CINXE.COM
Rosaceae in Flora of Pakistan @ efloras.org
<!DOCTYPE HTML PUBLIC "-//W3C//DTD HTML 4.0 Transitional//EN" > <HTML> <!--%@ OutputCache Duration=60 VaryByParam="flora_id;taxon_id" %--> <head> <!-- Google Tag Manager --> <script>(function(w,d,s,l,i){w[l]=w[l]||[];w[l].push({'gtm.start': new Date().getTime(),event:'gtm.js'});var f=d.getElementsByTagName(s)[0], j=d.createElement(s),dl=l!='dataLayer'?'&l='+l:'';j.async=true;j.src= 'https://www.googletagmanager.com/gtm.js?id='+i+dl;f.parentNode.insertBefore(j,f); })(window,document,'script','dataLayer','GTM-TGK4SMV');</script> <!-- End Google Tag Manager --> <title>Rosaceae in Flora of Pakistan @ efloras.org</title> <link rel="stylesheet" type="text/css" href='/templates/eflora.css' /> <script src="JavaScript/eflora.js" type="text/javascript" ></script> </head> <!-- body onload="formMOMap.submit()" --> <body> <TABLE id="TableMain" cellSpacing="1" cellPadding="1" width="640" border="0"> <TR> <TD><!--%@ Register TagPrefix="ucFlora" TagName="TaxonSearchBox" Src="TaxonSearchBox.ascx" %--> <script language="javascript"> function jsClearTextBox(sender) { // alert(formSearchBox["name_str"].id); // alert(sender.value); if (sender.value == "taxon name") { sender.value = ""; } } </script> <table id="ucEfloraHeader_tableHeaderWrapper" cellspacing="0" cellpadding="0" width="100%" border="0"> <tr> <TD style="WIDTH: 479px"><a id="ucEfloraHeader_linkFloraHome" title="Flora Homepage" href="flora_page.aspx?flora_id=5"><img src="images/flora/eflora-logo-5.gif" id="ucEfloraHeader_imageLogo" border="0" alt="Link to Flora of Pakistan home" /><br> </a></TD> <TD valign="bottom" align="right"> <div id="ucEfloraHeader_panelSearchBox"> <!-- begin of search box --> <FORM name="formSearchBox" action="browse.aspx" method="get"> <TABLE borderColor="#ccffcc" cellSpacing="0" cellPadding="2" width="200" border="1"> <TR> <TD> <span id="ucEfloraHeader_lblSearchBox"><INPUT TYPE='Hidden' NAME='flora_id' ' id='flora_id' VALUE='5' > <input id=name_str name=name_str value='' class=small title='Use % as the wildcard (e.g. pinus % or pinus a%)' onfocus='jsClearTextBox(this);' accessKey=s size=15> <input class=small type=submit value=Search name=btnSearch><br> <input name=chkAllFloras type=checkbox > <span class=small>All Floras <A HREF='search_page.aspx?flora_id=5' TARGET='_top' class='a.link.small'>Advanced Search</A></span> </span></TD> </TR> </TABLE> </FORM> <!-- end of search box --> </div> </TD> </tr> </table> </TD> </TR> <TR> <TD id="tdSubHeader"> <table id="FooterTableOut" width="100%" border="0" cellpadding="0" cellspacing="0"> <tr> <td bgcolor="#999999"><img src="~/images/layout/dot.gif" alt="" width="100%" height="1"></td> </tr> <tr> <TD> <table bgcolor="#ffffcc" id="footerTable" width="100%" border="0" cellpadding="2" cellspacing="0"> <tr> <td style="WIDTH: 291px"> Page 1</td> <TD align="right"> <span id="ucEfloraSubHeader_lblLinks"><a target='_top' href='/login.aspx'>Login</a> | <A HREF='/index.aspx' TARGET='_top' >eFloras Home</A> | <A HREF='/help_page.aspx' TARGET='_top' >Help</A></span></TD> </tr> </table> </TD> </tr> <tr> <td bgcolor="#999999"><img src="~/images/layout/dot.gif" alt="" width="100%" height="1"></td> </tr> </table> </TD> </TR> <TR> <TD id="tdBody"> <table id="tableBody" cellSpacing="0" cellPadding="0" width="100%" border="0"> <tr> <td id="tdLeftBar" valign="top" width="0"></td> <td id="tdContent" vAlign="top" width="1930"> <table width="100%" border="0"> <tr> <td vAlign="top"> <table id="tableTaxonChain" cellspacing="0" cellpadding="1" width="100%" border="0"> <tr> <td><span id="lblTaxonChain" style="font-size:9pt;"><A HREF='flora_page.aspx?flora_id=5' TARGET='_top' TITLE='Flora homepage' >Pakistan</A> | <A HREF='browse.aspx?flora_id=5' TARGET='_top' >Family List</A></span></td> </tr> </table> <P></P> <div id="panelTaxonTreatment"> <span id="lblTaxonDesc"><!