color_sensitivity_in_human_visual_cortical_area_V8"}">See full PDF</button><button class="ds2-5-button ds2-5-button--secondary js-swp-download-button" data-signup-modal="{"location":"download-pdf-button--work-card","attachmentId":89458184,"attachmentType":"pdf","workUrl":"https://www.academia.edu/84429963/Retinotopy_and_color_sensitivity_in_human_visual_cortical_area_V8"}"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">download</span>Download PDF</button></div></div></div></div><div data-auto_select="false" data-client_id="331998490334-rsn3chp12mbkiqhl6e7lu2q0mlbu0f1b" data-doc_id="89458184" data-landing_url="https://www.academia.edu/84429963/Retinotopy_and_color_sensitivity_in_human_visual_cortical_area_V8" data-login_uri="https://www.academia.edu/registrations/google_one_tap" data-moment_callback="onGoogleOneTapEvent" id="g_id_onload"></div><div class="ds-top-related-works--grid-container"><div class="ds-related-content--container ds-top-related-works--container"><h2 class="ds-related-content--heading">Related papers</h2><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="0" data-entity-id="56817585" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/56817585/The_architecture_of_the_colour_centre_in_the_human_visual_brain_new_results_and_a_review">The architecture of the colour centre in the human visual brain: new results and a review</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="35004922" href="https://independent.academia.edu/SemirZeki">Semir Zeki</a></div><p class="ds-related-work--metadata ds2-5-body-xs">European Journal of Neuroscience, 2000</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"The architecture of the colour centre in the human visual brain: new results and a review","attachmentId":72017491,"attachmentType":"pdf","work_url":"https://www.academia.edu/56817585/The_architecture_of_the_colour_centre_in_the_human_visual_brain_new_results_and_a_review","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/56817585/The_architecture_of_the_colour_centre_in_the_human_visual_brain_new_results_and_a_review"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="1" data-entity-id="12638649" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/12638649/Color_responsiveness_argues_against_a_dorsal_component_of_human_V4">Color responsiveness argues against a dorsal component of human V4.</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="31611829" href="https://optusnet.academia.edu/ErinGoddard">Erin Goddard</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Journal of Vision, 2011</p><p class="ds-related-work--abstract ds2-5-body-sm">The retinotopic organization, position, and functional responsiveness of some early visual cortical areas in human and non-human primates are consistent with their being homologous structures. The organization of other areas remains controversial. A critical debate concerns the potential human homologue of macaque area V4, an area very responsive to colored images: specifically, whether human V4 is divided between ventral and dorsal components, as in the macaque, or whether human V4 is confined to one ventral area. We used fMRI to define these areas retinotopically in human and to test the impact of image color on their responsivity. We found a robust preference for full-color movie segments over a luminance-matched achromatic version in ventral V4 but little or no preference in the vicinity of the putative dorsal counterpart. Contrary to previous reports that visual field coverage in the ventral part of V4 is deficient without the dorsal part, we found that coverage in ventral V4 extended to the lower vertical meridian, including the entire contralateral hemifield. Together these results provide evidence against a dorsal component of human V4. Instead, they are consistent with human V4 being a single, ventral region that is sensitive to the chromatic components of images.</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Color responsiveness argues against a dorsal component of human V4.","attachmentId":37755969,"attachmentType":"pdf","work_url":"https://www.academia.edu/12638649/Color_responsiveness_argues_against_a_dorsal_component_of_human_V4","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/12638649/Color_responsiveness_argues_against_a_dorsal_component_of_human_V4"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="2" data-entity-id="16521942" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/16521942/Functional_measurements_of_human_ventral_occipital_cortex_retinotopy_and_colour">Functional measurements of human ventral occipital cortex: retinotopy and colour</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="35770872" href="https://uci.academia.edu/AlyssaBrewer">Alyssa Brewer</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Philosophical transactions of the Royal Society of London. Series B, Biological sciences, 2002</p><p class="ds-related-work--abstract