-- Default Template --> <b>Rosaceae</b> <P> Sven Landrein, Renata Borosova and Joanna Osborne <script> function hide_DivAuthorInst() {DivAuthorInst.style.display='none' ;} </script> <a href='#' onMouseOver="DivAuthorInst.style.display='block';" onMouseOut="setTimeout('hide_DivAuthorInst()', 15000); " ><IMG SRC='images/layout/icon-info.gif' border='0' > </a><div id='DivAuthorInst' style="display='none'" class="divInfo"> <hr size=1>The Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, U.K.</div> <P> <Image:Begin> <TABLE cellSpacing="0" cellPadding="4" width="200" align="right" bgColor="#ffffcc" border="1" > <TR> <TD> <A HREF='object_page.aspx?object_id=118447&flora_id=5' ><IMG SRC='http://www.mobot.org/mobot/PakistanImages/216-Rosaceae/Potentilla_anserina.jpg' width=200 border=0 > </A><A HREF='florataxon.aspx?flora_id=5&taxon_id=200011027' TARGET='_top' TITLE='Treatment' >Potentilla anserina</A><br> Illustration <br> </TD> </TR> </TABLE> <Image:End> <P> <p> <p> Evergreen or deciduous trees, shrubs, sarmentous shrubs or perennial herbs. Leaves mostly alternate, simple, imparipinnate, interruptedly imparipinnate, trifoliolate, ternate or palmately to pinnately compound, stipules deciduous or persistent (absent in Spiraea and Aruncus). Glands often present on leaves in the genus Prunus. Stems sometimes spiny or bearing prickles. Flowers bisexual, rarely unisexual (plants dioecious in Sorbaria), actinomorphic, (4-)5-merous, solitary, in racemes or compound racemes. Hypanthium usually very conspicuous, campanulate with more or less obvious nectary disc on its rim. Disc lobed in some genera. Sepals (4-)5, imbricate, mostly borne on hypanthium, epicalyx often present outside calyx, petals (4-)5, imbricate, free, rarely 0. Stamens 4 to numerous, anthers 2 or 1-locular, dehiscing longitudinally, intrastaminal. Ovary superior to inferior, carpels 1 to many, free to variously connate, with mostly two ovules per carpel or locule, stylodia 1 per carpel, stigmas capitate (penicillate in Sanguisorba). Fruits diverse, drupe, drupetum, nuculanium, pome, pometum, follicetum, achenetum or glandetum. Seeds 1 to several in each carpel or locule, usually without endosperm.<p> A family consisting of 85 genera and about 3000 species, including many apomictic lines (Mabberley, Plant Book, 3rd. Ed. 2008), cosmopolitan, represented by 27 genera and about 160 species in Pakistan.<p> <p> <b>Infrafamilial classification</b> (Kalkman, Rosaceae in Kubitzki, Fam. Gen.Vasc. Pl. 6: 343-386. 2004).<p>Tribe Spiraeeae: <b>Spiraea, Aruncus</b><p>Tribe Gillenieae: <b>Sorbaria</b><p> Tribe Kerrieae: <b>Kerria</b><p>Tribe Ulmariaea: <b>Filipendula</b><p> Tribe Sanguisorbeae: <b>Agrimonia, Sanguisorba</b><p>Tribe Potentilleae: <b>Chamaerhodos, Potentilla, Sibbaldia, Duchesnea, Fragaria</b><p>Geum Group: <b>Geum, Acomastylis</b><p>Tribe Rubeae: <b>Rubus</b><p>Tribe Roseae: <b>Rosa</b><p>Alchemilla Group: <b>Alchemilla</b><p>Tribe Pruneae: <b>Prunus, Prinsepia</b><p>Cydonia Group: <b>Cydonia</b><p>Tribe Maleae: <b>Sorbus, Eriobotrya, Malus, Pyrus</b><p>Tribe Crataegeae: <b>Cotoneaster, Pyracantha, Crataegus</b><p>Note: The conventional groups in Rosaceae defined by Focke (subfam, Spiraoideae, Maloideae, Roisoideae and Prunoideae) are still much used but the phylogeny has not been worked out sufficiently and some groups are still unplaced in either Spiraeoideae, Maloideae or Rosoideae, Prunoideae. <p> <p> <b>Acknowledgements:</b> We are grateful to the Directors/ Curators of the following herbaria for herbarium and library facilities: A, B, BM, CGE, E, G, GH, KOR, KUH, KYO, LE, LINN, MICH, NA, PE, RAW, RNG, US, UZ, Z. We