ds2-5-body-sm">Human colour vision originates in the cone photoreceptors, whose spatial density peaks in the fovea and declines rapidly into the periphery. For this reason, one expects to find a large representation of the cone-rich fovea in those cortical locations that support colour perception. Human occipital cortex contains several distinct foveal representations including at least two that extend onto the ventral surface: a region thought to be critical for colour vision. To learn more about these ventral signals, we used functional magnetic resonance imaging to identify visual field maps and colour responsivity on the ventral surface. We found a visual map of the complete contralateral hemifield in a 4 cm(2) region adjacent to ventral V3; the foveal representation of this map is confluent with that of areas V1/2/3. Additionally, a distinct foveal representation is present on the ventral surface situated 3-5 cm anterior from the confluent V1/2/3 foveal representations. This organization is n...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Functional measurements of human ventral occipital cortex: retinotopy and colour","attachmentId":42448934,"attachmentType":"pdf","work_url":"https://www.academia.edu/16521942/Functional_measurements_of_human_ventral_occipital_cortex_retinotopy_and_colour","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/16521942/Functional_measurements_of_human_ventral_occipital_cortex_retinotopy_and_colour"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="3" data-entity-id="102666735" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/102666735/Cortical_area_V4_and_its_role_in_the_perception_of_color">Cortical area V4 and its role in the perception of color</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="40309924" href="https://independent.academia.edu/heywoodcharles">charles heywood</a></div><p class="ds-related-work--metadata ds2-5-body-xs">The Journal of Neuroscience, 1992</p><p class="ds-related-work--abstract ds2-5-body-sm">The color and lightness vision of three monkeys with bilateral removal of cortical area V4 and three unoperated controls were tested by measuring their ability to discriminate between two rows of colored or gray stimuli. In one row, the stimuli were ordered in terms of either chromaticity or luminance, whereas in the other row they were disordered. Their ability to select the odd-one-out in an array of colors or grays and to select the colored patch from an array of achromatic grays was also assessed. Unlike an achromatopsic patient tested previously in an identical fashion, monkeys with V4 lesions performed indistinguishably from controls in the oddity test. The animals lacking V4 were slightly impaired at discriminating between ordered and disordered arrays of colors or grays, but the color impairment was no more severe than the impairment with grays. These deficits were readily accounted for in terms of the conspicuous deficits in pattern discrimination apparent in a nine-choice ...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Cortical area V4 and its role in the perception of color","attachmentId":102876535,"attachmentType":"pdf","work_url":"https://www.academia.edu/102666735/Cortical_area_V4_and_its_role_in_the_perception_of_color","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/102666735/Cortical_area_V4_and_its_role_in_the_perception_of_color"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="4" data-entity-id="70019638" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/70019638/fMRI_measurements_of_color_in_macaque_and_human">fMRI measurements of color in macaque and human</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="9241526" href="https://independent.academia.edu/BrianWandell">Brian Wandell</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Journal of vision, 2008</p><p class="ds-related-work--abstract ds2-5-body-sm">We have used fMRI to measure responses to chromatic and achromatic contrast in retinotopically defined regions of macaque and human visual cortex. We make four observations. Firstly, the relative amplitudes of responses to color and luminance stimuli in macaque area V1 are similar to those previously observed in human fMRI experiments. Secondly, the dorsal and ventral subdivisions of macaque area V4 respond in a similar way to opponent (L--M)-cone chromatic contrast suggesting that they are part of a single functional area. Thirdly, we find that macaque area V4, like area V1, responds preferentially to chromatic contrast compared to luminance contrast and the degree of preference is strongly influenced by the temporal frequency of the stimulus. Finally, we observe that while macaque V4d is a region on the dorsal surface of the macaque visual cortex that responds robustly to chromatic stimuli, human chromatic responses to identical stimuli are largely confined to the ventral surface ...