thankfully acknowledge financial support for this publication from the Andrew Mellon Foundation and National Science Foundation (DEB 0103783), Washington, U.S.A. obtained through the Missouri Botanical Garden, St. Louis, U.S.A. We are grateful to Professor P. H. Raven of the Missouri Botanical Garden, St. Louis for his assistance in securing the support. Grateful thanks are due to Prof. Dr. Pirzada Qasim Raza Siddiqui, Vice-Chancellor, University of Karachi and Professor Anjum Parveen, Director, Institute of Plant Conservation for providing working facilities to the project and for their understanding and encouragement. It is a matter of pleasure to thank Dr. Shahina A. Ghazanfar (Kew) for kindly helping us from time to time.<p> <p> <p> <P> </span> <P><!-- Key --> <div id="panelKey"> <BR clear="all"> <table id="tableKey" cellspacing="0" cellpadding="4" width="100%" border="0"> <tr bgcolor="#d5ffff"> <TD width="347"> <P> <span id="lblKeyTitle" style="font-size:9pt;font-weight:bold;"></span></P> </TD> <TD nowrap="nowrap" align="right"> <P> <span id="lblKeyEditLink" style="font-size:8pt;"></span></P> </TD> </tr> <tr bgcolor="#f1ffff"> <TD colspan="2"> <P> <span id="lblKeyDesc" style="font-size:9pt;"></span></P> <P> <span id="lblKey"><TABLE WIDTH='100%' BORDER='0' > <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-1'>1</A></td> <td WIDTH=400 VALIGN=TOP>Carpels usually 5, free (if connate, ovary is superior), fruit a follicetum.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-2'>(2)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Carpels 1 to many, free to connate (if connate, ovary is inferior), fruit not a follicetum.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-4'>(4)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-2'>2</A> <A HREF='#KEY-1-1'>(1)</A></td> <td WIDTH=400 VALIGN=TOP>Leaves ternate or imparipinnate, disc on rim of the hypanthium slightly lobed or not, carpels connate or not.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-3'>(3)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Leaves simple, disc on rim of the hypanthium lobed, carpels free.</td> <td > </td> <td VALIGN=TOP NOWRAP>Spireaea</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-3'>3</A> <A HREF='#KEY-1-2'>(2)</A></td> <td WIDTH=400 VALIGN=TOP>Plants herbaceous without stipules, carpels free, disc slightly divided into lobes.</td> <td > </td> <td VALIGN=TOP NOWRAP>Aruncus</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Plants with woody branches, erect or prostrate with stipules, carpels connate, hypanthium closed at the mouth by the entire disc.</td> <td > </td> <td VALIGN=TOP NOWRAP>Sorbaria</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-4'>4</A> <A HREF='#KEY-1-1'>(1)</A></td> <td WIDTH=400 VALIGN=TOP>Herbs, shrubs or sarmentous shrubs. Carpels 1 to many and free. Fruit an achenetum (with drupaceous achenes in <i>Kerria</i>) , drupetum, pometum or glandetum.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-5'>(5)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Trees or shrubs. Carpels 1 or 3-5 and fused. Fruit a drupe, nuculanium or pome.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-18'>(18)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-5'>5</A> <A HREF='#KEY-1-4'>(4)</A></td> <td WIDTH=400 VALIGN=TOP>Epicalyx present.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-6'>(6)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Epicalyx absent.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-12'>(12)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-6'>6</A> <A HREF='#KEY-1-5'>(5)</A></td> <td WIDTH=400 VALIGN=TOP>Petals absent, hypanthium campanulate, almost closed by the disk at the mouth. Fruit a pometum.</td> <td > </td> <td VALIGN=TOP NOWRAP>Alchemilla</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Petals present, hypanthium saucer-shaped to obconic, not closed. Fruit an achenetum or glandetum.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-7'>(7)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-7'>7</A> <A HREF='#KEY-1-6'>(6)</A></td> <td WIDTH=400 VALIGN=TOP>Stylodia sub-terminal, lateral or sub-basal, leaves trifoliolate to imparipinnate.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-8'>(8)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Stylodia terminal, leaves imparipinnate.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-11'>(11)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-8'>8</A> <A HREF='#KEY-1-7'>(7)</A></td> <td WIDTH=400 VALIGN=TOP>Receptacle fleshy in fruit, enclosing the seeds (fruit a glandetum). Seeds superficial to prominent.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-9'>(9)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Receptacle not fleshy in fruit, fruit an achenetum. Seeds sunken in the receptacle.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-10'>(10)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-9'>9</A> <A HREF='#KEY-1-8'>(8)</A></td> <td WIDTH=400 VALIGN=TOP>Fruit tasteless, petals yellow, epicalyx broader than the sepals. Seeds prominent, red.</td> <td > </td> <td VALIGN=TOP NOWRAP>2 <A HREF='florataxon.aspx?flora_id=5&taxon_id=111019' TARGET='_top' >Duchesnea</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Fruit tasty, petals white, epicalyx narrower than the sepals. Seeds superficial, brown.</td> <td > </td> <td VALIGN=TOP NOWRAP>3 <A HREF='florataxon.aspx?flora_id=5&taxon_id=112975' TARGET='_top' >Fragaria</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-10'>10</A> <A HREF='#KEY-1-8'>(8)</A></td> <td WIDTH=400 VALIGN=TOP>Stamens 10 or more, carpels numerous.</td> <td > </td> <td VALIGN=TOP NOWRAP>4 <A HREF='florataxon.aspx?flora_id=5&taxon_id=126635' TARGET='_top' >Potentilla</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Stamens 4-5 (10), carpels 5-10 (12).</td> <td > </td> <td VALIGN=TOP NOWRAP>5 <A HREF='florataxon.aspx?flora_id=5&taxon_id=130287' TARGET='_top' >Sibbaldia</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-11'>11</A> <A HREF='#KEY-1-11'>(11)</A></td> <td WIDTH=400 VALIGN=TOP>Stylodia with caducous upper part, lower part persistent and hooked at the apex.</td> <td > </td> <td VALIGN=TOP NOWRAP>Geum</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Stylodia not hooked at apex, without caducous upper part.</td> <td > </td> <td VALIGN=TOP NOWRAP>Acomastylis</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-12'>12</A> <A HREF='#KEY-1-5'>(5)</A></td> <td WIDTH=400 VALIGN=TOP>Herbs.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-13'>(13)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Erect or sarmentous shrubs.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-16'>(16)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-13'>13</A> <A HREF='#KEY-1-12'>(12)</A></td> <td WIDTH=400 VALIGN=TOP>Hypanthium closed at the mouth, fruit a pometum.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-14'>(14)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Hypanthium open, fruit an achenetum.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-15'>(15)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-14'>14</A> <A HREF='#KEY-1-13'>(13)</A></td> <td WIDTH=400 VALIGN=TOP>Hypanthium armed with hooks, petals present, stigma capitate. Inflorescence a raceme. Leaves interruptedly imparipinnate.</td> <td > </td> <td VALIGN=TOP NOWRAP>Agrimonia</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Hypanthium unarmed, petals absent, stigma penicillate. Inflorescence in globular heads. Leaves imparipinnate.