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"fMRI measurements of color in macaque and human","attachmentId":79905247,"attachmentType":"pdf","work_url":"https://www.academia.edu/70019638/fMRI_measurements_of_color_in_macaque_and_human","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/70019638/fMRI_measurements_of_color_in_macaque_and_human"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="5" data-entity-id="12885901" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/12885901/Where_is_Dorsal_V4_in_Human_Visual_Cortex_Retinotopic_Topographic_and_Functional_Evidence">Where is 'Dorsal V4' in Human Visual Cortex? Retinotopic, Topographic and Functional Evidence</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="32026617" href="https://independent.academia.edu/NHadjikhani">Nouchine Hadjikhani</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Cerebral Cortex, 2001</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Where is 'Dorsal V4' in Human Visual Cortex? Retinotopic, Topographic and Functional Evidence","attachmentId":45866761,"attachmentType":"pdf","work_url":"https://www.academia.edu/12885901/Where_is_Dorsal_V4_in_Human_Visual_Cortex_Retinotopic_Topographic_and_Functional_Evidence","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/12885901/Where_is_Dorsal_V4_in_Human_Visual_Cortex_Retinotopic_Topographic_and_Functional_Evidence"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="6" data-entity-id="56916866" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/56916866/Color_Biased_Regions_of_the_Ventral_Visual_Pathway_Lie_between_Face_and_Place_Selective_Regions_in_Humans_as_in_Macaques">Color-Biased Regions of the Ventral Visual Pathway Lie between Face- and Place-Selective Regions in Humans, as in Macaques</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="20840822" href="https://wellesley.academia.edu/RosaLaferSousa">Rosa Lafer-Sousa</a></div><p class="ds-related-work--metadata ds2-5-body-xs">The Journal of neuroscience : the official journal of the Society for Neuroscience, 2016</p><p class="ds-related-work--abstract ds2-5-body-sm">The existence of color-processing regions in the human ventral visual pathway (VVP) has long been known from patient and imaging studies, but their location in the cortex relative to other regions, their selectivity for color compared with other properties (shape and object category), and their relationship to color-processing regions found in nonhuman primates remain unclear. We addressed these questions by scanning 13 subjects with fMRI while they viewed two versions of movie clips (colored, achromatic) of five different object classes (faces, scenes, bodies, objects, scrambled objects). We identified regions in each subject that were selective for color, faces, places, and object shape, and measured responses within these regions to the 10 conditions in independently acquired data. We report two key findings. First, the three previously reported color-biased regions (located within a band running posterior-anterior along the VVP, present in most of our subjects) were sandwiched b...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Color-Biased Regions of the Ventral Visual Pathway Lie between Face- and Place-Selective Regions in Humans, as in Macaques","attachmentId":72068111,"attachmentType":"pdf","work_url":"https://www.academia.edu/56916866/Color_Biased_Regions_of_the_Ventral_Visual_Pathway_Lie_between_Face_and_Place_Selective_Regions_in_Humans_as_in_Macaques","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/56916866/Color_Biased_Regions_of_the_Ventral_Visual_Pathway_Lie_between_Face_and_Place_Selective_Regions_in_Humans_as_in_Macaques"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="7" data-entity-id="20594718" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/20594718/Mapping_of_cortical_areas_involved_in_color_vision_in_non_human_primates">Mapping of cortical areas involved in color vision in non-human primates</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="7847439" href="https://nips.academia.edu/NorihiroSadato">Norihiro Sadato</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Neuroscience Letters, 1997</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Mapping of cortical areas involved in color vision in non-human primates","attachmentId":41455938,"attachmentType":"pdf","work_url":"https://www.academia.edu/20594718/Mapping_of_cortical_areas_involved_in_color_vision_in_non_human_primates","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/20594718/Mapping_of_cortical_areas_involved_in_color_vision_in_non_human_primates"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="8" data-entity-id="10060308" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/10060308/Colour_tuning_in_human_visual_cortex_measured_with_functional_magnetic_resonance_imaging">Colour tuning in human visual cortex measured with functional magnetic resonance