</td> <td > </td> <td VALIGN=TOP NOWRAP>Sanguisorba</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-15'>15</A> <A HREF='#KEY-1-13'>(13)</A></td> <td WIDTH=400 VALIGN=TOP>Stamens 5, stylodia lateral, 1 ovule per carpel. Leaves 1-3 times tripartite</td> <td > </td> <td VALIGN=TOP NOWRAP>1 <A HREF='florataxon.aspx?flora_id=5&taxon_id=106465' TARGET='_top' >Chamaerhodos</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Stamens 20-40, stylodia terminal, 2 ovules per carpel. Leaves interruptedly imparipinnate.</td> <td > </td> <td VALIGN=TOP NOWRAP>Filipendula</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-16'>16</A> <A HREF='#KEY-1-12'>(12)</A></td> <td WIDTH=400 VALIGN=TOP>Sarmentous shrubs, usually prickly. Leaves imparipinnate or palmately to pinnately compound. Fruit a pometum or drupetum with numerous achenes/ druplets.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-17'>(17)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Erect shrub, leaves simple, fruit an achenetum with 4-5 drupaceous achenes.</td> <td > </td> <td VALIGN=TOP NOWRAP>Kerria</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-17'>17</A> <A HREF='#KEY-1-16'>(16)</A></td> <td WIDTH=400 VALIGN=TOP>Hypanthium closed at the mouth, fruit a pometum.</td> <td > </td> <td VALIGN=TOP NOWRAP>16 <A HREF='florataxon.aspx?flora_id=5&taxon_id=128746' TARGET='_top' >Rosa</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Hypanthium open, fruit a drupetum.</td> <td > </td> <td VALIGN=TOP NOWRAP>Rubus</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-18'>18</A> <A HREF='#KEY-1-4'>(4)</A></td> <td WIDTH=400 VALIGN=TOP>Carpel 1, ovary superior, fruit a drupe or nuculanium.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-19'>(19)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Carpels 3-5, connate, ovary inferior, fruit a pome.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-20'>(20)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-19'>19</A> <A HREF='#KEY-1-18'>(18)</A></td> <td WIDTH=400 VALIGN=TOP>Style terminal, hypanthium cupular, stipules usually conspicuous or deciduous.</td> <td > </td> <td VALIGN=TOP NOWRAP>Prunus</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Style becoming sub-basal, hypanthium recurved, stipules minute.</td> <td > </td> <td VALIGN=TOP NOWRAP>Prinsepia</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-20'>20</A> <A HREF='#KEY-1-18'>(18)</A></td> <td WIDTH=400 VALIGN=TOP>Fruit a pome, mature fruit with stony endocarp.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-21'>(21)</A></td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Fruit a pome, mature fruit without a stony endocarp.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-23'>(23)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-21'>21</A> <A HREF='#KEY-1-20'>(20)</A></td> <td WIDTH=400 VALIGN=TOP>Leaves entire, plants without spines.</td> <td > </td> <td VALIGN=TOP NOWRAP>Cotoneaster</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Leaves crenate or lobed, plants with spines.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-22'>(22)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-22'>22</A> <A HREF='#KEY-1-21'>(21)</A></td> <td WIDTH=400 VALIGN=TOP>Evergreen shrub, stipules caducous, leaves simple.</td> <td > </td> <td VALIGN=TOP NOWRAP>Pyracantha</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Deciduous shrub, stipules persistent, leaves pinnately lobed.</td> <td > </td> <td VALIGN=TOP NOWRAP>Crataegus</td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-23'>23</A> <A HREF='#KEY-1-20'>(20)</A></td> <td WIDTH=400 VALIGN=TOP>Leaves imparipinnate.</td> <td > </td> <td VALIGN=TOP NOWRAP>Sorbus subgenus Aucuparia</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Leaves simple or lobed.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-24'>(24)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-24'>24</A> <A HREF='#KEY-1-23'>(23)</A></td> <td WIDTH=400 VALIGN=TOP>Flowers solitary, ovary with many ovules per locule.</td> <td > </td> <td VALIGN=TOP NOWRAP>Cydonia</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Flowers in axillary or terminal racemes, ovary with 2 ovules per locule.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-25'>(25)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-25'>25</A> <A HREF='#KEY-1-24'>(24)</A></td> <td WIDTH=400 VALIGN=TOP>Evergreen tree, inflorescence a compound raceme.</td> <td > </td> <td VALIGN=TOP NOWRAP>Eriobotrya</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Deciduous trees, inflorescence raceme or compound raceme.</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-26'>(26)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-26'>26</A> <A HREF='#KEY-1-25'>(25)</A></td> <td WIDTH=400 VALIGN=TOP>Leaves with secondary veins going straight to the margin. Inflorescence a compound raceme (not observed at fruiting stage).</td> <td > </td> <td VALIGN=TOP NOWRAP>Sorbus subgenus Aria</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Leaves with secondary veins not going straight to the margins. Inflorescence a simple raceme usually on a brachyblast (short branch).</td> <td > </td> <td VALIGN=TOP NOWRAP><A HREF='#KEY-1-27'>(27)</A></td> </TR> <TR > <td > </td> <td > </td> <td > </td> <td > </td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP><A NAME='KEY-1-27'>27</A> <A HREF='#KEY-1-26'>(26)</A></td> <td WIDTH=400 VALIGN=TOP>Styles fused at the base, pome without sclereids.</td> <td > </td> <td VALIGN=TOP NOWRAP>Malus</td> </TR> <TR > <td WIDTH=50 NOWRAP VALIGN=TOP>+</td> <td WIDTH=400 VALIGN=TOP>Styles free at the base emerging through a hole in the middle of the disc. Pome with sclereids, pyriform to globose.</td> <td > </td> <td VALIGN=TOP NOWRAP>Pyrus</td> </TR> </TABLE> </span></P> </TD> </tr> <tr bgcolor="#d5ffff"> <TD colspan="2"> <span id="lblKeyList" style="font-size:9pt;"></span></TD> </tr> </table> </div></P> <P> <span id="lblLowerTaxonList"><p> <B>Lower Taxa</B><br> <ul> <li><A HREF='florataxon.aspx?flora_id=5&taxon_id=106465' TITLE='Accepted Name' ><b>Chamaerhodos</b> Bunge</A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=111019' TITLE='Accepted Name' ><b>Duchesnea</b> Sm.</A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=112975' TITLE='Accepted Name' ><b>Fragaria</b> L.</A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=126635' TITLE='Accepted Name' ><b>Potentilla</b> L.</A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=128746' TITLE='Accepted Name' ><b>Rosa</b> L.</A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=250096892' TITLE='Accepted Name' ><b>Rosaceae</b> Tribe Potentilleae </A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=250096871' TITLE='Accepted Name' ><b>Rosaceae</b> Tribe Roseae </A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=130287' TITLE='Accepted Name' ><b>Sibbaldia</b> L.</A></ul> <p> <p> <B>Related Synonym(s):</B><br> <ul> <li><A HREF='florataxon.aspx?flora_id=5&taxon_id=250096058' TITLE='Synonym' ><i>Potentilla</i> <i>sect.</i> Duchesnea (Sm.) Dikshit & Panigrahi = <A HREF='florataxon.aspx?flora_id=5&taxon_id=111019' TITLE='Accepted Name' ><b>Duchesnea</b> Sm.</A></A><li><A HREF='florataxon.aspx?flora_id=5&taxon_id=250096059' TITLE='Synonym' ><i>Potentilla</i> <i>subgen.</i> <i>Duchesnea</i> (Sm.) M. Shah & Wilcock = <A HREF='florataxon.aspx?flora_id=5&taxon_id=111019' TITLE='Accepted Name' ><b>Duchesnea</b> Sm.