imaging</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="24493290" href="https://independent.academia.edu/StephenEngel">Stephen Engel</a></div><p class="ds-related-work--metadata ds2-5-body-xs">Nature, 1997</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"Colour tuning in human visual cortex measured with functional magnetic resonance imaging","attachmentId":36187991,"attachmentType":"pdf","work_url":"https://www.academia.edu/10060308/Colour_tuning_in_human_visual_cortex_measured_with_functional_magnetic_resonance_imaging","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/10060308/Colour_tuning_in_human_visual_cortex_measured_with_functional_magnetic_resonance_imaging"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div><div class="ds-related-work--container js-wsj-grid-card" data-collection-position="9" data-entity-id="102666736" data-sort-order="default"><a class="ds-related-work--title js-wsj-grid-card-title ds2-5-body-md ds2-5-body-link" href="https://www.academia.edu/102666736/On_the_role_of_cortical_area_V4_in_the_discrimination_of_hue_and_pattern_in_macaque_monkeys">On the role of cortical area V4 in the discrimination of hue and pattern in macaque monkeys</a><div class="ds-related-work--metadata"><a class="js-wsj-grid-card-author ds2-5-body-sm ds2-5-body-link" data-author-id="40309924" href="https://independent.academia.edu/heywoodcharles">charles heywood</a></div><p class="ds-related-work--metadata ds2-5-body-xs">The Journal of Neuroscience, 1987</p><p class="ds-related-work--abstract ds2-5-body-sm">Cortical visual area V4 in macaque monkeys has a large proportion of neurons that are sensitive to the wavelength or to the color of light. We tested its role in hue discrimination by removing it in macaque monkeys trained to discriminate small differences in hue. Hue discrimination thresholds were permanently elevated in 4 macaque monkeys in which V4 was removed bilaterally. In contrast, there was no impairment in achromatic intensity thresholds tested in an identical manner. However, the discrimination of pattern and orientation was also conspicuously impaired, indicating that area V4 is not concerned solely with processing information about wavelength. The multiple defect is consistent with evidence that V4 provides the major cortical visual input to the temporal lobe, where a large range of visual properties is registered. The performance of monkeys with V4 ablation was compared with that of unoperated control monkeys and monkeys with removal of cortex in the banks and floor of ...</p><div class="ds-related-work--ctas"><button class="ds2-5-text-link ds2-5-text-link--inline js-swp-download-button" data-signup-modal="{"location":"wsj-grid-card-download-pdf-modal","work_title":"On the role of cortical area V4 in the discrimination of hue and pattern in macaque monkeys","attachmentId":102876537,"attachmentType":"pdf","work_url":"https://www.academia.edu/102666736/On_the_role_of_cortical_area_V4_in_the_discrimination_of_hue_and_pattern_in_macaque_monkeys","alternativeTracking":true}"><span class="material-symbols-outlined" style="font-size: 18px" translate="no">download</span><span class="ds2-5-text-link__content">Download free PDF</span></button><a class="ds2-5-text-link ds2-5-text-link--inline js-wsj-grid-card-view-pdf" href="https://www.academia.edu/102666736/On_the_role_of_cortical_area_V4_in_the_discrimination_of_hue_and_pattern_in_macaque_monkeys"><span class="ds2-5-text-link__content">View PDF</span><span class="material-symbols-outlined" style="font-size: 18px" translate="no">chevron_right</span></a></div></div></div></div><div class="ds-sticky-ctas--wrapper js-loswp-sticky-ctas hidden"><div class="ds-sticky-ctas--grid-container"><div class="ds-sticky-ctas--container"><button class="ds2-5-button js-swp-download-button" data-signup-modal="{"location":"continue-reading-button--sticky-ctas","attachmentId":89458184,"attachmentType":"pdf","workUrl":null}">See full PDF</button><button class="ds2-5-button ds2-5-button--secondary js-swp-download-button" data-signup-modal="{"location":"download-pdf-button--sticky-ctas","attachmentId":89458184,"attachmentType":"pdf","workUrl":null}"><span class="material-symbols-outlined" style="font-size: 20px" translate="no">download</span>Download PDF</button></div></div></div><div class="ds-below-fold--grid-container"><div class="ds-work--container js-loswp-embedded-document"><div class="attachment_preview" data-attachment="Attachment_89458184" style="display: none"><div class="js-scribd-document-container"><div class="scribd--document-loading js-scribd-document-loader" style="display: block;"><img alt="Loading..." src="//a.academia-assets.com/images/loaders/paper-load.gif" /><p>Loading Preview</p></div></div><div style="text-align: center;"><div class="scribd--no-preview-alert js-preview-unavailable"><p>Sorry, preview is currently unavailable. 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