</A></A></ul> <p> </span></P> <BR clear="all"> <P> </P> <table id="tableObjectList" cellspacing="0" cellpadding="4" width="100%" border="1"> <tr bgcolor="#ffffcc"> <TD> <TABLE cellSpacing="0" cellPadding="0" width="100%" border="0"> <TR> <TD> <span id="Label1" style="font-size:9pt;"> <b>Related Objects</b> </span></TD> <TD align="right"> <span id="lblGalleryLink" style="font-size:8pt;"></span> </TD> </TR> </TABLE> </TD> </tr> <tr> <TD> <span id="lblObjectList"><p> <TABLE WIDTH='100%' BORDER='0' cellspacing=0> <TR > <td class='bold'>Chinese Plant Names</td> <td > </td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=47266&flora_id=3' TITLE='Image/JPEG' >Hesperomeles (Trees and shrubs of the Andes of Ecuador)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=47369&flora_id=3' TITLE='Image/JPEG' >Rubus (Trees and shrubs of the Andes of Ecuador)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td class='bold'>Flora of Chile</td> <td > </td> </TR> <TR > <td width=400><li><A HREF='https://images.mobot.org/efloras/FloraData/060/PDF/V02/vol2-36-Rosaceae.pdf' TITLE='PDF' >Family PDF</A></td> <td class='small'>PDF</td> </TR> <TR > <td class='bold'>Flora of China</td> <td > </td> </TR> <TR > <td width=400><li><A HREF='http://flora.huh.harvard.edu/china/mss/volume09/Rosaceae.PDF' TITLE='PDF' >PDF</A></td> <td class='small'>PDF</td> </TR> <TR > <td class='bold'>Interactive Keys by Xiangying Wen</td> <td > </td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=47880&flora_id=1001' TITLE='Image/JPEG' >Chaenomeles sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=47134&flora_id=1001' TITLE='Image/JPEG' >Hesperomeles (Trees and shrubs of the Andes of Ecuador)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=47724&flora_id=1001' TITLE='Image/JPEG' >Malus sargentii (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=47725&flora_id=1001' TITLE='Image/JPEG' >Malus sargentii (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48173&flora_id=1001' TITLE='Image/JPEG' >Malus sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48174&flora_id=1001' TITLE='Image/JPEG' >Pyrus sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48175&flora_id=1001' TITLE='Image/JPEG' >Rosa sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48176&flora_id=1001' TITLE='Image/JPEG' >Rosa sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48177&flora_id=1001' TITLE='Image/JPEG' >Rosa sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48178&flora_id=1001' TITLE='Image/JPEG' >Rosaceae sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48179&flora_id=1001' TITLE='Image/JPEG' >Rosaceae sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48180&flora_id=1001' TITLE='Image/JPEG' >Rosaceae sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48181&flora_id=1001' TITLE='Image/JPEG' >Rosaceae sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48182&flora_id=1001' TITLE='Image/JPEG' >Rosaceae sp. (Hong Song)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48183&flora_id=1001' TITLE='Image/JPEG' >Rosaceae sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=48172&flora_id=1001' TITLE='Image/JPEG' >Rosaceae sp. (Xiangying Wen)</A></td> <td class='small'>Image/JPEG</td> </TR> <TR > <td width=400><li><A HREF='object_page.aspx?object_id=47237&flora_id=1001' TITLE='Image/JPEG' >Rubus (Trees and shrubs of the Andes of Ecuador)</A></td> <td class='small'>Image/JPEG</td> </TR> </TABLE> </span></TD> </tr> </table> <P></P> </div> <P><div id="panelTaxonLinks"> </P> <P> <table id="tableLinkList" cellspacing="0" cellpadding="4" width="100%" border="1"> <tr bgcolor="#ffffcc"> <TD><span id="lblLinkTitle" style="font-size:9pt;"><b>Related Links</b> (opens in a new window)</span></TD> </tr> <tr> <TD> <P><span id="lblLinkList"><b>Treatments in Other Floras @ www.efloras.org</b><ul> <li><A HREF='florataxon.aspx?flora_id=110&taxon_id=10776' TARGET='_top' TITLE='treatment in Annotated Checklist of the Flowering Plants of Nepal' >Annotated Checklist of the Flowering Plants of Nepal</A><li><A HREF='florataxon.aspx?flora_id=3&taxon_id=10776' TARGET='_top' TITLE='treatment in Chinese Plant Names' >Chinese Plant Names</A><li><A HREF='florataxon.aspx?flora_id=60&taxon_id=10776' TARGET='_top' TITLE='treatment in Flora of Chile' >Flora of Chile</A><li><A HREF='florataxon.aspx?flora_id=2&taxon_id=10776' TARGET='_top' TITLE='treatment in Flora of China' >Flora of China</A><li><A HREF='florataxon.aspx?flora_id=11&taxon_id=10776' TARGET='_top' TITLE='treatment in Flora of Missouri' >Flora of Missouri</A><li><A HREF='florataxon.aspx?flora_id=1&taxon_id=10776' TARGET='_top' TITLE='treatment in Flora of North America' >Flora of North America</A><li><A HREF='florataxon.aspx?flora_id=101&taxon_id=10776' TARGET='_top' TITLE='treatment in Flora of Taiwan Checklist' >Flora of Taiwan Checklist</A><li><A HREF='florataxon.aspx?flora_id=1130&taxon_id=10776' TARGET='_top' TITLE='treatment in Interactive Keys by Anthony Brach' >Interactive Keys by Anthony Brach</A><li><A HREF='florataxon.aspx?flora_id=1001&taxon_id=10776' TARGET='_top' TITLE='treatment in Interactive Keys by Xiangying Wen' >Interactive Keys by Xiangying Wen</A><li><A HREF='florataxon.aspx?flora_id=120&taxon_id=10776' TARGET='_top' TITLE='treatment in Ornamental Plants From Russia And Adjacent States Of The Former Soviet Union' >Ornamental Plants From Russia And Adjacent States Of The Former Soviet Union</A><li><A HREF='florataxon.aspx?flora_id=201&taxon_id=10776' TARGET='_top' TITLE='treatment in Trees and shrubs of the Andes of Ecuador' >Trees and shrubs of the Andes of Ecuador</A></ul> <p> <b>Other Databases</b> <ul> <li><A HREF='http://mobot.mobot.org/cgi-bin/search_pick?name=Rosaceae' TARGET='FLORA_LINK' TITLE='W3TROPICOS' >W<SUP>3</SUP>TROPICOS</A> <li><A HREF='http://www.ipni.org/ipni/IpniServlet?family=Rosaceae&infrafamily=&genus=&infragenus=&species=&infraspecies=&author_abbrev=&publication_title=&is_apni_record=on&is_gci_record=on&is_ik_record=on&include_authors=on&include_basionym_authors=on&query_type=by_query' TARGET='FLORA_LINK' TITLE='IPNI' >IPNI</A></ul> </span></P> </TD> </tr> </table> </div></P> </td> </tr> <tr> <td> <!-- Links --> <P></P> <P><span id="lblEditLinks" style="font-size:9pt;"></span></P> <P> <!-- Comment List --></P> </td> </tr> </table> </td> <TD id="tdRightBar" valign="top"></TD> </tr> </table> </TD> <tr> <td> <table id="SubHeaderTableOut" width="100%" border="0" cellpadding="0" cellspacing="0"> <tr> <td bgcolor="#999999"><img src="~/images/layout/dot.gif" alt="" width="100%" height="1"></td> </tr> <tr> <TD> <table bgcolor="#99cc99" id="SubHeaderTable" width="100%" border="0" cellpadding="2" cellspacing="0"> <tr> <TD align="center" bgColor="#ffffcc"> | <a id="ucEfloraFooter_linkHome" href="index.aspx" target="_top">eFlora Home</a> | <a id="ucEfloraFooter_linkPeopleSearch" href="person_search.aspx" target="_top">People Search</a> | <a id="ucEfloraFooter_linkHelp" href="help_page.aspx" target="_top">Help</a> | <a id="ucEfloraFooter_linkActKey" href="http://flora.huh.harvard.edu/china/ActKey" target="_top">ActKey</a> | <a id="ucEfloraFooter_Hyperlink2" href="http://flora.huh.harvard.edu/HuCards/" target="_top">Hu Cards</a> | <a id="ucEfloraFooter_Hyperlink1" href="glossary.aspx" target="_top">Glossary</a> | </TD> </tr> </table> </TD> </tr> <tr> <td bgcolor="#999999"><img src="~/images/layout/dot.gif" alt="" width="100%" height="1"></td> </tr> </table> <p></p> </td> </tr> </TABLE> </body> </HTML>